pigmentation photos on Flickr

Canon 5DS-R_White Moggie_BZ011 by Barry Zee

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The white tiger is a pigmentation variant of the Bengal tiger, which is reported in the wild from time to time in the Indian states of Assam, West Bengal and Bihar in the Sunderbans region and especially in the former State of Rewa.

China - Kashgar - Streetlife - Fruitseller (Pomegranate) by Manfred Sommer

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The pomegranate (/ˈpɒmᵻɡrænᵻt/), botanical name Punica granatum, is a fruit-bearing deciduous shrub or small tree growing between 5 and 8 m tall.

In the Northern Hemisphere, the fruit is typically in season from September to February, and in the Southern Hemisphere from March to May. As intact arils or juice, pomegranates are used in cooking, baking, meal garnishes, juice blends, smoothies, and alcoholic beverages, such as cocktails and wine.

The pomegranate originated in the region of modern-day Iran and has been cultivated since ancient times throughout the Mediterranean region and northern India. It was introduced into America (Spanish America) in the late 16th century and California by Spanish settlers in 1769.

Today, it is widely cultivated throughout the Middle East and Caucasus region, north Africa and tropical Africa, the Indian subcontinent, Central Asia, the drier parts of southeast Asia, and parts of the Mediterranean Basin. It is also cultivated in parts of California and Arizona. In recent years, it has become more common in the commercial markets of Europe and the Western Hemisphere.

ETYMOLOGY

The name pomegranate derives from medieval Latin pōmum "apple" and grānātum "seeded". Perhaps stemming from the old French word for the fruit, pomme-grenade, the pomegranate was known in early English as "apple of Grenada" - a term which today survives only in heraldic blazons. This is a folk etymology, confusing Latin granatus with the name of the Spanish city of Granada, which derives from Arabic.

Garnet derives from Old French grenat by metathesis, from Medieval Latin granatum as used in a different meaning "of a dark red color". This derivation may have originated from pomum granatum describing the color of pomegranate pulp or from granum referring to "red dye, cochineal".

The French term for pomegranate, grenade, has given its name to the military grenade.

DESCRIPTION

A shrub or small tree growing 6 to 10 m high, the pomegranate has multiple spiny branches, and is extremely long-lived, with some specimens in France surviving for 200 years. P. granatum leaves are opposite or subopposite, glossy, narrow oblong, entire, 3–7 cm long and 2 cm broad. The flowers are bright red and 3 cm in diameter, with three to seven petals. Some fruitless varieties are grown for the flowers alone.

The edible fruit is a berry, intermediate in size between a lemon and a grapefruit, 5–12 cm in diameter with a rounded shape and thick, reddish skin. The number of seeds in a pomegranate can vary from 200 to about 1400. Each seed has a surrounding water-laden pulp — the edible sarcotesta that forms from the seed coat — ranging in color from white to deep red or purple. The seeds are "exarillate", i.e., unlike some other species in the order, Myrtales, no aril is present. The sarcotesta of pomegranate seeds consists of epidermis cells derived from the integument. The seeds are embedded in a white, spongy, astringent membrane.

CULTIVATION

P. granatum is grown for its fruit crop, and as ornamental trees and shrubs in parks and gardens. Mature specimens can develop sculptural twisted-bark multiple trunks and a distinctive overall form. Pomegranates are drought-tolerant, and can be grown in dry areas with either a Mediterranean winter rainfall climate or in summer rainfall climates. In wetter areas, they can be prone to root decay from fungal diseases. They can be tolerant of moderate frost, down to about −12 °C.

Insect pests of the pomegranate can include the pomegranate butterfly Virachola isocrates and the leaf-footed bug Leptoglossus zonatus, and fruit flies and ants are attracted to unharvested ripe fruit. Pomegranate grows easily from seed, but is commonly propagated from 25– to 50-cm hardwood cuttings to avoid the genetic variation of seedlings. Air layering is also an option for propagation, but grafting fails.

VARIETIES

P. granatum var. nana is a dwarf variety of P. granatum popularly planted as an ornamental plant in gardens and larger containers, and used as a bonsai specimen tree. It could well be a wild form with a distinct origin. It has gained the Royal Horticultural Society's Award of Garden Merit. The only other species in the genus Punica is the Socotran pomegranate (P. protopunica), which is endemic to the island of Socotra. It differs in having pink (not red) flowers and smaller, less sweet fruit.

CULTIVARS

P. granatum has more than 500 named cultivars, but evidently has considerable synonymy in which the same genotype is named differently across regions of the world.[15]

Several characteristics between pomegranate genotypes vary for identification, consumer preference, preferred use, and marketing, the most important of which are fruit size, exocarp color (ranging from yellow to purple, with pink and red most common), seed-coat color (ranging from white to red), hardness of seed, maturity, juice content and its acidity, sweetness, and astringency.

CULTURAL HISTORY

Pomegranate is native to a region from Iran to northern India. Pomegranates have been cultivated throughout the Middle East, South Asia, and Mediterranean region for several millennia, and also thrive in the drier climates of California and Arizona.

Carbonized exocarp of the fruit has been identified in early Bronze Age levels of Jericho in the West Bank, as well as late Bronze Age levels of Hala Sultan Tekke on Cyprus and Tiryns.[citation needed] A large, dry pomegranate was found in the tomb of Djehuty, the butler of Queen Hatshepsut in Egypt; Mesopotamian cuneiform records mention pomegranates from the mid-third millennium BC onwards.

It is also extensively grown in South China and in Southeast Asia, whether originally spread along the route of the Silk Road or brought by sea traders. Kandahar is famous in Afghanistan for its high-quality pomegranates.

Although not native to Korea or Japan, the pomegranate is widely grown there and many cultivars have been developed. It is widely used for bonsai because of its flowers and for the unusual twisted bark the older specimens can attain. The term "balaustine" (Latin: balaustinus) is also used for a pomegranate-red color.

The ancient city of Granada in Spain was renamed after the fruit during the Moorish period and today the province of Granada uses pomegranate as a charge in heraldry for its canting arms.

Spanish colonists later introduced the fruit to the Caribbean and America (Spanish America), but in the English colonies, it was less at home: "Don't use the pomegranate inhospitably, a stranger that has come so far to pay his respects to thee," the English Quaker Peter Collinson wrote to the botanizing John Bartram in Philadelphia, 1762. "Plant it against the side of thy house, nail it close to the wall. In this manner it thrives wonderfully with us, and flowers beautifully, and bears fruit this hot year. I have twenty-four on one tree... Doctor Fothergill says, of all trees this is most salutiferous to mankind."

The pomegranate had been introduced as an exotic to England the previous century, by John Tradescant the elder, but the disappointment that it did not set fruit there led to its repeated introduction to the American colonies, even New England. It succeeded in the South: Bartram received a barrel of pomegranates and oranges from a correspondent in Charleston, South Carolina, 1764. John Bartram partook of "delitious" pomegranates with Noble Jones at Wormsloe Plantation, near Savannah, Georgia, in September 1765. Thomas Jefferson planted pomegranates at Monticello in 1771: he had them from George Wythe of Williamsburg.

CULINARY USE

After the pomegranate is opened by scoring it with a knife and breaking it open, the seeds are separated from the peel and internal white pulp membranes. Separating the seeds is easier in a bowl of water because the seeds sink and the inedible pulp floats. Freezing the entire fruit also makes it easier to separate. Another effective way of quickly harvesting the seeds is to cut the pomegranate in half, score each half of the exterior rind four to six times, hold the pomegranate half over a bowl, and smack the rind with a large spoon. The seeds should eject from the pomegranate directly into the bowl, leaving only a dozen or more deeply embedded seeds to remove. The entire seed is consumed raw, though the watery, tasty sarcotesta is the desired part. The taste differs depending on the variety or cultivar of pomegranate and its ripeness.

Pomegranate juice can be sweet or sour, but most fruits are moderate in taste, with sour notes from the acidic tannins contained in the juice. Pomegranate juice has long been a popular drink in Europe, the Middle East and is now widely distributed in the United States and Canada.

Grenadine syrup long ago consisted of thickened and sweetened pomegranate juice, now is usually a sales name for a syrup based on various berries, citric acid, and food coloring, mainly used in cocktail mixing. In Europe, Bols still manufactures grenadine syrup with pomegranate. Before tomatoes, a New World fruit, arrived in the Middle East, pomegranate juice, molasses, and vinegar were widely used in many Iranian foods, and are still found in traditional recipes such as fesenjān, a thick sauce made from pomegranate juice and ground walnuts, usually spooned over duck or other poultry and rice, and in ash-e anar (pomegranate soup).

Pomegranate seeds are used as a spice known as anardana (from Persian: anar + dana‎‎, pomegranate + seed), most notably in Indian and Pakistani cuisine. Dried whole seeds can often be obtained in ethnic Indian subcontinent markets. These seeds are separated from the flesh, dried for 10–15 days, and used as an acidic agent for chutney and curry preparation. Ground anardana is also used, which results in a deeper flavoring in dishes and prevents the seeds from getting stuck in teeth. Seeds of the wild pomegranate variety known as daru from the Himalayas are regarded as quality sources for this spice.

Dried pomegranate seeds, found in some natural specialty food markets, still contain some residual water, maintaining a natural sweet and tart flavor. Dried seeds can be used in several culinary applications, such as trail mix, granola bars, or as a topping for salad, yogurt, or ice cream.

In the Caucasus, pomegranate is used mainly for juice. In Azerbaijan, a sauce from pomegranate juice narsharab, (from Persian: (a)nar + sharab‎‎, lit. "pomegranate wine") is usually served with fish or tika kabab. In Turkey, pomegranate sauce (Turkish: nar ekşisi) is used as a salad dressing, to marinate meat, or simply to drink straight. Pomegranate seeds are also used in salads and sometimes as garnish for desserts such as güllaç. Pomegranate syrup or molasses is used in muhammara, a roasted red pepper, walnut, and garlic spread popular in Syria and Turkey.

In Greece, pomegranate (Greek: ρόδι, rodi) is used in many recipes, including kollivozoumi, a creamy broth made from boiled wheat, pomegranates, and raisins, legume salad with wheat and pomegranate, traditional Middle Eastern lamb kebabs with pomegranate glaze, pomegranate eggplant relish, and avocado-pomegranate dip. Pomegranate is also made into a liqueur, and as a popular fruit confectionery used as ice cream topping, mixed with yogurt, or spread as jam on toast. In Cyprus and Greece, and among the Greek Orthodox Diaspora, ρόδι (Greek for pomegranate) is used to make koliva, a mixture of wheat, pomegranate seeds, sugar, almonds, and other seeds served at memorial services.

In Mexico, they are commonly used to adorn the traditional dish chiles en nogada, representing the red of the Mexican flag in the dish which evokes the green (poblano pepper), white (nogada sauce) and red (pomegranate seeds) tricolor.

IN TRADITIONAL MEDICINE

In the Indian subcontinent's ancient Ayurveda system of traditional medicine, the pomegranate is frequently described as an ingredient in remedies.

In folk medicine pomegranate has been thought a contraceptive and abortifacient when the seeds or rind are eaten, or when as a vaginal suppository.

NUTRITION

A 100-g serving of pomegranate seeds provides 12% of the Daily Value (DV) for vitamin C, 16% DV for vitamin K and 10% DV for folate (table).

Pomegranate seeds are an excellent source of dietary fiber (20% DV) which is entirely contained in the edible seeds. People who choose to discard the seeds forfeit nutritional benefits conveyed by the seed fiber and micronutrients.

Pomegranate seed oil contains punicic acid (65.3%), palmitic acid (4.8%), stearic acid (2.3%), oleic acid (6.3%), and linoleic acid (6.6%).

RESEARCH

JUICE

The most abundant phytochemicals in pomegranate juice are polyphenols, including the hydrolyzable tannins called ellagitannins formed when ellagic acid and/or gallic acid binds with a carbohydrate to form pomegranate ellagitannins, also known as punicalagins.

The red color of juice can be attributed to anthocyanins, such as delphinidin, cyanidin, and pelargonidin glycosides. Generally, an increase in juice pigmentation occurs during fruit ripening.

The phenolic content of pomegranate juice is adversely affected by processing and pasteurization techniques.

PEEL

Compared to the pulp, the inedible pomegranate peel contains as much as three times the total amount of polyphenols, including condensed tannins, catechins, gallocatechins and prodelphinidins.

The higher phenolic content of the peel yields extracts for use in dietary supplements and food preservatives.

Health claims

Despite limited research data, manufacturers and marketers of pomegranate juice have liberally used evolving research results for product promotion. In February 2010, the FDA issued a Warning Letter to one such manufacturer, POM Wonderful, for using published literature to make illegal claims of unproven anti-disease benefits.

SYMBOLISM

ANCIENT EGYPT

Ancient Egyptians regarded the pomegranate as a symbol of prosperity and ambition. According to the Ebers Papyrus, one of the oldest medical writings from around 1500 BC, Egyptians used the pomegranate for treatment of tapeworm and other infections.

ANCIENT GREECE

The Greeks were familiar with the fruit far before it was introduced to Rome via Carthage. In Ancient Greek mythology, the pomegranate was known as the "fruit of the dead", and believed to have sprung from the blood of Adonis.

The myth of Persephone, the goddess of the underworld, prominently features the pomegranate. In one version of Greek mythology, Persephone was kidnapped by Hades and taken off to live in the underworld as his wife. Her mother, Demeter (goddess of the Harvest), went into mourning for her lost daughter, thus all green things ceased to grow. Zeus, the highest-ranking of the Greek gods, could not allow the Earth to die, so he commanded Hades to return Persephone. It was the rule of the Fates that anyone who consumed food or drink in the underworld was doomed to spend eternity there. Persephone had no food, but Hades tricked her into eating six pomegranate seeds while she was still his prisoner, so she was condemned to spend six months in the underworld every year. During these six months, while Persephone sits on the throne of the underworld beside her husband Hades, her mother Demeter mourns and no longer gives fertility to the earth. This was an ancient Greek explanation for the seasons. Dante Gabriel Rossetti's painting Persephona depicts Persephone holding the fatal fruit. The number of seeds Persephone ate varies, depending on which version of the story is told. The number ranges from three to seven, which accounts for just one barren season if it is just three or four seeds, or two barren seasons (half the year) if she ate six or seven seeds.

The pomegranate also evoked the presence of the Aegean Triple Goddess who evolved into the Olympian Hera, who is sometimes represented offering the pomegranate, as in the Polykleitos' cult image of the Argive Heraion (see below). According to Carl A. P. Ruck and Danny Staples, the chambered pomegranate is also a surrogate for the poppy's narcotic capsule, with its comparable shape and chambered interior. On a Mycenaean seal illustrated in Joseph Campbell's Occidental Mythology 1964, figure 19, the seated Goddess of the double-headed axe (the labrys) offers three poppy pods in her right hand and supports her breast with her left. She embodies both aspects of the dual goddess, life-giving and death-dealing at once. The Titan Orion was represented as "marrying" Side, a name that in Boeotia means "pomegranate", thus consecrating the primal hunter to the Goddess. Other Greek dialects call the pomegranate rhoa; its possible connection with the name of the earth goddess Rhea, inexplicable in Greek, proved suggestive for the mythographer Karl Kerenyi, who suggested the consonance might ultimately derive from a deeper, pre-Indo-European language layer.

In the 5th century BC, Polycleitus took ivory and gold to sculpt the seated Argive Hera in her temple. She held a scepter in one hand and offered a pomegranate, like a 'royal orb', in the other. "About the pomegranate I must say nothing," whispered the traveller Pausanias in the 2nd century, "for its story is somewhat of a holy mystery." In the Orion story, Hera cast pomegranate-Side (an ancient city in Antalya) into dim Erebus — "for daring to rival Hera's beauty", which forms the probable point of connection with the older Osiris/Isis story.[citation needed] Since the ancient Egyptians identified the Orion constellation in the sky as Sah the "soul of Osiris", the identification of this section of the myth seems relatively complete. Hera wears, not a wreath nor a tiara nor a diadem, but clearly the calyx of the pomegranate that has become her serrated crown.[citation needed] The pomegranate has a calyx shaped like a crown. In Jewish tradition, it has been seen as the original "design" for the proper crown. In some artistic depictions, the pomegranate is found in the hand of Mary, mother of Jesus.

A pomegranate is displayed on coins from the ancient city of Side, Pamphylia.

Within the Heraion at the mouth of the Sele, near Paestum, Magna Graecia, is a chapel devoted to the Madonna del Granato, "Our Lady of the Pomegranate", "who by virtue of her epithet and the attribute of a pomegranate must be the Christian successor of the ancient Greek goddess Hera", observes the excavator of the Heraion of Samos, Helmut Kyrieleis.

In modern times, the pomegranate still holds strong symbolic meanings for the Greeks. On important days in the Greek Orthodox calendar, such as the Presentation of the Virgin Mary and on Christmas Day, it is traditional to have at the dinner table polysporia, also known by their ancient name panspermia, in some regions of Greece. In ancient times, they were offered to Demeter[citation needed] and to the other gods for fertile land, for the spirits of the dead and in honor of compassionate Dionysus.[citation needed] When one buys a new home, it is conventional for a house guest to bring as a first gift a pomegranate, which is placed under/near the ikonostasi (home altar) of the house, as a symbol of abundance, fertility, and good luck. Pomegranates are also prominent at Greek weddings and funerals.[citation needed] When Greeks commemorate their dead, they make kollyva as offerings, which consist of boiled wheat, mixed with sugar and decorated with pomegranate. It is also traditional in Greece to break a pomegranate on the ground at weddings and on New Years. Pomegranate decorations for the home are very common in Greece and sold in most home goods stores.

ANCIENT ISRAEL AND JUDAISM

Pomegranates were known in Ancient Israel as the fruits which the scouts brought to Moses to demonstrate the fertility of the "promised land". The Book of Exodus describes the me'il ("robe of the ephod") worn by the Hebrew high priest as having pomegranates embroidered on the hem alternating with golden bells which could be heard as the high priest entered and left the Holy of Holies. According to the Books of Kings, the capitals of the two pillars (Jachin and Boaz) that stood in front of Solomon's Temple in Jerusalem were engraved with pomegranates. Solomon is said to have designed his coronet based on the pomegranate's "crown" (calyx).

It is traditional to consume pomegranates on Rosh Hashana because, with its numerous seeds, it symbolizes fruitfulness. Also, it is said to have 613 seeds, which corresponds with the 613 mitzvot or commandments of the Torah.[61] This particular tradition is referred to in the opening pages of Ursula Dubosarsky's novel Theodora's Gift.

The pomegranate appeared on the ancient coins of Judea. When not in use, the handles of Torah scrolls are sometimes covered with decorative silver globes similar in shape to "pomegranates" (rimmonim). Some Jewish scholars believe the pomegranate was the forbidden fruit in the Garden of Eden.[60] Pomegranates are one of the Seven Species (Hebrew: שבעת המינים, Shiv'at Ha-Minim) of fruits and grains enumerated in the Hebrew Bible (Deuteronomy 8:8) as being special products of the Land of Israel. The pomegranate is mentioned in the Bible many times, including this quote from the Songs of Solomon, "Thy lips are like a thread of scarlet, and thy speech is comely: thy temples are like a piece of a pomegranate within thy locks." (Song of Solomon 4:3). Pomegranates also symbolize the mystical experience in the Jewish mystical tradition, or kabbalah, with the typical reference being to entering the "garden of pomegranates" or pardes rimonim; this is also the title of a book by the 16th-century mystic Moses ben Jacob Cordovero.

IN EUROPEAN CHRISTIAN MOTIFS

In the earliest incontrovertible appearance of Christ in a mosaic, a 4th-century floor mosaic from Hinton St Mary, Dorset, now in the British Museum, the bust of Christ and the chi rho are flanked by pomegranates. Pomegranates continue to be a motif often found in Christian religious decoration. They are often woven into the fabric of vestments and liturgical hangings or wrought in metalwork. Pomegranates figure in many religious paintings by the likes of Sandro Botticelli and Leonardo da Vinci, often in the hands of the Virgin Mary or the infant Jesus. The fruit, broken or bursting open, is a symbol of the fullness of Jesus' suffering and resurrection.

In the Eastern Orthodox Church, pomegranate seeds may be used in kolyva, a dish prepared for memorial services, as a symbol of the sweetness of the heavenly kingdom.

IN THE QUR´AN

According to the Qur'an, pomegranates grow in the gardens of paradise (55:68). The Qur'an also mentions pomegranates three times.(6:99, 6:141, 55:68)

AFGHANISTAN

Pomegranate, a favorite fall and winter fruit in Afghanistan, has mainly two varieties: one that is sweet and dark red with hard seeds growing in and around Kandhar province, and the other that has soft seeds with variable color growing in the central/northern region. The largest market for Afghan pomegranates is India followed by Pakistan, Russia, United Arab Emirates and Europe.

ARMENIA

The pomegranate is one of the main fruits in Armenian culture (the others being apricot and grapes). Its juice is famous with Armenians in food and heritage. The pomegranate is the symbol of Armenia and represents fertility, abundance and marriage. For example, the fruit played an integral role in a wedding custom widely practiced in ancient Armenia: a bride was given a pomegranate fruit, which she threw against a wall, breaking it into pieces. Scattered pomegranate seeds ensured the bride future children. In Karabakh, it was customary to put fruits next to the bridal couple during the first night of marriage, among them the pomegranate, which was said to ensure happiness. It is likely that newlyweds also enjoyed pomegranate wine. The symbolism of the pomegranate is that it protected a woman from infertility and protected a man's virility. Both homemade and commercial wine is made from pomegranate in Armenia. The Color of Pomegranates (1969) is a movie directed by Sergei Parajanov. It is a biography of the Armenian ashug Sayat-Nova (King of Song) which attempts to reveal the poet's life visually and poetically rather than literally.

AZERBAIJAN

Pomegranate is considered one of the symbols of Azerbaijan. Annually in October, a cultural festival is held in Goychay, Azerbaijan known as the Goychay Pomegranate Festival. The festival features Azerbaijani fruit-cuisine mainly the pomegranates from Goychay, which is famous for its pomegranate growing industry. At the festival, a parade is held with traditional Azerbaijani dances and Azerbaijani music. Pomegranate was depicted on the official logo of the 2015 European Games held in Azerbaijan. Nar the Pomegranate was one of the two mascots of these games. Pomegranates also featured on the jackets worn by Azerbaijani male athletes at the games' opening ceremony.

IRAN AND ANCIENT PERSIA

Pomegranate was the symbol of fertility in ancient Persian culture.[citation needed] In Persian mythology, Isfandiyar eats a pomegranate and becomes invincible. In the Greco-Persian Wars, Herodotus mentions golden pomegranates adorning the spears of warriors in the phalanx. Even in today's Iran, pomegranate may imply love and fertility.

Iran produces pomegranates as a common crop.[citation needed] Its juice and paste have a role in some Iranian cuisines, e.g. chicken, ghormas and refreshment bars. Pomegranate skins may be used to stain wool and silk in the carpet industry.

Pomegranate Festival is an annual cultural and artistic festival held during October in Tehran[citation needed] to exhibit and sell pomegranates, food products and handicrafts.

PAKISTAN

The pomegranate (known as "anār" in Urdu) is a popular fruit in Pakistan. It is grown in Pakistan and is also imported from Afghanistan.

INDIA

In some Hindu traditions, the pomegranate (Hindi: anār) symbolizes prosperity and fertility, and is associated with both Bhoomidevi (the earth goddess) and Lord Ganesha (the one fond of the many-seeded fruit). The Tamil name maadulampazham is a metaphor for a woman's mind. It is derived from, maadhu=woman, ullam=mind, which means as the seeds are hidden, it is not easy to decipher a woman's mind.

CHINA

Introduced to China during the Tang Dynasty (618-907 AD), the pomegranate (Chinese: 石榴; pinyin: shíliu) in olden times was considered an emblem of fertility and numerous progeny. This symbolism is a pun on the Chinese character 子 (zǐ) which, as well as meaning seed, also means "offspring" thus a fruit containing so many seeds is a sign of fecundity. Pictures of the ripe fruit with the seeds bursting forth were often hung in homes to bestow fertility and bless the dwelling with numerous offspring, an important facet of traditional Chinese culture.

WIKIPEDIA

Ophrys fuciflora by Jelltex

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FIXMAS.

6 months to Christmas.

After the hunt was cut short on Friday due to time constraints, I arranged to meet my friend, Terry, back at the site on Sunday morning.

And as Sunday was due to be the warmest, if not hottest day of the year, we were going to be there early.

Jools and I had coffee, then packed to leave, reaching the site via quiet lanes well before eight, but the temperature was already building.

A quick search of the original site, then a walk along the down to the newly found (by us) one, where could already see Terry high above us.

Also there was a guy from Devon, another Graham, and he had come to look to.

So, the four of us began to search the down for the little green bugger.

I am pretty sure we looked at every spike, at least once, and although we found some very nice spikes, not the one we were looking for.

A short drive away is the site I saw the hybrid last year, and this really was the last throw of the dice stuff. Its a long hike up the down, and across to the steep side.

Here, the spikes were already well past their best, with just a couple worth seeing and snapping, but certainly nothing of any note.

It was half eleven, twenty six degrees, and too hot for any more.

We all bailed.

Jools and I drove home, where once inside we both had a long, cold drink before brunch of fruit and croissants.

For three hours we sweltered inside, too hot for anything. But then came the cooking.

I asked Jen what she and Sylv would like to eat on Sunday: "roast" said Jen.

So, on the hottest day of the year, I roasted leg of lamb for some two hours, steamed vegetables, roasted potatoes and baked Yorkshire puddings.

By six, it was like I had crawled in the oven, but the meal was wonderful, and most cleared their plates.

But still hot.

We sat outside in the shade of the house for an hour, just talking, by which time it was eight and time for Jen and Sylv to go home, and for Jools and I to clear away the last of the washing up, have a brew, then go to bed.

Phew rock and roll.

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Classed as Vulnerable in the Red List of threatened orchids, the Late Spider-orchid is one of Britain's rarest orchids. It is now restricted to a few sites in Kent, where many of the plants are protected by cages to prevent damage by grazing animals and other predators. Like its close relative the Bee Orchid Ophrys apifera, Ophrys fuciflora has evolved to mimic the form of a pollinating insect, and this facilitates pollination. Interestingly, the particular species of bees that pollinate Late Spider-orchids on mainland Europe do not occur in the UK, and so what little pollination takes place here must be facilitated by other insects, possibly small beetles. Seed-set is very low, and the likely factor which saves the small populations of Late Spider-orchids in Kent is that fully developed plants live for a long time and therefore only a small number of new plants are required annually in order to keep the population stable. The Late Spider-orchid flowers from late May to late July. Ophrys fuciflora is widespread in other parts of Europe from France eastwards to Romania and south to Italy.

Plant: 5-30cm.

Leaves: 3-5 lanceolate leaves form a basal rosette of which the lower leaves lie flat on the ground. They are greenish-grey and prominantly veined. There are 2-3 smaller, more pointed leaves higher up and loosely sheathing the stem.

Bracts: lanceolate and grey-green.

Flowers: the sepals vary in colour from pale- to dark pink with a prominent green 'rib' on their outer surfaces, and they are green-veined on the inner surface. The triangular petals are much smaller than the sepals and sometimes have dark reddish swellings (auricles) towards the base. The lip is a rich velvety dark brown and its shape is noticeably square. Square 'shoulders' at the base of the lip vary in size and are often hairy. The speculum (mirror), which is extremely variable in colour and patterning, radiates from a semi-circular 'necklace' that surrounds the column.

The Late Spider-orchid belongs to the Ophrys genus. Its Latin name derives from the words 'focus' and 'flos' meaning 'bee-flowered'.

There are no subspecies, but there is such considerable range of colour and patterning of the lip as to invite numerous suggested variations, notably Ophrys fuciflora var. flavescens which lacks colour pigmentation and has white sepals and petals, a greenish lip with very faint markings. Two hybrids are recorded, Ophrys x albertiana is a hybrid with the Bee Orchid Ophrys apifera, while Ophrys x obscura is a hybrid with the Early Spider-orchid Ophrys sphegodes.

www.hardyorchidsociety.org.uk/hos%201012/orchidphotos/oph...

Postojna Cave #2 by Nabil z.a.

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Again, another shot of the Postojna Cave, a 20 km long limestone cave situated not far from Ljubljana, the capital city of Slovenia. This cave is the most visited cave in Slovenia, and some say in Europe as well. The cavern is carved and shaped by water, which dissolved the limestones to create holes, stalagmites and stalactites. Entrance is only by guided tour and the ticket is quite pricey in comparison to other attractions in Europe. And you might want to bring a jacket because the temperature down here is a constant 10C. And bring comfortable shoes as well as you have to walk for about an hour and a half within this cavern and the paths can get quite slippery. You can get more description and details on this cave in my previous post.

The booth look-alike structure in the picture above is actually an aquarium which is the end station of our tour.The aquarium displays a rare unique species that can only be found in Karst limestone region. The creature which is nicknamed the human fish is an amphibian living in the waters that flow through this extensive limestone region. The human fish or the ohm's (Proteus anguinus) body is similar to snakes but its skin lacks pigmentation. It eyes are undeveloped since it lives in complete darkness in its underground habitat. Look at the comment section for a picture link of the creature.

.

Vertorama from 2 horizontal shots

Shot with Tokina 11-16mm f/2.8 @11 mm

Aperture of f/6.3 with ISO value of 200

Color Transformation by Ruth and Joe Barrett

5

Powdery scales on the wings of the male butterfly Apatura ilia under optical microscopy reveal parallel fibers with a spacing of about 710 nm; these fibers, together with tiny transverse grooves, cause violet blue or blue iridescence due to interference of light on the wings that ordinarily have brown pigmentation, establishing the dependence of the sudden color changes on variable illumination and observation angle

Der Kleine Schillerfalter: Die Falter kommen in der Grundfarbe als braun-weiße (Blauschiller) und als rötlichbraune Form (Rotschiller) vor, wobei die Flächen der Flügeloberseiten schillern bei den Männchen blau

________________________________________________

Babenhausen 28AUG2019 – Lesser Purple Emperor Butterfly:

This male Apatura ilia form clytie butterfly flew onto my shoe! And as if that were not amazing enough, as it turned, its color changed from 42 shades of ordinary brown to become iridescent blue!

Billions of tiny scales – in 2 types – on the dorsal wing overlap like roof tiles and cover the membrane surface, to the naked eye appearing as dust, producing a brilliant color due to multilayer photonic crystal micro- and nanostructure, transforming a plain brownish hue to a stunning violet-blue, a phenomenon used to attract mates, camouflage, and startle predators: wingtop color changes in the wake of the variation of the incident light angle.

Even though this is the first time I've identified it, the Lesser Purple Emperor butterfly's range, the Palearctic [Palaearctic] (first identified in the 19th century and still used as the basis for zoogeographic classification), is the largest of the eight biogeographic realms on the Earth's surface, stretching across Europe, Asia north of the Himalayan foothills, North Africa, and northern and central parts of the Arabian Peninsula.

Hope you enjoy the 26% of 61 butterfly photos I took this day!

________________________________

– Album – Germany – 26AUG-11SEP2019:

Three days before leaving, I got my schedule rearranged to free up time for Joe & me to spend two weeks in and around Babenhausen!

LEGO Minifigures 'The Simpsons' Series #15 - Cheif Wiggum by Zed The Dragon

6 1

Lego Simpsons 71005

The Simpsons is an American adult animated sitcom created by Matt Groening for the Fox Broadcasting Company.The series is a satirical depiction of a middle class American lifestyle epitomized by its family of the same name, which consists of Homer, Marge, Bart, Lisa, and Maggie. The show is set in the fictional town of Springfield and parodies American culture, society, television, and many aspects of the human condition.

The family was conceived by Groening shortly before a solicitation for a series of animated shorts with the producer James L. Brooks. Groening created a dysfunctional family and named the characters after members of his own family, substituting Bart for his own name. The shorts became a part of The Tracey Ullman Show on April 19, 1987. After a three-season run, the sketch was developed into a half-hour prime time show and was an early hit for Fox, becoming the network's first series to land in the Top 30 ratings in a season (1989–1990).

Since its debut on December 17, 1989, the show has broadcast 548 episodes and the 25th season began on September 30, 2013. The Simpsons is the longest-running American sitcom, the longest-running American animated program, and in 2009 it surpassed Gunsmoke as the longest-running American primetime, scripted television series. The Simpsons Movie, a feature-length film, was released in theaters worldwide on July 26 and 27, 2007, and grossed over $527 million.

--------------

Les Simpson (The Simpsons) est une série télévisée d'animation américaine créée par Matt Groening et diffusée depuis le 17 décembre 1989 sur le réseau FOX.

Elle met en scène les Simpson, stéréotype d'une famille de classe moyenne. Leurs aventures servent une satire du mode de vie américain. Les membres de la famille, tous ayant la pigmentation de peau de couleur jaune, sont Homer, Marge, Bart, Lisa et Maggie.

[My Portfotolio] [My Google + ]

Thanks to all for visits and faves :)

[My GETTY Images @] [My MOST FAVE on Flickriver] [My RECENT on Fluidr] [My STREAM on Darckr]

4 common skin problems and how to fix them by alan broaderic

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Skin is the most visible and the most important part of your body. So, if you want to look good and beautiful, taking care of your skin is extremely important. In order to ensure that you always look fresh, lively and gorgeous, it is important that you adopt a regular skin care regime. This routine will keep your skin protected and prevent any major skin troubles. However, no matter how much you take care, there could be times when your skin does get affected with some very common skin related issues. These skin problems aren’t too big to be worried about – you simply need to understand them and take quick, corrective action.

Every woman is obsessed about her skin – some may be too obsessed while others are less. Skin is the biggest organ of your body and is also the biggest indicator of your beauty. Skin is also very prone to show the effects of weather changes. While there isn’t much you can do to prevent the bad impact of seasonal changes on your skin, there are still ways that will help you in keeping your skin in its best form. Here are some of the very common skin problems and how you can deal with them.

The 4 common skin problems

Redness

The basic cause of redness on your skin is over exposure to the sun’s UV rays. Constant exposure to sun causes your skin’s capillaries to burst and this is visible as the redness. Also, as you age your skin becomes thinner making the blood vessels underneath it more prominently visible. Rosacea is a skin condition that causes redness too. This is a chronic condition that involves the inflammation of the blood vessels under the skin. This redness comes and goes when the skin is exposed to sun, heat or stress.

Topical products that contain sulphur and azelaic acid are good to treat redness caused by rosacea. For the general redness, using a sunscreen before stepping out in sun will help a lot.

Sun damage and lines

As you age, the production of collagen and elastin gets reduced in your body. This directly impacts the elasticity of your skin. The result of this is the appearance of fine lines (read wrinkles) on your face. Over exposure to sun causes an increase in the number of pigment production cells in your skin and this causes the dark spots.

To take care of the fine lines, products that contain retinol build up collagen will help. For pigmentation, bleaching creams that contain about 2% hydroquinone or kojic acid are helpful.

Allergic and irritated skin

Eczema and dermatitis are common skin problems that are caused due to allergic reactions. Over the counter drugs and creams are helpful in dealing with most allergic reactions on skin. Hydrocortisone creams are good and drugs like ciclosporin can help in dealing with these skin conditions.

Bumps, lumps and growths

Another common issue with your skin is the appearance of some kinds of growths. One thing that you can be rest assured about is that not all growths are problematic. The most common types of growths are seborrheic keratoses, which are harmless black or brown growths that usually crop up as your skin ages. Warts are other common form of skin growths. These growths are usually painless so you don’t need any medications to treat them but if you want to remove them, you will have to visit a dermatologist.

Problems related to skin can be visually more disturbing than causing you pain. When you face these issues, try to stay calm and look for appropriate solutions as mentioned above.

The Humpbacks Are Back! by John C. Bruckman

5 2

The spring and early summer seasons are always exciting due to the miracle of the newborn marine mammals and birds in the Monterey Bay National Marine Sanctuary!

Cool Fact: Humpback individuals have a variable amount of white on their pectoral fins and belly. This variation is so distinctive that the pigmentation pattern on the undersides of their "flukes" is used to identify individual whales, similar to a human fingerprint. The boat captains photograph the tails and send them to a website that tracks the number and location of all Humpbacks whales on the West Coast.

Cool Fact: Humpback whales breathe voluntarily, unlike human beings. Since they have to remember to breathe, researchers believe humpback whales sleep by shutting off half of their brain at a time.

Whale Watching Boat Recommendation: Having been on almost every whale watch boat in the Monterey Bay, I highly recommend the "Sea Goddess" based in Moss Landing. They have the finest boat in the Bay and an excellent crew. You can contact them at: seagoddesswhalewatch.com/.

Lizzie by Patrick Marioné

4

le Mascaret, Rixensart

Antarctic Orcas by Tim Melling

30 16

These were two animals from a large pod of Orcas that we found adjacent to the sea ice on the Antarctic peninsula. Orcas in the Antarctica tend to look yellow rather than white but this is not their own pigmentation. This is a coating of algae known as diatoms, which gradually build up on cetaceans. The older the animal, the yellower it looks. It wasn't just Orcas that were stained. The pale right lower lip of Fin Whales appeared yellow, and even the Blue Whales looked a little greenish.

Orcas can be divided into types that are rather similar in appearance, yet are ecologically very different, and have not interbred with other Orca types for thousands of years. There are three types that occur in Antarctica and this one is a Type B (Pack Ice Orca) which have a very large eye patch and the rear of the body is paler than the front. They hunt the edge of the pack ice primarily for seals.

One hundred and thirty six by Jelltex

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There different thoughts on what is and isn't either var. alba.

In Orchis species, for me, the stem and hood has to be green, not just the interflorances be white. Straw coloured hoods means there is some pigmentation present.

Anyway, good to find my old friend growing in the same place deep in a beech wood this afternoon.

abstract by Patrick Marioné

6

Blow by Patrick Marioné

4

Brussels

How to minimize problems with bleaching skin by Skin Lightening Cream

via

The process of bleaching skin involves the use of substances, concoctions or physical treatments that result in lightening the color of the skin. The idea is to decrease the amount of melanin, a substance naturally found in the body that is responsible for the dark pigmentation of the skin. The market is saturated with numerous brands claiming the same thing so it’s necessary to select the product that is right for your needs.

Skincare tips of the day

What conditions do skin whitening products address?

Skin whitening can either be prescribed by a doctor resulting from certain medical conditions or in some cases, it can be purely cosmetic. Some conditions requiring whitening are as follows:

Dull skin due to overexposure to the sun

Dark spots – from acne, freckles, stress and/ or diet

Certain medical conditions

Dry and irritated skin

Chronic use of cosmetics causing blemishes

Acne scars

What are the possible side effects of skin whitening?

Due to the components used in the production of skin whitening products, there are several possible side effects that can occur. Experiencing a mild burning sensation, stinging, rashes, dryness and flaking are but a few of the possible risks even when using whitening products.

Is there a way to avoid these side effects?

Before heading to the counter to pay, here are some tips in choosing the best skin whitening cream. There are numerous products available on the market today and without the right knowledge, it’s not easy to choose the best product for your specific needs.

What’s your skin type?

Skin types range from normal, dry, oil or a combination thereof. There are some face whitening creams that use strong bleaching components that can cause drying, resulting in irritation. These components are recommended for oily skin types and not for dry or combination types.

Do I need to seek medical advice?

Prior to using any skin lightening creams, make sure to consult a doctor to avoid allergic reactions. Some of these products may contain inactive ingredients that can result in minor skin problems. Be sure to provide an accurate medical history, especially those related to skin conditions.

In order to avoid the risks associated with even the best skin bleaching cream, it is highly recommended to go for natural skin bleaching. This helps to avoid occurrences of allergic reactions and breakouts.

What results do you wish to achieve?

Overall, the main goal is to achieve a lighter complexion but some of the best skin whitening cream can also treat problems like the following:

Dark underarms, elbows and/ or knees

Birthmarks

Acne scars

Hyperpigmentation from keloids or scarring

What’s on the label?

When choosing a product, especially whitening cream for face, always make sure to check what is written on the label. Steer clear of products that contain ingredients such as steroids and mercury. These chemicals are toxic to the skin and can lead to very serious side effects. To get the right product, always be conscious of what’s ne the label.

Black and white bleaching cream for instance, is one of the brands available on the market today. It is very effective in lightening the skin color, resulting in an even skin tone. It can remove scars quickly and reduce the presence of freckles. Not only does it act as a whitening cream for face but also, the formulation makes it possible to use the product as a make-up base. It smoothens the skin, creating an even palate to work with when applying make-up. To achieve the best results from this skin whitening treatment, it is recommended to use it for about 2 to 3 weeks.

The active ingredient contained in this product is hydroquinone. It acts by reducing the level of melanin present in the skin resulting in a whiter complexion. The major risk in reduced melanin levels is that it can weaken the natural defense of the skin, leading to breakouts, irritation and other skin problems. But to address this, it is infused with vitamin C that enhances cell growth, resulting to supple and younger looking skin. In addition, it has powerful antioxidants that fight free radicals and slow down the signs of aging.

Never forget SPF!

Bleaching can result in extremely sensitive skin. This is because bleaching causes layers of the skin to “peel off” in order to produce an even skin tone. This causes the deeper layers of the skin to be exposed to the harsh environment, hence making it prone to freckles, sun spots, rashes and sunburn. The latter two can be extremely painful, especially if facial bleaching is involved.

To prevent these nasty conditions, always wear sunscreen to protect your skin from harsh elements. This will keep your skin tone even for a longer period of time.

Skin bleaching creams

Knowing the right product for your skin is essential in preventing the risks associated with whitening. Being informed on what your skin needs will give you a good idea on how to do it safely. When in doubt, always seek medical advice before buying a product. That way, you do not waste time, money and resources on any product. Bleaching skin can come with minimal risk when you know how to select the right products.

The post How to minimize problems with bleaching skin appeared first on Find The Best Skin Lightening Cream.

skinlighteningcreams.net/how-to-minimize-problems-with-bl...

White Bengal Tiger 2019 by James Arup Photography

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The white tiger or bleached tiger is a pigmentation variant of the Bengal tiger, which is reported in the wild from time to time in the Indian states of Madhya Pradesh, Assam, West Bengal and Bihar in the Sunderbans region and especially in the former State of Rewa. Such a tiger has the black stripes typical of the Bengal tiger, but carries a white or near-white coat.

Laguna Colorada (Red Lagoon), Bolivia by Ian Cowe

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Laguna Colorada (Red Lagoon) is a shallow salt lake in the southwest of the altiplano of Bolivia, within Eduardo Avaroa Andean Fauna National Reserve and close to the border with Chile.

The reddish color of its waters, which is caused by red sediments and pigmentation of some algae.

The birds which can be seen in some of the pictures are flamingos.

~Pre-moult Dicyrtomina ornata~ by Eddie The Bugman

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I think this is a Dicyrtomina ornata, I'm sure it's not a Dicyrtomina saundersi due to the more solid pigmentation pattern on the posterior and it seems too pigmented to be a Dicyrtomina minuta. This one was about 1.5mm in length.

Anyway, this is a pre-moult individual, it is showing the classic white feet and the mottled pigmentation on the abdomen. Eventually it will enter a pharate state and will begin its moult. This one was found in a woodland environment and I used a white card as a background. This was a x7 magnification handheld focus stack of 18 images using an F/5 aperture.

I hope to try for some more focus stacks this weekend, that's if the weather behaves, still its not too bad for the middle of November.

I hope everyone has a great weekend :o)

VIEW LARGE

This image is taken from United States Naval Medical Bulletin Vol. 25, Nos. 1-4, 1927 by Medical Heritage Library, Inc.

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Go to the Book with image in the Internet Archive

Title: United States Naval Medical Bulletin Vol. 25, Nos. 1-4, 1927

Creator: U.S. Navy. Bureau of Medicine and Surgery

Publisher:

Sponsor:

Contributor:

Date: 1927-01

Language: eng

Table of Contents

Number 1

PREFACE v

NOTICE TO SERVICE CONTRIBUTORS vi

SPECIAL ARTICLES:

Posture.

By Lieut. G. E. Mott, Medical Corps, United States Navy 1

Medical Tactics in Naval Warfare —Part III— Continued.

By Commander W. L. Mann, Medical Corps, United States Navy, and Maj. A.

D. Tuttle, Medical Corps, United States Army 20

Identification by the Teeth.

By J. H. Taylor, Identification Section, Bureau of Navigation, Navy

Department 49

An Analysis of the Annual Physical Examination of a Group of Officers.

By Lieut. Commander L. G. Roddis and Lieut. (Junior Grade) G. A.

Cooper, Medical Corps, United States Navy 54

Dressing Sterilizers with Special Reference to Temperature, Pressure,

and Chamber Air Exhaustion During the Process of Sterilization.

By Lieut. Commander J. Harper, Medical Corps, United States Navy 62

Calcium Hypochlorite for Lyster Bags.

By Lieut. E. M. Steger, Medical Corps, United States Navy 6S

The Use of Modified Milk in Infant Feeding.

By Lieut. Commander W. H. Short, Medical Corps, United States Navy 73

Aviation Crashes at Pensacola— 1925-26.

By Lieut. J. D. Benjamin, Medical Corps, United States Navy.. 86

Climatic Bubo.

By Lieut. Commander W. H. Whitmore, Medical Corps, United States Navy

89

Ideas on Recruiting.

By Lieut. J. G. Smith, Medical Corps, United States Navy 102

CLINICAL NOTES:

One Thousand Operations During a Shore Cruise.

By Lieut. Commander G. F. Cottle, Medical Corps, United States Navy 105

Filariasis.

By Lieut. Commander H. V. Hughens, Medical Corps, United States Navy. .

111

Bismuto-Yatren A and B in the Treatment of Yaws.

By Lieut. Commander R. P. Parsons, Medical Corps, United States Navy

117

Addison's Disease Without the Usual Pigmentation of the Skin.

By Commander E. C. White and Lieut. (Junior Grade) W. F. James, Medical

Corps, United States Navy 122

Scurvy.

By Lieut. Commander L. J. Roberts, Medical Corps, United States Navy

126

Early Pulmonary Tuberculosis With Negative X-ray Findings.

By Lieut. J. D. Blackwood, jr., Medical Corps, United States Navy 128

Cholecystography.

By Lieut. O. B. Spalding, Medical Corps, United States Navy.. 131

Syphilis.

By Lieut, (junior grade) B. W. Harris, Medical Corps, United States

Navy 134

Yaws Sera and the Kahn Precipitation Test, Experiments With.

By Pharmacist F. O. Huntsinger, United States Navy 135

NOTES AND COMMENTS:

Commendation for articles in the Bulletin—Treatment of burns — Parasitic

infections in China—Effect of elevation of temperature on spirochetes —Rabies

—-American Relief Administration in Russia, 1921-1923— Forecasting smallpox

epidemics in India —Helium-oxygen mixture in diving —Treatment of pernicious

anemia by diet —International meeting on cancer control, September 20-24, 1926

—American College of Surgeons —Resignation after special courses—Study courses

for Hospital Corps ratings 137-160

NURSE CORPS:

Laboratory Experiences with Epidemic Cerebrospinal Meningitis.

By Nurse S. Ruth Hassler, United States Navy 161

Help from the Laboratory.

By Nurse Ruth B. Meutzer, United States Navy 164

Some Interesting Laboratory Work.

By Nurse Frances C. Bonner, United States Navy 166

BOOK NOTICES 169

PREVENTIVE MEDICINE, STATISTICS:

Reactions incidental to the administration of 91,707 doses of

neoarsphenamine and other arsenical compounds in the United States Navy—Food

poisoning on board U. S. S. "Concord" May 13, 1926—Food poisoning

following a barbecue —Food poisoning at marine barracks, navy yard,

Philadelphia, Pa., May 21, 1926 — Influenza in Guam —Efficacy of B. typhosus

vaccine in controlling typhoid fever in Guam 177

Public Health Activities Against Tropical Diseases.

By Dr. Pedro N. Ortiz, commissioner of health, Porto Rico 208

Scarlet fever attack rate among contacts, Detroit, Mich.—Health of the

Navy— Statistics 220

Number 2

PREFACE---------------- --------- ------- - ------------- ------ vii

NOTICE TO SERVICE CONTRIBUTORS____________________________ viii

SPECIAL ARTICLES: .

MEDICINE IN TURKEY.

By Commander C. W. 0 . Bunker, Medical Corps, United States

Navy------------------------·------ 229

COLOR-BLINDNESS.

By Capt. E. H. H. Old, Medical Corps, United States Navy______ 253

ETIOLOGICAL FACTORS IN THE PRODUCTION OF MAXILLARY ANTRUM DISEASE.

By G. B. Trible, M. D., F. A. C. S., former commander, medical Corps,

United States Navy________ 266

BUREAU OF MEDICINE AND SURGERY'S EXHIBIT, SESQUICENTENNIAL,

PHILADELPHIA, 1926.

By Capt. R. C. Holcomb, Medical Corps, United States Navy____ 272

MALARIA IN HAITI.

By Capt. O. S. Butler, and Lieut. E. Peterson, Medical Corps, United

States Navy_____________ 278

INSTITUTIONAL POLICIES AS APPLIED TO A YEAR'S DENTAL ACTIVITIES.

By Lieut. Commander J. L. Brown, Dental Corps, United States

Navy________________ __________ 288

RECURRENCE OF INGUINAL HERNIA.

By Lieut. Commander K. E. Lowman, Medical Corps, United States Navy

------------------------- 300

SOME OBSERVATIONS ON CEREBROSPINAL FEVER.

By Lieut. Commander Paul Richmond, Medical Corps, United States

Navy__________________ 304

THEORIES CONCERNING THE MECHANISM OF THE INSULIN EFFECT ON CARBOHYDRATE

METABOLISM.

By Lieut. (Junior Grade) H. Phillips, Medical Corps, United States

Navy--------------------------- 309

THE WASSERMANN REACTION FOLLOWING THE USE OF BISMUTH IN THE TREATMENT

OF YAWS AND SYPHILIS.

By Lieut. (Junior Grade) C. D. Middlestadt, Medical Corps, United

States Navy _______________ 315.

THE TUBERCULOUS VETERANS' BUREAU PATIENT.

By Lieut. J. D. Blackwood, jr., Medical Corps, United States

Navy--------------------------------------- 319

CLINICAL NOTES:

OCULAR ENUCLEATION.

By Commander C. B. Camerer, and Lieut. Commander B. P. Davis, Medical

Corps, United States Navy________________ 325

TREATMENT OF AN IRREDUCIBLE DISLOCATED LOWER JAW OF 98 DAYS' DURATION.

By Lieut. Commander M. D. Willcutts, Medical Corps, United States Navy

------------------ 331

TRAUMATIC RUPTURE OF THE SPLEEN.

By Commander L. W. Johnson, Medical Corps, United States

Navy-------------------------------- 336

DIATHERMY IN SHOCK.

By Commander G. F. Cottle, Medical Corps, United States Navy_ 340

SPONTANEOUS RUPTURE OF THE HEART.

By Lieut. (Junior Grade) G. 'w. Cooper, Medical Corps, United States

Navy ------------------------------- 343

MERCUROCHROME-220 SOLUBLE, FOREIGN PROTEIN, AND SUGAR IN ACUTE

GONORRHEAL URETHRITIS, WITH A STUDY OF THE BLOOD CELLULAR CHANGES DURING THE

REACTION.

By Lieut. Commander L. H. Williams, and Lieut. (Junior Grade) G. F.

Cooper, Medical Corps, United States Navy _________ 352

GLANDULAR FEVER WITH INGUINAL ADENOPATHY.

By Lieut. (Junior Grade) C. C. Yanquell, Medical Corps, United States

Navy ___________358

A CASE OF ACQUIRED HYDROCEPHALUS.

By Capt. E. O. White, Medical Corps, United States Navy ____ 361

CARCINOMA OF THE STOMACH WITH LARGE RETROPERITONEAL TUMOR.

By Lieut. Commander W. H. Whitmore, Medical Corps, United States Navy----------------

365

HEXAMETHYLENAMINE IN MENINGEAL INFECTIONS.

By Lieut. (Junior Grade) A. E. Bruuschwig, Medical Corps, United States

Navy ________369

REINFECTION IN SYPHILIS.

By Lieut. O. P. Archambeault, Medical Corps, United States Navy--------------------------------------372

AN UNUSUAL BULLET WOUND. ·

By Lieut. (Junior Grade) P. H. Golberg, Medical Corps, United States

Navy __________374

THE NAVAL RESERVE -------------------------------------------- 375

NURSE CORPS:

THE PENSACOLA HURRICANE.

By Chief Nurse Mary J. McCloud, United States Navy ______ 379

MY FIRST EXPERIENCE IN A HURRICANE.

By Reserve Nurse Luama A. MacFarland, United States Navy _ 382

A HURRICANE THRILL.

By Nurse Ellen E. Wells and Nurse Mary Hennemeier, United States Navy

-------------------383

AN EPIDEMIC OF IMPETIGO CONTAGIOSA.

By Nurse Caroline W. Spofford, United States Navy _________ 385

NOTES AND COMMENTS:

A new section of the BULLETIN-Change in character of –examinations for

promotion-Hospital Corps instruction-Duty at sea – Handling poisons-Laboratory

technicians-The Henry S. Wellcome medal and prize for 1927-Micro-Kahn

reactions-Streptococcus cardioarthritidis--Progress in dermatology-Mitral

regurgitation- Cutaneous leishmaniasis

and the phlebotomus-Medicinals and dyes-Rectal feeding-Epidemic

encephalitis-Catarrhal jaundice- Excoriation of the skin about intestinal

fistulae-Ethylene

anesthesia-Sodoku in the treatment of general paresis—Carbuncles of the

neck-Gye's theory of cancer-$100,000 offered for conquest of cancer-The Sofie

A. Nordoff-Jung cancer prize ____387

BOOK NOTICES ------------------- 413

TOTAL FUEL REQUIREMENT IN HEALTH.

By Commander J. R. Phelps, Medical Corps, United States Navy_ 431

QUESTIONS AND ANSWERS ON SMALLPOX AND VACCINATION.

BY Surg. J. P. Leake, United States Public Health Service______ 461

A fatal case of food poisoning caused by fried oysters contaminated

with a paratyphoid B. bacillus-Outbreak of food poisoning in the wardroom mess

of the U. S. S. Richmond, caused by chicken

salad contaminated with B. enteritidis-------------------------- 475

R EPORT OF AN OUTBREAK OF FISH POISONING ON BOARD THE U. S. S. “CALIFORNIA.''

By Capt. J. L. Neilson, Medical Corps, United States Navy ------ 480

AN OUTBREAK OF FOOD POISONING CAUSED BY BOILED SMOKED TONGUE.

By Lieut. Commander W. A. Fort, Medical Corps, United States Navy -------------------------------------

484

Report of an outbreak of food poisoning caused by cheese-Outbreak of

food poisoning caused by corned-beef hash at the United States destroyer base,

San Diego, Calif. _______________ 486

REPORT OF POISONING BY TRINITROTOLUENE AMONG ENLISTED MEN ENGAGED IN

TRANSFERRING T. N. T. FROM STORAGE TO U. S. S. "NITRO."

By Lieut. Commander T. A. Fortescue, Medical Corps, United States

Navy_______________________________________________ 491

An epidemic of jaundice in San Diego, Calif.-Health of the Navy___ 494

Number 3

PREFACE ---------------------------------------------------------- v

NOTICE TO SERVICE CONTRIBUTORS--------------------------- vi

SPECIAL ARTICLES :

SOME OBSERVATIONS ON AVOIDABLE DROWNING.

By Lieut. Commander R. B. Miller, Medical Corps, United States Navy

___________505

FLIES AND THEIR ERADICATION.

By Lieut. Commander E. C. Carr, Medical Corps, United States

Navy----------------------------- 528

ACTIVITIES OF THE UNITED STATES NAVAL MEDICAL SUPPLY DEPOT.

By Capt. D. C. Cather, Medical Corps, United States Navy ___ 542

RETINITIS PIGMENTOSA.

By Commander C. B. Camerer, Medical Corps, United States

Navy---------------------------------- 562

PREMATURE CONTRACTIONS OF THE HEART.

By Capt. E. C. White, Medical Corps, United States Navy ___ 567

ROENTGEN RAY EXAMINATION IN SUSPECTED CHRONIC APPENDICITIS.

By Lieut. P. P. Maher, Medical Corps, United States Navy ____ 573

COMPARISON OF THE KAHN AND KOLMER REACTIONS.

By Lieut. Commander P. Richmond, jr., Medical Corps, United States

Navy--------------------585

COMMENTS ON THE ORIGIN OF DISABILITY.

By Lieut. Commander Lewis W. Johnson, Medical Corps, United States

Navy--------------------------------- 588

DEPARTMENT OF SANITATION, MARINE BARRACKS, QUANTICO, VA.

By Capt. W. M. Garton, Medical Corps, United States Navy ___593

CLINICAL NOTES :

CHOLECYSTOGRAPHY,

By Lieut. G. B. Larson, Medical Corps, United States Navy __597

THROMBO-ANGIITIS OBLITERANS.

By Lieut. (Junior Grade) R. A. Schneiders, Medical Corps, United States

Navy----------------------------605

THE DEBT OF SURGICAL DIAGNOSIS TO THE X RAY.

By Commander G. F. Cottle, Medical Corps, United States Navy_ 614

THE TREATMENT OF CHANOROIDS AND OTHER LESIONS WITH AMMONIACAL

SILVER NITRATE AND FORMALIN.

By Commander P. G. White, Dental Corps, and Lieut. (Junior Grade) J. Q.

Owsley, Medical Corps, United States Navy _____ 619

SUMMARY OF SEVENTY CASES OF GONOCOCCUS INFECTION TREATED WITH

MERCUROCHROME-220 SOLUBLE TOGETHER WITH SUGAR AND FOREIGN PROTEIN.

By Lieut. Commander L. H. Williams and Lieut. W. D. Small, Medical

Corps, United States Navy------621

TREATMENT OF ENCEPHALITIS LETHARGICA.

By Harold S. Hulbert, M. D. -------------------------------624

MULTIPLE UNERUPTED AND IMPACTED CUSPID AND BICUSPID TEETH.

By Lieut. Commander M. W. Mangold, Dental Corps, United States

Navy----------------- ------------------ 625

MIXED VENEREAL INFECTIONS.

By Lieut. (Junior Grade) G. F. Cooper, Medical Corps, United States

Navy-----------------------------626

CAISSON DISEASE DURING HELMET DIVING.

By Lieut. Commander W. M. Anderson, Medical Corps, United States Navy

--- ------------------------ 628

ANTERIO-VENOUS ANEURYSM OF THE COMMON CAROTID ARTERY AND INTERNAL

JUGULAR VEIN: OPERATION WITH CONSERVATION OF THE ARTERY.

By Commander F. X. Koltes, Medical Corps, United States Navy_ 630

CREEPING ERUPTION.

By Lieut. Commander H. L. Shinn, Medical Corps, United States Navy

___________________632

DEATH BY LIGHTNING.

By Lieut. J. D. Benjamin, Medical Corps, United States Navy _634

DEVICE FOR TRANSFERRING PATIENTS FROM SHIPS.

By Master Rigger Thomas Schofield _______________________ _ 635

AN ECONOMICAL AND PRACTICAL UTENSIL STERILIZER.

By Chief Pharmacist Charles Peek, United States Navy ______ _637

THE NAVAL RESERVE-------------------------------------------- 639

NURSE CORPS:

HOSPITAL HOUSEKEEPING.

By Chief Nurse Nell I. Disert, United States Navy ________ ___ _ 649

DIETS AND THE NURSE.

By J. Beatrice Bowman, Superintendent, Navy Nurse Corps __ 651

DIETETICS AT NAVAL HOSPITALS.

By Nurse Anna F. Patten, United States Navy _______________ 655

NOTES AND COMMENTS:

Comments on " Some observations on avoidable drowning

"-Hypertension- Unrecognized syphilitic myocarditis- Tuberculosis

hospitalization- Microbic dissociation-Bacteriological nomenclature - Carbon

dioxide in hiccough-Diathermy in pneumonia-Answering examination

questions-Medical ethics-Alcoholic content of brain-Electric shock-Acid and

alkali burns-Antiseptics and the nasal flora of rabbits-Annam swelling- The

metric system-Eye test for hypersensitiveness to serum-Research on

pharmacological

problems- Syphilis not caused by vaccination-Line of duty ____ 661

BOOK NOTICES--- - - --------- - - --- ------ - - - -------- ------

--------- 697

PREVENTIVE MEDICINE, STATISTICS :

Digest of the views of medical officers regarding venereal-disease

problems as recorded in various annual sanitary reports for 1926- Two deaths

following inoculation with B. typhosus vaccine- Report

of outbreak of cereb1'ospinal fever at United States naval training

station, San Diego, Calif., December, 1926-January, 1927-Report of outbreak of

scarlet fever at United States naval training station, Hampton Roads, Va.-Fatal

poisoning by arseniuretted hydrogen in the galvanizing shop at the navy yard,

Puget Sound, Wash.-Data useful in estimating the amount of food wasted by men

eating in hospital wards and in a general mess-Outbreak of food poisoning at

United States naval training station, Hampton Roads, Va., attributed to canned

Vienna sausages-Epidemic of influenza among natives of Samoa in August, 1926-

Health of the Navy --------------------------------------------

Number 4

PREFACE--------------------------------------- --------------- VII

NOTICE TO SERVICE CONTRIBUTORS----------------------------- viii

SPECIAL ARTICLES :

CHINESE MILITARY MEDICINE.

By Lieut. (Junior Grade) J. L. McClartney, Medical Corps (Vol. G),

United States Naval Reserve-------- 783

ABDOMINAL WOUNDS AND INJURIES.

By Commander Lucius W. Johnson, Medical Corps, United States Navy

---------------------------- 816

AVIATION HYGIENE.

By Commander R. G. Davis, Medical Corps, United States Navy_ 832

GOITER.

By Commander C. E. Henry, M. D., F. A. C. P., Medical Corps (Vol. S),

United States Naval Reserve __ 837

COMMENTS CONCERNING THE DUTIES OF THE COMMANDING OFFICER OF A NAVAL

HOSPITAL.

By Capt. A. Farenholt, Medical Corps, United States Navy______ 844

THE BUDGET AND THE BUREAU OF MEDICINE AND SURGERY ACCOUNTING SYSTEM.

By Chief Pharmacist B. E. Irwin, United States Navy_________ 851

ELECTROCARDIOGRAM IN THE DIAGNOSIS OF THE CARDIAC ARRHYTHMIAS.

By Lieut. Commander R. L. Nattkemper, Medical Corps, United States Navy

------------------- 862

SIPPY TREATMENT FOR PEPTIC ULCER IN NAVAL HOSPITALS.

By Lieut. (Junior Grade) A. E. Brunschwig, Medical Corps, United States

Navy ------ ------------------ 871

By Lieut. Commander W. H. Whitemore and Lieut. (Junior Grade) O. A.

Smith, Medical Corps, United States Navy______ 875

LOCAL ANESTHESIA IN EYE, EAR, NOSE, AND THROAT WORK.

By Lieut. Commander F. L. Young, Medical Corps (Vol. G), United States

Naval Reserve-------------- 879

ENDARTERITIS, ACUTE, FROM ELECTRIC SHOCK.

By Lieut. P. F. Dickens, Medical Corps, United States Navy____ 881

ACUTE OSTEOMYELITIS WITH METASTASIS.

By Lieut. G. G. Herman, Medical Corps, United States Navy____ 883

ABDOMINAL HODGKIN'S DISEASE.

By Lieut. (Junior Grade) W. D. C. Day, Medical Corps, United States

Navy--------------------- 886

A NEEDLE FOR ANESTHESIA OF THE MAXILLARY NERVE.

By Lieut. (Junior Grade) J. Connolly, Dental Corps, United States Navy

------- --- --- ----- 889

POISONING FROM SOAP-VINE.

By Lieut. (Junior Grade) G. F . Cooper, Medical Corps, United States

Navy - - ------- ---------- 892

THE DIRECT DIAGNOSIS OF PERICARDITIS WITH EFFUSION.

By Lieut. (Junior Grade) C. G. Dyke, Medical Corps, United States

Navy--- -------- - --- --- -- 894

WHAT SHOULD THE DIAGNOSIS BE?

By Lieut. Commander C. A. Andrus, Medical Corps, United States Navy-----

---------------------- 896

MULTIPLE SCROTAL FISTULAE FOLLOWING RUPTURE OF THE BLADDER WITH

STRICTURE OF URETHRA. OPERATION AND RESTORATION OF FUNCTION.

By Lieut. Commander L. H. Williams, Medical Corps, United States

Navy------ - - ----------------897

TRAUMATIC RUPTURE OF THE SPLEEN.

By Commander F. X. Koltes, Medical Corps, United States Navy_ 898

APPENDIX ABSCESS V. PYONEPHROSIs.

By Commander G. F. Cottle, Medical Corps, United States Navy_ 900

MERCUROCHROME IN A CASE WITHOUT DIAGNOSIS.

By Lieut. (Junior Grade) B. W. Harris, Medical Corps, United States Navy--------------------------903

GREASE RACK FOR AUTOMOBILES.

By Capt. A. Farenholt, Medical Corps, United States Navy ____ 908

NAVAL RESERVE:

Hospital units- Recent appointments _____________________________ 909

NURSE CORPS:

THE LECTURES ON ACCOUNTING FROM A NURSE'S POINT OF VIEW.

By Chief Nurse Mabel T. Cooper, United States Navy _________ 913

ADDRESS TO THE CLASS OF 1926 HAITIAN GENERAL HOSPITAL TRAINING SCHOOL

FOR NURSES.

By Capt. C. St. J. Butler, Medical Corps, United States Navy __ 918

A NAVY NURSE IN THE NEAR EAST.

By Mabell S. C. Smith--------------------------- 920

DIETETICS.

By Nurse Anna P. Smith, United States Navy ________________ 921

DIETETICS.

By Nurse Mary J. Miney, United States Navy ________________ 923

NOTES AND COMMENTS :

Correction-Report on treatment of Chinese wounded-Skin tests in asthma-

Treponematosis-The physiological effects of tropical climate-History of medical

practice in the State of Illinois-Mercurochrome- Water

supplies-Seasickness-Cholera in Shanghai in 1926--Fracture of the

skull-Senescence and senility-Bismuth in the treatment of syphilis-Pressure

method of vaccination - Ventilators- Neurosyphilis-Tennis leg-Tea in treatment

of burns-European influenza epidemic at end-Visit of the Relief to Washington-Aviation

instruction for Hospital Corpsmen_ ______ 927

BOOK NOTICES----------------------- ------------------------------ 959

PREVENTIVE MEDICINE, STATISTICS :

A STUDY OF TWO THOUSAND HEALTH RECORDS CONTAINING ENTRIES FOR SYPHILIS.

By Lieut. Commander H. H. Montgomery, Medical Corps, United States Navy-------------------------------973

MEASURES OF OUTSTANDING IMPORTANCE IN THE PREVENTION AND CONTROL OF

MALARIA.

By Commander M.A. Stuart, Medical Corps, United States Navy_ 996

Outbreak of infectious diarrhea in the Battle Fleet, April, 1927, not

investigated at the time of occurrence----An epidemic of malaria at the United

States Naval Station, Olongapo, P. I.- Two deaths from rabies contracted by

playing with a dog which was incubating the disease, in Hankow, China-Frequency

of rabies-An outbreak of mumps among midshipmen at the United States Naval

Academy-Health of the Navy_________ 1010

INDEX TO SUBJECTS ----------------------------------------- 1035

INDEX TO AUTHORS-------- ------------------------------------- 1047

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Germany - Lower Saxony - Corn Harvest - 6 by Manfred Sommer

2

Maize (/meɪz/ MAYZ; Zea mays subsp. mays, from Spanish: maíz after Taino: mahiz), also known as corn (North American and Australian English), is a cereal grain first domesticated by indigenous peoples in southern Mexico about 10,000 years ago. The leafy stalk of the plant produces pollen inflorescences and separate ovuliferous inflorescences called ears that yield kernels or seeds, which are fruits.

Maize has become a staple food in many parts of the world, with the total production of maize surpassing that of wheat or rice. In addition to being consumed directly by humans (often in the form of masa), maize is also used for corn ethanol, animal feed and other maize products, such as corn starch and corn syrup. The six major types of maize are dent corn, flint corn, pod corn, popcorn, flour corn, and sweet corn. Sugar-rich varieties called sweet corn are usually grown for human consumption as kernels, while field corn varieties are used for animal feed, various corn-based human food uses (including grinding into cornmeal or masa, pressing into corn oil, and fermentation and distillation into alcoholic beverages like bourbon whiskey), and as chemical feedstocks. Maize is also used in making ethanol and other biofuels.

Maize is widely cultivated throughout the world, and a greater weight of maize is produced each year than any other grain. In 2014, total world production was 1.04 billion tonnes. Maize is the most widely grown grain crop throughout the Americas, with 361 million metric tons grown in the United States alone in 2014. Genetically modified maize made up 85% of the maize planted in the United States in 2009. Subsidies in the United States help to account for its high level of cultivation of maize and its position as the largest producer in the world.

HISTORY

PRE-COLUMBIAN DEVELOPMENT

Maize is a cultigen; human intervention is required for it to propagate. Whether or not the kernels fall off the cob on their own is a key piece of evidence used in archaeology to distinguish domesticated maize from its naturally-propagating teosinte ancestor. Genetic evidence can also be used to determine when various lineages split.

Most historians believe maize was domesticated in the Tehuacán Valley of Mexico. Recent research in the early 21st century has modified this view somewhat; scholars now indicate the adjacent Balsas River Valley of south-central Mexico as the center of domestication.

An influential 2002 study by Matsuoka et al. has demonstrated that, rather than the multiple independent domestications model, all maize arose from a single domestication in southern Mexico about 9,000 years ago. The study also demonstrated that the oldest surviving maize types are those of the Mexican highlands. Later, maize spread from this region over the Americas along two major paths. This is consistent with a model based on the archaeological record suggesting that maize diversified in the highlands of Mexico before spreading to the lowlands.

Archaeologist Dolores Piperno has said:

A large corpus of data indicates that [maize] was dispersed into lower Central America by 7600 BP [5600 BC] and had moved into the inter-Andean valleys of Colombia between 7000 and 6000 BP [5000–4000 BC].

— Dolores Piperno, The Origins of Plant Cultivation and Domestication in the New World Tropics: Patterns, Process, and New Developments

Since then, even earlier dates have been published.

According to a genetic study by Embrapa, corn cultivation was introduced in South America from Mexico, in two great waves: the first, more than 6000 years ago, spread through the Andes. Evidence of cultivation in Peru has been found dating to about 6700 years ago. The second wave, about 2000 years ago, through the lowlands of South America.

The earliest maize plants grew only small, 25-millimetre-long (1 in) corn cobs, and only one per plant. In Jackson Spielvogel's view, many centuries of artificial selection (rather than the current view that maize was exploited by interplanting with teosinte) by the indigenous people of the Americas resulted in the development of maize plants capable of growing several cobs per plant, which were usually several centimetres/inches long each. The Olmec and Maya cultivated maize in numerous varieties throughout Mesoamerica; they cooked, ground and processed it through nixtamalization. It was believed that beginning about 2500 BC, the crop spread through much of the Americas. Research of the 21st century has established even earlier dates. The region developed a trade network based on surplus and varieties of maize crops.

Mapuches of south-central Chile cultivated maize along with quinoa and potatoes in pre-Hispanic times; however, potato was the staple food of most Mapuches, "specially in the southern and coastal [Mapuche] territories where maize did not reach maturity". Before the expansion of the Inca Empire maize was traded and transported as far south as 40°19' S in Melinquina, Lácar Department. In that location maize remains were found inside pottery dated to 730 ± 80 BP and 920 ± 60 BP. Probably this maize was brought across the Andes from Chile. The presence of maize in Guaitecas Archipelago (43°55' S), the southernmost outpost of pre-Hispanic agriculture, is reported by early Spanish explorers. However the Spanish may have misidentified the plant.

COLUMBIAN EXCHANGE

After the arrival of Europeans in 1492, Spanish settlers consumed maize, and explorers and traders carried it back to Europe and introduced it to other countries. Spanish settlers far preferred wheat bread to maize, cassava, or potatoes. Maize flour could not be substituted for wheat for communion bread, since in Christian belief only wheat could undergo transubstantiation and be transformed into the body of Christ. Some Spaniards worried that by eating indigenous foods, which they did not consider nutritious, they would weaken and risk turning into Indians. "In the view of Europeans, it was the food they ate, even more than the environment in which they lived, that gave Amerindians and Spaniards both their distinctive physical characteristics and their characteristic personalities." Despite these worries, Spaniards did consume maize. Archeological evidence from Florida sites indicate they cultivated it as well.

Maize spread to the rest of the world because of its ability to grow in diverse climates. It was cultivated in Spain just a few decades after Columbus's voyages and then spread to Italy, West Africa and elsewhere. Widespread cultivation most likely began in southern Spain in 1525, after which it quickly spread to the rest of the Spanish Empire including its territories in Italy (and, from there, to other Italian states). Maize had many advantages over wheat and barley; it yielded two and a half times the food energy per unit cultivated area, could be harvested in successive years from the same plot of land, and grew in wildly varying altitudes and climates, from relatively dry regions with only 250 mm (10 in) of annual rainfall to damp regions with over 5,000 mm (200 in). By the 17th century it was a common peasant food in Southwestern Europe, including Portugal, Spain, southern France, and Italy. By the 18th century, it was the chief food of the southern French and Italian peasantry, especially in the form of polenta in Italy.

Names

The word maize derives from the Spanish form of the indigenous Taíno word for the plant, mahiz. It is known by other names around the world.

The word "corn" outside the US, Canada, Australia, and New Zealand refers to any cereal crop, its meaning understood to vary geographically to refer to the local staple. In the United States, Canada, Australia, and New Zealand, corn primarily means maize; this usage started as a shortening of "Indian corn". "Indian corn" primarily means maize (the staple grain of indigenous Americans), but can refer more specifically to multicolored "flint corn" used for decoration.

In places outside the US, Canada, Australia, and New Zealand, corn often refers to maize in culinary contexts. The narrower meaning is usually indicated by some additional word, as in sweet corn, sweetcorn, corn on the cob, baby corn, the puffed confection known as popcorn and the breakfast cereal known as corn flakes.

In Southern Africa, maize is commonly called mielie (Afrikaans) or mealie (English), words derived from the Portuguese word for maize, milho.

Maize is preferred in formal, scientific, and international usage because it refers specifically to this one grain, unlike corn, which has a complex variety of meanings that vary by context and geographic region. Maize is used by agricultural bodies and research institutes such as the FAO and CSIRO. National agricultural and industry associations often include the word maize in their name even in English-speaking countries where the local, informal word is something other than maize; for example, the Maize Association of Australia, the Indian Maize Development Association, the Kenya Maize Consortium and Maize Breeders Network, the National Maize Association of Nigeria, the Zimbabwe Seed Maize Association.

STRUCTURE AND PHYSIOLOGY

The maize plant is often 3 m (10 ft) in height, though some natural strains can grow 13 m (43 ft). The stem is commonly composed of 20 internodes of 18 cm (7 in) length. The leaves arise from the nodes, alternately on opposite sides on the stalk. A leaf, which grows from each node, is generally 9 cm (3+1⁄2 in) in width and 120 cm (3 ft 11 in) in length.

Ears develop above a few of the leaves in the midsection of the plant, between the stem and leaf sheath, elongating by around 3 mm (1⁄8 in) per day, to a length of 18 cm (7 in) with 60 cm (24 in) being the maximum alleged in the subspecies. They are female inflorescences, tightly enveloped by several layers of ear leaves commonly called husks. Certain varieties of maize have been bred to produce many additional developed ears. These are the source of the "baby corn" used as a vegetable in Asian cuisine.

The apex of the stem ends in the tassel, an inflorescence of male flowers. When the tassel is mature and conditions are suitably warm and dry, anthers on the tassel dehisce and release pollen. Maize pollen is anemophilous (dispersed by wind), and because of its large settling velocity, most pollen falls within a few meters of the tassel.

Elongated stigmas, called silks, emerge from the whorl of husk leaves at the end of the ear. They are often pale yellow and 18 cm (7 in) in length, like tufts of hair in appearance. At the end of each is a carpel, which may develop into a "kernel" if fertilized by a pollen grain. The pericarp of the fruit is fused with the seed coat referred to as "caryopsis", typical of the grasses, and the entire kernel is often referred to as the "seed". The cob is close to a multiple fruit in structure, except that the individual fruits (the kernels) never fuse into a single mass. The grains are about the size of peas, and adhere in regular rows around a white, pithy substance, which forms the ear. The maximum size of kernels is reputedly 2.5 cm (1 in). An ear commonly holds 600 kernels. They are of various colors: blackish, bluish-gray, purple, green, red, white and yellow. When ground into flour, maize yields more flour with much less bran than wheat does. It lacks the protein gluten of wheat and, therefore, makes baked goods with poor rising capability. A genetic variant that accumulates more sugar and less starch in the ear is consumed as a vegetable and is called sweet corn. Young ears can be consumed raw, with the cob and silk, but as the plant matures (usually during the summer months), the cob becomes tougher and the silk dries to inedibility. By the end of the growing season, the kernels dry out and become difficult to chew without cooking them tender first in boiling water.

Planting density affects multiple aspects of maize. Modern farming techniques in developed countries usually rely on dense planting, which produces one ear per stalk. Stands of silage maize are yet denser,[citation needed] and achieve a lower percentage of ears and more plant matter.

Maize is a facultative short-day plant and flowers in a certain number of growing degree days > 10 °C (50 °F) in the environment to which it is adapted. The magnitude of the influence that long nights have on the number of days that must pass before maize flowers is genetically prescribed and regulated by the phytochrome system.

Photoperiodicity can be eccentric in tropical cultivars such that the long days characteristic of higher latitudes allow the plants to grow so tall that they do not have enough time to produce seed before being killed by frost. These attributes, however, may prove useful in using tropical maize for biofuels.

Immature maize shoots accumulate a powerful antibiotic substance, 2,4-dihydroxy-7-methoxy-1,4-benzoxazin-3-one (DIMBOA). DIMBOA is a member of a group of hydroxamic acids (also known as benzoxazinoids) that serve as a natural defense against a wide range of pests, including insects, pathogenic fungi and bacteria. DIMBOA is also found in related grasses, particularly wheat. A maize mutant (bx) lacking DIMBOA is highly susceptible to attack by aphids and fungi. DIMBOA is also responsible for the relative resistance of immature maize to the European corn borer (family Crambidae). As maize matures, DIMBOA levels and resistance to the corn borer decline.

Because of its shallow roots, maize is susceptible to droughts, intolerant of nutrient-deficient soils, and prone to be uprooted by severe winds.

While yellow maizes derive their color from lutein and zeaxanthin, in red-colored maizes, the kernel coloration is due to anthocyanins and phlobaphenes. These latter substances are synthesized in the flavonoids synthetic pathway from polymerization of flavan-4-ols by the expression of maize pericarp color1 (p1) gene which encodes an R2R3 myb-like transcriptional activator of the A1 gene encoding for the dihydroflavonol 4-reductase (reducing dihydroflavonols into flavan-4-ols) while another gene (Suppressor of Pericarp Pigmentation 1 or SPP1) acts as a suppressor. The p1 gene encodes an Myb-homologous transcriptional activator of genes required for biosynthesis of red phlobaphene pigments, while the P1-wr allele specifies colorless kernel pericarp and red cobs, and unstable factor for orange1 (Ufo1) modifies P1-wr expression to confer pigmentation in kernel pericarp, as well as vegetative tissues, which normally do not accumulate significant amounts of phlobaphene pigments. The maize P gene encodes a Myb homolog that recognizes the sequence CCT/AACC, in sharp contrast with the C/TAACGG bound by vertebrate Myb proteins.

The ear leaf is the leaf most closely associated with a particular developing ear. This leaf and above contribute 70% to 75% to 90% of grain fill. Therefore fungicide application is most important in that region in most disease environments.

ABNORMAL FLOWERS

Maize flowers may sometimes exhibit mutations that lead to the formation of female flowers in the tassel. These mutations, ts4 and Ts6, prohibit the development of the stamen while simultaneously promoting pistil development. This may cause inflorescences containing both male and female flowers, or hermaphrodite flowers.

GENETICS

Maize is an annual grass in the family Gramineae, which includes such plants as wheat, rye, barley, rice, sorghum, and sugarcane. There are two major species of the genus Zea (out of six total): Zea mays (maize) and Zea diploperennis, which is a perennial type of teosinte. The annual teosinte variety called Zea mays mexicana is the closest botanical relative to maize. It still grows in the wild as an annual in Mexico and Guatemala.

Many forms of maize are used for food, sometimes classified as various subspecies related to the amount of starch each has:

Flour corn: Zea mays var. amylacea

Popcorn: Zea mays var. everta

Dent corn : Zea mays var. indentata

Flint corn: Zea mays var. indurata

Sweet corn: Zea mays var. saccharata and Zea mays var. rugosa

Waxy corn: Zea mays var. ceratina

Amylomaize: Zea mays

Pod corn: Zea mays var. tunicata Larrañaga ex A. St. Hil.

Striped maize: Zea mays var. japonica

This system has been replaced (though not entirely displaced) over the last 60 years by multivariable classifications based on ever more data. Agronomic data were supplemented by botanical traits for a robust initial classification, then genetic, cytological, protein and DNA evidence was added. Now, the categories are forms (little used), races, racial complexes, and recently branches.

Maize is a diploid with 20 chromosomes (n=10). The combined length of the chromosomes is 1500 cM. Some of the maize chromosomes have what are known as "chromosomal knobs": highly repetitive heterochromatic domains that stain darkly. Individual knobs are polymorphic among strains of both maize and teosinte.

Barbara McClintock used these knob markers to validate her transposon theory of "jumping genes", for which she won the 1983 Nobel Prize in Physiology or Medicine. Maize is still an important model organism for genetics and developmental biology today.

The centromeres have two types of structural components, both of which are found only in the centromeres: Large arrays of CentC, a short satellite DNA; and a few of a family of retrotransposons. The B chromosome, unlike the others, contains an additional repeat which extends into neighboring areas of the chromosome. Centromeres can accidentally shrink during division and still function, although it is thought this will fail if it shrinks below a few hundred kilobase. Kinetochores contain RNA originating from centromeres. Centromere regions can become inactive, and can continue in that state if the chromosome still has another active one.

The Maize Genetics Cooperation Stock Center, funded by the USDA Agricultural Research Service and located in the Department of Crop Sciences at the University of Illinois at Urbana-Champaign, is a stock center of maize mutants. The total collection has nearly 80,000 samples. The bulk of the collection consists of several hundred named genes, plus additional gene combinations and other heritable variants. There are about 1000 chromosomal aberrations (e.g., translocations and inversions) and stocks with abnormal chromosome numbers (e.g., tetraploids). Genetic data describing the maize mutant stocks as well as myriad other data about maize genetics can be accessed at MaizeGDB, the Maize Genetics and Genomics Database.

In 2005, the US National Science Foundation (NSF), Department of Agriculture (USDA) and the Department of Energy (DOE) formed a consortium to sequence the B73 maize genome. The resulting DNA sequence data was deposited immediately into GenBank, a public repository for genome-sequence data. Sequences and genome annotations have also been made available throughout the project's lifetime at the project's official site.

Primary sequencing of the maize genome was completed in 2008. On November 20, 2009, the consortium published results of its sequencing effort in Science. The genome, 85% of which is composed of transposons, was found to contain 32,540 genes (By comparison, the human genome contains about 2.9 billion bases and 26,000 genes). Much of the maize genome has been duplicated and reshuffled by helitrons—group of rolling circle transposons.

In Z. mays and various other angiosperms the MADS-box motif is involved in floral development. Early study in several angiosperm models including Z. mays was the beginning of research into the molecular evolution of floral structure in general, as well as their role in nonflowering plants.

EVOLUTION

As with many plants and animals, Z. mays has a positive correlation between effective population size and the magnitude of selection pressure. Z. m. having an EPS of ~650,000, it clusters with others of about the same EPS, and has 79% of its amino acid sites under selection.

Recombination is a significant source of diversity in Z. mays. (Note that this finding supersedes previous studies which showed no such correlation.)

This recombination/diversity effect is seen throughout plants but is also found to not occur – or not as strongly – in regions of high gene density. This is likely the reason that domesticated Z. mays has not seen as much of an increase in diversity within areas of higher density as in regions of lower density, although there is more evidence in other plants.

Some lines of maize have undergone ancient polyploidy events, starting 11m years ago. Over that time ~72% of polyploid duplicated genes have been retained, which is higher than other plants with older polyploidy events. Thus maize may be due to lose more duplicate genes as time goes along, similar to the course followed by the genomes of other plants. If so - if gene loss has merely not occurred yet - that could explain the lack of observed positive selection and lower negative selection which are observed in otherwise similar plants, i.e. also naturally outcrossing and with similar effective population sizes.

Ploidy does not appear to influence EPS or magnitude of selection effect in maize.

BREEDING

Maize reproduces sexually each year. This randomly selects half the genes from a given plant to propagate to the next generation, meaning that desirable traits found in the crop (like high yield or good nutrition) can be lost in subsequent generations unless certain techniques are used.

Maize breeding in prehistory resulted in large plants producing large ears. Modern breeding began with individuals who selected highly productive varieties in their fields and then sold seed to other farmers. James L. Reid was one of the earliest and most successful developing Reid's Yellow Dent in the 1860s. These early efforts were based on mass selection. Later breeding efforts included ear to row selection (C. G. Hopkins c. 1896), hybrids made from selected inbred lines (G. H. Shull, 1909), and the highly successful double cross hybrids using four inbred lines (D. F. Jones c. 1918, 1922). University supported breeding programs were especially important in developing and introducing modern hybrids. By the 1930s, companies such as Pioneer devoted to production of hybrid maize had begun to influence long-term development. Internationally important seed banks such as the International Maize and Wheat Improvement Center (CIMMYT) and the US bank at the Maize Genetics Cooperation Stock Center University of Illinois at Urbana-Champaign maintain germplasm important for future crop development.

Since the 1940s the best strains of maize have been first-generation hybrids made from inbred strains that have been optimized for specific traits, such as yield, nutrition, drought, pest and disease tolerance. Both conventional cross-breeding and genetic modification have succeeded in increasing output and reducing the need for cropland, pesticides, water and fertilizer. There is conflicting evidence to support the hypothesis that maize yield potential has increased over the past few decades. This suggests that changes in yield potential are associated with leaf angle, lodging resistance, tolerance of high plant density, disease/pest tolerance, and other agronomic traits rather than increase of yield potential per individual plant.

Tropical landraces remain an important and underutilized source of resistance alleles for for disease and for herbivores. Notable discoveries of rare alleles for this purpose were made by Dao et al 2014 and Sood et al 2014.

GLOBAL PROGRAM

CIMMYT operates a conventional breeding program to provide optimized strains. The program began in the 1980s. Hybrid seeds are distributed in Africa by the Drought Tolerant Maize for Africa project.

GENETIC MODIFICATION

Genetically modified (GM) maize was one of the 26 GM crops grown commercially in 2016. The vast majority of this is Bt maize. Grown since 1997 in the United States and Canada, 92% of the US maize crop was genetically modified in 2016 and 33% of the worldwide maize crop was GM in 2016. As of 2011, Herbicide-tolerant maize varieties were grown in Argentina, Australia, Brazil, Canada, China, Colombia, El Salvador, the European Union, Honduras, Japan, Korea, Malaysia, Mexico, New Zealand, Philippines, the Russian Federation, Singapore, South Africa, Taiwan, Thailand, and the United States. Insect-resistant maize was grown in Argentina, Australia, Brazil, Canada, Chile, China, Colombia, Egypt, the European Union, Honduras, Japan, Korea, Malaysia, Mexico, New Zealand, Philippines, South Africa, Switzerland, Taiwan, the United States, and Uruguay.

In September 2000, up to $50 million worth of food products were recalled due to the presence of Starlink genetically modified corn, which had been approved only for animal consumption and had not been approved for human consumption, and was subsequently withdrawn from the market.

ORIGIN

Maize is the domesticated variant of teosinte. The two plants have dissimilar appearance, maize having a single tall stalk with multiple leaves and teosinte being a short, bushy plant. The difference between the two is largely controlled by differences in just two genes, called grassy tillers-1 (gt1, A0A317YEZ1) and teosinte branched-1 (tb1, Q93WI2).

Several theories had been proposed about the specific origin of maize in Mesoamerica:

It is a direct domestication of a Mexican annual teosinte, Zea mays ssp. parviglumis, native to the Balsas River valley in south-eastern Mexico, with up to 12% of its genetic material obtained from Zea mays ssp. mexicana through introgression.

It has been derived from hybridization between a small domesticated maize (a slightly changed form of a wild maize) and a teosinte of section Luxuriantes, either Z. luxurians or Z. diploperennis.

It has undergone two or more domestications either of a wild maize or of a teosinte. (The term "teosinte" describes all species and subspecies in the genus Zea, excluding Zea mays ssp. mays.)

It has evolved from a hybridization of Z. diploperennis by Tripsacum dactyloides.

In the late 1930s, Paul Mangelsdorf suggested that domesticated maize was the result of a hybridization event between an unknown wild maize and a species of Tripsacum, a related genus. This theory about the origin of maize has been refuted by modern genetic testing, which refutes Mangelsdorf's model and the fourth listed above.

The teosinte origin theory was proposed by the Russian botanist Nikolai Ivanovich Vavilov in 1931 and the later American Nobel Prize-winner George Beadle in 1932.: 10 It is supported experimentally and by recent studies of the plants' genomes. Teosinte and maize can cross-breed and produce fertile offspring. A number of questions remain concerning the species, among them:

how the immense diversity of the species of sect. Zea originated,

how the tiny archaeological specimens of 3500–2700 BC could have been selected from a teosinte, and

how domestication could have proceeded without leaving remains of teosinte or maize with teosintoid traits earlier than the earliest known until recently, dating from ca. 1100 BC.

The domestication of maize is of particular interest to researchers—archaeologists, geneticists, ethnobotanists, geographers, etc. The process is thought by some to have started 7,500 to 12,000 years ago. Research from the 1950s to 1970s originally focused on the hypothesis that maize domestication occurred in the highlands between the states of Oaxaca and Jalisco, because the oldest archaeological remains of maize known at the time were found there.

Connection with 'parviglumis' subspecies

Genetic studies, published in 2004 by John Doebley, identified Zea mays ssp. parviglumis, native to the Balsas River valley in Mexico's southwestern highlands, and also known as Balsas teosinte, as being the crop wild relative that is genetically most similar to modern maize. This was confirmed by further studies, which refined this hypothesis somewhat. Archaeobotanical studies, published in 2009, point to the middle part of the Balsas River valley as the likely location of early domestication; this river is not very long, so these locations are not very distant. Stone milling tools with maize residue have been found in an 8,700 year old layer of deposits in a cave not far from Iguala, Guerrero.

Doebley was part of the team that first published, in 2002, that maize had been domesticated only once, about 9,000 years ago, and then spread throughout the Americas.

A primitive corn was being grown in southern Mexico, Central America, and northern South America 7,000 years ago. Archaeological remains of early maize ears, found at Guila Naquitz Cave in the Oaxaca Valley, date back roughly 6,250 years; the oldest ears from caves near Tehuacan, Puebla, 5,450 B.P.

Maize pollen dated to 7,300 B.P. from San Andres, Tabasco, on the Caribbean coast has also been recovered.

As maize was introduced to new cultures, new uses were developed and new varieties selected to better serve in those preparations. Maize was the staple food, or a major staple – along with squash, Andean region potato, quinoa, beans, and amaranth – of most pre-Columbian North American, Mesoamerican, South American, and Caribbean cultures. The Mesoamerican civilization, in particular, was deeply interrelated with maize. Its traditions and rituals involved all aspects of maize cultivation – from the planting to the food preparation. Maize formed the Mesoamerican people's identity.

It is unknown what precipitated its domestication, because the edible portion of the wild variety is too small, and hard to obtain, to be eaten directly, as each kernel is enclosed in a very hard bivalve shell.

In 1939, George Beadle demonstrated that the kernels of teosinte are readily "popped" for human consumption, like modern popcorn. Some have argued it would have taken too many generations of selective breeding to produce large, compressed ears for efficient cultivation. However, studies of the hybrids readily made by intercrossing teosinte and modern maize suggest this objection is not well founded.

SPREADING TO THE NORTH

Around 4,500 ago, maize began to spread to the north; it was first cultivated in what is now the United States at several sites in New Mexico and Arizona, about 4,100 ago.

During the first millennium AD, maize cultivation spread more widely in the areas north. In particular, the large-scale adoption of maize agriculture and consumption in eastern North America took place about A.D. 900. Native Americans cleared large forest and grassland areas for the new crop.

In 2005, research by the USDA Forest Service suggested that the rise in maize cultivation 500 to 1,000 years ago in what is now the southeastern United States corresponded with a decline of freshwater mussels, which are very sensitive to environmental changes.

CULTIVATION

PLANTING

Because it is cold-intolerant, in the temperate zones maize must be planted in the spring. Its root system is generally shallow, so the plant is dependent on soil moisture. As a plant that uses C4 carbon fixation, maize is a considerably more water-efficient crop than plants that use C3 carbon fixation such as alfalfa and soybeans. Maize is most sensitive to drought at the time of silk emergence, when the flowers are ready for pollination. In the United States, a good harvest was traditionally predicted if the maize was "knee-high by the Fourth of July", although modern hybrids generally exceed this growth rate. Maize used for silage is harvested while the plant is green and the fruit immature. Sweet corn is harvested in the "milk stage", after pollination but before starch has formed, between late summer and early to mid-autumn. Field maize is left in the field until very late in the autumn to thoroughly dry the grain, and may, in fact, sometimes not be harvested until winter or even early spring. The importance of sufficient soil moisture is shown in many parts of Africa, where periodic drought regularly causes maize crop failure and consequent famine. Although it is grown mainly in wet, hot climates, it has been said to thrive in cold, hot, dry or wet conditions, meaning that it is an extremely versatile crop.

Maize was planted by the Native Americans in hills, in a complex system known to some as the Three Sisters. Maize provided support for beans, and the beans provided nitrogen derived from nitrogen-fixing rhizobia bacteria which live on the roots of beans and other legumes; and squashes provided ground cover to stop weeds and inhibit evaporation by providing shade over the soil. This method was replaced by single species hill planting where each hill 60–120 cm (2 ft 0 in–3 ft 11 in) apart was planted with three or four seeds, a method still used by home gardeners. A later technique was "checked maize", where hills were placed

1 m (40 in) apart in each direction, allowing cultivators to run through the field in two directions. In more arid lands, this was altered and seeds were planted in the bottom of 10–12 cm (4–4+1⁄2 in) deep furrows to collect water. Modern technique plants maize in rows which allows for cultivation while the plant is young, although the hill technique is still used in the maize fields of some Native American reservations. When maize is planted in rows, it also allows for planting of other crops between these rows to make more efficient use of land space.

In most regions today, maize grown in residential gardens is still often planted manually with a hoe, whereas maize grown commercially is no longer planted manually but rather is planted with a planter. In North America, fields are often planted in a two-crop rotation with a nitrogen-fixing crop, often alfalfa in cooler climates and soybeans in regions with longer summers. Sometimes a third crop, winter wheat, is added to the rotation.

Many of the maize varieties grown in the United States and Canada are hybrids. Often the varieties have been genetically modified to tolerate glyphosate or to provide protection against natural pests. Glyphosate is an herbicide which kills all plants except those with genetic tolerance. This genetic tolerance is very rarely found in nature.

In the midwestern United States, low-till or no-till farming techniques are usually used. In low-till, fields are covered once, maybe twice, with a tillage implement either ahead of crop planting or after the previous harvest. The fields are planted and fertilized. Weeds are controlled through the use of herbicides, and no cultivation tillage is done during the growing season. This technique reduces moisture evaporation from the soil, and thus provides more moisture for the crop. The technologies mentioned in the previous paragraph enable low-till and no-till farming. Weeds compete with the crop for moisture and nutrients, making them undesirable.

HARVESTING

Before the 20th century, all maize harvesting was by manual labour, by grazing, or by some combination of those. Whether the ears were hand-picked and the stover was grazed, or the whole plant was cut, gathered, and shocked, people and livestock did all the work. Between the 1890s and the 1970s, the technology of maize harvesting expanded greatly. Today, all such technologies, from entirely manual harvesting to entirely mechanized, are still in use to some degree, as appropriate to each farm's needs, although the thoroughly mechanized versions predominate, as they offer the lowest unit costs when scaled to large farm operations. For small farms, their unit cost can be too high, as their higher fixed cost cannot be amortized over as many units.[citation needed]

Before World War II, most maize in North America was harvested by hand. This involved a large number of workers and associated social events (husking or shucking bees). From the 1890s onward, some machinery became available to partially mechanize the processes, such as one- and two-row mechanical pickers (picking the ear, leaving the stover) and corn binders, which are reaper-binders designed specifically for maize (for example, Video on YouTube). The latter produce sheaves that can be shocked. By hand or mechanical picker, the entire ear is harvested, which then requires a separate operation of a maize sheller to remove the kernels from the ear. Whole ears of maize were often stored in corn cribs, and these whole ears are a sufficient form for some livestock feeding use. Today corn cribs with whole ears, and corn binders, are less common because most modern farms harvest the grain from the field with a combine and store it in bins. The combine with a corn head (with points and snap rolls instead of a reel) does not cut the stalk; it simply pulls the stalk down. The stalk continues downward and is crumpled into a mangled pile on the ground, where it usually is left to become organic matter for the soil. The ear of maize is too large to pass between slots in a plate as the snap rolls pull the stalk away, leaving only the ear and husk to enter the machinery. The combine separates the husk and the cob, keeping only the kernels.

When maize is a silage crop, the entire plant is usually chopped at once with a forage harvester (chopper) and ensiled in silos or polymer wrappers. Ensiling of sheaves cut by a corn binder was formerly common in some regions but has become uncommon. For storing grain in bins, the moisture of the grain must be sufficiently low to avoid spoiling. If the moisture content of the harvested grain is too high, grain dryers are used to reduce the moisture content by blowing heated air through the grain. This can require large amounts of energy in the form of combustible gases (propane or natural gas) and electricity to power the blowers.

PRODUCTION

Maize is widely cultivated throughout the world, and a greater weight of maize is produced each year than any other grain. In 2018, total world production was 1.15 billion tonnes, led by the United States with 34.2% of the total (table). China produced 22.4% of the global total.

UNITED STATES

In 2016, maize production was forecast to be over 380 million metric tons (15 billion bushels), an increase of 11% over 2014 American production. Based on conditions as of August 2016, the expected yield would be the highest ever for the United States. The area of harvested maize was forecast to be 35 million hectares (87 million acres), an increase of 7% over 2015. Maize is especially popular in Midwestern states such as Indiana, Iowa, and Illinois; in the latter, it was named the state's official grain in 2017.

STORAGE

Drying is vital to prevent or at least reduce mycotoxin contamination. Aspergillus and Fusarium spp. are the most common mycotoxin sources, but there are others. Altogether maize contaminants are so common, and this crop is so economically important, that maize mycotoxins are among the most important in agriculture in general.

USES

HUMAN FOOD

Maize and cornmeal (ground dried maize) constitute a staple food in many regions of the world. Maize is used to produce cornstarch, a common ingredient in home cooking and many industrialized food products. Maize starch can be hydrolyzed and enzymatically treated to produce syrups, particularly high fructose corn syrup, a sweetener; and also fermented and distilled to produce grain alcohol. Grain alcohol from maize is traditionally the source of Bourbon whiskey. Corn flour is used to make cornbread and other baked products.

In prehistoric times Mesoamerican women used a metate to process maize into ground cornmeal, allowing the preparation of foods that were more calorie dense than popcorn. After ceramic vessels were invented the Olmec people began to cook maize together with beans, improving the nutritional value of the staple meal. Although maize naturally contains niacin, an important nutrient, it was not bioavailable without the process of nixtamalization. The Maya used nixtamal meal to make varieties of porridges and tamales. The process was later used in the cuisine of the American South to prepare corn for grits and hominy.

Maize is a staple of Mexican cuisine. Masa (cornmeal treated with limewater) is the main ingredient for tortillas, atole and many other dishes of Central American food. It is the main ingredient of corn tortilla, tamales, pozole, atole and all the dishes based on them, like tacos, quesadillas, chilaquiles, enchiladas, tostadas and many more. In Mexico the fungus of maize, known as huitlacoche, is considered a delicacy.

Coarse maize meal is made into a thick porridge in many cultures: from the polenta of Italy, the angu of Brazil, the mămăligă of Romania, to cornmeal mush in the US (or hominy grits in the South) or the food called mieliepap in South Africa and sadza, nshima, ugali and other names in other parts of Africa. Introduced into Africa by the Portuguese in the 16th century, maize has become Africa's most important staple food crop. These are commonly eaten in the Southeastern United States, foods handed down from Native Americans, who called the dish sagamite.

Maize can also be harvested and consumed in the unripe state, when the kernels are fully grown but still soft. Unripe maize must usually be cooked to become palatable; this may be done by simply boiling or roasting the whole ears and eating the kernels right off the cob. Sweet corn, a genetic variety that is high in sugars and low in starch, is usually consumed in the unripe state. Such corn on the cob is a common dish in the United States, Canada, United Kingdom, Cyprus, some parts of South America, and the Balkans, but virtually unheard of in some European countries. Corn on the cob was hawked on the streets of early 19th-century New York City by poor, barefoot "Hot Corn Girls", who were thus the precursors of hot dog carts, churro wagons, and fruit stands seen on the streets of big cities today.

Within the United States, the usage of maize for human consumption constitutes only around 1/40th of the amount grown in the country. In the United States and Canada, maize is mostly grown to feed livestock, as forage, silage (made by fermentation of chopped green cornstalks), or grain. Maize meal is also a significant ingredient of some commercial animal food products.

NUTRITIONAL VALUE

Raw, yellow, sweet maize kernels are composed of 76% water, 19% carbohydrates, 3% protein, and 1% fat (table). In a 100-gram serving, maize kernels provide 86 calories and are a good source (10–19% of the Daily Value) of the B vitamins, thiamin, niacin (but see Pellagra warning below), pantothenic acid (B5) and folate (right table for raw, uncooked kernels, USDA Nutrient Database). In moderate amounts, they also supply dietary fiber and the essential minerals, magnesium and phosphorus whereas other nutrients are in low amounts (table).

Maize has suboptimal amounts of the essential amino acids tryptophan and lysine, which accounts for its lower status as a protein source. However, the proteins of beans and legumes complement those of maize.

FEED AND FODDER FOR LIVESTOCK

Maize is a major source of both grain feed and fodder for livestock. It is fed to the livestock in various ways. When it is used as a grain crop, the dried kernels are used as feed. They are often kept on the cob for storage in a corn crib, or they may be shelled off for storage in a grain bin. The farm that consumes the feed may produce it, purchase it on the market, or some of both. When the grain is used for feed, the rest of the plant (the corn stover) can be used later as fodder, bedding (litter), or soil amendment. When the whole maize plant (grain plus stalks and leaves) is used for fodder, it is usually chopped all at once and ensilaged, as digestibility and palatability are higher in the ensilaged form than in the dried form. Maize silage is one of the most valuable forages for ruminants. Before the advent of widespread ensilaging, it was traditional to gather the corn into shocks after harvesting, where it dried further. With or without a subsequent move to the cover of a barn, it was then stored for weeks to several months until fed to the livestock. Today ensilaging can occur not only in siloes but also in silage wrappers. However, in the tropics, maize can be harvested year-round and fed as green forage to the animals.

CHEMICALS

Starch from maize can also be made into plastics, fabrics, adhesives, and many other chemical products.

The corn steep liquor, a plentiful watery byproduct of maize wet milling process, is widely used in the biochemical industry and research as a culture medium to grow many kinds of microorganisms.

Chrysanthemin is found in purple corn and is used as a food coloring.

BIO-FUEL

"Feed maize" is being used increasingly for heating; specialized corn stoves (similar to wood stoves) are available and use either feed maize or wood pellets to generate heat. Maize cobs are also used as a biomass fuel source. Maize is relatively cheap and home-heating furnaces have been developed which use maize kernels as a fuel. They feature a large hopper that feeds the uniformly sized maize kernels (or wood pellets or cherry pits) into the fire.[citation needed]

Maize is increasingly used as a feedstock for the production of ethanol fuel. When considering where to construct an ethanol plant, one of the site selection criteria is to ensure there is locally available feedstock. Ethanol is mixed with gasoline to decrease the amount of pollutants emitted when used to fuel motor vehicles. High fuel prices in mid-2007 led to higher demand for ethanol, which in turn led to higher prices paid to farmers for maize. This led to the 2007 harvest being one of the most profitable maize crops in modern history for farmers. Because of the relationship between fuel and maize, prices paid for the crop now tend to track the price of oil.

The price of food is affected to a certain degree by the use of maize for biofuel production. The cost of transportation, production, and marketing are a large portion (80%) of the price of food in the United States. Higher energy costs affect these costs, especially transportation. The increase in food prices the consumer has been seeing is mainly due to the higher energy cost. The effect of biofuel production on other food crop prices is indirect. Use of maize for biofuel production increases the demand, and therefore price of maize. This, in turn, results in farm acreage being diverted from other food crops to maize production. This reduces the supply of the other food crops and increases their prices.

Maize is widely used in Germany as a feedstock for biogas plants. Here the maize is harvested, shredded then placed in silage clamps from which it is fed into the biogas plants. This process makes use of the whole plant rather than simply using the kernels as in the production of fuel ethanol.

A biomass gasification power plant in Strem near Güssing, Burgenland, Austria, began in 2005. Research is being done to make diesel out of the biogas by the Fischer Tropsch method.

Increasingly, ethanol is being used at low concentrations (10% or less) as an additive in gasoline (gasohol) for motor fuels to increase the octane rating, lower pollutants, and reduce petroleum use (what is nowadays also known as "biofuels" and has been generating an intense debate regarding the human beings' necessity of new sources of energy, on the one hand, and the need to maintain, in regions such as Latin America, the food habits and culture which has been the essence of civilizations such as the one originated in Mesoamerica; the entry, January 2008, of maize among the commercial agreements of NAFTA has increased this debate, considering the bad labor conditions of workers in the fields, and mainly the fact that NAFTA "opened the doors to the import of maize from the United States, where the farmers who grow it receive multimillion-dollar subsidies and other government supports. ... According to OXFAM UK, after NAFTA went into effect, the price of maize in Mexico fell 70% between 1994 and 2001. The number of farm jobs dropped as well: from 8.1 million in 1993 to 6.8 million in 2002. Many of those who found themselves without work were small-scale maize growers."). However, introduction in the northern latitudes of the US of tropical maize for biofuels, and not for human or animal consumption, may potentially alleviate this.

COMMODITY

Maize is bought and sold by investors and price speculators as a tradable commodity using corn futures contracts. These "futures" are traded on the Chicago Board of Trade (CBOT) under ticker symbol C. They are delivered every year in March, May, July, September, and December.

ORNAMENTAL AND OTHER USES

Some forms of the plant are occasionally grown for ornamental use in the garden. For this purpose, variegated and colored leaf forms as well as those with colorful ears are used.

Corncobs can be hollowed out and treated to make inexpensive smoking pipes, first manufactured in the United States in 1869.

An unusual use for maize is to create a "corn maze" (or "maize maze") as a tourist attraction. The idea of a maize maze was introduced by the American Maze Company who created a maze in Pennsylvania in 1993. Traditional mazes are most commonly grown using yew hedges, but these take several years to mature. The rapid growth of a field of maize allows a maze to be laid out using GPS at the start of a growing season and for the maize to grow tall enough to obstruct a visitor's line of sight by the start of the summer. In Canada and the US, these are popular in many farming communities.

Maize kernels can be used in place of sand in a sandboxlike enclosure for children's play.

Stigmas from female maize flowers, popularly called corn silk, are sold as herbal supplements.

Maize is used as a fish bait, called "dough balls". It is particularly popular in Europe for coarse fishing.

Additionally, feed corn is sometimes used by hunters to bait animals such as deer or wild hogs.

UNITED STATES USAGE BREAKDOWN

The breakdown of usage of the 12.1-billion-bushel (307-million-tonne) 2008 US maize crop was as follows, according to the World Agricultural Supply and Demand Estimates Report by the USDA.In the US since 2009/2010, maize feedstock use for ethanol production has somewhat exceeded direct use for livestock feed; maize use for fuel ethanol was 5,130 million bushels (130 million tonnes) in the 2013/2014 marketing year.A fraction of the maize feedstock dry matter used for ethanol production is usefully recovered as DDGS (dried distillers grains with solubles). In the 2010/2011 marketing year, about 29.1 million tonnes of DDGS were fed to US livestock and poultry. Because starch utilization in fermentation for ethanol production leaves other grain constituents more concentrated in the residue, the feed value per kg of DDGS, with regard to ruminant-metabolizable energy and protein, exceeds that of the grain. Feed value for monogastric animals, such as swine and poultry, is somewhat lower than for ruminants.

HAZARDS

PELLAGRA

When maize was first introduced into farming systems other than those used by traditional native-American peoples, it was generally welcomed with enthusiasm for its productivity. However, a widespread problem of malnutrition soon arose wherever maize was introduced as a staple food. This was a mystery, since these types of malnutrition were not normally seen among the indigenous Americans, for whom maize was the principal staple food.

It was eventually discovered that the indigenous Americans had learned to soak maize in alkali — water (the process now known as nixtamalization) — made with ashes and lime (calcium oxide) since at least 1200–1500 BC by Mesoamericans. They did this to liberate the corn hulls, but (unbeknownst to natives or colonists) it coincidentally liberates the B-vitamin niacin, the lack of which was the underlying cause of the condition known as pellagra.

Maize was introduced into the diet of non-indigenous Americans without the necessary cultural knowledge acquired over thousands of years in the Americas. In the late 19th century, pellagra reached epidemic proportions in parts of the southern US, as medical researchers debated two theories for its origin: the deficiency theory (which was eventually shown to be true) said that pellagra was due to a deficiency of some nutrient, and the germ theory said that pellagra was caused by a germ transmitted by stable flies. A third theory, promoted by the eugenicist Charles Davenport, held that people only contracted pellagra if they were susceptible to it due to certain "constitutional, inheritable" traits of the affected individual.

Once alkali processing and dietary variety were understood and applied, pellagra disappeared in the developed world. The development of high lysine maize and the promotion of a more balanced diet have also contributed to its demise. Pellagra still exists today in food-poor areas and refugee camps where people survive on donated maize.

ALLERGY

Maize contains lipid transfer protein, an indigestible protein that survives cooking. This protein has been linked to a rare and understudied allergy to maize in humans. The allergic reaction can cause skin rash, swelling or itching of mucous membranes, diarrhea, vomiting, asthma and, in severe cases, anaphylaxis. It is unclear how common this allergy is in the general population.

MYCOTOXINS

Fungicide application does not reduce fungal growth or mycotoxin dramatically, although it can be a part of a successful reduction strategy. Among the most common toxins are those produced by Aspergillus and Fusarium spp. The most common toxins are aflatoxins, fumonisins, zearalenone, and ochratoxin A. Bt maize discourages insect vectors and by so doing it dramatically reduces concentrations of fumonisins, significantly reduces aflatoxins, but only mildly reduces others.

ART

Maize has been an essential crop in the Andes since the pre-Columbian era. The Moche culture from Northern Peru made ceramics from earth, water, and fire. This pottery was a sacred substance, formed in significant shapes and used to represent important themes. Maize was represented anthropomorphically as well as naturally.

In the United States, maize ears along with tobacco leaves are carved into the capitals of columns in the United States Capitol building. Maize itself is sometimes used for temporary architectural detailing when the intent is to celebrate the fall season, local agricultural productivity and culture. Bundles of dried maize stalks are often displayed along with pumpkins, gourds and straw in autumnal displays outside homes and businesses. A well-known example of architectural use is the Corn Palace in Mitchell, South Dakota, which uses cobs and ears of colored maize to implement a mural design that is recycled annually. Another well-known example is the Field of Corn sculpture in Dublin, Ohio, where hundreds of concrete ears of corn stand in a grassy field.

A maize stalk with two ripe ears is depicted on the reverse of the Croatian 1 lipa coin, minted since 1993.

WIKIPEDIA

~Tis The Season For Stacking Springtails~ by Eddie The Bugman

30 47

A male Dicyrtomina saundersi today taken at x7 magnification, a handheld focus stack of 10 images at F/5.6. I'm surprised this worked out as it was so windy, I was expecting the antennae to be all over the shop, this one is a male, the dark cheek pigmentation on the head and darker pigmentation in general points to this. This one was about 2mm in size at a guess :o)

I really must try to find some different species to try focus stacking with, but these guys will do for now. The plentiful numbers mean that the odds of a stationary individual are in my favour :o)

VIEW ON BLACK

Happy Earth Day...:) by Keven Law

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Highest Explore Position #69 ~ On Friday April 23rd 2010.

Red Fox Club - British Wildlife Centre, Surrey, England - Sunday April 18th 2010.

Click here to see My most interesting images

Me and the Foxes are back for Earth Day....aren't you lucky..LOL...:))

Well, tiss been soooooooooo long since I last posted an image I'd almost forgotten how to upload..lol

Anyhoo...here's the first of many foxy images I captured at the Weekend, hope you all like them..:)

I could be made redundant from the job I've had for the last 18 Years in 12 weeks time and from next week I shall be on Gardening leave, whilst I look for another job...I may use some of my redundancy money to travel around the world for a while, whilst I figure out what to do next.........soooooooooo all those in Australia, New Zealand, Hong Kong and the USA....and a few places in between.....if your up for it I may pay you a visit...lol..:))

Have a great Earth Day then everybody and a wonderful upcoming Weekend..:))

Red Fox Cub ~ Fox is a name applied to any of roughly 27 species of small to medium-sized canids, characterized by possessing a long, narrow snout, and a bushy tail, or "brush". By far the most common and widespread species of fox is the red fox (Vulpes vulpes), although various species are found on almost every continent. The presence of fox-like carnivores all over the globe has led to their appearance in the popular culture and folklore of many nations, tribes, and other cultural groups.

Etymology ~ The Modern English "fox" is derived from Old English fox. The Old English word itself comes from the Proto-Germanic word *fukh – compare German Fuchs, Gothic fauho, Old Norse foa and Dutch vos. It corresponds to the Proto-Indo-European word *puke meaning "tail" (compare Sanskrit puccha, also "tail"). The bushy tail is also the source of the word for fox in Welsh: llwynog, from llwyn, "bush", Lithuanian: uodegis, from uodega, "tail", and Portuguese: raposa, from rabo, "tail".

General characteristics ~ Most foxes live 2 to 3 years, but they can survive for up to 10 years or even longer in captivity. Foxes are generally smaller than other members of the family Canidae such as wolves, jackals, and domestic dogs. Dogs (male foxes) weigh on average, 5.9kg and vixens (female foxes) weigh less, at 5.2kg (13 lbs and 11.5 lbs, respectively). Fox-like features typically include an acute muzzle (a "fox face") and bushy tail. Other physical characteristics vary according to their habitat. For example, the fennec fox (and other species of foxes adapted to life in the desert, such as the kit fox) has large ears and short fur, whereas the Arctic fox has small ears and thick, insulating fur.

Another example is the red fox which has a typical auburn pelt, the tail normally ending with white marking.

Unlike many canids, foxes are usually not pack animals. Typically, they are solitary, opportunistic feeders that hunt live prey (especially rodents). Using a pouncing technique practiced from an early age, they are usually able to kill their prey quickly. Foxes also gather a wide variety of other foods ranging from grasshoppers to fruit and berries.

Foxes are normally extremely wary of humans and are not kept as pets (with the exception of the fennec); however, the silver fox was successfully domesticated in Russia after a 45 year selective breeding program. This selective breeding also resulted in physical and behavioural traits appearing that are frequently seen in domestic cats, dogs, and other animals: pigmentation changes, floppy ears, and curly tails.

Diet ~ The diet of foxes comprises rodents, insects, worms, fruit, fish, birds, eggs, and all other kinds of small animals. The fox generally consumes around 1 kg of food every day. Foxes that live in neighborhoods mainly depend on household waste and even rodents and birds that keep moving around these areas. Foxes are known to cache their food, burying the excess for later consumption.

They mostly thrive in the higher latitudes, suburban and even urban environments both in Europe and in North America. They are found also in Eurasia, North Africa, India (Ladakh, Himalayas, Jammu and Kashmir, Rajasthan and Gujarat), China, Japan and in Australia.

Conservation ~ Foxes are readily found in cities and cultivated areas and (depending upon species) seem to adapt reasonably well to human presence.

Red foxes have been introduced into Australia and some other countries for hunting. Australia lacks similar carnivores, and the introduced foxes prey on native wildlife, some to the point of extinction. A similar introduction occurred in the seventeenth and eighteenth centuries in temperate North America, where European reds (Vulpes vulpes) were brought to the colonies for fox hunting, where they decimated the American red fox population through more aggressive hunting and breeding. Interbreeding with American reds, traits of the European red eventually pervaded the gene pool, leaving European and American foxes now virtually identical.

Other fox species do not adapt as well as the red fox, and are endangered in their native environments. Key among these are the crab-eating fox (Cerdocyon thous) and the African bat-eared fox. Other foxes such as fennec foxes, are not endangered, but will be if humans encroach further into their habitat.

Foxes have been successfully employed to control pests on fruit farms, where they leave the fruit intact.

Historians believe foxes were imported into non-native environments long before the colonial era. The first example of the introduction of the fox into a new habitat by humans seems to be Neolithic Cyprus. Stone carvings representing foxes have been found in the early settlement of Göbekli Tepe in eastern Turkey.

A White Lioness -- A White Big Cat II by Bob Gotham

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I've now got a Pro account now .. and have uploaded a full size different version here.

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It was a snapshot with my pocket camera taken from my drivers seat over my passengers lap and through my car window at arms length !

This was taken at Bewdley's (near Kidderminster) Safari Park, hence the pic taken from my car. There are about 3 or 4 white lions and tigers as well as normal lions and tigers too !

There are only 70 white lions in the world. 4 in the UK. They are not albino as they do have pigmentation, which is apparent in the eyes, paw pads and lip colour. This is a condition called leucism where there is loss of pigmentation in the skin and fur with almost normal eye colour. This occurs too in the white tiger.

You can read more about them here. .

NEWS

This picture will soon be seen in a role playing game guide for a character that is a werelion (I don't know if you have ever heard of Laurell K. Hamilton's Anita blake series or not but that's what the game is themed around) and that photo is pretty much dead on for what image they have in their head for the character when she is in lion form and they'd like to use it as a referance for other players to get an idea of what she would look like as well.

Jellyfish - Sea Jellies - Medusa (Subphylum Medusozoa) by PHOTOGRAPHY by DM & DBM.

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Jellyfish, also known sea jellies, are the medusa-phase of certain gelatinous members of the subphylum Medusozoa, which is a major part of the phylum Cnidaria.

Jellyfish are mainly free-swimming marine animals with umbrella-shaped bells and trailing tentacles, although a few are anchored to the seabed by stalks rather than being mobile. The bell can pulsate to provide propulsion for highly efficient locomotion. The tentacles are armed with stinging cells and may be used to capture prey and defend against predators. Jellyfish have a complex life cycle. The medusa is normally the sexual phase, which produces planula larvae; these then disperse widely and enter a sedentary polyp phase, before reaching sexual maturity.

Jellyfish are found all over the world, from surface waters to the deep sea. Scyphozoans (the "true jellyfish") are exclusively marine, but some hydrozoans with a similar appearance live in freshwater. Large, often colorful, jellyfish are common in coastal zones worldwide. The medusae of most species are fast-growing, and mature within a few months then die soon after breeding, but the polyp stage, attached to the seabed, may be much more long-lived. Jellyfish have been in existence for at least 500 million years, and possibly 700 million years or more, making them the oldest multi-organ animal group.

Jellyfish are eaten by humans in certain cultures. They are considered a delicacy in some Asian countries, where species in the Rhizostomeae order are pressed and salted to remove excess water. Australian researchers have described them as a "perfect food": sustainable and protein-rich but relatively low in food energy.

They are also used in research, where the green fluorescent protein used by some species to cause bioluminescence has been adapted as a fluorescent marker for genes inserted into other cells or organisms.

The stinging cells used by jellyfish to subdue their prey can injure humans. Thousands of swimmers worldwide are stung every year, with effects ranging from mild discomfort to serious injury or even death. When conditions are favourable, jellyfish can form vast swarms, which can be responsible for damage to fishing gear by filling fishing nets, and sometimes clog the cooling systems of power and desalination plants which draw their water from the sea.

Names

The name jellyfish, in use since 1796, has traditionally been applied to medusae and all similar animals including the comb jellies (ctenophores, another phylum). The term jellies or sea jellies is more recent, having been introduced by public aquaria in an effort to avoid use of the word "fish" with its modern connotation of an animal with a backbone, though shellfish, cuttlefish and starfish are not vertebrates either. In scientific literature, "jelly" and "jellyfish" have been used interchangeably. Many sources refer to only scyphozoans as "true jellyfish".

A group of jellyfish is called a "smack" or a "smuck".

Definition

The term jellyfish broadly corresponds to medusae, that is, a life-cycle stage in the Medusozoa. The American evolutionary biologist Paulyn Cartwright gives the following general definition:

Typically, medusozoan cnidarians have a pelagic, predatory jellyfish stage in their life cycle; staurozoans are the exceptions [as they are stalked].

The Merriam-Webster dictionary defines jellyfish as follows:

A free-swimming marine coelenterate that is the sexually reproducing form of a hydrozoan or scyphozoan and has a nearly transparent saucer-shaped body and extensible marginal tentacles studded with stinging cells.

Given that jellyfish is a common name, its mapping to biological groups is inexact. Some authorities have called the comb jellies and certain salps jellyfish, though other authorities state that neither of these are jellyfish, which they consider should be limited to certain groups within the medusozoa.

The non-medusozoan clades called jellyfish by some but not all authorities (both agreeing and disagreeing citations are given in each case) are indicated with on the following cladogram of the animal kingdom:

Jellyfish are not a clade, as they include most of the Medusozoa, barring some of the Hydrozoa. The medusozoan groups included by authorities are indicated on the following phylogenetic tree by the presence of citations. Names of included jellyfish, in English where possible, are shown in boldface; the presence of a named and cited example indicates that at least that species within its group has been called a jellyfish.

Taxonomy

The subphylum Medusozoa includes all cnidarians with a medusa stage in their life cycle. The basic cycle is egg, planula larva, polyp, medusa, with the medusa being the sexual stage. The polyp stage is sometimes secondarily lost. The subphylum include the major taxa, Scyphozoa (large jellyfish), Cubozoa (box jellyfish) and Hydrozoa (small jellyfish), and excludes Anthozoa (corals and sea anemones). This suggests that the medusa form evolved after the polyps. Medusozoans have tetramerous symmetry, with parts in fours or multiples of four.

The four major classes of medusozoan Cnidaria are:

Scyphozoa are sometimes called true jellyfish, though they are no more truly jellyfish than the others listed here. They have tetra-radial symmetry. Most have tentacles around the outer margin of the bowl-shaped bell, and long, oral arms around the mouth in the center of the subumbrella.

Cubozoa (box jellyfish) have a (rounded) box-shaped bell, and their velarium assists them to swim more quickly. Box jellyfish may be related more closely to scyphozoan jellyfish than either are to the Hydrozoa.

Hydrozoa medusae also have tetra-radial symmetry, nearly always have a velum (diaphragm used in swimming) attached just inside the bell margin, do not have oral arms, but a much smaller central stalk-like structure, the manubrium, with terminal mouth opening, and are distinguished by the absence of cells in the mesoglea. Hydrozoa show great diversity of lifestyle; some species maintain the polyp form for their entire life and do not form medusae at all (such as Hydra, which is hence not considered a jellyfish), and a few are entirely medusal and have no polyp form.

Staurozoa (stalked jellyfish) are characterized by a medusa form that is generally sessile, oriented upside down and with a stalk emerging from the apex of the "calyx" (bell), which attaches to the substrate. At least some Staurozoa also have a polyp form that alternates with the medusoid portion of the life cycle. Until recently, Staurozoa were classified within the Scyphozoa.

There are over 200 species of Scyphozoa, about 50 species of Staurozoa, about 50 species of Cubozoa, and the Hydrozoa includes about 1000–1500 species that produce medusae, but many more species that do not.

Fossil history

Since jellyfish have no hard parts, fossils are rare. The oldest unambiguous fossil of a free-swimming medusa is Burgessomedusa from the mid Cambrian Burgess Shale of Canada, which is likely either a stem group of box jellyfish (Cubozoa) or Acraspeda (the clade including Staurozoa, Cubozoa, and Scyphozoa). Other claimed records from the Cambrian of China and Utah in the United States are uncertain, and possibly represent ctenophores instead.

Anatomy

The main feature of a true jellyfish is the umbrella-shaped bell. This is a hollow structure consisting of a mass of transparent jelly-like matter known as mesoglea, which forms the hydrostatic skeleton of the animal. 95% or more of the mesogloea consists of water, but it also contains collagen and other fibrous proteins, as well as wandering amoebocytes which can engulf debris and bacteria. The mesogloea is bordered by the epidermis on the outside and the gastrodermis on the inside. The edge of the bell is often divided into rounded lobes known as lappets, which allow the bell to flex. In the gaps or niches between the lappets are dangling rudimentary sense organs known as rhopalia, and the margin of the bell often bears tentacles.

Anatomy of a scyphozoan jellyfish

On the underside of the bell is the manubrium, a stalk-like structure hanging down from the centre, with the mouth, which also functions as the anus, at its tip. There are often four oral arms connected to the manubrium, streaming away into the water below. The mouth opens into the gastrovascular cavity, where digestion takes place and nutrients are absorbed. This is subdivided by four thick septa into a central stomach and four gastric pockets. The four pairs of gonads are attached to the septa, and close to them four septal funnels open to the exterior, perhaps supplying good oxygenation to the gonads. Near the free edges of the septa, gastric filaments extend into the gastric cavity; these are armed with nematocysts and enzyme-producing cells and play a role in subduing and digesting the prey. In some scyphozoans, the gastric cavity is joined to radial canals which branch extensively and may join a marginal ring canal. Cilia in these canals circulate the fluid in a regular direction.

Discharge mechanism of a nematocyst

The box jellyfish is largely similar in structure. It has a squarish, box-like bell. A short pedalium or stalk hangs from each of the four lower corners. One or more long, slender tentacles are attached to each pedalium. The rim of the bell is folded inwards to form a shelf known as a velarium which restricts the bell's aperture and creates a powerful jet when the bell pulsates, allowing box jellyfish to swim faster than true jellyfish. Hydrozoans are also similar, usually with just four tentacles at the edge of the bell, although many hydrozoans are colonial and may not have a free-living medusal stage. In some species, a non-detachable bud known as a gonophore is formed that contains a gonad but is missing many other medusal features such as tentacles and rhopalia. Stalked jellyfish are attached to a solid surface by a basal disk, and resemble a polyp, the oral end of which has partially developed into a medusa with tentacle-bearing lobes and a central manubrium with four-sided mouth.

Most jellyfish do not have specialized systems for osmoregulation, respiration and circulation, and do not have a central nervous system. Nematocysts, which deliver the sting, are located mostly on the tentacles; true jellyfish also have them around the mouth and stomach. Jellyfish do not need a respiratory system because sufficient oxygen diffuses through the epidermis. They have limited control over their movement, but can navigate with the pulsations of the bell-like body; some species are active swimmers most of the time, while others largely drift. The rhopalia contain rudimentary sense organs which are able to detect light, water-borne vibrations, odour and orientation. A loose network of nerves called a "nerve net" is located in the epidermis. Although traditionally thought not to have a central nervous system, nerve net concentration and ganglion-like structures could be considered to constitute one in most species. A jellyfish detects stimuli, and transmits impulses both throughout the nerve net and around a circular nerve ring, to other nerve cells. The rhopalial ganglia contain pacemaker neurones which control swimming rate and direction.

In many species of jellyfish, the rhopalia include ocelli, light-sensitive organs able to tell light from dark. These are generally pigment spot ocelli, which have some of their cells pigmented. The rhopalia are suspended on stalks with heavy crystals at one end, acting like gyroscopes to orient the eyes skyward. Certain jellyfish look upward at the mangrove canopy while making a daily migration from mangrove swamps into the open lagoon, where they feed, and back again.

Box jellyfish have more advanced vision than the other groups. Each individual has 24 eyes, two of which are capable of seeing colour, and four parallel information processing areas that act in competition, supposedly making them one of the few kinds of animal to have a 360-degree view of its environment.

Box jellyfish eye

The study of jellyfish eye evolution is an intermediary to a better understanding of how visual systems evolved on Earth. Jellyfish exhibit immense variation in visual systems ranging from photoreceptive cell patches seen in simple photoreceptive systems to more derived complex eyes seen in box jellyfish. Major topics of jellyfish visual system research (with an emphasis on box jellyfish) include: the evolution of jellyfish vision from simple to complex visual systems), the eye morphology and molecular structures of box jellyfish (including comparisons to vertebrate eyes), and various uses of vision including task-guided behaviors and niche specialization.

Evolution

Experimental evidence for photosensitivity and photoreception in cnidarians antecedes the mid 1900s, and a rich body of research has since covered evolution of visual systems in jellyfish. Jellyfish visual systems range from simple photoreceptive cells to complex image-forming eyes. More ancestral visual systems incorporate extraocular vision (vision without eyes) that encompass numerous receptors dedicated to single-function behaviors. More derived visual systems comprise perception that is capable of multiple task-guided behaviors.

Although they lack a true brain, cnidarian jellyfish have a "ring" nervous system that plays a significant role in motor and sensory activity. This net of nerves is responsible for muscle contraction and movement and culminates the emergence of photosensitive structures. Across Cnidaria, there is large variation in the systems that underlie photosensitivity. Photosensitive structures range from non-specialized groups of cells, to more "conventional" eyes similar to those of vertebrates. The general evolutionary steps to develop complex vision include (from more ancestral to more derived states): non-directional photoreception, directional photoreception, low-resolution vision, and high-resolution vision. Increased habitat and task complexity has favored the high-resolution visual systems common in derived cnidarians such as box jellyfish.

Basal visual systems observed in various cnidarians exhibit photosensitivity representative of a single task or behavior. Extraocular photoreception (a form of non-directional photoreception), is the most basic form of light sensitivity and guides a variety of behaviors among cnidarians. It can function to regulate circadian rhythm (as seen in eyeless hydrozoans) and other light-guided behaviors responsive to the intensity and spectrum of light. Extraocular photoreception can function additionally in positive phototaxis (in planula larvae of hydrozoans), as well as in avoiding harmful amounts of UV radiation via negative phototaxis. Directional photoreception (the ability to perceive direction of incoming light) allows for more complex phototactic responses to light, and likely evolved by means of membrane stacking. The resulting behavioral responses can range from guided spawning events timed by moonlight to shadow responses for potential predator avoidance. Light-guided behaviors are observed in numerous scyphozoans including the common moon jelly, Aurelia aurita, which migrates in response to changes in ambient light and solar position even though they lack proper eyes.

The low-resolution visual system of box jellyfish is more derived than directional photoreception, and thus box jellyfish vision represents the most basic form of true vision in which multiple directional photoreceptors combine to create the first imaging and spatial resolution. This is different from the high-resolution vision that is observed in camera or compound eyes of vertebrates and cephalopods that rely on focusing optics. Critically, the visual systems of box jellyfish are responsible for guiding multiple tasks or behaviors in contrast to less derived visual systems in other jellyfish that guide single behavioral functions. These behaviors include phototaxis based on sunlight (positive) or shadows (negative), obstacle avoidance, and control of swim-pulse rate.

Box jellyfish possess "proper eyes" (similar to vertebrates) that allow them to inhabit environments that lesser derived medusae cannot. In fact, they are considered the only class in the clade Medusozoa that have behaviors necessitating spatial resolution and genuine vision. However, the lens in their eyes are more functionally similar to cup-eyes exhibited in low-resolution organisms, and have very little to no focusing capability. The lack of the ability to focus is due to the focal length exceeding the distance to the retina, thus generating unfocused images and limiting spatial resolution. The visual system is still sufficient for box jellyfish to produce an image to help with tasks such as object avoidance.

Utility as a model organism

Box jellyfish eyes are a visual system that is sophisticated in numerous ways. These intricacies include the considerable variation within the morphology of box jellyfishes' eyes (including their task/behavior specification), and the molecular makeup of their eyes including: photoreceptors, opsins, lenses, and synapses. The comparison of these attributes to more derived visual systems can allow for a further understanding of how the evolution of more derived visual systems may have occurred, and puts into perspective how box jellyfish can play the role as an evolutionary/developmental model for all visual systems.

Characteristics

Box jellyfish visual systems are both diverse and complex, comprising multiple photosystems. There is likely considerable variation in visual properties between species of box jellyfish given the significant inter-species morphological and physiological variation. Eyes tend to differ in size and shape, along with number of receptors (including opsins), and physiology across species of box jellyfish.

Box jellyfish have a series of intricate lensed eyes that are similar to those of more derived multicellular organisms such as vertebrates. Their 24 eyes fit into four different morphological categories. These categories consist of two large, morphologically different medial eyes (a lower and upper lensed eye) containing spherical lenses, a lateral pair of pigment slit eyes, and a lateral pair of pigment pit eyes. The eyes are situated on rhopalia (small sensory structures) which serve sensory functions of the box jellyfish and arise from the cavities of the exumbrella (the surface of the body) on the side of the bells of the jellyfish. The two large eyes are located on the mid-line of the club and are considered complex because they contain lenses. The four remaining eyes lie laterally on either side of each rhopalia and are considered simple. The simple eyes are observed as small invaginated cups of epithelium that have developed pigmentation. The larger of the complex eyes contains a cellular cornea created by a mono ciliated epithelium, cellular lens, homogenous capsule to the lens, vitreous body with prismatic elements, and a retina of pigmented cells. The smaller of the complex eyes is said to be slightly less complex given that it lacks a capsule but otherwise contains the same structure as the larger eye.

Box jellyfish have multiple photosystems that comprise different sets of eyes. Evidence includes immunocytochemical and molecular data that show photopigment differences among the different morphological eye types, and physiological experiments done on box jellyfish to suggest behavioral differences among photosystems. Each individual eye type constitutes photosystems that work collectively to control visually guided behaviors.

Box jellyfish eyes primarily use c-PRCs (ciliary photoreceptor cells) similar to that of vertebrate eyes. These cells undergo phototransduction cascades (process of light absorption by photoreceptors) that are triggered by c-opsins. Available opsin sequences suggest that there are two types of opsins possessed by all cnidarians including an ancient phylogenetic opsin, and a sister ciliary opsin to the c-opsins group. Box jellyfish could have both ciliary and cnidops (cnidarian opsins), which is something not previously believed to appear in the same retina. Nevertheless, it is not entirely evident whether cnidarians possess multiple opsins that are capable of having distinctive spectral sensitivities.

Comparison with other organisms

Comparative research on genetic and molecular makeup of box jellyfishes' eyes versus more derived eyes seen in vertebrates and cephalopods focuses on: lenses and crystallin composition, synapses, and Pax genes and their implied evidence for shared primordial (ancestral) genes in eye evolution.

Box jellyfish eyes are said to be an evolutionary/developmental model of all eyes based on their evolutionary recruitment of crystallins and Pax genes. Research done on box jellyfish including Tripedalia cystophora has suggested that they possess a single Pax gene, PaxB. PaxB functions by binding to crystallin promoters and activating them. PaxB in situ hybridization resulted in PaxB expression in the lens, retina, and statocysts. These results and the rejection of the prior hypothesis that Pax6 was an ancestral Pax gene in eyes has led to the conclusion that PaxB was a primordial gene in eye evolution, and that the eyes of all organisms likely share a common ancestor.

The lens structure of box jellyfish appears very similar to those of other organisms, but the crystallins are distinct in both function and appearance. Weak reactions were seen within the sera and there were very weak sequence similarities within the crystallins among vertebrate and invertebrate lenses. This is likely due to differences in lower molecular weight proteins and the subsequent lack of immunological reactions with antisera that other organisms' lenses exhibit.

All four of the visual systems of box jellyfish species investigated with detail (Carybdea marsupialis, Chiropsalmus quadrumanus, Tamoya haplonema and Tripedalia cystophora) have invaginated synapses, but only in the upper and lower lensed eyes. Different densities were found between the upper and lower lenses, and between species. Four types of chemical synapses have been discovered within the rhopalia which could help in understanding neural organization including: clear unidirectional, dense-core unidirectional, clear bidirectional, and clear and dense-core bidirectional. The synapses of the lensed eyes could be useful as markers to learn more about the neural circuit in box jellyfish retinal areas.

Evolution as a response to natural stimuli

The primary adaptive responses to environmental variation observed in box jellyfish eyes include pupillary constriction speeds in response to light environments, as well as photoreceptor tuning and lens adaptations to better respond to shifts between light environments and darkness. Interestingly, some box jellyfish species' eyes appear to have evolved more focused vision in response to their habitat.

Pupillary contraction appears to have evolved in response to variation in the light environment across ecological niches across three species of box jellyfish (Chironex fleckeri, Chiropsella bronzie, and Carukia barnesi). Behavioral studies suggest that faster pupil contraction rates allow for greater object avoidance, and in fact, species with more complex habitats exhibit faster rates. Ch. bronzie inhabit shallow beach fronts that have low visibility and very few obstacles, thus, faster pupil contraction in response to objects in their environment is not important. Ca. barnesi and Ch. fleckeri are found in more three-dimensionally complex environments like mangroves with an abundance of natural obstacles, where faster pupil contraction is more adaptive. Behavioral studies support the idea that faster pupillary contraction rates assist with obstacle avoidance as well as depth adjustments in response to differing light intensities.

Light/dark adaptation via pupillary light reflexes is an additional form of an evolutionary response to the light environment. This relates to the pupil's response to shifts between light intensity (generally from sunlight to darkness). In the process of light/dark adaptation, the upper and lower lens eyes of different box jellyfish species vary in specific function. The lower lens-eyes contain pigmented photoreceptors and long pigment cells with dark pigments that migrate on light/dark adaptation, while the upper-lens eyes play a concentrated role in light direction and phototaxis given that they face upward towards the water surface (towards the sun or moon). The upper lens of Ch. bronzie does not exhibit any considerable optical power while Tr. cystophora (a box jellyfish species that tends to live in mangroves) does. The ability to use light to visually guide behavior is not of as much importance to Ch. bronzie as it is to species in more obstacle-filled environments. Differences in visually guided behavior serve as evidence that species that share the same number and structure of eyes can exhibit differences in how they control behavior.

Largest and smallest

Jellyfish range from about one millimeter in bell height and diameter, to nearly 2 metres (6+1⁄2 ft) in bell height and diameter; the tentacles and mouth parts usually extend beyond this bell dimension.

The smallest jellyfish are the peculiar creeping jellyfish in the genera Staurocladia and Eleutheria, which have bell disks from 0.5 millimetres (1⁄32 in) to a few millimeters in diameter, with short tentacles that extend out beyond this, which these jellyfish use to move across the surface of seaweed or the bottoms of rocky pools; many of these tiny creeping jellyfish cannot be seen in the field without a hand lens or microscope. They can reproduce asexually by fission (splitting in half). Other very small jellyfish, which have bells about one millimeter, are the hydromedusae of many species that have just been released from their parent polyps; some of these live only a few minutes before shedding their gametes in the plankton and then dying, while others will grow in the plankton for weeks or months. The hydromedusae Cladonema radiatum and Cladonema californicum are also very small, living for months, yet never growing beyond a few mm in bell height and diameter.

The lion's mane jellyfish, Cyanea capillata, was long-cited as the largest jellyfish, and arguably the longest animal in the world, with fine, thread-like tentacles that may extend up to 36.5 m (119 ft 9 in) long (though most are nowhere near that large). They have a moderately painful, but rarely fatal, sting. The increasingly common giant Nomura's jellyfish, Nemopilema nomurai, found in some, but not all years in the waters of Japan, Korea and China in summer and autumn is another candidate for "largest jellyfish", in terms of diameter and weight, since the largest Nomura's jellyfish in late autumn can reach 2 m (6 ft 7 in) in bell (body) diameter and about 200 kg (440 lb) in weight, with average specimens frequently reaching 0.9 m (2 ft 11 in) in bell diameter and about 150 kg (330 lb) in weight. The large bell mass of the giant Nomura's jellyfish can dwarf a diver and is nearly always much greater than the Lion's Mane, whose bell diameter can reach 1 m (3 ft 3 in).

The rarely encountered deep-sea jellyfish Stygiomedusa gigantea is another candidate for "largest jellyfish", with its thick, massive bell up to 100 cm (3 ft 3 in) wide, and four thick, "strap-like" oral arms extending up to 6 m (19+1⁄2 ft) in length, very different from the typical fine, threadlike tentacles that rim the umbrella of more-typical-looking jellyfish, including the Lion's Mane.

Desmonema glaciale, which lives in the Antarctic region, can reach a very large size (several meters). Purple-striped jelly (Chrysaora colorata) can also be extremely long (up to 15 feet).

Life history and behavior

Life cycle

Jellyfish have a complex life cycle which includes both sexual and asexual phases, with the medusa being the sexual stage in most instances. Sperm fertilize eggs, which develop into larval planulae, become polyps, bud into ephyrae and then transform into adult medusae. In some species certain stages may be skipped.

Upon reaching adult size, jellyfish spawn regularly if there is a sufficient supply of food. In most species, spawning is controlled by light, with all individuals spawning at about the same time of day; in many instances this is at dawn or dusk. Jellyfish are usually either male or female (with occasional hermaphrodites). In most cases, adults release sperm and eggs into the surrounding water, where the unprotected eggs are fertilized and develop into larvae. In a few species, the sperm swim into the female's mouth, fertilizing the eggs within her body, where they remain during early development stages. In moon jellies, the eggs lodge in pits on the oral arms, which form a temporary brood chamber for the developing planula larvae.

The planula is a small larva covered with cilia. When sufficiently developed, it settles onto a firm surface and develops into a polyp. The polyp generally consists of a small stalk topped by a mouth that is ringed by upward-facing tentacles. The polyps resemble those of closely related anthozoans, such as sea anemones and corals. The jellyfish polyp may be sessile, living on the bottom, boat hulls or other substrates, or it may be free-floating or attached to tiny bits of free-living plankton or rarely, fish or other invertebrates. Polyps may be solitary or colonial. Most polyps are only millimetres in diameter and feed continuously. The polyp stage may last for years.

After an interval and stimulated by seasonal or hormonal changes, the polyp may begin reproducing asexually by budding and, in the Scyphozoa, is called a segmenting polyp, or a scyphistoma. Budding produces more scyphistomae and also ephyrae. Budding sites vary by species; from the tentacle bulbs, the manubrium (above the mouth), or the gonads of hydromedusae. In a process known as strobilation, the polyp's tentacles are reabsorbed and the body starts to narrow, forming transverse constrictions, in several places near the upper extremity of the polyp. These deepen as the constriction sites migrate down the body, and separate segments known as ephyra detach. These are free-swimming precursors of the adult medusa stage, which is the life stage that is typically identified as a jellyfish. The ephyrae, usually only a millimeter or two across initially, swim away from the polyp and grow. Limnomedusae polyps can asexually produce a creeping frustule larval form, which crawls away before developing into another polyp. A few species can produce new medusae by budding directly from the medusan stage. Some hydromedusae reproduce by fission.

Lifespan

Little is known of the life histories of many jellyfish as the places on the seabed where the benthic forms of those species live have not been found. However, an asexually reproducing strobila form can sometimes live for several years, producing new medusae (ephyra larvae) each year.

An unusual species, Turritopsis dohrnii, formerly classified as Turritopsis nutricula, might be effectively immortal because of its ability under certain circumstances to transform from medusa back to the polyp stage, thereby escaping the death that typically awaits medusae post-reproduction if they have not otherwise been eaten by some other organism. So far this reversal has been observed only in the laboratory.

Locomotion

Jellyfish locomotion is highly efficient. Muscles in the jellylike bell contract, setting up a start vortex and propelling the animal. When the contraction ends, the bell recoils elastically, creating a stop vortex with no extra energy input.

Using the moon jelly Aurelia aurita as an example, jellyfish have been shown to be the most energy-efficient swimmers of all animals. They move through the water by radially expanding and contracting their bell-shaped bodies to push water behind them. They pause between the contraction and expansion phases to create two vortex rings. Muscles are used for the contraction of the body, which creates the first vortex and pushes the animal forward, but the mesoglea is so elastic that the expansion is powered exclusively by relaxing the bell, which releases the energy stored from the contraction. Meanwhile, the second vortex ring starts to spin faster, sucking water into the bell and pushing against the centre of the body, giving a secondary and "free" boost forward. The mechanism, called passive energy recapture, only works in relatively small jellyfish moving at low speeds, allowing the animal to travel 30 percent farther on each swimming cycle. Jellyfish achieved a 48 percent lower cost of transport (food and oxygen intake versus energy spent in movement) than other animals in similar studies. One reason for this is that most of the gelatinous tissue of the bell is inactive, using no energy during swimming.

Ecology

Diet

Jellyfish are, like other cnidarians, generally carnivorous (or parasitic), feeding on planktonic organisms, crustaceans, small fish, fish eggs and larvae, and other jellyfish, ingesting food and voiding undigested waste through the mouth. They hunt passively using their tentacles as drift lines, or sink through the water with their tentacles spread widely; the tentacles, which contain nematocysts to stun or kill the prey, may then flex to help bring it to the mouth. Their swimming technique also helps them to capture prey; when their bell expands it sucks in water which brings more potential prey within reach of the tentacles.

A few species such as Aglaura hemistoma are omnivorous, feeding on microplankton which is a mixture of zooplankton and phytoplankton (microscopic plants) such as dinoflagellates. Others harbour mutualistic algae (Zooxanthellae) in their tissues; the spotted jellyfish (Mastigias papua) is typical of these, deriving part of its nutrition from the products of photosynthesis, and part from captured zooplankton. The upside-down jellyfish (Cassiopea andromeda) also has a symbiotic relationship with microalgae, but captures tiny animals to supplement their diet. This is done by releasing tiny balls of living cells composed of mesoglea. These use cilia to drive them through water and stinging cells which stun the prey. The blobs also seems to have digestive capabilities.

Predation

Other species of jellyfish are among the most common and important jellyfish predators. Sea anemones may eat jellyfish that drift into their range. Other predators include tunas, sharks, swordfish, sea turtles and penguins. Jellyfish washed up on the beach are consumed by foxes, other terrestrial mammals and birds. In general however, few animals prey on jellyfish; they can broadly be considered to be top predators in the food chain. Once jellyfish have become dominant in an ecosystem, for example through overfishing which removes predators of jellyfish larvae, there may be no obvious way for the previous balance to be restored: they eat fish eggs and juvenile fish, and compete with fish for food, preventing fish stocks from recovering.

Symbiosis

Some small fish are immune to the stings of the jellyfish and live among the tentacles, serving as bait in a fish trap; they are safe from potential predators and are able to share the fish caught by the jellyfish. The cannonball jellyfish has a symbiotic relationship with ten different species of fish, and with the longnose spider crab, which lives inside the bell, sharing the jellyfish's food and nibbling its tissues.

Main article: Jellyfish bloom

Jellyfish form large masses or blooms in certain environmental conditions of ocean currents, nutrients, sunshine, temperature, season, prey availability, reduced predation and oxygen concentration. Currents collect jellyfish together, especially in years with unusually high populations. Jellyfish can detect marine currents and swim against the current to congregate in blooms. Jellyfish are better able to survive in nutrient-rich, oxygen-poor water than competitors, and thus can feast on plankton without competition. Jellyfish may also benefit from saltier waters, as saltier waters contain more iodine, which is necessary for polyps to turn into jellyfish. Rising sea temperatures caused by climate change may also contribute to jellyfish blooms, because many species of jellyfish are able to survive in warmer waters. Increased nutrients from agricultural or urban runoff with nutrients including nitrogen and phosphorus compounds increase the growth of phytoplankton, causing eutrophication and algal blooms. When the phytoplankton die, they may create dead zones, so-called because they are hypoxic (low in oxygen). This in turn kills fish and other animals, but not jellyfish, allowing them to bloom. Jellyfish populations may be expanding globally as a result of land runoff and overfishing of their natural predators. Jellyfish are well placed to benefit from disturbance of marine ecosystems. They reproduce rapidly; they prey upon many species, while few species prey on them; and they feed via touch rather than visually, so they can feed effectively at night and in turbid waters. It may be difficult for fish stocks to re-establish themselves in marine ecosystems once they have become dominated by jellyfish, because jellyfish feed on plankton, which includes fish eggs and larvae.

As suspected at the turn of this century, jellyfish blooms are increasing in frequency. Between 2013 and 2020 the Mediterranean Science Commission monitored on a weekly basis the frequency of such outbreaks in coastal waters from Morocco to the Black Sea, revealing a relatively high frequency of these blooms nearly all year round, with peaks observed from March to July and often again in the autumn. The blooms are caused by different jellyfish species, depending on their localisation within the Basin: one observes a clear dominance of Pelagia noctiluca and Velella velella outbreaks in the western Mediterranean, of Rhizostoma pulmo and Rhopilema nomadica outbreaks in the eastern Mediterranean, and of Aurelia aurita and Mnemiopsis leidyi outbreaks in the Black Sea.

Some jellyfish populations that have shown clear increases in the past few decades are invasive species, newly arrived from other habitats: examples include the Black Sea, Caspian Sea, Baltic Sea, central and eastern Mediterranean, Hawaii, and tropical and subtropical parts of the West Atlantic (including the Caribbean, Gulf of Mexico and Brazil).

Jellyfish blooms can have significant impact on community structure. Some carnivorous jellyfish species prey on zooplankton while others graze on primary producers. Reductions in zooplankton and ichthyoplankton due to a jellyfish bloom can ripple through the trophic levels. High-density jellyfish populations can outcompete other predators and reduce fish recruitment. Increased grazing on primary producers by jellyfish can also interrupt energy transfer to higher trophic levels.

During blooms, jellyfish significantly alter the nutrient availability in their environment. Blooms require large amounts of available organic nutrients in the water column to grow, limiting availability for other organisms. Some jellyfish have a symbiotic relationship with single-celled dinoflagellates, allowing them to assimilate inorganic carbon, phosphorus, and nitrogen creating competition for phytoplankton. Their large biomass makes them an important source of dissolved and particulate organic matter for microbial communities through excretion, mucus production, and decomposition. The microbes break down the organic matter into inorganic ammonium and phosphate. However, the low carbon availability shifts the process from production to respiration creating low oxygen areas making the dissolved inorganic nitrogen and phosphorus largely unavailable for primary production.

These blooms have very real impacts on industries. Jellyfish can outcompete fish by utilizing open niches in over-fished fisheries. Catch of jellyfish can strain fishing gear and lead to expenses relating to damaged gear. Power plants have been shut down due to jellyfish blocking the flow of cooling water. Blooms have also been harmful for tourism, causing a rise in stings and sometimes the closure of beaches.

Jellyfish form a component of jelly-falls, events where gelatinous zooplankton fall to the seafloor, providing food for the benthic organisms there. In temperate and subpolar regions, jelly-falls usually follow immediately after a bloom.

Habitats

Most jellyfish are marine animals, although a few hydromedusae inhabit freshwater. The best known freshwater example is the cosmopolitan hydrozoan jellyfish, Craspedacusta sowerbii. It is less than an inch (2.5 cm) in diameter, colorless and does not sting. Some jellyfish populations have become restricted to coastal saltwater lakes, such as Jellyfish Lake in Palau. Jellyfish Lake is a marine lake where millions of golden jellyfish (Mastigias spp.) migrate horizontally across the lake daily.

Although most jellyfish live well off the ocean floor and form part of the plankton, a few species are closely associated with the bottom for much of their lives and can be considered benthic. The upside-down jellyfish in the genus Cassiopea typically lie on the bottom of shallow lagoons where they sometimes pulsate gently with their umbrella top facing down. Even some deep-sea species of hydromedusae and scyphomedusae are usually collected on or near the bottom. All of the stauromedusae are found attached to either seaweed or rocky or other firm material on the bottom.

Some species explicitly adapt to tidal flux. In Roscoe Bay, jellyfish ride the current at ebb tide until they hit a gravel bar, and then descend below the current. They remain in still waters until the tide rises, ascending and allowing it to sweep them back into the bay. They also actively avoid fresh water from mountain snowmelt, diving until they find enough salt.

Parasites

Jellyfish are hosts to a wide variety of parasitic organisms. They act as intermediate hosts of endoparasitic helminths, with the infection being transferred to the definitive host fish after predation. Some digenean trematodes, especially species in the family Lepocreadiidae, use jellyfish as their second intermediate hosts. Fish become infected by the trematodes when they feed on infected jellyfish.

Relation to humans

Jellyfish have long been eaten in some parts of the world. Fisheries have begun harvesting the American cannonball jellyfish, Stomolophus meleagris, along the southern Atlantic coast of the United States and in the Gulf of Mexico for export to Asia.

Jellyfish are also harvested for their collagen, which is being investigated for use in a variety of applications including the treatment of rheumatoid arthritis.

Aquaculture and fisheries of other species often suffer severe losses – and so losses of productivity – due to jellyfish.

Products

Main article: Jellyfish as food

In some countries, including China, Japan, and Korea, jellyfish are a delicacy. The jellyfish is dried to prevent spoiling. Only some 12 species of scyphozoan jellyfish belonging to the order Rhizostomeae are harvested for food, mostly in southeast Asia. Rhizostomes, especially Rhopilema esculentum in China (海蜇 hǎizhé, 'sea stingers') and Stomolophus meleagris (cannonball jellyfish) in the United States, are favored because of their larger and more rigid bodies and because their toxins are harmless to humans.

Traditional processing methods, carried out by a jellyfish master, involve a 20- to 40-day multi-phase procedure in which, after removing the gonads and mucous membranes, the umbrella and oral arms are treated with a mixture of table salt and alum, and compressed. Processing makes the jellyfish drier and more acidic, producing a crisp texture. Jellyfish prepared this way retain 7–10% of their original weight, and the processed product consists of approximately 94% water and 6% protein. Freshly processed jellyfish has a white, creamy color and turns yellow or brown during prolonged storage.

In China, processed jellyfish are desalted by soaking in water overnight and eaten cooked or raw. The dish is often served shredded with a dressing of oil, soy sauce, vinegar and sugar, or as a salad with vegetables. In Japan, cured jellyfish are rinsed, cut into strips and served with vinegar as an appetizer. Desalted, ready-to-eat products are also available.

Biotechnology

The hydromedusa Aequorea victoria was the source of green fluorescent protein, studied for its role in bioluminescence and later for use as a marker in genetic engineering.

Pliny the Elder reported in his Natural History that the slime of the jellyfish "Pulmo marinus" produced light when rubbed on a walking stick.

In 1961, Osamu Shimomura extracted green fluorescent protein (GFP) and another bioluminescent protein, called aequorin, from the large and abundant hydromedusa Aequorea victoria, while studying photoproteins that cause bioluminescence in this species. Three decades later, Douglas Prasher sequenced and cloned the gene for GFP. Martin Chalfie figured out how to use GFP as a fluorescent marker of genes inserted into other cells or organisms. Roger Tsien later chemically manipulated GFP to produce other fluorescent colors to use as markers. In 2008, Shimomura, Chalfie and Tsien won the Nobel Prize in Chemistry for their work with GFP. Man-made GFP became widely used as a fluorescent tag to show which cells or tissues express specific genes. The genetic engineering technique fuses the gene of interest to the GFP gene. The fused DNA is then put into a cell, to generate either a cell line or (via IVF techniques) an entire animal bearing the gene. In the cell or animal, the artificial gene turns on in the same tissues and the same time as the normal gene, making a fusion of the normal protein with GFP attached to the end, illuminating the animal or cell reveals what tissues express that protein—or at what stage of development. The fluorescence shows where the gene is expressed.

Aquarium display

Jellyfish are displayed in many public aquariums. Often the tank's background is blue and the animals are illuminated by side light, increasing the contrast between the animal and the background. In natural conditions, many jellies are so transparent that they are nearly invisible. Jellyfish are not adapted to closed spaces. They depend on currents to transport them from place to place. Professional exhibits as in the Monterey Bay Aquarium feature precise water flows, typically in circular tanks to avoid trapping specimens in corners. The outflow is spread out over a large surface area and the inflow enters as a sheet of water in front of the outflow, so the jellyfish do not get sucked into it. As of 2009, jellyfish were becoming popular in home aquariums, where they require similar equipment.

Stings

Jellyfish are armed with nematocysts, a type of specialized stinging cell. Contact with a jellyfish tentacle can trigger millions of nematocysts to pierce the skin and inject venom, but only some species' venom causes an adverse reaction in humans. In a study published in Communications Biology, researchers found a jellyfish species called Cassiopea xamachana which when triggered will release tiny balls of cells that swim around the jellyfish stinging everything in their path. Researchers described these as "self-propelling microscopic grenades" and named them cassiosomes.

The effects of stings range from mild discomfort to extreme pain and death. Most jellyfish stings are not deadly, but stings of some box jellyfish (Irukandji jellyfish), such as the sea wasp, can be deadly. Stings may cause anaphylaxis (a form of shock), which can be fatal. Jellyfish kill 20 to 40 people a year in the Philippines alone. In 2006 the Spanish Red Cross treated 19,000 stung swimmers along the Costa Brava.

Vinegar (3–10% aqueous acetic acid) may help with box jellyfish stings but not the stings of the Portuguese man o' war. Clearing the area of jelly and tentacles reduces nematocyst firing. Scraping the affected skin, such as with the edge of a credit card, may remove remaining nematocysts. Once the skin has been cleaned of nematocysts, hydrocortisone cream applied locally reduces pain and inflammation. Antihistamines may help to control itching. Immunobased antivenins are used for serious box jellyfish stings.

In Elba Island and Corsica dittrichia viscosa is now used by residents and tourists to heal stings from jellyfish, bees and wasps pressing fresh leaves on the skin with quick results.

Mechanical issues

Jellyfish in large quantities can fill and split fishing nets and crush captured fish. They can clog cooling equipment, having disabled power stations in several countries; jellyfish caused a cascading blackout in the Philippines in 1999, as well as damaging the Diablo Canyon Power Plant in California in 2008. They can also stop desalination plants and ships' engines.

Germany - Lower Saxony - Corn Harvest - 3 by Manfred Sommer