A bug I found last week, quite possibly only the 2nd reported case of the melanic form of Gorse shieldbug (Piezodorus lituratus) ever recorded anywhere! Melanism is a condition that turns animals black and is related to pigmentation like albinism and leucism.

El Greco The Martydom of St Maurice and the Theban (detail) oil on canvas 14'8" x 9'11" 1580-82

Painted for the Basilica of El Escorial but now in the picture gallery.El Greco (medieval Castilian for "the Greek") is the popular name for Δομήνικος Θεοτοκόπουλος Dominikos Theotokópoulos (1541, Fodele or Candia (now Heraklion), Crete – April 7, 1614, Toledo, Spain), a Cretan Greek painter, sculptor, and architect of the Spanish Renaissance. He is best known for tortuously elongated figures and often fantastic or phantasmagorical pigmentation.

He was a painter in Crete and first trained as an icon painter. At the age of twenty-six he journeyed to Venice where he is said to have studied western-style art under Titian and Tintoretto. He spent almost two years there before moving to Rome. In Rome, El Greco was influenced by the mannerist style as practiced by followers of Michelangelo. Mannerism appealed to him because of the talent and intelligence and virtuosity required to create the images. In 1577 he emigrated to Toledo — at the time the religious capital of Spain — where he produced his mature works. The Christian doctrines greatly influenced his life and his artwork, leading him to a successful career as a painter of altarpieces and portraits. Some works include The Annunciation, Laocoon, and The Repentent Peter. Many of El Greco's works are on display at Madrid's Museo del Prado; however others can be found in other places such as The Greco Museum and House in Toledo, The Museum of Santa Cruz in the same city, The Metropolitan Museum of Art in New York and The National Gallery of Art in Washington D.C.

   

Maize (/meɪz/ MAYZ; Zea mays subsp. mays, from Spanish: maíz after Taino: mahiz), also known as corn (North American and Australian English), is a cereal grain first domesticated by indigenous peoples in southern Mexico about 10,000 years ago. The leafy stalk of the plant produces pollen inflorescences and separate ovuliferous inflorescences called ears that yield kernels or seeds, which are fruits.

 

Maize has become a staple food in many parts of the world, with the total production of maize surpassing that of wheat or rice. In addition to being consumed directly by humans (often in the form of masa), maize is also used for corn ethanol, animal feed and other maize products, such as corn starch and corn syrup. The six major types of maize are dent corn, flint corn, pod corn, popcorn, flour corn, and sweet corn.[5] Sugar-rich varieties called sweet corn are usually grown for human consumption as kernels, while field corn varieties are used for animal feed, various corn-based human food uses (including grinding into cornmeal or masa, pressing into corn oil, and fermentation and distillation into alcoholic beverages like bourbon whiskey), and as chemical feedstocks. Maize is also used in making ethanol and other biofuels.

 

Maize is widely cultivated throughout the world, and a greater weight of maize is produced each year than any other grain. In 2014, total world production was 1.04 billion tonnes. Maize is the most widely grown grain crop throughout the Americas, with 361 million metric tons grown in the United States alone in 2014. Genetically modified maize made up 85% of the maize planted in the United States in 2009. Subsidies in the United States help to account for its high level of cultivation of maize and its position as the largest producer in the world.

 

HISTORY

PRE-COLUMBIAN DEVELOPMENT

Maize is a cultigen; human intervention is required for it to propagate. Whether or not the kernels fall off the cob on their own is a key piece of evidence used in archaeology to distinguish domesticated maize from its naturally-propagating teosinte ancestor. Genetic evidence can also be used to determine when various lineages split.

 

Most historians believe maize was domesticated in the Tehuacán Valley of Mexico. Recent research in the early 21st century has modified this view somewhat; scholars now indicate the adjacent Balsas River Valley of south-central Mexico as the center of domestication.

 

An influential 2002 study by Matsuoka et al. has demonstrated that, rather than the multiple independent domestications model, all maize arose from a single domestication in southern Mexico about 9,000 years ago. The study also demonstrated that the oldest surviving maize types are those of the Mexican highlands. Later, maize spread from this region over the Americas along two major paths. This is consistent with a model based on the archaeological record suggesting that maize diversified in the highlands of Mexico before spreading to the lowlands.

 

Archaeologist Dolores Piperno has said:

 

A large corpus of data indicates that [maize] was dispersed into lower Central America by 7600 BP [5600 BC] and had moved into the inter-Andean valleys of Colombia between 7000 and 6000 BP [5000–4000 BC].

— Dolores Piperno, The Origins of Plant Cultivation and Domestication in the New World Tropics: Patterns, Process, and New Developments

 

Since then, even earlier dates have been published.

 

According to a genetic study by Embrapa, corn cultivation was introduced in South America from Mexico, in two great waves: the first, more than 6000 years ago, spread through the Andes. Evidence of cultivation in Peru has been found dating to about 6700 years ago. The second wave, about 2000 years ago, through the lowlands of South America.

 

The earliest maize plants grew only small, 25-millimetre-long (1 in) corn cobs, and only one per plant. In Jackson Spielvogel's view, many centuries of artificial selection (rather than the current view that maize was exploited by interplanting with teosinte) by the indigenous people of the Americas resulted in the development of maize plants capable of growing several cobs per plant, which were usually several centimetres/inches long each. The Olmec and Maya cultivated maize in numerous varieties throughout Mesoamerica; they cooked, ground and processed it through nixtamalization. It was believed that beginning about 2500 BC, the crop spread through much of the Americas. Research of the 21st century has established even earlier dates. The region developed a trade network based on surplus and varieties of maize crops.

 

Mapuches of south-central Chile cultivated maize along with quinoa and potatoes in pre-Hispanic times; however, potato was the staple food of most Mapuches, "specially in the southern and coastal [Mapuche] territories where maize did not reach maturity". Before the expansion of the Inca Empire maize was traded and transported as far south as 40°19' S in Melinquina, Lácar Department. In that location maize remains were found inside pottery dated to 730 ± 80 BP and 920 ± 60 BP. Probably this maize was brought across the Andes from Chile. The presence of maize in Guaitecas Archipelago (43°55' S), the southernmost outpost of pre-Hispanic agriculture, is reported by early Spanish explorers. However the Spanish may have misidentified the plant.

 

COLUMBIAN EXCHANGE

After the arrival of Europeans in 1492, Spanish settlers consumed maize, and explorers and traders carried it back to Europe and introduced it to other countries. Spanish settlers far preferred wheat bread to maize, cassava, or potatoes. Maize flour could not be substituted for wheat for communion bread, since in Christian belief only wheat could undergo transubstantiation and be transformed into the body of Christ. Some Spaniards worried that by eating indigenous foods, which they did not consider nutritious, they would weaken and risk turning into Indians. "In the view of Europeans, it was the food they ate, even more than the environment in which they lived, that gave Amerindians and Spaniards both their distinctive physical characteristics and their characteristic personalities." Despite these worries, Spaniards did consume maize. Archeological evidence from Florida sites indicate they cultivated it as well.

 

Maize spread to the rest of the world because of its ability to grow in diverse climates. It was cultivated in Spain just a few decades after Columbus's voyages and then spread to Italy, West Africa and elsewhere. Widespread cultivation most likely began in southern Spain in 1525, after which it quickly spread to the rest of the Spanish Empire including its territories in Italy (and, from there, to other Italian states). Maize had many advantages over wheat and barley; it yielded two and a half times the food energy per unit cultivated area, could be harvested in successive years from the same plot of land, and grew in wildly varying altitudes and climates, from relatively dry regions with only 250 mm (10 in) of annual rainfall to damp regions with over 5,000 mm (200 in). By the 17th century it was a common peasant food in Southwestern Europe, including Portugal, Spain, southern France, and Italy. By the 18th century, it was the chief food of the southern French and Italian peasantry, especially in the form of polenta in Italy.

Names

 

The word maize derives from the Spanish form of the indigenous Taíno word for the plant, mahiz. It is known by other names around the world.

 

The word "corn" outside the US, Canada, Australia, and New Zealand refers to any cereal crop, its meaning understood to vary geographically to refer to the local staple. In the United States,[30] Canada, Australia, and New Zealand, corn primarily means maize; this usage started as a shortening of "Indian corn". "Indian corn" primarily means maize (the staple grain of indigenous Americans), but can refer more specifically to multicolored "flint corn" used for decoration.

 

In places outside the US, Canada, Australia, and New Zealand, corn often refers to maize in culinary contexts. The narrower meaning is usually indicated by some additional word, as in sweet corn, sweetcorn, corn on the cob, baby corn, the puffed confection known as popcorn and the breakfast cereal known as corn flakes.

 

In Southern Africa, maize is commonly called mielie (Afrikaans) or mealie (English), words derived from the Portuguese word for maize, milho.

 

Maize is preferred in formal, scientific, and international usage because it refers specifically to this one grain, unlike corn, which has a complex variety of meanings that vary by context and geographic region. Maize is used by agricultural bodies and research institutes such as the FAO and CSIRO. National agricultural and industry associations often include the word maize in their name even in English-speaking countries where the local, informal word is something other than maize; for example, the Maize Association of Australia, the Indian Maize Development Association, the Kenya Maize Consortium and Maize Breeders Network, the National Maize Association of Nigeria, the Zimbabwe Seed Maize Association.

 

STRUCTURE AND PHYSIOLOGY

The maize plant is often 3 m (10 ft) in height, though some natural strains can grow 13 m (43 ft). The stem is commonly composed of 20 internodes of 18 cm (7 in) length. The leaves arise from the nodes, alternately on opposite sides on the stalk. A leaf, which grows from each node, is generally 9 cm (3+1⁄2 in) in width and 120 cm (3 ft 11 in) in length.

 

Ears develop above a few of the leaves in the midsection of the plant, between the stem and leaf sheath, elongating by around 3 mm (1⁄8 in) per day, to a length of 18 cm (7 in) with 60 cm (24 in) being the maximum alleged in the subspecies. They are female inflorescences, tightly enveloped by several layers of ear leaves commonly called husks. Certain varieties of maize have been bred to produce many additional developed ears. These are the source of the "baby corn" used as a vegetable in Asian cuisine.

 

The apex of the stem ends in the tassel, an inflorescence of male flowers. When the tassel is mature and conditions are suitably warm and dry, anthers on the tassel dehisce and release pollen. Maize pollen is anemophilous (dispersed by wind), and because of its large settling velocity, most pollen falls within a few meters of the tassel.

 

Elongated stigmas, called silks, emerge from the whorl of husk leaves at the end of the ear. They are often pale yellow and 18 cm (7 in) in length, like tufts of hair in appearance. At the end of each is a carpel, which may develop into a "kernel" if fertilized by a pollen grain. The pericarp of the fruit is fused with the seed coat referred to as "caryopsis", typical of the grasses, and the entire kernel is often referred to as the "seed". The cob is close to a multiple fruit in structure, except that the individual fruits (the kernels) never fuse into a single mass. The grains are about the size of peas, and adhere in regular rows around a white, pithy substance, which forms the ear. The maximum size of kernels is reputedly 2.5 cm (1 in). An ear commonly holds 600 kernels. They are of various colors: blackish, bluish-gray, purple, green, red, white and yellow. When ground into flour, maize yields more flour with much less bran than wheat does. It lacks the protein gluten of wheat and, therefore, makes baked goods with poor rising capability. A genetic variant that accumulates more sugar and less starch in the ear is consumed as a vegetable and is called sweet corn. Young ears can be consumed raw, with the cob and silk, but as the plant matures (usually during the summer months), the cob becomes tougher and the silk dries to inedibility. By the end of the growing season, the kernels dry out and become difficult to chew without cooking them tender first in boiling water.

 

Planting density affects multiple aspects of maize. Modern farming techniques in developed countries usually rely on dense planting, which produces one ear per stalk. Stands of silage maize are yet denser,[citation needed] and achieve a lower percentage of ears and more plant matter.

 

Maize is a facultative short-day plant and flowers in a certain number of growing degree days > 10 °C (50 °F) in the environment to which it is adapted. The magnitude of the influence that long nights have on the number of days that must pass before maize flowers is genetically prescribed and regulated by the phytochrome system.

Photoperiodicity can be eccentric in tropical cultivars such that the long days characteristic of higher latitudes allow the plants to grow so tall that they do not have enough time to produce seed before being killed by frost. These attributes, however, may prove useful in using tropical maize for biofuels.

 

Immature maize shoots accumulate a powerful antibiotic substance, 2,4-dihydroxy-7-methoxy-1,4-benzoxazin-3-one (DIMBOA). DIMBOA is a member of a group of hydroxamic acids (also known as benzoxazinoids) that serve as a natural defense against a wide range of pests, including insects, pathogenic fungi and bacteria. DIMBOA is also found in related grasses, particularly wheat. A maize mutant (bx) lacking DIMBOA is highly susceptible to attack by aphids and fungi. DIMBOA is also responsible for the relative resistance of immature maize to the European corn borer (family Crambidae). As maize matures, DIMBOA levels and resistance to the corn borer decline.

 

Because of its shallow roots, maize is susceptible to droughts, intolerant of nutrient-deficient soils, and prone to be uprooted by severe winds.

 

While yellow maizes derive their color from lutein and zeaxanthin, in red-colored maizes, the kernel coloration is due to anthocyanins and phlobaphenes. These latter substances are synthesized in the flavonoids synthetic pathway from polymerization of flavan-4-ols by the expression of maize pericarp color1 (p1) gene which encodes an R2R3 myb-like transcriptional activator of the A1 gene encoding for the dihydroflavonol 4-reductase (reducing dihydroflavonols into flavan-4-ols) while another gene (Suppressor of Pericarp Pigmentation 1 or SPP1) acts as a suppressor. The p1 gene encodes an Myb-homologous transcriptional activator of genes required for biosynthesis of red phlobaphene pigments, while the P1-wr allele specifies colorless kernel pericarp and red cobs, and unstable factor for orange1 (Ufo1) modifies P1-wr expression to confer pigmentation in kernel pericarp, as well as vegetative tissues, which normally do not accumulate significant amounts of phlobaphene pigments. The maize P gene encodes a Myb homolog that recognizes the sequence CCT/AACC, in sharp contrast with the C/TAACGG bound by vertebrate Myb proteins.

 

The ear leaf is the leaf most closely associated with a particular developing ear. This leaf and above contribute 70%[57] to 75% to 90% of grain fill. Therefore fungicide application is most important in that region in most disease environments.

 

ABNORMAL FLOWERS

Maize flowers may sometimes exhibit mutations that lead to the formation of female flowers in the tassel. These mutations, ts4 and Ts6, prohibit the development of the stamen while simultaneously promoting pistil development. This may cause inflorescences containing both male and female flowers, or hermaphrodite flowers.

 

GENETICS

Maize is an annual grass in the family Gramineae, which includes such plants as wheat, rye, barley, rice, sorghum, and sugarcane. There are two major species of the genus Zea (out of six total): Zea mays (maize) and Zea diploperennis, which is a perennial type of teosinte. The annual teosinte variety called Zea mays mexicana is the closest botanical relative to maize. It still grows in the wild as an annual in Mexico and Guatemala.

 

Many forms of maize are used for food, sometimes classified as various subspecies related to the amount of starch each has:

 

Flour corn: Zea mays var. amylacea

Popcorn: Zea mays var. everta

Dent corn : Zea mays var. indentata

Flint corn: Zea mays var. indurata

Sweet corn: Zea mays var. saccharata and Zea mays var. rugosa

Waxy corn: Zea mays var. ceratina

Amylomaize: Zea mays

Pod corn: Zea mays var. tunicata Larrañaga ex A. St. Hil.

Striped maize: Zea mays var. japonica

 

This system has been replaced (though not entirely displaced) over the last 60 years by multivariable classifications based on ever more data. Agronomic data were supplemented by botanical traits for a robust initial classification, then genetic, cytological, protein and DNA evidence was added. Now, the categories are forms (little used), races, racial complexes, and recently branches.

 

Maize is a diploid with 20 chromosomes (n=10). The combined length of the chromosomes is 1500 cM. Some of the maize chromosomes have what are known as "chromosomal knobs": highly repetitive heterochromatic domains that stain darkly. Individual knobs are polymorphic among strains of both maize and teosinte.

 

Barbara McClintock used these knob markers to validate her transposon theory of "jumping genes", for which she won the 1983 Nobel Prize in Physiology or Medicine. Maize is still an important model organism for genetics and developmental biology today.

 

The centromeres have two types of structural components, both of which are found only in the centromeres: Large arrays of CentC, a short satellite DNA; and a few of a family of retrotransposons. The B chromosome, unlike the others, contains an additional repeat which extends into neighboring areas of the chromosome. Centromeres can accidentally shrink during division and still function, although it is thought this will fail if it shrinks below a few hundred kilobase. Kinetochores contain RNA originating from centromeres. Centromere regions can become inactive, and can continue in that state if the chromosome still has another active one.

 

The Maize Genetics Cooperation Stock Center, funded by the USDA Agricultural Research Service and located in the Department of Crop Sciences at the University of Illinois at Urbana-Champaign, is a stock center of maize mutants. The total collection has nearly 80,000 samples. The bulk of the collection consists of several hundred named genes, plus additional gene combinations and other heritable variants. There are about 1000 chromosomal aberrations (e.g., translocations and inversions) and stocks with abnormal chromosome numbers (e.g., tetraploids). Genetic data describing the maize mutant stocks as well as myriad other data about maize genetics can be accessed at MaizeGDB, the Maize Genetics and Genomics Database.

 

In 2005, the US National Science Foundation (NSF), Department of Agriculture (USDA) and the Department of Energy (DOE) formed a consortium to sequence the B73 maize genome. The resulting DNA sequence data was deposited immediately into GenBank, a public repository for genome-sequence data. Sequences and genome annotations have also been made available throughout the project's lifetime at the project's official site.

 

Primary sequencing of the maize genome was completed in 2008. On November 20, 2009, the consortium published results of its sequencing effort in Science. The genome, 85% of which is composed of transposons, was found to contain 32,540 genes (By comparison, the human genome contains about 2.9 billion bases and 26,000 genes). Much of the maize genome has been duplicated and reshuffled by helitrons—group of rolling circle transposons.

 

In Z. mays and various other angiosperms the MADS-box motif is involved in floral development. Early study in several angiosperm models including Z. mays was the beginning of research into the molecular evolution of floral structure in general, as well as their role in nonflowering plants.

 

EVOLUTION

As with many plants and animals, Z. mays has a positive correlation between effective population size and the magnitude of selection pressure. Z. m. having an EPS of ~650,000, it clusters with others of about the same EPS, and has 79% of its amino acid sites under selection.

 

Recombination is a significant source of diversity in Z. mays. (Note that this finding supersedes previous studies which showed no such correlation.)

 

This recombination/diversity effect is seen throughout plants but is also found to not occur – or not as strongly – in regions of high gene density. This is likely the reason that domesticated Z. mays has not seen as much of an increase in diversity within areas of higher density as in regions of lower density, although there is more evidence in other plants.

 

Some lines of maize have undergone ancient polyploidy events, starting 11m years ago. Over that time ~72% of polyploid duplicated genes have been retained, which is higher than other plants with older polyploidy events. Thus maize may be due to lose more duplicate genes as time goes along, similar to the course followed by the genomes of other plants. If so - if gene loss has merely not occurred yet - that could explain the lack of observed positive selection and lower negative selection which are observed in otherwise similar plants, i.e. also naturally outcrossing and with similar effective population sizes.

 

Ploidy does not appear to influence EPS or magnitude of selection effect in maize.

 

BREEDING

Maize reproduces sexually each year. This randomly selects half the genes from a given plant to propagate to the next generation, meaning that desirable traits found in the crop (like high yield or good nutrition) can be lost in subsequent generations unless certain techniques are used.

 

Maize breeding in prehistory resulted in large plants producing large ears. Modern breeding began with individuals who selected highly productive varieties in their fields and then sold seed to other farmers. James L. Reid was one of the earliest and most successful developing Reid's Yellow Dent in the 1860s. These early efforts were based on mass selection. Later breeding efforts included ear to row selection (C. G. Hopkins c. 1896), hybrids made from selected inbred lines (G. H. Shull, 1909), and the highly successful double cross hybrids using four inbred lines (D. F. Jones c. 1918, 1922). University supported breeding programs were especially important in developing and introducing modern hybrids. By the 1930s, companies such as Pioneer devoted to production of hybrid maize had begun to influence long-term development. Internationally important seed banks such as the International Maize and Wheat Improvement Center (CIMMYT) and the US bank at the Maize Genetics Cooperation Stock Center University of Illinois at Urbana-Champaign maintain germplasm important for future crop development.

 

Since the 1940s the best strains of maize have been first-generation hybrids made from inbred strains that have been optimized for specific traits, such as yield, nutrition, drought, pest and disease tolerance. Both conventional cross-breeding and genetic modification have succeeded in increasing output and reducing the need for cropland, pesticides, water and fertilizer. There is conflicting evidence to support the hypothesis that maize yield potential has increased over the past few decades. This suggests that changes in yield potential are associated with leaf angle, lodging resistance, tolerance of high plant density, disease/pest tolerance, and other agronomic traits rather than increase of yield potential per individual plant.

 

Tropical landraces remain an important and underutilized source of resistance alleles for for disease and for herbivores. Notable discoveries of rare alleles for this purpose were made by Dao et al 2014 and Sood et al 2014.

 

GLOBAL PROGRAM

CIMMYT operates a conventional breeding program to provide optimized strains. The program began in the 1980s. Hybrid seeds are distributed in Africa by the Drought Tolerant Maize for Africa project.

 

GENETIC MODIFICATION

Genetically modified (GM) maize was one of the 26 GM crops grown commercially in 2016. The vast majority of this is Bt maize. Grown since 1997 in the United States and Canada, 92% of the US maize crop was genetically modified in 2016 and 33% of the worldwide maize crop was GM in 2016. As of 2011, Herbicide-tolerant maize varieties were grown in Argentina, Australia, Brazil, Canada, China, Colombia, El Salvador, the European Union, Honduras, Japan, Korea, Malaysia, Mexico, New Zealand, Philippines, the Russian Federation, Singapore, South Africa, Taiwan, Thailand, and the United States. Insect-resistant maize was grown in Argentina, Australia, Brazil, Canada, Chile, China, Colombia, Egypt, the European Union, Honduras, Japan, Korea, Malaysia, Mexico, New Zealand, Philippines, South Africa, Switzerland, Taiwan, the United States, and Uruguay.

 

In September 2000, up to $50 million worth of food products were recalled due to the presence of Starlink genetically modified corn, which had been approved only for animal consumption and had not been approved for human consumption, and was subsequently withdrawn from the market.

 

ORIGIN

Maize is the domesticated variant of teosinte. The two plants have dissimilar appearance, maize having a single tall stalk with multiple leaves and teosinte being a short, bushy plant. The difference between the two is largely controlled by differences in just two genes, called grassy tillers-1 (gt1, A0A317YEZ1) and teosinte branched-1 (tb1, Q93WI2).

 

Several theories had been proposed about the specific origin of maize in Mesoamerica:

 

It is a direct domestication of a Mexican annual teosinte, Zea mays ssp. parviglumis, native to the Balsas River valley in south-eastern Mexico, with up to 12% of its genetic material obtained from Zea mays ssp. mexicana through introgression.

It has been derived from hybridization between a small domesticated maize (a slightly changed form of a wild maize) and a teosinte of section Luxuriantes, either Z. luxurians or Z. diploperennis.

It has undergone two or more domestications either of a wild maize or of a teosinte. (The term "teosinte" describes all species and subspecies in the genus Zea, excluding Zea mays ssp. mays.)

It has evolved from a hybridization of Z. diploperennis by Tripsacum dactyloides.

 

In the late 1930s, Paul Mangelsdorf suggested that domesticated maize was the result of a hybridization event between an unknown wild maize and a species of Tripsacum, a related genus. This theory about the origin of maize has been refuted by modern genetic testing, which refutes Mangelsdorf's model and the fourth listed above. 

 

The teosinte origin theory was proposed by the Russian botanist Nikolai Ivanovich Vavilov in 1931 and the later American Nobel Prize-winner George Beadle in 1932.: 10  It is supported experimentally and by recent studies of the plants' genomes. Teosinte and maize can cross-breed and produce fertile offspring. A number of questions remain concerning the species, among them:

 

how the immense diversity of the species of sect. Zea originated,

how the tiny archaeological specimens of 3500–2700 BC could have been selected from a teosinte, and

how domestication could have proceeded without leaving remains of teosinte or maize with teosintoid traits earlier than the earliest known until recently, dating from ca. 1100 BC.

 

The domestication of maize is of particular interest to researchers—archaeologists, geneticists, ethnobotanists, geographers, etc. The process is thought by some to have started 7,500 to 12,000 years ago. Research from the 1950s to 1970s originally focused on the hypothesis that maize domestication occurred in the highlands between the states of Oaxaca and Jalisco, because the oldest archaeological remains of maize known at the time were found there.

Connection with 'parviglumis' subspecies

Genetic studies, published in 2004 by John Doebley, identified Zea mays ssp. parviglumis, native to the Balsas River valley in Mexico's southwestern highlands, and also known as Balsas teosinte, as being the crop wild relative that is genetically most similar to modern maize. This was confirmed by further studies, which refined this hypothesis somewhat. Archaeobotanical studies, published in 2009, point to the middle part of the Balsas River valley as the likely location of early domestication; this river is not very long, so these locations are not very distant. Stone milling tools with maize residue have been found in an 8,700 year old layer of deposits in a cave not far from Iguala, Guerrero.

 

Doebley was part of the team that first published, in 2002, that maize had been domesticated only once, about 9,000 years ago, and then spread throughout the Americas.

 

A primitive corn was being grown in southern Mexico, Central America, and northern South America 7,000 years ago. Archaeological remains of early maize ears, found at Guila Naquitz Cave in the Oaxaca Valley, date back roughly 6,250 years; the oldest ears from caves near Tehuacan, Puebla, 5,450 B.P.

 

Maize pollen dated to 7,300 B.P. from San Andres, Tabasco, on the Caribbean coast has also been recovered.

 

As maize was introduced to new cultures, new uses were developed and new varieties selected to better serve in those preparations. Maize was the staple food, or a major staple – along with squash, Andean region potato, quinoa, beans, and amaranth – of most pre-Columbian North American, Mesoamerican, South American, and Caribbean cultures. The Mesoamerican civilization, in particular, was deeply interrelated with maize. Its traditions and rituals involved all aspects of maize cultivation – from the planting to the food preparation. Maize formed the Mesoamerican people's identity.

 

It is unknown what precipitated its domestication, because the edible portion of the wild variety is too small, and hard to obtain, to be eaten directly, as each kernel is enclosed in a very hard bivalve shell.

 

In 1939, George Beadle demonstrated that the kernels of teosinte are readily "popped" for human consumption, like modern popcorn.[91] Some have argued it would have taken too many generations of selective breeding to produce large, compressed ears for efficient cultivation. However, studies of the hybrids readily made by intercrossing teosinte and modern maize suggest this objection is not well founded.

 

SPREADING TO THE NORTH

Around 4,500 ago, maize began to spread to the north; it was first cultivated in what is now the United States at several sites in New Mexico and Arizona, about 4,100 ago.

 

During the first millennium AD, maize cultivation spread more widely in the areas north. In particular, the large-scale adoption of maize agriculture and consumption in eastern North America took place about A.D. 900. Native Americans cleared large forest and grassland areas for the new crop.

 

In 2005, research by the USDA Forest Service suggested that the rise in maize cultivation 500 to 1,000 years ago in what is now the southeastern United States corresponded with a decline of freshwater mussels, which are very sensitive to environmental changes.

 

CULTIVATION

PLANTING

Because it is cold-intolerant, in the temperate zones maize must be planted in the spring. Its root system is generally shallow, so the plant is dependent on soil moisture. As a plant that uses C4 carbon fixation, maize is a considerably more water-efficient crop than plants that use C3 carbon fixation such as alfalfa and soybeans. Maize is most sensitive to drought at the time of silk emergence, when the flowers are ready for pollination. In the United States, a good harvest was traditionally predicted if the maize was "knee-high by the Fourth of July", although modern hybrids generally exceed this growth rate. Maize used for silage is harvested while the plant is green and the fruit immature. Sweet corn is harvested in the "milk stage", after pollination but before starch has formed, between late summer and early to mid-autumn. Field maize is left in the field until very late in the autumn to thoroughly dry the grain, and may, in fact, sometimes not be harvested until winter or even early spring. The importance of sufficient soil moisture is shown in many parts of Africa, where periodic drought regularly causes maize crop failure and consequent famine. Although it is grown mainly in wet, hot climates, it has been said to thrive in cold, hot, dry or wet conditions, meaning that it is an extremely versatile crop.

 

Maize was planted by the Native Americans in hills, in a complex system known to some as the Three Sisters. Maize provided support for beans, and the beans provided nitrogen derived from nitrogen-fixing rhizobia bacteria which live on the roots of beans and other legumes; and squashes provided ground cover to stop weeds and inhibit evaporation by providing shade over the soil. This method was replaced by single species hill planting where each hill 60–120 cm (2 ft 0 in–3 ft 11 in) apart was planted with three or four seeds, a method still used by home gardeners. A later technique was "checked maize", where hills were placed

 

1 m (40 in) apart in each direction, allowing cultivators to run through the field in two directions. In more arid lands, this was altered and seeds were planted in the bottom of 10–12 cm (4–4+1⁄2 in) deep furrows to collect water. Modern technique plants maize in rows which allows for cultivation while the plant is young, although the hill technique is still used in the maize fields of some Native American reservations. When maize is planted in rows, it also allows for planting of other crops between these rows to make more efficient use of land space.

 

In most regions today, maize grown in residential gardens is still often planted manually with a hoe, whereas maize grown commercially is no longer planted manually but rather is planted with a planter. In North America, fields are often planted in a two-crop rotation with a nitrogen-fixing crop, often alfalfa in cooler climates and soybeans in regions with longer summers. Sometimes a third crop, winter wheat, is added to the rotation.

 

Many of the maize varieties grown in the United States and Canada are hybrids. Often the varieties have been genetically modified to tolerate glyphosate or to provide protection against natural pests. Glyphosate is an herbicide which kills all plants except those with genetic tolerance. This genetic tolerance is very rarely found in nature.

 

In the midwestern United States, low-till or no-till farming techniques are usually used. In low-till, fields are covered once, maybe twice, with a tillage implement either ahead of crop planting or after the previous harvest. The fields are planted and fertilized. Weeds are controlled through the use of herbicides, and no cultivation tillage is done during the growing season. This technique reduces moisture evaporation from the soil, and thus provides more moisture for the crop. The technologies mentioned in the previous paragraph enable low-till and no-till farming. Weeds compete with the crop for moisture and nutrients, making them undesirable.

 

HARVESTING

Before the 20th century, all maize harvesting was by manual labour, by grazing, or by some combination of those. Whether the ears were hand-picked and the stover was grazed, or the whole plant was cut, gathered, and shocked, people and livestock did all the work. Between the 1890s and the 1970s, the technology of maize harvesting expanded greatly. Today, all such technologies, from entirely manual harvesting to entirely mechanized, are still in use to some degree, as appropriate to each farm's needs, although the thoroughly mechanized versions predominate, as they offer the lowest unit costs when scaled to large farm operations. For small farms, their unit cost can be too high, as their higher fixed cost cannot be amortized over as many units.[citation needed]

 

Before World War II, most maize in North America was harvested by hand. This involved a large number of workers and associated social events (husking or shucking bees). From the 1890s onward, some machinery became available to partially mechanize the processes, such as one- and two-row mechanical pickers (picking the ear, leaving the stover) and corn binders, which are reaper-binders designed specifically for maize (for example, Video on YouTube). The latter produce sheaves that can be shocked. By hand or mechanical picker, the entire ear is harvested, which then requires a separate operation of a maize sheller to remove the kernels from the ear. Whole ears of maize were often stored in corn cribs, and these whole ears are a sufficient form for some livestock feeding use. Today corn cribs with whole ears, and corn binders, are less common because most modern farms harvest the grain from the field with a combine and store it in bins. The combine with a corn head (with points and snap rolls instead of a reel) does not cut the stalk; it simply pulls the stalk down. The stalk continues downward and is crumpled into a mangled pile on the ground, where it usually is left to become organic matter for the soil. The ear of maize is too large to pass between slots in a plate as the snap rolls pull the stalk away, leaving only the ear and husk to enter the machinery. The combine separates the husk and the cob, keeping only the kernels.

When maize is a silage crop, the entire plant is usually chopped at once with a forage harvester (chopper) and ensiled in silos or polymer wrappers. Ensiling of sheaves cut by a corn binder was formerly common in some regions but has become uncommon. For storing grain in bins, the moisture of the grain must be sufficiently low to avoid spoiling. If the moisture content of the harvested grain is too high, grain dryers are used to reduce the moisture content by blowing heated air through the grain. This can require large amounts of energy in the form of combustible gases (propane or natural gas) and electricity to power the blowers.

 

PRODUCTION

Maize is widely cultivated throughout the world, and a greater weight of maize is produced each year than any other grain. In 2018, total world production was 1.15 billion tonnes, led by the United States with 34.2% of the total (table). China produced 22.4% of the global total.

 

UNITED STATES

In 2016, maize production was forecast to be over 380 million metric tons (15 billion bushels), an increase of 11% over 2014 American production. Based on conditions as of August 2016, the expected yield would be the highest ever for the United States. The area of harvested maize was forecast to be 35 million hectares (87 million acres), an increase of 7% over 2015. Maize is especially popular in Midwestern states such as Indiana, Iowa, and Illinois; in the latter, it was named the state's official grain in 2017.

 

STORAGE

Drying is vital to prevent or at least reduce mycotoxin contamination. Aspergillus and Fusarium spp. are the most common mycotoxin sources, but there are others. Altogether maize contaminants are so common, and this crop is so economically important, that maize mycotoxins are among the most important in agriculture in general.

 

USES

HUMAN FOOD

Maize and cornmeal (ground dried maize) constitute a staple food in many regions of the world. Maize is used to produce cornstarch, a common ingredient in home cooking and many industrialized food products. Maize starch can be hydrolyzed and enzymatically treated to produce syrups, particularly high fructose corn syrup, a sweetener; and also fermented and distilled to produce grain alcohol. Grain alcohol from maize is traditionally the source of Bourbon whiskey. Corn flour is used to make cornbread and other baked products.

 

In prehistoric times Mesoamerican women used a metate to process maize into ground cornmeal, allowing the preparation of foods that were more calorie dense than popcorn. After ceramic vessels were invented the Olmec people began to cook maize together with beans, improving the nutritional value of the staple meal. Although maize naturally contains niacin, an important nutrient, it was not bioavailable without the process of nixtamalization. The Maya used nixtamal meal to make varieties of porridges and tamales. The process was later used in the cuisine of the American South to prepare corn for grits and hominy.

 

Maize is a staple of Mexican cuisine. Masa (cornmeal treated with limewater) is the main ingredient for tortillas, atole and many other dishes of Central American food. It is the main ingredient of corn tortilla, tamales, pozole, atole and all the dishes based on them, like tacos, quesadillas, chilaquiles, enchiladas, tostadas and many more. In Mexico the fungus of maize, known as huitlacoche, is considered a delicacy.

 

Coarse maize meal is made into a thick porridge in many cultures: from the polenta of Italy, the angu of Brazil, the mămăligă of Romania, to cornmeal mush in the US (or hominy grits in the South) or the food called mieliepap in South Africa and sadza, nshima, ugali and other names in other parts of Africa. Introduced into Africa by the Portuguese in the 16th century, maize has become Africa's most important staple food crop. These are commonly eaten in the Southeastern United States, foods handed down from Native Americans, who called the dish sagamite.

 

Maize can also be harvested and consumed in the unripe state, when the kernels are fully grown but still soft. Unripe maize must usually be cooked to become palatable; this may be done by simply boiling or roasting the whole ears and eating the kernels right off the cob. Sweet corn, a genetic variety that is high in sugars and low in starch, is usually consumed in the unripe state. Such corn on the cob is a common dish in the United States, Canada, United Kingdom, Cyprus, some parts of South America, and the Balkans, but virtually unheard of in some European countries. Corn on the cob was hawked on the streets of early 19th-century New York City by poor, barefoot "Hot Corn Girls", who were thus the precursors of hot dog carts, churro wagons, and fruit stands seen on the streets of big cities today.

 

Within the United States, the usage of maize for human consumption constitutes only around 1/40th of the amount grown in the country. In the United States and Canada, maize is mostly grown to feed livestock, as forage, silage (made by fermentation of chopped green cornstalks), or grain. Maize meal is also a significant ingredient of some commercial animal food products.

 

NUTRITIONAL VALUE

Raw, yellow, sweet maize kernels are composed of 76% water, 19% carbohydrates, 3% protein, and 1% fat (table). In a 100-gram serving, maize kernels provide 86 calories and are a good source (10–19% of the Daily Value) of the B vitamins, thiamin, niacin (but see Pellagra warning below), pantothenic acid (B5) and folate (right table for raw, uncooked kernels, USDA Nutrient Database). In moderate amounts, they also supply dietary fiber and the essential minerals, magnesium and phosphorus whereas other nutrients are in low amounts (table).

 

Maize has suboptimal amounts of the essential amino acids tryptophan and lysine, which accounts for its lower status as a protein source. However, the proteins of beans and legumes complement those of maize.

 

FEED AND FODDER FOR LIVESTOCK

Maize is a major source of both grain feed and fodder for livestock. It is fed to the livestock in various ways. When it is used as a grain crop, the dried kernels are used as feed. They are often kept on the cob for storage in a corn crib, or they may be shelled off for storage in a grain bin. The farm that consumes the feed may produce it, purchase it on the market, or some of both. When the grain is used for feed, the rest of the plant (the corn stover) can be used later as fodder, bedding (litter), or soil amendment. When the whole maize plant (grain plus stalks and leaves) is used for fodder, it is usually chopped all at once and ensilaged, as digestibility and palatability are higher in the ensilaged form than in the dried form. Maize silage is one of the most valuable forages for ruminants. Before the advent of widespread ensilaging, it was traditional to gather the corn into shocks after harvesting, where it dried further. With or without a subsequent move to the cover of a barn, it was then stored for weeks to several months until fed to the livestock. Today ensilaging can occur not only in siloes but also in silage wrappers. However, in the tropics, maize can be harvested year-round and fed as green forage to the animals.

 

CHEMICALS

Starch from maize can also be made into plastics, fabrics, adhesives, and many other chemical products.

 

The corn steep liquor, a plentiful watery byproduct of maize wet milling process, is widely used in the biochemical industry and research as a culture medium to grow many kinds of microorganisms.

 

Chrysanthemin is found in purple corn and is used as a food coloring.

 

BIO-FUEL

"Feed maize" is being used increasingly for heating; specialized corn stoves (similar to wood stoves) are available and use either feed maize or wood pellets to generate heat. Maize cobs are also used as a biomass fuel source. Maize is relatively cheap and home-heating furnaces have been developed which use maize kernels as a fuel. They feature a large hopper that feeds the uniformly sized maize kernels (or wood pellets or cherry pits) into the fire.[citation needed]

 

Maize is increasingly used as a feedstock for the production of ethanol fuel.[120] When considering where to construct an ethanol plant, one of the site selection criteria is to ensure there is locally available feedstock. Ethanol is mixed with gasoline to decrease the amount of pollutants emitted when used to fuel motor vehicles. High fuel prices in mid-2007 led to higher demand for ethanol, which in turn led to higher prices paid to farmers for maize. This led to the 2007 harvest being one of the most profitable maize crops in modern history for farmers. Because of the relationship between fuel and maize, prices paid for the crop now tend to track the price of oil.

 

The price of food is affected to a certain degree by the use of maize for biofuel production. The cost of transportation, production, and marketing are a large portion (80%) of the price of food in the United States. Higher energy costs affect these costs, especially transportation. The increase in food prices the consumer has been seeing is mainly due to the higher energy cost. The effect of biofuel production on other food crop prices is indirect. Use of maize for biofuel production increases the demand, and therefore price of maize. This, in turn, results in farm acreage being diverted from other food crops to maize production. This reduces the supply of the other food crops and increases their prices.

 

Maize is widely used in Germany as a feedstock for biogas plants. Here the maize is harvested, shredded then placed in silage clamps from which it is fed into the biogas plants. This process makes use of the whole plant rather than simply using the kernels as in the production of fuel ethanol.

 

A biomass gasification power plant in Strem near Güssing, Burgenland, Austria, began in 2005. Research is being done to make diesel out of the biogas by the Fischer Tropsch method.

 

Increasingly, ethanol is being used at low concentrations (10% or less) as an additive in gasoline (gasohol) for motor fuels to increase the octane rating, lower pollutants, and reduce petroleum use (what is nowadays also known as "biofuels" and has been generating an intense debate regarding the human beings' necessity of new sources of energy, on the one hand, and the need to maintain, in regions such as Latin America, the food habits and culture which has been the essence of civilizations such as the one originated in Mesoamerica; the entry, January 2008, of maize among the commercial agreements of NAFTA has increased this debate, considering the bad labor conditions of workers in the fields, and mainly the fact that NAFTA "opened the doors to the import of maize from the United States, where the farmers who grow it receive multimillion-dollar subsidies and other government supports. ... According to OXFAM UK, after NAFTA went into effect, the price of maize in Mexico fell 70% between 1994 and 2001. The number of farm jobs dropped as well: from 8.1 million in 1993 to 6.8 million in 2002. Many of those who found themselves without work were small-scale maize growers."). However, introduction in the northern latitudes of the US of tropical maize for biofuels, and not for human or animal consumption, may potentially alleviate this.

 

COMMODITY

Maize is bought and sold by investors and price speculators as a tradable commodity using corn futures contracts. These "futures" are traded on the Chicago Board of Trade (CBOT) under ticker symbol C. They are delivered every year in March, May, July, September, and December.

Ornamental and other uses

 

Some forms of the plant are occasionally grown for ornamental use in the garden. For this purpose, variegated and colored leaf forms as well as those with colorful ears are used.

 

Corncobs can be hollowed out and treated to make inexpensive smoking pipes, first manufactured in the United States in 1869.

 

An unusual use for maize is to create a "corn maze" (or "maize maze") as a tourist attraction. The idea of a maize maze was introduced by the American Maze Company who created a maze in Pennsylvania in 1993. Traditional mazes are most commonly grown using yew hedges, but these take several years to mature. The rapid growth of a field of maize allows a maze to be laid out using GPS at the start of a growing season and for the maize to grow tall enough to obstruct a visitor's line of sight by the start of the summer. In Canada and the US, these are popular in many farming communities.

 

Maize kernels can be used in place of sand in a sandboxlike enclosure for children's play.

 

Stigmas from female maize flowers, popularly called corn silk, are sold as herbal supplements.

 

Maize is used as a fish bait, called "dough balls". It is particularly popular in Europe for coarse fishing.

 

Additionally, feed corn is sometimes used by hunters to bait animals such as deer or wild hogs.

 

UNITED STATES USAGE BREAKDOWN

The breakdown of usage of the 12.1-billion-bushel (307-million-tonne) 2008 US maize crop was as follows, according to the World Agricultural Supply and Demand Estimates Report by the USDA.In the US since 2009/2010, maize feedstock use for ethanol production has somewhat exceeded direct use for livestock feed; maize use for fuel ethanol was 5,130 million bushels (130 million tonnes) in the 2013/2014 marketing year.A fraction of the maize feedstock dry matter used for ethanol production is usefully recovered as DDGS (dried distillers grains with solubles). In the 2010/2011 marketing year, about 29.1 million tonnes of DDGS were fed to US livestock and poultry. Because starch utilization in fermentation for ethanol production leaves other grain constituents more concentrated in the residue, the feed value per kg of DDGS, with regard to ruminant-metabolizable energy and protein, exceeds that of the grain. Feed value for monogastric animals, such as swine and poultry, is somewhat lower than for ruminants.

 

HAZARDS

PELLAGRA

When maize was first introduced into farming systems other than those used by traditional native-American peoples, it was generally welcomed with enthusiasm for its productivity. However, a widespread problem of malnutrition soon arose wherever maize was introduced as a staple food. This was a mystery, since these types of malnutrition were not normally seen among the indigenous Americans, for whom maize was the principal staple food.

 

It was eventually discovered that the indigenous Americans had learned to soak maize in alkali — water (the process now known as nixtamalization) — made with ashes and lime (calcium oxide) since at least 1200–1500 BC by Mesoamericans. They did this to liberate the corn hulls, but (unbeknownst to natives or colonists) it coincidentally liberates the B-vitamin niacin, the lack of which was the underlying cause of the condition known as pellagra.

 

Maize was introduced into the diet of non-indigenous Americans without the necessary cultural knowledge acquired over thousands of years in the Americas. In the late 19th century, pellagra reached epidemic proportions in parts of the southern US, as medical researchers debated two theories for its origin: the deficiency theory (which was eventually shown to be true) said that pellagra was due to a deficiency of some nutrient, and the germ theory said that pellagra was caused by a germ transmitted by stable flies. A third theory, promoted by the eugenicist Charles Davenport, held that people only contracted pellagra if they were susceptible to it due to certain "constitutional, inheritable" traits of the affected individual.

 

Once alkali processing and dietary variety were understood and applied, pellagra disappeared in the developed world. The development of high lysine maize and the promotion of a more balanced diet have also contributed to its demise. Pellagra still exists today in food-poor areas and refugee camps where people survive on donated maize.

 

ALLERGY

Maize contains lipid transfer protein, an indigestible protein that survives cooking. This protein has been linked to a rare and understudied allergy to maize in humans. The allergic reaction can cause skin rash, swelling or itching of mucous membranes, diarrhea, vomiting, asthma and, in severe cases, anaphylaxis. It is unclear how common this allergy is in the general population.

 

MYCOTOXINS

Fungicide application does not reduce fungal growth or mycotoxin dramatically, although it can be a part of a successful reduction strategy. Among the most common toxins are those produced by Aspergillus and Fusarium spp. The most common toxins are aflatoxins, fumonisins, zearalenone, and ochratoxin A. Bt maize discourages insect vectors and by so doing it dramatically reduces concentrations of fumonisins, significantly reduces aflatoxins, but only mildly reduces others.

 

ART

Maize has been an essential crop in the Andes since the pre-Columbian era. The Moche culture from Northern Peru made ceramics from earth, water, and fire. This pottery was a sacred substance, formed in significant shapes and used to represent important themes. Maize was represented anthropomorphically as well as naturally.

 

In the United States, maize ears along with tobacco leaves are carved into the capitals of columns in the United States Capitol building. Maize itself is sometimes used for temporary architectural detailing when the intent is to celebrate the fall season, local agricultural productivity and culture. Bundles of dried maize stalks are often displayed along with pumpkins, gourds and straw in autumnal displays outside homes and businesses. A well-known example of architectural use is the Corn Palace in Mitchell, South Dakota, which uses cobs and ears of colored maize to implement a mural design that is recycled annually. Another well-known example is the Field of Corn sculpture in Dublin, Ohio, where hundreds of concrete ears of corn stand in a grassy field.

 

A maize stalk with two ripe ears is depicted on the reverse of the Croatian 1 lipa coin, minted since 1993.

 

WIKIPEDIA

le Mascaret, Rixensart

The peregrine falcon (Falco peregrinus), also known simply as the peregrine, and historically as the duck hawk in North America, is a cosmopolitan bird of prey (raptor) in the family Falconidae. A large, crow-sized falcon, it has a blue-grey back, barred white underparts, and a black head. The peregrine is renowned for its speed. It can reach over 320 km/h (200 mph) during its characteristic hunting stoop (high-speed dive), making it the fastest member of the animal kingdom. According to a National Geographic TV program, the highest measured speed of a peregrine falcon is 389 km/h (242 mph). As is typical for bird-eating (avivore) raptors, peregrine falcons are sexually dimorphic, with females being considerably larger than males.

 

The peregrine's breeding range includes land regions from the Arctic tundra to the tropics. It can be found nearly everywhere on Earth, except extreme polar regions, very high mountains, and most tropical rainforests; the only major ice-free landmass from which it is entirely absent is New Zealand. This makes it the world's most widespread raptor and one of the most widely found wild bird species. In fact, the only land-based bird species found over a larger geographic area owes its success to human-led introduction; the domestic and feral pigeons are both domesticate forms of the rock dove, which are a major prey species for Eurasian Peregrine populations. Due to their prevalence over most other bird species in cities, feral pigeons support many peregrine populations as a staple food source, especially in urban settings.

 

The peregrine is a highly successful example of urban wildlife in much of its range, taking advantage of tall buildings as nest sites and an abundance of prey such as pigeons and ducks. Both the English and scientific names of this species mean "wandering falcon", referring to the migratory habits of many northern populations. Experts recognize 17 to 19 subspecies, which vary in appearance and range; disagreement exists over whether the distinctive Barbary falcon is represented by two subspecies of Falco peregrinus or is a separate species, F. pelegrinoides. The two species' divergence is relatively recent, during the time of the last ice age, therefore the genetic differential between them (and also the difference in their appearance) is relatively tiny. They are only about 0.6–0.8% genetically differentiated.

 

Although its diet consists almost exclusively of medium-sized birds, the peregrine will sometimes hunt small mammals, small reptiles, or even insects. Reaching sexual maturity at one year, it mates for life and nests in a scrape, normally on cliff edges or, in recent times, on tall human-made structures. The peregrine falcon became an endangered species in many areas because of the widespread use of certain pesticides, especially DDT. Since the ban on DDT from the early 1970s, populations have recovered, supported by large-scale protection of nesting places and releases to the wild.

 

The peregrine falcon is a well-respected falconry bird due to its strong hunting ability, high trainability, versatility, and availability via captive breeding. It is effective on most game bird species, from small to large. It has also been used as a religious, royal, or national symbol across multiple eras and areas of human civilization.

 

Description

 

Falco peregrinus. Royal National Park, New South Wales, Australia

The peregrine falcon has a body length of 34 to 58 cm (13–23 in) and a wingspan from 74 to 120 cm (29–47 in). The male and female have similar markings and plumage but, as with many birds of prey, the peregrine falcon displays marked sexual dimorphism in size, with the female measuring up to 30% larger than the male. Males weigh 330 to 1,000 g (12–35 oz) and the noticeably larger females weigh 700 to 1,500 g (25–53 oz). In most subspecies, males weigh less than 700 g (25 oz) and females weigh more than 800 g (28 oz), and cases of females weighing about 50% more than their male breeding mates are not uncommon. The standard linear measurements of peregrines are: the wing chord measures 26.5 to 39 cm (10.4–15.4 in), the tail measures 13 to 19 cm (5.1–7.5 in) and the tarsus measures 4.5 to 5.6 cm (1.8–2.2 in).

 

The back and the long pointed wings of the adult are usually bluish black to slate grey with indistinct darker barring (see "Subspecies" below); the wingtips are black. The white to rusty underparts are barred with thin clean bands of dark brown or black. The tail, coloured like the back but with thin clean bars, is long, narrow, and rounded at the end with a black tip and a white band at the very end. The top of the head and a "moustache" along the cheeks are black, contrasting sharply with the pale sides of the neck and white throat. The cere is yellow, as are the feet, and the beak and claws are black. The upper beak is notched near the tip, an adaptation which enables falcons to kill prey by severing the spinal column at the neck. An immature bird is much browner, with streaked, rather than barred, underparts, and has a pale bluish cere and orbital ring.

 

A study shows that their black malar stripe exists to reduce glare from solar radiation, allowing them to see better. Photos from The Macaulay Library and iNaturalist showed that the malar stripe is thicker where there is more solar radiation. That supports the solar glare hypothesis.

 

Taxonomy and systematics

 

Falco peregrinus was first described under its current binomial name by English ornithologist Marmaduke Tunstall in his 1771 work Ornithologia Britannica. The scientific name Falco peregrinus is a Medieval Latin phrase that was used by Albertus Magnus in 1225. Peregrinus is Latin, meaning "one from abroad" or "coming from foreign parts". It is likely the name was used as juvenile birds were taken while journeying to their breeding location (rather than from the nest), as falcon nests are often difficult to get at. The Latin term for falcon, falco, is related to falx, meaning "sickle", in reference to the silhouette of the falcon's long, pointed wings in flight.

 

The peregrine falcon belongs to a genus whose lineage includes the hierofalcon and the prairie falcon (F. mexicanus). This lineage probably diverged from other falcons towards the end of the Late Miocene or in the Late Pliocene, about 3–8 million years ago (mya). As the peregrine-hierofalcon group includes both Old World and North American species, it is likely that the lineage originated in western Eurasia or Africa. Its relationship to other falcons is not clear, as the issue is complicated by widespread hybridization confounding mtDNA sequence analyses. One genetic lineage of the saker falcon (F. cherrug) is known to have originated from a male saker ancestor producing fertile young with a female peregrine ancestor, and the descendants further breeding with sakers.

 

Today, peregrines are regularly paired in captivity with other species such as the lanner falcon (F. biarmicus) to produce the "perilanner", a somewhat popular bird in falconry as it combines the peregrine's hunting skill with the lanner's hardiness, or the gyrfalcon to produce large, strikingly coloured birds for the use of falconers. As can be seen, the peregrine is still genetically close to the hierofalcons, though their lineages diverged in the Late Pliocene (maybe some 2.5–2 mya in the Gelasian).

 

Subspecies

Numerous subspecies of Falco peregrinus have been described, with 19 accepted by the 1994 Handbook of the Birds of the World, which considers the Barbary falcon of the Canary Islands and coastal North Africa to be two subspecies (pelegrinoides and babylonicus) of Falco peregrinus, rather than a distinct species, F. pelegrinoides. The following map shows the general ranges of these 19 subspecies.

 

A map of the world, green shows on several continents, but there are also several big bare spots marked with E for extinct

Breeding ranges of the 19 subspecies

 

Falco peregrinus anatum, described by Bonaparte in 1838, is known as the American peregrine falcon or "duck hawk"; its scientific name means "duck peregrine falcon". At one time, it was partly included in leucogenys. It is mainly found in the Rocky Mountains. It was formerly common throughout North America between the tundra and northern Mexico, where current reintroduction efforts are being made to restore the population. Most mature anatum, except those that breed in more northern areas, winter in their breeding range. Most vagrants that reach western Europe seem to belong to the more northern and strongly migratory tundrius, only considered distinct since 1968. It is similar to the nominate subspecies but is slightly smaller; adults are somewhat paler and less patterned below, but juveniles are darker and more patterned below. Males weigh 500 to 700 g (1.1–1.5 lb), while females weigh 800 to 1,100 g (1.8–2.4 lb). It has become extinct in eastern North America and populations there are hybrids as a result of reintroductions of birds from elsewhere.

Falco peregrinus babylonicus, described by P.L. Sclater in 1861, is found in eastern Iran along the Hindu Kush and the Tian Shan to the Mongolian Altai ranges. A few birds winter in northern and northwestern India, mainly in dry semi-desert habitats. It is paler than pelegrinoides and somewhat similar to a small, pale lanner falcon (Falco biarmicus). Males weigh 330 to 400 grams (12 to 14 oz), while females weigh 513 to 765 grams (18.1 to 27.0 oz).

Falco peregrinus brookei, described by Sharpe in 1873, is also known as the Mediterranean peregrine falcon or the Maltese falcon. It includes caucasicus and most specimens of the proposed race punicus, though others may be pelegrinoides (Barbary falcons), or perhaps the rare hybrids between these two which might occur around Algeria. They occur from the Iberian Peninsula around the Mediterranean, except in arid regions, to the Caucasus. They are non-migratory. It is smaller than the nominate subspecies and the underside usually has a rusty hue. Males weigh around 445 g (0.981 lb), while females weigh up to 920 g (2.03 lb).

Falco peregrinus calidus, described by John Latham in 1790, it was formerly called leucogenys and includes caeruleiceps. It breeds in the Arctic tundra of Eurasia from Murmansk Oblast to roughly Yana and Indigirka Rivers, Siberia. It is completely migratory and travels south in winter as far as South Asia and sub-Saharan Africa. It is often seen around wetland habitats. It is paler than the nominate subspecies, especially on the crown. Males weigh 588 to 740 g (1.296–1.631 lb), while females weigh 925 to 1,333 g (2.039–2.939 lb).

Falco peregrinus cassini, described by Sharpe in 1873, is also known as the austral peregrine falcon. It includes kreyenborgi, the pallid falcon, a leucistic colour morph occurring in southernmost South America, which was long believed to be a distinct species. Its range includes South America from Ecuador through Bolivia, northern Argentina and Chile to Tierra del Fuego and the Falkland Islands. It is non-migratory. It is similar to the nominate subspecies, but slightly smaller with a black ear region. The pallid falcon morph kreyenborgi is medium grey above, has little barring below and has a head pattern like the saker falcon (Falco cherrug), but the ear region is white.

Falco peregrinus ernesti, described by Sharpe in 1894, is found from the Sunda Islands to the Philippines and south to eastern New Guinea and the nearby Bismarck Archipelago. Its geographical separation from nesiotes requires confirmation. It is non-migratory. It differs from the nominate subspecies in the very dark, dense barring on its underside and its black ear coverts.

Falco peregrinus furuitii, described by Momiyama in 1927, is found on the Izu and Ogasawara Islands south of Honshū, Japan. It is non-migratory. It is very rare and may only remain on a single island. It is a dark form, resembling pealei in colour, but darker, especially on the tail.

Falco peregrinus japonensis, described by Gmelin in 1788, includes kleinschmidti, pleskei, and harterti, and seems to refer to intergrades with calidus. It is found from northeast Siberia to Kamchatka (though it is possibly replaced by pealei on the coast there) and Japan. Northern populations are migratory, while those of Japan are resident. It is similar to the nominate subspecies, but the young are even darker than those of anatum.

Falco peregrinus macropus, described by Swainson in 1837, is the Australian peregrine falcon. It is found in Australia in all regions except the southwest. It is non-migratory. It is similar to brookei in appearance, but is slightly smaller and the ear region is entirely black. The feet are proportionally large.

Falco peregrinus madens, described by Ripley and Watson in 1963, is unusual in having some sexual dichromatism. If the Barbary falcon (see below) is considered a distinct species, it is sometimes placed therein. It is found in the Cape Verde Islands and is non-migratory; it is also endangered, with only six to eight pairs surviving. Males have a rufous wash on the crown, nape, ears and back; the underside is conspicuously washed pinkish-brown. Females are tinged rich brown overall, especially on the crown and nape.

 

Falco peregrinus minor, first described by Bonaparte in 1850. It was formerly often known as perconfusus. It is sparsely and patchily distributed throughout much of sub-Saharan Africa and widespread in Southern Africa. It apparently reaches north along the Atlantic coast as far as Morocco. It is non-migratory and dark-coloured. This is the smallest subspecies, with smaller males weighing as little as approximately 300 g (11 oz).

Falco peregrinus nesiotes, described by Mayr in 1941, is found in Fiji and probably also Vanuatu and New Caledonia. It is non-migratory.

Falco peregrinus pealei, described by Ridgway in 1873, is Peale's falcon and includes rudolfi. It is found in the Pacific Northwest of North America, northwards from Puget Sound along the British Columbia coast (including the Haida Gwaii), along the Gulf of Alaska and the Aleutian Islands to the far eastern Bering Sea coast of Russia, and may also occur on the Kuril Islands and the coasts of Kamchatka. It is non-migratory. It is the largest subspecies and it looks like an oversized and darker tundrius or like a strongly barred and large anatum. The bill is very wide. Juveniles occasionally have pale crowns. Males weigh 700 to 1,000 g (1.5–2.2 lb), while females weigh 1,000 to 1,500 g (2.2–3.3 lb).

Falco peregrinus pelegrinoides, first described by Temminck in 1829, is found in the Canary Islands through North Africa and the Near East to Mesopotamia. It is most similar to brookei, but is markedly paler above, with a rusty neck, and is a light buff with reduced barring below. It is smaller than the nominate subspecies; females weigh around 610 g (1.34 lb).

Falco peregrinus peregrinator, described by Sundevall in 1837, is known as the Indian peregrine falcon, black shaheen, Indian shaheen or shaheen falcon. It was formerly sometimes known as Falco atriceps or Falco shaheen. Its range includes South Asia from across the Indian subcontinent to Sri Lanka and southeastern China. In India, the shaheen falcon is reported from all states except Uttar Pradesh, mainly from rocky and hilly regions. The shaheen falcon is also reported from the Andaman and Nicobar Islands in the Bay of Bengal. It has a clutch size of 3 to 4 eggs, with the chicks fledging time of 48 days with an average nesting success of 1.32 chicks per nest. In India, apart from nesting on cliffs, it has also been recorded as nesting on man-made structures such as buildings and cellphone transmission towers.[36] A population estimate of 40 breeding pairs in Sri Lanka was made in 1996. It is non-migratory and is small and dark, with rufous underparts. In Sri Lanka this species is found to favour the higher hills, while the migrant calidus is more often seen along the coast.

Falco peregrinus peregrinus, the nominate (first-named) subspecies, described by Tunstall in 1771, breeds over much of temperate Eurasia between the tundra in the north and the Pyrenees, Mediterranean region and Alpide belt in the south. It is mainly non-migratory in Europe, but migratory in Scandinavia and Asia. Males weigh 580 to 750 g (1.28–1.65 lb), while females weigh 925 to 1,300 g (2.039–2.866 lb). It includes brevirostris, germanicus, rhenanus and riphaeus.

Falco peregrinus radama, described by Hartlaub in 1861, is found in Madagascar and the Comoros. It is non-migratory.

Falco peregrinus submelanogenys, described by Mathews in 1912, is the Southwest Australian peregrine falcon. It is found in southwestern Australia and is non-migratory.

Falco peregrinus tundrius, described by C.M. White in 1968, was at one time included in leucogenys. It is found in the Arctic tundra of North America to Greenland, and migrates to wintering grounds in Central and South America. Most vagrants that reach western Europe belong to this subspecies, which was previously considered synonymous with anatum. It is the New World equivalent to calidus. It is smaller and paler than anatum; most have a conspicuous white forehead and white in ear region, but the crown and "moustache" are very dark, unlike in calidus. Juveniles are browner and less grey than in calidus and paler, sometimes almost sandy, than in anatum. Males weigh 500 to 700 g (1.1–1.5 lb), while females weigh 800 to 1,100 g (1.8–2.4 lb). Despite its current recognition as a valid subspecies, a population genetic study of both pre-decline (i.e., museum) and recovered contemporary populations failed to distinguish genetically the anatum and tundrius subspecies.

Barbary falcon

Main article: Barbary falcon

The Barbary falcon is a subspecies of the peregrine falcon that inhabits parts of North Africa; namely, from the Canary Islands to the Arabian Peninsula. There is discussion concerning the taxonomic status of the bird, with some considering it a subspecies of the peregrine falcon and others considering it a full species with two subspecies (White et al. 2013). Compared to the other peregrine falcon subspecies, Barbary falcons sport a slimmer body and a distinct plumage color pattern. Despite numbers and range of these birds throughout the Canary Islands generally increasing, they are considered endangered, with human interference through falconry and shooting threatening their well-being. Falconry can further complicate the speciation and genetics of these Canary Islands falcons, as the practice promotes genetic mixing between individuals from outside the islands with those originating from the islands. Population density of the Barbary falcons on Tenerife, the biggest of the seven major Canary Islands, was found to be 1.27 pairs/100 km², with the mean distance between pairs being 5869 ± 3338 m. The falcons were only observed near large and natural cliffs with a mean altitude of 697.6 m. Falcons show an affinity for tall cliffs away from human-mediated establishments and presence.

 

Barbary falcons have a red neck patch, but otherwise differ in appearance from the peregrine falcon proper merely according to Gloger's rule, relating pigmentation to environmental humidity. The Barbary falcon has a peculiar way of flying, beating only the outer part of its wings like fulmars sometimes do; this also occurs in the peregrine falcon, but less often and far less pronounced. The Barbary falcon's shoulder and pelvis bones are stout by comparison with the peregrine falcon and its feet are smaller. Barbary falcons breed at different times of year than neighboring peregrine falcon subspecies, but they are capable of interbreeding. There is a 0.6–0.7% genetic distance in the peregrine falcon-Barbary falcon ("peregrinoid") complex.

 

Ecology and behaviour

The peregrine falcon lives mostly along mountain ranges, river valleys, coastlines, and increasingly in cities. In mild-winter regions, it is usually a permanent resident, and some individuals, especially adult males, will remain on the breeding territory. Only populations that breed in Arctic climates typically migrate great distances during the northern winter.

 

The peregrine falcon reaches faster speeds than any other animal on the planet when performing the stoop, which involves soaring to a great height and then diving steeply at speeds of over 320 km/h (200 mph), hitting one wing of its prey so as not to harm itself on impact. The air pressure from such a dive could possibly damage a bird's lungs, but small bony tubercles on a falcon's nostrils are theorized to guide the powerful airflow away from the nostrils, enabling the bird to breathe more easily while diving by reducing the change in air pressure. To protect their eyes, the falcons use their nictitating membranes (third eyelids) to spread tears and clear debris from their eyes while maintaining vision. The distinctive malar stripe or 'moustache', a dark area of feathers below the eyes, is thought to reduce solar glare and improve contrast sensitivity when targeting fast moving prey in bright light condition; the malar stripe has been found to be wider and more pronounced in regions of the world with greater solar radiation supporting this solar glare hypothesis. Peregrine falcons have a flicker fusion frequency of 129 Hz (cycles per second), very fast for a bird of its size, and much faster than mammals. A study testing the flight physics of an "ideal falcon" found a theoretical speed limit at 400 km/h (250 mph) for low-altitude flight and 625 km/h (388 mph) for high-altitude flight. In 2005, Ken Franklin recorded a falcon stooping at a top speed of 389 km/h (242 mph).

 

The life span of peregrine falcons in the wild is up to 19 years 9 months. Mortality in the first year is 59–70%, declining to 25–32% annually in adults. Apart from such anthropogenic threats as collision with human-made objects, the peregrine may be killed by larger hawks and owls.

 

The peregrine falcon is host to a range of parasites and pathogens. It is a vector for Avipoxvirus, Newcastle disease virus, Falconid herpesvirus 1 (and possibly other Herpesviridae), and some mycoses and bacterial infections. Endoparasites include Plasmodium relictum (usually not causing malaria in the peregrine falcon), Strigeidae trematodes, Serratospiculum amaculata (nematode), and tapeworms. Known peregrine falcon ectoparasites are chewing lice, Ceratophyllus garei (a flea), and Hippoboscidae flies (Icosta nigra, Ornithoctona erythrocephala).

 

In the Arctic Peregrine falcons chasing away small rodent predators from their nesting territory and Rough-legged Buzzards (Buteo lagopus) could use these hot spots as a nesting territory.

 

Feeding

The peregrine falcon's diet varies greatly and is adapted to available prey in different regions. However, it typically feeds on medium-sized birds such as pigeons and doves, waterfowl, gamebirds, songbirds, parrots, seabirds, and waders. Worldwide, it is estimated that between 1,500 and 2,000 bird species, or roughly a fifth of the world's bird species, are predated somewhere by these falcons.The peregrine falcon preys on the most diverse range of bird species of any raptor in North America, with over 300 species and including nearly 100 shorebirds. Its prey can range from 3 g (0.11 oz) hummingbirds (Selasphorus and Archilochus ssp.) to the 3.1 kg (6.8 lb) sandhill crane, although most prey taken by peregrines weigh between 20 g (0.71 oz) (small passerines) and 1,100 g (2.4 lb) (ducks, geese, loons, gulls, capercaillies, ptarmigans and other grouse). Smaller hawks (such as sharp-shinned hawks) and owls are regularly predated, as well as smaller falcons such as the American kestrel, merlin and, rarely, other peregrines.

 

In urban areas, where it tends to nest on tall buildings or bridges, it subsists mostly on a variety of pigeons. Among pigeons, the rock or feral pigeon comprises 80% or more of the dietary intake of peregrines. Other common city birds are also taken regularly, including mourning doves, common wood pigeons, common swifts, northern flickers, common starlings, American robins, common blackbirds, and corvids such as magpies, jays or carrion, house, and American crows. Coastal populations of the large subspecies pealei feed almost exclusively on seabirds. In the Brazilian mangrove swamp of Cubatão, a wintering falcon of the subspecies tundrius was observed successfully hunting a juvenile scarlet ibis.

 

Among mammalian prey species, bats in the genera Eptesicus, Myotis, Pipistrellus and Tadarida are the most common prey which taken at night. Though peregrines generally do not prefer terrestrial mammalian prey, in Rankin Inlet, peregrines largely take northern collared lemmings (Dicrostonyx groenlandicus) along with a few Arctic ground squirrels (Urocitellus parryii). Other small mammals including shrews, mice, rats, voles, and squirrels are more seldom taken. Peregrines occasionally take rabbits, mainly young individuals and juvenile hares. Additionally, remains of red fox kits and adult female American marten were found among prey remains. Insects and reptiles such as small snakes make up a small proportion of the diet, and salmonid fish have been taken by peregrines.

 

The peregrine falcon hunts most often at dawn and dusk, when prey are most active, but also nocturnally in cities, particularly during migration periods when hunting at night may become prevalent. Nocturnal migrants taken by peregrines include species as diverse as yellow-billed cuckoo, black-necked grebe, virginia rail, and common quail. The peregrine requires open space in order to hunt, and therefore often hunts over open water, marshes, valleys, fields, and tundra, searching for prey either from a high perch or from the air. Large congregations of migrants, especially species that gather in the open like shorebirds, can be quite attractive to a hunting peregrine. Once prey is spotted, it begins its stoop, folding back the tail and wings, with feet tucked. Prey is typically struck and captured in mid-air; the peregrine falcon strikes its prey with a clenched foot, stunning or killing it with the impact, then turns to catch it in mid-air. If its prey is too heavy to carry, a peregrine will drop it to the ground and eat it there. If they miss the initial strike, peregrines will chase their prey in a twisting flight.

 

Although previously thought rare, several cases of peregrines contour-hunting, i.e., using natural contours to surprise and ambush prey on the ground, have been reported and even rare cases of prey being pursued on foot. In addition, peregrines have been documented preying on chicks in nests, from birds such as kittiwakes. Prey is plucked before consumption. A 2016 study showed that the presence of peregrines benefits non-preferred species while at the same time causing a decline in its preferred prey. As of 2018, the fastest recorded falcon was at 242 mph (nearly 390 km/h). Researchers at the University of Groningen in the Netherlands and at Oxford University used 3D computer simulations in 2018 to show that the high speed allows peregrines to gain better maneuverability and precision in strikes.

 

Reproduction

The peregrine falcon is sexually mature at one to three years of age, but in larger populations they breed after two to three years of age. A pair mates for life and returns to the same nesting spot annually. The courtship flight includes a mix of aerial acrobatics, precise spirals, and steep dives. The male passes prey it has caught to the female in mid-air. To make this possible, the female actually flies upside-down to receive the food from the male's talons.

 

During the breeding season, the peregrine falcon is territorial; nesting pairs are usually more than 1 km (0.62 mi) apart, and often much farther, even in areas with large numbers of pairs. The distance between nests ensures sufficient food supply for pairs and their chicks. Within a breeding territory, a pair may have several nesting ledges; the number used by a pair can vary from one or two up to seven in a 16-year period.

 

The peregrine falcon nests in a scrape, normally on cliff edges. The female chooses a nest site, where she scrapes a shallow hollow in the loose soil, sand, gravel, or dead vegetation in which to lay eggs. No nest materials are added. Cliff nests are generally located under an overhang, on ledges with vegetation. South-facing sites are favoured. In some regions, as in parts of Australia and on the west coast of northern North America, large tree hollows are used for nesting. Before the demise of most European peregrines, a large population of peregrines in central and western Europe used the disused nests of other large birds. In remote, undisturbed areas such as the Arctic, steep slopes and even low rocks and mounds may be used as nest sites. In many parts of its range, peregrines now also nest regularly on tall buildings or bridges; these human-made structures used for breeding closely resemble the natural cliff ledges that the peregrine prefers for its nesting locations.

 

The pair defends the chosen nest site against other peregrines, and often against ravens, herons, and gulls, and if ground-nesting, also such mammals as foxes, wolverines, felids, bears, wolves, and mountain lions. Both nests and (less frequently) adults are predated by larger-bodied raptorial birds like eagles, large owls, or gyrfalcons. The most serious predators of peregrine nests in North America and Europe are the great horned owl and the Eurasian eagle-owl. When reintroductions have been attempted for peregrines, the most serious impediments were these two species of owls routinely picking off nestlings, fledglings and adults by night. Peregrines defending their nests have managed to kill raptors as large as golden eagles and bald eagles (both of which they normally avoid as potential predators) that have come too close to the nest by ambushing them in a full stoop. In one instance, when a snowy owl killed a newly fledged peregrine, the larger owl was in turn killed by a stooping peregrine parent.

 

The date of egg-laying varies according to locality, but is generally from February to March in the Northern Hemisphere, and from July to August in the Southern Hemisphere, although the Australian subspecies macropus may breed as late as November, and equatorial populations may nest anytime between June and December. If the eggs are lost early in the nesting season, the female usually lays another clutch, although this is extremely rare in the Arctic due to the short summer season. Generally three to four eggs, but sometimes as few as one or as many as five, are laid in the scrape. The eggs are white to buff with red or brown markings. They are incubated for 29 to 33 days, mainly by the female, with the male also helping with the incubation of the eggs during the day, but only the female incubating them at night. The average number of young found in nests is 2.5, and the average number that fledge is about 1.5, due to the occasional production of infertile eggs and various natural losses of nestlings.

 

After hatching, the chicks (called "eyases") are covered with creamy-white down and have disproportionately large feet. The male (called the "tiercel") and the female (simply called the "falcon") both leave the nest to gather prey to feed the young. The hunting territory of the parents can extend a radius of 19 to 24 km (12 to 15 mi) from the nest site. Chicks fledge 42 to 46 days after hatching, and remain dependent on their parents for up to two months.

 

Relationship with humans

The peregrine falcon is a highly admired falconry bird, and has been used in falconry for more than 3,000 years, beginning with nomads in central Asia. Its advantages in falconry include not only its athleticism and eagerness to hunt, but an equable disposition that leads to it being one of the easier falcons to train. The peregrine falcon has the additional advantage of a natural flight style of circling above the falconer ("waiting on") for game to be flushed, and then performing an effective and exciting high-speed diving stoop to take the quarry. The speed of the stoop not only allows the falcon to catch fast flying birds, it also enhances the falcon's ability to execute maneuvers to catch highly agile prey, and allows the falcon to deliver a knockout blow with a fist-like clenched talon against game that may be much larger than itself.

 

Additionally the versatility of the species, with agility allowing capture of smaller birds and a strength and attacking style allowing capture of game much larger than themselves, combined with the wide size range of the many peregrine subspecies, means there is a subspecies suitable to almost any size and type of game bird. This size range, evolved to fit various environments and prey species, is from the larger females of the largest subspecies to the smaller males of the smallest subspecies, approximately five to one (approximately 1500 g to 300 g). The males of smaller and medium-sized subspecies, and the females of the smaller subspecies, excel in the taking of swift and agile small game birds such as dove, quail, and smaller ducks. The females of the larger subspecies are capable of taking large and powerful game birds such as the largest of duck species, pheasant, and grouse.

 

Peregrine falcons handled by falconers are also occasionally used to scare away birds at airports to reduce the risk of bird-plane strikes, improving air-traffic safety. They were also used to intercept homing pigeons during World War II.

 

Peregrine falcons have been successfully bred in captivity, both for falconry and for release into the wild. Until 2004 nearly all peregrines used for falconry in the US were captive-bred from the progeny of falcons taken before the US Endangered Species Act was enacted and from those few infusions of wild genes available from Canada and special circumstances. Peregrine falcons were removed from the United States' endangered species list in 1999. The successful recovery program was aided by the effort and knowledge of falconers – in collaboration with The Peregrine Fund and state and federal agencies – through a technique called hacking. Finally, after years of close work with the US Fish and Wildlife Service, a limited take of wild peregrines was allowed in 2004, the first wild peregrines taken specifically for falconry in over 30 years.

 

The development of captive breeding methods has led to peregrines being commercially available for falconry use, thus mostly eliminating the need to capture wild birds for support of falconry. The main reason for taking wild peregrines at this point is to maintain healthy genetic diversity in the breeding lines. Hybrids of peregrines and gyrfalcons are also available that can combine the best features of both species to create what many consider to be the ultimate falconry bird for the taking of larger game such as the sage-grouse. These hybrids combine the greater size, strength, and horizontal speed of the gyrfalcon with the natural propensity to stoop and greater warm weather tolerance of the peregrine.

 

Decline due to pesticides

The peregrine falcon became an endangered species over much of its range because of the use of organochlorine pesticides, especially DDT, during the 1950s, '60s, and '70s. Pesticide biomagnification caused organochlorine to build up in the falcons' fat tissues, reducing the amount of calcium in their eggshells. With thinner shells, fewer falcon eggs survived until hatching. In addition, the PCB concentrations found in these falcons is dependent upon the age of the falcon. While high levels are still found in young birds (only a few months old) and even higher concentrations are found in more mature falcons, further increasing in adult peregrine falcons. These pesticides caused falcon prey to also have thinner eggshells (one example of prey being the Black Petrels). In several parts of the world, such as the eastern United States and Belgium, this species became extirpated (locally extinct) as a result. An alternate point of view is that populations in the eastern North America had vanished due to hunting and egg collection. Following the ban of organochlorine pesticides, the reproductive success of Peregrines increased in Scotland in terms of territory occupancy and breeding success, although spatial variation in recovery rates indicate that in some areas Peregrines were also impacted by other factors such as persecution.

 

Recovery efforts

Peregrine falcon recovery teams breed the species in captivity. The chicks are usually fed through a chute or with a hand puppet mimicking a peregrine's head, so they cannot see to imprint on the human trainers. Then, when they are old enough, the rearing box is opened, allowing the bird to train its wings. As the fledgling gets stronger, feeding is reduced, forcing the bird to learn to hunt. This procedure is called hacking back to the wild. To release a captive-bred falcon, the bird is placed in a special cage at the top of a tower or cliff ledge for some days or so, allowing it to acclimate itself to its future environment.

 

Worldwide recovery efforts have been remarkably successful. The widespread restriction of DDT use eventually allowed released birds to breed successfully. The peregrine falcon was removed from the U.S. Endangered Species list on 25 August 1999.

 

Some controversy has existed over the origins of captive breeding stock used by the Peregrine Fund in the recovery of peregrine falcons throughout the contiguous United States. Several peregrine subspecies were included in the breeding stock, including birds of Eurasian origin. Due to the extirpation of the eastern population of Falco peregrinus anatum, the near-extirpation of anatum in the Midwest and the limited gene pool within North American breeding stock, the inclusion of non-native subspecies was justified to optimize the genetic diversity found within the species as a whole.

 

During the 1970s, peregrine falcons in Finland experienced a population bottleneck as a result of large declines associated with bio-accumulation of organochloride pesticides. However, the genetic diversity of peregrines in Finland is similar to other populations, indicating that high dispersal rates have maintained the genetic diversity of this species.

 

Since peregrine falcon eggs and chicks are still often targeted by illegal poachers, it is common practice not to publicize unprotected nest locations.

 

Current status

Populations of the peregrine falcon have bounced back in most parts of the world. In the United Kingdom, there has been a recovery of populations since the crash of the 1960s. This has been greatly assisted by conservation and protection work led by the Royal Society for the Protection of Birds. The RSPB estimated that there were 1,402 breeding pairs in the UK in 2011. In Canada, where peregrines were identified as endangered in 1978 (in the Yukon territory of northern Canada that year, only a single breeding pair was identified), the Committee on the Status of Endangered Wildlife in Canada declared the species no longer at risk in December 2017.

 

Peregrines now breed in many mountainous and coastal areas, especially in the west and north, and nest in some urban areas, capitalising on the urban feral pigeon populations for food. Additionally, falcons benefit from artificial illumination, which allows the raptors to extend their hunting periods into the dusk when natural illumination would otherwise be too low for them to pursue prey. In England, this has allowed them to prey on nocturnal migrants such as redwings, fieldfares, starlings, and woodcocks.

 

In many parts of the world peregrine falcons have adapted to urban habitats, nesting on cathedrals, skyscraper window ledges, tower blocks, and the towers of suspension bridges. Many of these nesting birds are encouraged, sometimes gathering media attention and often monitored by cameras.

 

In England, peregrine falcons have become increasingly urban in distribution, particularly in southern areas where inland cliffs suitable as nesting sites are scarce. The first recorded urban breeding pair was observed nesting on the Swansea Guildhall in the 1980s. In Southampton, a nest prevented restoration of mobile telephony services for several months in 2013, after Vodafone engineers despatched to repair a faulty transmitter mast discovered a nest in the mast, and were prevented by the Wildlife and Countryside Act – on pain of a possible prison sentence – from proceeding with repairs until the chicks fledged.

 

In Oregon, Portland houses ten percent of the state's peregrine nests, despite only covering around 0.1 percent of the state's land area.

 

Cultural significance

Due to its striking hunting technique, the peregrine has often been associated with aggression and martial prowess. The Ancient Egyptian solar deity Ra was often represented as a man with the head of a peregrine falcon adorned with the solar disk, although most Egyptologists agree that it's most likely a Lanner falcon. Native Americans of the Mississippian culture (c. 800–1500) used the peregrine, along with several other birds of prey, in imagery as a symbol of "aerial (celestial) power" and buried men of high status in costumes associating to the ferocity of raptorial birds. In the late Middle Ages, the Western European nobility that used peregrines for hunting, considered the bird associated with princes in formal hierarchies of birds of prey, just below the gyrfalcon associated with kings. It was considered "a royal bird, more armed by its courage than its claws". Terminology used by peregrine breeders also used the Old French term gentil, "of noble birth; aristocratic", particularly with the peregrine.

 

The peregrine falcon is the national animal of the United Arab Emirates. Since 1927, the peregrine falcon has been the official mascot of Bowling Green State University in Bowling Green, Ohio. The 2007 U.S. Idaho state quarter features a peregrine falcon. The peregrine falcon has been designated the official city bird of Chicago.

 

The Peregrine, by J. A. Baker, is widely regarded as one of the best nature books in English written in the twentieth century. Admirers of the book include Robert Macfarlane, Mark Cocker, who regards the book as "one of the most outstanding books on nature in the twentieth century" and Werner Herzog, who called it "the one book I would ask you to read if you want to make films", and said elsewhere "it has prose of the calibre that we have not seen since Joseph Conrad". In the book, Baker recounts, in diary form, his detailed observations of peregrines (and their interaction with other birds) near his home in Chelmsford, Essex, over a single winter from October to April.

 

An episode of the hour-long TV series Starman in 1986 titled "Peregrine" was about an injured peregrine falcon and the endangered species program. It was filmed with the assistance of the University of California's peregrine falcon project in Santa Cruz.

part of a large group of fallow deer Fallow of the white variety are not albinos but have normal eye pigmentation, although their hooves and noses might be somewhat paler than the other colour varieties. Unusually pale coats in any deer are usually a result of under-production of melanin, the chemical responsible for skin pigmentation (over-production results in darker than normal coats).

  

It is said to be lucky to see a white buck.

Leucistic

American Robin AMRO (Turdus migratorius)

 

Patricia Bay

North Saanich BC

 

DSCN8993 Cropped

 

Jeremy G first noticed this bird which i bird-nerded on to photo doc

 

Leucism - Wikipedia

en.wikipedia.org/wiki/Leucism

Leucism (/ˈljuːkɪzəm/; or /ˈluːsɪzəm/) is a condition in which there is partial loss of pigmentation in an animal resulting in white, pale, or patchy coloration of the skin, hair, feathers, scales or cuticle, but not the eyes. Unlike albinism, it is caused by a reduction in multiple types of pigment, not just melanin.

 

Leucism & albinism

www.bto.org/volunteer-surveys/gbw/gardens-

White squirrels are almost always a white version of the eastern grey squirrel. There are a few types of genetic aberrations that cause the white coats. The first is albinism, caused by a mutation on a gene that codes for pigmentation. Albinos have red eyes.

Lego Simpsons 71005

 

The Simpsons is an American adult animated sitcom created by Matt Groening for the Fox Broadcasting Company.The series is a satirical depiction of a middle class American lifestyle epitomized by its family of the same name, which consists of Homer, Marge, Bart, Lisa, and Maggie. The show is set in the fictional town of Springfield and parodies American culture, society, television, and many aspects of the human condition.

  

The family was conceived by Groening shortly before a solicitation for a series of animated shorts with the producer James L. Brooks. Groening created a dysfunctional family and named the characters after members of his own family, substituting Bart for his own name. The shorts became a part of The Tracey Ullman Show on April 19, 1987. After a three-season run, the sketch was developed into a half-hour prime time show and was an early hit for Fox, becoming the network's first series to land in the Top 30 ratings in a season (1989–1990).

  

Since its debut on December 17, 1989, the show has broadcast 548 episodes and the 25th season began on September 30, 2013. The Simpsons is the longest-running American sitcom, the longest-running American animated program, and in 2009 it surpassed Gunsmoke as the longest-running American primetime, scripted television series. The Simpsons Movie, a feature-length film, was released in theaters worldwide on July 26 and 27, 2007, and grossed over $527 million.

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Les Simpson (The Simpsons) est une série télévisée d'animation américaine créée par Matt Groening et diffusée depuis le 17 décembre 1989 sur le réseau FOX.

  

Elle met en scène les Simpson, stéréotype d'une famille de classe moyenne. Leurs aventures servent une satire du mode de vie américain. Les membres de la famille, tous ayant la pigmentation de peau de couleur jaune, sont Homer, Marge, Bart, Lisa et Maggie.

  

[My Portfotolio] [My Google + ]

  

Thanks to all for visits and faves :)

  

[My GETTY Images @] [My MOST FAVE on Flickriver] [My RECENT on Fluidr] [My STREAM on Darckr]

 

Nyah is part of Inamorata Vitiligo collection that celebrates the beauty of this unique type of pigmentation. The collection consists of three dolls in Chocolate resin: Nyah (Nnaji sculpt), Nala (Nnaji sculpt) and Imani (Nubia sculpt).

 

Nala has one blue and one brown eye, black lashes, red glossy lips and vitiligo pigmentation. The white lingerie is from Inamorata Cherub LE30 from 2013.

 

The jewellery and dolls are available for sale in my shop at emiliacouture.com/shop/

Dactylorhiza fuchsii var. rhodochila.

 

When I was away, a friend sent me news of after several months of searching, he had found this striking and rare variant of the CSO.

 

So today, I met with him and we set off from a housing estate into the woods, and there it was.

 

There they were.

 

Four spikes, one frazzled, but three in good shape, and well worth the hour long drive to get there.

 

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The Common Spotted-orchid is one of the few species that are still thriving in Britain despite the ongoing environmental damage and habitat destruction that has caused so many other orchids to decline. Its secret is an adaptability that enables it to colonise new areas. Few of us can have failed to spot the vast numbers of Dactylorhiza fuchsii on urban roadside verges and roundabouts - this orchid is living among us. It lives up to its common name on both counts: this orchid is indeed common, and its leaves are profusely marked with dark spots and blotches. Although best known from alkaline habitats such as fens, dune slacks, old quarries and lime pits, the Common Spotted-orchid is also tolerant of mildly acidic substrates and so can sometimes be found on heathland among heather. In Britain and Ireland this orchid flowers from mid May to the end of July. On mainland Europe the range of Common Spotted-orchid extends from Scandinavia in the north through central Europe and southwards to the Mediterranean.

 

Plant: 7 to 30cm, occasionally to 50cm, stem flushed pink-to-purple towards the top.

Leaves: up to 7 semi-erect lanceolate-oval in basal rosette, blue-green and unspotted.

Bracts: 2 to 3 smaller bract-like leaves below the flower.

Flowers: Relatively few and quite large, mainly pink-purple but often paler pink or almost white. The sepals and petals form a 'hood' that encloses the column. The lip is broad and divided into 3 lobes, often with dark purple spots in the centre. The spur is the same colour as the lip and flattened towards the tip. The flowers are reputed to have a delicate scent reminiscent of vanilla.

 

The specific name 'fuchsii' refers to the German botanist Leonard Fuchs.

Subspecies: There is one subspecies found in Britain: Dactylorhiza fuchsii subsp. hebridensis which is found in Scotland, the Outer Hebrides and western Ireland.

Varieties: Dactylorhiza fuchsii var. albiflora has unmarked white flowers and is fairly widespread. Dactylorhiza fuchsii var. alpina has small, darker flowers and is found in Scotland and parts of northern England; it has also been reported from Wales. Dactylorhiza fuchsii var. cornubiensis is a smaller plant with a relatively large infloresence and larger flowers; it is found in Cornwall. Dactylorhiza fuchsii var. albiflora has unmarked white flowers and unspotted leaves; it is widespread but uncommon. Dactylorhiza fuchsii var. okellyii is somewhat contentious: it resembles Dactylorhiza fuchsii var. albiflora but the leaves can be either spotted or unspotted; this variety is thought to be confined to western Ireland, the Isle of Man and parts of Scotland. Dactylorhiza fuchsii var. rhodochila is a widespread but rare hyperchromatic variant with excessive pigmentation; the lips of its flowers are reddish purple with a paler border, and the leaves can have either heavily marked or completely purple leaves.

Hybrids: Dactylorhiza x transiens is the hybrid with Heath Spotted-orchid Dactylorhiza maculata. Dactylorhiza x kernerorum is the hybrid with Early Marsh-orchid Dactylorhiza incarnata. Dactylorhiza x mixtum is a rare hybrid with the Frog Orchid Dactylorhiza viridis. Dactylorhiza x venusta is the hybrid with Northern Marsh-orchid Dactylorhiza purpurella. Dactylorhiza x grandis is the hybrid with Southern Marsh-orchid and is widespread in southern Britain. Dactylorhiza x silvae-gabretae is the hybrid with Narrow-leaved Marsh-orchid Dactylorhiza traunsteinerioides. Dactylorhiza x braunii is the hybrid with Irish Marsh-orchid Dactylorhiza occidentalis and is recorded from County Clare.

There is also an intergeneric hybrid. X Dactylodenia st-quintinii is the hybrid with Fragrant Orchid Gymnadenia conopsea and possibly with Britain's other two Fragrant Orchids

 

www.hardyorchidsociety.org.uk/hos%201012/orchidphotos/dac...

The black or “melanistic” Chipmunk is opposite to an albino Chipmunk (an albino has a complete lack of pigmentation whereas the black chipmunk has far too much pigmentation.) This is the first time I have ever seen one. Depending on the internet site, they can be spotted infrequently or can be quite rare. Canon 7D, 400 f5.6, Ottawa, Ontario.

  

Brussels

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One of the artistic goals I've set for 2016 is for my photos to have a better representation of "me" in them. I've found that while many of my photos look nice, they don't truly show who I am as a person, photographer, or artist. Consider this the start of more personality in my work.

 

When CoverGirl announced that they were creating a line of Star Wars makeup in honor of The Force Awakens, I lost it. Makeup is one of my favorite things, as is Star Wars. I've managed to collect 3 of the lipsticks, and only one mascara (truth be told, I was not impressed with the mascara, but couldn't refuse the packaging!). My mornings always begin with a cup of coffee, and since seeing the film, this Stormtrooper mug is a new favorite. After coffee is glam time, and I keep reaching for these lightsaber looking lipsticks - the nude, shimmer-y one in the back (70) is my absolute favorite. High pigmentation, long lasting, and the tube says Star Wars on it. You really can't go wrong!

 

Thanks for your views, comments, and favorites!

  

Típica pigmentación de algunos elefantes indios.

Built in 1919-1920, this Chicago School and Sullivanesque-style building was designed by Louis Sullivan for the Farmers and Merchants Union Bank in Columbus, Wisconsin as one of his late-career “jewel box” bank buildings that are largely located in smaller communities throughout the midwest. The building was the last “jewel box bank” designed by Sullivan, and the second-to-last commission of his career, and was intended to communicate the bank as a modern and progressive institution, rather than employing the stodgier and more traditional Classical design found on most other banks of the era. The bank was commissioned by the president of the bank, J. Russell Wheeler, whose wife, Anna May Wheeler, pushed him to commission Sullivan to design a new home for the bank. In addition to Louis Sullivan, the building’s stained glass windows, were designed by architectural decorator Louis J. Millet, and the terra cotta by clay modeler Kristian Schneider, whom developed moulds for the building’s terra cotta, metal, and plaster details. The two artisans worked alongside Sullivan on several other bank projects. The building was heavily documented in Sullivan’s 1924 “A System of Architectural Ornament”, published shortly before he died.

 

The building is clad in red tapestry brick, which features blue and green mixed with the red clay mixture in some bricks, creating variation in color and texture across the facade. The brick creates a backdrop to some of the best terra cotta on any of Sullivan’s projects. The terra cotta features many of the floral and geometric motifs found on Sullivan’s other works, and is arranged similarly to other Sullivan banks that utilized brick cladding. The building features two principal facades, with a narrower facade along James Street, and a broader facade facing Dickason Boulevard. The James Street facade features two openings close to ground level, with the eastern bay housing a large plate glass window, and the western bay housing a doorway flanked by skylights, both of which are recessed under a large terra cotta architrave and flanked by square pilasters with decorative Sullivanesque ornament panels at the capitals. The architrave above the doorway and window is divided into three segments by vertical terra cotta elements that feature floral motifs and, like many Sullivan buildings, appear like plants with roots, branches, and crowns. The outer panels of the architrave feature circular cartouches with hexagonal trim, leaves, and geometric elements, with circular central medallions featuring the years 1861, when the bank was founded, and 1919, when the bank was completed. The central panel is clad in marble with the words “Farmers & Merchants Union Bank” and “Louis Sullivan, Architect” engraved into the stone with yellow pigmentation, contrasting against the white and green marble background. Atop the two vertical elements on either side of the central panel are griffin sculptures holding shields, a common element on many of Sullivan’s “Jewel Box Banks,” while the base of the outer vertical elements features the initials of the bank at the base. Above the architrave is an arched bay that houses a stained glass window, trimmed with decorative terra cotta at the inner and outer rings of the arch, with the bay becoming more recessed after each concentric arch, much like the entrances to medieval Romanesque churches. Besides a band of belt coursing that runs on either side of the architrave and wraps the corner to a tapered buttress on the Dickason Boulevard facade, the only other adornment is an eagle sculpture on a vertical trim element at the center of the parapet, which terminates many brick courses above the arched opening below, and another band of terra cotta trim along the top of the parapet, which forms a cap on the parapet around the perimeter of the building’s low-slope roof. On the Dickason Boulevard facade, the building features five recessed clerestory arched bays housing stained glass windows, flanked by tapered buttresses. Surrounding the arched tops of the windows are decorative trim panels with floral motifs, which begin just below the base of the arches, and extend up above the top of the arches, terminating in a band of belt coursing. Atop the buttresses at either end are trim elements featuring large spheres atop rectilinear legs with floral motifs below, undulating in and out with the brick below. Additionally, a band of belt coursing, which wraps the corner of Dickson Boulevard and James Street, runs beneath the windows, only interrupted by the buttresses. Toward the back, on the building’s original rear wing, there are three windows at eye level in the original building, with bands of belt coursing below and at the top of the parapet. The rear window is a recessed bay window flanked by two pilasters with sullivanesque terra cotta panels, while the smaller windows are flanked by sullivanesque relief panels. The rear wing features a roof at multiple heights, and was extended in 1961 with a matching addition by Law, Potter and Nystrom, since removed. The rear of the taller portion of the building features a simple recessed bay with an arched window, and a similar eagle sculpture and vertical trim piece as on the front facade.

 

Inside, the front wing of the building features a tall banking hall with brick cladding on the walls up to the level of the windows, where it terminates at a wooden sill. The space is split down the middle by a row of brick piers and low walls framing the teller cages, which terminate at the sill line of the windows, dividing the space while still allowing it to read as a single continuous lofty space. The brick forms piers at the teller’s cages, pilasters separating desks on the exterior wall, and low brick walls with marble caps. The upper portion of the walls and the coffered ceiling in this space is finished with white plaster, which gives the space a very vertical and airy feeling, as do the cream-colored terrazzo floors, which feature black edges at the base of the walls, tying the space together. The space features a terra cotta water fountain, or bubbler, also designed by sullivan, which features intricate ornament by Schneider. The space also features two mezzanine balconies with metal railings that run below the arched windows at the front and rear of the space, allowing managers to observe the activities in the lobby and teller area below from the rear balcony, while the front balcony exists solely to balance the space and keep it symmetrical. An office for private conferences with customers was originally located near the front of the space, along with a manager’s office, allowing convenience for customers seeking a meeting with the bank management. The teller’s side of the space also housed the bank’s two vaults and several other private offices. The bank originally featured a large meeting room in the one-story rear wing, behind the vaults, with a women’s waiting room sitting along the Dickason Boulevard side of the rear wing, featuring a bay window and a restroom. The building’s interior has changed in function somewhat due to the growth of the bank, changes in bank operations, and expansion of the building with new additions to house offices and a drive-through in the rear.

 

The building was listed on the National Register of Historic Places in 1972, was designated a National Historic Landmark in 1976, and is a contributing structure in the Columbus Downtown Historic District, listed on the National Register of Historic Places in 1992. The building saw an addition in 2006, clad in buff brick, which replicated a historic building that formerly stood to the east, and wraps the building to the rear, with a two-story section behind a one-story annex that connects the one-story rear wing of the bank to the new building. This wing replaced older additions made in 1961, which matched the one-story rear wing of the historic building, and 1980, which was modern in appearance and slightly recessed along James Street to give precedence to the historic building. The building still functions as the main office branch of the Farmers and Merchants Union Bank, which has grown substantially. The building has been long considered to be among the best of Sullivan’s “Jewel Box Banks,” and has been kept in excellent condition by the bank’s careful and caring generational stewardship.

In the center (left to right) and about 4 "rows" of birds from the front is a mostly white shearwater. Calling a bird "leucistic" means that it is lacking, to some degree, its normal pigmentation. The greenish color on the (normally white) undersides of these birds is due to reflection from the water.

 

(Primarily) Black-vented Shearwaters, Puffinus opisthomelas. This is a minuscule portion of the entire flock, which stretched for miles.

 

Offshore, San Luis Obispo County, California, USA.

 

The use of any of my photos, of any file size, for any purpose, is subject to approval by me. Contact me for permission. Image files are available upon request. My email address is available at my Flickr profile page. Or send me a FlickrMail.

 

We bought this flowering plant in a hanging planter at the hardware store - the label says it is a Viola. Viola is a genus with 600 species, so to be more specific I think it is a Viola tricolor, which is a common European wild flower, or a hybrid cultivar of similar appearance. The flower is unusual with two upper petals of solid color, and three lower petals with contrasting varigated pigmentation. Common names include wild pansy, Johnny Jump up, heartsease, heart's delight, tickle-my-fancy, Jack-jump-up-and-kiss-me, come-and-cuddle-me, and three faces in a hood. It has a long history in both folk medicine, folklore, and literature.

Could the Ancestral Puebloan used this boulder to clean their freshly killed game? Or is this fire retardant that was used when the wild fire broke out?

I didn't used to like freckles on the nose, lips and eyelinings of orange cats. They looked like blemishes to me. Now I have a cat who has freckles and I think they're cute. Which is a good thing because Mack's getting more and more the older he gets.

 

View on black. In fact, it looks much better on black - the freckles show up much better. This morning I tried to figure out how to add black frames or borders to my pictures in Photoshop Elements. I guess it's obvious I didn't succeed.

 

Explored June 25, 2012

Victor Hasselblad Sea Turtle Research And Conservation Centre

Kosgoda

Sri Lanka

 

Albinism and leucism are two types of color mutations involving color loss producing different phenotypes. There are different definitions of leucism, with some authors referring to it as partial albinism

However, true albinos are easily differentiated, because they lack pigmentation and feature red or pinkish eyes, while leucistic animals have reduced/absent coloration but with normal eye pigmentation (Krecsak 2008; Turner 2011).

 

They referred to this turtle as albino,but I guess this one is rather leucistic than albino as the eyes were dark and not red or pinkish.

The equivalent of albinism in animals, erythrism results from the inheritance of two recessive genes for the absence of pigmentation. Normally the katydid colour palette runs the gamut of greens, browns and yellows, colours which keep them camouflaged and aid in their survival. Although it has been hypothesized that pink coloration may increase survival rates amongst red vegetation it is much more likely that the genetic anomaly decreases fitness by increasing the insect's visibility to predators. Therefore it is likely that most individuals with this condition don't survive long and rarely make it to adulthood, which made this discovery all the more noteworthy. Found during a night hike in Vohimana reserve, Madagascar.

Scientific Name: Ursus maritimus

 

Description : Polar bears are considered the largest land carnivores in the world, matched only by very large individual Kodiak brown bears. Both sexes differ in size throughout their range; males being much larger than females and continuing to grow for a longer period of time. These bears have long, massive skulls, necks and bodies with long legs and large paws. Ears and tail are short. The nose is more prominent or “Roman†with a black rhinarium (nose pad). The tongue is black and the eyes brown. The surface of the skin is also black. Fur colour varies slightly with the season: new coats grown just prior to the winter season are very white appearing as slightly creamy white against the stark white of their icy winter environment. This pelage is thick, coarse and long with dense underfur. Guard hairs, found throughout the pelage, are shiny, almost glossy, oily and waterproof and have hollow shafts. Polar bears moult annually between the end of May and August. The coat becomes thinner and has a yellow wash or is almost a golden colour.

 

Male polar bears weigh between 400 – 600 kg, and have been recorded up to a maximum of 800 kg. Females are smaller than the males weighing up to 300 kg, and when pregnant up to 460 kg. Polar bears are 2.5 – 3.5 m long.

 

Distribution : The polar bear is circumpolar in distribution, inhabiting all Arctic seas and coastlines. It is found on the pack-ice off the Alaskan coast north of Bering Strait, off the coasts of Greenland and along the Eurasian Arctic coast from Spitsbergen to Wrangell Island. Rare stragglers reach Iceland. Individual bears have been seen on the frozen Arctic Ocean as far as latitude 88 degrees North, only 2 degrees from the North Pole. In Canada, they are found along the Arctic coasts from Alaska to Labrador and from the tip of James Bay to northern Ellesmere Island. Polar bears do roam as far as 150 kilometres inland into the coniferous forests, where they live very differently from the polar bears which belong to the high Arctic.

 

Habitat : They prefer areas of annual ice, which they use as a hunting platform and protective cover. This includes snow-drifted pressure ridges, refrozen cracks and areas of open water surrounded by ice. In areas where the pack ice melts by mid to late summer they come inland and live in coniferous forest areas. Here they remain until the ice re-freezes. Areas of solidly frozen sea ice and the open seas are avoided. Generally they are most common along coastal areas. Some do enter the permanent pack ice.

 

Food : The ringed seal is by far the most common prey. They also eat bearded seal, harp seal and hooded seal. Young walrus are sometimes taken. During the summer months they feed upon the shoreline carrion, fish, mussels, crabs, starfish, lemmings and the eggs and nestling young of waterfowl and cliff-dwelling birds. They will also graze on kelp, grasses and eat mushrooms and crowberries.

 

Reproduction and Development : Normally they are solitary animals outside the breeding season, the exception being a mother with cubs. Polar bears mate in mid-summer. Females first start to breed at 3 to 5 years of age. In April and May adult females are in oestrus and ready to accept a mate. They are polyandrous, meaning one female will mate with more than one male in one breeding season. Males fight among themselves for the female’s attention and a couple will pair off for a period of a few days to two weeks. With females, delayed implantation occurs, the fertilized egg does not implant in the uterus until mid-September to mid-October. Embryonic development begins at this time. Gestation periods, therefore, vary a great deal when including this period of delay. Females choose suitable locations to build their maternity dens in mid-October and retreat to them for the winter season. They give birth to one to four cubs somewhere between late November and early January. Twins are most common. The newborns are small; 25 to 30 cm long and weigh less than 1 kg. They are covered with very fine hair, appearing almost naked and their eyes are closed. Their eyes open at 6 weeks. Growth is very quick; at two months their fur has thickened; they weigh about 5 kg and move about the den. By mid-March to early April, when the den is opened, the cubs weigh about 10 kg and are surprisingly strong. The cubs suckle for nine months, occasionally one year. They are very dependent on their mother and stay with her for two years. At that time they weigh 90 - 180 kg and are half grown.

 

Adaptations : Polar bears are wonderfully adapted to their Arctic surroundings.

 

Locomotion. On land a shuffling walk may be increased to a rolling gallop of 40 km/h and can outrun caribou over a short distance. Bears are often seen standing high on their hind legs, necks stretched to scan the landscape. On thin ice, legs are spread to distribute body mass. Thickly padded and furred soles allow the bear to move quietly as well as providing good traction. Small bumps and cavities on the soles act like suction cups keeping bears from slipping on the ice. The claws are used to dig into icy slopes and to grip prey. They are strong swimmers, paddling with their forefeet only and trailing their hind feet which act as a rudder. They can stay submerged for over one minute, keeping their eyes open. They swim at a speed of approximately 6.4 km/h, often covering long distances.

 

Insulation. Polar bears have a thick layer of sub-cutaneous fat and very dense underfur with several layers of glossy guard hair on the outside. Their pelt is much thicker in winter and provides excellent insulation. The fat layer also adds to buoyancy in the water. Water is shed easily from the oily waterproof fur. Small, furry ears have a heavy network of blood vessels, keeping them warm and conserving heat. The tail is short and rounded also conserving heat. Fur is very dense around the soles of the feet.

 

Pelage. The creamy white appearance of the coat allows the bear to be inconspicuous when hunting seal. Each hair is similar to an optical fibre; colourless and hollow. Being translucent, it reflects the heat from the sun down to the base of the hair, where it is absorbed by the black skin. Whiteness comes from reflection of light rather than pigmentation.

 

Hunting. Bears use their keen sense of smell to detect seal breathing holes. These can be up to a kilometer away and covered by a layer of snow and ice. They will stand or lie by the seal’s blowhole in the ice for hours; they may swim towards seals resting on the ice flows with only their nose showing above the water. They will dive quietly, then swim up to the ice edge and jump out on the seal, and will also crawl towards a sunbathing seal using every piece of raised ice to conceal the approach.

 

Denning. Both sexes occupy dens for shelter. Topographic factors influence the den sites. In Canadian core areas, dens frequently occur on south-facing slopes where northerly prevailing winds create the best drifts, where the wind-chill is least and insulation from received solar radiation is greatest. One of the three largest denning areas worldwide is in Canada. There are three main types of winter refuges: maternity dens, temporary dens and winter shelters. During the winter any bear may dig a temporary den and use it for a few days during a storm, or take shelter in a natural cavity. Winter shelters are used for longer periods of time as resting places. This type of shelter is usually roomier with additional features such as alcoves, porches and ventilation holes. Bears do not hibernate in the strict sense of the word; they have the ability to slow down their metabolism to conserve energy at any time of year. The state of self-induced lethargy while in the shelter allows them to preserve their vital fat reserves. During this time, the body temperature of the polar bear decreases by a few degrees from normal and the respiration rates are markedly reduced. Maternity dens are built and occupied by pregnant females and can vary in size. The denning chamber is at the upper end of an entrance tunnel 1 to 2 m long. It averages 1.5 m in diameter with a height of 90 to 100cm in the middle. Drifting snow seals the entrance. The chamber is higher than the tunnel, trapping bear body heat inside. Dens not only provide a safe place to give birth to her cubs, but are also a place of protection for the cubs during their first few months. During this time she does not leave the den, remaining with her cubs and living on her reserves of fat. In the spring, with her fat severely depleted she must leave to find food to sustain herself and her cubs. After they leave the maternity den she will build temporary refuges in which to nurse, rest and shelter her young cubs and warms them as they all sleep together. She heads with her cubs towards the nearest supply of food, usually towards pack ice.

 

Sight and sounds. Polar bears have good eyesight. Their eyes have inner eyelids that keep the glare of the sun on snow and ice from blinding them. When defending a food source from other bears they use a deep growl. They hiss and snort to show aggression. Angry bears use loud roars and growls. Mothers scold cubs with a low growl.

 

Threats to Survival : Polar bears are one of the animals most threatened by global warming. They depend entirely on sea ice as a platform from which to hunt seals. Reduction of the total ice cover in the Arctic is a serious concern globally. When the ice does not form or forms too late in the season many polar bears starve. In Hudson Bay, scientists have found the main cause of death for cubs to be either lack of food or lack of fat on nursing mothers. Exploitation of minerals and fossil fuels in the Arctic pose a continuous threat. Of the oil and natural gas deposits globally, 20% are located in the Arctic. As the ice cap recedes these become more accessible. Countries are competing which each other in claiming ownership of Arctic and its resources. This can only result in further and more drastic impact on polar bear habitat.

 

Status : IUCN: Vulnerable; CITES: Appendix II; COSEWIC: Special Concern

 

Zoo Diet : Toronto Zoo carnivore diet, dog chow, jumbo smelt and herring, carrots, bean sprouts, Vitamin E and Thiamine supplements.

 

Toronto Zoo Website

Jellyfish, also known sea jellies, are the medusa-phase of certain gelatinous members of the subphylum Medusozoa, which is a major part of the phylum Cnidaria.

 

Jellyfish are mainly free-swimming marine animals with umbrella-shaped bells and trailing tentacles, although a few are anchored to the seabed by stalks rather than being mobile. The bell can pulsate to provide propulsion for highly efficient locomotion. The tentacles are armed with stinging cells and may be used to capture prey and defend against predators. Jellyfish have a complex life cycle. The medusa is normally the sexual phase, which produces planula larvae; these then disperse widely and enter a sedentary polyp phase, before reaching sexual maturity.

 

Jellyfish are found all over the world, from surface waters to the deep sea. Scyphozoans (the "true jellyfish") are exclusively marine, but some hydrozoans with a similar appearance live in freshwater. Large, often colorful, jellyfish are common in coastal zones worldwide. The medusae of most species are fast-growing, and mature within a few months then die soon after breeding, but the polyp stage, attached to the seabed, may be much more long-lived. Jellyfish have been in existence for at least 500 million years, and possibly 700 million years or more, making them the oldest multi-organ animal group.

 

Jellyfish are eaten by humans in certain cultures. They are considered a delicacy in some Asian countries, where species in the Rhizostomeae order are pressed and salted to remove excess water. Australian researchers have described them as a "perfect food": sustainable and protein-rich but relatively low in food energy.

 

They are also used in research, where the green fluorescent protein used by some species to cause bioluminescence has been adapted as a fluorescent marker for genes inserted into other cells or organisms.

 

The stinging cells used by jellyfish to subdue their prey can injure humans. Thousands of swimmers worldwide are stung every year, with effects ranging from mild discomfort to serious injury or even death. When conditions are favourable, jellyfish can form vast swarms, which can be responsible for damage to fishing gear by filling fishing nets, and sometimes clog the cooling systems of power and desalination plants which draw their water from the sea.

  

Names

The name jellyfish, in use since 1796, has traditionally been applied to medusae and all similar animals including the comb jellies (ctenophores, another phylum). The term jellies or sea jellies is more recent, having been introduced by public aquaria in an effort to avoid use of the word "fish" with its modern connotation of an animal with a backbone, though shellfish, cuttlefish and starfish are not vertebrates either. In scientific literature, "jelly" and "jellyfish" have been used interchangeably. Many sources refer to only scyphozoans as "true jellyfish".

 

A group of jellyfish is called a "smack" or a "smuck".

 

Definition

The term jellyfish broadly corresponds to medusae, that is, a life-cycle stage in the Medusozoa. The American evolutionary biologist Paulyn Cartwright gives the following general definition:

 

Typically, medusozoan cnidarians have a pelagic, predatory jellyfish stage in their life cycle; staurozoans are the exceptions [as they are stalked].

 

The Merriam-Webster dictionary defines jellyfish as follows:

 

A free-swimming marine coelenterate that is the sexually reproducing form of a hydrozoan or scyphozoan and has a nearly transparent saucer-shaped body and extensible marginal tentacles studded with stinging cells.

 

Given that jellyfish is a common name, its mapping to biological groups is inexact. Some authorities have called the comb jellies and certain salps jellyfish, though other authorities state that neither of these are jellyfish, which they consider should be limited to certain groups within the medusozoa.

 

The non-medusozoan clades called jellyfish by some but not all authorities (both agreeing and disagreeing citations are given in each case) are indicated with on the following cladogram of the animal kingdom:

 

Jellyfish are not a clade, as they include most of the Medusozoa, barring some of the Hydrozoa. The medusozoan groups included by authorities are indicated on the following phylogenetic tree by the presence of citations. Names of included jellyfish, in English where possible, are shown in boldface; the presence of a named and cited example indicates that at least that species within its group has been called a jellyfish.

 

Taxonomy

The subphylum Medusozoa includes all cnidarians with a medusa stage in their life cycle. The basic cycle is egg, planula larva, polyp, medusa, with the medusa being the sexual stage. The polyp stage is sometimes secondarily lost. The subphylum include the major taxa, Scyphozoa (large jellyfish), Cubozoa (box jellyfish) and Hydrozoa (small jellyfish), and excludes Anthozoa (corals and sea anemones). This suggests that the medusa form evolved after the polyps. Medusozoans have tetramerous symmetry, with parts in fours or multiples of four.

 

The four major classes of medusozoan Cnidaria are:

Scyphozoa are sometimes called true jellyfish, though they are no more truly jellyfish than the others listed here. They have tetra-radial symmetry. Most have tentacles around the outer margin of the bowl-shaped bell, and long, oral arms around the mouth in the center of the subumbrella.

Cubozoa (box jellyfish) have a (rounded) box-shaped bell, and their velarium assists them to swim more quickly. Box jellyfish may be related more closely to scyphozoan jellyfish than either are to the Hydrozoa.

Hydrozoa medusae also have tetra-radial symmetry, nearly always have a velum (diaphragm used in swimming) attached just inside the bell margin, do not have oral arms, but a much smaller central stalk-like structure, the manubrium, with terminal mouth opening, and are distinguished by the absence of cells in the mesoglea. Hydrozoa show great diversity of lifestyle; some species maintain the polyp form for their entire life and do not form medusae at all (such as Hydra, which is hence not considered a jellyfish), and a few are entirely medusal and have no polyp form.

Staurozoa (stalked jellyfish) are characterized by a medusa form that is generally sessile, oriented upside down and with a stalk emerging from the apex of the "calyx" (bell), which attaches to the substrate. At least some Staurozoa also have a polyp form that alternates with the medusoid portion of the life cycle. Until recently, Staurozoa were classified within the Scyphozoa.

There are over 200 species of Scyphozoa, about 50 species of Staurozoa, about 50 species of Cubozoa, and the Hydrozoa includes about 1000–1500 species that produce medusae, but many more species that do not.

 

Fossil history

Since jellyfish have no hard parts, fossils are rare. The oldest unambiguous fossil of a free-swimming medusa is Burgessomedusa from the mid Cambrian Burgess Shale of Canada, which is likely either a stem group of box jellyfish (Cubozoa) or Acraspeda (the clade including Staurozoa, Cubozoa, and Scyphozoa). Other claimed records from the Cambrian of China and Utah in the United States are uncertain, and possibly represent ctenophores instead.

 

Anatomy

The main feature of a true jellyfish is the umbrella-shaped bell. This is a hollow structure consisting of a mass of transparent jelly-like matter known as mesoglea, which forms the hydrostatic skeleton of the animal. 95% or more of the mesogloea consists of water, but it also contains collagen and other fibrous proteins, as well as wandering amoebocytes which can engulf debris and bacteria. The mesogloea is bordered by the epidermis on the outside and the gastrodermis on the inside. The edge of the bell is often divided into rounded lobes known as lappets, which allow the bell to flex. In the gaps or niches between the lappets are dangling rudimentary sense organs known as rhopalia, and the margin of the bell often bears tentacles.

  

Anatomy of a scyphozoan jellyfish

On the underside of the bell is the manubrium, a stalk-like structure hanging down from the centre, with the mouth, which also functions as the anus, at its tip. There are often four oral arms connected to the manubrium, streaming away into the water below. The mouth opens into the gastrovascular cavity, where digestion takes place and nutrients are absorbed. This is subdivided by four thick septa into a central stomach and four gastric pockets. The four pairs of gonads are attached to the septa, and close to them four septal funnels open to the exterior, perhaps supplying good oxygenation to the gonads. Near the free edges of the septa, gastric filaments extend into the gastric cavity; these are armed with nematocysts and enzyme-producing cells and play a role in subduing and digesting the prey. In some scyphozoans, the gastric cavity is joined to radial canals which branch extensively and may join a marginal ring canal. Cilia in these canals circulate the fluid in a regular direction.

  

Discharge mechanism of a nematocyst

The box jellyfish is largely similar in structure. It has a squarish, box-like bell. A short pedalium or stalk hangs from each of the four lower corners. One or more long, slender tentacles are attached to each pedalium. The rim of the bell is folded inwards to form a shelf known as a velarium which restricts the bell's aperture and creates a powerful jet when the bell pulsates, allowing box jellyfish to swim faster than true jellyfish. Hydrozoans are also similar, usually with just four tentacles at the edge of the bell, although many hydrozoans are colonial and may not have a free-living medusal stage. In some species, a non-detachable bud known as a gonophore is formed that contains a gonad but is missing many other medusal features such as tentacles and rhopalia. Stalked jellyfish are attached to a solid surface by a basal disk, and resemble a polyp, the oral end of which has partially developed into a medusa with tentacle-bearing lobes and a central manubrium with four-sided mouth.

 

Most jellyfish do not have specialized systems for osmoregulation, respiration and circulation, and do not have a central nervous system. Nematocysts, which deliver the sting, are located mostly on the tentacles; true jellyfish also have them around the mouth and stomach. Jellyfish do not need a respiratory system because sufficient oxygen diffuses through the epidermis. They have limited control over their movement, but can navigate with the pulsations of the bell-like body; some species are active swimmers most of the time, while others largely drift. The rhopalia contain rudimentary sense organs which are able to detect light, water-borne vibrations, odour and orientation. A loose network of nerves called a "nerve net" is located in the epidermis. Although traditionally thought not to have a central nervous system, nerve net concentration and ganglion-like structures could be considered to constitute one in most species. A jellyfish detects stimuli, and transmits impulses both throughout the nerve net and around a circular nerve ring, to other nerve cells. The rhopalial ganglia contain pacemaker neurones which control swimming rate and direction.

 

In many species of jellyfish, the rhopalia include ocelli, light-sensitive organs able to tell light from dark. These are generally pigment spot ocelli, which have some of their cells pigmented. The rhopalia are suspended on stalks with heavy crystals at one end, acting like gyroscopes to orient the eyes skyward. Certain jellyfish look upward at the mangrove canopy while making a daily migration from mangrove swamps into the open lagoon, where they feed, and back again.

 

Box jellyfish have more advanced vision than the other groups. Each individual has 24 eyes, two of which are capable of seeing colour, and four parallel information processing areas that act in competition, supposedly making them one of the few kinds of animal to have a 360-degree view of its environment.

 

Box jellyfish eye

The study of jellyfish eye evolution is an intermediary to a better understanding of how visual systems evolved on Earth. Jellyfish exhibit immense variation in visual systems ranging from photoreceptive cell patches seen in simple photoreceptive systems to more derived complex eyes seen in box jellyfish. Major topics of jellyfish visual system research (with an emphasis on box jellyfish) include: the evolution of jellyfish vision from simple to complex visual systems), the eye morphology and molecular structures of box jellyfish (including comparisons to vertebrate eyes), and various uses of vision including task-guided behaviors and niche specialization.

 

Evolution

Experimental evidence for photosensitivity and photoreception in cnidarians antecedes the mid 1900s, and a rich body of research has since covered evolution of visual systems in jellyfish. Jellyfish visual systems range from simple photoreceptive cells to complex image-forming eyes. More ancestral visual systems incorporate extraocular vision (vision without eyes) that encompass numerous receptors dedicated to single-function behaviors. More derived visual systems comprise perception that is capable of multiple task-guided behaviors.

 

Although they lack a true brain, cnidarian jellyfish have a "ring" nervous system that plays a significant role in motor and sensory activity. This net of nerves is responsible for muscle contraction and movement and culminates the emergence of photosensitive structures. Across Cnidaria, there is large variation in the systems that underlie photosensitivity. Photosensitive structures range from non-specialized groups of cells, to more "conventional" eyes similar to those of vertebrates. The general evolutionary steps to develop complex vision include (from more ancestral to more derived states): non-directional photoreception, directional photoreception, low-resolution vision, and high-resolution vision. Increased habitat and task complexity has favored the high-resolution visual systems common in derived cnidarians such as box jellyfish.

 

Basal visual systems observed in various cnidarians exhibit photosensitivity representative of a single task or behavior. Extraocular photoreception (a form of non-directional photoreception), is the most basic form of light sensitivity and guides a variety of behaviors among cnidarians. It can function to regulate circadian rhythm (as seen in eyeless hydrozoans) and other light-guided behaviors responsive to the intensity and spectrum of light. Extraocular photoreception can function additionally in positive phototaxis (in planula larvae of hydrozoans), as well as in avoiding harmful amounts of UV radiation via negative phototaxis. Directional photoreception (the ability to perceive direction of incoming light) allows for more complex phototactic responses to light, and likely evolved by means of membrane stacking. The resulting behavioral responses can range from guided spawning events timed by moonlight to shadow responses for potential predator avoidance. Light-guided behaviors are observed in numerous scyphozoans including the common moon jelly, Aurelia aurita, which migrates in response to changes in ambient light and solar position even though they lack proper eyes.

 

The low-resolution visual system of box jellyfish is more derived than directional photoreception, and thus box jellyfish vision represents the most basic form of true vision in which multiple directional photoreceptors combine to create the first imaging and spatial resolution. This is different from the high-resolution vision that is observed in camera or compound eyes of vertebrates and cephalopods that rely on focusing optics. Critically, the visual systems of box jellyfish are responsible for guiding multiple tasks or behaviors in contrast to less derived visual systems in other jellyfish that guide single behavioral functions. These behaviors include phototaxis based on sunlight (positive) or shadows (negative), obstacle avoidance, and control of swim-pulse rate.

 

Box jellyfish possess "proper eyes" (similar to vertebrates) that allow them to inhabit environments that lesser derived medusae cannot. In fact, they are considered the only class in the clade Medusozoa that have behaviors necessitating spatial resolution and genuine vision. However, the lens in their eyes are more functionally similar to cup-eyes exhibited in low-resolution organisms, and have very little to no focusing capability. The lack of the ability to focus is due to the focal length exceeding the distance to the retina, thus generating unfocused images and limiting spatial resolution. The visual system is still sufficient for box jellyfish to produce an image to help with tasks such as object avoidance.

 

Utility as a model organism

Box jellyfish eyes are a visual system that is sophisticated in numerous ways. These intricacies include the considerable variation within the morphology of box jellyfishes' eyes (including their task/behavior specification), and the molecular makeup of their eyes including: photoreceptors, opsins, lenses, and synapses. The comparison of these attributes to more derived visual systems can allow for a further understanding of how the evolution of more derived visual systems may have occurred, and puts into perspective how box jellyfish can play the role as an evolutionary/developmental model for all visual systems.

 

Characteristics

Box jellyfish visual systems are both diverse and complex, comprising multiple photosystems. There is likely considerable variation in visual properties between species of box jellyfish given the significant inter-species morphological and physiological variation. Eyes tend to differ in size and shape, along with number of receptors (including opsins), and physiology across species of box jellyfish.

 

Box jellyfish have a series of intricate lensed eyes that are similar to those of more derived multicellular organisms such as vertebrates. Their 24 eyes fit into four different morphological categories. These categories consist of two large, morphologically different medial eyes (a lower and upper lensed eye) containing spherical lenses, a lateral pair of pigment slit eyes, and a lateral pair of pigment pit eyes. The eyes are situated on rhopalia (small sensory structures) which serve sensory functions of the box jellyfish and arise from the cavities of the exumbrella (the surface of the body) on the side of the bells of the jellyfish. The two large eyes are located on the mid-line of the club and are considered complex because they contain lenses. The four remaining eyes lie laterally on either side of each rhopalia and are considered simple. The simple eyes are observed as small invaginated cups of epithelium that have developed pigmentation. The larger of the complex eyes contains a cellular cornea created by a mono ciliated epithelium, cellular lens, homogenous capsule to the lens, vitreous body with prismatic elements, and a retina of pigmented cells. The smaller of the complex eyes is said to be slightly less complex given that it lacks a capsule but otherwise contains the same structure as the larger eye.

 

Box jellyfish have multiple photosystems that comprise different sets of eyes. Evidence includes immunocytochemical and molecular data that show photopigment differences among the different morphological eye types, and physiological experiments done on box jellyfish to suggest behavioral differences among photosystems. Each individual eye type constitutes photosystems that work collectively to control visually guided behaviors.

 

Box jellyfish eyes primarily use c-PRCs (ciliary photoreceptor cells) similar to that of vertebrate eyes. These cells undergo phototransduction cascades (process of light absorption by photoreceptors) that are triggered by c-opsins. Available opsin sequences suggest that there are two types of opsins possessed by all cnidarians including an ancient phylogenetic opsin, and a sister ciliary opsin to the c-opsins group. Box jellyfish could have both ciliary and cnidops (cnidarian opsins), which is something not previously believed to appear in the same retina. Nevertheless, it is not entirely evident whether cnidarians possess multiple opsins that are capable of having distinctive spectral sensitivities.

 

Comparison with other organisms

Comparative research on genetic and molecular makeup of box jellyfishes' eyes versus more derived eyes seen in vertebrates and cephalopods focuses on: lenses and crystallin composition, synapses, and Pax genes and their implied evidence for shared primordial (ancestral) genes in eye evolution.

 

Box jellyfish eyes are said to be an evolutionary/developmental model of all eyes based on their evolutionary recruitment of crystallins and Pax genes. Research done on box jellyfish including Tripedalia cystophora has suggested that they possess a single Pax gene, PaxB. PaxB functions by binding to crystallin promoters and activating them. PaxB in situ hybridization resulted in PaxB expression in the lens, retina, and statocysts. These results and the rejection of the prior hypothesis that Pax6 was an ancestral Pax gene in eyes has led to the conclusion that PaxB was a primordial gene in eye evolution, and that the eyes of all organisms likely share a common ancestor.

 

The lens structure of box jellyfish appears very similar to those of other organisms, but the crystallins are distinct in both function and appearance. Weak reactions were seen within the sera and there were very weak sequence similarities within the crystallins among vertebrate and invertebrate lenses. This is likely due to differences in lower molecular weight proteins and the subsequent lack of immunological reactions with antisera that other organisms' lenses exhibit.

 

All four of the visual systems of box jellyfish species investigated with detail (Carybdea marsupialis, Chiropsalmus quadrumanus, Tamoya haplonema and Tripedalia cystophora) have invaginated synapses, but only in the upper and lower lensed eyes. Different densities were found between the upper and lower lenses, and between species. Four types of chemical synapses have been discovered within the rhopalia which could help in understanding neural organization including: clear unidirectional, dense-core unidirectional, clear bidirectional, and clear and dense-core bidirectional. The synapses of the lensed eyes could be useful as markers to learn more about the neural circuit in box jellyfish retinal areas.

 

Evolution as a response to natural stimuli

The primary adaptive responses to environmental variation observed in box jellyfish eyes include pupillary constriction speeds in response to light environments, as well as photoreceptor tuning and lens adaptations to better respond to shifts between light environments and darkness. Interestingly, some box jellyfish species' eyes appear to have evolved more focused vision in response to their habitat.

 

Pupillary contraction appears to have evolved in response to variation in the light environment across ecological niches across three species of box jellyfish (Chironex fleckeri, Chiropsella bronzie, and Carukia barnesi). Behavioral studies suggest that faster pupil contraction rates allow for greater object avoidance, and in fact, species with more complex habitats exhibit faster rates. Ch. bronzie inhabit shallow beach fronts that have low visibility and very few obstacles, thus, faster pupil contraction in response to objects in their environment is not important. Ca. barnesi and Ch. fleckeri are found in more three-dimensionally complex environments like mangroves with an abundance of natural obstacles, where faster pupil contraction is more adaptive. Behavioral studies support the idea that faster pupillary contraction rates assist with obstacle avoidance as well as depth adjustments in response to differing light intensities.

 

Light/dark adaptation via pupillary light reflexes is an additional form of an evolutionary response to the light environment. This relates to the pupil's response to shifts between light intensity (generally from sunlight to darkness). In the process of light/dark adaptation, the upper and lower lens eyes of different box jellyfish species vary in specific function. The lower lens-eyes contain pigmented photoreceptors and long pigment cells with dark pigments that migrate on light/dark adaptation, while the upper-lens eyes play a concentrated role in light direction and phototaxis given that they face upward towards the water surface (towards the sun or moon). The upper lens of Ch. bronzie does not exhibit any considerable optical power while Tr. cystophora (a box jellyfish species that tends to live in mangroves) does. The ability to use light to visually guide behavior is not of as much importance to Ch. bronzie as it is to species in more obstacle-filled environments. Differences in visually guided behavior serve as evidence that species that share the same number and structure of eyes can exhibit differences in how they control behavior.

 

Largest and smallest

Jellyfish range from about one millimeter in bell height and diameter, to nearly 2 metres (6+1⁄2 ft) in bell height and diameter; the tentacles and mouth parts usually extend beyond this bell dimension.

 

The smallest jellyfish are the peculiar creeping jellyfish in the genera Staurocladia and Eleutheria, which have bell disks from 0.5 millimetres (1⁄32 in) to a few millimeters in diameter, with short tentacles that extend out beyond this, which these jellyfish use to move across the surface of seaweed or the bottoms of rocky pools; many of these tiny creeping jellyfish cannot be seen in the field without a hand lens or microscope. They can reproduce asexually by fission (splitting in half). Other very small jellyfish, which have bells about one millimeter, are the hydromedusae of many species that have just been released from their parent polyps; some of these live only a few minutes before shedding their gametes in the plankton and then dying, while others will grow in the plankton for weeks or months. The hydromedusae Cladonema radiatum and Cladonema californicum are also very small, living for months, yet never growing beyond a few mm in bell height and diameter.

 

The lion's mane jellyfish, Cyanea capillata, was long-cited as the largest jellyfish, and arguably the longest animal in the world, with fine, thread-like tentacles that may extend up to 36.5 m (119 ft 9 in) long (though most are nowhere near that large). They have a moderately painful, but rarely fatal, sting. The increasingly common giant Nomura's jellyfish, Nemopilema nomurai, found in some, but not all years in the waters of Japan, Korea and China in summer and autumn is another candidate for "largest jellyfish", in terms of diameter and weight, since the largest Nomura's jellyfish in late autumn can reach 2 m (6 ft 7 in) in bell (body) diameter and about 200 kg (440 lb) in weight, with average specimens frequently reaching 0.9 m (2 ft 11 in) in bell diameter and about 150 kg (330 lb) in weight. The large bell mass of the giant Nomura's jellyfish can dwarf a diver and is nearly always much greater than the Lion's Mane, whose bell diameter can reach 1 m (3 ft 3 in).

 

The rarely encountered deep-sea jellyfish Stygiomedusa gigantea is another candidate for "largest jellyfish", with its thick, massive bell up to 100 cm (3 ft 3 in) wide, and four thick, "strap-like" oral arms extending up to 6 m (19+1⁄2 ft) in length, very different from the typical fine, threadlike tentacles that rim the umbrella of more-typical-looking jellyfish, including the Lion's Mane.

 

Desmonema glaciale, which lives in the Antarctic region, can reach a very large size (several meters). Purple-striped jelly (Chrysaora colorata) can also be extremely long (up to 15 feet).

 

Life history and behavior

Life cycle

Jellyfish have a complex life cycle which includes both sexual and asexual phases, with the medusa being the sexual stage in most instances. Sperm fertilize eggs, which develop into larval planulae, become polyps, bud into ephyrae and then transform into adult medusae. In some species certain stages may be skipped.

 

Upon reaching adult size, jellyfish spawn regularly if there is a sufficient supply of food. In most species, spawning is controlled by light, with all individuals spawning at about the same time of day; in many instances this is at dawn or dusk. Jellyfish are usually either male or female (with occasional hermaphrodites). In most cases, adults release sperm and eggs into the surrounding water, where the unprotected eggs are fertilized and develop into larvae. In a few species, the sperm swim into the female's mouth, fertilizing the eggs within her body, where they remain during early development stages. In moon jellies, the eggs lodge in pits on the oral arms, which form a temporary brood chamber for the developing planula larvae.

 

The planula is a small larva covered with cilia. When sufficiently developed, it settles onto a firm surface and develops into a polyp. The polyp generally consists of a small stalk topped by a mouth that is ringed by upward-facing tentacles. The polyps resemble those of closely related anthozoans, such as sea anemones and corals. The jellyfish polyp may be sessile, living on the bottom, boat hulls or other substrates, or it may be free-floating or attached to tiny bits of free-living plankton or rarely, fish or other invertebrates. Polyps may be solitary or colonial. Most polyps are only millimetres in diameter and feed continuously. The polyp stage may last for years.

 

After an interval and stimulated by seasonal or hormonal changes, the polyp may begin reproducing asexually by budding and, in the Scyphozoa, is called a segmenting polyp, or a scyphistoma. Budding produces more scyphistomae and also ephyrae. Budding sites vary by species; from the tentacle bulbs, the manubrium (above the mouth), or the gonads of hydromedusae. In a process known as strobilation, the polyp's tentacles are reabsorbed and the body starts to narrow, forming transverse constrictions, in several places near the upper extremity of the polyp. These deepen as the constriction sites migrate down the body, and separate segments known as ephyra detach. These are free-swimming precursors of the adult medusa stage, which is the life stage that is typically identified as a jellyfish. The ephyrae, usually only a millimeter or two across initially, swim away from the polyp and grow. Limnomedusae polyps can asexually produce a creeping frustule larval form, which crawls away before developing into another polyp. A few species can produce new medusae by budding directly from the medusan stage. Some hydromedusae reproduce by fission.

 

Lifespan

Little is known of the life histories of many jellyfish as the places on the seabed where the benthic forms of those species live have not been found. However, an asexually reproducing strobila form can sometimes live for several years, producing new medusae (ephyra larvae) each year.

 

An unusual species, Turritopsis dohrnii, formerly classified as Turritopsis nutricula, might be effectively immortal because of its ability under certain circumstances to transform from medusa back to the polyp stage, thereby escaping the death that typically awaits medusae post-reproduction if they have not otherwise been eaten by some other organism. So far this reversal has been observed only in the laboratory.

 

Locomotion

Jellyfish locomotion is highly efficient. Muscles in the jellylike bell contract, setting up a start vortex and propelling the animal. When the contraction ends, the bell recoils elastically, creating a stop vortex with no extra energy input.

Using the moon jelly Aurelia aurita as an example, jellyfish have been shown to be the most energy-efficient swimmers of all animals. They move through the water by radially expanding and contracting their bell-shaped bodies to push water behind them. They pause between the contraction and expansion phases to create two vortex rings. Muscles are used for the contraction of the body, which creates the first vortex and pushes the animal forward, but the mesoglea is so elastic that the expansion is powered exclusively by relaxing the bell, which releases the energy stored from the contraction. Meanwhile, the second vortex ring starts to spin faster, sucking water into the bell and pushing against the centre of the body, giving a secondary and "free" boost forward. The mechanism, called passive energy recapture, only works in relatively small jellyfish moving at low speeds, allowing the animal to travel 30 percent farther on each swimming cycle. Jellyfish achieved a 48 percent lower cost of transport (food and oxygen intake versus energy spent in movement) than other animals in similar studies. One reason for this is that most of the gelatinous tissue of the bell is inactive, using no energy during swimming.

 

Ecology

Diet

Jellyfish are, like other cnidarians, generally carnivorous (or parasitic), feeding on planktonic organisms, crustaceans, small fish, fish eggs and larvae, and other jellyfish, ingesting food and voiding undigested waste through the mouth. They hunt passively using their tentacles as drift lines, or sink through the water with their tentacles spread widely; the tentacles, which contain nematocysts to stun or kill the prey, may then flex to help bring it to the mouth. Their swimming technique also helps them to capture prey; when their bell expands it sucks in water which brings more potential prey within reach of the tentacles.

 

A few species such as Aglaura hemistoma are omnivorous, feeding on microplankton which is a mixture of zooplankton and phytoplankton (microscopic plants) such as dinoflagellates. Others harbour mutualistic algae (Zooxanthellae) in their tissues; the spotted jellyfish (Mastigias papua) is typical of these, deriving part of its nutrition from the products of photosynthesis, and part from captured zooplankton. The upside-down jellyfish (Cassiopea andromeda) also has a symbiotic relationship with microalgae, but captures tiny animals to supplement their diet. This is done by releasing tiny balls of living cells composed of mesoglea. These use cilia to drive them through water and stinging cells which stun the prey. The blobs also seems to have digestive capabilities.

 

Predation

Other species of jellyfish are among the most common and important jellyfish predators. Sea anemones may eat jellyfish that drift into their range. Other predators include tunas, sharks, swordfish, sea turtles and penguins. Jellyfish washed up on the beach are consumed by foxes, other terrestrial mammals and birds. In general however, few animals prey on jellyfish; they can broadly be considered to be top predators in the food chain. Once jellyfish have become dominant in an ecosystem, for example through overfishing which removes predators of jellyfish larvae, there may be no obvious way for the previous balance to be restored: they eat fish eggs and juvenile fish, and compete with fish for food, preventing fish stocks from recovering.

 

Symbiosis

Some small fish are immune to the stings of the jellyfish and live among the tentacles, serving as bait in a fish trap; they are safe from potential predators and are able to share the fish caught by the jellyfish. The cannonball jellyfish has a symbiotic relationship with ten different species of fish, and with the longnose spider crab, which lives inside the bell, sharing the jellyfish's food and nibbling its tissues.

 

Main article: Jellyfish bloom

Jellyfish form large masses or blooms in certain environmental conditions of ocean currents, nutrients, sunshine, temperature, season, prey availability, reduced predation and oxygen concentration. Currents collect jellyfish together, especially in years with unusually high populations. Jellyfish can detect marine currents and swim against the current to congregate in blooms. Jellyfish are better able to survive in nutrient-rich, oxygen-poor water than competitors, and thus can feast on plankton without competition. Jellyfish may also benefit from saltier waters, as saltier waters contain more iodine, which is necessary for polyps to turn into jellyfish. Rising sea temperatures caused by climate change may also contribute to jellyfish blooms, because many species of jellyfish are able to survive in warmer waters. Increased nutrients from agricultural or urban runoff with nutrients including nitrogen and phosphorus compounds increase the growth of phytoplankton, causing eutrophication and algal blooms. When the phytoplankton die, they may create dead zones, so-called because they are hypoxic (low in oxygen). This in turn kills fish and other animals, but not jellyfish, allowing them to bloom. Jellyfish populations may be expanding globally as a result of land runoff and overfishing of their natural predators. Jellyfish are well placed to benefit from disturbance of marine ecosystems. They reproduce rapidly; they prey upon many species, while few species prey on them; and they feed via touch rather than visually, so they can feed effectively at night and in turbid waters. It may be difficult for fish stocks to re-establish themselves in marine ecosystems once they have become dominated by jellyfish, because jellyfish feed on plankton, which includes fish eggs and larvae.

 

As suspected at the turn of this century, jellyfish blooms are increasing in frequency. Between 2013 and 2020 the Mediterranean Science Commission monitored on a weekly basis the frequency of such outbreaks in coastal waters from Morocco to the Black Sea, revealing a relatively high frequency of these blooms nearly all year round, with peaks observed from March to July and often again in the autumn. The blooms are caused by different jellyfish species, depending on their localisation within the Basin: one observes a clear dominance of Pelagia noctiluca and Velella velella outbreaks in the western Mediterranean, of Rhizostoma pulmo and Rhopilema nomadica outbreaks in the eastern Mediterranean, and of Aurelia aurita and Mnemiopsis leidyi outbreaks in the Black Sea.

 

Some jellyfish populations that have shown clear increases in the past few decades are invasive species, newly arrived from other habitats: examples include the Black Sea, Caspian Sea, Baltic Sea, central and eastern Mediterranean, Hawaii, and tropical and subtropical parts of the West Atlantic (including the Caribbean, Gulf of Mexico and Brazil).

 

Jellyfish blooms can have significant impact on community structure. Some carnivorous jellyfish species prey on zooplankton while others graze on primary producers. Reductions in zooplankton and ichthyoplankton due to a jellyfish bloom can ripple through the trophic levels. High-density jellyfish populations can outcompete other predators and reduce fish recruitment. Increased grazing on primary producers by jellyfish can also interrupt energy transfer to higher trophic levels.

 

During blooms, jellyfish significantly alter the nutrient availability in their environment. Blooms require large amounts of available organic nutrients in the water column to grow, limiting availability for other organisms. Some jellyfish have a symbiotic relationship with single-celled dinoflagellates, allowing them to assimilate inorganic carbon, phosphorus, and nitrogen creating competition for phytoplankton. Their large biomass makes them an important source of dissolved and particulate organic matter for microbial communities through excretion, mucus production, and decomposition. The microbes break down the organic matter into inorganic ammonium and phosphate. However, the low carbon availability shifts the process from production to respiration creating low oxygen areas making the dissolved inorganic nitrogen and phosphorus largely unavailable for primary production.

 

These blooms have very real impacts on industries. Jellyfish can outcompete fish by utilizing open niches in over-fished fisheries. Catch of jellyfish can strain fishing gear and lead to expenses relating to damaged gear. Power plants have been shut down due to jellyfish blocking the flow of cooling water. Blooms have also been harmful for tourism, causing a rise in stings and sometimes the closure of beaches.

 

Jellyfish form a component of jelly-falls, events where gelatinous zooplankton fall to the seafloor, providing food for the benthic organisms there. In temperate and subpolar regions, jelly-falls usually follow immediately after a bloom.

 

Habitats

Most jellyfish are marine animals, although a few hydromedusae inhabit freshwater. The best known freshwater example is the cosmopolitan hydrozoan jellyfish, Craspedacusta sowerbii. It is less than an inch (2.5 cm) in diameter, colorless and does not sting. Some jellyfish populations have become restricted to coastal saltwater lakes, such as Jellyfish Lake in Palau. Jellyfish Lake is a marine lake where millions of golden jellyfish (Mastigias spp.) migrate horizontally across the lake daily.

 

Although most jellyfish live well off the ocean floor and form part of the plankton, a few species are closely associated with the bottom for much of their lives and can be considered benthic. The upside-down jellyfish in the genus Cassiopea typically lie on the bottom of shallow lagoons where they sometimes pulsate gently with their umbrella top facing down. Even some deep-sea species of hydromedusae and scyphomedusae are usually collected on or near the bottom. All of the stauromedusae are found attached to either seaweed or rocky or other firm material on the bottom.

 

Some species explicitly adapt to tidal flux. In Roscoe Bay, jellyfish ride the current at ebb tide until they hit a gravel bar, and then descend below the current. They remain in still waters until the tide rises, ascending and allowing it to sweep them back into the bay. They also actively avoid fresh water from mountain snowmelt, diving until they find enough salt.

  

Parasites

Jellyfish are hosts to a wide variety of parasitic organisms. They act as intermediate hosts of endoparasitic helminths, with the infection being transferred to the definitive host fish after predation. Some digenean trematodes, especially species in the family Lepocreadiidae, use jellyfish as their second intermediate hosts. Fish become infected by the trematodes when they feed on infected jellyfish.

 

Relation to humans

Jellyfish have long been eaten in some parts of the world. Fisheries have begun harvesting the American cannonball jellyfish, Stomolophus meleagris, along the southern Atlantic coast of the United States and in the Gulf of Mexico for export to Asia.

 

Jellyfish are also harvested for their collagen, which is being investigated for use in a variety of applications including the treatment of rheumatoid arthritis.

 

Aquaculture and fisheries of other species often suffer severe losses – and so losses of productivity – due to jellyfish.

 

Products

Main article: Jellyfish as food

In some countries, including China, Japan, and Korea, jellyfish are a delicacy. The jellyfish is dried to prevent spoiling. Only some 12 species of scyphozoan jellyfish belonging to the order Rhizostomeae are harvested for food, mostly in southeast Asia. Rhizostomes, especially Rhopilema esculentum in China (海蜇 hǎizhé, 'sea stingers') and Stomolophus meleagris (cannonball jellyfish) in the United States, are favored because of their larger and more rigid bodies and because their toxins are harmless to humans.

 

Traditional processing methods, carried out by a jellyfish master, involve a 20- to 40-day multi-phase procedure in which, after removing the gonads and mucous membranes, the umbrella and oral arms are treated with a mixture of table salt and alum, and compressed. Processing makes the jellyfish drier and more acidic, producing a crisp texture. Jellyfish prepared this way retain 7–10% of their original weight, and the processed product consists of approximately 94% water and 6% protein. Freshly processed jellyfish has a white, creamy color and turns yellow or brown during prolonged storage.

 

In China, processed jellyfish are desalted by soaking in water overnight and eaten cooked or raw. The dish is often served shredded with a dressing of oil, soy sauce, vinegar and sugar, or as a salad with vegetables. In Japan, cured jellyfish are rinsed, cut into strips and served with vinegar as an appetizer. Desalted, ready-to-eat products are also available.

 

Biotechnology

The hydromedusa Aequorea victoria was the source of green fluorescent protein, studied for its role in bioluminescence and later for use as a marker in genetic engineering.

Pliny the Elder reported in his Natural History that the slime of the jellyfish "Pulmo marinus" produced light when rubbed on a walking stick.

 

In 1961, Osamu Shimomura extracted green fluorescent protein (GFP) and another bioluminescent protein, called aequorin, from the large and abundant hydromedusa Aequorea victoria, while studying photoproteins that cause bioluminescence in this species. Three decades later, Douglas Prasher sequenced and cloned the gene for GFP. Martin Chalfie figured out how to use GFP as a fluorescent marker of genes inserted into other cells or organisms. Roger Tsien later chemically manipulated GFP to produce other fluorescent colors to use as markers. In 2008, Shimomura, Chalfie and Tsien won the Nobel Prize in Chemistry for their work with GFP. Man-made GFP became widely used as a fluorescent tag to show which cells or tissues express specific genes. The genetic engineering technique fuses the gene of interest to the GFP gene. The fused DNA is then put into a cell, to generate either a cell line or (via IVF techniques) an entire animal bearing the gene. In the cell or animal, the artificial gene turns on in the same tissues and the same time as the normal gene, making a fusion of the normal protein with GFP attached to the end, illuminating the animal or cell reveals what tissues express that protein—or at what stage of development. The fluorescence shows where the gene is expressed.

 

Aquarium display

Jellyfish are displayed in many public aquariums. Often the tank's background is blue and the animals are illuminated by side light, increasing the contrast between the animal and the background. In natural conditions, many jellies are so transparent that they are nearly invisible. Jellyfish are not adapted to closed spaces. They depend on currents to transport them from place to place. Professional exhibits as in the Monterey Bay Aquarium feature precise water flows, typically in circular tanks to avoid trapping specimens in corners. The outflow is spread out over a large surface area and the inflow enters as a sheet of water in front of the outflow, so the jellyfish do not get sucked into it. As of 2009, jellyfish were becoming popular in home aquariums, where they require similar equipment.

 

Stings

Jellyfish are armed with nematocysts, a type of specialized stinging cell. Contact with a jellyfish tentacle can trigger millions of nematocysts to pierce the skin and inject venom, but only some species' venom causes an adverse reaction in humans. In a study published in Communications Biology, researchers found a jellyfish species called Cassiopea xamachana which when triggered will release tiny balls of cells that swim around the jellyfish stinging everything in their path. Researchers described these as "self-propelling microscopic grenades" and named them cassiosomes.

 

The effects of stings range from mild discomfort to extreme pain and death. Most jellyfish stings are not deadly, but stings of some box jellyfish (Irukandji jellyfish), such as the sea wasp, can be deadly. Stings may cause anaphylaxis (a form of shock), which can be fatal. Jellyfish kill 20 to 40 people a year in the Philippines alone. In 2006 the Spanish Red Cross treated 19,000 stung swimmers along the Costa Brava.

 

Vinegar (3–10% aqueous acetic acid) may help with box jellyfish stings but not the stings of the Portuguese man o' war. Clearing the area of jelly and tentacles reduces nematocyst firing. Scraping the affected skin, such as with the edge of a credit card, may remove remaining nematocysts. Once the skin has been cleaned of nematocysts, hydrocortisone cream applied locally reduces pain and inflammation. Antihistamines may help to control itching. Immunobased antivenins are used for serious box jellyfish stings.

 

In Elba Island and Corsica dittrichia viscosa is now used by residents and tourists to heal stings from jellyfish, bees and wasps pressing fresh leaves on the skin with quick results.

 

Mechanical issues

Jellyfish in large quantities can fill and split fishing nets and crush captured fish. They can clog cooling equipment, having disabled power stations in several countries; jellyfish caused a cascading blackout in the Philippines in 1999, as well as damaging the Diablo Canyon Power Plant in California in 2008. They can also stop desalination plants and ships' engines.

Foxes - British Wildlife Centre, Kent, England - Sunday August 17th 2008.

Click here to see the Larger image

 

Fox is a name applied to any of roughly 27 species of small to medium-sized canids, characterized by possessing a long, narrow snout, and a bushy tail, or "brush". By far the most common and widespread species of fox is the red fox (Vulpes vulpes), although various species are found on almost every continent. The presence of fox-like carnivores all over the globe has led to their appearance in the popular culture and folklore of many nations, tribes, and other cultural groups.

 

Etymology ~ The Modern English "fox" is derived from Old English fox. The Old English word itself comes from the Proto-Germanic word *fukh – compare German Fuchs, Gothic fauho, Old Norse foa and Dutch vos. It corresponds to the Proto-Indo-European word *puke meaning "tail" (compare Sanskrit puccha, also "tail"). The bushy tail is also the source of the word for fox in Welsh: llwynog, from llwyn, "bush", Lithuanian: uodegis, from uodega, "tail", and Portuguese: raposa, from rabo, "tail".

 

General characteristics ~ Most foxes live 2 to 3 years, but they can survive for up to 10 years or even longer in captivity. Foxes are generally smaller than other members of the family Canidae such as wolves, jackals, and domestic dogs. Dogs (male foxes) weigh on average, 5.9kg and vixens (female foxes) weigh less, at 5.2kg (13 lbs and 11.5 lbs, respectively). Fox-like features typically include an acute muzzle (a "fox face") and bushy tail. Other physical characteristics vary according to their habitat. For example, the fennec fox (and other species of foxes adapted to life in the desert, such as the kit fox) has large ears and short fur, whereas the Arctic fox has small ears and thick, insulating fur.

 

Another example is the red fox which has a typical auburn pelt, the tail normally ending with white marking.

Unlike many canids, foxes are usually not pack animals. Typically, they are solitary, opportunistic feeders that hunt live prey (especially rodents). Using a pouncing technique practiced from an early age, they are usually able to kill their prey quickly. Foxes also gather a wide variety of other foods ranging from grasshoppers to fruit and berries.

 

Foxes are normally extremely wary of humans and are not kept as pets (with the exception of the fennec); however, the silver fox was successfully domesticated in Russia after a 45 year selective breeding program. This selective breeding also resulted in physical and behavioural traits appearing that are frequently seen in domestic cats, dogs, and other animals: pigmentation changes, floppy ears, and curly tails.

 

Diet ~ The diet of foxes comprises rodents, insects, worms, fruit, fish, birds, eggs, and all other kinds of small animals. The fox generally consumes around 1 kg of food every day. Foxes that live in neighborhoods mainly depend on household waste and even rodents and birds that keep moving around these areas. Foxes are known to cache their food, burying the excess for later consumption.

 

They mostly thrive in the higher latitudes, suburban and even urban environments both in Europe and in North America. They are found also in Eurasia, North Africa, India (Ladakh, Himalayas, Jammu and Kashmir, Rajasthan and Gujarat), China, Japan and in Australia.

 

Conservation ~ Foxes are readily found in cities and cultivated areas and (depending upon species) seem to adapt reasonably well to human presence.

 

Red foxes have been introduced into Australia and some other countries for hunting. Australia lacks similar carnivores, and the introduced foxes prey on native wildlife, some to the point of extinction. A similar introduction occurred in the seventeenth and eighteenth centuries in temperate North America, where European reds (Vulpes vulpes) were brought to the colonies for fox hunting, where they decimated the American red fox population through more aggressive hunting and breeding. Interbreeding with American reds, traits of the European red eventually pervaded the gene pool, leaving European and American foxes now virtually identical.

 

Other fox species do not adapt as well as the red fox, and are endangered in their native environments. Key among these are the crab-eating fox (Cerdocyon thous) and the African bat-eared fox. Other foxes such as fennec foxes, are not endangered, but will be if humans encroach further into their habitat.

 

Foxes have been successfully employed to control pests on fruit farms, where they leave the fruit intact.

 

Historians believe foxes were imported into non-native environments long before the colonial era. The first example of the introduction of the fox into a new habitat by humans seems to be Neolithic Cyprus. Stone carvings representing foxes have been found in the early settlement of Göbekli Tepe in eastern Turkey.

Bronica SQ-A, Zenzanon 50mm f/3.5 PS, Kodak Ektachrome 100GX, CS8800F.

A game of great charm in the adoption of mathematical measurements to the timing of human movements, the exactitudes and adjustments of physical ability to hazardous chance. The speed of the legs, the dexterity of the body, the grace of the swing, the elusiveness of the slide - these are the features that make Americans everywhere forget the last syllable of a man's last name or the pigmentation of his skin.

 

~ Branch Rickey, May 1960

SEE UPGRADE 2023

 

www.flickr.com/photos/193116254@N07/52986354418/in/datepo...

  

Nous sommes vraiment à la fin du tirage sur verre ambrotype. 1870, dans le nord de la France.

( Sommes ? )

Quoiqu'aujourd’hui encore des stages en France proposent ce procédé de prise de vues et développement.

Il est vrai que l'on ne peut que rester admiratif devant la qualité et le rendu de ce tirage, plus une stéréoscopie exemplaire nous apprend au moins que ce photographe anonyme était un vrai professionnel !

Ce procédé pouvait aussi se faire sur une plaque de fer peinte en noir.( ferrotypie) Sur cuir, pierre…

Cette ambrotype n'a pas été verni au pinceau comme les autres de cette série, ce qui visuellement avant traitement informatique donne une vue qui avec les poussières et les rayures recto verso ressemble plus à une vue prise à Londres la nuit sans aucun éclairage, par temps de pluie et brouillard...

Pour voir l'image en positif, il suffit de mettre la plaque devant un fond sombre, la plupart des plaques étaient d'ailleurs peintes en noir, et c'est la sur-pigmentation de celui-ci qui est souvent la cause de détérioration des ambrotypes.

 

We are really at the end of the ambrotype glass print. 1870, in the north of France.

( Somme ? ) Although still today internships in France offer this process of shooting and development.

 

It is true that we can only remain admiring the quality and the rendering of this print, plus an exemplary stereoscopy tells us at least that this anonymous photographer was a true professional!

This process could also be done on an iron plate painted in black.( ferrotypia) On leather, stone....

This ambrotype has not been varnished with brush like the others in this series, which visually before computer processing gives a view that with the dust and scratches two-sided looks more to a view taken in London at night without any lighting, in rain and fog...

To see the image in positive, just put the plate in front of a dark background, most of the plates were moreover painted in black, and it is the over-pigmentation of it that is often the cause of deterioration of the ambrotypes.

 

L’invention de l’ambrotype s’est faite par chance et par tâtonnement. Son histoire débute dans les années 1850, en même temps que l’invention des négatifs sur plaque de verre (dit : procédé au collodion humide). À cette époque, les pionniers de la photographie s’ingénient à découvrir de nouveaux procédés. On constate alors rapidement que certains des négatifs sous exposés peuvent être vus en positif. Il faut pour cela les placer sur un fond sombre. La technique est petit-à-petit affinée. Les négatifs sont par exemple blanchis chimiquement lors du développement. Ils sont ainsi davantage visibles lorsqu’on les dispose sur leur fond sombre.

Le daguerréotypiste Marcus Aurelius Root donna le nom d’ambrotype à un procédé photographique inventé par James Ambrose Cutting. Il fut d’abord développé entre 1851 et 1854 par Frederick Scott ARCHER puis Adolphe MARTIN sous le nom d’amphitype. Il est finalement breveté en 1854 par CUTTING et son associé Isaac REHN. Le terme d’ambrotype provient du grec ambrotos, signifiant « immortel ».

 

Le daguerréotype et l’ambrotype se sont concurrencés. L’ambrotype permettait une rapidité d'obtention des images. Il était aussi moins coûteux. L’ambrotype est une image négative unique, associée à un fond noir pour apparaître en positif. Un côté d’une plaque de verre est recouvert d’une fine couche de collodion iodé. La plaque est ensuite plongée dans une solution acide de nitrate d’argent pour homogénéifier sa sensibilité. Elle est installée encore humide dans l’appareil et exposée à la lumière pendant quelques secondes, selon la luminosité. La plaque est ensuite développée et fixée, produisant ainsi une image en négatif. Après l’ajout d’un vernis, un fond noir est glissé derrière ce négatif, révélant ainsi l’image en positif.

 

Très utilisé pour les portraits et les paysages, l'ambrotype se présente généralement encadré comme le daguerréotype. Ils sont d’ailleurs parfois confondus. Ambrotypes et daguerréotypes étaient habituellement présentés dans un écrin, semblable à de véritables petits bijoux. L’ambrotype fut supplanté dans les années 1870 par l’apparition de nouvelles techniques.

S:

www.gadcollection.com/fr/blog/p-ambrotype

Laguna Colorada (Red Lagoon) is a shallow salt lake in the southwest of the altiplano of Bolivia, within Eduardo Avaroa Andean Fauna National Reserve and close to the border with Chile.

 

The reddish color of its waters, which is caused by red sediments and pigmentation of some algae.

 

The birds which can be seen in some of the pictures are flamingos.

Kaloplocamus acutus is characterized by a mantle which ranges in colour from pale yellow-orange to a deep orange red. The branched papillae have translucent white stalks with some scattered opaque white pigmentation and the pointed branches are bright orange or orange-red. There are scattered white specks on the body. The six branched papillae on the head shield or veil are pretty long when extended, longer than in Plocamopherus. There are four branched papillae down each side of the body and a slight ridge between them denotes the mantle edge. Found in Anilao, South Luzon, Philippines

Built in 1919-1920, this Chicago School and Sullivanesque-style building was designed by Louis Sullivan for the Farmers and Merchants Union Bank in Columbus, Wisconsin as one of his late-career “jewel box” bank buildings that are largely located in smaller communities throughout the midwest. The building was the last “jewel box bank” designed by Sullivan, and the second-to-last commission of his career, and was intended to communicate the bank as a modern and progressive institution, rather than employing the stodgier and more traditional Classical design found on most other banks of the era. The bank was commissioned by the president of the bank, J. Russell Wheeler, whose wife, Anna May Wheeler, pushed him to commission Sullivan to design a new home for the bank. In addition to Louis Sullivan, the building’s stained glass windows, were designed by architectural decorator Louis J. Millet, and the terra cotta by clay modeler Kristian Schneider, whom developed moulds for the building’s terra cotta, metal, and plaster details. The two artisans worked alongside Sullivan on several other bank projects. The building was heavily documented in Sullivan’s 1924 “A System of Architectural Ornament”, published shortly before he died.

 

The building is clad in red tapestry brick, which features blue and green mixed with the red clay mixture in some bricks, creating variation in color and texture across the facade. The brick creates a backdrop to some of the best terra cotta on any of Sullivan’s projects. The terra cotta features many of the floral and geometric motifs found on Sullivan’s other works, and is arranged similarly to other Sullivan banks that utilized brick cladding. The building features two principal facades, with a narrower facade along James Street, and a broader facade facing Dickason Boulevard. The James Street facade features two openings close to ground level, with the eastern bay housing a large plate glass window, and the western bay housing a doorway flanked by skylights, both of which are recessed under a large terra cotta architrave and flanked by square pilasters with decorative Sullivanesque ornament panels at the capitals. The architrave above the doorway and window is divided into three segments by vertical terra cotta elements that feature floral motifs and, like many Sullivan buildings, appear like plants with roots, branches, and crowns. The outer panels of the architrave feature circular cartouches with hexagonal trim, leaves, and geometric elements, with circular central medallions featuring the years 1861, when the bank was founded, and 1919, when the bank was completed. The central panel is clad in marble with the words “Farmers & Merchants Union Bank” and “Louis Sullivan, Architect” engraved into the stone with yellow pigmentation, contrasting against the white and green marble background. Atop the two vertical elements on either side of the central panel are griffin sculptures holding shields, a common element on many of Sullivan’s “Jewel Box Banks,” while the base of the outer vertical elements features the initials of the bank at the base. Above the architrave is an arched bay that houses a stained glass window, trimmed with decorative terra cotta at the inner and outer rings of the arch, with the bay becoming more recessed after each concentric arch, much like the entrances to medieval Romanesque churches. Besides a band of belt coursing that runs on either side of the architrave and wraps the corner to a tapered buttress on the Dickason Boulevard facade, the only other adornment is an eagle sculpture on a vertical trim element at the center of the parapet, which terminates many brick courses above the arched opening below, and another band of terra cotta trim along the top of the parapet, which forms a cap on the parapet around the perimeter of the building’s low-slope roof. On the Dickason Boulevard facade, the building features five recessed clerestory arched bays housing stained glass windows, flanked by tapered buttresses. Surrounding the arched tops of the windows are decorative trim panels with floral motifs, which begin just below the base of the arches, and extend up above the top of the arches, terminating in a band of belt coursing. Atop the buttresses at either end are trim elements featuring large spheres atop rectilinear legs with floral motifs below, undulating in and out with the brick below. Additionally, a band of belt coursing, which wraps the corner of Dickson Boulevard and James Street, runs beneath the windows, only interrupted by the buttresses. Toward the back, on the building’s original rear wing, there are three windows at eye level in the original building, with bands of belt coursing below and at the top of the parapet. The rear window is a recessed bay window flanked by two pilasters with sullivanesque terra cotta panels, while the smaller windows are flanked by sullivanesque relief panels. The rear wing features a roof at multiple heights, and was extended in 1961 with a matching addition by Law, Potter and Nystrom, since removed. The rear of the taller portion of the building features a simple recessed bay with an arched window, and a similar eagle sculpture and vertical trim piece as on the front facade.

 

Inside, the front wing of the building features a tall banking hall with brick cladding on the walls up to the level of the windows, where it terminates at a wooden sill. The space is split down the middle by a row of brick piers and low walls framing the teller cages, which terminate at the sill line of the windows, dividing the space while still allowing it to read as a single continuous lofty space. The brick forms piers at the teller’s cages, pilasters separating desks on the exterior wall, and low brick walls with marble caps. The upper portion of the walls and the coffered ceiling in this space is finished with white plaster, which gives the space a very vertical and airy feeling, as do the cream-colored terrazzo floors, which feature black edges at the base of the walls, tying the space together. The space features a terra cotta water fountain, or bubbler, also designed by sullivan, which features intricate ornament by Schneider. The space also features two mezzanine balconies with metal railings that run below the arched windows at the front and rear of the space, allowing managers to observe the activities in the lobby and teller area below from the rear balcony, while the front balcony exists solely to balance the space and keep it symmetrical. An office for private conferences with customers was originally located near the front of the space, along with a manager’s office, allowing convenience for customers seeking a meeting with the bank management. The teller’s side of the space also housed the bank’s two vaults and several other private offices. The bank originally featured a large meeting room in the one-story rear wing, behind the vaults, with a women’s waiting room sitting along the Dickason Boulevard side of the rear wing, featuring a bay window and a restroom. The building’s interior has changed in function somewhat due to the growth of the bank, changes in bank operations, and expansion of the building with new additions to house offices and a drive-through in the rear.

 

The building was listed on the National Register of Historic Places in 1972, was designated a National Historic Landmark in 1976, and is a contributing structure in the Columbus Downtown Historic District, listed on the National Register of Historic Places in 1992. The building saw an addition in 2006, clad in buff brick, which replicated a historic building that formerly stood to the east, and wraps the building to the rear, with a two-story section behind a one-story annex that connects the one-story rear wing of the bank to the new building. This wing replaced older additions made in 1961, which matched the one-story rear wing of the historic building, and 1980, which was modern in appearance and slightly recessed along James Street to give precedence to the historic building. The building still functions as the main office branch of the Farmers and Merchants Union Bank, which has grown substantially. The building has been long considered to be among the best of Sullivan’s “Jewel Box Banks,” and has been kept in excellent condition by the bank’s careful and caring generational stewardship.

The Hawthorn Shield Bug up close and personal, I think it was mesmerised by my camera as it stayed perfectly still for a 19 frame handheld focus stack, I was very impressed. I used the flash fill in technique for this one with an ISO of 500, an aperture of F/6.3 and a shutter speed of 1/60. (Camera in aperture priority and flash in ETTL with -2/3 bias)

 

Here is what Wiki says

 

The Hawthorn Shield Bug, Acanthosoma haemorrhoidale, is a common European shield bug. Its chief food is haws, the fruit of the hawthorn tree, but adults can overwinter on a diet of leaves, and individuals can be found on many potential food plants, including pedunculate oak, sessile oak and whitebeam. They may grow up to 17 mm long, and are camouflaged in shades of green and brown. Like many so-called "stink bugs", they may release unpleasant odours when disturbed.

 

The dark green scutellum band is trapezoidal in shape, and extends from the dorsal pronotum to the forewing hemelytral membrane which accounts for the final third of the main body length. The front wings lie flat without flight, with the scleratized pronotum and corium regions, coloured in a dark red pigmentation. The compound eyes are also pigmented red. There is a speckled pigmentation extending from the upper dorsal throax to the abodomen region. The tarsi are 2-segmented and antennae 5-segmented. As a heteropteran phytophagyte of mainly green leafed trees and red berries, the species feeds through the anterior section of the rostrum, which suctions liquidised plant tissue, semi-digested by the delivery of saliva enzymes from the posterior sector of the rostrum. The rostrum beak acts as a modified proboscis formed by the interlocking of mandibular and maxillary stylet into a double-tubed elongation covered by the labium. The structure is needle shaped and penetrates beyond the cuticle and epidermis layer to access the vascular tissue of the mesophyll layer of the leaf. The rostrum pricks into berries to access fruit sugars. As a stress response, an orange secretion is secreted from gland openings in the thorax.

 

Hope everyone has a grand Bank Holiday weekend :o)

 

VIEW ON BLACK, BEST VIEWED ORIGINAL OR LARGE SIZE

Appley IOW RSPB Love Nature ! This is a leucistic blackbird :) Leucism is a genetic condition that can cause white patches due to a lack of pigmentation. Blackbirds are among the most common species to be affected

Laguna Colorada means Red Lagoon and it is found in the Eduardo Avaroa Andean Fauna National Reserve. The reddish color of its waters is caused by red sediments and pigmentation of son algae.

 

La Laguna Colorada está ubicada en la reserva natural Eduardo Avaroa. El color rojo del agua se debe a sedimentos rojos y pigmentos de algas.

le Mascaret, Rixensart

Polar Bear

 

"Region: America

Class: Mammalia

Order: Carnivora

Family: Ursidae

Genus: Ursus

Scientific Name: Ursus maritimus

 

Description : Polar bears are considered the largest land carnivores in the world, matched only by very large individual Kodiak brown bears. Both sexes differ in size throughout their range; males being much larger than females and continuing to grow for a longer period of time. These bears have long, massive skulls, necks and bodies with long legs and large paws. Ears and tail are short. The nose is more prominent or “Roman” with a black rhinarium (nose pad). The tongue is black and the eyes brown. The surface of the skin is also black. Fur colour varies slightly with the season: new coats grown just prior to the winter season are very white appearing as slightly creamy white against the stark white of their icy winter environment. This pelage is thick, coarse and long with dense underfur. Guard hairs, found throughout the pelage, are shiny, almost glossy, oily and waterproof and have hollow shafts. Polar bears moult annually between the end of May and August. The coat becomes thinner and has a yellow wash or is almost a golden colour. Male polar bears weigh between 400 – 600 kg, and have been recorded up to a maximum of 800 kg. Females are smaller than the males weighing up to 300 kg, and when pregnant up to 460 kg. Polar bears are 2.5 – 3.5 m long.

 

Distribution : The polar bear is circumpolar in distribution, inhabiting all Arctic seas and coastlines. It is found on the pack-ice off the Alaskan coast north of Bering Strait, off the coasts of Greenland and along the Eurasian Arctic coast from Spitsbergen to Wrangell Island. Rare stragglers reach Iceland. Individual bears have been seen on the frozen Arctic Ocean as far as latitude 88 degrees North, only 2 degrees from the North Pole. In Canada, they are found along the Arctic coasts from Alaska to Labrador and from the tip of James Bay to northern Ellesmere Island. Polar bears do roam as far as 150 kilometres inland into the coniferous forests, where they live very differently from the polar bears which belong to the high Arctic.

 

Habitat : They prefer areas of annual ice, which they use as a hunting platform and protective cover. This includes snow-drifted pressure ridges, refrozen cracks and areas of open water surrounded by ice. In areas where the pack ice melts by mid to late summer they come inland and live in coniferous forest areas. Here they remain until the ice re-freezes. Areas of solidly frozen sea ice and the open seas are avoided. Generally they are most common along coastal areas. Some do enter the permanent pack ice.

 

Food : The ringed seal is by far the most common prey. They also eat bearded seal, harp seal and hooded seal. Young walrus are sometimes taken. During the summer months they feed upon the shoreline carrion, fish, mussels, crabs, starfish, lemmings and the eggs and nestling young of waterfowl and cliff-dwelling birds. They will also graze on kelp, grasses and eat mushrooms and crowberries.

 

Reproduction and Development : Normally they are solitary animals outside the breeding season, the exception being a mother with cubs. Polar bears mate in mid-summer. Females first start to breed at 3 to 5 years of age. In April and May adult females are in oestrus and ready to accept a mate. They are polyandrous, meaning one female will mate with more than one male in one breeding season. Males fight among themselves for the female’s attention and a couple will pair off for a period of a few days to two weeks. With females, delayed implantation occurs, the fertilized egg does not implant in the uterus until mid-September to mid-October. Embryonic development begins at this time. Gestation periods, therefore, vary a great deal when including this period of delay. Females choose suitable locations to build their maternity dens in mid-October and retreat to them for the winter season. They give birth to one to four cubs somewhere between late November and early January. Twins are most common. The newborns are small; 25 to 30 cm long and weigh less than 1 kg. They are covered with very fine hair, appearing almost naked and their eyes are closed. Their eyes open at 6 weeks. Growth is very quick; at two months their fur has thickened; they weigh about 5 kg and move about the den. By mid-March to early April, when the den is opened, the cubs weigh about 10 kg and are surprisingly strong. The cubs suckle for nine months, occasionally one year. They are very dependent on their mother and stay with her for two years. At that time they weigh 90 - 180 kg and are half grown.

 

Adaptations : Polar bears are wonderfully adapted to their Arctic surroundings.

 

Locomotion. On land a shuffling walk may be increased to a rolling gallop of 40 km/h and can outrun caribou over a short distance. Bears are often seen standing high on their hind legs, necks stretched to scan the landscape. On thin ice, legs are spread to distribute body mass. Thickly padded and furred soles allow the bear to move quietly as well as providing good traction. Small bumps and cavities on the soles act like suction cups keeping bears from slipping on the ice. The claws are used to dig into icy slopes and to grip prey. They are strong swimmers, paddling with their forefeet only and trailing their hind feet which act as a rudder. They can stay submerged for over one minute, keeping their eyes open. They swim at a speed of approximately 6.4 km/h, often covering long distances.

 

Insulation. Polar bears have a thick layer of sub-cutaneous fat and very dense underfur with several layers of glossy guard hair on the outside. Their pelt is much thicker in winter and provides excellent insulation. The fat layer also adds to buoyancy in the water. Water is shed easily from the oily waterproof fur. Small, furry ears have a heavy network of blood vessels, keeping them warm and conserving heat. The tail is short and rounded also conserving heat. Fur is very dense around the soles of the feet.

 

Pelage. The creamy white appearance of the coat allows the bear to be inconspicuous when hunting seal. Each hair is similar to an optical fibre; colourless and hollow. Being translucent, it reflects the heat from the sun down to the base of the hair, where it is absorbed by the black skin. Whiteness comes from reflection of light rather than pigmentation.

 

Hunting. Bears use their keen sense of smell to detect seal breathing holes. These can be up to a kilometer away and covered by a layer of snow and ice. They will stand or lie by the seal’s blowhole in the ice for hours; they may swim towards seals resting on the ice flows with only their nose showing above the water. They will dive quietly, then swim up to the ice edge and jump out on the seal, and will also crawl towards a sunbathing seal using every piece of raised ice to conceal the approach.

 

Denning. Both sexes occupy dens for shelter. Topographic factors influence the den sites. In Canadian core areas, dens frequently occur on south-facing slopes where northerly prevailing winds create the best drifts, where the wind-chill is least and insulation from received solar radiation is greatest. One of the three largest denning areas worldwide is in Canada. There are three main types of winter refuges: maternity dens, temporary dens and winter shelters. During the winter any bear may dig a temporary den and use it for a few days during a storm, or take shelter in a natural cavity. Winter shelters are used for longer periods of time as resting places. This type of shelter is usually roomier with additional features such as alcoves, porches and ventilation holes. Bears do not hibernate in the strict sense of the word; they have the ability to slow down their metabolism to conserve energy at any time of year. The state of self-induced lethargy while in the shelter allows them to preserve their vital fat reserves. During this time, the body temperature of the polar bear decreases by a few degrees from normal and the respiration rates are markedly reduced. Maternity dens are built and occupied by pregnant females and can vary in size. The denning chamber is at the upper end of an entrance tunnel 1 to 2 m long. It averages 1.5 m in diameter with a height of 90 to 100cm in the middle. Drifting snow seals the entrance. The chamber is higher than the tunnel, trapping bear body heat inside. Dens not only provide a safe place to give birth to her cubs, but are also a place of protection for the cubs during their first few months. During this time she does not leave the den, remaining with her cubs and living on her reserves of fat. In the spring, with her fat severely depleted she must leave to find food to sustain herself and her cubs. After they leave the maternity den she will build temporary refuges in which to nurse, rest and shelter her young cubs and warms them as they all sleep together. She heads with her cubs towards the nearest supply of food, this is usually pack ice.

 

Sight and sounds. Polar bears have good eyesight. Their eyes have inner eyelids that keep the glare of the sun on snow and ice from blinding them. When defending a food source from other bears they use a deep growl. They hiss and snort to show aggression. Angry bears use loud roars and growls. Mothers scold cubs with a low growl.

 

Threats to Survival : Polar bears are one of the animals most threatened by global warming. They depend entirely on sea ice as a platform from which to hunt seals. Reduction of the total ice cover in the Arctic is a serious concern globally. When the ice does not form or forms too late in the season many polar bears starve. In Hudson Bay, scientists have found the main cause of death for cubs to be either lack of food or lack of fat on nursing mothers. Exploitation of minerals and fossil fuels in the Arctic pose a continuous threat. Of the oil and natural gas deposits globally, 20% are located in the Arctic. As the ice cap recedes these become more accessible. Countries are competing which each other in claiming ownership of Arctic and its resources. This can only result in further and more drastic impact on polar bear habitat.

 

Status : IUCN: Vulnerable; CITES: Appendix II; COSEWIC: Special Concern

 

Zoo Diet : Toronto Zoo carnivore diet, dog chow, jumbo smelt and herring, carrots, bean sprouts, Vitamin E and Thiamine supplements."

- Courtesy of the Metro Toronto Zoo

 

© All Rights Reserved - Miles Away Photography

Please! No usage allowed without the consent of Mandi A. Miles

Based out of Flesherton, ON, please write for prices and information!

Miles Away Photography

 

aka Michael Jacksons disease

 

in

Kolkata

   

Photography’s new conscience

linktr.ee/GlennLosack

linktr.ee/GlennLosack

  

glosack.wixsite.com/tbws

 

Polar Bear

 

"Region: America

Class: Mammalia

Order: Carnivora

Family: Ursidae

Genus: Ursus

Scientific Name: Ursus maritimus

 

Description : Polar bears are considered the largest land carnivores in the world, matched only by very large individual Kodiak brown bears. Both sexes differ in size throughout their range; males being much larger than females and continuing to grow for a longer period of time. These bears have long, massive skulls, necks and bodies with long legs and large paws. Ears and tail are short. The nose is more prominent or “Roman” with a black rhinarium (nose pad). The tongue is black and the eyes brown. The surface of the skin is also black. Fur colour varies slightly with the season: new coats grown just prior to the winter season are very white appearing as slightly creamy white against the stark white of their icy winter environment. This pelage is thick, coarse and long with dense underfur. Guard hairs, found throughout the pelage, are shiny, almost glossy, oily and waterproof and have hollow shafts. Polar bears moult annually between the end of May and August. The coat becomes thinner and has a yellow wash or is almost a golden colour. Male polar bears weigh between 400 – 600 kg, and have been recorded up to a maximum of 800 kg. Females are smaller than the males weighing up to 300 kg, and when pregnant up to 460 kg. Polar bears are 2.5 – 3.5 m long.

 

Distribution : The polar bear is circumpolar in distribution, inhabiting all Arctic seas and coastlines. It is found on the pack-ice off the Alaskan coast north of Bering Strait, off the coasts of Greenland and along the Eurasian Arctic coast from Spitsbergen to Wrangell Island. Rare stragglers reach Iceland. Individual bears have been seen on the frozen Arctic Ocean as far as latitude 88 degrees North, only 2 degrees from the North Pole. In Canada, they are found along the Arctic coasts from Alaska to Labrador and from the tip of James Bay to northern Ellesmere Island. Polar bears do roam as far as 150 kilometres inland into the coniferous forests, where they live very differently from the polar bears which belong to the high Arctic.

 

Habitat : They prefer areas of annual ice, which they use as a hunting platform and protective cover. This includes snow-drifted pressure ridges, refrozen cracks and areas of open water surrounded by ice. In areas where the pack ice melts by mid to late summer they come inland and live in coniferous forest areas. Here they remain until the ice re-freezes. Areas of solidly frozen sea ice and the open seas are avoided. Generally they are most common along coastal areas. Some do enter the permanent pack ice.

 

Food : The ringed seal is by far the most common prey. They also eat bearded seal, harp seal and hooded seal. Young walrus are sometimes taken. During the summer months they feed upon the shoreline carrion, fish, mussels, crabs, starfish, lemmings and the eggs and nestling young of waterfowl and cliff-dwelling birds. They will also graze on kelp, grasses and eat mushrooms and crowberries.

 

Reproduction and Development : Normally they are solitary animals outside the breeding season, the exception being a mother with cubs. Polar bears mate in mid-summer. Females first start to breed at 3 to 5 years of age. In April and May adult females are in oestrus and ready to accept a mate. They are polyandrous, meaning one female will mate with more than one male in one breeding season. Males fight among themselves for the female’s attention and a couple will pair off for a period of a few days to two weeks. With females, delayed implantation occurs, the fertilized egg does not implant in the uterus until mid-September to mid-October. Embryonic development begins at this time. Gestation periods, therefore, vary a great deal when including this period of delay. Females choose suitable locations to build their maternity dens in mid-October and retreat to them for the winter season. They give birth to one to four cubs somewhere between late November and early January. Twins are most common. The newborns are small; 25 to 30 cm long and weigh less than 1 kg. They are covered with very fine hair, appearing almost naked and their eyes are closed. Their eyes open at 6 weeks. Growth is very quick; at two months their fur has thickened; they weigh about 5 kg and move about the den. By mid-March to early April, when the den is opened, the cubs weigh about 10 kg and are surprisingly strong. The cubs suckle for nine months, occasionally one year. They are very dependent on their mother and stay with her for two years. At that time they weigh 90 - 180 kg and are half grown.

 

Adaptations : Polar bears are wonderfully adapted to their Arctic surroundings.

 

Locomotion. On land a shuffling walk may be increased to a rolling gallop of 40 km/h and can outrun caribou over a short distance. Bears are often seen standing high on their hind legs, necks stretched to scan the landscape. On thin ice, legs are spread to distribute body mass. Thickly padded and furred soles allow the bear to move quietly as well as providing good traction. Small bumps and cavities on the soles act like suction cups keeping bears from slipping on the ice. The claws are used to dig into icy slopes and to grip prey. They are strong swimmers, paddling with their forefeet only and trailing their hind feet which act as a rudder. They can stay submerged for over one minute, keeping their eyes open. They swim at a speed of approximately 6.4 km/h, often covering long distances.

 

Insulation. Polar bears have a thick layer of sub-cutaneous fat and very dense underfur with several layers of glossy guard hair on the outside. Their pelt is much thicker in winter and provides excellent insulation. The fat layer also adds to buoyancy in the water. Water is shed easily from the oily waterproof fur. Small, furry ears have a heavy network of blood vessels, keeping them warm and conserving heat. The tail is short and rounded also conserving heat. Fur is very dense around the soles of the feet.

 

Pelage. The creamy white appearance of the coat allows the bear to be inconspicuous when hunting seal. Each hair is similar to an optical fibre; colourless and hollow. Being translucent, it reflects the heat from the sun down to the base of the hair, where it is absorbed by the black skin. Whiteness comes from reflection of light rather than pigmentation.

 

Hunting. Bears use their keen sense of smell to detect seal breathing holes. These can be up to a kilometer away and covered by a layer of snow and ice. They will stand or lie by the seal’s blowhole in the ice for hours; they may swim towards seals resting on the ice flows with only their nose showing above the water. They will dive quietly, then swim up to the ice edge and jump out on the seal, and will also crawl towards a sunbathing seal using every piece of raised ice to conceal the approach.

 

Denning. Both sexes occupy dens for shelter. Topographic factors influence the den sites. In Canadian core areas, dens frequently occur on south-facing slopes where northerly prevailing winds create the best drifts, where the wind-chill is least and insulation from received solar radiation is greatest. One of the three largest denning areas worldwide is in Canada. There are three main types of winter refuges: maternity dens, temporary dens and winter shelters. During the winter any bear may dig a temporary den and use it for a few days during a storm, or take shelter in a natural cavity. Winter shelters are used for longer periods of time as resting places. This type of shelter is usually roomier with additional features such as alcoves, porches and ventilation holes. Bears do not hibernate in the strict sense of the word; they have the ability to slow down their metabolism to conserve energy at any time of year. The state of self-induced lethargy while in the shelter allows them to preserve their vital fat reserves. During this time, the body temperature of the polar bear decreases by a few degrees from normal and the respiration rates are markedly reduced. Maternity dens are built and occupied by pregnant females and can vary in size. The denning chamber is at the upper end of an entrance tunnel 1 to 2 m long. It averages 1.5 m in diameter with a height of 90 to 100cm in the middle. Drifting snow seals the entrance. The chamber is higher than the tunnel, trapping bear body heat inside. Dens not only provide a safe place to give birth to her cubs, but are also a place of protection for the cubs during their first few months. During this time she does not leave the den, remaining with her cubs and living on her reserves of fat. In the spring, with her fat severely depleted she must leave to find food to sustain herself and her cubs. After they leave the maternity den she will build temporary refuges in which to nurse, rest and shelter her young cubs and warms them as they all sleep together. She heads with her cubs towards the nearest supply of food, this is usually pack ice.

 

Sight and sounds. Polar bears have good eyesight. Their eyes have inner eyelids that keep the glare of the sun on snow and ice from blinding them. When defending a food source from other bears they use a deep growl. They hiss and snort to show aggression. Angry bears use loud roars and growls. Mothers scold cubs with a low growl.

 

Threats to Survival : Polar bears are one of the animals most threatened by global warming. They depend entirely on sea ice as a platform from which to hunt seals. Reduction of the total ice cover in the Arctic is a serious concern globally. When the ice does not form or forms too late in the season many polar bears starve. In Hudson Bay, scientists have found the main cause of death for cubs to be either lack of food or lack of fat on nursing mothers. Exploitation of minerals and fossil fuels in the Arctic pose a continuous threat. Of the oil and natural gas deposits globally, 20% are located in the Arctic. As the ice cap recedes these become more accessible. Countries are competing which each other in claiming ownership of Arctic and its resources. This can only result in further and more drastic impact on polar bear habitat.

 

Status : IUCN: Vulnerable; CITES: Appendix II; COSEWIC: Special Concern

 

Zoo Diet : Toronto Zoo carnivore diet, dog chow, jumbo smelt and herring, carrots, bean sprouts, Vitamin E and Thiamine supplements."

- Courtesy of the Metro Toronto Zoo

 

© All Rights Reserved - Miles Away Photography

Please! No usage allowed without the consent of Mandi A. Miles

Based out of Flesherton, ON, please write for prices and information!

Miles Away Photography

Life as an albino can be tough in Africa since there is a wide belief that body parts of albinos have magical power.

In Africa many people are affected by albinism characterized by the lack of pigmentation. In addition to sight problems and cancer risks that come with the condition, albinos are subject to persistent beliefs in Africa. In spite of educational campaigns, human sacrifices still occur on the continent. It is not that unusual to find the murder of an albino in the columns of an African newspaper.

Photo is taken in Ganvie in Bénin, West Africa

 

he benefits of niacinamide in skincare

 

Skin Supporting Niacinamide…

 

- is an active form of vitamin B3

- is pH balanced and non-irritating

- is antimicrobial so it balances acne-causing bacteria

- is anti-inflammatory so calms active acne breakouts

- has proven to be as effective as topical antibiotics for acne without the risk of bacterial resistance

- is gentle unlike retinoids and benzoyl peroxide

- has proven to balance oily skin and it doesn’t just mop up excess oil, it slows down how much sebum is released after about 4 weeks of continued use

- strengthens sensitive skin by restoring the skin’s natural barrier function (compromised lipid barrier? Get some niacinamide in your routine!)

- fights against dehydration

- fights free radicals (aka it’s an “anti-aging” ingredient)

- prevents oxidative stress (aka it’s an anti-pollution skincare ingredient)

- has proven to fade pigmentation including acne scars and post-acne spots

- helps calm down redness for acne-prone skin

- helps keep collagen flexible, which helps reduce fine lines – something acne-prone skin is often prone to due to dehydration or damage from past breakouts.

Leucistic Female

American Wigeon AMWI (Anas americana)

 

Martindale Flats

 

Central Saanich BC

 

DSCN3408

at dusk ,heavily cropped

 

Leucism (/ˈljuːkɪzəm/; or /ˈluːsɪzəm/) is a condition in which there is partial loss of pigmentation in an animal resulting in white, pale, or patchy coloration of the skin, hair, feathers, scales or cuticle, but not the eyes. Unlike albinism, it is caused by a reduction in multiple types of pigment, not just melanin.

 

Leucism - Wikipedia

  

Leucism | Definition of Leucism by Merriam-Webster

www.merriam-webster.com/dictionary/leucism

: an abnormal condition of reduced pigmentation affecting various animals (such as birds, mammals, and reptiles) that is marked by overall pale color or patches of reduced coloring and is caused by a genetic mutation which inhibits melanin and other pigments from being deposited in feathers, hair, or skin.

An example of Bird Leucism….

317/366

Radicchio - which tastes as good as it looks. :-)

I didn't know:

>>Pliny the Elder claimed radicchio was useful as a blood purifier and an aid for insomniacs in Naturalis Historia. In fact, radicchio contains intybin, a sedative/analgesic, as well as a type of flavonoid called anthocyanin which is used for making dye-sensitized solar cells.

 

Modern cultivation of the plant began in the fifteenth century, in the Veneto, Friuli Venezia Giulia and Trentino regions of Italy, but the deep-red radicchio of today was engineered in 1860 by the Belgian agronomist Francesco Van den Borre, who used a technique called imbianchimento (whitening), preforcing, or blanching to create the dark red, white-veined leaves: radicchio plants are taken from the ground and placed in water in darkened sheds, where lack of light and ensuing inhibition of chlorophyll production cause the plants to lose their green pigmentation.<<

from: en.wikipedia.org/wiki/Radicchio

Built in 1919-1920, this Chicago School and Sullivanesque-style building was designed by Louis Sullivan for the Farmers and Merchants Union Bank in Columbus, Wisconsin as one of his late-career “jewel box” bank buildings that are largely located in smaller communities throughout the midwest. The building was the last “jewel box bank” designed by Sullivan, and the second-to-last commission of his career, and was intended to communicate the bank as a modern and progressive institution, rather than employing the stodgier and more traditional Classical design found on most other banks of the era. The bank was commissioned by the president of the bank, J. Russell Wheeler, whose wife, Anna May Wheeler, pushed him to commission Sullivan to design a new home for the bank. In addition to Louis Sullivan, the building’s stained glass windows, were designed by architectural decorator Louis J. Millet, and the terra cotta by clay modeler Kristian Schneider, whom developed moulds for the building’s terra cotta, metal, and plaster details. The two artisans worked alongside Sullivan on several other bank projects. The building was heavily documented in Sullivan’s 1924 “A System of Architectural Ornament”, published shortly before he died.

 

The building is clad in red tapestry brick, which features blue and green mixed with the red clay mixture in some bricks, creating variation in color and texture across the facade. The brick creates a backdrop to some of the best terra cotta on any of Sullivan’s projects. The terra cotta features many of the floral and geometric motifs found on Sullivan’s other works, and is arranged similarly to other Sullivan banks that utilized brick cladding. The building features two principal facades, with a narrower facade along James Street, and a broader facade facing Dickason Boulevard. The James Street facade features two openings close to ground level, with the eastern bay housing a large plate glass window, and the western bay housing a doorway flanked by skylights, both of which are recessed under a large terra cotta architrave and flanked by square pilasters with decorative Sullivanesque ornament panels at the capitals. The architrave above the doorway and window is divided into three segments by vertical terra cotta elements that feature floral motifs and, like many Sullivan buildings, appear like plants with roots, branches, and crowns. The outer panels of the architrave feature circular cartouches with hexagonal trim, leaves, and geometric elements, with circular central medallions featuring the years 1861, when the bank was founded, and 1919, when the bank was completed. The central panel is clad in marble with the words “Farmers & Merchants Union Bank” and “Louis Sullivan, Architect” engraved into the stone with yellow pigmentation, contrasting against the white and green marble background. Atop the two vertical elements on either side of the central panel are griffin sculptures holding shields, a common element on many of Sullivan’s “Jewel Box Banks,” while the base of the outer vertical elements features the initials of the bank at the base. Above the architrave is an arched bay that houses a stained glass window, trimmed with decorative terra cotta at the inner and outer rings of the arch, with the bay becoming more recessed after each concentric arch, much like the entrances to medieval Romanesque churches. Besides a band of belt coursing that runs on either side of the architrave and wraps the corner to a tapered buttress on the Dickason Boulevard facade, the only other adornment is an eagle sculpture on a vertical trim element at the center of the parapet, which terminates many brick courses above the arched opening below, and another band of terra cotta trim along the top of the parapet, which forms a cap on the parapet around the perimeter of the building’s low-slope roof. On the Dickason Boulevard facade, the building features five recessed clerestory arched bays housing stained glass windows, flanked by tapered buttresses. Surrounding the arched tops of the windows are decorative trim panels with floral motifs, which begin just below the base of the arches, and extend up above the top of the arches, terminating in a band of belt coursing. Atop the buttresses at either end are trim elements featuring large spheres atop rectilinear legs with floral motifs below, undulating in and out with the brick below. Additionally, a band of belt coursing, which wraps the corner of Dickson Boulevard and James Street, runs beneath the windows, only interrupted by the buttresses. Toward the back, on the building’s original rear wing, there are three windows at eye level in the original building, with bands of belt coursing below and at the top of the parapet. The rear window is a recessed bay window flanked by two pilasters with sullivanesque terra cotta panels, while the smaller windows are flanked by sullivanesque relief panels. The rear wing features a roof at multiple heights, and was extended in 1961 with a matching addition by Law, Potter and Nystrom, since removed. The rear of the taller portion of the building features a simple recessed bay with an arched window, and a similar eagle sculpture and vertical trim piece as on the front facade.

 

Inside, the front wing of the building features a tall banking hall with brick cladding on the walls up to the level of the windows, where it terminates at a wooden sill. The space is split down the middle by a row of brick piers and low walls framing the teller cages, which terminate at the sill line of the windows, dividing the space while still allowing it to read as a single continuous lofty space. The brick forms piers at the teller’s cages, pilasters separating desks on the exterior wall, and low brick walls with marble caps. The upper portion of the walls and the coffered ceiling in this space is finished with white plaster, which gives the space a very vertical and airy feeling, as do the cream-colored terrazzo floors, which feature black edges at the base of the walls, tying the space together. The space features a terra cotta water fountain, or bubbler, also designed by sullivan, which features intricate ornament by Schneider. The space also features two mezzanine balconies with metal railings that run below the arched windows at the front and rear of the space, allowing managers to observe the activities in the lobby and teller area below from the rear balcony, while the front balcony exists solely to balance the space and keep it symmetrical. An office for private conferences with customers was originally located near the front of the space, along with a manager’s office, allowing convenience for customers seeking a meeting with the bank management. The teller’s side of the space also housed the bank’s two vaults and several other private offices. The bank originally featured a large meeting room in the one-story rear wing, behind the vaults, with a women’s waiting room sitting along the Dickason Boulevard side of the rear wing, featuring a bay window and a restroom. The building’s interior has changed in function somewhat due to the growth of the bank, changes in bank operations, and expansion of the building with new additions to house offices and a drive-through in the rear.

 

The building was listed on the National Register of Historic Places in 1972, was designated a National Historic Landmark in 1976, and is a contributing structure in the Columbus Downtown Historic District, listed on the National Register of Historic Places in 1992. The building saw an addition in 2006, clad in buff brick, which replicated a historic building that formerly stood to the east, and wraps the building to the rear, with a two-story section behind a one-story annex that connects the one-story rear wing of the bank to the new building. This wing replaced older additions made in 1961, which matched the one-story rear wing of the historic building, and 1980, which was modern in appearance and slightly recessed along James Street to give precedence to the historic building. The building still functions as the main office branch of the Farmers and Merchants Union Bank, which has grown substantially. The building has been long considered to be among the best of Sullivan’s “Jewel Box Banks,” and has been kept in excellent condition by the bank’s careful and caring generational stewardship.

high iso

On one of my regular cetacean watches at Strumble Head recently I was astonished to capture this very rare Leucistic porpoise. How rare you ask? Well, if my research is correct there has only been seven recorded to this date around the UK coast in over a hundred years. This would therefore be Number eight. Unlike an albino where all pigmentation is lost, in this instance only partial pigment loss occurs and as you can see here the fin and tail are normally pigmented. Very pleased to capture these shots as it was quite a distance from the shore. Interestingly it had a calf with it which is also partially pigmented but not as severely as it's mother.

Amnéville

HUMPBACK WHALES: 2015

 

COOL FACTS:

•Males sing complex songs on wintering grounds in Hawaii that can last up to 20 minutes and be heard 20 miles (30 km) away!

•In the Pacific, humpbacks migrate seasonally from Alaska to Hawaii--they can complete the 3,000-mile (4,830 km) trip in as few as 36 days!

•Humpback whales are well known for their long pectoral fins, which can be up to 15 feet (4.6 m) in length. Their scientific name, Megaptera novaeangliae, means "big-winged New Englander" as the New England population was the one best known to Europeans. These long fins give them increased maneuverability; they can be used to slow down or even go backwards.

•Several hunting methods involve using air bubbles to herd, corral, or disorient fish. One highly complex variant, called "bubble netting" is unique to humpbacks. This technique is often performed in groups with defined roles for distracting, scaring, and herding before whales lunge at prey corralled near the surface.

•Their body coloration is primarily dark grey, but individuals have a variable amount of white on their pectoral fins and belly. This variation is so distinctive that the pigmentation pattern on the undersides of their "flukes" is used to identify individual whales, similar to a human fingerprint.

•Humpback whales are the favorite of whale watchers, as they frequently perform aerial displays, such as breaching (jumping out of the water), or slapping the surface with their pectoral fins, tails, or heads.

•Humpback whales travel great distances during their seasonal migration, the farthest migration of any mammal. The longest recorded migration was 5,160 miles (8,300 km); seven animals, including a calf, completed this trek from Costa Rica to Antarctica.

•Also on wintering grounds, males sing complex songs that can last up to 20 minutes and be heard 20 miles (30 km) away. A male may sing for hours, repeating the song several times. All males in a population sing the same song, but that song continually evolves over time. Humpback whale singing has been studied for decades, but scientists still understand very little about its function.

Source: www.nmfs.noaa.gov/pr/species/mammals/whales/humpback-whal...

   

1101_2011_Update

 

Thanks for the German AtmoSAFE Company choosing this zebrafish photo named "Female Zebrafish-01_Worth $200 US dollars" on their official website homepage.

The homegape URL is " www.atmosafe.net/de.html ".

The application article is "Der Zebrafisch mag keinen Stress" and its URL is " www.atmosafe.net/de/anwendungen/bebrueten-und-zuechten/ze... "

 

0428.2011 Update

 

Thanks for the Anaspec Company choosing this zebrafish photo named "Female Zebrafish-01_worth $200 US dollars" on one Z-Fish Antibodies ad in the 2011 zebrafish meeting brochure (www.union.wisc.edu/zebrafish/).

 

The zebrafish meeting is "4th Strategic Conference of Zebrafish Investigators" to be held January 29th - February 2nd, 2011 at Asilomar Conference Center in Pacific Grove, California.

 

0214.2010 Update

Thanks for the Notre Dame University's NDeRC (Notre Dame extended Research Community) choose this photo as the main photo along their BioEyes website (erc.nd.edu/blogs/bioeyes/) and thier Collaborations website (erc.nd.edu/collaborations/).

 

0422.2009 Update

Thanks for the CBCnews Canada choosing this zebrafish photo named "Female Zebrafish-01_Worth $200 US dollars" on their official website.

The homegape URL is "http://www.cbc.ca/news".

The application article is "The eyes have it" and its URL is "http://www.cbc.ca/news/canada/the-eyes-have-it-1.791619"

 

Acknowledgement :

Thanks for the "Lin Li-Yih Lab"* supplied the zebrafish.

* Lin Li-Yih Lab, The Department of Life Science, The National Taiwan Normal University, ROC.

 

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The following descriptions of zebrafish quote from wikipedia website (URL: en.wikipedia.org/wiki/Zebrafish). All Rights are belonging to "Wikipedia website".

 

Zebrafish:

 

1.Introduce: The zebrafish, Danio rerio, is a tropical freshwater fish belonging to the minnow family (Cyprinidae). It is a popular aquarium fish, frequently sold under the trade name zebra danio, and is an important vertebrate model organism in scientific research.

 

2.Distribution: The zebrafish is native to the streams of the southeastern Himalayan region., including the countries Pakistan, Bangladesh, Nepal, and Myanmar. It arose in the Ganges region in Eastern India. It commonly inhabits streams, canals, ditches, ponds, and slow-moving to stagnant water bodies, including rice fields. Zebrafish have been introduced to parts of the United States, presumably by deliberate release or by escape from fish farms. They have also been sighted in Colombia.

 

3.Description: The fish is named for the five uniform, pigmented, horizontal blue stripes on the side of the body, all of which extend to the end of the caudal fin. Its shape can be described as fusiform and laterally compressed, with its mouth directed upwards. Males are torpedo shaped and have gold stripes between the blue stripes; females have a larger, whitish belly and have silver stripes instead of gold. Adult females will exhibit a small genital papilla in front of the anal fin origin. The zebrafish can grow to 6.4 centimetres (2.5 in), although it is uncommon for them to grow past 4 centimetres in captivity.

The approximate generation time for the Danio is 3–4 months. It has been observed that there must be a male present in order for ovulation and spawning of eggs to occur. Females are able to spawn as often as 2–3 days with hundreds of eggs being laid in each clutch. Upon release from the mother, developmental steps will be made, however without the presence of sperm growth will stop after the first few embryonic cleavages. Fertilized eggs will almost immediately become transparent, which is an important characteristic yielding D. rerio as a convenient research model. Development rapidly progresses, with precursors to all major organs appearing within 36 hours of fertilization. Hatching will take place anywhere from 48–72 hours post-fertilization, depending on the internal conditions of the embryo itself and the external temperature (ideally 28.5 °C). Swimming and feeding behavior are observed to occur approximately 72 hours post-fertilization. The sex of juvenile zebrafish cannot be distinguished except by dissection, and the genetic sex determinants are not clearly understood. The range of life-span for a zebrafish in captivity is around 2–3 years, although in ideal conditions, they may live up to 5 years. The zebrafish is omnivorous, and it primarily eats zooplankton, insects, and phytoplankton. It can eat a variety of foods if its main sources are not readily available.

 

4.Model organism for development and genetics: Zebrafish chromatophores, shown here mediating background adaptation, are studied by scientists D. rerio are a common and useful model organism for studies of vertebrate development and gene function. They may supplement higher vertebrate models, such as rats and mice. Pioneering work of George Streisinger at the University of Oregon established the zebrafish as a model organism; its importance was consolidated by large scale forward genetic screens (commonly referred to as the Tübingen/Boston screens). The scholarly journal Development devoted an issue to research using the fish in celebration of this landmark. An online database of zebrafish genetic, genomic, and developmental information, the Zebrafish Information Network (ZFIN), has been established. D. rerio is one of the few species of fish to have been flown into space.

A Zebrafish Pigment Mutant. The mutant called bleached blond was produced by insertional mutagenesis. The embryos in the picture are four days old. At the top is a wild-type embryo, below is the mutant. The mutant lacks black pigment in the melanocytes because it fails to synthesise melanin properly.

Research with D. rerio has allowed advances in the fields of developmental biology, oncology, toxicology, reproductive studies, teratology, genetics, neurobiology, environmental sciences, stem cell and regenerative medicine, and evolutionary theory. Perhaps its greatest advantages for use in the laboratory as a model system come from its now sequenced genetic code, well understood, easily observable and testable developmental behaviors, and the availability of well-characterized mutants. Zebrafish embryonic development provides advantages over other vertebrate model organisms as well. Although the overall generation time of zebrafish is comparable to that of mice, zebrafish embryos develop rapidly, progressing from eggs to larvae in under three days. The embryos are large, robust, and transparent and develop externally to the mother, characteristics which all facilitate experimental manipulation and observation. Their nearly constant size during early development facilitates simple staining techniques, and drugs may be administered by adding directly to the tank. Unfertilized eggs can be made to divide, and the two-celled embryo fused into a single cell, creating a fully homozygous embryo.

See link for pigmentation mutants of D rerio: www.nature.com/hdy/journal/v97/n3/fig_tab/6800867f5.html#...

A common reverse genetics technique is to reduce gene expression or modify splicing in zebrafish using Morpholino antisense technology. Morpholino oligonucleotides are stable, synthetic macromolecules that contain the same bases as DNA or RNA; by binding to complementary RNA sequences, they reduce the expression of specific genes. The journal Genesis devoted an issue to research using Morpholino oligos, mostly in D. rerio. Morpholino oligonucleotides can be injected into one cell of a zebrafish embryo after the 32-cell stage, producing an organism in which gene expression is reduced in only the cells descended from the injected cell. However, cells in the early embryo (<32 cells) are interpermeable to large molecules, allowing diffusion of Morpholinos between cells. A known problem with gene knockdowns in zebrafish is that, because the genome underwent a duplication after the divergence of ray-finned fishes and lobe-finned fishes, it is not always easy to silence the activity one of the two gene paralogs reliably due to complementation by the other paralog.

Despite the complications of the zebrafish genome a number of commercially available global platforms for analysis of both gene expression by microarrays and promoter regulation using ChIP-on-chip exist.

Zebrafish have the ability to regenerate fins, skin, the heart, and the brain (in larval stages). Zebrafish have also been found to regenerate photoreceptors and retinal neurons following injury. The mechanisms of this regeneration are unknown, but are currently being studied. Researchers frequently cut the dorsal and ventral tail fins and analyze their regrowth to test for mutations. This research is leading the scientific community in the understanding of healing/repair mechanisms in vertebrates.

 

5.Recent developments: In October 2001, researchers from the University of Oklahoma published the complete mitochondrial DNA sequence of D. rerio. The length of the zebrafish mitochondrial genome is 16,596 base pairs. This is within 100 base pairs of other related species of fish, and it is notably only 18 bp longer than the goldfish (Carassius auratus) and 21 bp longer than the carp (Cyprinus carpio). The zebrafish gene order and content is identical to the common vertebrate form of mitochondrial DNA. It contains 13 protein-coding genes and a noncoding control region containing the origin of replication for the heavy strand. In between a grouping of five tRNA genes, a sequence resembling vertebrate origin of light strand replication is found. In comparing the nucleotide sequence to other vertebrates it is difficult to draw any evolutionary conclusions because it is difficult to determine as to whether base pair changes have adaptive significance.

In December 2005, a study of the golden strain identified the gene responsible for the unusual pigmentation of this strain as SLC24A5, a solute carrier that appeared to be required for melanin production, and confirmed its function with a Morpholino knockdown. The orthologous gene was then characterized in humans and a one base pair difference was found to segregate strongly between fair-skinned Europeans and dark-skinned Africans. This study featured on the cover of the academic journal Science and demonstrates the power of zebrafish as a model organism in the relatively new field of comparative genomics.

In January 2007, Chinese researchers at Fudan University raised genetically modified fish that can detect estrogen pollution in lakes and rivers, showing environmental officials what waterways need to be treated for the substance, which is linked to male infertility. Song Houyan and Zhong Tao, professors at Fudan's molecular medicine lab, spent three years cloning estrogen-sensitive genes and injecting them into the fertile eggs of zebrafish. The modified fish turn green if they are placed into water that is polluted by estrogen.

On August 1, 2007, researchers at University College London said they had grown in the laboratory a type of adult stem cell found in the eyes of fish and mammals that develops into neurons in the retina. These cells could be injected in the eye to treat all diseases where the retinal neurons are damaged — nearly every disease of the eye, including macular degeneration, glaucoma, and diabetes-related blindness. Damage to the retina — the part of the eye that sends messages to the brain — is responsible for most cases of sight loss. The researchers studied Müller glial cells in the eyes of humans aged from 18 months to 91 years and were able to develop them into all types of neurons found in the retina. They were also able to grow them easily in the lab, they reported in the journal Stem Cells. The cells were tested in rats with diseased retinas, where they successfully migrated into the retina and took on the characteristics of the surrounding neurons. Now the team is working on the same approach in humans.

In February 2008, researchers at Children's Hospital Boston reported in the journal Cell Stem Cell the development of a new strain of zebrafish, named Casper, with see-through bodies. This allows for detailed visualization of individual blood stem cells and metastasizing (spreading) cancer cells within a living adult organism. Because the function of many genes are shared between fish and humans, this tool is expected to yield insight into human diseases such as leukemia and other cancers.

In April 2009, Researchers at the Institute of Genomics and Integrative Biology, Delhi announced the sequencing of the wild-type strain of Zebrafish, complete with about 1.7 billion genetic alphabets.

 

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Nikon AF-D 60mm F2.8 Macro

 

可以到以下網址察看斑馬魚資料，資料取自維基百科！

 

en.wikipedia.org/wiki/Zebrafish

 

阿鶴已經上傳一套斑馬魚胚胎發育照片，從剛受精到4.5天的仔魚照片都有。，請點擊以下網址：

 

www.flickr.com/photos/chenhowen/sets/72157618669794787/

 

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There is one biochemistry company pay our lab $200 (US dollars) to get the rights to put this photo on their website and their product fliers. 0808.2009

 

有一家美國生技公司花200美金（約台幣6600元）向實驗室買使用權，之後這張斑馬魚照片會放在該公司的網站和產品封面。 0830.2009

Macclesfield Canal - Scholar Green, Cheshire, UK

 

he orange tip, is a butterfly in the family Pieridae, which contains about 1,100 species. A. cardamines is mainly found throughout Europe and temperate Asia The males feature wings with a signature orange pigmentation, which is the origin of A. cardamines' common name.