pigmentation photos on Flickr

Indian Corn Close Up. It is thought that we named this colorful corn after the indigenous people that first introduced us to corn. This corn is now grown purely for its interesting pigmentation and rarely eaten. The dramatic and varied pigmentation we see by David Williams

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via Instagram ift.tt/2dLDKqO

BJ2B2435 Passage of time , by Jan Zoltonos

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Baby boy 10 month and Great-Grand-Father 87 years , his old skin is age relating skin pigmentation , they are a family of four generation .

Archaeological Museum, Heraklion, Crete ~ Minoan Art ~ Ladies in Blue by Nieuw Album

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Heads and upper torsos of three women with long tresses, headbands, flounced dresses of open bodice. Blue is the predominant color.

From: en.wikipedia.org/wiki/List_of_Aegean_frescos

Overview of rooms in the museum:

The museum, as it is now, is completely renewed. Old information about rooms and their numbers are still not updated, not even in Wikipedia. They mention twenty rooms, and their names, but there are 27 rooms, XXVII

Maybe wiki will update their page soon, as it is in October 2015, it is not updated.

en.wikipedia.org/wiki/Heraklion_Archaeological_Museum

~

This serial of photos offers an impression of details of the collection with Minoan Art, belonging to the Palace of Knossos. the Archaeological Museum in Heraklion has been completely renewed. Lots of amazing information can be read next to art objects.

It is not allowed to use flash when making photos. This, and the many visitors made it hardly possible to make really sharp photos in some seconds.

My camera is a rather cheap one and creates curved lines.

Some photos are not sharp, but I kept them anyway.

Altogether it has been a deeply impressing visit, moving, deeply moving because of the mystical, spiritual, mental and emotional depth of the Art.

The Minoans were utterly creative.

Their art is comparable with our modern art. Their use of colors makes the art characteristic: pastel colors (modest in pigmentation), terra colors, with blue, green and ochre.

Often I edited the photos in several ways. Or cropped them, to attract the attention for details.

Enjoy the collection of photos. If you want to read more about the Minoans:

www.heraklion-crete.org/archaeological-museum/

Leucistic Dunnock - Prunella Modularis in Killamarsh Derbyshire 2022. by Alan Weaver

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These are brown. This is a very rare albino or leucistic, lacking pigmentation, it stayed around for 3 days.

Leucistic Grey Squirrel by Stephen Stephen

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We have lots of Grey and Black Phase Grey Squirrels in Ottawa but I've never seen a white (Leucistic) one until I photographed this one in Parc Lafontaine in Montreal this past June.

Leucistic animals are not Albinos but rather the condition is characterised by reduced skin pigmentation of all types not just melanin.

I would have liked to have gotten a few closeups of this squirrel but I only had my Sigma 17-70mm lens with me.

lumière d'automne by Patrick Marioné

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Bruxelles

mobile pigmentation by julie ames

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I oddly find this picture rather attractive.

British White by Chris, Norfolk

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This animal lives at Jimmy's Farm at Wherstead in Suffolk.

The British White is a breed of medium sized cattle, with cows typically weighing between 1,210 and 1,545 lb. (550 and 700 kg), and bulls weighing in at 1,985 to 2,425 lb. (900 to 1100 kg). They have a predominantly white coat, complemented by black ,or red, points on their muzzle, ears and feet.

Known for their docile nature, the British White is polled, meaning they naturally lack horns, making them easy to handle and manage.

Cows of this breed have long lifespans, 20 years plus, and exhibit high fertility, continuing to produce calves well into their teenage years. The breed is known for easy calving, displaying good maternal instincts, and giving birth to robust calves.

British Whites are adept at withstanding winter conditions, making them suitable for out-wintering in various environments. They are also known for their heat tolerance, with some cattle within the breed have white hair overlying dark skin pigmentation which increases their tolerance of hot climates

Legend has it that the breed has ties to Whalley Abbey in Lancashire. During the 17th. century, the Assheton family, who acquired the Abbey following the dissolution of the monasteries, possessed polled white cattle. Through marriage and subsequent breeding, these cattle gradually spread from the Whalley Abbey region to East Anglia, which became the primary center for the breed from the late 1700's until the 1960's. While there are still significant herds in East Anglia, the British White breed has now expanded across the entire country and is growing in popularity.

The Rare Breeds Survival Trust rates the British White as 'At Risk'.

Pink Meadow Grasshopper by JTW Aviation Images

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Due to a pigmentation called "erythrism", some Meadow Grasshoppers appear pink, and apparently the chances of you seeing one are 1%, so these are the 1%!

Photos taken outside RAF Valley, Anglesey, on Monday 5th September 2022. ©JTW Aviation Images.

Farmers and Merchants Union Bank, James Street and Dickason Boulevard, Columbus, WI by Warren LeMay

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Built in 1919-1920, this Chicago School and Sullivanesque-style building was designed by Louis Sullivan for the Farmers and Merchants Union Bank in Columbus, Wisconsin as one of his late-career “jewel box” bank buildings that are largely located in smaller communities throughout the midwest. The building was the last “jewel box bank” designed by Sullivan, and the second-to-last commission of his career, and was intended to communicate the bank as a modern and progressive institution, rather than employing the stodgier and more traditional Classical design found on most other banks of the era. The bank was commissioned by the president of the bank, J. Russell Wheeler, whose wife, Anna May Wheeler, pushed him to commission Sullivan to design a new home for the bank. In addition to Louis Sullivan, the building’s stained glass windows, were designed by architectural decorator Louis J. Millet, and the terra cotta by clay modeler Kristian Schneider, whom developed moulds for the building’s terra cotta, metal, and plaster details. The two artisans worked alongside Sullivan on several other bank projects. The building was heavily documented in Sullivan’s 1924 “A System of Architectural Ornament”, published shortly before he died.

The building is clad in red tapestry brick, which features blue and green mixed with the red clay mixture in some bricks, creating variation in color and texture across the facade. The brick creates a backdrop to some of the best terra cotta on any of Sullivan’s projects. The terra cotta features many of the floral and geometric motifs found on Sullivan’s other works, and is arranged similarly to other Sullivan banks that utilized brick cladding. The building features two principal facades, with a narrower facade along James Street, and a broader facade facing Dickason Boulevard. The James Street facade features two openings close to ground level, with the eastern bay housing a large plate glass window, and the western bay housing a doorway flanked by skylights, both of which are recessed under a large terra cotta architrave and flanked by square pilasters with decorative Sullivanesque ornament panels at the capitals. The architrave above the doorway and window is divided into three segments by vertical terra cotta elements that feature floral motifs and, like many Sullivan buildings, appear like plants with roots, branches, and crowns. The outer panels of the architrave feature circular cartouches with hexagonal trim, leaves, and geometric elements, with circular central medallions featuring the years 1861, when the bank was founded, and 1919, when the bank was completed. The central panel is clad in marble with the words “Farmers & Merchants Union Bank” and “Louis Sullivan, Architect” engraved into the stone with yellow pigmentation, contrasting against the white and green marble background. Atop the two vertical elements on either side of the central panel are griffin sculptures holding shields, a common element on many of Sullivan’s “Jewel Box Banks,” while the base of the outer vertical elements features the initials of the bank at the base. Above the architrave is an arched bay that houses a stained glass window, trimmed with decorative terra cotta at the inner and outer rings of the arch, with the bay becoming more recessed after each concentric arch, much like the entrances to medieval Romanesque churches. Besides a band of belt coursing that runs on either side of the architrave and wraps the corner to a tapered buttress on the Dickason Boulevard facade, the only other adornment is an eagle sculpture on a vertical trim element at the center of the parapet, which terminates many brick courses above the arched opening below, and another band of terra cotta trim along the top of the parapet, which forms a cap on the parapet around the perimeter of the building’s low-slope roof. On the Dickason Boulevard facade, the building features five recessed clerestory arched bays housing stained glass windows, flanked by tapered buttresses. Surrounding the arched tops of the windows are decorative trim panels with floral motifs, which begin just below the base of the arches, and extend up above the top of the arches, terminating in a band of belt coursing. Atop the buttresses at either end are trim elements featuring large spheres atop rectilinear legs with floral motifs below, undulating in and out with the brick below. Additionally, a band of belt coursing, which wraps the corner of Dickson Boulevard and James Street, runs beneath the windows, only interrupted by the buttresses. Toward the back, on the building’s original rear wing, there are three windows at eye level in the original building, with bands of belt coursing below and at the top of the parapet. The rear window is a recessed bay window flanked by two pilasters with sullivanesque terra cotta panels, while the smaller windows are flanked by sullivanesque relief panels. The rear wing features a roof at multiple heights, and was extended in 1961 with a matching addition by Law, Potter and Nystrom, since removed. The rear of the taller portion of the building features a simple recessed bay with an arched window, and a similar eagle sculpture and vertical trim piece as on the front facade.

Inside, the front wing of the building features a tall banking hall with brick cladding on the walls up to the level of the windows, where it terminates at a wooden sill. The space is split down the middle by a row of brick piers and low walls framing the teller cages, which terminate at the sill line of the windows, dividing the space while still allowing it to read as a single continuous lofty space. The brick forms piers at the teller’s cages, pilasters separating desks on the exterior wall, and low brick walls with marble caps. The upper portion of the walls and the coffered ceiling in this space is finished with white plaster, which gives the space a very vertical and airy feeling, as do the cream-colored terrazzo floors, which feature black edges at the base of the walls, tying the space together. The space features a terra cotta water fountain, or bubbler, also designed by sullivan, which features intricate ornament by Schneider. The space also features two mezzanine balconies with metal railings that run below the arched windows at the front and rear of the space, allowing managers to observe the activities in the lobby and teller area below from the rear balcony, while the front balcony exists solely to balance the space and keep it symmetrical. An office for private conferences with customers was originally located near the front of the space, along with a manager’s office, allowing convenience for customers seeking a meeting with the bank management. The teller’s side of the space also housed the bank’s two vaults and several other private offices. The bank originally featured a large meeting room in the one-story rear wing, behind the vaults, with a women’s waiting room sitting along the Dickason Boulevard side of the rear wing, featuring a bay window and a restroom. The building’s interior has changed in function somewhat due to the growth of the bank, changes in bank operations, and expansion of the building with new additions to house offices and a drive-through in the rear.

The building was listed on the National Register of Historic Places in 1972, was designated a National Historic Landmark in 1976, and is a contributing structure in the Columbus Downtown Historic District, listed on the National Register of Historic Places in 1992. The building saw an addition in 2006, clad in buff brick, which replicated a historic building that formerly stood to the east, and wraps the building to the rear, with a two-story section behind a one-story annex that connects the one-story rear wing of the bank to the new building. This wing replaced older additions made in 1961, which matched the one-story rear wing of the historic building, and 1980, which was modern in appearance and slightly recessed along James Street to give precedence to the historic building. The building still functions as the main office branch of the Farmers and Merchants Union Bank, which has grown substantially. The building has been long considered to be among the best of Sullivan’s “Jewel Box Banks,” and has been kept in excellent condition by the bank’s careful and caring generational stewardship.

Blow by Patrick Marioné

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Brussels

The Vestal Moth (Rhodometra sacraria) © by Nick Dobbs

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The Vestal Moth (Rhodometra sacraria) is a migrant species, mainly occurring in southern England in varying numbers. In good immigration years, several hundred may appear, and some may turn up almost anywhere in Britain.

The amount and intensity of the pink pigmentation is somewhat variable, as can be seen from the photographs.

The species breeds in southern Europe and North Africa, and the caterpillars live on knotgrass (Polygonum), dock (Rumex) and other low plants.

It flies mainly at night and is regularly attracted to light. Photo by Nick Dobbs, Malta 21-08-2023

Historical Marker, Farmers and Merchants Union Bank, James Street and Dickason Boulevard, Columbus, WI by Warren LeMay

1

Built in 1919-1920, this Chicago School and Sullivanesque-style building was designed by Louis Sullivan for the Farmers and Merchants Union Bank in Columbus, Wisconsin as one of his late-career “jewel box” bank buildings that are largely located in smaller communities throughout the midwest. The building was the last “jewel box bank” designed by Sullivan, and the second-to-last commission of his career, and was intended to communicate the bank as a modern and progressive institution, rather than employing the stodgier and more traditional Classical design found on most other banks of the era. The bank was commissioned by the president of the bank, J. Russell Wheeler, whose wife, Anna May Wheeler, pushed him to commission Sullivan to design a new home for the bank. In addition to Louis Sullivan, the building’s stained glass windows, were designed by architectural decorator Louis J. Millet, and the terra cotta by clay modeler Kristian Schneider, whom developed moulds for the building’s terra cotta, metal, and plaster details. The two artisans worked alongside Sullivan on several other bank projects. The building was heavily documented in Sullivan’s 1924 “A System of Architectural Ornament”, published shortly before he died.

The building is clad in red tapestry brick, which features blue and green mixed with the red clay mixture in some bricks, creating variation in color and texture across the facade. The brick creates a backdrop to some of the best terra cotta on any of Sullivan’s projects. The terra cotta features many of the floral and geometric motifs found on Sullivan’s other works, and is arranged similarly to other Sullivan banks that utilized brick cladding. The building features two principal facades, with a narrower facade along James Street, and a broader facade facing Dickason Boulevard. The James Street facade features two openings close to ground level, with the eastern bay housing a large plate glass window, and the western bay housing a doorway flanked by skylights, both of which are recessed under a large terra cotta architrave and flanked by square pilasters with decorative Sullivanesque ornament panels at the capitals. The architrave above the doorway and window is divided into three segments by vertical terra cotta elements that feature floral motifs and, like many Sullivan buildings, appear like plants with roots, branches, and crowns. The outer panels of the architrave feature circular cartouches with hexagonal trim, leaves, and geometric elements, with circular central medallions featuring the years 1861, when the bank was founded, and 1919, when the bank was completed. The central panel is clad in marble with the words “Farmers & Merchants Union Bank” and “Louis Sullivan, Architect” engraved into the stone with yellow pigmentation, contrasting against the white and green marble background. Atop the two vertical elements on either side of the central panel are griffin sculptures holding shields, a common element on many of Sullivan’s “Jewel Box Banks,” while the base of the outer vertical elements features the initials of the bank at the base. Above the architrave is an arched bay that houses a stained glass window, trimmed with decorative terra cotta at the inner and outer rings of the arch, with the bay becoming more recessed after each concentric arch, much like the entrances to medieval Romanesque churches. Besides a band of belt coursing that runs on either side of the architrave and wraps the corner to a tapered buttress on the Dickason Boulevard facade, the only other adornment is an eagle sculpture on a vertical trim element at the center of the parapet, which terminates many brick courses above the arched opening below, and another band of terra cotta trim along the top of the parapet, which forms a cap on the parapet around the perimeter of the building’s low-slope roof. On the Dickason Boulevard facade, the building features five recessed clerestory arched bays housing stained glass windows, flanked by tapered buttresses. Surrounding the arched tops of the windows are decorative trim panels with floral motifs, which begin just below the base of the arches, and extend up above the top of the arches, terminating in a band of belt coursing. Atop the buttresses at either end are trim elements featuring large spheres atop rectilinear legs with floral motifs below, undulating in and out with the brick below. Additionally, a band of belt coursing, which wraps the corner of Dickson Boulevard and James Street, runs beneath the windows, only interrupted by the buttresses. Toward the back, on the building’s original rear wing, there are three windows at eye level in the original building, with bands of belt coursing below and at the top of the parapet. The rear window is a recessed bay window flanked by two pilasters with sullivanesque terra cotta panels, while the smaller windows are flanked by sullivanesque relief panels. The rear wing features a roof at multiple heights, and was extended in 1961 with a matching addition by Law, Potter and Nystrom, since removed. The rear of the taller portion of the building features a simple recessed bay with an arched window, and a similar eagle sculpture and vertical trim piece as on the front facade.

Inside, the front wing of the building features a tall banking hall with brick cladding on the walls up to the level of the windows, where it terminates at a wooden sill. The space is split down the middle by a row of brick piers and low walls framing the teller cages, which terminate at the sill line of the windows, dividing the space while still allowing it to read as a single continuous lofty space. The brick forms piers at the teller’s cages, pilasters separating desks on the exterior wall, and low brick walls with marble caps. The upper portion of the walls and the coffered ceiling in this space is finished with white plaster, which gives the space a very vertical and airy feeling, as do the cream-colored terrazzo floors, which feature black edges at the base of the walls, tying the space together. The space features a terra cotta water fountain, or bubbler, also designed by sullivan, which features intricate ornament by Schneider. The space also features two mezzanine balconies with metal railings that run below the arched windows at the front and rear of the space, allowing managers to observe the activities in the lobby and teller area below from the rear balcony, while the front balcony exists solely to balance the space and keep it symmetrical. An office for private conferences with customers was originally located near the front of the space, along with a manager’s office, allowing convenience for customers seeking a meeting with the bank management. The teller’s side of the space also housed the bank’s two vaults and several other private offices. The bank originally featured a large meeting room in the one-story rear wing, behind the vaults, with a women’s waiting room sitting along the Dickason Boulevard side of the rear wing, featuring a bay window and a restroom. The building’s interior has changed in function somewhat due to the growth of the bank, changes in bank operations, and expansion of the building with new additions to house offices and a drive-through in the rear.

The building was listed on the National Register of Historic Places in 1972, was designated a National Historic Landmark in 1976, and is a contributing structure in the Columbus Downtown Historic District, listed on the National Register of Historic Places in 1992. The building saw an addition in 2006, clad in buff brick, which replicated a historic building that formerly stood to the east, and wraps the building to the rear, with a two-story section behind a one-story annex that connects the one-story rear wing of the bank to the new building. This wing replaced older additions made in 1961, which matched the one-story rear wing of the historic building, and 1980, which was modern in appearance and slightly recessed along James Street to give precedence to the historic building. The building still functions as the main office branch of the Farmers and Merchants Union Bank, which has grown substantially. The building has been long considered to be among the best of Sullivan’s “Jewel Box Banks,” and has been kept in excellent condition by the bank’s careful and caring generational stewardship.

Farmers and Merchants Union Bank, James Street and Dickason Boulevard, Columbus, WI by Warren LeMay

2

Built in 1919-1920, this Chicago School and Sullivanesque-style building was designed by Louis Sullivan for the Farmers and Merchants Union Bank in Columbus, Wisconsin as one of his late-career “jewel box” bank buildings that are largely located in smaller communities throughout the midwest. The building was the last “jewel box bank” designed by Sullivan, and the second-to-last commission of his career, and was intended to communicate the bank as a modern and progressive institution, rather than employing the stodgier and more traditional Classical design found on most other banks of the era. The bank was commissioned by the president of the bank, J. Russell Wheeler, whose wife, Anna May Wheeler, pushed him to commission Sullivan to design a new home for the bank. In addition to Louis Sullivan, the building’s stained glass windows, were designed by architectural decorator Louis J. Millet, and the terra cotta by clay modeler Kristian Schneider, whom developed moulds for the building’s terra cotta, metal, and plaster details. The two artisans worked alongside Sullivan on several other bank projects. The building was heavily documented in Sullivan’s 1924 “A System of Architectural Ornament”, published shortly before he died.

The building is clad in red tapestry brick, which features blue and green mixed with the red clay mixture in some bricks, creating variation in color and texture across the facade. The brick creates a backdrop to some of the best terra cotta on any of Sullivan’s projects. The terra cotta features many of the floral and geometric motifs found on Sullivan’s other works, and is arranged similarly to other Sullivan banks that utilized brick cladding. The building features two principal facades, with a narrower facade along James Street, and a broader facade facing Dickason Boulevard. The James Street facade features two openings close to ground level, with the eastern bay housing a large plate glass window, and the western bay housing a doorway flanked by skylights, both of which are recessed under a large terra cotta architrave and flanked by square pilasters with decorative Sullivanesque ornament panels at the capitals. The architrave above the doorway and window is divided into three segments by vertical terra cotta elements that feature floral motifs and, like many Sullivan buildings, appear like plants with roots, branches, and crowns. The outer panels of the architrave feature circular cartouches with hexagonal trim, leaves, and geometric elements, with circular central medallions featuring the years 1861, when the bank was founded, and 1919, when the bank was completed. The central panel is clad in marble with the words “Farmers & Merchants Union Bank” and “Louis Sullivan, Architect” engraved into the stone with yellow pigmentation, contrasting against the white and green marble background. Atop the two vertical elements on either side of the central panel are griffin sculptures holding shields, a common element on many of Sullivan’s “Jewel Box Banks,” while the base of the outer vertical elements features the initials of the bank at the base. Above the architrave is an arched bay that houses a stained glass window, trimmed with decorative terra cotta at the inner and outer rings of the arch, with the bay becoming more recessed after each concentric arch, much like the entrances to medieval Romanesque churches. Besides a band of belt coursing that runs on either side of the architrave and wraps the corner to a tapered buttress on the Dickason Boulevard facade, the only other adornment is an eagle sculpture on a vertical trim element at the center of the parapet, which terminates many brick courses above the arched opening below, and another band of terra cotta trim along the top of the parapet, which forms a cap on the parapet around the perimeter of the building’s low-slope roof. On the Dickason Boulevard facade, the building features five recessed clerestory arched bays housing stained glass windows, flanked by tapered buttresses. Surrounding the arched tops of the windows are decorative trim panels with floral motifs, which begin just below the base of the arches, and extend up above the top of the arches, terminating in a band of belt coursing. Atop the buttresses at either end are trim elements featuring large spheres atop rectilinear legs with floral motifs below, undulating in and out with the brick below. Additionally, a band of belt coursing, which wraps the corner of Dickson Boulevard and James Street, runs beneath the windows, only interrupted by the buttresses. Toward the back, on the building’s original rear wing, there are three windows at eye level in the original building, with bands of belt coursing below and at the top of the parapet. The rear window is a recessed bay window flanked by two pilasters with sullivanesque terra cotta panels, while the smaller windows are flanked by sullivanesque relief panels. The rear wing features a roof at multiple heights, and was extended in 1961 with a matching addition by Law, Potter and Nystrom, since removed. The rear of the taller portion of the building features a simple recessed bay with an arched window, and a similar eagle sculpture and vertical trim piece as on the front facade.

Inside, the front wing of the building features a tall banking hall with brick cladding on the walls up to the level of the windows, where it terminates at a wooden sill. The space is split down the middle by a row of brick piers and low walls framing the teller cages, which terminate at the sill line of the windows, dividing the space while still allowing it to read as a single continuous lofty space. The brick forms piers at the teller’s cages, pilasters separating desks on the exterior wall, and low brick walls with marble caps. The upper portion of the walls and the coffered ceiling in this space is finished with white plaster, which gives the space a very vertical and airy feeling, as do the cream-colored terrazzo floors, which feature black edges at the base of the walls, tying the space together. The space features a terra cotta water fountain, or bubbler, also designed by sullivan, which features intricate ornament by Schneider. The space also features two mezzanine balconies with metal railings that run below the arched windows at the front and rear of the space, allowing managers to observe the activities in the lobby and teller area below from the rear balcony, while the front balcony exists solely to balance the space and keep it symmetrical. An office for private conferences with customers was originally located near the front of the space, along with a manager’s office, allowing convenience for customers seeking a meeting with the bank management. The teller’s side of the space also housed the bank’s two vaults and several other private offices. The bank originally featured a large meeting room in the one-story rear wing, behind the vaults, with a women’s waiting room sitting along the Dickason Boulevard side of the rear wing, featuring a bay window and a restroom. The building’s interior has changed in function somewhat due to the growth of the bank, changes in bank operations, and expansion of the building with new additions to house offices and a drive-through in the rear.

The building was listed on the National Register of Historic Places in 1972, was designated a National Historic Landmark in 1976, and is a contributing structure in the Columbus Downtown Historic District, listed on the National Register of Historic Places in 1992. The building saw an addition in 2006, clad in buff brick, which replicated a historic building that formerly stood to the east, and wraps the building to the rear, with a two-story section behind a one-story annex that connects the one-story rear wing of the bank to the new building. This wing replaced older additions made in 1961, which matched the one-story rear wing of the historic building, and 1980, which was modern in appearance and slightly recessed along James Street to give precedence to the historic building. The building still functions as the main office branch of the Farmers and Merchants Union Bank, which has grown substantially. The building has been long considered to be among the best of Sullivan’s “Jewel Box Banks,” and has been kept in excellent condition by the bank’s careful and caring generational stewardship.

Farmers and Merchants Union Bank, James Street and Dickason Boulevard, Columbus, WI by Warren LeMay

1

Built in 1919-1920, this Chicago School and Sullivanesque-style building was designed by Louis Sullivan for the Farmers and Merchants Union Bank in Columbus, Wisconsin as one of his late-career “jewel box” bank buildings that are largely located in smaller communities throughout the midwest. The building was the last “jewel box bank” designed by Sullivan, and the second-to-last commission of his career, and was intended to communicate the bank as a modern and progressive institution, rather than employing the stodgier and more traditional Classical design found on most other banks of the era. The bank was commissioned by the president of the bank, J. Russell Wheeler, whose wife, Anna May Wheeler, pushed him to commission Sullivan to design a new home for the bank. In addition to Louis Sullivan, the building’s stained glass windows, were designed by architectural decorator Louis J. Millet, and the terra cotta by clay modeler Kristian Schneider, whom developed moulds for the building’s terra cotta, metal, and plaster details. The two artisans worked alongside Sullivan on several other bank projects. The building was heavily documented in Sullivan’s 1924 “A System of Architectural Ornament”, published shortly before he died.

The building is clad in red tapestry brick, which features blue and green mixed with the red clay mixture in some bricks, creating variation in color and texture across the facade. The brick creates a backdrop to some of the best terra cotta on any of Sullivan’s projects. The terra cotta features many of the floral and geometric motifs found on Sullivan’s other works, and is arranged similarly to other Sullivan banks that utilized brick cladding. The building features two principal facades, with a narrower facade along James Street, and a broader facade facing Dickason Boulevard. The James Street facade features two openings close to ground level, with the eastern bay housing a large plate glass window, and the western bay housing a doorway flanked by skylights, both of which are recessed under a large terra cotta architrave and flanked by square pilasters with decorative Sullivanesque ornament panels at the capitals. The architrave above the doorway and window is divided into three segments by vertical terra cotta elements that feature floral motifs and, like many Sullivan buildings, appear like plants with roots, branches, and crowns. The outer panels of the architrave feature circular cartouches with hexagonal trim, leaves, and geometric elements, with circular central medallions featuring the years 1861, when the bank was founded, and 1919, when the bank was completed. The central panel is clad in marble with the words “Farmers & Merchants Union Bank” and “Louis Sullivan, Architect” engraved into the stone with yellow pigmentation, contrasting against the white and green marble background. Atop the two vertical elements on either side of the central panel are griffin sculptures holding shields, a common element on many of Sullivan’s “Jewel Box Banks,” while the base of the outer vertical elements features the initials of the bank at the base. Above the architrave is an arched bay that houses a stained glass window, trimmed with decorative terra cotta at the inner and outer rings of the arch, with the bay becoming more recessed after each concentric arch, much like the entrances to medieval Romanesque churches. Besides a band of belt coursing that runs on either side of the architrave and wraps the corner to a tapered buttress on the Dickason Boulevard facade, the only other adornment is an eagle sculpture on a vertical trim element at the center of the parapet, which terminates many brick courses above the arched opening below, and another band of terra cotta trim along the top of the parapet, which forms a cap on the parapet around the perimeter of the building’s low-slope roof. On the Dickason Boulevard facade, the building features five recessed clerestory arched bays housing stained glass windows, flanked by tapered buttresses. Surrounding the arched tops of the windows are decorative trim panels with floral motifs, which begin just below the base of the arches, and extend up above the top of the arches, terminating in a band of belt coursing. Atop the buttresses at either end are trim elements featuring large spheres atop rectilinear legs with floral motifs below, undulating in and out with the brick below. Additionally, a band of belt coursing, which wraps the corner of Dickson Boulevard and James Street, runs beneath the windows, only interrupted by the buttresses. Toward the back, on the building’s original rear wing, there are three windows at eye level in the original building, with bands of belt coursing below and at the top of the parapet. The rear window is a recessed bay window flanked by two pilasters with sullivanesque terra cotta panels, while the smaller windows are flanked by sullivanesque relief panels. The rear wing features a roof at multiple heights, and was extended in 1961 with a matching addition by Law, Potter and Nystrom, since removed. The rear of the taller portion of the building features a simple recessed bay with an arched window, and a similar eagle sculpture and vertical trim piece as on the front facade.

Inside, the front wing of the building features a tall banking hall with brick cladding on the walls up to the level of the windows, where it terminates at a wooden sill. The space is split down the middle by a row of brick piers and low walls framing the teller cages, which terminate at the sill line of the windows, dividing the space while still allowing it to read as a single continuous lofty space. The brick forms piers at the teller’s cages, pilasters separating desks on the exterior wall, and low brick walls with marble caps. The upper portion of the walls and the coffered ceiling in this space is finished with white plaster, which gives the space a very vertical and airy feeling, as do the cream-colored terrazzo floors, which feature black edges at the base of the walls, tying the space together. The space features a terra cotta water fountain, or bubbler, also designed by sullivan, which features intricate ornament by Schneider. The space also features two mezzanine balconies with metal railings that run below the arched windows at the front and rear of the space, allowing managers to observe the activities in the lobby and teller area below from the rear balcony, while the front balcony exists solely to balance the space and keep it symmetrical. An office for private conferences with customers was originally located near the front of the space, along with a manager’s office, allowing convenience for customers seeking a meeting with the bank management. The teller’s side of the space also housed the bank’s two vaults and several other private offices. The bank originally featured a large meeting room in the one-story rear wing, behind the vaults, with a women’s waiting room sitting along the Dickason Boulevard side of the rear wing, featuring a bay window and a restroom. The building’s interior has changed in function somewhat due to the growth of the bank, changes in bank operations, and expansion of the building with new additions to house offices and a drive-through in the rear.

The building was listed on the National Register of Historic Places in 1972, was designated a National Historic Landmark in 1976, and is a contributing structure in the Columbus Downtown Historic District, listed on the National Register of Historic Places in 1992. The building saw an addition in 2006, clad in buff brick, which replicated a historic building that formerly stood to the east, and wraps the building to the rear, with a two-story section behind a one-story annex that connects the one-story rear wing of the bank to the new building. This wing replaced older additions made in 1961, which matched the one-story rear wing of the historic building, and 1980, which was modern in appearance and slightly recessed along James Street to give precedence to the historic building. The building still functions as the main office branch of the Farmers and Merchants Union Bank, which has grown substantially. The building has been long considered to be among the best of Sullivan’s “Jewel Box Banks,” and has been kept in excellent condition by the bank’s careful and caring generational stewardship.

Polar Bear - Waving Hello! by Mandi A. Miles

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Polar Bear

"Region: America

Class: Mammalia

Order: Carnivora

Family: Ursidae

Genus: Ursus

Scientific Name: Ursus maritimus

Description : Polar bears are considered the largest land carnivores in the world, matched only by very large individual Kodiak brown bears. Both sexes differ in size throughout their range; males being much larger than females and continuing to grow for a longer period of time. These bears have long, massive skulls, necks and bodies with long legs and large paws. Ears and tail are short. The nose is more prominent or “Roman” with a black rhinarium (nose pad). The tongue is black and the eyes brown. The surface of the skin is also black. Fur colour varies slightly with the season: new coats grown just prior to the winter season are very white appearing as slightly creamy white against the stark white of their icy winter environment. This pelage is thick, coarse and long with dense underfur. Guard hairs, found throughout the pelage, are shiny, almost glossy, oily and waterproof and have hollow shafts. Polar bears moult annually between the end of May and August. The coat becomes thinner and has a yellow wash or is almost a golden colour. Male polar bears weigh between 400 – 600 kg, and have been recorded up to a maximum of 800 kg. Females are smaller than the males weighing up to 300 kg, and when pregnant up to 460 kg. Polar bears are 2.5 – 3.5 m long.

Distribution : The polar bear is circumpolar in distribution, inhabiting all Arctic seas and coastlines. It is found on the pack-ice off the Alaskan coast north of Bering Strait, off the coasts of Greenland and along the Eurasian Arctic coast from Spitsbergen to Wrangell Island. Rare stragglers reach Iceland. Individual bears have been seen on the frozen Arctic Ocean as far as latitude 88 degrees North, only 2 degrees from the North Pole. In Canada, they are found along the Arctic coasts from Alaska to Labrador and from the tip of James Bay to northern Ellesmere Island. Polar bears do roam as far as 150 kilometres inland into the coniferous forests, where they live very differently from the polar bears which belong to the high Arctic.

Habitat : They prefer areas of annual ice, which they use as a hunting platform and protective cover. This includes snow-drifted pressure ridges, refrozen cracks and areas of open water surrounded by ice. In areas where the pack ice melts by mid to late summer they come inland and live in coniferous forest areas. Here they remain until the ice re-freezes. Areas of solidly frozen sea ice and the open seas are avoided. Generally they are most common along coastal areas. Some do enter the permanent pack ice.

Food : The ringed seal is by far the most common prey. They also eat bearded seal, harp seal and hooded seal. Young walrus are sometimes taken. During the summer months they feed upon the shoreline carrion, fish, mussels, crabs, starfish, lemmings and the eggs and nestling young of waterfowl and cliff-dwelling birds. They will also graze on kelp, grasses and eat mushrooms and crowberries.

Reproduction and Development : Normally they are solitary animals outside the breeding season, the exception being a mother with cubs. Polar bears mate in mid-summer. Females first start to breed at 3 to 5 years of age. In April and May adult females are in oestrus and ready to accept a mate. They are polyandrous, meaning one female will mate with more than one male in one breeding season. Males fight among themselves for the female’s attention and a couple will pair off for a period of a few days to two weeks. With females, delayed implantation occurs, the fertilized egg does not implant in the uterus until mid-September to mid-October. Embryonic development begins at this time. Gestation periods, therefore, vary a great deal when including this period of delay. Females choose suitable locations to build their maternity dens in mid-October and retreat to them for the winter season. They give birth to one to four cubs somewhere between late November and early January. Twins are most common. The newborns are small; 25 to 30 cm long and weigh less than 1 kg. They are covered with very fine hair, appearing almost naked and their eyes are closed. Their eyes open at 6 weeks. Growth is very quick; at two months their fur has thickened; they weigh about 5 kg and move about the den. By mid-March to early April, when the den is opened, the cubs weigh about 10 kg and are surprisingly strong. The cubs suckle for nine months, occasionally one year. They are very dependent on their mother and stay with her for two years. At that time they weigh 90 - 180 kg and are half grown.

Adaptations : Polar bears are wonderfully adapted to their Arctic surroundings.

Locomotion. On land a shuffling walk may be increased to a rolling gallop of 40 km/h and can outrun caribou over a short distance. Bears are often seen standing high on their hind legs, necks stretched to scan the landscape. On thin ice, legs are spread to distribute body mass. Thickly padded and furred soles allow the bear to move quietly as well as providing good traction. Small bumps and cavities on the soles act like suction cups keeping bears from slipping on the ice. The claws are used to dig into icy slopes and to grip prey. They are strong swimmers, paddling with their forefeet only and trailing their hind feet which act as a rudder. They can stay submerged for over one minute, keeping their eyes open. They swim at a speed of approximately 6.4 km/h, often covering long distances.

Insulation. Polar bears have a thick layer of sub-cutaneous fat and very dense underfur with several layers of glossy guard hair on the outside. Their pelt is much thicker in winter and provides excellent insulation. The fat layer also adds to buoyancy in the water. Water is shed easily from the oily waterproof fur. Small, furry ears have a heavy network of blood vessels, keeping them warm and conserving heat. The tail is short and rounded also conserving heat. Fur is very dense around the soles of the feet.

Pelage. The creamy white appearance of the coat allows the bear to be inconspicuous when hunting seal. Each hair is similar to an optical fibre; colourless and hollow. Being translucent, it reflects the heat from the sun down to the base of the hair, where it is absorbed by the black skin. Whiteness comes from reflection of light rather than pigmentation.

Hunting. Bears use their keen sense of smell to detect seal breathing holes. These can be up to a kilometer away and covered by a layer of snow and ice. They will stand or lie by the seal’s blowhole in the ice for hours; they may swim towards seals resting on the ice flows with only their nose showing above the water. They will dive quietly, then swim up to the ice edge and jump out on the seal, and will also crawl towards a sunbathing seal using every piece of raised ice to conceal the approach.

Denning. Both sexes occupy dens for shelter. Topographic factors influence the den sites. In Canadian core areas, dens frequently occur on south-facing slopes where northerly prevailing winds create the best drifts, where the wind-chill is least and insulation from received solar radiation is greatest. One of the three largest denning areas worldwide is in Canada. There are three main types of winter refuges: maternity dens, temporary dens and winter shelters. During the winter any bear may dig a temporary den and use it for a few days during a storm, or take shelter in a natural cavity. Winter shelters are used for longer periods of time as resting places. This type of shelter is usually roomier with additional features such as alcoves, porches and ventilation holes. Bears do not hibernate in the strict sense of the word; they have the ability to slow down their metabolism to conserve energy at any time of year. The state of self-induced lethargy while in the shelter allows them to preserve their vital fat reserves. During this time, the body temperature of the polar bear decreases by a few degrees from normal and the respiration rates are markedly reduced. Maternity dens are built and occupied by pregnant females and can vary in size. The denning chamber is at the upper end of an entrance tunnel 1 to 2 m long. It averages 1.5 m in diameter with a height of 90 to 100cm in the middle. Drifting snow seals the entrance. The chamber is higher than the tunnel, trapping bear body heat inside. Dens not only provide a safe place to give birth to her cubs, but are also a place of protection for the cubs during their first few months. During this time she does not leave the den, remaining with her cubs and living on her reserves of fat. In the spring, with her fat severely depleted she must leave to find food to sustain herself and her cubs. After they leave the maternity den she will build temporary refuges in which to nurse, rest and shelter her young cubs and warms them as they all sleep together. She heads with her cubs towards the nearest supply of food, this is usually pack ice.

Sight and sounds. Polar bears have good eyesight. Their eyes have inner eyelids that keep the glare of the sun on snow and ice from blinding them. When defending a food source from other bears they use a deep growl. They hiss and snort to show aggression. Angry bears use loud roars and growls. Mothers scold cubs with a low growl.

Threats to Survival : Polar bears are one of the animals most threatened by global warming. They depend entirely on sea ice as a platform from which to hunt seals. Reduction of the total ice cover in the Arctic is a serious concern globally. When the ice does not form or forms too late in the season many polar bears starve. In Hudson Bay, scientists have found the main cause of death for cubs to be either lack of food or lack of fat on nursing mothers. Exploitation of minerals and fossil fuels in the Arctic pose a continuous threat. Of the oil and natural gas deposits globally, 20% are located in the Arctic. As the ice cap recedes these become more accessible. Countries are competing which each other in claiming ownership of Arctic and its resources. This can only result in further and more drastic impact on polar bear habitat.

Status : IUCN: Vulnerable; CITES: Appendix II; COSEWIC: Special Concern

Zoo Diet : Toronto Zoo carnivore diet, dog chow, jumbo smelt and herring, carrots, bean sprouts, Vitamin E and Thiamine supplements."

- Courtesy of the Metro Toronto Zoo

Please! No usage allowed without the consent of Mandi A. Miles

Based out of Flesherton, ON, please write for prices and information!

Miles Away Photography

20180106...IMGP5014cm Fullerton.CA.us Blood Orange by Ranong Payakapan

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Our blood orange starts to show its color.

Abstract

BACKGROUND

Anthocyanins are water-soluble pigments belonging to the flavonoid family. They are typically present in the flesh and peel in the blood orange cultivars. Although blood orange young shoots and flowers are not anthocyanin-colored, lemon, citron, rangpur lime, and Meyer lemon young shoots and flowers exhibit marked pigmentation as a result of anthocyanins, demonstrating that anthocyanin biosynthesis in the Citrus genus is both tissue- and genotype-dependent. The present study aimed to examine the qualitative and quantitative anthocyanin profile of fruit and other tissues from different Citrus species.

RESULTS

The presence of anthocyanin-pigmented stigmas in the young flowers of a blood orange tree (cv. ‘Moro’) was characterized and reported for the first time. The dominant pigments in blood orange fruits were cyanidin 3-glucoside and cyanidin 3-(6′′-malonyl-glucoside), whereas different patterns were observed in the young shoots, flowers and peel tissues of different Citrus species.

CONCLUSION

The present study is the first to report differentially expressed anthocyanin pigmentation patterns in different organs from several species of the genus Citrus. The results obtained could also represent a starting point for further investigations that aim to better understand which regulatory genes are involved in anthocyanin biosynthesis in the fruits, shoots and floral tissues of different Citrus species. © 2016 Society of Chemical Industry

onlinelibrary.wiley.com/doi/10.1002/jsfa.7916/abstract

thoughtful, sad, tired by tropicaLiving - Jessy Eykendorp

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White Bengal Tiger

Location: Bali Safari and Marine Park

Genus: Panthera

Species: P. tigris

Subspecies: P. t. tigris

Conservation Status: Endangered

White Bengal Tiger have stripes all over their body. Their stripes are like fingerprints. No two are the same. The stripes are not only in the tigers fur, but are a pigmentation of the skin. They have a white spot on the back of their ears,whick looks like eyes.

White Bengal tiger is very patient, it watches its prey and waits till the right time to make its move. It has very strong jaws; it catches the prey by its neck and the strong jaws and teeth help him to kill its prey. Bengal tiger has a strong sense of smell. It also has a great vision which helps it to hunt during the night time. Bengal tigers can also swim and move quite swiftly in the water. They have the ability to shed or grow body hair according to how cold or hot it gets. They are one of the only two species of cats that like water.

Ooh; what have we here? by Ed Phillips

6 7

I've been working on a series of shots for FransJanssens@www.collembola.org, to establish the size and differences between the various instars of a Katiannid springtail (Katiannidae Genus nov.1 sp. nov.). There's a colony in our Staffordshire garden.

Well, I went out this morning to see what I could find. I photographed what I thought were two mature females. These typically have lots of black pigmentation. The males often show deep orange/red. At the time, I thought this individual was particularly dark. It also looked rather small for a mature female. Once I looked on my PC, I could see it looked a bit different from usual. It has an atypical colouration pattern, is definitely small for an adult female (0.85 mm) and the final abdominal segment appears dark (it's usually non pigmented). Perhaps just an unusual colour form. I'll wait and see what Frans thinks!

Male Pigmentation by USFWS Midwest Region

5

Male coaster brook trouts are easily identified by their vibrant colors. Photo by Katie Steiger-Meister/USFWS

This image is taken from United States Naval Medical Bulletin Vol. 25, Nos. 1-4, 1927 by Medical Heritage Library, Inc.

2

Go to the Book with image in the Internet Archive

Title: United States Naval Medical Bulletin Vol. 25, Nos. 1-4, 1927

Creator: U.S. Navy. Bureau of Medicine and Surgery

Publisher:

Sponsor:

Contributor:

Date: 1927-01

Language: eng

Table of Contents

Number 1

PREFACE v

NOTICE TO SERVICE CONTRIBUTORS vi

SPECIAL ARTICLES:

Posture.

By Lieut. G. E. Mott, Medical Corps, United States Navy 1

Medical Tactics in Naval Warfare —Part III— Continued.

By Commander W. L. Mann, Medical Corps, United States Navy, and Maj. A.

D. Tuttle, Medical Corps, United States Army 20

Identification by the Teeth.

By J. H. Taylor, Identification Section, Bureau of Navigation, Navy

Department 49

An Analysis of the Annual Physical Examination of a Group of Officers.

By Lieut. Commander L. G. Roddis and Lieut. (Junior Grade) G. A.

Cooper, Medical Corps, United States Navy 54

Dressing Sterilizers with Special Reference to Temperature, Pressure,

and Chamber Air Exhaustion During the Process of Sterilization.

By Lieut. Commander J. Harper, Medical Corps, United States Navy 62

Calcium Hypochlorite for Lyster Bags.

By Lieut. E. M. Steger, Medical Corps, United States Navy 6S

The Use of Modified Milk in Infant Feeding.

By Lieut. Commander W. H. Short, Medical Corps, United States Navy 73

Aviation Crashes at Pensacola— 1925-26.

By Lieut. J. D. Benjamin, Medical Corps, United States Navy.. 86

Climatic Bubo.

By Lieut. Commander W. H. Whitmore, Medical Corps, United States Navy

89

Ideas on Recruiting.

By Lieut. J. G. Smith, Medical Corps, United States Navy 102

CLINICAL NOTES:

One Thousand Operations During a Shore Cruise.

By Lieut. Commander G. F. Cottle, Medical Corps, United States Navy 105

Filariasis.

By Lieut. Commander H. V. Hughens, Medical Corps, United States Navy. .

111

Bismuto-Yatren A and B in the Treatment of Yaws.

By Lieut. Commander R. P. Parsons, Medical Corps, United States Navy

117

Addison's Disease Without the Usual Pigmentation of the Skin.

By Commander E. C. White and Lieut. (Junior Grade) W. F. James, Medical

Corps, United States Navy 122

Scurvy.

By Lieut. Commander L. J. Roberts, Medical Corps, United States Navy

126

Early Pulmonary Tuberculosis With Negative X-ray Findings.

By Lieut. J. D. Blackwood, jr., Medical Corps, United States Navy 128

Cholecystography.

By Lieut. O. B. Spalding, Medical Corps, United States Navy.. 131

Syphilis.

By Lieut, (junior grade) B. W. Harris, Medical Corps, United States

Navy 134

Yaws Sera and the Kahn Precipitation Test, Experiments With.

By Pharmacist F. O. Huntsinger, United States Navy 135

NOTES AND COMMENTS:

Commendation for articles in the Bulletin—Treatment of burns — Parasitic

infections in China—Effect of elevation of temperature on spirochetes —Rabies

—-American Relief Administration in Russia, 1921-1923— Forecasting smallpox

epidemics in India —Helium-oxygen mixture in diving —Treatment of pernicious

anemia by diet —International meeting on cancer control, September 20-24, 1926

—American College of Surgeons —Resignation after special courses—Study courses

for Hospital Corps ratings 137-160

NURSE CORPS:

Laboratory Experiences with Epidemic Cerebrospinal Meningitis.

By Nurse S. Ruth Hassler, United States Navy 161

Help from the Laboratory.

By Nurse Ruth B. Meutzer, United States Navy 164

Some Interesting Laboratory Work.

By Nurse Frances C. Bonner, United States Navy 166

BOOK NOTICES 169

PREVENTIVE MEDICINE, STATISTICS:

Reactions incidental to the administration of 91,707 doses of

neoarsphenamine and other arsenical compounds in the United States Navy—Food

poisoning on board U. S. S. "Concord" May 13, 1926—Food poisoning

following a barbecue —Food poisoning at marine barracks, navy yard,

Philadelphia, Pa., May 21, 1926 — Influenza in Guam —Efficacy of B. typhosus

vaccine in controlling typhoid fever in Guam 177

Public Health Activities Against Tropical Diseases.

By Dr. Pedro N. Ortiz, commissioner of health, Porto Rico 208

Scarlet fever attack rate among contacts, Detroit, Mich.—Health of the

Navy— Statistics 220

Number 2

PREFACE---------------- --------- ------- - ------------- ------ vii

NOTICE TO SERVICE CONTRIBUTORS____________________________ viii

SPECIAL ARTICLES: .

MEDICINE IN TURKEY.

By Commander C. W. 0 . Bunker, Medical Corps, United States

Navy------------------------·------ 229

COLOR-BLINDNESS.

By Capt. E. H. H. Old, Medical Corps, United States Navy______ 253

ETIOLOGICAL FACTORS IN THE PRODUCTION OF MAXILLARY ANTRUM DISEASE.

By G. B. Trible, M. D., F. A. C. S., former commander, medical Corps,

United States Navy________ 266

BUREAU OF MEDICINE AND SURGERY'S EXHIBIT, SESQUICENTENNIAL,

PHILADELPHIA, 1926.

By Capt. R. C. Holcomb, Medical Corps, United States Navy____ 272

MALARIA IN HAITI.

By Capt. O. S. Butler, and Lieut. E. Peterson, Medical Corps, United

States Navy_____________ 278

INSTITUTIONAL POLICIES AS APPLIED TO A YEAR'S DENTAL ACTIVITIES.

By Lieut. Commander J. L. Brown, Dental Corps, United States

Navy________________ __________ 288

RECURRENCE OF INGUINAL HERNIA.

By Lieut. Commander K. E. Lowman, Medical Corps, United States Navy

------------------------- 300

SOME OBSERVATIONS ON CEREBROSPINAL FEVER.

By Lieut. Commander Paul Richmond, Medical Corps, United States

Navy__________________ 304

THEORIES CONCERNING THE MECHANISM OF THE INSULIN EFFECT ON CARBOHYDRATE

METABOLISM.

By Lieut. (Junior Grade) H. Phillips, Medical Corps, United States

Navy--------------------------- 309

THE WASSERMANN REACTION FOLLOWING THE USE OF BISMUTH IN THE TREATMENT

OF YAWS AND SYPHILIS.

By Lieut. (Junior Grade) C. D. Middlestadt, Medical Corps, United

States Navy _______________ 315.

THE TUBERCULOUS VETERANS' BUREAU PATIENT.

By Lieut. J. D. Blackwood, jr., Medical Corps, United States

Navy--------------------------------------- 319

CLINICAL NOTES:

OCULAR ENUCLEATION.

By Commander C. B. Camerer, and Lieut. Commander B. P. Davis, Medical

Corps, United States Navy________________ 325

TREATMENT OF AN IRREDUCIBLE DISLOCATED LOWER JAW OF 98 DAYS' DURATION.

By Lieut. Commander M. D. Willcutts, Medical Corps, United States Navy

------------------ 331

TRAUMATIC RUPTURE OF THE SPLEEN.

By Commander L. W. Johnson, Medical Corps, United States

Navy-------------------------------- 336

DIATHERMY IN SHOCK.

By Commander G. F. Cottle, Medical Corps, United States Navy_ 340

SPONTANEOUS RUPTURE OF THE HEART.

By Lieut. (Junior Grade) G. 'w. Cooper, Medical Corps, United States

Navy ------------------------------- 343

MERCUROCHROME-220 SOLUBLE, FOREIGN PROTEIN, AND SUGAR IN ACUTE

GONORRHEAL URETHRITIS, WITH A STUDY OF THE BLOOD CELLULAR CHANGES DURING THE

REACTION.

By Lieut. Commander L. H. Williams, and Lieut. (Junior Grade) G. F.

Cooper, Medical Corps, United States Navy _________ 352

GLANDULAR FEVER WITH INGUINAL ADENOPATHY.

By Lieut. (Junior Grade) C. C. Yanquell, Medical Corps, United States

Navy ___________358

A CASE OF ACQUIRED HYDROCEPHALUS.

By Capt. E. O. White, Medical Corps, United States Navy ____ 361

CARCINOMA OF THE STOMACH WITH LARGE RETROPERITONEAL TUMOR.

By Lieut. Commander W. H. Whitmore, Medical Corps, United States Navy----------------

365

HEXAMETHYLENAMINE IN MENINGEAL INFECTIONS.

By Lieut. (Junior Grade) A. E. Bruuschwig, Medical Corps, United States

Navy ________369

REINFECTION IN SYPHILIS.

By Lieut. O. P. Archambeault, Medical Corps, United States Navy--------------------------------------372

AN UNUSUAL BULLET WOUND. ·

By Lieut. (Junior Grade) P. H. Golberg, Medical Corps, United States

Navy __________374

THE NAVAL RESERVE -------------------------------------------- 375

NURSE CORPS:

THE PENSACOLA HURRICANE.

By Chief Nurse Mary J. McCloud, United States Navy ______ 379

MY FIRST EXPERIENCE IN A HURRICANE.

By Reserve Nurse Luama A. MacFarland, United States Navy _ 382

A HURRICANE THRILL.

By Nurse Ellen E. Wells and Nurse Mary Hennemeier, United States Navy

-------------------383

AN EPIDEMIC OF IMPETIGO CONTAGIOSA.

By Nurse Caroline W. Spofford, United States Navy _________ 385

NOTES AND COMMENTS:

A new section of the BULLETIN-Change in character of –examinations for

promotion-Hospital Corps instruction-Duty at sea – Handling poisons-Laboratory

technicians-The Henry S. Wellcome medal and prize for 1927-Micro-Kahn

reactions-Streptococcus cardioarthritidis--Progress in dermatology-Mitral

regurgitation- Cutaneous leishmaniasis

and the phlebotomus-Medicinals and dyes-Rectal feeding-Epidemic

encephalitis-Catarrhal jaundice- Excoriation of the skin about intestinal

fistulae-Ethylene

anesthesia-Sodoku in the treatment of general paresis—Carbuncles of the

neck-Gye's theory of cancer-$100,000 offered for conquest of cancer-The Sofie

A. Nordoff-Jung cancer prize ____387

BOOK NOTICES ------------------- 413

TOTAL FUEL REQUIREMENT IN HEALTH.

By Commander J. R. Phelps, Medical Corps, United States Navy_ 431

QUESTIONS AND ANSWERS ON SMALLPOX AND VACCINATION.

BY Surg. J. P. Leake, United States Public Health Service______ 461

A fatal case of food poisoning caused by fried oysters contaminated

with a paratyphoid B. bacillus-Outbreak of food poisoning in the wardroom mess

of the U. S. S. Richmond, caused by chicken

salad contaminated with B. enteritidis-------------------------- 475

R EPORT OF AN OUTBREAK OF FISH POISONING ON BOARD THE U. S. S. “CALIFORNIA.''

By Capt. J. L. Neilson, Medical Corps, United States Navy ------ 480

AN OUTBREAK OF FOOD POISONING CAUSED BY BOILED SMOKED TONGUE.

By Lieut. Commander W. A. Fort, Medical Corps, United States Navy -------------------------------------

484

Report of an outbreak of food poisoning caused by cheese-Outbreak of

food poisoning caused by corned-beef hash at the United States destroyer base,

San Diego, Calif. _______________ 486

REPORT OF POISONING BY TRINITROTOLUENE AMONG ENLISTED MEN ENGAGED IN

TRANSFERRING T. N. T. FROM STORAGE TO U. S. S. "NITRO."

By Lieut. Commander T. A. Fortescue, Medical Corps, United States

Navy_______________________________________________ 491

An epidemic of jaundice in San Diego, Calif.-Health of the Navy___ 494

Number 3

PREFACE ---------------------------------------------------------- v

NOTICE TO SERVICE CONTRIBUTORS--------------------------- vi

SPECIAL ARTICLES :

SOME OBSERVATIONS ON AVOIDABLE DROWNING.

By Lieut. Commander R. B. Miller, Medical Corps, United States Navy

___________505

FLIES AND THEIR ERADICATION.

By Lieut. Commander E. C. Carr, Medical Corps, United States

Navy----------------------------- 528

ACTIVITIES OF THE UNITED STATES NAVAL MEDICAL SUPPLY DEPOT.

By Capt. D. C. Cather, Medical Corps, United States Navy ___ 542

RETINITIS PIGMENTOSA.

By Commander C. B. Camerer, Medical Corps, United States

Navy---------------------------------- 562

PREMATURE CONTRACTIONS OF THE HEART.

By Capt. E. C. White, Medical Corps, United States Navy ___ 567

ROENTGEN RAY EXAMINATION IN SUSPECTED CHRONIC APPENDICITIS.

By Lieut. P. P. Maher, Medical Corps, United States Navy ____ 573

COMPARISON OF THE KAHN AND KOLMER REACTIONS.

By Lieut. Commander P. Richmond, jr., Medical Corps, United States

Navy--------------------585

COMMENTS ON THE ORIGIN OF DISABILITY.

By Lieut. Commander Lewis W. Johnson, Medical Corps, United States

Navy--------------------------------- 588

DEPARTMENT OF SANITATION, MARINE BARRACKS, QUANTICO, VA.

By Capt. W. M. Garton, Medical Corps, United States Navy ___593

CLINICAL NOTES :

CHOLECYSTOGRAPHY,

By Lieut. G. B. Larson, Medical Corps, United States Navy __597

THROMBO-ANGIITIS OBLITERANS.

By Lieut. (Junior Grade) R. A. Schneiders, Medical Corps, United States

Navy----------------------------605

THE DEBT OF SURGICAL DIAGNOSIS TO THE X RAY.

By Commander G. F. Cottle, Medical Corps, United States Navy_ 614

THE TREATMENT OF CHANOROIDS AND OTHER LESIONS WITH AMMONIACAL

SILVER NITRATE AND FORMALIN.

By Commander P. G. White, Dental Corps, and Lieut. (Junior Grade) J. Q.

Owsley, Medical Corps, United States Navy _____ 619

SUMMARY OF SEVENTY CASES OF GONOCOCCUS INFECTION TREATED WITH

MERCUROCHROME-220 SOLUBLE TOGETHER WITH SUGAR AND FOREIGN PROTEIN.

By Lieut. Commander L. H. Williams and Lieut. W. D. Small, Medical

Corps, United States Navy------621

TREATMENT OF ENCEPHALITIS LETHARGICA.

By Harold S. Hulbert, M. D. -------------------------------624

MULTIPLE UNERUPTED AND IMPACTED CUSPID AND BICUSPID TEETH.

By Lieut. Commander M. W. Mangold, Dental Corps, United States

Navy----------------- ------------------ 625

MIXED VENEREAL INFECTIONS.

By Lieut. (Junior Grade) G. F. Cooper, Medical Corps, United States

Navy-----------------------------626

CAISSON DISEASE DURING HELMET DIVING.

By Lieut. Commander W. M. Anderson, Medical Corps, United States Navy

--- ------------------------ 628

ANTERIO-VENOUS ANEURYSM OF THE COMMON CAROTID ARTERY AND INTERNAL

JUGULAR VEIN: OPERATION WITH CONSERVATION OF THE ARTERY.

By Commander F. X. Koltes, Medical Corps, United States Navy_ 630

CREEPING ERUPTION.

By Lieut. Commander H. L. Shinn, Medical Corps, United States Navy

___________________632

DEATH BY LIGHTNING.

By Lieut. J. D. Benjamin, Medical Corps, United States Navy _634

DEVICE FOR TRANSFERRING PATIENTS FROM SHIPS.

By Master Rigger Thomas Schofield _______________________ _ 635

AN ECONOMICAL AND PRACTICAL UTENSIL STERILIZER.

By Chief Pharmacist Charles Peek, United States Navy ______ _637

THE NAVAL RESERVE-------------------------------------------- 639

NURSE CORPS:

HOSPITAL HOUSEKEEPING.

By Chief Nurse Nell I. Disert, United States Navy ________ ___ _ 649

DIETS AND THE NURSE.

By J. Beatrice Bowman, Superintendent, Navy Nurse Corps __ 651

DIETETICS AT NAVAL HOSPITALS.

By Nurse Anna F. Patten, United States Navy _______________ 655

NOTES AND COMMENTS:

Comments on " Some observations on avoidable drowning

"-Hypertension- Unrecognized syphilitic myocarditis- Tuberculosis

hospitalization- Microbic dissociation-Bacteriological nomenclature - Carbon

dioxide in hiccough-Diathermy in pneumonia-Answering examination

questions-Medical ethics-Alcoholic content of brain-Electric shock-Acid and

alkali burns-Antiseptics and the nasal flora of rabbits-Annam swelling- The

metric system-Eye test for hypersensitiveness to serum-Research on

pharmacological

problems- Syphilis not caused by vaccination-Line of duty ____ 661

BOOK NOTICES--- - - --------- - - --- ------ - - - -------- ------

--------- 697

PREVENTIVE MEDICINE, STATISTICS :

Digest of the views of medical officers regarding venereal-disease

problems as recorded in various annual sanitary reports for 1926- Two deaths

following inoculation with B. typhosus vaccine- Report

of outbreak of cereb1'ospinal fever at United States naval training

station, San Diego, Calif., December, 1926-January, 1927-Report of outbreak of

scarlet fever at United States naval training station, Hampton Roads, Va.-Fatal

poisoning by arseniuretted hydrogen in the galvanizing shop at the navy yard,

Puget Sound, Wash.-Data useful in estimating the amount of food wasted by men

eating in hospital wards and in a general mess-Outbreak of food poisoning at

United States naval training station, Hampton Roads, Va., attributed to canned

Vienna sausages-Epidemic of influenza among natives of Samoa in August, 1926-

Health of the Navy --------------------------------------------

Number 4

PREFACE--------------------------------------- --------------- VII

NOTICE TO SERVICE CONTRIBUTORS----------------------------- viii

SPECIAL ARTICLES :

CHINESE MILITARY MEDICINE.

By Lieut. (Junior Grade) J. L. McClartney, Medical Corps (Vol. G),

United States Naval Reserve-------- 783

ABDOMINAL WOUNDS AND INJURIES.

By Commander Lucius W. Johnson, Medical Corps, United States Navy

---------------------------- 816

AVIATION HYGIENE.

By Commander R. G. Davis, Medical Corps, United States Navy_ 832

GOITER.

By Commander C. E. Henry, M. D., F. A. C. P., Medical Corps (Vol. S),

United States Naval Reserve __ 837

COMMENTS CONCERNING THE DUTIES OF THE COMMANDING OFFICER OF A NAVAL

HOSPITAL.

By Capt. A. Farenholt, Medical Corps, United States Navy______ 844

THE BUDGET AND THE BUREAU OF MEDICINE AND SURGERY ACCOUNTING SYSTEM.

By Chief Pharmacist B. E. Irwin, United States Navy_________ 851

ELECTROCARDIOGRAM IN THE DIAGNOSIS OF THE CARDIAC ARRHYTHMIAS.

By Lieut. Commander R. L. Nattkemper, Medical Corps, United States Navy

------------------- 862

SIPPY TREATMENT FOR PEPTIC ULCER IN NAVAL HOSPITALS.

By Lieut. (Junior Grade) A. E. Brunschwig, Medical Corps, United States

Navy ------ ------------------ 871

By Lieut. Commander W. H. Whitemore and Lieut. (Junior Grade) O. A.

Smith, Medical Corps, United States Navy______ 875

LOCAL ANESTHESIA IN EYE, EAR, NOSE, AND THROAT WORK.

By Lieut. Commander F. L. Young, Medical Corps (Vol. G), United States

Naval Reserve-------------- 879

ENDARTERITIS, ACUTE, FROM ELECTRIC SHOCK.

By Lieut. P. F. Dickens, Medical Corps, United States Navy____ 881

ACUTE OSTEOMYELITIS WITH METASTASIS.

By Lieut. G. G. Herman, Medical Corps, United States Navy____ 883

ABDOMINAL HODGKIN'S DISEASE.

By Lieut. (Junior Grade) W. D. C. Day, Medical Corps, United States

Navy--------------------- 886

A NEEDLE FOR ANESTHESIA OF THE MAXILLARY NERVE.

By Lieut. (Junior Grade) J. Connolly, Dental Corps, United States Navy

------- --- --- ----- 889

POISONING FROM SOAP-VINE.

By Lieut. (Junior Grade) G. F . Cooper, Medical Corps, United States

Navy - - ------- ---------- 892

THE DIRECT DIAGNOSIS OF PERICARDITIS WITH EFFUSION.

By Lieut. (Junior Grade) C. G. Dyke, Medical Corps, United States

Navy--- -------- - --- --- -- 894

WHAT SHOULD THE DIAGNOSIS BE?

By Lieut. Commander C. A. Andrus, Medical Corps, United States Navy-----

---------------------- 896

MULTIPLE SCROTAL FISTULAE FOLLOWING RUPTURE OF THE BLADDER WITH

STRICTURE OF URETHRA. OPERATION AND RESTORATION OF FUNCTION.

By Lieut. Commander L. H. Williams, Medical Corps, United States

Navy------ - - ----------------897

TRAUMATIC RUPTURE OF THE SPLEEN.

By Commander F. X. Koltes, Medical Corps, United States Navy_ 898

APPENDIX ABSCESS V. PYONEPHROSIs.

By Commander G. F. Cottle, Medical Corps, United States Navy_ 900

MERCUROCHROME IN A CASE WITHOUT DIAGNOSIS.

By Lieut. (Junior Grade) B. W. Harris, Medical Corps, United States Navy--------------------------903

GREASE RACK FOR AUTOMOBILES.

By Capt. A. Farenholt, Medical Corps, United States Navy ____ 908

NAVAL RESERVE:

Hospital units- Recent appointments _____________________________ 909

NURSE CORPS:

THE LECTURES ON ACCOUNTING FROM A NURSE'S POINT OF VIEW.

By Chief Nurse Mabel T. Cooper, United States Navy _________ 913

ADDRESS TO THE CLASS OF 1926 HAITIAN GENERAL HOSPITAL TRAINING SCHOOL

FOR NURSES.

By Capt. C. St. J. Butler, Medical Corps, United States Navy __ 918

A NAVY NURSE IN THE NEAR EAST.

By Mabell S. C. Smith--------------------------- 920

DIETETICS.

By Nurse Anna P. Smith, United States Navy ________________ 921

DIETETICS.

By Nurse Mary J. Miney, United States Navy ________________ 923

NOTES AND COMMENTS :

Correction-Report on treatment of Chinese wounded-Skin tests in asthma-

Treponematosis-The physiological effects of tropical climate-History of medical

practice in the State of Illinois-Mercurochrome- Water

supplies-Seasickness-Cholera in Shanghai in 1926--Fracture of the

skull-Senescence and senility-Bismuth in the treatment of syphilis-Pressure

method of vaccination - Ventilators- Neurosyphilis-Tennis leg-Tea in treatment

of burns-European influenza epidemic at end-Visit of the Relief to Washington-Aviation

instruction for Hospital Corpsmen_ ______ 927

BOOK NOTICES----------------------- ------------------------------ 959

PREVENTIVE MEDICINE, STATISTICS :

A STUDY OF TWO THOUSAND HEALTH RECORDS CONTAINING ENTRIES FOR SYPHILIS.

By Lieut. Commander H. H. Montgomery, Medical Corps, United States Navy-------------------------------973

MEASURES OF OUTSTANDING IMPORTANCE IN THE PREVENTION AND CONTROL OF

MALARIA.

By Commander M.A. Stuart, Medical Corps, United States Navy_ 996

Outbreak of infectious diarrhea in the Battle Fleet, April, 1927, not

investigated at the time of occurrence----An epidemic of malaria at the United

States Naval Station, Olongapo, P. I.- Two deaths from rabies contracted by

playing with a dog which was incubating the disease, in Hankow, China-Frequency

of rabies-An outbreak of mumps among midshipmen at the United States Naval

Academy-Health of the Navy_________ 1010

INDEX TO SUBJECTS ----------------------------------------- 1035

INDEX TO AUTHORS-------- ------------------------------------- 1047

If you have questions concerning reproductions, please contact the Contributing Library.

Note: The colors, contrast and appearance of these illustrations are unlikely to be true to life. They are derived from scanned images that have been enhanced for machine interpretation and have been altered from their originals.

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RED! by Claudia Domenig

6 23

Pompadour Cotinga (Xipholena punicea), Amazon & Beyond Cloud Forest Aviary,

Miami Metrozoo, Miami, FL

Thanks for the ID, Buckeye.. So this is a Pompadour Cotinga, I had no clue. Very posh name, but then it's an amazing looking bird. I love the color. Now I really want to have red hair again... LOL.

I saw this bird sitting in the shade and took a shot, more because it was sitting there and posing than anything else. I couldn't really make out it's color, it looked kinda brownish to me, until I looked at the LCD screen. Wow!

It came out to the feeder later and it was really that red, I didn't fake the colors.

View On Black

The Pompadour Cotingas are South American birds from the Cotingidae family. The coloration of the Cotingas is from true pigmentation, not the more common prismatic feather structure. They are usually seen alone but pairs are common. They generally do not fly with mixed-species flocks.

The Pompadour Cotinga averages 20 cm in length and 65 gm in weight. The eyes are white to yellowish white. Both sexes are plump and short-tailed. The males are a glistening wine red color with white flight feathers narrowly tipped in black. The elongated and stiff greater wing coverts are wine red with white shafts partly covering the flight feathers. The females are mainly gray and paler below. The throat and belly are grayish white, with the wings and tail dusky. Greater wing coverts and inner flight feathers are broadly edged in white. The juveniles are like the female but with dark eyes.

Farmers and Merchants Union Bank, James Street and Dickason Boulevard, Columbus, WI by Warren LeMay

2

Built in 1919-1920, this Chicago School and Sullivanesque-style building was designed by Louis Sullivan for the Farmers and Merchants Union Bank in Columbus, Wisconsin as one of his late-career “jewel box” bank buildings that are largely located in smaller communities throughout the midwest. The building was the last “jewel box bank” designed by Sullivan, and the second-to-last commission of his career, and was intended to communicate the bank as a modern and progressive institution, rather than employing the stodgier and more traditional Classical design found on most other banks of the era. The bank was commissioned by the president of the bank, J. Russell Wheeler, whose wife, Anna May Wheeler, pushed him to commission Sullivan to design a new home for the bank. In addition to Louis Sullivan, the building’s stained glass windows, were designed by architectural decorator Louis J. Millet, and the terra cotta by clay modeler Kristian Schneider, whom developed moulds for the building’s terra cotta, metal, and plaster details. The two artisans worked alongside Sullivan on several other bank projects. The building was heavily documented in Sullivan’s 1924 “A System of Architectural Ornament”, published shortly before he died.

The building is clad in red tapestry brick, which features blue and green mixed with the red clay mixture in some bricks, creating variation in color and texture across the facade. The brick creates a backdrop to some of the best terra cotta on any of Sullivan’s projects. The terra cotta features many of the floral and geometric motifs found on Sullivan’s other works, and is arranged similarly to other Sullivan banks that utilized brick cladding. The building features two principal facades, with a narrower facade along James Street, and a broader facade facing Dickason Boulevard. The James Street facade features two openings close to ground level, with the eastern bay housing a large plate glass window, and the western bay housing a doorway flanked by skylights, both of which are recessed under a large terra cotta architrave and flanked by square pilasters with decorative Sullivanesque ornament panels at the capitals. The architrave above the doorway and window is divided into three segments by vertical terra cotta elements that feature floral motifs and, like many Sullivan buildings, appear like plants with roots, branches, and crowns. The outer panels of the architrave feature circular cartouches with hexagonal trim, leaves, and geometric elements, with circular central medallions featuring the years 1861, when the bank was founded, and 1919, when the bank was completed. The central panel is clad in marble with the words “Farmers & Merchants Union Bank” and “Louis Sullivan, Architect” engraved into the stone with yellow pigmentation, contrasting against the white and green marble background. Atop the two vertical elements on either side of the central panel are griffin sculptures holding shields, a common element on many of Sullivan’s “Jewel Box Banks,” while the base of the outer vertical elements features the initials of the bank at the base. Above the architrave is an arched bay that houses a stained glass window, trimmed with decorative terra cotta at the inner and outer rings of the arch, with the bay becoming more recessed after each concentric arch, much like the entrances to medieval Romanesque churches. Besides a band of belt coursing that runs on either side of the architrave and wraps the corner to a tapered buttress on the Dickason Boulevard facade, the only other adornment is an eagle sculpture on a vertical trim element at the center of the parapet, which terminates many brick courses above the arched opening below, and another band of terra cotta trim along the top of the parapet, which forms a cap on the parapet around the perimeter of the building’s low-slope roof. On the Dickason Boulevard facade, the building features five recessed clerestory arched bays housing stained glass windows, flanked by tapered buttresses. Surrounding the arched tops of the windows are decorative trim panels with floral motifs, which begin just below the base of the arches, and extend up above the top of the arches, terminating in a band of belt coursing. Atop the buttresses at either end are trim elements featuring large spheres atop rectilinear legs with floral motifs below, undulating in and out with the brick below. Additionally, a band of belt coursing, which wraps the corner of Dickson Boulevard and James Street, runs beneath the windows, only interrupted by the buttresses. Toward the back, on the building’s original rear wing, there are three windows at eye level in the original building, with bands of belt coursing below and at the top of the parapet. The rear window is a recessed bay window flanked by two pilasters with sullivanesque terra cotta panels, while the smaller windows are flanked by sullivanesque relief panels. The rear wing features a roof at multiple heights, and was extended in 1961 with a matching addition by Law, Potter and Nystrom, since removed. The rear of the taller portion of the building features a simple recessed bay with an arched window, and a similar eagle sculpture and vertical trim piece as on the front facade.

Inside, the front wing of the building features a tall banking hall with brick cladding on the walls up to the level of the windows, where it terminates at a wooden sill. The space is split down the middle by a row of brick piers and low walls framing the teller cages, which terminate at the sill line of the windows, dividing the space while still allowing it to read as a single continuous lofty space. The brick forms piers at the teller’s cages, pilasters separating desks on the exterior wall, and low brick walls with marble caps. The upper portion of the walls and the coffered ceiling in this space is finished with white plaster, which gives the space a very vertical and airy feeling, as do the cream-colored terrazzo floors, which feature black edges at the base of the walls, tying the space together. The space features a terra cotta water fountain, or bubbler, also designed by sullivan, which features intricate ornament by Schneider. The space also features two mezzanine balconies with metal railings that run below the arched windows at the front and rear of the space, allowing managers to observe the activities in the lobby and teller area below from the rear balcony, while the front balcony exists solely to balance the space and keep it symmetrical. An office for private conferences with customers was originally located near the front of the space, along with a manager’s office, allowing convenience for customers seeking a meeting with the bank management. The teller’s side of the space also housed the bank’s two vaults and several other private offices. The bank originally featured a large meeting room in the one-story rear wing, behind the vaults, with a women’s waiting room sitting along the Dickason Boulevard side of the rear wing, featuring a bay window and a restroom. The building’s interior has changed in function somewhat due to the growth of the bank, changes in bank operations, and expansion of the building with new additions to house offices and a drive-through in the rear.

The building was listed on the National Register of Historic Places in 1972, was designated a National Historic Landmark in 1976, and is a contributing structure in the Columbus Downtown Historic District, listed on the National Register of Historic Places in 1992. The building saw an addition in 2006, clad in buff brick, which replicated a historic building that formerly stood to the east, and wraps the building to the rear, with a two-story section behind a one-story annex that connects the one-story rear wing of the bank to the new building. This wing replaced older additions made in 1961, which matched the one-story rear wing of the historic building, and 1980, which was modern in appearance and slightly recessed along James Street to give precedence to the historic building. The building still functions as the main office branch of the Farmers and Merchants Union Bank, which has grown substantially. The building has been long considered to be among the best of Sullivan’s “Jewel Box Banks,” and has been kept in excellent condition by the bank’s careful and caring generational stewardship.

Albinism,,, Haji Malang Beggars by firoze shakir photographerno1

3 2

From Wikipedia, the free encyclopedia

Albinism (from Latin albus, "white"; see extended etymology, also called achromia, achromasia, or achromatosis) is a congenital disorder characterized by the complete or partial absence of pigment in the skin, hair and eyes due to absence or defect of tyrosinase, a copper-containing enzyme involved in the production of melanin. Albinism results from inheritance of recessive gene alleles and is known to affect all vertebrates, including humans. While an organism with complete absence of melanin is called an albino (US /ælˈbaɪnoʊ/,[1] or UK /ælˈbiːnoʊ/)[2] an organism with only a diminished amount of melanin is described as albinoid.[3]

Albinism is associated with a number of vision defects, such as photophobia, nystagmus and astigmatism. Lack of skin pigmentation makes for more susceptibility to sunburn and skin cancers. In rare cases such as Chédiak–Higashi syndrome, albinism may be associated with deficiencies in the transportation of melanin granules. This also affects essential granules present in immune cells leading to increased susceptibility to infection.[4]

In humans, there are two principal types of albinism, oculocutaneous, affecting the eyes, skin and hair, and ocular affecting the eyes only.

Most oculocutaenous albinistic humans appear white or very pale as the melanin pigments responsible for brown, black, and some yellow colorations are not present. Ocular albinism results in pale blue eyes, and may require genetic testing to diagnose.

Because individuals with albinism have skin that entirely lacks the dark pigment melanin, which helps protect the skin from the sun's ultraviolet radiation, their skin can burn more easily from overexposure.[5]

The human eye normally produces enough pigment to color the iris blue, green or brown and lend opacity to the eye. However, there are cases in which the eyes of an albinistic person appear red, pink or purple, depending on the amount of pigment present, due to the red of retina being visible through the iris. Lack of pigment in the eyes also results in problems with vision, both related and unrelated to photosensitivity.

The albinistic are generally as healthy as the rest of the population (but see related disorders below), with growth and development occurring as normal, and albinism by itself does not cause mortality,[6] although the lack of pigment blocking ultraviolet radiation increases the risk of skin cancer and other problems.

Visual problems[edit source | editbeta]

Malian singer Salif Keita

Development of the optical system is highly dependent on the presence of melanin, and the reduction or absence of this pigment in albinistic individuals may lead to

Misrouting of the retinogeniculate projections, resulting in abnormal decussation (crossing) of optic nerve fibres[5]

Photophobia and decreased visual acuity due to light scattering within the eye (ocular straylight)[5][7]

Reduced visual acuity due to foveal hypoplasia and possibly light-induced retinal damage[5]

Eye conditions common in albinism include:

Nystagmus, irregular rapid movement of the eyes back and forth, or in circular motion.[5]

Astigmatism, irregular shaped cornea requiring additional cylindrical corrective lenses in spectacles.[8]

Amblyopia, decrease in acuity of one or both eyes due to poor transmission to the brain, often due to other conditions such as strabismus.[5]

Optic nerve hypoplasia, underdevelopment of the optic nerve

Some of the visual problems associated with albinism arise from a poorly developed retinal pigment epithelium (RPE) due to the lack of melanin.[citation needed] This degenerate RPE causes foveal hypoplasia (a failure in the development of normal foveae), which results in eccentric fixation and lower visual acuity, and often a minor level of strabismus.

The iris is a sphincter formed from pigmented tissue that contracts when the eye is exposed to bright light, to protect the retina by limiting the amount of light passing through the pupil. In low light conditions the iris relaxes to allow more light to enter the eye. In albinistic subjects, the iris does not have enough pigment to block the light, thus the decrease in pupil diameter is only partially successful in reducing the amount of light entering the eye.[citation needed] Additionally, the improper development of the RPE, which in normal eyes absorbs most of the reflected sunlight, further increases glare due to light scattering within the eye.[9] The resulting sensitivity (photophobia) generally leads to discomfort in bright light, but this can be reduced by the use of sunglasses and/or brimmed hats.[10]

Genetics[edit source | editbeta]

Most forms of albinism are the result of the biological inheritance of genetically recessive alleles (genes) passed from both parents of an individual, though some rare forms are inherited from only one parent. There are other genetic mutations which are proven to be associated with albinism. All alterations, however, lead to changes in melanin production in the body.[6][11]

The chance of offspring with albinism resulting from the pairing of an organism with albinism and one without albinism is low. However, because organisms can be carriers of genes for albinism without exhibiting any traits, albinistic offspring can be produced by two non-albinistic parents. Albinism usually occurs with equal frequency in both sexes.[6] An exception to this is ocular albinism, which it is passed on to offspring through X-linked inheritance. Thus, ocular albinism occurs more frequently in males as they have a single X and Y chromosome, unlike females, whose genetics are characterized by two X chromosomes.[12]

There are two different forms of albinism: a partial lack of the melanin is known as hypomelanism, or hypomelanosis and the total absence of melanin is known as amelanism or amelanosis.

Diagnosis[edit source | editbeta]

This section needs additional citations for verification. Please help improve this article by adding citations to reliable sources. Unsourced material may be challenged and removed. (November 2007)

Genetic testing can confirm albinism and what variety it is, but offers no medical benefits except in the cases of non-OCA disorders (see below) that cause albinism along with other medical problems which may be treatable. The symptoms of albinism can be treated by various methods detailed below.

Treatment[edit source | editbeta]

For the most part, treatment of the eye conditions consists of visual rehabilitation. Surgery is possible on the ocular muscles to decrease nystagmus, strabismus and common refractive errors like astigmatism.[5] Nystagmus-damping surgery can also be performed, to reduce the "shaking" of the eyes back and forth.[13] The effectiveness of all these procedures varies greatly and depends on individual circumstances.

Glasses and other vision aids, large-print materials as well as bright but angled reading lights, can help individuals with albinism, even though their vision cannot be corrected completely. Some people with albinism do well using bifocals (with a strong reading lens), prescription reading glasses, and/or hand-held devices such as magnifiers or monoculars.[10] Contact lenses may be colored to block light transmission through the iris. But in the case of nystagmus this is not possible, due to the irritation that is caused by the movement of the eyes. Some use bioptics, glasses which have small telescopes mounted on, in, or behind their regular lenses, so that they can look through either the regular lens or the telescope. Newer designs of bioptics use smaller light-weight lenses. Some US states allow the use of bioptic telescopes for driving motor vehicles. (See also NOAH bulletin "Low Vision Aids".)

Epidemiology[edit source | editbeta]

Albinism affects people of all ethnic backgrounds; its frequency worldwide is estimated to be approximately one in 17,000. Prevalence of the different forms of albinism varies considerably by population, and is highest overall in people of sub-Saharan African descent.[14]

Society and culture[edit source | editbeta]

Main articles: Persecution of people with albinism and Albinism in popular culture

In physical terms, humans with albinism commonly have visual problems and need sun protection. They often face social and cultural challenges (even threats), as the condition is often a source of ridicule, discrimination, or even fear and violence. Many cultures around the world have developed beliefs regarding people with albinism.

In African countries such as Tanzania[15] and Burundi,[16][17] there has been an unprecedented rise in witchcraft-related killings of albino people in recent years, because their body parts are used in potions sold by witchdoctors. Numerous authenticated incidents have occurred in Africa during the 21st Century.[18][19][20][21] For example, in Tanzania, in September 2009, three men were convicted of killing a 14-year-old albino boy and severing his legs in order to sell them for witchcraft purposes.[22] Again in Tanzania and Burundi in 2010, the murder and dismemberment of a kidnapped albino child was reported from the courts,[16] as part of a continuing problem. National Geographic estimates that in Tanzania a complete set of albino body parts is worth $75,000.[1]

Another harmful and false belief is that sex with an albinistic woman will cure a man of HIV. This has led, for example in Zimbabwe, to rapes (and subsequent HIV infection).[23]

Certain ethnic groups and insular areas exhibit heightened susceptibility to albinism, presumably due to genetic factors. These include notably the Native American Kuna and Zuni nations (respectively of Panama and New Mexico); Japan, in which one particular form of albinism is unusually common; and Ukerewe Island, the population of which shows a very high incidence of albinism.[24]

Famous people with albinism include historical figures such as Oxford don William Archibald Spooner; actor-comedian Victor Varnado; musicians such as Johnny and Edgar Winter, Salif Keita, Winston "Yellowman" Foster, Brother Ali, Sivuca, Willie "Piano Red" Perryman; and fashion models Connie Chiu and Shaun Ross. Emperor Seinei of Japan is thought to have been an albino because he was said to have been born with white hair.

In animals[edit source | editbeta]

Many animals with albinism lack their protective camouflage and are unable to conceal themselves from their predators or prey; the survival rate of animals with albinism in the wild is usually quite low.[25][26] However the novelty of albino animals has occasionally led to their protection by groups such as the Albino Squirrel Preservation Society.

In what used to be called "partial albinism" but is more often termed leucism there can be a single patch or patches of skin that lack melanin. Especially in albinistic birds and reptiles, ruddy and yellow hues or other colors may be present on the entire body or in patches (as is common among pigeons), because of the presence of other pigments unaffected by albinism such as porphyrins, pteridines and psittacins, as well as carotenoid pigments derived from the diet.

Intentionally bred albinistic strains of some animal species are commonly used as model organisms in biomedical study and experimentation, although some researchers have argued that they are not always the best choice.[27] Examples include the BALB/c mouse and Wistar and Sprague Dawley rat strains, while albino rabbits were historically used for Draize toxicity testing.[28] The yellow mutation in fruit flies is their version of albinism.

The incidence of albinism can be artificially increased in fish by exposing the eggs to heavy metals (arsenic, cadmium, copper, mercury, selenium, zinc).[29]

The eyes of an albino animal appear red because the colour of the red blood cells in the underlying retinal blood vessels shows through where there is no pigment to obscure it.

Famous albino animals include Migaloo, a humpback whale off the coast of Australia; Pinky, a bottlenose dolphin living in and around in Calcasieu Lake, Louisiana; Snowflake, a Barcelona Zoo gorilla; Snowflake, a Bristol Zoo penguin; and Mahpiya Ska (Sioux for "White Cloud"), a buffalo in Jamestown, North Dakota.[30] The inspiration for Herman Melville's novel Moby-Dick was a sperm whale known as Mocha Dick.

~Dicyrtoma Chow Down~ by Eddie The Bugman

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A Dicyrtoma fusca chowing down on something, not exactly sure what, certainly not a full Turkey dinner LOL. This is the colour variant with distinct middorsal dark line, and yellowish lateral pigmentation or to those in the know Var.1. This was a 7 image focus stack at x7 magnification and shows the abdominal patterning quite nicely, see comments. You can also see how they can blend in quite nicely with their environment, I often struggle to see these guys first off, these guys are typically about 1.5-2.5mm in length at a guess, similar in size to Dicyrtomina species, incidentally Dicyrtomina means small Dicyrtoma :0)

Anyway, hope you are all having a cracking Christmas, I did manage to get out today, but everywhere is waterlogged, the local park has a pond which had well and truly burst its banks :o)

VIEW ON BLACK

Grey Parrot - Congo Grey Parrot - Congo African Grey Parrot - African Grey Parrot (Psittacus Erithacus) by PHOTOGRAPHY by DM & DBM.

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The grey parrot (Psittacus erithacus), also known as the Congo grey parrot, Congo African grey parrot or African grey parrot, is an Old World parrot in the family Psittacidae. The Timneh parrot (Psittacus timneh) once was identified as a subspecies of the grey parrot, but has since been elevated to a full species.

Taxonomy

The grey parrot was formally described in 1758 by the Swedish naturalist Carl Linnaeus in the tenth edition of his Systema Naturae. He placed it with all the other parrots in the genus Psittacus and coined the binomial name Psittacus erithacus. Linnaeus erroneously specified the type locality as "Guinea": the locality was later designated as Ghana in West Africa. The genus name is Latin for "parrot". The specific epithet erithacus is Latin and is derived from the Ancient Greek εριθακος (erithakos) for an unknown bird that was said to mimic human sounds, perhaps the black redstart. The species is monotypic: no subspecies are recognised.

The Timneh parrot was formerly treated as a subspecies of the grey parrot but is now considered to be a separate species based mainly on the results from a genetic and morphological study published in 2007. Although Linnaeus placed all the parrots known to him in the genus Psittacus, only the grey parrot and the Timneh parrot are now assigned to this genus.

Description

The grey parrot is a medium-sized, predominantly grey, black-billed parrot. Its typical weight is 400 g (14 oz), with an approximate length of 33 cm (13 in), and a wingspan of 46–52 cm (18–20+1⁄2 in). The grey colour on the head and wings is generally darker than its body. The head and body feathers have slight white edges. The tail feathers are red.

Due to selection by parrot breeders, some grey parrots are partly or completely red. Both sexes appear similar. The colouration of juveniles is similar to that of adults, but typically their eyes are dark grey to black, in comparison to the yellow irises around dark pupils of the adult birds, and their undertail coverts are tinged with grey. Adults weigh 418–526 g (14+3⁄4–18+1⁄2 oz).

Grey parrots may live for 40–60 years in captivity, although their mean lifespan in the wild appears to be shorter—approximately 23 years. They start breeding at an age of 3–5 years and lay 3-5 eggs per brood.

Distribution and habitat

The grey parrot is native to equatorial Africa, including Angola, Cameroon, the Congo, Gabon, Ivory Coast, Ghana, Kenya, and Uganda. The species is found inside a range from Kenya to the eastern part of the Ivory Coast. Current estimates for the global population are uncertain and range from 630,000 to 13 million birds. Populations are decreasing worldwide. The species seems to favor dense forests, but can also be found at forest edges and in more open vegetation types, such as gallery and savanna forests.

A population study published in 2015 found that the species had been "virtually eliminated" from Ghana with numbers declining 90 to 99% since 1992. They were found in only 10 of 42 forested areas, and three roosts that once held 700–1200 birds each, now had only 18 in total. Local people mainly blamed the pet trade and the felling of timber for the decline. Populations are thought to be stable in Cameroon. In the Congo, an estimated 15,000 are taken every year for the pet trade, from the eastern part of the country, although the annual quota is stated to be 5,000.

Grey parrots have escaped or been deliberately released into Florida, U.S., but no evidence indicates that the population is breeding naturally.

Behaviour and ecology in the wild

Little is known about the behaviour and activities of these birds in the wild. In addition to a lack of research funding, it can be particularly difficult to study these birds in wild situations due to their status as prey animals, which leads them to have rather secretive personalities. It has been shown that wild greys may also imitate a wide variety of sounds they hear, much like their captive relatives. In the Democratic Republic of the Congo, two greys sound-recorded while roosting reportedly had a repertoire of over 200 different calls, including nine imitations of other wild bird songs and one of a bat.

Feeding

Grey parrots are mainly frugivorous, with most of their diet consisting of fruit, nuts, and seeds, including oil palm fruit. They sometimes also eat flowers and tree bark, as well as insects and snails. In the wild, the grey parrot is partly a ground feeder.

Breeding

Grey parrots are monogamous breeders who nest in tree cavities. Each mated pair of parrots needs their own tree for their nest. The hen lays three to five eggs, which she incubates for 30 days while being fed by her mate. The adults defend their nesting sites.

Grey parrot chicks require feeding and care from their parents in the nest. The parents take care of them until 4–5 weeks after they are fledged. Young leave the nest at the age of 12 weeks. Little is known about the courtship behaviour of this species in the wild.[9] They weigh 12–14 g (7⁄16–1⁄2 oz) at hatching and 372–526 g (13+1⁄8–18+1⁄2 oz) when they leave their parents.

Conservation

Natural predators for this species include palm-nut vultures and several raptors. Monkeys target eggs and the young for food.

Humans are by far the largest threat to wild grey populations. Between 1994 and 2003, more than 359,000 grey parrots were traded on the international market. Approximately 21% of the wild population was being harvested every year. Mortality rates are extremely high between the time they are captured and they reach the market, ranging from 60 to 66%. This species also is hunted for its meat and for its body parts, which are used in traditional medicines. As a result of the extensive harvest of wild birds, in addition to habitat loss, this species is believed to be undergoing a rapid decline in the wild and therefore, has been rated as endangered by the International Union for Conservation of Nature.

In October 2016, the Convention on the International Trade of Endangered Fauna and Flora (CITES) extended the highest level of protection to grey parrots by listing the species under Appendix 1, which regulates international trade in the species.

In 2021, the Kenyan government held a short amnesty, during which grey parrot owners could pay a fee to obtain a permit for their birds and facilitate legal ownership. Following the expiry of this time period, it is now illegal to own this species without a permit.

In captivity

The species is common in captivity and regularly kept by humans as a companion parrot, prized for its ability to mimic human speech, which makes it one of the most popular avian pets. An escaped pet in Japan was returned to his owner after repeating the owner's name and address.

Grey parrots are notorious for mimicking noises heard in their environment and using them tirelessly. They are highly intelligent birds, needing extensive behavioral and social enrichment as well as extensive attention in captivity or else they may become distressed. Feather plucking is a common symptom seen among such distressed grey parrots, affecting up to 40% of captive individuals. They may also be prone to behavioural problems due to their sensitive nature. Social isolation hastens stress and aging.

The grey parrot is a highly social species which relies on a flock-type structure, even when raised in captivity. Because they are so dependent on the other birds within their flock, much of their speech and vocal ability is acquired through interaction with the humans with whom they reside. Both wild and captive parrots have been shown to use contact calls, which allow them to interact with their flock mates and communicate information about their location, detection of predators, availability of food, and safety status. In addition, contact calls are used to form strong social bonds with their flock mates, or in the case of captive greys, with their human housemates. In captivity, they have been shown to display communicative competence, meaning they not only use human language correctly, but also in such a way that is appropriate for the social situation which they are in.

Diet

In captivity, they may be fed bird pellets, a variety of fruits such as pear, orange, pomegranate, apple, and banana, and vegetables such as carrot, cooked sweet potato, celery, fresh kale, peas, and green beans. They also need a source of calcium.

Disease

Grey parrots in captivity have been observed to be susceptible to fungal infections, bacterial infections, nutritional insufficiency, malignant tumors, psittacine beak and feather disease, tapeworms, and blood-worms. Young grey parrots are more commonly infected by psittacine beak and feather disease than adults. Infected birds show symptoms such as loss of appetite, fluffy feathers, sluggishness, and reduced walking abilities due to brittle bones.

Grey parrots are more likely to have rhinitis,[clarification needed] an inflammatory and infectious disease of the nasal cavity. Birds may exhibit signs like wheezing, sneezing, nasal snuffling, and swelling or occlusion of the nares. Treatment options include gentle debridement and nasal irrigation.

Intelligence and cognition

Grey parrots are highly intelligent and are considered by many to be one of the most intelligent species of psittacines. Many individuals have been shown to perform at the cognitive level of a four- to six-year-old human child in some tasks. Several studies have been conducted, indicating a suite of higher-level cognitive abilities. Experiments have shown that grey parrots can learn number sequences and can learn to associate human voices with the faces of the humans who create them. It has been reported that grey parrots are capable of using existing known English words to create new labels for objects when the bird does not know the name of the object. For example "banerry" ("banana" + "cherry") for "apple", "banana crackers" for "dried banana chips" or "yummy bread" for "cake".

The American scientist Irene Pepperberg's research with Alex the parrot showed his ability to learn more than 100 words, differentiating between objects, colours, materials and shapes. Pepperberg spent several decades working with Alex, and wrote numerous scientific papers on experiments performed, indicating his advanced cognitive abilities. One such study found that Alex had the ability to add numbers as well as having a zero-like concept, similar to that of young children and apes.

In addition to their striking cognitive abilities, grey parrots have displayed altruistic behavior and concern for others. Researchers found that while blue-headed macaws were unlikely to share a nut with other members of their own species, grey parrots would actively give their conspecific partner a nut, even if it meant that they would not be able to get one themselves. When the roles were reversed, their partners were overwhelmingly likely to return the favor, foregoing their own nut to their partner's benefits. This indicates not only a display of selflessness but also an act of reciprocity.

A 2012 study demonstrated that captive grey parrots have individual musical preferences. When presented with the opportunity to choose between two different pieces of music via a touch screen monitor located in their cage, the two birds in the test consistently chose different songs, to which they then danced and sang along. Some pet grey parrots have also been observed using the music feature of smart speakers (such as Alexa or Amazon Echo) to verbally request playback of specific favored songs.

Some research has shown that foot preference can be linked to the number of words a particular parrot may know and use. Researchers found that grey parrots who prefer to use their right foot showed a marked increase in the number of words within their lexicon as compared to parrots who were left-footed. Scientists postulate that parrots may have lateralization of brain function, much like mammals do.

In two murder trials, one in 1993 and another in 2017, there was consideration to use the deceased victim's pet grey parrot's "testimony" as evidence due to the pet parrot's witnessing and repeating the victim's last words. In the 1993 murder trial of Gary Joseph Rasp, the defendant was accused of murdering Jane Gill. Public defender Charles Ogulnik wanted to use Jane's pet grey parrot Max as evidence to prove Gary's innocence due to Max repeating Jane's last words "Richard, no, no, no!". In the 2017 murder trial of Glenna Duram, the defendant is accused of murdering her husband Martin Duram. The prosecutor was exploring the possibility of using the couple's pet parrot Bud as evidence when Bud kept repeating Martin's last words "Don't fucking shoot."

Mutations

Grey mutations occur naturally in the wild, such as the Blue Ino (albino), the Incomplete Ino, and the Blue varietals. The Blue Ino is all white. The Incomplete Ino has light pigmentation. The Blue has a white tail.

Breeders from South Africa, Australia, New Zealand, and Scandinavia have bred greys intensively since the 1800s. These bred varieties include the Red Pied, F2 Pied, Grizzles, Ino, Incomplete, Parino, Lutino, Cinnamon, and Red Factor. South African bird breeder Von van Antwerpen and New Zealand partner Jaco Bosman selected F2 Pieds and created the first Red Factor Greys. They are rare, may be predominantly red-pigmented, and vary in price depending upon the extent of the red plumage displayed.

History

The domestication of grey parrots has a history dating to 2000 B.C., depicting native birds in Egyptian hieroglyphics as pets. They were used for values by the Greeks and the Romans who kept them in birdcages. The grey parrots, due to recent years of illegal trading, have been classified as Endangered in 2016 by the IUCN Red List.

Vicious B's Pigmentations by Vanny

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portrait by Patrick Marioné

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Brussels

At Laguna Colorada by Northern Adventures

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Laguna Colorada (Red Lagoon) is a shallow salt lake in the southwest of the altiplano of Bolivia, within Eduardo Avaroa Andean Fauna National Reserve and close to the border with Chile.

The lake contains borax islands, whose white color contrasts with the reddish color of its waters, which is caused by red sediments and pigmentation of some algae.

Laguna Colorada is one of the Ramsar Wetlands of International Importance under the Ramsar Convention signed in 1971.

James's Flamingos abound in the area. Also it is possible to find Andean and Chilean flamingos, but in a minor quantity.

(Wikipedia)

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We took the backward route from Tupiza northwards to the Reserva Nacional de Fauna Andina Eduardo Avaroa and further to Salar de Uyuni salt plain; snow-covered volcanoes at the horizon line, red-coloured rocks and sand, colorful lagunas, and slow 4WD journey through the land of thin air - that's Bolivia southern part of Altiplano.

Laguna Colorada is most likely the most stunning of the colorful lakes; we spent there a night and thus got a chance for slow silent walks around, admiring its unforgettable stunning beauty...

Jellyfish - Sea Jellies - Medusa (Subphylum Medusozoa) by PHOTOGRAPHY by DM & DBM.

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Jellyfish, also known sea jellies, are the medusa-phase of certain gelatinous members of the subphylum Medusozoa, which is a major part of the phylum Cnidaria.

Jellyfish are mainly free-swimming marine animals with umbrella-shaped bells and trailing tentacles, although a few are anchored to the seabed by stalks rather than being mobile. The bell can pulsate to provide propulsion for highly efficient locomotion. The tentacles are armed with stinging cells and may be used to capture prey and defend against predators. Jellyfish have a complex life cycle. The medusa is normally the sexual phase, which produces planula larvae; these then disperse widely and enter a sedentary polyp phase, before reaching sexual maturity.

Jellyfish are found all over the world, from surface waters to the deep sea. Scyphozoans (the "true jellyfish") are exclusively marine, but some hydrozoans with a similar appearance live in freshwater. Large, often colorful, jellyfish are common in coastal zones worldwide. The medusae of most species are fast-growing, and mature within a few months then die soon after breeding, but the polyp stage, attached to the seabed, may be much more long-lived. Jellyfish have been in existence for at least 500 million years, and possibly 700 million years or more, making them the oldest multi-organ animal group.

Jellyfish are eaten by humans in certain cultures. They are considered a delicacy in some Asian countries, where species in the Rhizostomeae order are pressed and salted to remove excess water. Australian researchers have described them as a "perfect food": sustainable and protein-rich but relatively low in food energy.

They are also used in research, where the green fluorescent protein used by some species to cause bioluminescence has been adapted as a fluorescent marker for genes inserted into other cells or organisms.

The stinging cells used by jellyfish to subdue their prey can injure humans. Thousands of swimmers worldwide are stung every year, with effects ranging from mild discomfort to serious injury or even death. When conditions are favourable, jellyfish can form vast swarms, which can be responsible for damage to fishing gear by filling fishing nets, and sometimes clog the cooling systems of power and desalination plants which draw their water from the sea.

Names

The name jellyfish, in use since 1796, has traditionally been applied to medusae and all similar animals including the comb jellies (ctenophores, another phylum). The term jellies or sea jellies is more recent, having been introduced by public aquaria in an effort to avoid use of the word "fish" with its modern connotation of an animal with a backbone, though shellfish, cuttlefish and starfish are not vertebrates either. In scientific literature, "jelly" and "jellyfish" have been used interchangeably. Many sources refer to only scyphozoans as "true jellyfish".

A group of jellyfish is called a "smack" or a "smuck".

Definition

The term jellyfish broadly corresponds to medusae, that is, a life-cycle stage in the Medusozoa. The American evolutionary biologist Paulyn Cartwright gives the following general definition:

Typically, medusozoan cnidarians have a pelagic, predatory jellyfish stage in their life cycle; staurozoans are the exceptions [as they are stalked].

The Merriam-Webster dictionary defines jellyfish as follows:

A free-swimming marine coelenterate that is the sexually reproducing form of a hydrozoan or scyphozoan and has a nearly transparent saucer-shaped body and extensible marginal tentacles studded with stinging cells.

Given that jellyfish is a common name, its mapping to biological groups is inexact. Some authorities have called the comb jellies and certain salps jellyfish, though other authorities state that neither of these are jellyfish, which they consider should be limited to certain groups within the medusozoa.

The non-medusozoan clades called jellyfish by some but not all authorities (both agreeing and disagreeing citations are given in each case) are indicated with on the following cladogram of the animal kingdom:

Jellyfish are not a clade, as they include most of the Medusozoa, barring some of the Hydrozoa. The medusozoan groups included by authorities are indicated on the following phylogenetic tree by the presence of citations. Names of included jellyfish, in English where possible, are shown in boldface; the presence of a named and cited example indicates that at least that species within its group has been called a jellyfish.

Taxonomy

The subphylum Medusozoa includes all cnidarians with a medusa stage in their life cycle. The basic cycle is egg, planula larva, polyp, medusa, with the medusa being the sexual stage. The polyp stage is sometimes secondarily lost. The subphylum include the major taxa, Scyphozoa (large jellyfish), Cubozoa (box jellyfish) and Hydrozoa (small jellyfish), and excludes Anthozoa (corals and sea anemones). This suggests that the medusa form evolved after the polyps. Medusozoans have tetramerous symmetry, with parts in fours or multiples of four.

The four major classes of medusozoan Cnidaria are:

Scyphozoa are sometimes called true jellyfish, though they are no more truly jellyfish than the others listed here. They have tetra-radial symmetry. Most have tentacles around the outer margin of the bowl-shaped bell, and long, oral arms around the mouth in the center of the subumbrella.

Cubozoa (box jellyfish) have a (rounded) box-shaped bell, and their velarium assists them to swim more quickly. Box jellyfish may be related more closely to scyphozoan jellyfish than either are to the Hydrozoa.

Hydrozoa medusae also have tetra-radial symmetry, nearly always have a velum (diaphragm used in swimming) attached just inside the bell margin, do not have oral arms, but a much smaller central stalk-like structure, the manubrium, with terminal mouth opening, and are distinguished by the absence of cells in the mesoglea. Hydrozoa show great diversity of lifestyle; some species maintain the polyp form for their entire life and do not form medusae at all (such as Hydra, which is hence not considered a jellyfish), and a few are entirely medusal and have no polyp form.

Staurozoa (stalked jellyfish) are characterized by a medusa form that is generally sessile, oriented upside down and with a stalk emerging from the apex of the "calyx" (bell), which attaches to the substrate. At least some Staurozoa also have a polyp form that alternates with the medusoid portion of the life cycle. Until recently, Staurozoa were classified within the Scyphozoa.

There are over 200 species of Scyphozoa, about 50 species of Staurozoa, about 50 species of Cubozoa, and the Hydrozoa includes about 1000–1500 species that produce medusae, but many more species that do not.

Fossil history

Since jellyfish have no hard parts, fossils are rare. The oldest unambiguous fossil of a free-swimming medusa is Burgessomedusa from the mid Cambrian Burgess Shale of Canada, which is likely either a stem group of box jellyfish (Cubozoa) or Acraspeda (the clade including Staurozoa, Cubozoa, and Scyphozoa). Other claimed records from the Cambrian of China and Utah in the United States are uncertain, and possibly represent ctenophores instead.

Anatomy

The main feature of a true jellyfish is the umbrella-shaped bell. This is a hollow structure consisting of a mass of transparent jelly-like matter known as mesoglea, which forms the hydrostatic skeleton of the animal. 95% or more of the mesogloea consists of water, but it also contains collagen and other fibrous proteins, as well as wandering amoebocytes which can engulf debris and bacteria. The mesogloea is bordered by the epidermis on the outside and the gastrodermis on the inside. The edge of the bell is often divided into rounded lobes known as lappets, which allow the bell to flex. In the gaps or niches between the lappets are dangling rudimentary sense organs known as rhopalia, and the margin of the bell often bears tentacles.

Anatomy of a scyphozoan jellyfish

On the underside of the bell is the manubrium, a stalk-like structure hanging down from the centre, with the mouth, which also functions as the anus, at its tip. There are often four oral arms connected to the manubrium, streaming away into the water below. The mouth opens into the gastrovascular cavity, where digestion takes place and nutrients are absorbed. This is subdivided by four thick septa into a central stomach and four gastric pockets. The four pairs of gonads are attached to the septa, and close to them four septal funnels open to the exterior, perhaps supplying good oxygenation to the gonads. Near the free edges of the septa, gastric filaments extend into the gastric cavity; these are armed with nematocysts and enzyme-producing cells and play a role in subduing and digesting the prey. In some scyphozoans, the gastric cavity is joined to radial canals which branch extensively and may join a marginal ring canal. Cilia in these canals circulate the fluid in a regular direction.

Discharge mechanism of a nematocyst

The box jellyfish is largely similar in structure. It has a squarish, box-like bell. A short pedalium or stalk hangs from each of the four lower corners. One or more long, slender tentacles are attached to each pedalium. The rim of the bell is folded inwards to form a shelf known as a velarium which restricts the bell's aperture and creates a powerful jet when the bell pulsates, allowing box jellyfish to swim faster than true jellyfish. Hydrozoans are also similar, usually with just four tentacles at the edge of the bell, although many hydrozoans are colonial and may not have a free-living medusal stage. In some species, a non-detachable bud known as a gonophore is formed that contains a gonad but is missing many other medusal features such as tentacles and rhopalia. Stalked jellyfish are attached to a solid surface by a basal disk, and resemble a polyp, the oral end of which has partially developed into a medusa with tentacle-bearing lobes and a central manubrium with four-sided mouth.

Most jellyfish do not have specialized systems for osmoregulation, respiration and circulation, and do not have a central nervous system. Nematocysts, which deliver the sting, are located mostly on the tentacles; true jellyfish also have them around the mouth and stomach. Jellyfish do not need a respiratory system because sufficient oxygen diffuses through the epidermis. They have limited control over their movement, but can navigate with the pulsations of the bell-like body; some species are active swimmers most of the time, while others largely drift. The rhopalia contain rudimentary sense organs which are able to detect light, water-borne vibrations, odour and orientation. A loose network of nerves called a "nerve net" is located in the epidermis. Although traditionally thought not to have a central nervous system, nerve net concentration and ganglion-like structures could be considered to constitute one in most species. A jellyfish detects stimuli, and transmits impulses both throughout the nerve net and around a circular nerve ring, to other nerve cells. The rhopalial ganglia contain pacemaker neurones which control swimming rate and direction.

In many species of jellyfish, the rhopalia include ocelli, light-sensitive organs able to tell light from dark. These are generally pigment spot ocelli, which have some of their cells pigmented. The rhopalia are suspended on stalks with heavy crystals at one end, acting like gyroscopes to orient the eyes skyward. Certain jellyfish look upward at the mangrove canopy while making a daily migration from mangrove swamps into the open lagoon, where they feed, and back again.

Box jellyfish have more advanced vision than the other groups. Each individual has 24 eyes, two of which are capable of seeing colour, and four parallel information processing areas that act in competition, supposedly making them one of the few kinds of animal to have a 360-degree view of its environment.

Box jellyfish eye

The study of jellyfish eye evolution is an intermediary to a better understanding of how visual systems evolved on Earth. Jellyfish exhibit immense variation in visual systems ranging from photoreceptive cell patches seen in simple photoreceptive systems to more derived complex eyes seen in box jellyfish. Major topics of jellyfish visual system research (with an emphasis on box jellyfish) include: the evolution of jellyfish vision from simple to complex visual systems), the eye morphology and molecular structures of box jellyfish (including comparisons to vertebrate eyes), and various uses of vision including task-guided behaviors and niche specialization.

Evolution

Experimental evidence for photosensitivity and photoreception in cnidarians antecedes the mid 1900s, and a rich body of research has since covered evolution of visual systems in jellyfish. Jellyfish visual systems range from simple photoreceptive cells to complex image-forming eyes. More ancestral visual systems incorporate extraocular vision (vision without eyes) that encompass numerous receptors dedicated to single-function behaviors. More derived visual systems comprise perception that is capable of multiple task-guided behaviors.

Although they lack a true brain, cnidarian jellyfish have a "ring" nervous system that plays a significant role in motor and sensory activity. This net of nerves is responsible for muscle contraction and movement and culminates the emergence of photosensitive structures. Across Cnidaria, there is large variation in the systems that underlie photosensitivity. Photosensitive structures range from non-specialized groups of cells, to more "conventional" eyes similar to those of vertebrates. The general evolutionary steps to develop complex vision include (from more ancestral to more derived states): non-directional photoreception, directional photoreception, low-resolution vision, and high-resolution vision. Increased habitat and task complexity has favored the high-resolution visual systems common in derived cnidarians such as box jellyfish.

Basal visual systems observed in various cnidarians exhibit photosensitivity representative of a single task or behavior. Extraocular photoreception (a form of non-directional photoreception), is the most basic form of light sensitivity and guides a variety of behaviors among cnidarians. It can function to regulate circadian rhythm (as seen in eyeless hydrozoans) and other light-guided behaviors responsive to the intensity and spectrum of light. Extraocular photoreception can function additionally in positive phototaxis (in planula larvae of hydrozoans), as well as in avoiding harmful amounts of UV radiation via negative phototaxis. Directional photoreception (the ability to perceive direction of incoming light) allows for more complex phototactic responses to light, and likely evolved by means of membrane stacking. The resulting behavioral responses can range from guided spawning events timed by moonlight to shadow responses for potential predator avoidance. Light-guided behaviors are observed in numerous scyphozoans including the common moon jelly, Aurelia aurita, which migrates in response to changes in ambient light and solar position even though they lack proper eyes.

The low-resolution visual system of box jellyfish is more derived than directional photoreception, and thus box jellyfish vision represents the most basic form of true vision in which multiple directional photoreceptors combine to create the first imaging and spatial resolution. This is different from the high-resolution vision that is observed in camera or compound eyes of vertebrates and cephalopods that rely on focusing optics. Critically, the visual systems of box jellyfish are responsible for guiding multiple tasks or behaviors in contrast to less derived visual systems in other jellyfish that guide single behavioral functions. These behaviors include phototaxis based on sunlight (positive) or shadows (negative), obstacle avoidance, and control of swim-pulse rate.

Box jellyfish possess "proper eyes" (similar to vertebrates) that allow them to inhabit environments that lesser derived medusae cannot. In fact, they are considered the only class in the clade Medusozoa that have behaviors necessitating spatial resolution and genuine vision. However, the lens in their eyes are more functionally similar to cup-eyes exhibited in low-resolution organisms, and have very little to no focusing capability. The lack of the ability to focus is due to the focal length exceeding the distance to the retina, thus generating unfocused images and limiting spatial resolution. The visual system is still sufficient for box jellyfish to produce an image to help with tasks such as object avoidance.

Utility as a model organism

Box jellyfish eyes are a visual system that is sophisticated in numerous ways. These intricacies include the considerable variation within the morphology of box jellyfishes' eyes (including their task/behavior specification), and the molecular makeup of their eyes including: photoreceptors, opsins, lenses, and synapses. The comparison of these attributes to more derived visual systems can allow for a further understanding of how the evolution of more derived visual systems may have occurred, and puts into perspective how box jellyfish can play the role as an evolutionary/developmental model for all visual systems.

Characteristics

Box jellyfish visual systems are both diverse and complex, comprising multiple photosystems. There is likely considerable variation in visual properties between species of box jellyfish given the significant inter-species morphological and physiological variation. Eyes tend to differ in size and shape, along with number of receptors (including opsins), and physiology across species of box jellyfish.

Box jellyfish have a series of intricate lensed eyes that are similar to those of more derived multicellular organisms such as vertebrates. Their 24 eyes fit into four different morphological categories. These categories consist of two large, morphologically different medial eyes (a lower and upper lensed eye) containing spherical lenses, a lateral pair of pigment slit eyes, and a lateral pair of pigment pit eyes. The eyes are situated on rhopalia (small sensory structures) which serve sensory functions of the box jellyfish and arise from the cavities of the exumbrella (the surface of the body) on the side of the bells of the jellyfish. The two large eyes are located on the mid-line of the club and are considered complex because they contain lenses. The four remaining eyes lie laterally on either side of each rhopalia and are considered simple. The simple eyes are observed as small invaginated cups of epithelium that have developed pigmentation. The larger of the complex eyes contains a cellular cornea created by a mono ciliated epithelium, cellular lens, homogenous capsule to the lens, vitreous body with prismatic elements, and a retina of pigmented cells. The smaller of the complex eyes is said to be slightly less complex given that it lacks a capsule but otherwise contains the same structure as the larger eye.

Box jellyfish have multiple photosystems that comprise different sets of eyes. Evidence includes immunocytochemical and molecular data that show photopigment differences among the different morphological eye types, and physiological experiments done on box jellyfish to suggest behavioral differences among photosystems. Each individual eye type constitutes photosystems that work collectively to control visually guided behaviors.

Box jellyfish eyes primarily use c-PRCs (ciliary photoreceptor cells) similar to that of vertebrate eyes. These cells undergo phototransduction cascades (process of light absorption by photoreceptors) that are triggered by c-opsins. Available opsin sequences suggest that there are two types of opsins possessed by all cnidarians including an ancient phylogenetic opsin, and a sister ciliary opsin to the c-opsins group. Box jellyfish could have both ciliary and cnidops (cnidarian opsins), which is something not previously believed to appear in the same retina. Nevertheless, it is not entirely evident whether cnidarians possess multiple opsins that are capable of having distinctive spectral sensitivities.

Comparison with other organisms

Comparative research on genetic and molecular makeup of box jellyfishes' eyes versus more derived eyes seen in vertebrates and cephalopods focuses on: lenses and crystallin composition, synapses, and Pax genes and their implied evidence for shared primordial (ancestral) genes in eye evolution.

Box jellyfish eyes are said to be an evolutionary/developmental model of all eyes based on their evolutionary recruitment of crystallins and Pax genes. Research done on box jellyfish including Tripedalia cystophora has suggested that they possess a single Pax gene, PaxB. PaxB functions by binding to crystallin promoters and activating them. PaxB in situ hybridization resulted in PaxB expression in the lens, retina, and statocysts. These results and the rejection of the prior hypothesis that Pax6 was an ancestral Pax gene in eyes has led to the conclusion that PaxB was a primordial gene in eye evolution, and that the eyes of all organisms likely share a common ancestor.

The lens structure of box jellyfish appears very similar to those of other organisms, but the crystallins are distinct in both function and appearance. Weak reactions were seen within the sera and there were very weak sequence similarities within the crystallins among vertebrate and invertebrate lenses. This is likely due to differences in lower molecular weight proteins and the subsequent lack of immunological reactions with antisera that other organisms' lenses exhibit.

All four of the visual systems of box jellyfish species investigated with detail (Carybdea marsupialis, Chiropsalmus quadrumanus, Tamoya haplonema and Tripedalia cystophora) have invaginated synapses, but only in the upper and lower lensed eyes. Different densities were found between the upper and lower lenses, and between species. Four types of chemical synapses have been discovered within the rhopalia which could help in understanding neural organization including: clear unidirectional, dense-core unidirectional, clear bidirectional, and clear and dense-core bidirectional. The synapses of the lensed eyes could be useful as markers to learn more about the neural circuit in box jellyfish retinal areas.

Evolution as a response to natural stimuli

The primary adaptive responses to environmental variation observed in box jellyfish eyes include pupillary constriction speeds in response to light environments, as well as photoreceptor tuning and lens adaptations to better respond to shifts between light environments and darkness. Interestingly, some box jellyfish species' eyes appear to have evolved more focused vision in response to their habitat.

Pupillary contraction appears to have evolved in response to variation in the light environment across ecological niches across three species of box jellyfish (Chironex fleckeri, Chiropsella bronzie, and Carukia barnesi). Behavioral studies suggest that faster pupil contraction rates allow for greater object avoidance, and in fact, species with more complex habitats exhibit faster rates. Ch. bronzie inhabit shallow beach fronts that have low visibility and very few obstacles, thus, faster pupil contraction in response to objects in their environment is not important. Ca. barnesi and Ch. fleckeri are found in more three-dimensionally complex environments like mangroves with an abundance of natural obstacles, where faster pupil contraction is more adaptive. Behavioral studies support the idea that faster pupillary contraction rates assist with obstacle avoidance as well as depth adjustments in response to differing light intensities.

Light/dark adaptation via pupillary light reflexes is an additional form of an evolutionary response to the light environment. This relates to the pupil's response to shifts between light intensity (generally from sunlight to darkness). In the process of light/dark adaptation, the upper and lower lens eyes of different box jellyfish species vary in specific function. The lower lens-eyes contain pigmented photoreceptors and long pigment cells with dark pigments that migrate on light/dark adaptation, while the upper-lens eyes play a concentrated role in light direction and phototaxis given that they face upward towards the water surface (towards the sun or moon). The upper lens of Ch. bronzie does not exhibit any considerable optical power while Tr. cystophora (a box jellyfish species that tends to live in mangroves) does. The ability to use light to visually guide behavior is not of as much importance to Ch. bronzie as it is to species in more obstacle-filled environments. Differences in visually guided behavior serve as evidence that species that share the same number and structure of eyes can exhibit differences in how they control behavior.

Largest and smallest

Jellyfish range from about one millimeter in bell height and diameter, to nearly 2 metres (6+1⁄2 ft) in bell height and diameter; the tentacles and mouth parts usually extend beyond this bell dimension.

The smallest jellyfish are the peculiar creeping jellyfish in the genera Staurocladia and Eleutheria, which have bell disks from 0.5 millimetres (1⁄32 in) to a few millimeters in diameter, with short tentacles that extend out beyond this, which these jellyfish use to move across the surface of seaweed or the bottoms of rocky pools; many of these tiny creeping jellyfish cannot be seen in the field without a hand lens or microscope. They can reproduce asexually by fission (splitting in half). Other very small jellyfish, which have bells about one millimeter, are the hydromedusae of many species that have just been released from their parent polyps; some of these live only a few minutes before shedding their gametes in the plankton and then dying, while others will grow in the plankton for weeks or months. The hydromedusae Cladonema radiatum and Cladonema californicum are also very small, living for months, yet never growing beyond a few mm in bell height and diameter.

The lion's mane jellyfish, Cyanea capillata, was long-cited as the largest jellyfish, and arguably the longest animal in the world, with fine, thread-like tentacles that may extend up to 36.5 m (119 ft 9 in) long (though most are nowhere near that large). They have a moderately painful, but rarely fatal, sting. The increasingly common giant Nomura's jellyfish, Nemopilema nomurai, found in some, but not all years in the waters of Japan, Korea and China in summer and autumn is another candidate for "largest jellyfish", in terms of diameter and weight, since the largest Nomura's jellyfish in late autumn can reach 2 m (6 ft 7 in) in bell (body) diameter and about 200 kg (440 lb) in weight, with average specimens frequently reaching 0.9 m (2 ft 11 in) in bell diameter and about 150 kg (330 lb) in weight. The large bell mass of the giant Nomura's jellyfish can dwarf a diver and is nearly always much greater than the Lion's Mane, whose bell diameter can reach 1 m (3 ft 3 in).

The rarely encountered deep-sea jellyfish Stygiomedusa gigantea is another candidate for "largest jellyfish", with its thick, massive bell up to 100 cm (3 ft 3 in) wide, and four thick, "strap-like" oral arms extending up to 6 m (19+1⁄2 ft) in length, very different from the typical fine, threadlike tentacles that rim the umbrella of more-typical-looking jellyfish, including the Lion's Mane.

Desmonema glaciale, which lives in the Antarctic region, can reach a very large size (several meters). Purple-striped jelly (Chrysaora colorata) can also be extremely long (up to 15 feet).

Life history and behavior

Life cycle

Jellyfish have a complex life cycle which includes both sexual and asexual phases, with the medusa being the sexual stage in most instances. Sperm fertilize eggs, which develop into larval planulae, become polyps, bud into ephyrae and then transform into adult medusae. In some species certain stages may be skipped.

Upon reaching adult size, jellyfish spawn regularly if there is a sufficient supply of food. In most species, spawning is controlled by light, with all individuals spawning at about the same time of day; in many instances this is at dawn or dusk. Jellyfish are usually either male or female (with occasional hermaphrodites). In most cases, adults release sperm and eggs into the surrounding water, where the unprotected eggs are fertilized and develop into larvae. In a few species, the sperm swim into the female's mouth, fertilizing the eggs within her body, where they remain during early development stages. In moon jellies, the eggs lodge in pits on the oral arms, which form a temporary brood chamber for the developing planula larvae.

The planula is a small larva covered with cilia. When sufficiently developed, it settles onto a firm surface and develops into a polyp. The polyp generally consists of a small stalk topped by a mouth that is ringed by upward-facing tentacles. The polyps resemble those of closely related anthozoans, such as sea anemones and corals. The jellyfish polyp may be sessile, living on the bottom, boat hulls or other substrates, or it may be free-floating or attached to tiny bits of free-living plankton or rarely, fish or other invertebrates. Polyps may be solitary or colonial. Most polyps are only millimetres in diameter and feed continuously. The polyp stage may last for years.

After an interval and stimulated by seasonal or hormonal changes, the polyp may begin reproducing asexually by budding and, in the Scyphozoa, is called a segmenting polyp, or a scyphistoma. Budding produces more scyphistomae and also ephyrae. Budding sites vary by species; from the tentacle bulbs, the manubrium (above the mouth), or the gonads of hydromedusae. In a process known as strobilation, the polyp's tentacles are reabsorbed and the body starts to narrow, forming transverse constrictions, in several places near the upper extremity of the polyp. These deepen as the constriction sites migrate down the body, and separate segments known as ephyra detach. These are free-swimming precursors of the adult medusa stage, which is the life stage that is typically identified as a jellyfish. The ephyrae, usually only a millimeter or two across initially, swim away from the polyp and grow. Limnomedusae polyps can asexually produce a creeping frustule larval form, which crawls away before developing into another polyp. A few species can produce new medusae by budding directly from the medusan stage. Some hydromedusae reproduce by fission.

Lifespan

Little is known of the life histories of many jellyfish as the places on the seabed where the benthic forms of those species live have not been found. However, an asexually reproducing strobila form can sometimes live for several years, producing new medusae (ephyra larvae) each year.

An unusual species, Turritopsis dohrnii, formerly classified as Turritopsis nutricula, might be effectively immortal because of its ability under certain circumstances to transform from medusa back to the polyp stage, thereby escaping the death that typically awaits medusae post-reproduction if they have not otherwise been eaten by some other organism. So far this reversal has been observed only in the laboratory.

Locomotion

Jellyfish locomotion is highly efficient. Muscles in the jellylike bell contract, setting up a start vortex and propelling the animal. When the contraction ends, the bell recoils elastically, creating a stop vortex with no extra energy input.

Using the moon jelly Aurelia aurita as an example, jellyfish have been shown to be the most energy-efficient swimmers of all animals. They move through the water by radially expanding and contracting their bell-shaped bodies to push water behind them. They pause between the contraction and expansion phases to create two vortex rings. Muscles are used for the contraction of the body, which creates the first vortex and pushes the animal forward, but the mesoglea is so elastic that the expansion is powered exclusively by relaxing the bell, which releases the energy stored from the contraction. Meanwhile, the second vortex ring starts to spin faster, sucking water into the bell and pushing against the centre of the body, giving a secondary and "free" boost forward. The mechanism, called passive energy recapture, only works in relatively small jellyfish moving at low speeds, allowing the animal to travel 30 percent farther on each swimming cycle. Jellyfish achieved a 48 percent lower cost of transport (food and oxygen intake versus energy spent in movement) than other animals in similar studies. One reason for this is that most of the gelatinous tissue of the bell is inactive, using no energy during swimming.

Ecology

Diet

Jellyfish are, like other cnidarians, generally carnivorous (or parasitic), feeding on planktonic organisms, crustaceans, small fish, fish eggs and larvae, and other jellyfish, ingesting food and voiding undigested waste through the mouth. They hunt passively using their tentacles as drift lines, or sink through the water with their tentacles spread widely; the tentacles, which contain nematocysts to stun or kill the prey, may then flex to help bring it to the mouth. Their swimming technique also helps them to capture prey; when their bell expands it sucks in water which brings more potential prey within reach of the tentacles.

A few species such as Aglaura hemistoma are omnivorous, feeding on microplankton which is a mixture of zooplankton and phytoplankton (microscopic plants) such as dinoflagellates. Others harbour mutualistic algae (Zooxanthellae) in their tissues; the spotted jellyfish (Mastigias papua) is typical of these, deriving part of its nutrition from the products of photosynthesis, and part from captured zooplankton. The upside-down jellyfish (Cassiopea andromeda) also has a symbiotic relationship with microalgae, but captures tiny animals to supplement their diet. This is done by releasing tiny balls of living cells composed of mesoglea. These use cilia to drive them through water and stinging cells which stun the prey. The blobs also seems to have digestive capabilities.

Predation

Other species of jellyfish are among the most common and important jellyfish predators. Sea anemones may eat jellyfish that drift into their range. Other predators include tunas, sharks, swordfish, sea turtles and penguins. Jellyfish washed up on the beach are consumed by foxes, other terrestrial mammals and birds. In general however, few animals prey on jellyfish; they can broadly be considered to be top predators in the food chain. Once jellyfish have become dominant in an ecosystem, for example through overfishing which removes predators of jellyfish larvae, there may be no obvious way for the previous balance to be restored: they eat fish eggs and juvenile fish, and compete with fish for food, preventing fish stocks from recovering.

Symbiosis

Some small fish are immune to the stings of the jellyfish and live among the tentacles, serving as bait in a fish trap; they are safe from potential predators and are able to share the fish caught by the jellyfish. The cannonball jellyfish has a symbiotic relationship with ten different species of fish, and with the longnose spider crab, which lives inside the bell, sharing the jellyfish's food and nibbling its tissues.

Main article: Jellyfish bloom

Jellyfish form large masses or blooms in certain environmental conditions of ocean currents, nutrients, sunshine, temperature, season, prey availability, reduced predation and oxygen concentration. Currents collect jellyfish together, especially in years with unusually high populations. Jellyfish can detect marine currents and swim against the current to congregate in blooms. Jellyfish are better able to survive in nutrient-rich, oxygen-poor water than competitors, and thus can feast on plankton without competition. Jellyfish may also benefit from saltier waters, as saltier waters contain more iodine, which is necessary for polyps to turn into jellyfish. Rising sea temperatures caused by climate change may also contribute to jellyfish blooms, because many species of jellyfish are able to survive in warmer waters. Increased nutrients from agricultural or urban runoff with nutrients including nitrogen and phosphorus compounds increase the growth of phytoplankton, causing eutrophication and algal blooms. When the phytoplankton die, they may create dead zones, so-called because they are hypoxic (low in oxygen). This in turn kills fish and other animals, but not jellyfish, allowing them to bloom. Jellyfish populations may be expanding globally as a result of land runoff and overfishing of their natural predators. Jellyfish are well placed to benefit from disturbance of marine ecosystems. They reproduce rapidly; they prey upon many species, while few species prey on them; and they feed via touch rather than visually, so they can feed effectively at night and in turbid waters. It may be difficult for fish stocks to re-establish themselves in marine ecosystems once they have become dominated by jellyfish, because jellyfish feed on plankton, which includes fish eggs and larvae.

As suspected at the turn of this century, jellyfish blooms are increasing in frequency. Between 2013 and 2020 the Mediterranean Science Commission monitored on a weekly basis the frequency of such outbreaks in coastal waters from Morocco to the Black Sea, revealing a relatively high frequency of these blooms nearly all year round, with peaks observed from March to July and often again in the autumn. The blooms are caused by different jellyfish species, depending on their localisation within the Basin: one observes a clear dominance of Pelagia noctiluca and Velella velella outbreaks in the western Mediterranean, of Rhizostoma pulmo and Rhopilema nomadica outbreaks in the eastern Mediterranean, and of Aurelia aurita and Mnemiopsis leidyi outbreaks in the Black Sea.

Some jellyfish populations that have shown clear increases in the past few decades are invasive species, newly arrived from other habitats: examples include the Black Sea, Caspian Sea, Baltic Sea, central and eastern Mediterranean, Hawaii, and tropical and subtropical parts of the West Atlantic (including the Caribbean, Gulf of Mexico and Brazil).

Jellyfish blooms can have significant impact on community structure. Some carnivorous jellyfish species prey on zooplankton while others graze on primary producers. Reductions in zooplankton and ichthyoplankton due to a jellyfish bloom can ripple through the trophic levels. High-density jellyfish populations can outcompete other predators and reduce fish recruitment. Increased grazing on primary producers by jellyfish can also interrupt energy transfer to higher trophic levels.

During blooms, jellyfish significantly alter the nutrient availability in their environment. Blooms require large amounts of available organic nutrients in the water column to grow, limiting availability for other organisms. Some jellyfish have a symbiotic relationship with single-celled dinoflagellates, allowing them to assimilate inorganic carbon, phosphorus, and nitrogen creating competition for phytoplankton. Their large biomass makes them an important source of dissolved and particulate organic matter for microbial communities through excretion, mucus production, and decomposition. The microbes break down the organic matter into inorganic ammonium and phosphate. However, the low carbon availability shifts the process from production to respiration creating low oxygen areas making the dissolved inorganic nitrogen and phosphorus largely unavailable for primary production.

These blooms have very real impacts on industries. Jellyfish can outcompete fish by utilizing open niches in over-fished fisheries. Catch of jellyfish can strain fishing gear and lead to expenses relating to damaged gear. Power plants have been shut down due to jellyfish blocking the flow of cooling water. Blooms have also been harmful for tourism, causing a rise in stings and sometimes the closure of beaches.

Jellyfish form a component of jelly-falls, events where gelatinous zooplankton fall to the seafloor, providing food for the benthic organisms there. In temperate and subpolar regions, jelly-falls usually follow immediately after a bloom.

Habitats

Most jellyfish are marine animals, although a few hydromedusae inhabit freshwater. The best known freshwater example is the cosmopolitan hydrozoan jellyfish, Craspedacusta sowerbii. It is less than an inch (2.5 cm) in diameter, colorless and does not sting. Some jellyfish populations have become restricted to coastal saltwater lakes, such as Jellyfish Lake in Palau. Jellyfish Lake is a marine lake where millions of golden jellyfish (Mastigias spp.) migrate horizontally across the lake daily.

Although most jellyfish live well off the ocean floor and form part of the plankton, a few species are closely associated with the bottom for much of their lives and can be considered benthic. The upside-down jellyfish in the genus Cassiopea typically lie on the bottom of shallow lagoons where they sometimes pulsate gently with their umbrella top facing down. Even some deep-sea species of hydromedusae and scyphomedusae are usually collected on or near the bottom. All of the stauromedusae are found attached to either seaweed or rocky or other firm material on the bottom.

Some species explicitly adapt to tidal flux. In Roscoe Bay, jellyfish ride the current at ebb tide until they hit a gravel bar, and then descend below the current. They remain in still waters until the tide rises, ascending and allowing it to sweep them back into the bay. They also actively avoid fresh water from mountain snowmelt, diving until they find enough salt.

Parasites

Jellyfish are hosts to a wide variety of parasitic organisms. They act as intermediate hosts of endoparasitic helminths, with the infection being transferred to the definitive host fish after predation. Some digenean trematodes, especially species in the family Lepocreadiidae, use jellyfish as their second intermediate hosts. Fish become infected by the trematodes when they feed on infected jellyfish.

Relation to humans

Jellyfish have long been eaten in some parts of the world. Fisheries have begun harvesting the American cannonball jellyfish, Stomolophus meleagris, along the southern Atlantic coast of the United States and in the Gulf of Mexico for export to Asia.

Jellyfish are also harvested for their collagen, which is being investigated for use in a variety of applications including the treatment of rheumatoid arthritis.

Aquaculture and fisheries of other species often suffer severe losses – and so losses of productivity – due to jellyfish.

Products

Main article: Jellyfish as food

In some countries, including China, Japan, and Korea, jellyfish are a delicacy. The jellyfish is dried to prevent spoiling. Only some 12 species of scyphozoan jellyfish belonging to the order Rhizostomeae are harvested for food, mostly in southeast Asia. Rhizostomes, especially Rhopilema esculentum in China (海蜇 hǎizhé, 'sea stingers') and Stomolophus meleagris (cannonball jellyfish) in the United States, are favored because of their larger and more rigid bodies and because their toxins are harmless to humans.

Traditional processing methods, carried out by a jellyfish master, involve a 20- to 40-day multi-phase procedure in which, after removing the gonads and mucous membranes, the umbrella and oral arms are treated with a mixture of table salt and alum, and compressed. Processing makes the jellyfish drier and more acidic, producing a crisp texture. Jellyfish prepared this way retain 7–10% of their original weight, and the processed product consists of approximately 94% water and 6% protein. Freshly processed jellyfish has a white, creamy color and turns yellow or brown during prolonged storage.

In China, processed jellyfish are desalted by soaking in water overnight and eaten cooked or raw. The dish is often served shredded with a dressing of oil, soy sauce, vinegar and sugar, or as a salad with vegetables. In Japan, cured jellyfish are rinsed, cut into strips and served with vinegar as an appetizer. Desalted, ready-to-eat products are also available.

Biotechnology

The hydromedusa Aequorea victoria was the source of green fluorescent protein, studied for its role in bioluminescence and later for use as a marker in genetic engineering.

Pliny the Elder reported in his Natural History that the slime of the jellyfish "Pulmo marinus" produced light when rubbed on a walking stick.

In 1961, Osamu Shimomura extracted green fluorescent protein (GFP) and another bioluminescent protein, called aequorin, from the large and abundant hydromedusa Aequorea victoria, while studying photoproteins that cause bioluminescence in this species. Three decades later, Douglas Prasher sequenced and cloned the gene for GFP. Martin Chalfie figured out how to use GFP as a fluorescent marker of genes inserted into other cells or organisms. Roger Tsien later chemically manipulated GFP to produce other fluorescent colors to use as markers. In 2008, Shimomura, Chalfie and Tsien won the Nobel Prize in Chemistry for their work with GFP. Man-made GFP became widely used as a fluorescent tag to show which cells or tissues express specific genes. The genetic engineering technique fuses the gene of interest to the GFP gene. The fused DNA is then put into a cell, to generate either a cell line or (via IVF techniques) an entire animal bearing the gene. In the cell or animal, the artificial gene turns on in the same tissues and the same time as the normal gene, making a fusion of the normal protein with GFP attached to the end, illuminating the animal or cell reveals what tissues express that protein—or at what stage of development. The fluorescence shows where the gene is expressed.

Aquarium display

Jellyfish are displayed in many public aquariums. Often the tank's background is blue and the animals are illuminated by side light, increasing the contrast between the animal and the background. In natural conditions, many jellies are so transparent that they are nearly invisible. Jellyfish are not adapted to closed spaces. They depend on currents to transport them from place to place. Professional exhibits as in the Monterey Bay Aquarium feature precise water flows, typically in circular tanks to avoid trapping specimens in corners. The outflow is spread out over a large surface area and the inflow enters as a sheet of water in front of the outflow, so the jellyfish do not get sucked into it. As of 2009, jellyfish were becoming popular in home aquariums, where they require similar equipment.

Stings

Jellyfish are armed with nematocysts, a type of specialized stinging cell. Contact with a jellyfish tentacle can trigger millions of nematocysts to pierce the skin and inject venom, but only some species' venom causes an adverse reaction in humans. In a study published in Communications Biology, researchers found a jellyfish species called Cassiopea xamachana which when triggered will release tiny balls of cells that swim around the jellyfish stinging everything in their path. Researchers described these as "self-propelling microscopic grenades" and named them cassiosomes.

The effects of stings range from mild discomfort to extreme pain and death. Most jellyfish stings are not deadly, but stings of some box jellyfish (Irukandji jellyfish), such as the sea wasp, can be deadly. Stings may cause anaphylaxis (a form of shock), which can be fatal. Jellyfish kill 20 to 40 people a year in the Philippines alone. In 2006 the Spanish Red Cross treated 19,000 stung swimmers along the Costa Brava.

Vinegar (3–10% aqueous acetic acid) may help with box jellyfish stings but not the stings of the Portuguese man o' war. Clearing the area of jelly and tentacles reduces nematocyst firing. Scraping the affected skin, such as with the edge of a credit card, may remove remaining nematocysts. Once the skin has been cleaned of nematocysts, hydrocortisone cream applied locally reduces pain and inflammation. Antihistamines may help to control itching. Immunobased antivenins are used for serious box jellyfish stings.

In Elba Island and Corsica dittrichia viscosa is now used by residents and tourists to heal stings from jellyfish, bees and wasps pressing fresh leaves on the skin with quick results.

Mechanical issues

Jellyfish in large quantities can fill and split fishing nets and crush captured fish. They can clog cooling equipment, having disabled power stations in several countries; jellyfish caused a cascading blackout in the Philippines in 1999, as well as damaging the Diablo Canyon Power Plant in California in 2008. They can also stop desalination plants and ships' engines.

Winter Sparrow by mahar15

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We have a couple of sparrows that are missing pigmentation in their feathers. They seem to blend into the snowy landscape.