Panelists include Joyce Parker, Co-Founder & Executive Director, Citizens for a Better Greenville, Jack Wuest, Alternative Schools Network, Chicago, and Rodney Skager, Professor Emeritus, UCLA Graduate School of Education.

Snakes are elongated, limbless, carnivorous reptiles of the suborder Serpentes Like all other squamates, snakes are ectothermic, amniote vertebrates covered in overlapping scales. Many species of snakes have skulls with several more joints than their lizard ancestors, enabling them to swallow prey much larger than their heads (cranial kinesis). To accommodate their narrow bodies, snakes' paired organs (such as kidneys) appear one in front of the other instead of side by side, and most have only one functional lung. Some species retain a pelvic girdle with a pair of vestigial claws on either side of the cloaca. Lizards have independently evolved elongate bodies without limbs or with greatly reduced limbs at least twenty-five times via convergent evolution, leading to many lineages of legless lizards. These resemble snakes, but several common groups of legless lizards have eyelids and external ears, which snakes lack, although this rule is not universal (see Amphisbaenia, Dibamidae, and Pygopodidae).

 

Living snakes are found on every continent except Antarctica, and on most smaller land masses; exceptions include some large islands, such as Ireland, Iceland, Greenland, the Hawaiian archipelago, and the islands of New Zealand, as well as many small islands of the Atlantic and central Pacific oceans. Additionally, sea snakes are widespread throughout the Indian and Pacific oceans. Around thirty families are currently recognized, comprising about 520 genera and about 3,900 species. They range in size from the tiny, 10.4 cm-long (4.1 in) Barbados threadsnake to the reticulated python of 6.95 meters (22.8 ft) in length. The fossil species Titanoboa cerrejonensis was 12.8 meters (42 ft) long. Snakes are thought to have evolved from either burrowing or aquatic lizards, perhaps during the Jurassic period, with the earliest known fossils dating to between 143 and 167 Ma ago. The diversity of modern snakes appeared during the Paleocene epoch (c. 66 to 56 Ma ago, after the Cretaceous–Paleogene extinction event). The oldest preserved descriptions of snakes can be found in the Brooklyn Papyrus.

 

Most species of snake are nonvenomous and those that have venom use it primarily to kill and subdue prey rather than for self-defense. Some possess venom that is potent enough to cause painful injury or death to humans. Nonvenomous snakes either swallow prey alive or kill by constriction.

 

Etymology

The English word snake comes from Old English snaca, itself from Proto-Germanic *snak-an- (cf. Germanic Schnake 'ring snake', Swedish snok 'grass snake'), from Proto-Indo-European root *(s)nēg-o- 'to crawl to creep', which also gave sneak as well as Sanskrit nāgá 'snake'. The word ousted adder, as adder went on to narrow in meaning, though in Old English næddre was the general word for snake. The other term, serpent, is from French, ultimately from Indo-European *serp- 'to creep', which also gave Ancient Greek ἕρπω (hérpō) 'I crawl' and Sanskrit sarpá ‘snake’.

 

The fossil record of snakes is relatively poor because snake skeletons are typically small and fragile making fossilization uncommon. Fossils readily identifiable as snakes (though often retaining hind limbs) first appear in the fossil record during the Cretaceous period. The earliest known true snake fossils (members of the crown group Serpentes) come from the marine simoliophiids, the oldest of which is the Late Cretaceous (Cenomanian age) Haasiophis terrasanctus from the West Bank, dated to between 112 and 94 million years old.

 

Based on comparative anatomy, there is consensus that snakes descended from lizards. Pythons and boas—primitive groups among modern snakes—have vestigial hind limbs: tiny, clawed digits known as anal spurs, which are used to grasp during mating The families Leptotyphlopidae and Typhlopidae also possess remnants of the pelvic girdle, appearing as horny projections when visible.

 

Front limbs are nonexistent in all known snakes. This is caused by the evolution of their Hox genes, controlling limb morphogenesis. The axial skeleton of the snakes' common ancestor, like most other tetrapods, had regional specializations consisting of cervical (neck), thoracic (chest), lumbar (lower back), sacral (pelvic), and caudal (tail) vertebrae. Early in snake evolution, the Hox gene expression in the axial skeleton responsible for the development of the thorax became dominant. As a result, the vertebrae anterior to the hindlimb buds (when present) all have the same thoracic-like identity (except from the atlas, axis, and 1–3 neck vertebrae). In other words, most of a snake's skeleton is an extremely extended thorax. Ribs are found exclusively on the thoracic vertebrae. Neck, lumbar and pelvic vertebrae are very reduced in number (only 2–10 lumbar and pelvic vertebrae are present), while only a short tail remains of the caudal vertebrae. However, the tail is still long enough to be of important use in many species, and is modified in some aquatic and tree-dwelling species.

 

Many modern snake groups originated during the Paleocene, alongside the adaptive radiation of mammals following the extinction of (non-avian) dinosaurs. The expansion of grasslands in North America also led to an explosive radiation among snakes. Previously, snakes were a minor component of the North American fauna, but during the Miocene, the number of species and their prevalence increased dramatically with the first appearances of vipers and elapids in North America and the significant diversification of Colubridae (including the origin of many modern genera such as Nerodia, Lampropeltis, Pituophis, and Pantherophis).

 

Fossils

There is fossil evidence to suggest that snakes may have evolved from burrowing lizards, during the Cretaceous Period. An early fossil snake relative, Najash rionegrina, was a two-legged burrowing animal with a sacrum, and was fully terrestrial. One extant analog of these putative ancestors is the earless monitor Lanthanotus of Borneo (though it also is semiaquatic). Subterranean species evolved bodies streamlined for burrowing, and eventually lost their limbs. According to this hypothesis, features such as the transparent, fused eyelids (brille) and loss of external ears evolved to cope with fossorial difficulties, such as scratched corneas and dirt in the ears. Some primitive snakes are known to have possessed hindlimbs, but their pelvic bones lacked a direct connection to the vertebrae. These include fossil species like Haasiophis, Pachyrhachis and Eupodophis, which are slightly older than Najash.

 

This hypothesis was strengthened in 2015 by the discovery of a 113-million-year-old fossil of a four-legged snake in Brazil that has been named Tetrapodophis amplectus. It has many snake-like features, is adapted for burrowing and its stomach indicates that it was preying on other animals. It is currently uncertain if Tetrapodophis is a snake or another species, in the squamate order, as a snake-like body has independently evolved at least 26 times. Tetrapodophis does not have distinctive snake features in its spine and skull. A study in 2021 places the animal in a group of extinct marine lizards from the Cretaceous period known as dolichosaurs and not directly related to snakes.

 

An alternative hypothesis, based on morphology, suggests the ancestors of snakes were related to mosasaurs—extinct aquatic reptiles from the Cretaceous—forming the clade Pythonomorpha. According to this hypothesis, the fused, transparent eyelids of snakes are thought to have evolved to combat marine conditions (corneal water loss through osmosis), and the external ears were lost through disuse in an aquatic environment. This ultimately led to an animal similar to today's sea snakes. In the Late Cretaceous, snakes recolonized land, and continued to diversify into today's snakes. Fossilized snake remains are known from early Late Cretaceous marine sediments, which is consistent with this hypothesis; particularly so, as they are older than the terrestrial Najash rionegrina. Similar skull structure, reduced or absent limbs, and other anatomical features found in both mosasaurs and snakes lead to a positive cladistical correlation, although some of these features are shared with varanids.

 

Genetic studies in recent years have indicated snakes are not as closely related to monitor lizards as was once believed—and therefore not to mosasaurs, the proposed ancestor in the aquatic scenario of their evolution. However, more evidence links mosasaurs to snakes than to varanids. Fragmented remains found from the Jurassic and Early Cretaceous indicate deeper fossil records for these groups, which may potentially refute either hypothesis.

 

Genetic basis of snake evolution

Main article: Limb development

Both fossils and phylogenetic studies demonstrate that snakes evolved from lizards, hence the question became which genetic changes led to limb loss in the snake ancestor. Limb loss is actually very common in extant reptiles and has happened dozens of times within skinks, anguids, and other lizards.

 

In 2016, two studies reported that limb loss in snakes is associated with DNA mutations in the Zone of Polarizing Activity Regulatory Sequence (ZRS), a regulatory region of the sonic hedgehog gene which is critically required for limb development. More advanced snakes have no remnants of limbs, but basal snakes such as pythons and boas do have traces of highly reduced, vestigial hind limbs. Python embryos even have fully developed hind limb buds, but their later development is stopped by the DNA mutations in the ZRS.

 

Distribution

There are about 3,900 species of snakes, ranging as far northward as the Arctic Circle in Scandinavia and southward through Australia. Snakes can be found on every continent except Antarctica, as well as in the sea, and as high as 16,000 feet (4,900 m) in the Himalayan Mountains of Asia. There are numerous islands from which snakes are absent, such as Ireland, Iceland, and New Zealand (although New Zealand's northern waters are infrequently visited by the yellow-bellied sea snake and the banded sea krait).

 

Taxonomy

All modern snakes are grouped within the suborder Serpentes in Linnean taxonomy, part of the order Squamata, though their precise placement within squamates remains controversial.

 

The two infraorders of Serpentes are Alethinophidia and Scolecophidia. This separation is based on morphological characteristics and mitochondrial DNA sequence similarity. Alethinophidia is sometimes split into Henophidia and Caenophidia, with the latter consisting of "colubroid" snakes (colubrids, vipers, elapids, hydrophiids, and atractaspids) and acrochordids, while the other alethinophidian families comprise Henophidia. While not extant today, the Madtsoiidae, a family of giant, primitive, python-like snakes, was around until 50,000 years ago in Australia, represented by genera such as Wonambi.

 

There are numerous debates in the systematics within the group. For instance, many sources classify Boidae and Pythonidae as one family, while some keep the Elapidae and Hydrophiidae (sea snakes) separate for practical reasons despite their extremely close relation.

 

Recent molecular studies support the monophyly of the clades of modern snakes, scolecophidians, typhlopids + anomalepidids, alethinophidians, core alethinophidians, uropeltids (Cylindrophis, Anomochilus, uropeltines), macrostomatans, booids, boids, pythonids and caenophidians.

 

Legless lizards

Main article: Legless lizard

While snakes are limbless reptiles, evolved from (and grouped with) lizards, there are many other species of lizards that have lost their limbs independently but which superficially look similar to snakes. These include the slowworm and glass snake.

 

Other serpentine tetrapods that are unrelated to snakes include caecilians (amphibians), amphisbaenians (near-lizard squamates), and the extinct aistopods (amphibians).

 

Biology

The now extinct Titanoboa cerrejonensis was 12.8 m (42 ft) in length. By comparison, the largest extant snakes are the reticulated python, measuring about 6.95 m (22.8 ft) long, and the green anaconda, which measures about 5.21 m (17.1 ft) long and is considered the heaviest snake on Earth at 97.5 kg (215 lb).

 

At the other end of the scale, the smallest extant snake is Leptotyphlops carlae, with a length of about 10.4 cm (4.1 in). Most snakes are fairly small animals, approximately 1 m (3.3 ft) in length.

 

Perception

Pit vipers, pythons, and some boas have infrared-sensitive receptors in deep grooves on the snout, allowing them to "see" the radiated heat of warm-blooded prey. In pit vipers, the grooves are located between the nostril and the eye in a large "pit" on each side of the head. Other infrared-sensitive snakes have multiple, smaller labial pits lining the upper lip, just below the nostrils.

 

A snake tracks its prey using smell, collecting airborne particles with its forked tongue, then passing them to the vomeronasal organ or Jacobson's organ in the mouth for examination. The fork in the tongue provides a sort of directional sense of smell and taste simultaneously. The snake's tongue is constantly in motion, sampling particles from the air, ground, and water, analyzing the chemicals found, and determining the presence of prey or predators in the local environment. In water-dwelling snakes, such as the anaconda, the tongue functions efficiently underwater.

 

The underside of a snake is very sensitive to vibration, allowing the snake to detect approaching animals by sensing faint vibrations in the ground. Despite the lack of outer ears, they are also able to detect airborne sounds.

 

Snake vision varies greatly between species. Some have keen eyesight and others are only able to distinguish light from dark, but the important trend is that a snake's visual perception is adequate enough to track movements. Generally, vision is best in tree-dwelling snakes and weakest in burrowing snakes. Some have binocular vision, where both eyes are capable of focusing on the same point, an example of this being the Asian vine snake. Most snakes focus by moving the lens back and forth in relation to the retina. Diurnal snakes have round pupils and many nocturnal snakes have slit pupils. Most species possess three visual pigments and are probably able to see two primary colors in daylight. The annulated sea snake and the genus Helicops appears to have regained much of their color vision as an adaption to the marine environment they live in. It has been concluded that the last common ancestors of all snakes had UV-sensitive vision, but most snakes that depend on their eyesight to hunt in daylight have evolved lenses that act like sunglasses for filtering out the UV-light, which probably also sharpens their vision by improving the contrast.

 

Skin

The skin of a snake is covered in scales. Contrary to the popular notion of snakes being slimy (because of possible confusion of snakes with worms), snakeskin has a smooth, dry texture. Most snakes use specialized belly scales to travel, allowing them to grip surfaces. The body scales may be smooth, keeled, or granular. The eyelids of a snake are transparent "spectacle" scales, also known as brille, which remain permanently closed.

 

The shedding of scales is called ecdysis (or in normal usage, molting or sloughing). Snakes shed the complete outer layer of skin in one piece. Snake scales are not discrete, but extensions of the epidermis—hence they are not shed separately but as a complete outer layer during each molt, akin to a sock being turned inside out.

 

Snakes have a wide diversity of skin coloration patterns which are often related to behavior, such as the tendency to have to flee from predators. Snakes that are at a high risk of predation tend to be plain, or have longitudinal stripes, providing few reference points to predators, thus allowing the snake to escape without being noticed. Plain snakes usually adopt active hunting strategies, as their pattern allows them to send little information to prey about motion. Blotched snakes usually use ambush-based strategies, likely because it helps them blend into an environment with irregularly shaped objects, like sticks or rocks. Spotted patterning can similarly help snakes to blend into their environment.

 

The shape and number of scales on the head, back, and belly are often characteristic and used for taxonomic purposes. Scales are named mainly according to their positions on the body. In "advanced" (Caenophidian) snakes, the broad belly scales and rows of dorsal scales correspond to the vertebrae, allowing these to be counted without the need for dissection.

 

Molting

Molting (or "ecdysis") serves a number of purposes. It allows old, worn skin to be replaced and it can remove parasites such as mites and ticks that live in the skin. It has also been observed in snakes that molting can be synced to mating cycles. Shedding skin can release pheromones and revitalize color and patterns of the skin to increase attraction of mates. Renewal of the skin by molting supposedly allows growth in some animals such as insects, but this has been disputed in the case of snakes.

 

Molting occurs periodically throughout the life of a snake. Before each molt, the snake stops eating and often hides or moves to a safe place. Just before shedding, the skin becomes dull and dry looking and the snake's eyes turn cloudy or blue-colored. The inner surface of the old skin liquefies, causing it to separate from the new skin beneath it. After a few days, the eyes become clear and the snake "crawls" out of its old skin, which splits close to the snake's mouth. The snake rubs its body against rough surfaces to aid in the shedding of its old skin. In many cases, the cast skin peels backward over the body from head to tail in one piece, like pulling a sock off inside-out, revealing a new, larger, brighter layer of skin which has formed underneath.

 

A young snake that is still growing may shed its skin up to four times a year, but an older snake may shed only once or twice a year. The discarded skin carries a perfect imprint of the scale pattern, so it is usually possible to identify the snake from the cast skin if it is reasonably intact. This periodic renewal has led to the snake being a symbol of healing and medicine, as pictured in the Rod of Asclepius.

 

Scale counts can sometimes be used to identify the sex of a snake when the species is not distinctly sexually dimorphic. A probe is fully inserted into the cloaca, marked at the point where it stops, then removed and measured against the subcaudal scales. The scalation count determines whether the snake is a male or female, as the hemipenes of a male will probe to a different depth (usually longer) than the cloaca of a female.

 

Skeleton

The skeletons of snakes are radically different from those of most other reptiles (as compared with the turtle here, for example), consisting almost entirely of an extended ribcage.

The skeleton of most snakes consists solely of the skull, hyoid, vertebral column, and ribs, though henophidian snakes retain vestiges of the pelvis and rear limbs.

 

The skull consists of a solid and complete neurocranium, to which many of the other bones are only loosely attached, particularly the highly mobile jaw bones, which facilitate manipulation and ingestion of large prey items. The left and right sides of the lower jaw are joined only by a flexible ligament at the anterior tips, allowing them to separate widely, and the posterior end of the lower jaw bones articulate with a quadrate bone, allowing further mobility. The mandible and quadrate bones can pick up ground-borne vibrations; because the sides of the lower jaw can move independently of one another, a snake resting its jaw on a surface has sensitive stereo auditory perception, used for detecting the position of prey. The jaw–quadrate–stapes pathway is capable of detecting vibrations on the angstrom scale, despite the absence of an outer ear and the lack of an impedance matching mechanism—provided by the ossicles in other vertebrates—for receiving vibrations from the air.

 

The hyoid is a small bone located posterior and ventral to the skull, in the 'neck' region, which serves as an attachment for the muscles of the snake's tongue, as it does in all other tetrapods.

 

The vertebral column consists of between 200 and 400 vertebrae, or sometimes more. The body vertebrae each have two ribs articulating with them. The tail vertebrae are comparatively few in number (often less than 20% of the total) and lack ribs. The vertebrae have projections that allow for strong muscle attachment, enabling locomotion without limbs.

 

Caudal autotomy (self-amputation of the tail), a feature found in some lizards, is absent in most snakes. In the rare cases where it does exist in snakes, caudal autotomy is intervertebral (meaning the separation of adjacent vertebrae), unlike that in lizards, which is intravertebral, i.e. the break happens along a predefined fracture plane present on a vertebra.

 

In some snakes, most notably boas and pythons, there are vestiges of the hindlimbs in the form of a pair of pelvic spurs. These small, claw-like protrusions on each side of the cloaca are the external portion of the vestigial hindlimb skeleton, which includes the remains of an ilium and femur.

 

Snakes are polyphyodonts with teeth that are continuously replaced

 

Snakes and other non-archosaur (crocodilians, dinosaurs + birds and allies) reptiles have a three-chambered heart that controls the circulatory system via the left and right atrium, and one ventricle. Internally, the ventricle is divided into three interconnected cavities: the cavum arteriosum, the cavum pulmonale, and the cavum venosum. The cavum venosum receives deoxygenated blood from the right atrium and the cavum arteriosum receives oxygenated blood from the left atrium. Located beneath the cavum venosum is the cavum pulmonale, which pumps blood to the pulmonary trunk.

 

The snake's heart is encased in a sac, called the pericardium, located at the bifurcation of the bronchi. The heart is able to move around, owing to the lack of a diaphragm; this adjustment protects the heart from potential damage when large ingested prey is passed through the esophagus. The spleen is attached to the gall bladder and pancreas and filters the blood. The thymus, located in fatty tissue above the heart, is responsible for the generation of immune cells in the blood. The cardiovascular system of snakes is unique for the presence of a renal portal system in which the blood from the snake's tail passes through the kidneys before returning to the heart.

 

The vestigial left lung is often small or sometimes even absent, as snakes' tubular bodies require all of their organs to be long and thin.[71] In the majority of species, only one lung is functional. This lung contains a vascularized anterior portion and a posterior portion that does not function in gas exchange. This 'saccular lung' is used for hydrostatic purposes to adjust buoyancy in some aquatic snakes and its function remains unknown in terrestrial species. Many organs that are paired, such as kidneys or reproductive organs, are staggered within the body, one located ahead of the other.

 

Snakes have no lymph nodes.

 

Venom

Cobras, vipers, and closely related species use venom to immobilize, injure, or kill their prey. The venom is modified saliva, delivered through fangs. The fangs of 'advanced' venomous snakes like viperids and elapids are hollow, allowing venom to be injected more effectively, and the fangs of rear-fanged snakes such as the boomslang simply have a groove on the posterior edge to channel venom into the wound. Snake venoms are often prey-specific, and their role in self-defense is secondary.

 

Venom, like all salivary secretions, is a predigestant that initiates the breakdown of food into soluble compounds, facilitating proper digestion. Even nonvenomous snakebites (like any animal bite) cause tissue damage.

 

Certain birds, mammals, and other snakes (such as kingsnakes) that prey on venomous snakes have developed resistance and even immunity to certain venoms.Venomous snakes include three families of snakes, and do not constitute a formal taxonomic classification group.

 

The colloquial term "poisonous snake" is generally an incorrect label for snakes. A poison is inhaled or ingested, whereas venom produced by snakes is injected into its victim via fangs. There are, however, two exceptions: Rhabdophis sequesters toxins from the toads it eats, then secretes them from nuchal glands to ward off predators; and a small unusual population of garter snakes in the US state of Oregon retains enough toxins in their livers from ingested newts to be effectively poisonous to small local predators (such as crows and foxes).

 

Snake venoms are complex mixtures of proteins, and are stored in venom glands at the back of the head. In all venomous snakes, these glands open through ducts into grooved or hollow teeth in the upper jaw. The proteins can potentially be a mix of neurotoxins (which attack the nervous system), hemotoxins (which attack the circulatory system), cytotoxins (which attack the cells directly), bungarotoxins (related to neurotoxins, but also directly affect muscle tissue), and many other toxins that affect the body in different ways. Almost all snake venom contains hyaluronidase, an enzyme that ensures rapid diffusion of the venom.

 

Venomous snakes that use hemotoxins usually have fangs in the front of their mouths, making it easier for them to inject the venom into their victims. Some snakes that use neurotoxins (such as the mangrove snake) have fangs in the back of their mouths, with the fangs curled backwards. This makes it difficult both for the snake to use its venom and for scientists to milk them. Elapids, however, such as cobras and kraits are proteroglyphous—they possess hollow fangs that cannot be erected toward the front of their mouths, and cannot "stab" like a viper. They must actually bite the victim.

 

It has been suggested that all snakes may be venomous to a certain degree, with harmless snakes having weak venom and no fangs. According to this theory, most snakes that are labelled "nonvenomous" would be considered harmless because they either lack a venom delivery method or are incapable of delivering enough to endanger a human. The theory postulates that snakes may have evolved from a common lizard ancestor that was venomous, and also that venomous lizards like the gila monster, beaded lizard, monitor lizards, and the now-extinct mosasaurs, may have derived from this same common ancestor. They share this "venom clade" with various other saurian species.

 

Venomous snakes are classified in two taxonomic families:

Elapids – cobras including king cobras, kraits, mambas, Australian copperheads, sea snakes, and coral snakes.

Viperids – vipers, rattlesnakes, copperheads/cottonmouths, and bushmasters.

There is a third family containing the opistoglyphous (rear-fanged) snakes (as well as the majority of other snake species):

 

Colubrids – boomslangs, tree snakes, vine snakes, cat snakes, although not all colubrids are venomous.

 

Reproduction

Although a wide range of reproductive modes are used by snakes, all employ internal fertilization. This is accomplished by means of paired, forked hemipenes, which are stored, inverted, in the male's tail. The hemipenes are often grooved, hooked, or spined—designed to grip the walls of the female's cloaca. The clitoris of the female snake consists of two structures located between the cloaca and the scent glands.

 

Most species of snakes lay eggs which they abandon shortly after laying. However, a few species (such as the king cobra) construct nests and stay in the vicinity of the hatchlings after incubation. Most pythons coil around their egg-clutches and remain with them until they hatch. A female python will not leave the eggs, except to occasionally bask in the sun or drink water. She will even "shiver" to generate heat to incubate the eggs.

 

Some species of snake are ovoviviparous and retain the eggs within their bodies until they are almost ready to hatch. Several species of snake, such as the boa constrictor and green anaconda, are fully viviparous, nourishing their young through a placenta as well as a yolk sac; this is highly unusual among reptiles, and normally found in requiem sharks or placental mammals. Retention of eggs and live birth are most often associated with colder environments.

 

Sexual selection in snakes is demonstrated by the 3,000 species that each use different tactics in acquiring mates. Ritual combat between males for the females they want to mate with includes topping, a behavior exhibited by most viperids in which one male will twist around the vertically elevated fore body of its opponent and force it downward. It is common for neck-biting to occur while the snakes are entwined.

 

Facultative parthenogenesis

Parthenogenesis is a natural form of reproduction in which growth and development of embryos occur without fertilization. Agkistrodon contortrix (copperhead) and Agkistrodon piscivorus (cottonmouth) can reproduce by facultative parthenogenesis, meaning that they are capable of switching from a sexual mode of reproduction to an asexual mode. The most likely type of parthenogenesis to occur is automixis with terminal fusion, a process in which two terminal products from the same meiosis fuse to form a diploid zygote. This process leads to genome-wide homozygosity, expression of deleterious recessive alleles, and often to developmental abnormalities. Both captive-born and wild-born copperheads and cottonmouths appear to be capable of this form of parthenogenesis.

 

Reproduction in squamate reptiles is almost exclusively sexual. Males ordinarily have a ZZ pair of sex-determining chromosomes, and females a ZW pair. However, the Colombian Rainbow boa (Epicrates maurus) can also reproduce by facultative parthenogenesis, resulting in production of WW female progeny. The WW females are likely produced by terminal automixis.

 

Embryonic Development

Snake embryonic development initially follows similar steps as any vertebrate embryo. The snake embryo begins as a zygote, undergoes rapid cell division, forms a germinal disc, also called a blastodisc, then undergoes gastrulation, neurulation, and organogenesis. Cell division and proliferation continues until an early snake embryo develops and the typical body shape of a snake can be observed. Multiple features differentiate the embryologic development of snakes from other vertebrates, two significant factors being the elongation of the body and the lack of limb development.

 

The elongation in snake body is accompanied by a significant increase in vertebra count (mice have 60 vertebrae, whereas snakes may have over 300). This increase in vertebrae is due to an increase in somites during embryogenesis, leading to an increased number of vertebrae which develop. Somites are formed at the presomitic mesoderm due to a set of oscillatory genes that direct the somitogenesis clock. The snake somitogenesis clock operates at a frequency 4 times that of a mouse (after correction for developmental time), creating more somites, and therefore creating more vertebrae. This difference in clock speed is believed to be caused by differences in Lunatic fringe gene expression, a gene involved in the somitogenesis clock.

 

There is ample literature focusing on the limb development/lack of development in snake embryos and the gene expression associated with the different stages. In basal snakes, such as the python, embryos in early development exhibit a hind limb bud that develops with some cartilage and a cartilaginous pelvic element, however this degenerates before hatching. This presence of vestigial development suggests that some snakes are still undergoing hind limb reduction before they are eliminated. There is no evidence in basal snakes of forelimb rudiments and no examples of snake forelimb bud initiation in embryo, so little is known regarding the loss of this trait. Recent studies suggests that hind limb reduction could be due to mutations in enhancers for the SSH gene, however other studies suggested that mutations within the Hox Genes or their enhancers could contribute to snake limblessness. Since multiple studies have found evidence suggesting different genes played a role in the loss of limbs in snakes, it is likely that multiple gene mutations had an additive effect leading to limb loss in snakes.

 

Behavior

Snake coiled on a stick in Oklahoma. It was brumating in a large pile of wood chips, found by this landscaper after he bulldozed the pile in late autumn 2018.

In regions where winters are too cold for snakes to tolerate while remaining active, local species will enter a period of brumation. Unlike hibernation, in which the dormant mammals are actually asleep, brumating reptiles are awake but inactive. Individual snakes may brumate in burrows, under rock piles, or inside fallen trees, or large numbers of snakes may clump together in hibernacula.

 

Feeding and diet

All snakes are strictly carnivorous, preying on small animals including lizards, frogs, other snakes, small mammals, birds, eggs, fish, snails, worms, and insects. Snakes cannot bite or tear their food to pieces so must swallow their prey whole. The eating habits of a snake are largely influenced by body size; smaller snakes eat smaller prey. Juvenile pythons might start out feeding on lizards or mice and graduate to small deer or antelope as an adult, for example.

 

The snake's jaw is a complex structure. Contrary to the popular belief that snakes can dislocate their jaws, they have an extremely flexible lower jaw, the two halves of which are not rigidly attached, and numerous other joints in the skull, which allow the snake to open its mouth wide enough to swallow prey whole, even if it is larger in diameter than the snake itself. For example, the African egg-eating snake has flexible jaws adapted for eating eggs much larger than the diameter of its head.  This snake has no teeth, but does have bony protrusions on the inside edge of its spine, which it uses to break the shell when eating eggs.

 

The majority of snakes eat a variety of prey animals, but there is some specialization in certain species. King cobras and the Australian bandy-bandy consume other snakes. Species of the family Pareidae have more teeth on the right side of their mouths than on the left, as they mostly prey on snails and the shells usually spiral clockwise.

 

Some snakes have a venomous bite, which they use to kill their prey before eating it. Other snakes kill their prey by constriction, while some swallow their prey when it is still alive.

 

After eating, snakes become dormant to allow the process of digestion to take place; this is an intense activity, especially after consumption of large prey. In species that feed only sporadically, the entire intestine enters a reduced state between meals to conserve energy. The digestive system is then 'up-regulated' to full capacity within 48 hours of prey consumption. Being ectothermic ("cold-blooded"), the surrounding temperature plays an important role in the digestion process. The ideal temperature for snakes to digest food is 30 °C (86 °F). There is a huge amount of metabolic energy involved in a snake's digestion, for example the surface body temperature of the South American rattlesnake (Crotalus durissus) increases by as much as 1.2 °C (2.2 °F) during the digestive process. If a snake is disturbed after having eaten recently, it will often regurgitate its prey to be able to escape the perceived threat. When undisturbed, the digestive process is highly efficient; the snake's digestive enzymes dissolve and absorb everything but the prey's hair (or feathers) and claws, which are excreted along with waste.

 

Hooding and spitting

Hooding (expansion of the neck area) is a visual deterrent, mostly seen in cobras (elapids), and is primarily controlled by rib muscles.[98] Hooding can be accompanied by spitting venom towards the threatening object,[99] and producing a specialized sound; hissing. Studies on captive cobras showed that 13 to 22% of the body length is raised during hooding.

 

Locomotion

The lack of limbs does not impede the movement of snakes. They have developed several different modes of locomotion to deal with particular environments. Unlike the gaits of limbed animals, which form a continuum, each mode of snake locomotion is discrete and distinct from the others; transitions between modes are abrupt.

 

Lateral undulation

Lateral undulation is the sole mode of aquatic locomotion, and the most common mode of terrestrial locomotion In this mode, the body of the snake alternately flexes to the left and right, resulting in a series of rearward-moving "waves". While this movement appears rapid, snakes have rarely been documented moving faster than two body-lengths per second, often much less. This mode of movement has the same net cost of transport (calories burned per meter moved) as running in lizards of the same mass.

 

Terrestrial lateral undulation is the most common mode of terrestrial locomotion for most snake species. In this mode, the posteriorly moving waves push against contact points in the environment, such as rocks, twigs, irregularities in the soil, etc. Each of these environmental objects, in turn, generates a reaction force directed forward and towards the midline of the snake, resulting in forward thrust while the lateral components cancel out. The speed of this movement depends upon the density of push-points in the environment, with a medium density of about 8[clarification needed] along the snake's length being ideal. The wave speed is precisely the same as the snake speed, and as a result, every point on the snake's body follows the path of the point ahead of it, allowing snakes to move through very dense vegetation and small openings.

 

When swimming, the waves become larger as they move down the snake's body, and the wave travels backwards faster than the snake moves forwards. Thrust is generated by pushing their body against the water, resulting in the observed slip. In spite of overall similarities, studies show that the pattern of muscle activation is different in aquatic versus terrestrial lateral undulation, which justifies calling them separate modes. All snakes can laterally undulate forward (with backward-moving waves), but only sea snakes have been observed reversing the motion (moving backwards with forward-moving waves).

 

Sidewinding

Most often employed by colubroid snakes (colubrids, elapids, and vipers) when the snake must move in an environment that lacks irregularities to push against (rendering lateral undulation impossible), such as a slick mud flat, or a sand dune, sidewinding is a modified form of lateral undulation in which all of the body segments oriented in one direction remain in contact with the ground, while the other segments are lifted up, resulting in a peculiar "rolling" motion. This mode of locomotion overcomes the slippery nature of sand or mud by pushing off with only static portions on the body, thereby minimizing slipping. The static nature of the contact points can be shown from the tracks of a sidewinding snake, which show each belly scale imprint, without any smearing. This mode of locomotion has very low caloric cost, less than 1⁄3 of the cost for a lizard to move the same distance. Contrary to popular belief, there is no evidence that sidewinding is associated with the sand being hot.

 

Concertina

When push-points are absent, but there is not enough space to use sidewinding because of lateral constraints, such as in tunnels, snakes rely on concertina locomotion. In this mode, the snake braces the posterior portion of its body against the tunnel wall while the front of the snake extends and straightens. The front portion then flexes and forms an anchor point, and the posterior is straightened and pulled forwards. This mode of locomotion is slow and very demanding, up to seven times the cost of laterally undulating over the same distance. This high cost is due to the repeated stops and starts of portions of the body as well as the necessity of using active muscular effort to brace against the tunnel walls.

 

Arboreal

The movement of snakes in arboreal habitats has only recently been studied. While on tree branches, snakes use several modes of locomotion depending on species and bark texture. In general, snakes will use a modified form of concertina locomotion on smooth branches, but will laterally undulate if contact points are available. Snakes move faster on small branches and when contact points are present, in contrast to limbed animals, which do better on large branches with little 'clutter'.

 

Gliding snakes (Chrysopelea) of Southeast Asia launch themselves from branch tips, spreading their ribs and laterally undulating as they glide between trees. These snakes can perform a controlled glide for hundreds of feet depending upon launch altitude and can even turn in midair.

 

Rectilinear

The slowest mode of snake locomotion is rectilinear locomotion, which is also the only one where the snake does not need to bend its body laterally, though it may do so when turning. In this mode, the belly scales are lifted and pulled forward before being placed down and the body pulled over them. Waves of movement and stasis pass posteriorly, resulting in a series of ripples in the skin. The ribs of the snake do not move in this mode of locomotion and this method is most often used by large pythons, boas, and vipers when stalking prey across open ground as the snake's movements are subtle and harder to detect by their prey in this manner.

 

Interactions with humans

Snakes do not ordinarily prey on humans. Unless startled or injured, most snakes prefer to avoid contact and will not attack humans. With the exception of large constrictors, nonvenomous snakes are not a threat to humans. The bite of a nonvenomous snake is usually harmless; their teeth are not adapted for tearing or inflicting a deep puncture wound, but rather grabbing and holding. Although the possibility of infection and tissue damage is present in the bite of a nonvenomous snake, venomous snakes present far greater hazard to humans.  The World Health Organization (WHO) lists snakebite under the "other neglected conditions" category.

 

Documented deaths resulting from snake bites are uncommon. Nonfatal bites from venomous snakes may result in the need for amputation of a limb or part thereof. Of the roughly 725 species of venomous snakes worldwide, only 250 are able to kill a human with one bite. Australia averages only one fatal snake bite per year. In India, 250,000 snakebites are recorded in a single year, with as many as 50,000 recorded initial deaths. The WHO estimates that on the order of 100,000 people die each year as a result of snake bites, and around three times as many amputations and other permanent disabilities are caused by snakebites annually.

 

The treatment for a snakebite is as variable as the bite itself. The most common and effective method is through antivenom (or antivenin), a serum made from the venom of the snake. Some antivenom is species-specific (monovalent) while some is made for use with multiple species in mind (polyvalent). In the United States for example, all species of venomous snakes are pit vipers, with the exception of the coral snake. To produce antivenom, a mixture of the venoms of the different species of rattlesnakes, copperheads, and cottonmouths is injected into the body of a horse in ever-increasing dosages until the horse is immunized. Blood is then extracted from the immunized horse. The serum is separated and further purified and freeze-dried. It is reconstituted with sterile water and becomes antivenom. For this reason, people who are allergic to horses are more likely to have an allergic reaction to antivenom. Antivenom for the more dangerous species (such as mambas, taipans, and cobras) is made in a similar manner in South Africa, Australia , and India, although these antivenoms are species-specific.

 

Snake charmers

In some parts of the world, especially in India, snake charming is a roadside show performed by a charmer. In such a show, the snake charmer carries a basket containing a snake that he seemingly charms by playing tunes with his flutelike musical instrument, to which the snake responds. The snake is in fact responding to the movement of the flute, not the sound it makes, as snakes lack external ears (though they do have internal ears).

 

The Wildlife Protection Act of 1972 in India technically prohibits snake charming on the grounds of reducing animal cruelty. Other types of snake charmers use a snake and mongoose show, where the two animals have a mock fight; however, this is not very common, as the animals may be seriously injured or killed. Snake charming as a profession is dying out in India because of competition from modern forms of entertainment and environment laws proscribing the practice. Many Indians have never seen snake charming and it is becoming a folktale of the past.

 

Trapping

The Irulas tribe of Andhra Pradesh and Tamil Nadu in India have been hunter-gatherers in the hot, dry plains forests, and have practiced the art of snake catching for generations. They have a vast knowledge of snakes in the field. They generally catch the snakes with the help of a simple stick. Earlier, the Irulas caught thousands of snakes for the snake-skin industry. After the complete ban of the snake-skin industry in India and protection of all snakes under the Indian Wildlife (Protection) Act 1972, they formed the Irula Snake Catcher's Cooperative and switched to catching snakes for removal of venom, releasing them in the wild after four extractions. The venom so collected is used for producing life-saving antivenom, biomedical research and for other medicinal products. The Irulas are also known to eat some of the snakes they catch and are very useful in rat extermination in the villages.

 

Despite the existence of snake charmers, there have also been professional snake catchers or wranglers. Modern-day snake trapping involves a herpetologist using a long stick with a V-shaped end. Some television show hosts, like Bill Haast, Austin Stevens, Steve Irwin, and Jeff Corwin, prefer to catch them using bare hands.

 

Consumption

Although snakes are not commonly thought of as food, their consumption is acceptable in some cultures and may even be considered a delicacy. Snake soup is popular in Cantonese cuisine, consumed by locals in the autumn to warm their bodies. Western cultures document the consumption of snakes only under extreme circumstances of hunger, with the exception of cooked rattlesnake meat, which is commonly consumed in Texas and parts of the Midwestern United States.

 

In Asian countries such as China, Taiwan, Thailand, Indonesia, Vietnam, and Cambodia, drinking the blood of a snake—particularly the cobra—is believed to increase sexual virility. When possible, the blood is drained while the cobra is still alive, and it is usually mixed with some form of liquor to improve the taste.

 

The use of snakes in alcohol is accepted in some Asian countries. In such cases, one or more snakes are left to steep in a jar or container of liquor, as this is claimed to make the liquor stronger (as well as more expensive). One example of this is the Habu snake, which is sometimes placed in the Okinawan liqueur Habushu (ハブ酒), also known as "Habu Sake".

 

Snake wine (蛇酒) is an alcoholic beverage produced by infusing whole snakes in rice wine or grain alcohol. First recorded as being consumed in China during the Western Zhou dynasty, this drink is considered an important curative and is believed to reinvigorate a person according to traditional Chinese medicine

 

Pets

In the Western world, some snakes are kept as pets, especially docile species such as the ball python and corn snake. To meet the demand, a captive breeding industry has developed. Snakes bred in captivity are considered preferable to specimens caught in the wild and tend to make better pets. Compared with more traditional types of companion animal, snakes can be very low-maintenance pets; they require minimal space, as most common species do not exceed 5 feet (1.5 m) in length, and can be fed relatively infrequently—usually once every five to 14 days. Certain snakes have a lifespan of more than 40 years if given proper care.

 

Symbolism

In ancient Mesopotamia, Nirah, the messenger god of Ištaran, was represented as a serpent on kudurrus, or boundary stones. Representations of two intertwined serpents are common in Sumerian art and Neo-Sumerian artwork and still appear sporadically on cylinder seals and amulets until as late as the thirteenth century BC. The horned viper (Cerastes cerastes) appears in Kassite and Neo-Assyrian kudurrus and is invoked in Assyrian texts as a magical protective entity. A dragon-like creature with horns, the body and neck of a snake, the forelegs of a lion, and the hind-legs of a bird appears in Mesopotamian art from the Akkadian Period until the Hellenistic Period (323 BC–31 BC). This creature, known in Akkadian as the mušḫuššu, meaning "furious serpent", was used as a symbol for particular deities and also as a general protective emblem. It seems to have originally been the attendant of the Underworld god Ninazu, but later became the attendant to the Hurrian storm-god Tishpak, as well as, later, Ninazu's son Ningishzida, the Babylonian national god Marduk, the scribal god Nabu, and the Assyrian national god Ashur.

 

In Egyptian history, the snake occupies a primary role with the Nile cobra adorning the crown of the pharaoh in ancient times. It was worshipped as one of the gods and was also used for sinister purposes: murder of an adversary and ritual suicide (Cleopatra). The ouroboros was a well-known ancient Egyptian symbol of a serpent swallowing its own tail. The precursor to the ouroboros was the "Many-Faced", a serpent with five heads, who, according to the Amduat, the oldest surviving Book of the Afterlife, was said to coil around the corpse of the sun god Ra protectively. The earliest surviving depiction of a "true" ouroboros comes from the gilded shrines in the tomb of Tutankhamun. In the early centuries AD, the ouroboros was adopted as a symbol by Gnostic Christians and chapter 136 of the Pistis Sophia, an early Gnostic text, describes "a great dragon whose tail is in its mouth". In medieval alchemy, the ouroboros became a typical western dragon with wings, legs, and a tail.

 

In the Bible, King Nahash of Ammon, whose name means "Snake", is depicted very negatively, as a particularly cruel and despicable enemy of the ancient Hebrews.

 

The ancient Greeks used the Gorgoneion, a depiction of a hideous face with serpents for hair, as an apotropaic symbol to ward off evil. In a Greek myth described by Pseudo-Apollodorus in his Bibliotheca, Medusa was a Gorgon with serpents for hair whose gaze turned all those who looked at her to stone and was slain by the hero Perseus. In the Roman poet Ovid's Metamorphoses, Medusa is said to have once been a beautiful priestess of Athena, whom Athena turned into a serpent-haired monster after she was raped by the god Poseidon in Athena's temple. In another myth referenced by the Boeotian poet Hesiod and described in detail by Pseudo-Apollodorus, the hero Heracles is said to have slain the Lernaean Hydra, a multiple-headed serpent which dwelt in the swamps of Lerna.

 

The legendary account of the foundation of Thebes mentioned a monster snake guarding the spring from which the new settlement was to draw its water. In fighting and killing the snake, the companions of the founder Cadmus all perished – leading to the term "Cadmean victory" (i.e. a victory involving one's own ruin).

 

Three medical symbols involving snakes that are still used today are Bowl of Hygieia, symbolizing pharmacy, and the Caduceus and Rod of Asclepius, which are symbols denoting medicine in general.

 

One of the etymologies proposed for the common female first name Linda is that it might derive from Old German Lindi or Linda, meaning a serpent.

 

India is often called the land of snakes and is steeped in tradition regarding snakes. Snakes are worshipped as gods even today with many women pouring milk on snake pits (despite snakes' aversion for milk). The cobra is seen on the neck of Shiva and Vishnu is depicted often as sleeping on a seven-headed snake or within the coils of a serpent. There are also several temples in India solely for cobras sometimes called Nagraj (King of Snakes) and it is believed that snakes are symbols of fertility. There is a Hindu festival called Nag Panchami each year on which day snakes are venerated and prayed to. See also Nāga.

 

In India there is another mythology about snakes. Commonly known in Hindi as "Ichchhadhari" snakes. Such snakes can take the form of any living creature, but prefer human form. These mythical snakes possess a valuable gem called "Mani", which is more brilliant than diamond. There are many stories in India about greedy people trying to possess this gem and ending up getting killed.

 

The snake is one of the 12 celestial animals of Chinese zodiac, in the Chinese calendar.

 

Many ancient Peruvian cultures worshipped nature. They emphasized animals and often depicted snakes in their art.

 

Religion

Snakes are used in Hinduism as a part of ritual worship. In the annual Nag Panchami festival, participants worship either live cobras or images of Nāgas. Lord Shiva is depicted in most images with a snake coiled around his neck. Puranic literature includes various stories associated with snakes, for example Shesha is said to hold all the planets of the Universe on his hoods and to constantly sing the glories of Vishnu from all his mouths. Other notable snakes in Hinduism are Vasuki, Takshaka, Karkotaka, and Pingala. The term Nāga is used to refer to entities that take the form of large snakes in Hinduism and Buddhism.

 

Snakes have been widely revered in many cultures, such as in ancient Greece where the serpent was seen as a healer.[148] Asclepius carried a serpent wound around his wand, a symbol seen today on many ambulances. In Judaism, the snake of brass is also a symbol of healing, of one's life being saved from imminent death.

 

In religious terms, the snake and jaguar were arguably the most important animals in ancient Mesoamerica. "In states of ecstasy, lords dance a serpent dance; great descending snakes adorn and support buildings from Chichen Itza to Tenochtitlan, and the Nahuatl word coatl meaning serpent or twin, forms part of primary deities such as Mixcoatl, Quetzalcoatl, and Coatlicue." In the Maya and Aztec calendars, the fifth day of the week was known as Snake Day.

 

In some parts of Christianity, the redemptive work of Jesus Christ is compared to saving one's life through beholding the Nehushtan (serpent of brass). Snake handlers use snakes as an integral part of church worship, to demonstrate their faith in divine protection. However, more commonly in Christianity, the serpent has been depicted as a representative of evil and sly plotting, as seen in the description in Genesis of a snake tempting Eve in the Garden of Eden. Saint Patrick is purported to have expelled all snakes from Ireland while converting the country to Christianity in the 5th century, thus explaining the absence of snakes there.

 

In Christianity and Judaism, the snake makes its infamous appearance in the first book of the Bible when a serpent appears before Adam and Eve and tempts them with the forbidden fruit from the Tree of Knowledge. The snake returns in the Book of Exodus when Moses turns his staff into a snake as a sign of God's power, and later when he makes the Nehushtan, a bronze snake on a pole that when looked at cured the people of bites from the snakes that plagued them in the desert. The serpent makes its final appearance symbolizing Satan in the Book of Revelation: "And he laid hold on the dragon the old serpent, which is the devil and Satan, and bound him for a thousand years."

 

In Neo-Paganism and Wicca, the snake is seen as a symbol of wisdom and knowledge. Additionally, snakes are sometimes associated with Hecate, the Greek goddess of witchcraft.

 

Medicine

Several compounds from snake venoms are being researched as potential treatments or preventatives for pain, cancers, arthritis, stroke, heart disease, hemophilia, and hypertension, and to control bleeding (e.g. during surgery).

Flowers from my dad's garden, taken during a visit on Father's Day.

Blue skies at times for today's Paddle Round the Pier event in Brighton - making everyone happy. One for this week's challenge of an unusual viewpoint

Songs include hits such as 'Excuse Me' - 'Under The Influence' - 'Act Naturally' & 'Loves Gonna Live Here' + loads more .

Games to play while railfanning include various forms of tossing ballast under the generic term of "Rocks" The rules have to do with tossing rocks between and hitting rails, but one thing had never happened before that a rock landed on top of the rail. So in true Calvin & Hobbes Calvinball a new rule was made at that time.

Includes both Leon and Claires GD-ROMs, along with the demo disc for Code: Veronica

i used to include the white squares when i started doing my "inspired by" nods ( which have become my friday faves ) but stopped because i felt happier sharing photos with my contacts, not just links.

 

but i am making an exception today.

i just think this looks nice,lol.

and it actually speaks to the intent of the photo that is NOT appearing in the center...so take a sec and look at it ( it is one of maite's ) but also appreciate the quietness it shares with all the other photos i have loved this week:)

 

1. [ trece ], 2. sister, sister, 3. progress report, 4. Life Is Beautiful, 5. Untitled, 6. outtake, in which i open the door and hide my giant hands and feet but remained unconvinced of what the hell i am going for here, 7. what did you learn from the time, 8. Mammoth, Yosemite, HOME, 9. Hey Baby, Come Have A Seat Beside Me On My New Couch, 10. Can you Colourize My Life, I'm So Sick of Black and White, 11. Mine, 12. "shall i pose for you?", 13. Untitled14. Not available15. Not available16. Not available

 

Created with fd's Flickr Toys

The church yard of St Mary de Haura on a warm, still, evening. I guess the phrase "pushing up the daisies" is very adapt for this one

modeling her new outfit which includes her "Colvin" mini skirt from Becky Colvin of SewCool99. Bolero/shrug knit with US size 2 circulars and double points using Schoeller Fortissima Socka yarn.

Fluffed up against the cold at Woods Mill

Ships in this shot include the USS Ramage and Normandy, HNLMS Van Amstel and De Zeven Provinciën, Álvaro de Bazán, and HDMS Peter Willemoes

 

It was a beautiful late autumn day for a walk (and some flights!) around Halifax Harbour. The city was buzzing with the arrival of the US' latest aircraft carrier (and first of its class), the Gerald R. Ford, two cruise ships, HalCon, and nice weather. In the midst of some errands and hanging out with friends, I sought vantage points to take in the fleet.

 

I got terrible pictures the last time a carrier was in port, so it was great to have a 150-600mm with me this time, as well as a better idea where to go for lighting. My drone also added a really fun dimension. I was a little apprehensive, as some people were being a bit irresponsible in getting too close to the carrier, but I checked all my flights on Transport Canada's NavDrone, and I tried to keep to what seemed like the exclusion zone around the carrier in particular. For the relatively close shots of the shipyard, I launched from the Casino and avoided sliding over the shipyard itself.

More from a wander around the paddock on Friday at the Goodwood Revival

A gala dinner to complete the foundation of the Australian Media Hall of Fame.

 

Another 48 greats of national journalism were inducted into the Australian Media Hall of Fame at the 2018 Melbourne dinner.

 

They include world-famous war cameramen Neil Davis and David Brill, TV legends Ray Martin and Geraldine Doogue, magazine queen Dulcie Boling, Pulitzer Prize-winning cartoonist Pat Oliphant and ground-breaking Aboriginal journalist John Newfong.

 

Also honoured was the celebrated former political editor of The Age and The Australian, Michael Gordon, a champion of social justice journalism who died suddenly in February this year.

 

The inductions were announced at a gala dinner in Melbourne that completed the national foundation of the Media Hall of Fame – a decade-long project of the Melbourne Press Club.

 

Launched in Victoria in 2012, the Media Hall of Fame last year expanded to include NSW. This year’s inductees include greats from Queensland, Western Australia, South Australia, Tasmania and the territories.

 

The Australian Media Hall of Fame seeks to demonstrate the importance and public benefit of journalism in the face of threats from digital disruption and political interference.

 

“The achievements of these extraordinary men and women shows the abiding importance of great journalism for our democracy and why we should celebrate and defend it,” Melbourne Press Club CEO Mark Baker said.

 

The Advisory Panel for this year’s nominations was former editor of The Age Michael Smith, former editor-in-chief of The Australian Chris Mitchell, media historian Bridget Griffen-Foley of Macquarie University and Mike Bowers, photographer-at-large for The Guardian.

 

This event was proudly supported by Visit Victoria.

Theme MY COUNTRY - which must include the novels of Jane Austen. We have had two very significant holidays here, so it is also a perfect jigsaw for the theme HOLIDAY DESTINATIONS.

 

June 2017:

It has been a while since I've uploaded any photos of jigsaws - I've been on holiday and have also been busy listing and packing them away. (We got back from Washington DC the night before the most recent BCD Meeting so couldn't go to that either - the theme was Map puzzles so perhaps I should tackle one or two map jigsaws soon to take to the next meeting.)

 

This beautiful painting of rainy pavements outside Bath Abbey by Irene Marsh is a lovely contrast to the heatwave in Washington and back home in England. The view is almost identical to that obtained from the window of a Landmark Trust property we stayed in with friends three years ago (Marshall Wade's House, on the left opposite the Pump Rooms).

 

www.landmarktrust.org.uk/search-and-book/properties/marsh...

 

For more examples of Irene's watercolours visit her website. To date Wentworth have issued jigsaws of Bath Abbey and The Circus II.

irenemarsh.co.uk/shop/

 

Irene says, "I want my work to show life as it is today, right down to the last puddle. I love the reflections you see on traffic, paintwork and windows but most of all I love old, wet paving stones.”

 

Lots of bikes in the side streets of the old city of Lucca, Tuscany

Includes Hardee's

From Friday night's gig at Sticky Mike's Frog Bar in Brighton, where Deaf from Behind were headlining for the first time.

 

Deaf from Behind and friends after the gig www.reverbnation.com/deaffrombehind

Spring Lambs by the River Adur this evening.

Discover fun facts about elephants in Thailand, care and feeding them with love! Feeding, mud SPA and shower with the elephants, make paper from elephant poop, take funny photos with elephants, the price includes lunch, free photographs, free transfer...

Details and reservation online: thai-online.tours

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Read in Russian language: thai-online.org/

Around the world excursions and guided tours: www.7stars-tours.com. Use the link to search best deals and online reservations with the lowest prices!

 

ALL THINGS TO DO IN PATTAYA

 

All the best, newest, popular and not expensive excursions in Pattaya - on our THAI-ONLINE website. Can read and download the price with all of our proposals.

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Page from Keighley and the Bronte Country Official Handbook, (Ed. J. Burrow & Co. Ltd., 1960). This page includes an advert for Precimax Precision Machine Tools, manufactured by Landis Lund Ltd., of Cross Hills.

 

The handbook was published for the Corporation of Keighley and served as an official guide to the town, its history, and the surrounding area, and as an industrial directory, listing and featuring adverts from many of the key businesses in the area. The book measures approximately 190 mm by 245 mm and runs to 76 pages. It came with a pull-out street plan of the town.

 

The handbook belongs to Joyce Newton and was loaned to the Keighley and District Local History Society for scanning in February 2020.

Taman Mini Indonesia Indah includes an open-air ethnographic museum showing the various regional architecture styles from throughout Indonesia, along with cultural performances and regional foods and handicrafts. It also includes series of topical museums (military, stamps, telecommunications, etc.), an aquarium, an aviary, and a museum dedicated to the fauna of Komodo, among other attractions.

A blog post that includes these photos lives here: likeafishinwater.com/2017/04/01/pilgrimage-to-tokyo-for-k...

 

My company: www.thirdplacemedia.com - Research, content development and communications strategy focused on transit, walkability, placemaking and environment issues

 

My blog: likeafishinwater.com

Some of the early finishers in this morning Sussex Classic - Ocean Surfski race, off the beach at Lancing. The water looked almost tropical. This veteran entrant was the clear winner

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A classic Ford GT40 down the pit straight during practice for the Alan Mann Trophy at the 74th Members' Meeting, Goodwood

Hindu deities are the gods and goddesses in Hinduism. The terms and epithets found in Indian culture, that are translated as deity, varies with the text and diverse traditions within Hinduism, and include Deva, Devi, Ishvara, Bhagavan and Bhagavathi.[1][2][note 1]

 

The deities of Hinduism have evolved from Vedic era (2nd millennium BCE) through medieval era (1st millennium CE), regionally within India and in southeast Asia, and across Hinduism's diverse traditions.[3][4] The Hindu deity concept varies from a personal god as in Yoga school of Hindu philosophy,[5][6] to 33 Vedic deities,[7] to hundreds of Puranic deities, to millions of deities in Tantra traditions of Hinduism.[8] Illustrations of major deities include Vishnu, Sri (Lakshmi), Shiva, Parvati (Durga), Brahma and Saraswati. These deities have distinct and complex personalities, yet often viewed as aspects of the same Ultimate Reality called Brahman.[9][note 2] From ancient times, the idea of equivalence has been cherished in Hinduism, in its texts and in early 1st millennium sculpture with concepts such as Harihara (half Shiva, half Vishnu),[10] Ardhanarishvara (half Shiva, half Parvati) or Vaikuntha Kamalaja (half Vishnu, half Lakshmi),[11] with mythologies and temples that feature them together, declaring they are the same.[12][13][14] Major deities have inspired their own Hindu traditions, such as Vaishnavism, Shaivism and Shaktism, but with shared mythology, ritual grammar, theosophy, axiology and polycentrism.[15][16][17] Some Hindu traditions such as Smartism from mid 1st millennium CE, have included multiple major deities as henotheistic manifestations of Saguna Brahman, and as a means to realizing Nirguna Brahman.[18][19][20]

 

Hindu deities are represented with various icons and anicons, in paintings and sculptures, called Murtis and Pratimas.[21][22][23] Some Hindu traditions, such as ancient Charvakas rejected all deities and concept of god or goddess,[24][25][26] while 19th-century British colonial era movements such as the Arya Samaj and Brahmo Samaj rejected deities and adopted monotheistic concepts similar to Abrahamic religions.[27][28] Hindu deities have been adopted in other religions such as Jainism,[29] and in regions outside India such as predominantly Buddhist Thailand and Japan where they continue to be revered in regional temples or arts.[30][31][32]

 

In ancient and medieval era texts of Hinduism, the human body is described as a temple,[33][34] and deities are described to be parts residing within it,[35][36] while the Brahman (Absolute Reality, God)[18][37] is described to be the same, or of similar nature, as the Atman (self, soul), which Hindus believe is eternal and within every living being.[38][39][40] Deities in Hinduism are as diverse as its traditions, and a Hindu can choose to be polytheistic, pantheistic, monotheistic, monistic, agnostic, atheistic or humanist.[41][42][43]

Deities in Hinduism are referred to as Deva (masculine) and Devi (feminine).[44][45][46] The root of these terms mean "heavenly, divine, anything of excellence".[47] According to Douglas Harper, the etymological roots of Deva mean "a shining one," from *div- "to shine," and it is a cognate with Greek dios "divine" and Zeus, and Latin deus (Old Latin deivos).[48]

 

In the earliest Vedic literature, all supernatural beings are called Asuras.[49][50] By the late Vedic period (~500 BCE), benevolent supernatural beings are referred to as Deva-Asuras. In post-Vedic texts, such as the Puranas and the Itihasas of Hinduism, the Devas represent the good, and the Asuras the bad.[3][4] In some medieval Indian literature, Devas are also referred to as Suras and contrasted with their equally powerful, but malevolent half-brothers referred to as the Asuras.[51]

 

Hindu deities are part of Indian mythology, both Devas and Devis feature in one of many cosmological theories in Hinduism.[52][53]

 

Characteristics of Vedic era deities[edit]

In Vedic literature, Devas and Devis represent the forces of nature and some represent moral values (such as the Adityas, Varuna, and Mitra), each symbolizing the epitome of a specialized knowledge, creative energy, exalted and magical powers (Siddhis).[54][55]

The most referred to Devas in the Rig Veda are Indra, Agni (fire) and Soma, with "fire deity" called the friend of all humanity, it and Soma being the two celebrated in a yajna fire ritual that marks major Hindu ceremonies. Savitr, Vishnu, Rudra (later given the exclusive epithet of Shiva), and Prajapati (later Brahma) are gods and hence Devas.[30]

 

The Vedas describes a number of significant Devis such as Ushas (dawn), Prithvi (earth), Aditi (cosmic moral order), Saraswati (river, knowledge), Vāc (sound), Nirṛti (destruction), Ratri (night), Aranyani (forest), and bounty goddesses such as Dinsana, Raka, Puramdhi, Parendi, Bharati, Mahi among others are mentioned in the Rigveda.[58] Sri, also called Lakshmi, appears in late Vedic texts dated to be pre-Buddhist, but verses dedicated to her do not suggest that her characteristics were fully developed in the Vedic era.[59] All gods and goddesses are distinguished in the Vedic times, but in the post-Vedic texts (~500 BCE to 200 CE), and particularly in the early medieval era literature, they are ultimately seen as aspects or manifestations of one Brahman, the Supreme power.[59][60]

 

Ananda Coomaraswamy states that Devas and Asuras in the Vedic lore are similar to Angels-Theoi-Gods and Titans of Greek mythology, both are powerful but have different orientations and inclinations, the Devas representing the powers of Light and the Asuras representing the powers of Darkness in Hindu mythology.[61][62] According to Coomaraswamy's interpretation of Devas and Asuras, both these natures exist in each human being, the tyrant and the angel is within each being, the best and the worst within each person struggles before choices and one's own nature, and the Hindu formulation of Devas and Asuras is an eternal dance between these within each person.[63][64]

 

The Devas and Asuras, Angels and Titans, powers of Light and powers of Darkness in Rigveda, although distinct and opposite in operation, are in essence consubstantial, their distinction being a matter not of essence but of orientation, revolution or transformation. In this case, the Titan is potentially an Angel, the Angel still by nature a Titan; the Darkness in actu is Light, the Light in potentia Darkness; whence the designations Asura and Deva may be applied to one and the same Person according to the mode of operation, as in Rigveda 1.163.3, "Trita art thou (Agni) by interior operation".

 

— Ananda Coomaraswamy, Journal of the American Oriental Society[65]

Characteristics of medieval era deities[edit]

In the Puranas and the Itihasas with the embedded Bhagavad Gita, the Devas represent the good, and the Asuras the bad.[3][4] According to the Bhagavad Gita (16.6-16.7), all beings in the universe have both the divine qualities (daivi sampad) and the demonic qualities (asuri sampad) within each.[4][66] The sixteenth chapter of the Bhagavad Gita states that pure god-like saints are rare and pure demon-like evil are rare among human beings, and the bulk of humanity is multi-charactered with a few or many faults.[4] According to Jeaneane Fowler, the Gita states that desires, aversions, greed, needs, emotions in various forms "are facets of ordinary lives", and it is only when they turn to lust, hate, cravings, arrogance, conceit, anger, harshness, hypocrisy, violence, cruelty and such negativity- and destruction-inclined that natural human inclinations metamorphose into something demonic (Asura).[4][66]

The Epics and medieval era texts, particularly the Puranas, developed extensive and richly varying mythologies associated with Hindu deities, including their genealogies.[67][68][69] Several of the Purana texts are named after major Hindu deities such as Vishnu, Shiva and Devi.[67] Other texts and commentators such as Adi Shankara explain that Hindu deities live or rule over the cosmic body as well in the temple of human body.[33][70] They remark that the Sun deity is the giver of vision, the Vayu deity the nose, the Prajapati the sexual organs, the Lokapalas (directions) are the ears, moon deity the mind, Mitra deity is the inward breath, Varuna deity is the outward breath, Indra deity the arms, Brhaspati the speech, Vishnu whose stride is great is the feet, and Maya is the smile.[70]

 

Symbolism[edit]

Edelmann states that gods and anti-gods of Hinduism are symbolism for spiritual concepts. For example, god Indra (a Deva) and the antigod Virocana (an Asura) question a sage for insights into the knowledge of the self.[71] Virocana leaves with the first given answer, believing now he can use the knowledge as a weapon. In contrast, Indra keeps pressing the sage, churning the ideas, and learning about means to inner happiness and power. Edelmann suggests that the Deva-Asura dichotomies in Hindu mythology may be seen as "narrative depictions of tendencies within our selves".[71] Hindu deities in Vedic era, states Mahoney, are those artists with "powerfully inward transformative, effective and creative mental powers".[72]

 

In Hindu mythology, everyone starts as an Asura, born of the same father. "Asuras who remain Asura" share the character of powerful beings craving for more power, more wealth, ego, anger, unprincipled nature, force and violence.[73][74] The "Asuras who become Devas" in contrast are driven by an inner voice, seek understanding and meaning, prefer moderation, principled behavior, aligned with Ṛta and Dharma, knowledge and harmony.[73][74][75]

 

The god (Deva) and antigod (Asura), states Edelmann, are also symbolically the contradictory forces that motivate each individual and people, and thus Deva-Asura dichotomy is a spiritual concept rather than mere genealogical category or species of being.[76] In the Bhāgavata Purana, saints and gods are born in families of Asuras, such as Mahabali and Prahlada, conveying the symbolism that motivations, beliefs and actions rather than one's birth and family circumstances define whether one is Deva-like or Asura-like.[76]

Another Hindu term that is sometimes translated as deity is Ishvara, or alternatively various deities are described, state Sorajjakool et al., as "the personifications of various aspects of one and the same Ishvara".[77] The term Ishvara has a wide range of meanings that depend on the era and the school of Hinduism.[78][79][80] In ancient texts of Indian philosophy, Ishvara means supreme soul, Brahman (Highest Reality), ruler, king or husband depending on the context.[78] In medieval era texts, Ishvara means God, Supreme Being, personal god, or special Self depending on the school of Hinduism.[2][80][81]

 

Among the six systems of Hindu philosophy, Samkhya and Mimamsa do not consider the concept of Ishvara, i.e., a supreme being, relevant. Yoga, Vaisheshika, Vedanta and Nyaya schools of Hinduism discuss Ishvara, but assign different meanings.

 

Early Nyaya school scholars considered the hypothesis of a deity as a creator God with the power to grant blessings, boons and fruits; but these early Nyaya scholars then rejected this hypothesis, and were non-theistic or atheists.[25][82] Later scholars of Nyaya school reconsidered this question and offered counter arguments for what is Ishvara and various arguments to prove the existence of omniscient, omnipresent, omnipotent deity (God).[83]

 

Vaisheshika school of Hinduism, as founded by Kanada in 1st millennium BC, neither required nor relied on creator deity.[84][85] Later Vaisheshika school adopted the concept of Ishvara, states Klaus Klostermaier, but as an eternal God who co-exists in the universe with eternal substances and atoms, but He "winds up the clock, and lets it run its course".[84]

 

Ancient Mimamsa scholars of Hinduism questioned what is Ishvara (deity, God)?[86] They considered deity concept unnecessary for a consistent philosophy and moksha (soteriology).[86][87]

 

In Samkhya school of Hindu philosophy, Isvara is neither a creator-God, nor a savior-God.[88] This is called one of the several major atheistic schools of Hinduism by some scholars.[89][90][91] Others, such as Jacobsen, state that Samkhya is more accurately described as non-theistic.[92] Deity is considered an irrelevant concept, neither defined nor denied, in Samkhya school of Hindu philosophy.[93]

 

In Yoga school of Hinduism, it is any "personal deity" (Ishta Deva or Ishta Devata)[94] or "spiritual inspiration", but not a creator God.[81][89] Whicher explains that while Patanjali's terse verses in the Yogasutras can be interpreted both as theistic or non-theistic, Patanjali's concept of Isvara in Yoga philosophy functions as a "transformative catalyst or guide for aiding the yogin on the path to spiritual emancipation".[95]

 

The Advaita Vedanta school of Hinduism asserted that there is no dualistic existence of deity (or deities).[96][97] There is no otherness nor distinction between Jiva and Ishvara.[98][99] God (Ishvara, Brahman) is identical with the Atman (soul) within each human being in Advaita Vedanta school,[100] and there is a monistic Universal Absolute Oneness that connects everyone and everything, states this school of Hinduism.[39][99][101] This school, states Anantanand Rambachan, has "perhaps exerted the most widespread influence".[102]

 

The Dvaita sub-school of Vedanta Hinduism, founded in medieval era, Ishvara is defined as a creator God that is distinct from Jiva (individual souls in living beings).[40] In this school, God creates individual souls, but the individual soul never was and never will become one with God; the best it can do is to experience bliss by getting infinitely close to God.[20]

 

Number of deities[edit]

Yāska, the earliest known language scholar of India (~ 500 BCE), notes Wilkins, mentions that there are three deities (Devas) according to the Vedas, "Agni (fire), whose place is on the earth; Vayu (wind), whose place is the air; and Surya (sun), whose place is in the sky".[107] This principle of three worlds (or zones), and its multiples is found thereafter in many ancient texts. The Samhitas, which are the oldest layer of text in Vedas enumerate 33 devas,[note 3] either 11 each for the three worlds, or as 12 Adityas, 11 Rudras, 8 Vasus and 2 Ashvins in the Brahmanas layer of Vedic texts.[7][47]

 

The Rigveda states in hymn 1.139.11,

 

ये देवासो दिव्येकादश स्थ पृथिव्यामध्येकादश स्थ ।

अप्सुक्षितो महिनैकादश स्थ ते देवासो यज्ञमिमं जुषध्वम् ॥११॥[111]

 

O ye eleven gods whose home is heaven, O ye eleven who make earth your dwelling,

Ye who with might, eleven, live in waters, accept this sacrifice, O gods, with pleasure.

– Translated by Ralph T. H. Griffith[112]

 

Gods who are eleven in heaven; who are eleven on earth;

and who are eleven dwelling with glory in mid-air; may ye be pleased with this our sacrifice.

– Translated by HH Wilson[113]

 

— Rigveda 1.139.11

Millions, one or one-ness?[edit]

Thirty-three divinities are mentioned in other ancient texts, such as the Yajurveda,[114] however, there is no fixed "number of deities" in Hinduism any more than a standard representation of "deity".[115] There is, however, a popular perception stating that there are 330 million (or "33 crore") deities in Hinduism.[116] Most, by far, are goddesses, state Foulston and Abbott, suggesting "how important and popular goddesses are" in Hindu culture.[115] No one has a list of the 330 million goddesses and gods, but all deities, state scholars, are typically viewed in Hinduism as "emanations or manifestation of genderless principle called Brahman, representing the many facets of Ultimate Reality".[115][116][117]

 

This concept of Brahman is not the same as the monotheistic separate God found in Abrahamic religions, where God is considered, states Brodd, as "creator of the world, above and independent of human existence", while in Hinduism "God, the universe, human beings and all else is essentially one thing" and everything is connected oneness, the same god is in every human being as Atman, the eternal Self.[117][118]

Hinduism has an ancient and extensive iconography tradition, particularly in the form of Murti (Sanskrit: मूर्ति, IAST: Mūrti), or Vigraha or Pratima.[22] A Murti is itself not the god in Hinduism, but it is an image of god and represents emotional and religious value.[124] A literal translation of Murti as idol is incorrect, states Jeaneane Fowler, when idol is understood as superstitious end in itself.[124] Just like the photograph of a person is not the real person, a Murti is an image in Hinduism but not the real thing, but in both cases the image reminds of something of emotional and real value to the viewer.[124] When a person worships a Murti, it is assumed to be a manifestation of the essence or spirit of the deity, the worshipper's spiritual ideas and needs are meditated through it, yet the idea of ultimate reality or Brahman is not confined in it.[124]

 

A Murti of a Hindu deity is typically made by carving stone, wood working, metal casting or through pottery. Medieval era texts describing their proper proportions, positions and gestures include the Puranas, Agamas and Samhitas particularly the Shilpa Shastras.[21] The expressions in a Murti vary in diverse Hindu traditions, ranging from Ugra symbolism to express destruction, fear and violence (Durga, Kali), as well as Saumya symbolism to express joy, knowledge and harmony (Saraswati, Lakshmi). Saumya images are most common in Hindu temples.[125] Other Murti forms found in Hinduism include the Linga.[126]

 

A Murti is an embodiment of the divine, the Ultimate Reality or Brahman to some Hindus.[21] In religious context, they are found in Hindu temples or homes, where they may be treated as a beloved guest and serve as a participant of Puja rituals in Hinduism.[127] A murti is installed by priests, in Hindu temples, through the Prana Pratishtha ceremony,[128] whereby state Harold Coward and David Goa, the "divine vital energy of the cosmos is infused into the sculpture" and then the divine is welcomed as one would welcome a friend.[129] In other occasions, it serves as the center of attention in annual festive processions and these are called Utsava Murti.[130]

In Hinduism, deities and their icons may be hosted in a Hindu temple, within a home or as an amulet. The worship performed by Hindus is known by a number of regional names, such as Puja.[134] This practice in front of a murti may be elaborate in large temples, or be a simple song or mantra muttered in home, or offering made to sunrise or river or symbolic anicon of a deity.[135][136][137] Archaeological evidence of deity worship in Hindu temples trace Puja rituals to Gupta Empire era (~4th century CE).[138][139] In Hindu temples, various pujas may be performed daily at various times of the day; in other temples, it may be occasional.[140][141]

 

The Puja practice is structured as an act of welcoming, hosting, honoring the deity of one's choice as one's honored guest,[142] and remembering the spiritual and emotional significance the deity represents the devotee.[124][134] Jan Gonda, as well as Diana L. Eck, states that a typical Puja involves one or more of 16 steps (Shodasha Upachara) traceable to ancient times: the deity is invited as a guest, the devotee hosts and takes care of the deity as an honored guest, praise (hymns) with Dhupa or Aarti along with food (Naivedhya) is offered to the deity, after an expression of love and respect the host takes leave, and with affection expresses good bye to the deity.[143][144] The worship practice may also involve reflecting on spiritual questions, with image serving as support for such meditation.[145]

 

Deity worship (Bhakti), visiting temples and Puja rituals are not mandatory and is optional in Hinduism; it is the choice of a Hindu, it may be a routine daily affair for some Hindus, periodic ritual or infrequent for some.[146][147] Worship practices in Hinduism are as diverse as its traditions, and a Hindu can choose to be polytheistic, pantheistic, monotheistic, monistic, agnostic, atheistic or humanist.[41]

 

Examples[edit]

Main articles: List of Hindu deities and Rigvedic deities

Major deities have inspired a vast genre of literature such as the Puranas and Agama texts as well their own Hindu traditions, but with shared mythology, ritual grammar, theosophy, axiology and polycentrism.[16][17] Vishnu and his avatars are at the foundation of Vaishnavism, Shiva for Shaivism, Devi for Shaktism, and some Hindu traditions such as Smarta traditions who revere multiple major deities (five) as henotheistic manifestations of Brahman (absolute metaphysical Reality).[116][148][149]

 

While there are diverse deities in Hinduism, states Lawrence, "Exclusivism – which maintains that only one's own deity is real" is rare in Hinduism.[116] Julius Lipner, and other scholars, state that pluralism and "polycentrism" – where other deities are recognized and revered by members of different "denominations", has been the Hindu ethos and way of life.[16][150]

 

Trimurti and Tridevi[edit]

The concept of Triad (or Trimurti, Trinity) makes a relatively late appearance in Hindu literature, or in the second half of 1st millennium BCE.[151] The idea of triad, playing three roles in the cosmic affairs, is typically associated with Brahma, Vishnu and Shiva (also called Mahesh); however, this is not the only triad in Hindu literature.[152] Other triads include Tridevi, of three goddesses – Lakshmi, Saraswati and Durga in the text Devi Mahatmya, in the Shakta tradition, who further assert that Devi is the Brahman (Ultimate Reality) and it is her energy that empowers Brahma, Vishnu and Shiva.[151] The other triads, formulated as deities in ancient Indian literature, include Sun (creator), Air (sustainer) and Fire (destroyer); Prana (creator), Food (sustainer) and Time (destroyer).[151] These triads, states Jan Gonda, are in some mythologies grouped together without forming a Trinity, and in other times represented as equal, a unity and manifestations of one Brahman.[151] In the Puranas, for example, this idea of threefold "hypostatization" is expressed as follows,

 

They [Brahma, Vishnu, Shiva] exist through each other, and uphold each other; they are parts of one another; they subsist through one another; they are not for a moment separated; they never abandon one another.

 

— Vayu Purana, 5.17, Translated by Jan Gonda[151]

The triad appears in Maitrayaniya Upanishad, for the first time in recognized roles known ever since, where they are deployed to present the concept of three Guṇa – the innate nature, tendencies and inner forces found within every being and everything, whose balance transform and keeps changing the individual and the world.[152][153] It is in the medieval Puranic texts, Trimurti concepts appears in various context, from rituals to spiritual concepts.[151] The Bhagavad Gita, in verses 9.18, 10.21-23 and 11.15, asserts that the triad or trinity is manifestation of one Brahman, which Krishna affirms himself to be.[154] However, suggests Bailey, the mythology of triad is "not the influence nor the most important one" in Hindu traditions, rather the ideologies and spiritual concepts develop on their own foundations.[152]

 

Avatars of Hindu deities[edit]

Hindu mythology has nurtured the concept of Avatar, which represents the descent of a deity on earth.[155][156] This concept is commonly translated as "incarnation",[155] and is an "appearance" or "manifestation".[157][158]

 

The concept of Avatar is most developed in Vaishnavism tradition, and associated with Vishnu, particularly with Rama and Krishna.[159][160] Vishnu takes numerous avatars in Hindu mythology. He becomes female, during the Samudra manthan, in the form of Mohini, to resolve a conflict between the Devas and Asuras. His male avatars include Matsya, Kurma, Varaha, Narasimha, Vamana, Parashurama, Rama, Krishna, Buddha, and Kalki.[160] Various texts, particularly the Bhagavad Gita, discuss the idea of Avatar of Vishnu appearing to restore the cosmic balance whenever the power of evil becomes excessive and causes persistent oppression in the world.[156]

 

In Shaktism traditions, the concept appears in its legends as the various manifestations of Devi, the Divine Mother principal in Hinduism.[161] The avatars of Devi or Parvati include Durga and Kali, who are particularly revered in eastern states of India, as well as Tantra traditions.[162][163][164] Twenty one avatars of Shiva are also described in Shaivism texts, but unlike Vaishnava traditions, Shaiva traditions have focussed directly on Shiva rather than the Avatar concept.[155]

 

Introducing you to this amazing musical game which is Kids Piano. This game includes many baby instruments with amazing sounds for kids to play and enjoy their best times using this amazing game.

 

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Maltby Street Food Market SE1: Ropewalk -

 

A small but bustling market offering fresh produce, including breads, cakes, meats and cheeses. Popular sellers include African Volcano, Waffle On, Tozino, Bad Brownie, Finest Fayre Scotch Egg & Pies, The Cheese Truck, Maltby Greek, The Modern Beer Bar, Avocado Cafe, The Gay Farmer, Monty's Deli sandwiches, St John Bakery's sourdough, fresh filled doughnuts and many more. The Kernel Brewery is also open during market time. The market is open Saturdays between 9am-4pm and Sundays between 11am-4pm - information from Maltby Street Market.

 

Southbank Centre & Food Village -

 

Southbank Centre has become a food-lovers destination and one of London’s most famous food market venues. Try, buy, eat and shop throughout the year.

Most weekends, Southbank Centre Square hosts Real Food Market, with tasty treats and fresh produce to sample and buy. We also host KERB and will have a new-look Christmas market this Winter - info from Southbank Centre.

 

Shoreditch Street Kitchen -

 

Street Kitchen parks up in east London ahead of Shoreditch Village development

 

By Luke Nicholls, 26-Mar-2013

 

Street Kitchen – the Airstream caravan concept from Jun Tanaka and Mark Jankel – has just opened up a third venue on Shoreditch High Street, which will run until the end of the year ahead of a potential permanent site within the upcoming Shoreditch Village development.

 

www.bighospitality.co.uk/Venues/Street-Kitchen-parks-up-i...

 

COPYRIGHT – Please be aware that all the images on this blog are the property of – © Enrique Guadiz Photography.

 

Feel free to share but do not download, copy, manipulate or use in any other way.

If you wish to use one or more then please contact me and we can discuss use and costs, if any.

This is the first of my posters that includes three recently dropped photos

 

1. Kiss - P1070599, 2. Coburg - IMG_6665, 3. Creidlitz - IMG_4714, 4. Coburg Rathaus - IMG_3885, 5. Klein Schneen - IMG_2189-1, 6. Misty Morning at the Ochsenkopf Slopes, 7. Günter - IMG_1930, 8. Arkaden in Coburg - IMG_4734,

 

9. First Snow - IMG_1610-1, 10. Feuer Akrobatin - IMG_0282, 11. Russia: Michurinskoye - Squared Fire - IMG_8223-3, 12. Rainha da Bateria Dance Competition - Tanz Wettbewerb Râinha da Bateria - IMG_1058, 13. Tulpen - IMG_4490_b, 14. Schloss Peterhof - St. Petersburg - P6160009, 15. Tulips in April - Weinheim Hermannshof Garden - P4170031, 16. 2001-09-12 Sommer (12) Bellagio Lago di Como Italien 19,

 

17. 2001-09-12 Sommer (12) Bellagio Lago di Como Italien 19-1, 18. View from the Rosskopf near Tegernsee - pano - Oberbayern Suttengebiet 2005-07, 19. P1110235 - Dahlie quadratisch, 20. Morgens am Ochsenkopf - 25 September 2005 - P1090070, 21. Foto Serie Stiftskirche Mariä Geburt Rottenbuch P1060514, 22. Street Parade - A Photo Exhibition from the Sambafestival Coburg 2005, 23. Germany: Brocken, 24. St. Petersburg Eremitage P6120143

 

Created with fd's Flickr Toys.

Jan. 21, 2010. Boston, MA.

Gavin McCarthy.

Transgender Equality Lobby Day”sponsored by The Massachusetts Transgender Political Coalition. Transgender people and allies rallied at the Massachusetts State House to urge legislators to ban discrimination in the Commonwealth based on gender identity and expression by passing the bill H1728/S1687 “An Act Relative To Gender-Based Discrimination and Hate Crimes. Participants then visited their legislators to tell them their stories and ask them to support the bill.

Speakers at “Transgender Equality Lobby Day” include Chairman of House

Ways and Means Committee Charles Murphy, Co-Chair of the Joint

Committee on the Judiciary Senator Cynthia Creem, Senator Ben Downing,

Representative Byron Rushing, Representative Carl Sciortino, Jr.,

MTPC’s Executive Director Gunner Scott and community members Jesse

Pack, Marion Freedman-Gurspan, and Lorelei McLaughlin.

One of the speakers, Lorelei McLaughlin, struggled for many years before

she was finally able to find employment. “I would walk into a place

with a help wanted sign in the window, only to be told that, well;

they weren’t actually hiring right now but would be happy to put my

application on file. I couldn’t even get anyone to look at my

resume.”

  

From a stroll along Hove seafront as the sun was going down.

sheridan, wyoming

1972

 

all american indian days

 

set includes pictures taken during a trip through the blacks hills of the western united states; to include shots of the all-american indian days celebration in sheridan, wyoming, devil's tower national monument, and the "days of '76" festival in deadwood, south dakota.

 

part of an archival project, featuring the photographs of nick dewolf

 

© the Nick DeWolf Foundation

Image-use requests are welcome via flickrmail or nickdewolfphotoarchive [at] gmail [dot] com

A few more from the Brighton Pride Parade - before the start ot the event down on Madeira Drive

More from last weekend's trip to the velodrome for the last day of the 3 Day London.

The spectacular sapphire tiara includes 655 diamonds and 33 sapphires, It was a gift from King Willem III for his wife Queen Emma in 1881. The larger removable sapphire in the middle is a legacy of Anna Pavlovna (wife of king Willem II, daughter of the Tsar of Russia).

Polo action from the Cooch Behar Cup match at Cowdray, between Clarke & Green and King Power Foxes

Thursday at the 2021 Goodwood Festival of Speed, which went ahead as a government Covid secure test event.

The Eurasian eagle-owl (Bubo bubo) is a species of eagle-owl that resides in much of Eurasia. It is also called the Uhu and it is occasionally abbreviated to just the eagle-owl in Europe. It is one of the largest species of owl, and females can grow to a total length of 75 cm (30 in), with a wingspan of 188 cm (6 ft 2 in), with males being slightly smaller. This bird has distinctive ear tufts, with upper parts that are mottled with darker blackish colouring and tawny. The wings and tail are barred. The underparts are a variably hued buff, streaked with darker colouring. The facial disc is not very defined and the orange eyes are distinctive.

 

Eurasian eagle-owls are found in many habitats, but are mostly birds of mountainous regions or other rocky areas, often those near varied woodland edge and shrubby areas with openings or wetlands to hunt a majority of their prey. Additionally, they inhabit coniferous forests, steppes, and other areas at varied elevations that are typically relatively remote. Eurasian eagle-owls are occasionally found amongst farmland and in park-like settings within European cities, even rarely within busier urban areas. The eagle-owl is mostly a nocturnal predator, hunting for a range of different prey species. Predominantly, their diet is composed of small mammals such as rodents and rabbits, but they also prey on larger mammals and birds of varying sizes. Other secondary prey can include reptiles, amphibians, fish, large insects and other assorted invertebrates. The species typically breeds on cliff ledges, in gullies, among rocks, or in other concealed locations. The nest is a scrape containing a clutch of 2–4 eggs typically, which are laid at intervals and hatch at different times. The female incubates the eggs and broods the young, and the male provides food for her, and when they hatch, for the nestlings, as well. Continuing parental care for the young is provided by both adults for about five months. At least 12 subspecies of the Eurasian eagle-owl are described.

 

In addition to being one of the largest living species of owl, the Eurasian eagle-owl is also one of the most widely distributed.[9] With a total range in Europe and Asia of about 51.4 million km2 (19.8 million sq mi) and a total population estimated to be between 100,000 and 500,000 individuals, the IUCN lists the bird's conservation status as being of least concern, although the trend is listed as decreasing. The vast majority of eagle-owls live in Continental Europe, Scandinavia, Russia (which is almost certainly where the peak numbers and diversity of race occurs), and Central Asia. Additional minor populations exist in Anatolia, the northern Middle East, the montane upper part of South Asia, China, Korea and in Japan; in addition, an estimated 12 to 40 pairs are thought to reside in the United Kingdom as of 2016 (where they are arguably non-native), a number which may be on the rise, and have successfully bred in the UK since at least 1996. Tame eagle-owls have occasionally been used in pest control because of their size to deter large birds such as gulls from nesting.

 

Description

The Eurasian eagle-owl is among the larger birds of prey, smaller than the golden eagle (Aquila chrysaetos), but larger than the snowy owl (Bubo scandiacus), despite some overlap in size with both of those species. It is sometimes referred to as the world's largest owl, although Blakiston's fish owl (B. blakistoni) is slightly heavier on average and the much lighter weight great grey owl (Strix nebulosa) is slightly longer on average. Heimo Mikkola reported the largest specimens of eagle-owl as having the same upper body mass, 4.6 kg (10 lb), as the largest Blakiston’s fish owl and attained a length around 3 cm (1.2 in) longer. In terms of average weight and wing size, the Blakiston’s is the slightly larger species seemingly, even averaging a bit larger in these aspects than the biggest eagle-owl races from Russia. Also, although 9 cm (3.5 in) shorter than the largest of the latter species, the Eurasian eagle-owl can weigh well more than twice as much as the largest great grey owl. The Eurasian eagle-owl typically has a wingspan of 131–188 cm (4 ft 4 in – 6 ft 2 in), with the largest specimens possibly attaining 2 m (6 ft 7 in). The total length of the species can vary from 56 to 75 cm (22 to 30 in). Females can weigh from 1.75 to 4.6 kg (3.9 to 10.1 lb), and males can weigh from 1.2 to 3.2 kg (2.6 to 7.1 lb). In comparison, the barn owl (Tyto alba), the world's most widely distributed owl species, weighs about 0.5 kg (1.1 lb) and the great horned owl (B. virginianus), which fills the eagle-owl's ecological niche in North America, weighs around 1.4 kg (3.1 lb).

  

Eurasian eagle-owl in captivity

Besides the female being larger, little external sexual dimorphism is seen in the Eurasian eagle-owl, although the ear tufts of males reportedly tend to be more upright than those of females. When an eagle-owl is seen on its own in the field, distinguishing the individual’s sex is generally not possible. Gender determination by size is possible by in-hand measurements. In some populations, the female typically may be slightly darker than the male. The plumage coloration across at least 13 accepted subspecies can be highly variable. The upper parts may be brown-black to tawny-buff to pale creamy gray, typically showing dense freckling on the forehead and crown, stripes on the nape, sides, and back of the neck, and dark splotches on the pale ground colour of the back, mantle, and scapulars. A narrow buff band, freckled with brown or buff, often runs up from the base of the bill, above the inner part of the eye, and along the inner edge of the black-brown ear tufts. The rump and upper tail-coverts are delicately patterned with dark vermiculations and fine, wavy barring, the extent of which varies with subspecies. The underwing coverts and undertail coverts are similar, but tend to be more strongly barred in brownish-black.

 

The primaries and secondaries are brown with broad, dark brown bars and dark brown tips, and grey or buff irregular lines. A complete moult takes place each year between July and December. The facial disc is tawny-buff, speckled with black-brown, so densely on the outer edge of the disc as to form a "frame" around the face. The chin and throat are white with a brownish central streak. The feathers of the upper breast generally have brownish-black centres and reddish-brown edges except for the central ones, which have white edges. The chin and throat may appear white continuing down the center of the upper breast. The lower breast and belly feathers are creamy-brown to tawny buff to off-white with a variable amount of fine dark wavy barring, on a tawny-buff ground colour. The legs and feet (which are feathered almost to the talons) are likewise marked on a buff ground colour but more faintly. The tail is tawny-buff, mottled dark grey-brown with about six black-brown bars. The bill and feet are black. The iris is most often orange but is fairly variable. In some European birds, the iris is a bright reddish, blood-orange colour but then in subspecies found in arid, desert-like habitats, the iris can range into an orange-yellow colour (most closely related species generally have yellowish irises, excluding the Indian eagle-owl).

 

Standard measurements and physiology

Among standard measurements for the Eurasian eagle-owl, the wing chord measures 378 to 518 mm (14.9 to 20.4 in), the tail measures 229–310 mm (9.0–12.2 in) long, the tarsus measures 64.5–112 mm (2.54–4.41 in), and the total length of the bill is 38.9–59 mm (1.53–2.32 in). The wings are reportedly the smallest in proportion to the body weight of any European owl, when measured by the weight per area of wing size, was found to be 0.72 g/cm2. Thus, they have quite high wing loading. The great horned owl has even smaller wings (0.8 g/cm2) relative to its body size. The golden eagle has slightly lower wing loading proportionately (0.65 g/cm2), so the aerial abilities of the two species (beyond the eagle’s spectacular ability to stoop) may not be as disparate as expected. Some other owls, such as barn owls, short-eared owls (Asio flammeus), and even the related snowy owls have lower wing loading relative to their size, so are presumably able to fly faster, with more agility, and for more extended periods than the Eurasian eagle-owl. In the relatively small race B. b. hispanus, the middle claw, the largest talon, (as opposed to rear hallux-claw, which is the largest in accipitrids) was found to measure from 21.6 to 40.1 mm (0.85 to 1.58 in) in length. A 3.82 kg (8.4 lb) female examined in Britain (origins unspecified) had a middle claw measuring 57.9 mm (2.28 in), on par in length with a large female golden eagle hallux-claw. Generally, owls do not have talons as proportionately large as those of accipitrids, but have stronger, more robust feet relative to their size. Accipitrids use their talons to inflict organ damage and blood loss, whereas typical owls use their feet to constrict their prey to death, the talons serving only to hold the prey in place or provide incidental damage. The talons of the Eurasian eagle-owl are very large and not often exceeded in size by diurnal raptors. Unlike the great horned owls, the overall foot size and strength of the Eurasian eagle-owl is not known to have been tested, but the considerably smaller horned owl has one of the strongest grips ever measured in a bird.

 

The feathers of the ear tufts in Spanish birds (when not damaged) were found to measure from 63.3 to 86.6 mm (2.49 to 3.41 in).[26] The ear openings (covered in feathers as in all birds) are relatively uncomplicated for an owl, but are also large, being larger on the right than on the left as in most owls, and proportionately larger than those of the great horned owl. In the female, the ear opening averages 31.7 mm (1.25 in) on the right and 27.4 mm (1.08 in) on the left, and in males, averages 26.8 mm (1.06 in) on the right and 24.4 mm (0.96 in) on the left. The depth of the facial disc and the size and complexity of the ear opening are directly correlated to the importance of sound in an owl’s hunting behaviour. Examples of owls with more complicated ear structures and deeper facial disc are barn owls, long-eared owls (Asio otus), and boreal owls (Aegolius funereus). Given the uncomplicated structure of their ear openings and relatively shallow, undefined facial discs, hunting by ear is secondary to hunting by sight in eagle-owls; this seems to be true for Bubo in general. More sound-based hunters such as the aforementioned species likely focus their hunting activity in more complete darkness. Also, owls with white throat patches such as the Eurasian eagle-owl are more likely to be active in low-light conditions in the hours before and after sunrise and sunset rather than the darkest times in the middle of the night. The boreal and barn owls, to extend these examples, lack obvious visual cues such as white throat patches (puffed up in displaying eagle-owls), again indicative of primary activity being in darker periods.

 

Distinguishing from other species

The great size, bulky, barrel-shaped build, erect ear tufts, and orange eyes render this as a distinctive species. Other than general morphology, the above features differ markedly from those of two of the next largest subarctic owl species in Europe and western Asia, which are the great grey owl and the greyish to chocolate-brown Ural owl (Strix uralensis), both of which have no ear tufts and have a distinctly rounded head, rather than the blocky shape of the eagle-owl’s head. The snowy owl is obviously distinctive from most eagle-owls, but during winter the palest Eurasian eagle-owl race (B. b. sibiricus) can appear off-white. Nevertheless, the latter is still distinctively an ear-tufted Eurasian eagle-owl and lacks the pure white background colour and variable blackish spotting of the slightly smaller species (which has relatively tiny, vestigial ear tufts that have only been observed to have flared on rare occasions).

  

Unique camouflage pattern

The long-eared owl has a somewhat similar plumage to the eagle-owl, but is considerably smaller (an average female eagle-owl may be twice as long and 10 times heavier than an average long-eared owl). Long-eared owls in Eurasia have vertical striping like that of the Eurasian eagle-owl, while long-eared owls in North America show a more horizontal striping like that of great horned owls. Whether these are examples of mimicry either way is unclear but it is known that both Bubo owls are serious predators of long-eared owls. The same discrepancy in underside streaking has also been noted in the Eurasian and American representations of the grey owl. A few other related species overlap minimally in range in Asia, mainly in East Asia and the southern reaches of the Eurasian eagle-owl’s range. Three fish owls appear to overlap in range, the brown (Ketupa zeylonensis) in at least northern Pakistan, probably Kashmir, and discontinuously in southern Turkey, the tawny (K. flavipes) through much of eastern China, and Blakiston's fish owl in the Russian Far East, northeastern China, and Hokkaido. Fish owls are distinctively different looking, possessing more scraggy ear tufts that hang to the side rather than sit erect on top of the head, and generally have more uniform, brownish plumages without the contrasting darker streaking of an eagle-owl. The brown fish owl has no feathering on the tarsus or feet, and the tawny has feathering only on the upper portion of the tarsi, but the Blakiston’s is nearly as extensively feathered on the tarsi and feet as the eagle-owl. Tawny and brown fish owls are both slightly smaller than co-occurring Eurasian eagle-owls, and Blakiston’s fish owls are similar or slightly larger than co-occurring large northern eagle-owls. Fish owls, being tied to the edges of fresh water, where they hunt mainly fish and crabs, also have slightly differing, and more narrow, habitat preferences.

 

In the lower Himalayas of northern Pakistan and Jammu and Kashmir, along with the brown fish owl, the Eurasian eagle-owl at the limit of its distribution may co-exist with at least two to three other eagle-owls. One of these, the dusky eagle-owl (B. coromandus) is smaller, with more uniform tan-brownish plumage, untidy uniform light streaking rather than the Eurasian’s dark streaking below and an even less well-defined facial disc. The dusky is usually found in slightly more enclosed woodland areas than Eurasian eagle-owls. Another is possibly the spot-bellied eagle-owl (B. nipalensis), which is strikingly different looking, with stark brown plumage, rather than the warm hues typical of the Eurasian, bold spotting on a whitish background on the belly, and somewhat askew ear tufts that are bold white with light brown crossbars on the front. Both species may occur in some parts of the Himalayan foothills, but they are not currently verified to occur in the same area, in part because of the spot-bellied’s preference for dense, primary forest. Most similar, with basically the same habitat preferences and the only one verified to co-occur with the Eurasian eagle-owls of the race B. b. turcomanus in Kashmir is the Indian eagle-owl (B. bengalensis). The Indian species is smaller, with a bolder, blackish facial disc border, more rounded and relatively smaller wings, and partially unfeathered toes. Far to the west, the pharaoh eagle-owl (B. ascalaphus) also seemingly overlaps in range with the Eurasian, at least in Jordan. Although also relatively similar to the Eurasian eagle-owl, the pharaoh eagle-owl is distinguished by its smaller size, paler, more washed-out plumage, and the diminished size of its ear tufts.

 

Moulting

The Eurasian eagle-owls’ feathers are lightweight and robust, but nevertheless need to be replaced periodically as they become worn. In the Eurasian eagle-owl, this happens in stages, and the first moult starts the year after hatching with some body feathers and wing coverts being replaced. The next year, the three central secondaries on each wing and three middle tail feathers are shed and regrow, and the following year, two or three primaries and their coverts are lost. In the final year of this postjuvenile moult, the remaining primaries are moulted and all the juvenile feathers will have been replaced. Another moult takes place during years 6-12 of the bird's life. This happens between June and October after the conclusion of the breeding season, and again it is a staged process with six to nine main flight feathers being replaced each year. Such a moulting pattern lasting several years is repeated throughout the bird's life.

 

Taxonomy

The Eurasian eagle-owl was formally described by the Swedish naturalist Carl Linnaeus in 1758 in the tenth edition of his Systema Naturae under the binomial name Strix bulbo. Although Linnaeus specified the "habitat" as "Europa" the type locality is restricted to Sweden. The Eurasian eagle-owl is now placed in the genus Bubo that was introduced by André Duméril in 1805.

 

The genus Bubo with 20 extant species includes most of the larger owl species in the world today. Based on an extensive fossil record and a central distribution of extant species on that continent, Bubo appears to have evolved into existence in Africa, although early radiations seem to branch from southern Asia, as well. Two genera belonging to the scops owls complex, the giant scops owls (Otus gurneyi) found in Asia and the Ptilopsis or the white-faced scops owl found in Africa, although firmly ensconced in the scops owl group, appear to share some characteristics with the eagle-owls. The Strix genus is also related to Bubo, and is considered a "sister complex", with Pulsatrix possibly being intermediate between the two. The Eurasian eagle-owl appears to represent an expansion of the genus Bubo into the Eurasian continent. A few of the other species of Bubo seem to have been derived from the Eurasian eagle-owl, making it a "paraspecies", or they at least share a relatively recent common ancestor.

 

The pharaoh eagle-owl, distributed in the Arabian Peninsula and sections of the Sahara Desert through North Africa where rocky outcrops are found, was until recently considered a subspecies of the Eurasian eagle-owl. The pharaoh eagle-owl apparently differs about 3.8% in mitochondrial DNA from the Eurasian eagle-owl, well past the minimum genetic difference to differentiate species of 1.5%. Smaller and paler than Eurasian eagle-owls, the pharaoh eagle-owl can also be considered a distinct species largely due to its higher-pitched and more descending call, and the observation that Eurasian eagle-owls formerly found in Morocco (B. b. hispanus) apparently did not breed with the co-existing pharaoh eagle-owls. On the contrary, the race still found together with the pharaoh eagle-owl in the wild (B. b. interpositus) in the central Middle East has been found to interbreed in the wild with the pharaoh eagle-owl, although genetical materials have indicated B. b. interpositus may itself be a distinct species from the Eurasian eagle-owl, as it differs from the nominate subspecies of the Eurasian eagle-owl by 2.8% in mitochondrial DNA. For three Asian Eurasian eagle-owl subspecies (B. b. ussuriensis, B. b. kiautschensis and B. b. hemachlana, respectively), it was found that they met the criterion for subspecies well, with a high haplotype diversity and in spite of a relatively recent common ancestor and low genetic diversity. The Indian eagle-owl (B. bengalensis) was also considered a subspecies of the Eurasian eagle-owl until recently, but its smaller size, distinct voice (more clipped and high-pitched than the Eurasian), and the fact that it is largely allopatric in distribution (filling out the Indian subcontinent) with other Eurasian eagle-owl races has led to it being considered a distinct species. The mitochondrial DNA of the Indian species also appears considerably distinct from the Eurasian species. The Cape eagle-owl (B. capensis) appears to represent a return of this genetic line back into the African continent, where it leads a lifestyle similar to Eurasian eagle-owls, albeit far to the south. Another offshoot of the northern Bubo group is the snowy owl. It appears to have separated from other Bubo species at least 4 million years ago.

 

The fourth and most famous derivation of the evolutionary line that includes the Eurasian eagle-owl is the great horned owl, which appears to have been the result of primitive eagle-owls spreading into North America. According to some authorities, the great horned owls and Eurasian eagle-owls are barely distinct as species, with a similar level of divergence in their plumages as the Eurasian and North American representations of the great grey owl or the long-eared owl. More outward physical differences exist between the great horned owl and the Eurasian eagle-owl than in those two examples, including a great size difference favoring the Eurasian species, the great horned owl’s horizontal rather than vertical underside barring, yellow rather than orange eyes, and a much stronger black bracket to the facial disc, not to mention a number of differences in their reproductive behaviour and distinctive voices. Furthermore, genetic research has revealed that the snowy owl is more closely related to the great horned owl than are Eurasian eagle-owls. The most closely related species beyond the pharaoh, Indian, and Cape eagle-owls to the Eurasian eagle-owl is the smaller, less powerful and African spotted eagle-owl (B. africanus), which was likely to have divided from the line before they radiated away from Africa. Somehow, genetic materials indicate the spotted eagle-owl appears to share a more recent ancestor with the Indian eagle-owl than with the Eurasian eagle-owl or even the sympatric Cape eagle-owl. Eurasian eagle-owls in captivity have produced apparently healthy hybrids with both the Indian eagle-owl and the great horned owl. The pharaoh, Indian, and Cape eagle-owls and the great horned owl are all broadly similar in size to each other, but all are considerably smaller than the Eurasian eagle-owl, which averages at least 15–30% larger in linear dimensions and 30–50% larger in body mass than these other related species, possibly as the eagle-owls adapted to warmer climates and smaller prey. Fossils from southern France have indicated that during the Middle Pleistocene, Eurasian eagle-owls (this paleosubspecies is given the name B. b. davidi) were larger than they are today, even larger were those found in Azerbaijan and in the Caucasus (either B. b. bignadensis or B. bignadensis), which were deemed to date to the Late Pleistocene. About 12 subspecies are recognized today.

 

Habitat

Eagle-owls are distributed somewhat sparsely, but can potentially inhabit a wide range of habitats, with a partiality for irregular topography. They have been found in habitats as diverse as northern coniferous forests to the edge of vast deserts. Essentially, Eurasian eagle-owls have been found living in almost every climatic and environmental condition on the Eurasian continent, excluding the greatest extremities, i.e. they are absent from humid rainforest in Southeast Asia, and the high Arctic tundra, both of which they are more or less replaced by other species of Bubo owls. They are often found in the largest numbers in areas where cliffs and ravines are surrounded by a scattering of trees and bushes. Grassland areas such as alpine meadows or desert-like steppe can also host them so long as they have the cover and protection of rocky areas. The preference of eagle-owls for places with irregular topography has been reported in most known studies. The obvious benefit of such nesting locations is that both nests and daytime roosts located in rocky areas and/or steep slopes would be less accessible to predators, including man. Also, they may be attracted to the vicinity of riparian or wetlands areas, because the soft soil of wet areas is conducive to burrowing by the small, terrestrial mammals normally preferred in the diet, such as voles and rabbits.

 

Due to their preference for rocky areas, the species is often found in mountainous areas, and can be found up to elevations of 2,100 m (6,900 ft) in the Alps, 4,500 m (14,800 ft) in the Himalayas, and 4,700 m (15,400 ft) in the adjacent Tibetan Plateau. They can also be found living at sea level and may nest amongst rocky sea cliffs. Despite their success in areas such as subarctic zones and mountains that are frigid for much of the year, warmer conditions seem to result in more successful breeding attempts per studies in the Eifel region of Germany. In a study from Spain, areas primarily consisting of woodlands (52% of study area being forested) were preferred with pine trees predominating the oaks in habitats used, as opposed to truly mixed pine-oak woodland. Pine and other coniferous stands are often preferred in great horned owls, as well, due to the constant density, which make overlooking the large birds more likely. In mountainous forest, they are not generally found in enclosed wooded areas, as is the tawny owl (Strix alucco), instead usually near forest edge. Only 2.7% of the habitat included in the territorial ranges for eagle-owls per the habitat study in Spain consisted of cultivated or agricultural land. Compared to golden eagles, though, they can visit cultivated land more regularly in hunting forays due to their nocturnal habits, which allow them to largely evade human activity. Other accounts make clear that farmland is only frequented where its less intensively farmed, holds more extensive treed and bushy areas, and often has limited to no irrigation; farmland areas with fallow or abandoned fields are more likely to hold more prey, so are prone to less frequent human disturbance. In the Italian Alps, almost no pristine habitat remained, and eagle-owls nested locally in the vicinity of towns, villages, and ski resorts.

 

Although found in the largest numbers in areas sparsely populated by humans, farmland is sometimes inhabited, and they even have been observed living in park-like or other quiet settings within European cities. Since 2005, at least five pairs have nested in Helsinki. This is due in part to feral European rabbits (Oryctolagus cuniculus) having recently populated the Helsinki area, originally from pet rabbits released to the wild. The number is expected to increase due to the growth of the European rabbit population in Helsinki. European hares (Lepus europaeus), the often preferred prey species by biomass of the eagle-owls in their natural habitat, live only in rural areas of Finland, not in the city centre. In June 2007, an eagle-owl nicknamed 'Bubi' landed in the crowded Helsinki Olympic Stadium during the European Football Championship qualification match between Finland and Belgium. The match was interrupted for six minutes. After tiring of the match, following Jonathan Johansson's opening goal for Finland, the bird left the scene. Finland's national football team have had the nickname Huuhkajat (Finnish for "Eurasian eagle-owls") ever since. The owl was named "Helsinki Citizen of the Year" in December 2007. In 2020, a brood of three eagle-owl chicks was raised by their mother on a large, well-foliaged planter on an apartment window in the city centre of Geel, Belgium.

 

Distribution

The Eurasian eagle-owl is one of the most widely distributed of all owl species, although it is far less wide-ranging than the barn owl, the short-eared owl (Asio flammeus) and long-eared owl and lacks the circumpolar range of boreal species such as great grey owl, boreal owl and northern hawk owl (Surnia ulula). This eagle-owl reaches its westernmost range in the Iberian Peninsula, both almost throughout Spain and more spottily in Portugal. From there, the Eurasian eagle-owl ranges widely in the south of France from Toulouse to Monaco and as far north into the central part of the country as in Allier. Farther north, they are found sporadically and discontinuously in Luxembourg, southern and western Belgium and scarcely into the Netherlands. It is infrequently found in southern and central United Kingdom. In Germany, the eagle-owl can be found in large but highly discontinuous areas, mostly in the south and central areas but is almost entirely absent in areas such as Brandenburg. Across from its south German range, this species range is nearly continuous into the Czech Republic, Slovakia, northern and eastern Hungary and very spottily into Poland. In the fairly montane countries of Switzerland and Austria, the eagle-owl can be found fairly broadly. In Italy, the Eurasian eagle-owl is found where the habitat is favorable in much of the northern, western and central portions down to as far south Melito di Porto Salvo. From Italy, this species sweeps quite broadly along the Mediterranean coast in Southeastern Europe from Slovenia mostly continuously to most of Greece and Bulgaria. In eastern Europe, the Eurasian eagle-owl is found essentially throughout from central Romania to Estonia. The species also occupies a majority of Finland and Scandinavia, where most broadly found in Norway, somewhat more spottily in Sweden and in Denmark it is found widely in Jutland (absent from the islands).

 

The Eurasian eagle-owl's range in Russia is truly massive, with the species apparently nearly unbound by habitat, with their distribution only excluding them from the true Arctic zone, i.e. their range stops around the tree line. If not the most densely populated species, they almost certainly stand as Russia's most widely distributed owl species. From Russia, they are found throughout Central Asia, residing continuously in each nation from Kazakhstan down to Afghanistan. In Asia Minor, they are found broadly in Georgia, Azerbaijan and somewhat so in western and southern Turkey but is quite sporadic in distribution overall in Turkey. A spotty range also exists in the Middle East in Syria, Iraq, Lebanon, Israel, Jordan and western Iran, the species being found broadly only in north and western Iran. In South Asia, the Eurasian eagle-owl is found mostly often in northern Pakistan, northern Nepal and Bhutan and more marginally into far northern India. This species resides throughout Mongolia, almost the entirety of China (mainly absent only from southern Yunnan and southern Guangxi). From China and eastern Russia, the Eurasian eagle-owl is found throughout Korea, Sakhalin, the Kuril Islands and rarely into Japan in northern Hokkaido. Besides the Kurils, the farthest eastern part of the range for this species is in Magadan in the Russian Far East.

 

Behaviour

The Eurasian eagle-owl is largely nocturnal in activity, as are most owl species, with its activity focused in the first few hours after sunset and the last few hours before sunrise. In the northern stretches of its range, partial diurnal behaviour has been recorded, including active hunting in broad daylight during the late afternoon. In such areas, full nightfall is essentially non-existent at the peak of summer, so eagle-owls must presumably hunt and actively brood at the nest during daylight. The Eurasian eagle-owl has a number of vocalizations that are used at different times. It will usually select obvious topographic features such as rocky pinnacles, stark ridges and mountain peaks to use as regular song posts. These are dotted along the outer edges of the eagle-owl's territory and they are visited often but only for a few minutes at a time.

 

Vocal activity is almost entirely confined to the colder months from late fall through winter, with vocal activity in October through December mainly having territorial purposes and from January to February being primarily oriented towards courtship and mating purposes. Vocalizations in a Spanish study begin no sooner than 29 minutes after sunset and end no later than 55 minutes before sunrise. The territorial song, which can be heard at great distance, is a deep resonant ooh-hu with emphasis on the first syllable for the male, and a more high-pitched and slightly more drawn-out uh-hu for the female. It is not uncommon for a pair to perform an antiphonal duet. The widely used name in Germany as well as some other sections of Europe for this species is uhu due to its song. At 250–350 Hz, the Eurasian eagle-owls territorial song or call is deeper, farther-carrying and is often considering "more impressive" than the territorial songs of the great horned owl or even that of the slightly larger Blakiston's fish owl, although the horned owl’s call averages slightly longer in duration and the Blakiston's call is typically deeper.[7] Other calls include a rather faint, laughter-like OO-OO-oo and a harsh kveck-kveck. Intruding eagle-owls and other potential dangers may be met with a "terrifying", extremely loud hooo. Raucous barks not unlike those of ural owls or long-eared owls have been recorded but are deeper and more powerful than those species’ barks. Annoyance at close quarters is expressed by bill-clicking and cat-like spitting, and a defensive posture involves lowering the head, ruffling the back feathers, fanning the tail and spreading the wings.

 

The Eurasian eagle-owl rarely assumes the so-called "tall-thin position", which is when an owl adopts an upright stance with plumage closely compressed and may stand tightly beside a tree trunk. Among others, the long-eared owl is among the most often reported to sit with this pose. The great horned owl has been more regularly recorded using the tall-thin, if not as consistently as some Strix and Asio owls, and it is commonly thought to aid camouflage if encountering a threatening or novel animal or sound. The Eurasian eagle-owl is a broad-winged species and engages in a strong, direct flight, usually consisting of shallow wing beats and long, surprisingly fast glides. It has, unusually for an owl, also been known to soar on updrafts on rare occasions. The latter method of flight has led them to be mistaken for Buteos, which are smaller and quite differently proportioned. Usually when seen flying during the day, it is due to being disturbed or displaced from its roost by humans or mobbing animals, such as crows. Eurasian eagle-owls are highly sedentary, normally maintaining a single territory throughout their adult lives.

 

Eurasian eagle-owl are considered a completely non-migratory bird, as are all members of the Bubo genus excluding the snowy owl. Even those near the northern limits of their range, where winters are harsh and likely to bear little in food, the eagle-owl does not leave its native range. In 2020, a study presented evidence of a short distance distribution by adult eagle-owls in the fall subsequent to breeding, with 5 adults found to move over 20 km (12 mi) away from their nests. There are additionally claimed cases from Russia of Eurasian eagle-owls moving south for the winter, as the icebound, infamously harsh climate there may be too severe even for these hardy birds and their prey. Similarly, Eurasian eagle-owls living in the Tibetan highlands and Himalayas may in some anecdotal cases vacate their normal territories when winter hits and move south. In both of those examples, these are old, unverified reports and there is no evidence whatsoever of consistent, annual migration by Eurasian eagle-owls and the birds may eke out a living on their normal territories even in the sparsest times.

 

Dietary biology

Eurasian eagle-owls are strictly territorial and will defend their territories from interloping eagle-owls year around, but territorial calling appears to peak around October to early January. Territory size is similar or occasionally slightly greater than great horned owl: averaging 15 to 80 km2 (5.8 to 30.9 sq mi). Territories are established by the male eagle-owl, who selected the highest points in the territory from which to sing. The high prominence of singing perches allows their song to be heard at greater distances and lessens the need for potentially dangerous physical confrontations in the areas where territories may meet. Nearly as important in territorial behaviour as vocalization is the white throat patch. When taxidermied specimens with flared white throats were placed around the perimeter of eagle-owl territories, male eagle-owls reacted quite strongly and often attacked the stuffed owl, reacting more mildly to a stuffed eagle-owl with a non-flared white throat. Females were less likely to be aggressive to mounted specimens and did not seem to vary in their response whether exposed to the specimens with or without the puffed up white patch. In January and February, the primary function for vocalization becomes for the purpose of courtship. More often than not, eagle-owls will pair for life but usually engage in courtship rituals annually, most likely to re-affirm pair bonds. When calling for the purposes of courtship, males tend to bow and hoot loudly but do so in a less contorted manner than the male great horned owl. Courtship in the Eurasian eagle-owl may involve bouts of "duetting", with the male sitting upright and the female bowing as she calls. There may be mutual bowing, billing and fondling before the female flies to a perch where coitus occurs, usually taking place several times over the course of a few minutes.

 

Nests

The male selects breeding sites and advertises their potential to the female by flying to them and kneading out a small depression (if soil is present) and making staccato notes and clucking noises. Several potential sites may be presented, with the female selecting one. In Baden-Wurttenberg, Germany, the amount of male nest site visits were found to increase in time spent over the pre-laying breeding season from a mean of 29 minutes to 3 hours with frequent incubation like sitting by the male. Like all owls, Eurasian eagle-owls do not build nests or add material but nest on the surface or material already present. Eurasian eagle-owls normally nest on rocks or boulders, most often utilizing cliff ledges and steep slopes, as well as crevices, gullies, holes or caves. Rocky areas that also prove concealing woodlots as well as, for hunting purposes, that border river valleys and grassy scrubland may be especially attractive. If only low rubble is present, they will nest on the ground between rocks. Often, in more densely forested areas, they've been recorded nesting on the ground, often among roots of trees, under large bushes and under fallen tree trunks. Steep slopes with dense vegetation are preferred if nesting on the ground, although some ground nests are surprisingly exposed or in flat spots such as in open spots of the taiga, steppe, ledges of river banks and between wide tree trunks. All Eurasian eagle-owl nests in the largely forested Altai Krai region of Russia were found to be on the ground, usually at the base of pines. This species does not often use other bird’s nests as does the great horned owl, which often prefers nests built by other animals over any other nesting site. The Eurasian eagle-owl has been recorded in singular cases using nests built by common buzzards (Buteo buteo), golden eagle, greater spotted (Clanga clanga) and white-tailed eagles (Haliaeetus albicilla), common ravens (Corvus corax) and black storks (Ciconia nigra). Among the eagle-owls of the fairly heavily wooded wildlands of Belarus, they more commonly utilize nests built by other birds than most eagle-owls, i.e. stork or accipitrid nests, but a majority of nests are still located on the ground. This is contrary to the indication that ground nests are selected only if rocky areas or other bird nests are unavailable, as many will utilize ground nests even where large bird nests seem to be accessible. Tree holes being used for nesting sites are even more rarely recorded than nests constructed by other birds. While it may be assumed that the eagle-owl is too large to utilize tree hollows, when other large species like the great grey owl have never been recorded nesting in one, the even more robust Blakiston's fish owl nests exclusively in cavernous hollows. The Eurasian eagle-owl often uses the same nest site year after year.

 

Parental behaviour

In Engadin, Switzerland, the male eagle-owl alone hunts until the young are 4 to 5 weeks old and the female spends all her time brooding at the nest. After this point, the female gradually resumes hunting from both herself and the young and thus provides a greater range of food for the young. While it may seem contrary to the species’ highly territorial nature, there is one verified cases of polygamy in Germany, with a male apparently mating with two females, and cooperative brooding in Spain, with a third adult of undetermined sex helping a breeding pair care for the chicks. The response of Eurasian eagle-owls to humans approaching at the nest is quite variable. The species is often rather less aggressive than some other owls, including related species like the spot-bellied eagle-, great horned and snowy owls, many of the northern Strix species, and even some rather smaller owl species, which often fearlessly attack any person found to be nearing their nests. Occasionally, if a person climbs to an active nest, the adult female eagle-owl will do a distraction display, in which they feign an injury. This is an uncommon behavior in most owls and is most often associated with small birds trying to falsely draw the attention of potential predators away from their offspring. More commonly, the adults withdraw to a safe distance, as their nests are usually well-camouflaged. Occasionally if cornered both adults and nestlings will do an elaborate threat display, also rare in owls in general, in which the eagle-owls raise their wings into a semi-circle and puff up their feathers, followed by a snapping of their bills. Apparently, eagle-owls of uncertain and probably exotic origin in Britain are likely to react aggressively to humans approaching the nest. Also, aggressive encounters involving eagle-owls around their nest, despite being historically uncommon, apparently have increased in recent decades in Scandinavia. The discrepancy of aggressiveness at the nest between the Eurasian eagle-owl and its Nearctic counterpart may be correlated to variation in the extent of nest predation that the species endured during the evolutionary process.

 

Eggs and offspring development

The eggs are normally laid at intervals of three days and are incubated only by the female. Laying generally begins in late winter but may be later in the year in colder habitats. During the incubation period, the female is brought food at the nest by her mate. A single clutch of white eggs is laid; each egg can measure from 56 to 73 mm (2.2 to 2.9 in) long by 44.2 to 53 mm (1.74 to 2.09 in) in width, and will usually weigh about 75 to 80 g (2.6 to 2.8 oz). In Central Europe, eggs average 59.8 mm × 49.5 mm (2.35 in × 1.95 in), and in Siberia, eggs average 59.4 mm × 50.1 mm (2.34 in × 1.97 in). Their eggs are only slightly larger than those of snowy owls and the nominate subspecies of great horned owl, while similar in size to those of spot-bellied eagle-owls and Blakiston's fish owls. The Eurasian eagle-owl’s eggs are noticeably larger than those of Indian eagle-owl and pharaoh eagle-owls. Usually clutch size is one or two, rarely three or four, and exceptionally to six. The average number of eggs laid varies with latitude in Europe. Clutch size ranges from 2.02 to 2.14 in Spain and the massifs of France, and 1.82 to 1.89 in central Europe and the eastern Alps; in Sweden and Finland, the mean clutch size is 1.56 and 1.87, respectively. While variation based on climate is not unusual for different wide-ranging palearctic species, the higher clutch size of western Mediterranean eagle-owls is also probably driven by the presence of lagomorphs in the diet, which provide high nutritional value than most other regular prey. The average clutch size, attributed as 2.7, was the lowest of any European owl per one study. One species was attributed with an even lower clutch size in North America, the great grey owl with a mean of 2.6, but the mean clutch size was much higher for the same species in Europe, at 4.05.

 

In Spain, incubation is from mid-January to mid-March, hatching and early nestling period is from late March to early April, fledging and postfledging dependence can range from mid-April to August, and territorial/courtship is anytime hereafter; i.e. the period between the beginning of juvenile dispersal to egg laying; from September to early January. The same general date parameters were followed in southern France. In the Italian Alps, the mean egg-laying date was similarly February 27, but the young were more likely to be dependent later, as all fledglings were still being cared for by the end of August, and some even lingered under parental care until October. In northern climes, the breeding season shifts somewhat later by as much as a month so that egg laying may be as late as late March or early April. Nonetheless, the Eurasian eagle-owl is one of the earliest nesting bird species in Europe or northern, temperate Asia.

 

The first egg hatches after 31 to 36 days of incubation. The eggs hatch successively; although the average interval between egg-laying is 3 days, the young tend to hatch no more than a day or two apart. Like all owls that nest in the open, the downy young are often a mottled grey with some white and buff, which provides camouflage. They open their eyes at 4 days of age. The chicks grow rapidly, being able to consume small prey whole after roughly 3 weeks. In Andalusia, the most noticeable development of the young before they leave the nest was the increase of body size, which was the highest growth rate of any studied owl and faster than either snowy or great horned owls. Body mass increased fourteen times over from 5 days old to 60 days old in this study. The male continues to bring prey, leaving it on or around the nest, and the female feeds the nestlings, tearing up the food into suitably sized pieces. The female resumes hunting after about 3 weeks, which increases the food supply to the chicks. Many nesting attempts produce two fledglings, indicating that siblicide is not as common as in other birds of prey, especially a few species of eagles. In Spain, males are thought to be the first egg laid to reduce the likelihood of sibling aggression due to the size difference, thus the younger female hatchling is less likely to be killed since it is similar in size to its older sibling.

 

Apparently, the point at which the chicks venture out of the nest is driven by the location of the nest. In elevated nest sites, chicks usually wander out of the nest at 5 to as late as 7 weeks of age, but have been recorded leaving the nest if the nest is on the ground as early as 22 to 25 days old. The chicks can walk well at 5 weeks of age and by 7 weeks are taking short flights. Hunting and flying skills are not tested prior to the young eagle-owls leaving the nest. Young Eurasian eagle-owls leave the nest by 5–6 weeks of age and typically can be flying weakly (a few metres) by about 7–8 weeks of age. Normally, they are cared for at least another month. By the end of the month, the young eagle-owls are quite assured fliers. A few cases have been confirmed of adult eagle-owls in Spain feeding and caring for postfledgling juvenile eagle-owls that were not their own.

  

Like many large owls, Eurasian eagle-owls leave the nest while still in a functionally flightless state and with large amounts of second down still present, but will fly shortly thereafter.

A study from southern France found the mean number of fledglings per nest was 1.67. In central Europe, the mean number of fledglings per nest was between 1.8 and 1.9. The mean fledgling rate in the Italian Alps was 1.89, thus being similar. In the Italian Alps, heavier rainfall during breeding decreased fledgling success because it inhibited the ability of the parents to hunt and potentially exposed nestlings to hypothermia. In the reintroduced population of eagle-owls in Eifel, Germany, occupied territories produced an average of 1.17 fledglings, but not all occupying pairs attempted to breed, with about 23% of those attempting to breed being unsuccessful. In slightly earlier studies, possibly due to higher persecution rates, the mean number of young leaving the nest was often lower, such as 1.77 in Bavaria, Germany, 1.1 in lower Austria, and 0.6 in southern Sweden. An experimental supplemental feeding program to young eagle-owls on two small Norwegian islands were found to increase mean numbers of fledglings from a mean of about 1.2 to 1.7 despite evidence that increased human activity near the nest decreased owlet survivability. While sibling owls are close in the stage between leaving the nest and fully fledged, about 20 days after leaving the nest, the family unit seems to dissolve and the young disperse quickly and directly. All told, the dependence of young eagle-owls on their parents lasts for 20 to 24 weeks. Independence in central Europe is from September to November. The young leave their parents' care normally on their own, but are also sometimes chased away by their parents. The young Eurasian eagle-owls reach sexual maturity by the following year, but do not normally breed until they can establish a territory at around 2–3 years old. Until they are able to establish their own territories, young eagle-owls spend their lives as nomadic "floaters", and while they also call, select inconspicuous perch sites unlike breeding birds. Male floaters are especially wary about intrusion into an established territory to avoid potential conspecific aggression.

 

Status

he Eurasian eagle-owl has a very wide range across much of Europe and Asia, estimated to be about 32,000,000 km2 (12,000,000 sq mi). In Europe, the population is estimated at 19,000 to 38,000 breeding pairs, and in the whole world around 250,000 to 2,500,000 individual birds. The population trend is thought to be decreasing because of human activities, but with such a large range and large total population, the International Union for Conservation of Nature has rated the bird as being of least concern. Although roughly equal in adaptability and wideness of distribution, the great horned owl, with a total estimated population up to 5.3 million individuals, apparently has a total population that is roughly twice that of the Eurasian eagle-owl. Numerous factors, including a shorter history of systematic persecution, lesser sensitivity to human disturbance while nesting, somewhat greater ability to adapt to marginal habitats and widespread urbanization, and slightly smaller territories may play into the horned owls greater numbers in modern times. Eurasian eagle-owls are listed in Appendix II of the Convention on International Trade in Endangered Species (CITES) meaning international trade (including in parts and derivatives) is regulated.

 

Longevity

The Eurasian eagle-owl surely is one of the longest-living owls on average. The eagle-owl can live for up to 20 years in the wild. At one time, the oldest ringed eagle-owl was considered a 19-year-old specimen. Some studies posited that in protected areas, lifespans ranging up to 15–20 years may not be uncommon. A record-breaking specimen banded in the wild was subsequently found to survive to be 27 years and 9 months old. Like many other bird species in captivity, they can live much longer without having to endure difficult natural conditions, and have possibly survived up to 68 years in zoo collections. Healthy adults normally have no natural predators, thus are considered apex predators. The leading causes of death for this species are man-made; electrocution, traffic accidents, and shooting frequently claim the lives of eagle-owls.

 

Anthropogenic mortality

Electrocution was the greatest cause of mortality in 68% of 25 published studies, and accounted, on average, for 38.2% of the reported eagle-owl deaths. This was particularly true in the Italian Alps, where the number of dangerous, uninsulated pylons near nests was extremely high, but is highly problematic almost throughout the species’ European distribution. In one telemetry study, 55% of 27 dispersing young were electrocuted within 1 year of their release from captivity, while electrocution rates of wild-born young are even higher. Mortality in the Swiss Rhine Valley was variable, in radio-tagged, released individuals, most died as a result of starvation (48%) rather than human-based causes, but 93% of the wild, untagged individuals found dead were due to human activities, 46% due to electrocution, and 43% due to collision with vehicles or trains. Insulation of pylons is thought to result in a stabilisation of the local population due to floaters taking up residence in unoccupied territories that formerly held deceased eagle-owls. Eurasian eagle-owls from Finland were found mainly to die due to electrocution (39%) and collisions with vehicles (22%). Wind turbine collisions can also be a serious cause of mortality locally.

 

Eagle-owls have been singled out historically as a threat to game species, thus to the economic well-being of landowners, game-keepers, and even governmental agencies, and as such, have been singled out for widespread persecution. Local extinctions of Eurasian eagle-owls have been primarily due to persecution. Examples of this include northern Germany in 1830, the Netherlands sometimes in the late 19th century, Luxembourg in 1903, Belgium in 1943, and central and western Germany in the 1960s. In trying to determine causes of death for 1476 eagle-owls from Spain, most were unknown and undetermined types of trauma. The largest group that could be determined, 411 birds, was due to collisions, more than half of which were from electrocution, while 313 were due to persecution, and merely 85 were directly attributable to natural causes. Clearly, while pylon safety is perhaps the most serious factor to be addressed in Spain, persecution continues to be a massive problem for Spanish eagle-owls. Of seven European nations where modern Eurasian eagle-owl mortality is well-studied, continual persecution is by far the largest problem in Spain, although also continues to be serious (often comprising at least half of studied mortality) in France. From France and Spain, nearly equal numbers of eagle-owls are poisoned (for which raptors might not be the main target), or shot intentionally.

 

Conservation and reintroductions

While the eagle-owl remains reasonably numerous in some parts of its habitat where nature is still relatively little disturbed by human activity, such as the sparsely populated regions of Russia and Scandinavia, concern has been expressed about the future of the Eurasian eagle-owl in Western and Central Europe. There, very few areas are not heavily modified by human civilisation, thus exposing the birds to the risk of collisions with deadly man-made objects (e.g. pylons) and a depletion of native prey numbers due to ongoing habitat degradation and urbanisation.

 

In Spain, long-term governmental protection of the Eurasian eagle-owl seems to have no positive effect on reducing the persecution of eagle-owls. Therefore, Spanish conservationists have recommended to boost education and stewardship programs to protect eagle-owls from direct killing by local residents. Unanimously, biologists studying eagle-owl mortality and conservation factors have recommended to proceed with the proper insulation of electric wires and pylons in areas where the species is present. As this measure is labour-intensive and therefore rather expensive, few efforts have actually been made to insulate pylons in areas with few fiscal resources devoted to conservation such as rural Spain. In Sweden, a mitigation project was launched to insulate transformers that are frequently damaged by eagle-owl electrocution.

 

Large reintroduction programs were instituted in Germany after the eagle-owl was deemed extinct in the country as a breeding species by the 1960s, as a result of a long period of heavy persecution. The largest reintroduction there occurred from the 1970s to the 1990s in the Eifel region, near the border with Belgium and Luxembourg. The success of this measure, consisting in more than a thousand eagle-owls being reintroduced at an average cost of US$1,500 per bird, is a subject of controversy. Those eagle-owls reintroduced in the Eifel region appear to be able to breed successfully, and enjoy nesting success comparable with wild eagle-owls from elsewhere in Europe. Mortality levels in the Eifel region, though, appear to remain quite high due to anthropogenic factors. Also, concerns exist about a lack of genetic diversity of the species in this part of Germany. Apparently, the German reintroductions have allowed eagle-owls to repopulate neighbouring parts of Europe, as the breeding populations now occurring in the Low Countries (the Netherlands, Belgium, and Luxembourg) are believed to be the result of influx from regions further to the east. Smaller reintroductions have been done elsewhere, and the current breeding population in Sweden is believed to be primarily the result of a series of reintroductions. Conversely to numerous threats and declines incurred by Eurasian eagle-owls, areas where human-dependent, non-native prey species such as brown rats (Rattus norvegicus) and rock pigeons (Columba livia) have flourished, have given the eagle-owls a primary food source and allowed them occupy regions where they were once marginalized or absent.

 

Occurrence in Great Britain

The Eurasian eagle-owl at one time occurred naturally in Great Britain. Some, including the RSPB, have claimed that it had disappeared about 10,000–9,000 years ago, after the last ice age, but fossil remains found in Meare Lake Village indicate the eagle-owl occurring as recently as roughly 2,000 years ago in the fossil record. The lack of presence of the Eurasian eagle-owl in British folklore or writings in recent millennium may indicate the lack of occurrence by this species there. The flooding of the land bridge between Britain and continental Europe may have been responsible for their extirpation as they only disperse over limited distances, although early human persecution presumably played a role as well. Some reportages of eagle-owls in Britain have been revealed to actually be great horned owls or Indian eagle-owls, the latter a particularly popular owl in falconry circuits.[110] Some breeding pairs do still occur in Britain, though the exact number of pairs and individuals is not definitely known. The World Owl Trust stated that they believe some eagle-owls occurring in North England and Scotland are naturally occurring, making the flight of roughly 350 to 400 km (220 to 250 mi) from the west coast of Norway to Shetland and the east coast of Scotland, as well as possibly from the coasts of the Netherlands and Belgium to the south. Although not migratory, eagle-owls can disperse some notable distances in young birds seeking a territory.

 

Prior studies of eagle-owl distribution have indicated a strong reluctance to cross large bodies of water in the species. Many authorities state that the Eurasian eagle-owls occurring in Britain are individuals that have escaped from captivity. While, until the 19th century, wealthy collectors may have released unwanted eagle-owls, despite press to the contrary, no evidence of any organization or individual intentionally releasing eagle-owls recently with the intent to establish a breeding population has been found. Many feel that the eagle-owl would be classified as an "alien" species. Due to its predatory abilities, many, especially those in the press, have expressed alarm of their effect on "native" species. From 1994 to 2007, 73 escaped eagle-owls were not registered as returned, while 50 escapees were recaptured. Several recorded breeding attempts have been studied, and most were unsuccessful, due in large part to incidental disturbance by humans and some due to direct persecution, with eggs having been smashed.

 

Effect on conservation-dependent species

As highly opportunistic predators, Eurasian eagle-owls hunt almost any appropriately sized prey they encounter. Most often, they take whatever prey is locally common and can take a large number of species considered harmful to human financial interests, such as rats, mice, and pigeons. Eurasian eagle-owls do take rare or endangered species, as well. Among the species considered at least vulnerable (up to critically endangered as in the mink and eel, both heavily overexploited by humans) to extinction known to be hunted by Eurasian eagle-owls are Russian desman (Desmana moschata) Pyrenean desman (Galemys pyrenaicus), barbastelle (Barbastella barbastellus), European ground squirrel (Spermophilus citellus), southwestern water vole (Arvicola sapidus), European mink (Mustela lutreola), marbled polecat (Vormela peregusna), lesser white-fronted goose (Anser erythrops), Egyptian vulture (Neophron percnopterus), greater spotted eagle (Clanga clanga), eastern imperial eagle (Aquila heliaca), saker falcon (Falco cherrug), houbara bustard (Chlamydotis undulata), great bustard (Otis tarda), spur-thighed tortoise (Testudo graeca), Atlantic cod (Gadus morhua), European eel (Anguilla anguilla) and lumpfish (Cyclopterus lumpus).

Includes Battle Japan, Tyranno Ranger, Red Racer, Hurricane Red and Gosei Red

Action in the rain from Friday at the 2017 Goodwood Revival

Since I didn't include him in the name post, I thought he should get equal time. This is also his before picture. Paul and I are going to give him a haircut tomorrow. We're fairly comfortable with his body but his head/face make me nervous, especially since he's not good about holding still. We just got tired of paying $50+ for bad haircuts. If only Annie lived here.

 

About his name, those of you who are Facebook friends may have noticed that Cooper was my maiden name. So it seemed like a no-brainer that would be his name. My dad always thought we named Cooper after him to harass him because he didn't like dogs. Actually, it would have served him right.

 

You may notice that Cooper has a runny nose. Since we moved to Arizona, both he and Ruby have developed allergies. We heard through a friend that her canine allergist said it's common for adult dogs who move to a new area to suffer from allergies. So far it hasn't been bad, just occasional runny noses and slightly itchy skin. We've been trying local honey for all of us since that's supposed to help.

U.S. 27 Phase 1 - Begins March 27, Finishes Fall 2017

 

The first phase of construction will include:

~ straightening the alignment of southbound U.S. 27 near the intersection of North C Street,

~ rebuilding State Road 121/North J Street approaching U.S. 27, and

~ rebuilding both directions of U.S. 27 between North B Street and the Whitewater River, excluding the bridge over the railroad.

Work will reduce U.S. 27 to one lane in each direction between North B Street and the Whitewater River bridge. Multiple traffic shifts will be required to maintain one lane in each direction and rebuild the roadway one side at a time. Access to and from intersecting streets will be temporarily closed while that side of the roadway is being rebuilt.

 

U.S. 27 Phase 2 - Begins Late April, Finishes Summer 2018

The second phase of construction will include rebuilding the west half of southbound U.S. 27 between North C Street and the South O Street overpass. Traffic will be reduced to one lane and shifted to the east side of the road. A short section between South C and F streets will temporarily remain shifted to the west half of the road.

 

Multiple traffic lane shifts will be required to maintain one open lane and rebuild the roadway one side at a time. Access to and from intersecting streets will be temporarily closed while that side of the roadway is being rebuilt.

 

Weather permitting, contractors intend to finish one half of the southbound U.S. 27 rebuild this year. Once complete, one lane of traffic will be shifted to the west side of the road and the eastern half will be finished in 2018.

 

U.S. 27 construction will require lane closures and traffic shifts on U.S. 40 (North A and South A streets) as the intersections are rebuilt later this summer.

Back at the Goodwood Festival of Speed today, a free ticket as a reward for those involved in a work project that went very well. Bigger crowds for Sunday, which I used for these hit-and-miss panning shots