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Known for their huge bodies, unique trunks, and long lives, elephants are the largest land animals in the world. They live in social groups called herds which are composed of related females, called cows, and their offspring. The leader of the herd is a female, usually the oldest, most experienced member of the herd. She decides the herd’s activities. The females stay with their female relatives for life, but young males leave the herd when they reach puberty, either traveling alone or in bachelor groups. Females will practice motherhood in the herd by being “aunts” to the calves.
Elephants will display dominance by raising their head, trunk and ears.They will also shake their heads and make trumpeting noises, followed by loud rumbles. They use their huge ears as fans, flapping them to cool themselves off, and their trunks perform a variety of jobs from spraying water and dust to picking up tiny objects to heavy lifting. Their tusks are actually their incisor teeth and they use them to dig, lift things, and protect themselves.
DIET
Grass, herbs, bark of various trees, leaves and frui
By far the most impressive feature of the hippo is its enormous head, with its mammoth mouth opening to reveal a set of long, razor-sharp teeth. No one is going to mess with this little guy...
The mouth of an adult hippo is roomy enough for 36 teeth, including tusks that can grow up to 2 feet long. Incisors and bottom incisors thrust vertically to the front of the mouth to stab foes. More commonly, male hippos size one another up by opening their mouth up to 150 degrees, stretching it to its full four to five feet length.
(Source: Jodi Thornton O'Connell)
Tusks are teeth and just like our teeth, if one is broken, it stays broken. But unlike our teeth, a tusk can continue growing from the root if that isn't damaged. It's not unusual to see an elephant with only one tusk because the other was injured to the point that it stopped growing.
The tusk is the equivalent of our incisor teeth . It is made of ivory, a material soft enough to be carved, which is the root of the poaching problem.
Back from a 2 weeks trip in Omo Valley, in Ethiopia. 2500 km in the mud, sand, and bush to see the tribes, and to avoid the too touristic ones. on this first picture, a Mursi woman, the most agressive tribe i met.
Agressive in the way that when you come to see them (after hours of 4x4 on a very bad track), they only think about the money they can get from you! The craziest thing is that they ask for a fee to park the car (20 euros!) at the entrance of the village!!
They use those wild pigs tusks as a decoration. the main problem is that they tend to use more and more things to attract photographers!
You may have seen or read books about this area by great photographers those last years, and many of the people inside are just "disguised"...
But at the end, those tribes really live like in the primitives times, whitout anything around apart their catle, and still fighting with other tribes to catch cows and women...
This is a daily reality in this area of the world!
7 000 tourists visit the mursis every year. It makes an average of 20 by day in a very big area, so it's very few.But as 70% are from Spain and pay a low fee to the tour operator, some tribes start to settle close to "hotels" or campings to get money from the photographers.
I had the chance to go in Omo valley with a guide who avoid those touristic spots.
Mursi woman with her giant lip plate, a sign of beauty in Mursi tribe, like in Surma one. When they are ready to marry, they start to make a hole in the lip with a wood stick.
It will be kept for one night , and is removed to put a bigger one. This is very painful at this time... Few months after, the lip plate has its full size, and the girl is seen as beautiful by the men.
The lip plate made of wood or terracotta, and they have to remove the lower incisors to let some space for the disc. it's amazing to see them speak without any trouble, put it and remove it as a classic jewel.
Sometimes the lip is broken by the pressure of the lip plate. This is a very big problem for the girl cos men will consider her as ugly, she won't be able to marry anyone in the tribe apart the old men or the sick people...
The women are shaved, like the men, cos they hate hairiness!
© Eric Lafforgue
The Caucasian squirrel - Sciurus anomalus ; The Caucasian squirrel or Persian squirrel, is a tree squirrel in the genus Sciurus found in temperate broadleaf and mixed forests in south-western Asia.
The species is usually said to have first been described in 1778 by Johann Friedrich Gmelin in the 13th edition of Systema Naturae,and named Sciurus anomalus. However, some authors argue that this work was actually published in 1788, and that the true first description was made by Johann Anton Güldenstädt in 1785.
Description -
Caucasian squirrels are small tree squirrels, with a total length of 32 to 36 cm (13 to 14 in), including the 13 to 18 cm (5.1 to 7.1 in) tail, and weighing 250 to 410 g (8.8 to 14.5 oz). The color of the upper body fur ranges from greyish brown to pale grey, depending on the subspecies, while that of the underparts is rusty brown to yellowish, and that of the tail, yellow brown to deep red. The claws are relatively short, compared with those of other tree squirrels, and females have either eight or ten teats.
Samuel Griswold Goodrich described the Caucasian squirrel in 1885 as "Its color is grayish-brown above, and yellowish-brown below".
Physical Description -
Caucasian squirrels have a dental formula of incisors 1/1, canines 0/0, premolars 1/1, and molars 3/3, totaling 20. They have four fingered fore feet and five fingered hind feet. Sex differences in body length or mass are not evident.
Distribution and habitat -
Caucasian squirrels are native to south-western Asia, where they are found from Turkey, and the islands of Gökçeada and Lesbos in the west, Iran in the southeast, and as far as Israel and Jordan in the south.It is one of only two species of the genus Sciurus to be found on Mediterranean islands,and, although Eurasian red squirrels have been recently introduced to some areas, is the only species of Sciurus native to the wider region.
The species mainly lives in forested areas dominated by oak, pine, and pistachio, up to altitudes of 2,000 metres (6,600 ft).
Biology and behavior -
The squirrels are diurnal, and solitary, although temporary groups may forage where food is plentiful. Their diet includes nuts, seeds, tree shoots, and buds,with the seeds of oak and pine being particularly favored. Like many other squirrels, they cache their food within tree cavities or loose soil, with some larders containing up to 6 kg (13 lb) of seeds. They live in trees, where they make their dens, but frequently forage on the ground, and are considered less arboreal than Eurasian red squirrels. They commonly nest in tree hollows lined with moss and leaves, and located 5 to 14 m (16 to 46 ft) above the ground, but nests are also sometimes found under rocks or tree roots. Their alarm call is high-pitched, and said to resemble the call of the European green woodpecker, and they mark their territories with urine and dung.
Breeding occurs throughout the year, but is more common in spring or autumn. Litters range from two to seven, with three or four being typical, and the young are fully mature by five or six months of age.
Conservation -
A survey in 2008 found that the species remained abundant within Turkey, however declines are noted in population within the Levant region. The guides for a survey in 1993 in Israel stated that they considered the species to be nearly extinct within the area studied. Whilst the Caucasian squirrel is threatened by poaching and deforestation, the declines recorded are not sufficient to qualify them as anything other than "Least Concern" by the International Union for Conservation of Nature.[1] Hunting of the species is banned by the Central Hunting Commission, and the Caucasian squirrel is protected by the Bern Convention and the EU Habitats Directive.
This information is sourced from "Wikipedia".
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Surma or Suri are sedentary pastoral people living in south west of Ethiopia, on the western bank of the Omo River, in the Kibish and Tulgit area.
These breeders tribal groups have a cattle centred culture. They grow cabbage, beans, yams, tobacco and coffee and breed their cattle, mostly cows, on their traditional lands, located in the Omo Valley. Cows are tremendously important for the Suris. They do not see cattle simply as a material asset but as a life sustaining and meaningful companion. Suri even sing songs for them and make fires to warm them. These cows are not bred for their meat and are usually not killed unless they are needed for ceremonial purposes. They use their milk and their blood, which they both drink. Cows also have a social and symbolic meaning in Suri’s society. Suri men are judged on how much cattle they own. In desperate times, Suri men can risk their lives to steal cattle from other tribes. The average male in the Suri tribe owns 40 cows. Every young male is named after their cattle, which they have to look after since the age of 8. Cows are given to the bride’s family after the wedding ceremony. Usually 20 cows and they offer also a Kalashnikov as wedding gift.
This central role of the cow in their way of life accounts for the fierce independence they want to preserve and explains their warlike culture. Indeed, it’s quite common to see men and even women carrying Kalashnikovs, which are part of the daily life. Their remote homeland has always been a place of traditional rivalries with the neighbouring tribes such as the Bume (Nyangatom) or the Toposa from Sudan who regularly team up to raid the Suri’s cattle. These fights, and even sometimes battles, have become quite bloody since automatic firearms have become available from the parties in the Sudanese Civil War. This conflict has pushed neighbouring tribes into Suri’s land and is a constant competition to keep and protect their territory and owns.
Like their neighbours, the Surma also paint their bodies. They create a variety of designs on their necked bodies using their fingertips, which helps them to expose their dark skins. The painting could have both a beautifying and opponent frightening purpose. As one studies these body paintings whirls, stripes, flower and star designs are noticeable. Surma men who are generally believed to be expert artists also paint the girls.
A ritual chief in the villages known as the Komoru, dressed in colourful robes and wearing a crown of baboon fur leads the Suri. Village life is largely communal, sharing the produce of the cattle (milk and blood). The men in an assembly take decisions of the village.
Although their traditional remoteness and autarky is threatened, only few Surma are familiar with Amharic, the official language of Ethiopia, and their literacy level is very low. Lip plate and Donga stick fight are the two typical distinctive features of these people, which they share with the neighbouring Mursi people.
Suri women wear giant lip plate, a sign of beauty, like in Mursi tribe, and also a prime attraction for tourists which help to sustain a view of them, in guidebooks and travel articles, as an untouched people, living in one of the last wildernesses of Africa. When they are ready to marry, teenagers start to make a hole in the lower lip with a wood stick. It will be kept for one night, and is removed to put a bigger one. This is very painful at this time. Few months after, the lip plate has its full size, and the men see the girl as beautiful. The lip plate is made of wood or terracotta. They have to remove the lower incisors to let some space for the disc. Sometimes the pressure of the plate breaks the lip. This is a big problem for the girl because men will consider her as ugly, she won't be able to marry anyone in the tribe apart the old men or the sick people.
The Sagenai, called also donga ritual is a combat that brings both wounds and honor to both the winner and loser. The men bodies are decorated with ritual drawings and their heads are protected by a sort of helmet. For the boys participating to the donga, this challenge is a true moment of glory. The combat is taking place in a middle of a circle made by the crowd. The rules are simple and can be summed up as follows: the person who manages to stay on his feet is the winner, and one must absolutely not kill his opponent. The winner will be honored by the entire tribe and can choose girls to date.
The lands of the Suri are stolen by the Ethiopian government to be rented to foreign companies. A Malaysian company, Lim Slow Jin, runs the Koka plantation near Kibish on the east side of the Omo. The lands are confiscated and rented out for 1 euro per hectare for a year.
© Eric Lafforgue
There is a common dental procedure that nearly every dentist will tell you is completely safe, despite the fact that scientists have been warning of its dangers for more than 100 years.
What is this dental procedure?
The root canal.
Root-canaled teeth are essentially “dead” teeth that can become silent incubators for highly toxic anaerobic bacteria that can, under certain conditions, make their way into your bloodstream to cause a number of serious medical conditions—many not appearing until decades later.
Most of these toxic teeth feel and look fine for many years, which make their role in systemic disease even harder to trace back.
Sadly, the vast majority of dentists are oblivious to the serious potential health risks they are exposing their patients to, risks that persist for the rest of their patients’ lives.
The American Dental Association claims root canals have been proven safe, but they have NO published data or actual research to substantiate this claim.
Dr. Weston Price, regarded by many as the greatest dentist of all time, who, more than a century ago, made the connection between root-canaled teeth and disease.
Dr. Price was a dentist and researcher who traveled the world to study the teeth, bones, and diets of native populations living without the “benefit” of modern food. Around the year 1900, Price had been treating persistent root canal infections and became suspicious that root-canaled teeth always remained infected, in spite of treatments. Then one day, he recommended to a woman, wheelchair bound for six years, to have her root canal tooth extracted, even though it appeared to be fine.
She agreed, so he extracted her tooth and then implanted it under the skin of a rabbit. The rabbit amazingly developed the same crippling arthritis as the woman and died from the infection 10 days later. But the woman, now free of the toxic tooth, immediately recovered from her arthritis and could now walk without even the assistance of a cane.
Price discovered that it’s mechanically impossible to sterilize a root-canaled (e.g. root-filled) tooth. He then went on to show that many chronic degenerative diseases originate from root-filled teeth—the most frequent being heart and circulatory diseases. He actually found 16 different causative bacterial agents for these conditions. But there were also strong correlations between root-filled teeth and diseases of the joints, brain and nervous system. Dr. Price went on to write two groundbreaking books in 1922 detailing his research into the link between dental pathology and chronic illness. Unfortunately, his work was deliberately buried for 70 years, until finally one endodontist named George Meinig recognized the importance of Price’s work and sought to explain the truth.
Dr. Meinig, a native of Chicago, was a captain in the U.S. Army during World War II before moving to Hollywood to become a dentist for the stars. He eventually became one of the founding members of the American Association of Endodontists (root canal specialists).
In the 1990s, he spent 18 months immersed in Dr. Price’s research. In June of 1993, Dr. Meinig published the book Root Canal Cover-Up, which continues to be the most comprehensive reference on this topic today.
Your teeth are made of the hardest substances in your body.
In the middle of each tooth is the pulp chamber, a soft living inner structure that houses blood vessels and nerves. Surrounding the pulp chamber is the dentin, which is made of living cells that secrete a hard mineral substance. The outermost and hardest layer of your tooth is the white enamel, which encases the dentin.
The roots of each tooth descend into your jawbone and are held in place by the periodontal ligament. In dental school, dentists are taught that each tooth has one to four major canals. However, there are accessory canals that are never mentioned. Literally miles of them!
Just as your body has large blood vessels that branch down into very small capillaries, each of your teeth has a maze of very tiny tubules that, if stretched out, would extend for three miles. Weston Price identified as many as 75 separate accessory canals in a single central incisor (front tooth). Microscopic organisms regularly move in and around these tubules, like gophers in underground tunnels.
When a dentist performs a root canal, he or she hollows out the tooth, then fills the hollow chamber with a substance (called guttapercha), which cuts off the tooth from its blood supply, so fluid can no longer circulate through the tooth. But the maze of tiny tubules remains. And bacteria, cut off from their food supply, hide out in these tunnels where they are remarkably safe from antibiotics and your own body’s immune defenses.
Under the stresses of oxygen and nutrient deprivation, these formerly friendly organisms morph into stronger, more virulent anaerobes that produce a variety of potent toxins. What were once ordinary, friendly oral bacteria mutate into highly toxic pathogens lurking in the tubules of the dead tooth, just awaiting an opportunity to spread.
No amount of sterilization has been found effective in reaching these tubules—and just about every single root-canaled tooth has been found colonized by these bacteria, especially around the apex and in the periodontal ligament. Oftentimes, the infection extends down into the jawbone where it creates cavitations—areas of necrotic tissue in the jawbone itself.
Cavitations are areas of unhealed bone, often accompanied by pockets of infected tissue and gangrene. Sometimes they form after a tooth extraction (such as a wisdom tooth extraction), but they can also follow a root canal. According to Weston Price Foundation, in the records of 5,000 surgical cavitation cleanings, only two were found healed.
And all of this occurs with few, if any, accompanying symptoms. So you may have an abscessed dead tooth and not know it. This focal infection in the immediate area of the root-canaled tooth is bad enough, but the damage doesn’t stop there.
As long as your immune system remains strong, any bacteria that stray away from the infected tooth are captured and destroyed. But once your immune system is weakened by something like an accident or illness or other trauma, your immune system may be unable to keep the infection in check.
These bacteria can migrate out into surrounding tissues by hitching a ride into your blood stream, where they are transported to new locations to set up camp. The new location can be any organ or gland or tissue.
Dr. Price was able to transfer diseases harbored by humans to rabbits, by implanting fragments of root-canaled teeth, as mentioned above. He found that root canal fragments from a person who had suffered a heart attack, when implanted into a rabbit, would cause a heart attack in the rabbit within a few weeks.
He discovered he could transfer heart disease to the rabbit 100 percent of the time! Other diseases were more than 80 percent transferable by this method. Nearly every chronic degenerative disease has been linked with root canals, including:
Heart disease
Kidney disease
Arthritis, joint, and rheumatic diseases
Neurological diseases (including ALS and MS)
Autoimmune diseases (Lupus and more)
There may also be a cancer connection. Dr. Robert Jones, a researcher of the relationship between root canals and breast cancer, found an extremely high correlation between root canals and breast cancer. He claims to have found the following correlations in a five-year study of 300 breast cancer cases:
93 percent of women with breast cancer had root canals
7 percent had other oral pathology
Tumors, in the majority of cases, occurred on the same side of the body as the root canal(s) or other oral pathology.
Dr. Jones claims that toxins from the bacteria in an infected tooth or jawbone are able to inhibit the proteins that suppress tumor development. A German physician reported similar findings. Dr. Josef Issels reported that, in his 40 years of treating “terminal” cancer patients, 97 percent of his cancer patients had root canals. If these physicians are correct, the cure for cancer may be as simple as having a tooth pulled, then rebuilding your immune system.
Giovanni Angelo Canini
Giovanni Angelo Canini (Roma, 1609 – Roma, 1666) è stato un pittore e incisore italiano del periodo barocco. È anche noto come Giovanni Agnolo Canini o Giannangiolo.
Biografia
Figlio di uno scalpellino di nome Vincenzo, aveva due fratelli, tutti diventati artisti. Il fratello maggiore era un pittore, mentre quello minore, Marcantonio, era uno scultore. Da bambino fu allievo di Domenichino e quando quest'ultimo andò a Napoli nel 1631, Canini divenne allievo di Antonio Barbalonga. Nel 1634, su richiesta di Domenichino, fu incaricato di restaurare i dipinti di Passignano sulle pareti della cappella di San Sebastiano, nella Villa Aldobrandini di Frascati.
Dipinse alcuni pezzi d'altare a Roma, tra cui due immagini per la Basilica dei Santi Silvestro e Martino ai Monti. Dopo aver accompagnato il cardinale Chigi in Francia, fu incoraggiato dal ministro Colbert ad eseguire il suo progetto che consisteva nel disegnare una serie di ritratti di personaggi più illustri dell'antichità ripresi da medaglie, gemme antiche e altri oggetti simili. Ma poco dopo l'inizio dell'impresa Canini morì a Roma. Il lavoro, tuttavia, fu perseguito dal fratello Marcantonio, che, con l'aiuto di Picard e Valet, lo completò e pubblicò nel 1699, con il nome di Iconografia di Giovanni Agnolo Canini. L'opera contiene 150 incisioni. Una ristampa in lingua italiana e francese apparve ad Amsterdam nel 1731.
Da Wikipedia, l'enciclopedia libera.
Di Giovanni Angelo Canini;
Raccolta Foto de Alvariis;
Sagui
Callitrichinae (também chamada Hapalinae) é uma subfamília de Macacos do Novo Mundo, da família Cebidae. Popularmente, são conhecidos por saguis, soim ou sauim, apesar de que para o gênero Leontopithecus, é mais comum o termo mico-leão.
Sagui-de-tufos-pretos
Um sagui[1][2] (do tupi sauín), soim ou mico são as designações comuns dadas a várias espécies de pequenos macacos pertencentes à família Callitrichidae. A palavra sagui tem origem no tupi e sua pronúncia é feita observando-se o som da vogal "u".
Estes primatas são representados por várias espécies em território brasileiro. Todos os quais possuem o dedo polegar da mão muito curto e não oponível, as unhas em forma de garras, e dentes molares de fórmula 2/2. São espécies de pequeno porte e de cauda longa.
São os menores símios do mundo, estão dispersos por toda a América do Sul e vivem geralmente em bandos que se hospedam em árvores, como os esquilos. Travessos e ágeis, movem-se em saltos bruscos, emitindo guinchos e assobios que são ouvidos de longe.
The black-tufted marmoset (Callithrix penicillata), also known as Mico-estrela in Portuguese, is a species of New World monkey that lives primarily in the Neo-tropical gallery forests of the Brazilian Central Plateau. It ranges from Bahia to Paraná,[3] and as far inland as Goiás, between 14 and 25 degrees south of the equator. This marmoset typically resides in rainforests, living an arboreal life high in the trees, but below the canopy. They are only rarely spotted near the ground.
Physical description:
The black-tufted marmoset is characterized by black tufts of hair around their ears. It typically has some sparse white hairs on its face. It usually has a brown or black head and its limbs and upper body are gray, as well as its abdomen, while its rump and underside are usually black. Its tail is ringed with black and white and is not prehensile, but is used for balance. It does not have an opposable thumb and its nails tend to have a claw-like appearance. The black-tufted marmoset reaches a size of 19 to 22 cm and weighs up to 350 g.
Behavior:
Diurnal and arboreal, the black-tufted marmoset has a lifestyle very similar to other marmosets. It typically lives in family groups of 2 to 14. The groups usually consist of a reproductive couple and their offspring. Twins are very common among this species and the males, as well as juvenile offspring, often assist the female in the raising of the young.
Though the black-tufted marmoset lives in small family groups, it is believed that they share their food source, sap trees, with other marmoset groups. Scent marking does occur within these groups, but it is believed that the marking is to deter other species rather than other black-tufted marmoset groups, because other groups typically ignore these markings. They also appear to be migratory, often moving in relation to the wet or dry seasons, however, the extent of their migration is unknown.
Though communication between black-tufted marmosets has not been studied thoroughly, it is believed that it communicates through vocalizations. It has known predator-specific cries and appears to vocalize frequently outside of predator cries.
Food and predation:
The Black-tufted Marmoset diet consists primarily of tree sap which it gets by nibbling the bark with its long lower incisors. In periods of drought, it will also include fruit and insects in its diet. In periods of serious drought it has also been known to eat small arthropods, molluscs, bird eggs, baby birds and small vertebrates.
Large birds of prey are the greatest threat to the black-tufted marmoset, however, snakes and wild cats also pose a danger to them. Predator-specific vocalizations and visual scanning are its only anti-predation techniques.
Reproduction:
The black-tufted marmoset is monogamous and lives in family groups. It reproduces twice a year, producing 1 to 4 offspring, though most often just twins. Its gestation period is 150 days and offspring are weaned after 8 weeks. There is considerable parental investment by this species, with both parents, as well as older juveniles, helping to raise the young. The offspring are extremely dependent on their parents and though they are sexually mature at 18 months, they typically do not mate until much later, staying with their family group until they do.
Ecosystem roles and conservation status:
The black-tufted marmoset is a mutualist with many species of fruit trees because it distributes the seeds from the fruit it consumes throughout the forests. However, it is a parasite on other species of trees because it creates sores in trees in order to extract sap, while offering no apparent benefit to the trees. Though this marmoset is not a main food source to any specific species, it is a food source to a number of different species, specifically large birds of prey, wild cats, and snakes.
While there are no known negative effects of marmosets towards humans, it carries specific positive effects by being a highly valuable exotic pet. It is also used in zoo exhibits and scientific research.
The black-tufted marmoset is listed as having no special status on the IUCN Red List or the United States Endangered Species Act List. It is listed in Appendix II of CITES and is not currently considered an endangered or threatened species.
These are two of the four elephants that live at Colchester Zoo’s, who's charity, Action for the Wild, supports the Elephant Orphanage Project in Zambia, with its mission to rescue, rehabilitate and release orphaned elephants into protected reserves. The elephants they look after have become orphans due to the illegal ivory trade.
African elephants are the largest animals walking the earth. The word 'elephant' is Latin for 'huge arch'.
The average African elephant will stand between 8 ft. 2 in. to 13 ft. (2.5 to 4 m) at the shoulder and weigh between 2.25 to 6.25 tons (5,000 to 14,000 lb. - 2,270 to 6,350 kg), Male elephants can grow to be significantly larger than female which grow to between 9 and 13 ft. ( and weigh from 2.68 to 3.57 tons (6,000 to 8,000 lb. - 2,720 to 3,630 kg). The trunk alone can weigh as much as 310 lb. (140 kg) and can be a used as a deadly weapon. It can hold 2.65 gal. (12 litres) of water. There are about 50,000 muscles in the trunk, made up of 6 muscle groups and no bones.
Both male and female African elephants have tusks, which are elongated incisor teeth that first appear at two around years of age. The male's tusks are longer and heavier, weighing between 110 and 175 lb. (50 and 79 kg) each, with females' tusks weigh approximately 40 lb. (18 kg) each.
Elephants have large ears which are designed to help keep them cool by circulating air around the large blood vessels of the inner ears. The more the ears flap, the hotter the elephant is.
The elephant is an extremely intelligent creatures and they have brains that can weigh as much as 8.8 to 13.25 lb. (4 to 6 kg).
The skin of an elephant can be up to 1 in. (2.5 cm) thick in places. The folds and wrinkles in their skin can retain up to 10 times more water than flat skin does, which helps to cool them down. They keep their skin clean and protect themselves from sunburn by taking regular dust and mud baths.
Elephant have six sets of teeth that grow one set after another, throughout their lives. By the time they reach their 50's, most elephant have started to use their final set. After the elephant runs out of teeth, they cannot chew food and their condition deteriorates and they either fall prey to disease or predators or stave to death.
To feed an animal of such proportions takes a lot of food. Their average sleep time was found to be only 2 hours in 24 hour and most of their day is spent eating. Their daily food intake is as much as 4 to 7% of their body weight, which means they can eat as much as 330 Ib. (150 kg) of food per day. Elephants are herbivores and only eat grasses, herbs, fruit, plants and trees. They cause quite a lot of damage to their surroundings while feeding, especially when they use their tusks to tear off branches from trees. They also need 35 gal.(160 litres) of water a day.
Despite their size, they are actually pretty nimble and on average walk 15.5 miles (25 km) a day. They can also run faster than you would expect, easily reaching speeds of 40 mph (64 km/h).
African elephant live in matriarchal family units led by an older cow and made up of adult cows with their daughters and sub-adult sons. As soon as a young male becomes a bull he leaves the family group and may form an alliance with other bachelors. An elephant pregnancy lasts around 22 months. A baby elephant, called a calf, can weigh between 200 and 300 lbs (90 and 135 Kg) and stand about 3 ft. (1 m) tall. Calves are able to stand within 20 minutes of being born and can walk within 1 hour. After two days, they can keep up with the herd. This incredible survival technique means that herds of elephants can keep migrating to find food and water to thrive. Typically, elephants only give birth to one calf at a time, although twins do sometimes occur. In her lifetime a female elephant can give birth to as many as 12 calves. African elephants practice what is known as allomothering, whereby the calves are cared for not only by their mothers but also by other young cows in the family. A calf can expect to have an average life span of around 70 years.
Once elephants roamed across the whole of Africa, they are now limited to conservation areas and the Savannah. African elephants can currently be found in 37 African countries. In the 1970's there were 1.3 million African elephants. Less than a third of that number, an estimated 400,000, are left in the wild now. On average 96 elephants are poached a day for their valuable tusks. Moreover, humans are responsible for huge areas of habit loss of African elephants. As the human population in Africa increases, more and more land is converted to agriculture. A smaller habit has meant that elephants and humans are coming into contact with one another more than they did historically, and this is causing conflict. Elephants are very intelligent creature and sometimes raid farmers’ crop for food. This can lead to both humans and elephants being killed in the conflict.
African bush elephant
Afrikanischer Elefant
Kruger National Park is one of the largest game reserves in Africa. It covers an area of 19,485 km2 (7,523 sq mi) in the provinces of Limpopo and Mpumalanga in northeastern South Africa, and extends 360 km (220 mi) from north to south and 65 km (40 mi) from east to west. The administrative headquarters are in Skukuza. Areas of the park were first protected by the government of the South African Republic in 1898, and it became South Africa's first national park in 1926.
To the west and south of the Kruger National Park are the two South African provinces of Limpopo and Mpumalanga. In the north is Zimbabwe, and to the east is Mozambique. It is now part of the Great Limpopo Transfrontier Park, a peace park that links Kruger National Park with the Gonarezhou National Park in Zimbabwe, and with the Limpopo National Park in Mozambique.
The park is part of the Kruger to Canyons Biosphere an area designated by the United Nations Educational, Scientific and Cultural Organization (UNESCO) as an International Man and Biosphere Reserve (the "Biosphere").
The park has nine main gates allowing entrance to the different camps.
(Wikipedia)
Name
African Elephant or African Bush Elephant [Loxodonta africana]
Introduction
The Elephant is the world's largest land mammal, and weighs up to 7 tonnes and reaches heights of 3.3 m at the shoulder. Elephants can live to a potential age of 70 years. The massive tusks of older bulls can weigh up to 50 or 60 kilograms, but tusks weighing up to 90 kilograms have been recorded.
Appearance
What is the trunk and what is it used for?
The Elephant's trunk is a modified nose which is very sensitive and can even detect water under ground. There are as many as 50 000 muscles in an Elephant trunk. The sensitive finger-like appendages at the tip of the trunk enables them to pick the smallest twig or flower, pull the toughest reed of grass or even pick out a thorn from their feet.
Do elephants have knees or elbows?
The joints that are perceived as 'knees', are in fact wrists. This is a common misunderstanding due to the belief that a leg joint that bends between the foot and the body must be a knee. The main difference between us and the elephants is that our foot bones and hand bones are separate, whereas those of the elephant are one in the same, and have evolved to suit this four-legged mammal.
Why do elephants have tusks?
The tusks are used for obtaining food, fighting (amongst males) and for self defence. They are actually their upper incisors, and grow continuously until they die at around 60 years old. Although their skin is up to 3cm (1 inch) thick, it is quite sensitive.
Diet
Elephants are voracious feeders which in a day consume up to 272 kg (600 pounds) of grass, tender shoots and bark from trees. An adult Elephant can drink up to 200 litres of water in a single session. A single Elephant deposits up to 150kg (330 pounds) of dung every day - about one dollop every 15 minutes!
Breeding
African Elephant are not seasonal breeders. Generally they produce one calf every 3 to 4 years. The gestation period is about 22 months. At birth calves weigh about 100 kg (220 pounds) and are fully weaned between 18-24 months. An orphaned calf will usually be adopted by one of the family's lactating females or suckled by various females. Elephants are very attentive mothers, and because most Elephant behavior has to be learned, they keep their offspring with them for many years. Tusks erupt at 16 months but do not show externally until 30 months. Once weaned, usually at age 4 or 5, the calf still remains in the maternal group. Females mature at about 11 years and stay in the group, while the males, which mature between 12 and 15, are usually expelled from the maternal herd. Even though these young males are sexually mature, they do not breed until they are in their mid, or late 20s or even older and have moved up in the social hierarchy.
Behaviour
Mature males form bachelor groups and become solitary bulls. Elephant form strong family units of cows, calves and young offspring. Such herds are always led by an old female. Apart from drinking large quantities of water they also love wading or swimming in it. Elephants clearly relish mud baths.
It was once thought that family groups were led by old bull elephants, but these males are most often solitary. The female family groups are often visited by mature males checking for females in oestrus. Several interrelated family groups may inhabit an area and know each other well.
How do you tell an elephant's mock charge from a serious one?
It is imperative to keep in mind that Elephant are extremely intelligent, and each individual has a distinct character. Although there will be exceptions to the rules, the common signs of a mock charge are bush-bashing, dust-throwing, trumpeting and other vocalizations, open ears and an intimidating presence, can be considered a mock-display. Aggressive or startled elephants usually make sudden headshakes and flap their large ears against their head. Serious charges usually occur after all attempts to intimidate have failed, and the Elephant feels threatened. The ears are pinned back and head and trunk are lowered. Ultimately, the key lies in the intelligence of the animal and how they will react to the 'target' and unfamiliar actions, and a conscious decision is made.
Why do elephants rhythmically flap their ears?
Contrary to common belief, it is not an expression of anger. Being an animal of such a large size, with no sweat glands and a dark body colour, elephants flap their ears to cool the body and rid themselves of irritating insects.
Where are they found?
Once ranging across most of Africa the Elephant population has declined dramatically across the continent. In South Africa the Addo Elephant and Kruger National Park protect large herds. Due to rigorous conservation measures the Elephant population in South Africa has grown from a estimated 120 in 1920 in 4 locations, to 10 000 at 40 locations to date.
Notes
The African Elephant has recently been classified into two separate species, the more common African Bush Elephant [Loxodonta Africana] and the smaller African Forest Elephant [Loxodonta cyclotis] of the rainforest of Central Africa.
(krugerpark.co.za)
Der Kruger-Nationalpark (deutsch häufig falsch Krüger-Nationalpark) ist das größte Wildschutzgebiet Südafrikas. Er liegt im Nordosten des Landes in der Landschaft des Lowveld auf dem Gebiet der Provinz Limpopo sowie des östlichen Abschnitts von Mpumalanga. Seine Fläche erstreckt sich vom Crocodile-River im Süden bis zum Limpopo, dem Grenzfluss zu Simbabwe, im Norden. Die Nord-Süd-Ausdehnung beträgt etwa 350 km, in Ost-West-Richtung ist der Park durchschnittlich 54 km breit und umfasst eine Fläche von rund 20.000 Quadratkilometern. Damit gehört er zu den größten Nationalparks in Afrika.
Das Schutzgebiet wurde am 26. März 1898 unter dem Präsidenten Paul Kruger als Sabie Game Reserve zum Schutz der Wildnis gegründet. 1926 erhielt das Gebiet den Status Nationalpark und wurde in seinen heutigen Namen umbenannt. Im Park leben 147 Säugetierarten inklusive der „Big Five“, außerdem etwa 507 Vogelarten und 114 Reptilienarten, 49 Fischarten und 34 Amphibienarten.
(Wikipedia)
Der Afrikanische Elefant (Loxodonta africana), auch Afrikanischer Steppenelefant oder Afrikanischer Buschelefant, ist eine Art aus der Familie der Elefanten. Er ist das größte gegenwärtig lebende Landsäugetier und gleichzeitig das größte rezente landbewohnende Tier der Erde. Herausragende Kennzeichen sind neben den Stoßzähnen und dem markanten Rüssel die großen Ohren und die säulenförmigen Beine. In zahlreichen morphologischen und anatomischen Merkmalen unterscheidet sich der Afrikanische Elefant von seinen etwas kleineren Verwandten, dem Waldelefanten und dem Asiatischen Elefanten. Das Verbreitungsgebiet umfasst heute große Teile von Afrika südlich der Sahara. Die Tiere haben sich dort an zahlreiche unterschiedliche Lebensräume angepasst, die von geschlossenen Wäldern über offene Savannenlandschaften bis hin zu Sumpfgebieten und wüstenartigen Regionen reichen. Insgesamt ist das Vorkommen aber stark fragmentiert.
Die Lebensweise des Afrikanischen Elefanten ist durch intensive Studien gut erforscht. Sie wird durch einen stark sozialen Charakter geprägt. Weibliche Tiere und ihr Nachwuchs leben in Familienverbänden (Herden). Diese formieren sich wiederum zu einem enger verwandten Clan. Die einzelnen Herden treffen sich zu bestimmten Gelegenheiten und trennen sich danach wieder. Die männlichen Tiere bilden Junggesellengruppen. Die verschiedenen Verbände nutzen Aktionsräume, in denen sie teils im Jahreszyklus herumwandern. Für die Kommunikation untereinander nutzen die Tiere verschiedene Töne im niedrigen Frequenzbereich. Anhand der Lautgebung, aber auch durch bestimmte chemische Signale können sich die einzelnen Individuen untereinander erkennen. Darüber hinaus besteht ein umfangreiches Repertoire an Gesten. Hervorzuheben sind auch die kognitiven Fähigkeiten des Afrikanischen Elefanten.
Die Nahrung besteht sowohl aus weicher wie auch harter Pflanzenkost. Die genaue Zusammensetzung variiert dabei regional und jahreszeitlich. Generell verbringt der Afrikanische Elefant einen großen Teil seiner Tagesaktivitäten mit der Nahrungsaufnahme. Die Fortpflanzung erfolgt ganzjährig, regional gibt es Tendenzen zu einer stärkeren Saisonalisierung. Bullen kommen einmal jährlich in die Musth, während deren sie auf Wanderung zur Suche nach fortpflanzungswilligen Kühen gehen. Während der Musth ist die Aggressivität gesteigert, es finden dann auch Rivalenkämpfe statt. Der Sexualzyklus der Kühe dauert vergleichsweise lange und weist einen für Säugetiere untypischen Verlauf auf. Nach erfolgter Geburt setzt er in der Regel mehrere Jahre aus. Zumeist wird nach fast zweijähriger Tragzeit ein Jungtier geboren, das in der mütterlichen Herde aufwächst. Junge weibliche Tiere verbleiben später in der Herde, die jungen männlichen verlassen diese.
Die wissenschaftliche Erstbeschreibung des Afrikanischen Elefanten erfolgte im Jahr 1797 mit einer formalen artlichen Trennung des Afrikanischen vom Asiatischen Elefanten. Der heute gebräuchliche Gattungsname Loxodonta wurde offiziell erst dreißig Jahre später eingeführt. Die Bezeichnung bezieht sich auf markante Zahnunterschiede zwischen den asiatischen und den afrikanischen Elefanten. Im Verlauf des 20. Jahrhunderts wurden mehrere Unterarten unterschieden, darunter auch der Waldelefant des zentralen Afrikas. Letzterer gilt heute genetischen Untersuchungen zufolge als eigenständige Art, die weiteren Unterarten sind nicht anerkannt. Stammesgeschichtlich lässt sich der Afrikanische Elefant erstmals im beginnenden Mittleren Pleistozän belegen. Der Gesamtbestand gilt als gefährdet. Ursachen hierfür sind hauptsächlich die Jagd nach Elfenbein und Lebensraumverlust durch die zunehmend wachsende menschliche Bevölkerung. Der Afrikanische Elefant zählt zu den sogenannten „Big Five“ von Großwildjagd und Safari.
(Wikipedia)
African bush elephant
Afrikanischer Elefant
Kruger National Park is one of the largest game reserves in Africa. It covers an area of 19,485 km2 (7,523 sq mi) in the provinces of Limpopo and Mpumalanga in northeastern South Africa, and extends 360 km (220 mi) from north to south and 65 km (40 mi) from east to west. The administrative headquarters are in Skukuza. Areas of the park were first protected by the government of the South African Republic in 1898, and it became South Africa's first national park in 1926.
To the west and south of the Kruger National Park are the two South African provinces of Limpopo and Mpumalanga. In the north is Zimbabwe, and to the east is Mozambique. It is now part of the Great Limpopo Transfrontier Park, a peace park that links Kruger National Park with the Gonarezhou National Park in Zimbabwe, and with the Limpopo National Park in Mozambique.
The park is part of the Kruger to Canyons Biosphere an area designated by the United Nations Educational, Scientific and Cultural Organization (UNESCO) as an International Man and Biosphere Reserve (the "Biosphere").
The park has nine main gates allowing entrance to the different camps.
(Wikipedia)
Name
African Elephant or African Bush Elephant [Loxodonta africana]
Introduction
The Elephant is the world's largest land mammal, and weighs up to 7 tonnes and reaches heights of 3.3 m at the shoulder. Elephants can live to a potential age of 70 years. The massive tusks of older bulls can weigh up to 50 or 60 kilograms, but tusks weighing up to 90 kilograms have been recorded.
Appearance
What is the trunk and what is it used for?
The Elephant's trunk is a modified nose which is very sensitive and can even detect water under ground. There are as many as 50 000 muscles in an Elephant trunk. The sensitive finger-like appendages at the tip of the trunk enables them to pick the smallest twig or flower, pull the toughest reed of grass or even pick out a thorn from their feet.
Do elephants have knees or elbows?
The joints that are perceived as 'knees', are in fact wrists. This is a common misunderstanding due to the belief that a leg joint that bends between the foot and the body must be a knee. The main difference between us and the elephants is that our foot bones and hand bones are separate, whereas those of the elephant are one in the same, and have evolved to suit this four-legged mammal.
Why do elephants have tusks?
The tusks are used for obtaining food, fighting (amongst males) and for self defence. They are actually their upper incisors, and grow continuously until they die at around 60 years old. Although their skin is up to 3cm (1 inch) thick, it is quite sensitive.
Diet
Elephants are voracious feeders which in a day consume up to 272 kg (600 pounds) of grass, tender shoots and bark from trees. An adult Elephant can drink up to 200 litres of water in a single session. A single Elephant deposits up to 150kg (330 pounds) of dung every day - about one dollop every 15 minutes!
Breeding
African Elephant are not seasonal breeders. Generally they produce one calf every 3 to 4 years. The gestation period is about 22 months. At birth calves weigh about 100 kg (220 pounds) and are fully weaned between 18-24 months. An orphaned calf will usually be adopted by one of the family's lactating females or suckled by various females. Elephants are very attentive mothers, and because most Elephant behavior has to be learned, they keep their offspring with them for many years. Tusks erupt at 16 months but do not show externally until 30 months. Once weaned, usually at age 4 or 5, the calf still remains in the maternal group. Females mature at about 11 years and stay in the group, while the males, which mature between 12 and 15, are usually expelled from the maternal herd. Even though these young males are sexually mature, they do not breed until they are in their mid, or late 20s or even older and have moved up in the social hierarchy.
Behaviour
Mature males form bachelor groups and become solitary bulls. Elephant form strong family units of cows, calves and young offspring. Such herds are always led by an old female. Apart from drinking large quantities of water they also love wading or swimming in it. Elephants clearly relish mud baths.
It was once thought that family groups were led by old bull elephants, but these males are most often solitary. The female family groups are often visited by mature males checking for females in oestrus. Several interrelated family groups may inhabit an area and know each other well.
How do you tell an elephant's mock charge from a serious one?
It is imperative to keep in mind that Elephant are extremely intelligent, and each individual has a distinct character. Although there will be exceptions to the rules, the common signs of a mock charge are bush-bashing, dust-throwing, trumpeting and other vocalizations, open ears and an intimidating presence, can be considered a mock-display. Aggressive or startled elephants usually make sudden headshakes and flap their large ears against their head. Serious charges usually occur after all attempts to intimidate have failed, and the Elephant feels threatened. The ears are pinned back and head and trunk are lowered. Ultimately, the key lies in the intelligence of the animal and how they will react to the 'target' and unfamiliar actions, and a conscious decision is made.
Why do elephants rhythmically flap their ears?
Contrary to common belief, it is not an expression of anger. Being an animal of such a large size, with no sweat glands and a dark body colour, elephants flap their ears to cool the body and rid themselves of irritating insects.
Where are they found?
Once ranging across most of Africa the Elephant population has declined dramatically across the continent. In South Africa the Addo Elephant and Kruger National Park protect large herds. Due to rigorous conservation measures the Elephant population in South Africa has grown from a estimated 120 in 1920 in 4 locations, to 10 000 at 40 locations to date.
Notes
The African Elephant has recently been classified into two separate species, the more common African Bush Elephant [Loxodonta Africana] and the smaller African Forest Elephant [Loxodonta cyclotis] of the rainforest of Central Africa.
(krugerpark.co.za)
Der Kruger-Nationalpark (deutsch häufig falsch Krüger-Nationalpark) ist das größte Wildschutzgebiet Südafrikas. Er liegt im Nordosten des Landes in der Landschaft des Lowveld auf dem Gebiet der Provinz Limpopo sowie des östlichen Abschnitts von Mpumalanga. Seine Fläche erstreckt sich vom Crocodile-River im Süden bis zum Limpopo, dem Grenzfluss zu Simbabwe, im Norden. Die Nord-Süd-Ausdehnung beträgt etwa 350 km, in Ost-West-Richtung ist der Park durchschnittlich 54 km breit und umfasst eine Fläche von rund 20.000 Quadratkilometern. Damit gehört er zu den größten Nationalparks in Afrika.
Das Schutzgebiet wurde am 26. März 1898 unter dem Präsidenten Paul Kruger als Sabie Game Reserve zum Schutz der Wildnis gegründet. 1926 erhielt das Gebiet den Status Nationalpark und wurde in seinen heutigen Namen umbenannt. Im Park leben 147 Säugetierarten inklusive der „Big Five“, außerdem etwa 507 Vogelarten und 114 Reptilienarten, 49 Fischarten und 34 Amphibienarten.
(Wikipedia)
Der Afrikanische Elefant (Loxodonta africana), auch Afrikanischer Steppenelefant oder Afrikanischer Buschelefant, ist eine Art aus der Familie der Elefanten. Er ist das größte gegenwärtig lebende Landsäugetier und gleichzeitig das größte rezente landbewohnende Tier der Erde. Herausragende Kennzeichen sind neben den Stoßzähnen und dem markanten Rüssel die großen Ohren und die säulenförmigen Beine. In zahlreichen morphologischen und anatomischen Merkmalen unterscheidet sich der Afrikanische Elefant von seinen etwas kleineren Verwandten, dem Waldelefanten und dem Asiatischen Elefanten. Das Verbreitungsgebiet umfasst heute große Teile von Afrika südlich der Sahara. Die Tiere haben sich dort an zahlreiche unterschiedliche Lebensräume angepasst, die von geschlossenen Wäldern über offene Savannenlandschaften bis hin zu Sumpfgebieten und wüstenartigen Regionen reichen. Insgesamt ist das Vorkommen aber stark fragmentiert.
Die Lebensweise des Afrikanischen Elefanten ist durch intensive Studien gut erforscht. Sie wird durch einen stark sozialen Charakter geprägt. Weibliche Tiere und ihr Nachwuchs leben in Familienverbänden (Herden). Diese formieren sich wiederum zu einem enger verwandten Clan. Die einzelnen Herden treffen sich zu bestimmten Gelegenheiten und trennen sich danach wieder. Die männlichen Tiere bilden Junggesellengruppen. Die verschiedenen Verbände nutzen Aktionsräume, in denen sie teils im Jahreszyklus herumwandern. Für die Kommunikation untereinander nutzen die Tiere verschiedene Töne im niedrigen Frequenzbereich. Anhand der Lautgebung, aber auch durch bestimmte chemische Signale können sich die einzelnen Individuen untereinander erkennen. Darüber hinaus besteht ein umfangreiches Repertoire an Gesten. Hervorzuheben sind auch die kognitiven Fähigkeiten des Afrikanischen Elefanten.
Die Nahrung besteht sowohl aus weicher wie auch harter Pflanzenkost. Die genaue Zusammensetzung variiert dabei regional und jahreszeitlich. Generell verbringt der Afrikanische Elefant einen großen Teil seiner Tagesaktivitäten mit der Nahrungsaufnahme. Die Fortpflanzung erfolgt ganzjährig, regional gibt es Tendenzen zu einer stärkeren Saisonalisierung. Bullen kommen einmal jährlich in die Musth, während deren sie auf Wanderung zur Suche nach fortpflanzungswilligen Kühen gehen. Während der Musth ist die Aggressivität gesteigert, es finden dann auch Rivalenkämpfe statt. Der Sexualzyklus der Kühe dauert vergleichsweise lange und weist einen für Säugetiere untypischen Verlauf auf. Nach erfolgter Geburt setzt er in der Regel mehrere Jahre aus. Zumeist wird nach fast zweijähriger Tragzeit ein Jungtier geboren, das in der mütterlichen Herde aufwächst. Junge weibliche Tiere verbleiben später in der Herde, die jungen männlichen verlassen diese.
Die wissenschaftliche Erstbeschreibung des Afrikanischen Elefanten erfolgte im Jahr 1797 mit einer formalen artlichen Trennung des Afrikanischen vom Asiatischen Elefanten. Der heute gebräuchliche Gattungsname Loxodonta wurde offiziell erst dreißig Jahre später eingeführt. Die Bezeichnung bezieht sich auf markante Zahnunterschiede zwischen den asiatischen und den afrikanischen Elefanten. Im Verlauf des 20. Jahrhunderts wurden mehrere Unterarten unterschieden, darunter auch der Waldelefant des zentralen Afrikas. Letzterer gilt heute genetischen Untersuchungen zufolge als eigenständige Art, die weiteren Unterarten sind nicht anerkannt. Stammesgeschichtlich lässt sich der Afrikanische Elefant erstmals im beginnenden Mittleren Pleistozän belegen. Der Gesamtbestand gilt als gefährdet. Ursachen hierfür sind hauptsächlich die Jagd nach Elfenbein und Lebensraumverlust durch die zunehmend wachsende menschliche Bevölkerung. Der Afrikanische Elefant zählt zu den sogenannten „Big Five“ von Großwildjagd und Safari.
(Wikipedia)
The polar bear (Ursus maritimus) is a large bear native to the Arctic and nearby areas. It is closely related to the brown bear, and the two species can interbreed. The polar bear is the largest extant species of bear and land carnivore, with adult males weighing 300–800 kg (660–1,760 lb). The species is sexually dimorphic, as adult females are much smaller. The polar bear is white- or yellowish-furred with black skin and a thick layer of fat. It is more slender than the brown bear, with a narrower skull, longer neck and lower shoulder hump. Its teeth are sharper and more adapted to cutting meat. The paws are large and allow the bear to walk on ice and paddle in the water.
Polar bears are both terrestrial and pagophilic (ice-living) and are considered to be marine mammals due to their dependence on marine ecosystems. They prefer the annual sea ice but live on land when the ice melts in the summer. They are mostly carnivorous and specialized for preying on seals, particularly ringed seals. Such prey is typically taken by ambush; the bear may stalk its prey on the ice or in the water, but also will stay at a breathing hole or ice edge to wait for prey to swim by. The bear primarily feeds on the seal's energy-rich blubber. Other prey include walruses, beluga whales and some terrestrial animals. Polar bears are usually solitary but can be found in groups when on land. During the breeding season, male bears guard females and defend them from rivals. Mothers give birth to cubs in maternity dens during the winter. Young stay with their mother for up to two and a half years.
The polar bear is considered to be a vulnerable species by the International Union for Conservation of Nature (IUCN) with an estimated total population of 22,000 to 31,000 individuals. Its biggest threats are climate change, pollution and energy development. Climate change has caused a decline in sea ice, giving the polar bear less access to its favoured prey and increasing the risk of malnutrition and starvation. Less sea ice also means that the bears must spend more time on land, increasing conflicts with people. Polar bears have been hunted, both by native and non-native peoples, for their coats, meat and other items. They have been kept in captivity in zoos and circuses and are prevalent in art, folklore, religion and modern culture.
Naming
The polar bear was given its common name by Thomas Pennant in A Synopsis of Quadrupeds (1771). It was known as the "white bear" in Europe between the 13th and 18th centuries, as well as "ice bear", "sea bear" and "Greenland bear". The Norse referred to it as isbjørn ("ice bear") and hvitebjørn ("white bear"). The bear is called nanook by the Inuit. The Netsilik cultures additionally have different names for bears based on certain factors, such as sex and age: these include adult males (anguraq), single adult females (tattaq), gestating females (arnaluk), newborns (hagliaqtug), large adolescents (namiaq) and dormant bears (apitiliit). The scientific name Ursus maritimus is Latin for "sea bear".
Taxonomy
Carl Linnaeus classified the polar bear as a type of brown bear (Ursus arctos), labelling it as Ursus maritimus albus-major, articus in the 1758 edition of his work Systema Naturae. Constantine John Phipps formally described the polar bear as a distinct species, Ursus maritimus in 1774, following his 1773 voyage towards the North Pole. Due to its adaptations to a marine environment, some taxonomists like Theodore Knottnerus-Meyer have placed the polar bear in its genus Thalarctos. However Ursus is widely considered to be the valid genus for the species based on the fossil record and the fact that it can breed with the brown bear.
Different subspecies have been proposed including Ursus maritimus maritimus and U. m. marinus. However these are not supported and the polar bear is considered to be monotypic. One possible fossil subspecies, U. m. tyrannus, was posited in 1964 by Björn Kurtén, who reconstructed the subspecies from a single fragment of an ulna which was approximately 20 percent larger than expected for a polar bear. However, re-evaluation in the 21st century has indicated that the fragment likely comes from a giant brown bear.
Evolution
The polar bear is one of eight extant species in the bear family Ursidae and of six extant species in the subfamily Ursinae. A possible phylogeny of extant bear species is shown in a cladogram based on complete mitochondrial DNA sequences from Yu et al. (2007). The polar bear and the brown bear form a close grouping, while the relationships of the other species are not very well resolved.
Giant panda (Ailuropoda melanoleuca)
Spectacled bear (Tremarctos ornatus)
Ursinae
Sloth bear (Melursus ursinus)
Sun bear (Helarctos malayanus)
Asian black bear (Ursus thibetanus)
American black bear (Ursus americanus)
Polar bear (Ursus maritimus)
Brown bear (Ursus arctos)
A more recent phylogeny below is based on a 2017 genetic study. The study concludes that Ursine bears originated around 5 million years ago and show extensive hybridization of species in their lineage.
Ursidae
Giant panda (Ailuropoda melanoleuca)
Spectacled bear (Tremarctos ornatus)
Ursinae
Sloth bear (Melursus ursinus)
Sun bear (Helarctos malayanus)
Asian black bear (Ursus thibetanus)
American black bear (Ursus americanus)
Polar bear (Ursus maritimus)
Brown bear (Ursus arctos)
Fossils of polar bears are uncommon. The oldest known fossil is a 130,000- to 110,000-year-old jaw bone, found on Prince Charles Foreland, Norway, in 2004. Scientists in the 20th century surmised that polar bears directly descended from a population of brown bears, possibly in eastern Siberia or Alaska. Mitochondrial DNA studies in the 1990s and 2000s supported the status of the polar bear as a derivative of the brown bear, finding that some brown bear populations were more closely related to polar bears than to other brown bears, particularly the ABC Islands bears of Southeast Alaska. A 2010 study estimated that the polar bear lineage split from other brown bears around 150,000 years ago.
More extensive genetic studies have refuted the idea that polar bears are directly descended from brown bears and found that the two species are separate sister lineages. The genetic similarities between polar bears and some brown bears were found to be the result of interbreeding A 2012 study estimated the split between polar and brown bears as occurring around 600,000 years ago. A 2022 study estimated the divergence as occurring even earlier at over one million years ago. Glaciation events over hundreds of thousands of years led to both the origin of polar bears and their subsequent interactions and hybridizations with brown bears.
Studies in 2011 and 2012 concluded that gene flow went from brown bears to polar bears during hybridization. In particular, a 2011 study concluded that living polar bear populations derived their maternal lines from now-extinct Irish brown bears. Later studies have clarified that gene flow went from polar to brown bears rather than the reverse. Up to 9 percent of the genome of ABC bears was transferred from polar bears, while Irish bears had up to 21.5 percent polar bear origin. Mass hybridization between the two species appears to have stopped around 200,000 years ago. Modern hybrids are relatively rare in the wild.
Analysis of the number of variations of gene copies in polar bears compared with brown bears and American black bears shows distinct adaptions. Polar bears have a less diverse array of olfactory receptor genes, a result of there being fewer odours in their Arctic habitat. With its carnivorous, high-fat diet the species has fewer copies of the gene involved in making amylase, an enzyme that breaks down starch, and more selection for genes for fatty acid breakdown and a more efficient circulatory system. The polar bear's thicker coat is the result of more copies of genes involved in keratin-creating proteins.
Characteristics
The polar bear is the largest living species of bear and land carnivore, though some brown bear subspecies like the Kodiak bear can rival it in size. Males are generally 200–250 cm (6.6–8.2 ft) long with a weight of 300–800 kg (660–1,760 lb). Females are smaller at 180–200 cm (5.9–6.6 ft) with a weight of 150–300 kg (330–660 lb). Sexual dimorphism in the species is particularly high compared with most other mammals. Male polar bears also have proportionally larger heads than females. The weight of polar bears fluctuates during the year, as they can bulk up on fat and increase their mass by 50 percent. A fattened, pregnant female can weigh as much as 500 kg (1,100 lb). Adults may stand 130–160 cm (4.3–5.2 ft) tall at the shoulder. The tail is 76–126 mm (3.0–5.0 in) long. The largest polar bear on record, reportedly weighing 1,002 kg (2,209 lb), was a male shot at Kotzebue Sound in northwestern Alaska in 1960.
Compared with the brown bear, this species has a more slender build, with a narrower, flatter and smaller skull, a longer neck, and a lower shoulder hump. The snout profile is curved, resembling a "Roman nose". They have 34–42 teeth including 12 incisors, 4 canines, 8–16 premolars and 10 molars. The teeth are adapted for a more carnivorous diet than that of the brown bear, having longer, sharper and more spaced out canines, and smaller, more pointed cheek teeth (premolars and molars). The species has a large space or diastema between the canines and cheek teeth, which may allow it to better bite into prey. Since it normally preys on animals much smaller than it, the polar bear does not have a particularly strong bite Polar bears have large paws, with the front paws being broader than the back. The feet are hairier than in other bear species, providing warmth and friction when stepping on snow and sea ice. The claws are small but sharp and hooked and are used both to snatch prey and climb onto ice.
Polar bear jumping on floating ice at Svalbard
The coat consists of dense underfur around 5 cm (2.0 in) long and guard hairs around 15 cm (5.9 in) long. Males have long hairs on their forelegs, which is thought to signal their fitness to females. The outer surface of the hairs has a scaly appearance, and the guard hairs are hollow, which allows the animals to trap heat and float in the water. The transparent guard hairs forward scatter ultraviolet light between the underfur and the skin, leading to a cycle of absorption and re-emission, keeping them warm. The fur appears white due to the backscatter of incident light and the absence of pigment. Polar bears gain a yellowish colouration as they are exposed more to the sun. This is reversed after they moult. It can also be grayish or brownish. Their light fur provides camouflage in their snowy environment. After emerging from the water, the bear can easily shake itself dry before freezing since the hairs are resistant to tangling when wet. The skin, including the nose and lips, is black and absorbs heat. Polar bears have a 5–10 cm (2.0–3.9 in) thick layer of fat underneath the skin, which provides both warmth and energy. Polar bears maintain their core body temperature at about 36.9 °C (98 °F). Overheating is countered by a layer of highly vascularized striated muscle tissue and finely controlled blood vessels. Bears also cool off by entering the water.
The eyes of a polar bear are close to the top of the head, which may allow them to stay out of the water when the animal is swimming at the surface. They are relatively small, which may be an adaption against blowing snow and snow blindness. Polar bears are dichromats, and lack the cone cells for seeing green. They have many rod cells which allow them to see at night. The ears are small, allowing them to retain heat and not get frostbitten. They can hear best at frequencies of 11.2–22.5 kHz, a wider frequency range than expected given that their prey mostly makes low-frequency sounds. The nasal concha creates a large surface area, so more warm air can move through the nasal passages. Their olfactory system is also large and adapted for smelling prey over vast distances. The animal has reniculate kidneys which filter out the salt in their food.
Distribution and habitat
Polar bears inhabit the Arctic and adjacent areas. Their range includes Greenland, Canada, Alaska, Russia and the Svalbard Archipelago of Norway. Polar bears have been recorded 25 km (16 mi) from the North Pole. The southern limits of their range include James Bay and Newfoundland and Labrador in Canada and St. Matthew Island and the Pribilof Islands of Alaska. They are not permanent residents of Iceland but have been recorded visiting there if they can reach it via sea ice. Due to minimal human encroachment on the bears' remote habitat, they can still be found in much of their original range, more so than any other large land carnivore.
Polar bears have been divided into at least 18 subpopulations labelled East Greenland (ES), Barents Sea (BS), Kara Sea (KS), Laptev Sea (LVS), Chukchi Sea (CS), northern and southern Beaufort Sea (SBS and NBS), Viscount Melville (VM), M'Clintock Channel (MC), Gulf of Boothia (GB), Lancaster Sound (LS), Norwegian Bay (NB), Kane Basin (KB), Baffin Bay (BB), Davis Strait (DS), Foxe Basin (FB) and the western and southern Hudson Bay (WHB and SHB) populations. Bears in and around the Queen Elizabeth Islands have been proposed as a subpopulation but this is not universally accepted. A 2022 study has suggested that the bears in southeast Greenland should be considered a different subpopulation based on their geographic isolation and genetics. Polar bear populations can also be divided into four gene clusters: Southern Canadian, Canadian Archipelago, Western Basin (northwestern Canada west to the Russian Far East) and Eastern Basin (Greenland east to Siberia).
The polar bear is dependent enough on the ocean to be considered a marine mammal. It is pagophilic and mainly inhabits annual sea ice covering continental shelves and between islands of archipelagos. These areas, known as the "Arctic Ring of Life", have high biological productivity. The species tends to frequent areas where sea ice meets water, such as polynyas and leads, to hunt the seals that make up most of its diet. Polar bears travel in response to changes in ice cover throughout the year. They are forced onto land in summer when the sea ice disappears. Terrestrial habitats used by polar bears include forests, mountains, rocky areas, lakeshores and creeks. In the Chukchi and Beaufort seas, where the sea ice breaks off and floats north during the summer, polar bears generally stay on the ice, though a large portion of the population (15–40%) has been observed spending all summer on land since the 1980s. Some areas have thick multiyear ice that does not completely melt and the bears can stay on all year, though this type of ice has fewer seals and allows for less productivity in the water.
Behaviour and ecology
Polar bears may travel areas as small as 3,500 km2 (1,400 sq mi) to as large as 38,000 km2 (15,000 sq mi) in a year, while drifting ice allows them to move further. Depending on ice conditions, a bear can travel an average of 12 km (7.5 mi) per day. These movements are powered by their energy-rich diet. Polar bears move by walking and galloping and do not trot. Walking bears tilt their front paws towards each other. They can run at estimated speeds of up to 40 km/h (25 mph) but typically move at around 5.5 km/h (3.4 mph). Polar bears are also capable swimmers and can swim at up to 6 km/h (3.7 mph). One study found they can swim for an average of 3.4 days at a time and travel an average of 154.2 km (95.8 mi). They can dive for as long as three minutes. When swimming, the broad front paws do the paddling, while the hind legs play a role in steering and diving.
Mother bear and cubs sleeping
Most polar bears are active year-round. Hibernation occurs only among pregnant females. Non-hibernating bears typically have a normal 24-hour cycle even during days of all darkness or all sunlight, though cycles less than a day are more common during the former. The species is generally diurnal, being most active early in the day. Polar bears sleep close to eight hours a day on average. They will sleep in various positions, including curled up, sitting up, lying on one side, on the back with limbs spread, or on the belly with the rump elevated. On sea ice, polar bears snooze at pressure ridges where they dig on the sheltered side and lie down. After a snowstorm, a bear may rest under the snow for hours or days. On land, the bears may dig a resting spot on gravel or sand beaches. They will also sleep on rocky outcrops. In mountainous areas on the coast, mothers and subadults will sleep on slopes where they can better spot another bear coming. Adult males are less at risk from other bears and can sleep nearly anywhere.
Social life
Polar bears are typically solitary, aside from mothers with cubs and mating pairs. On land, they are found closer together and gather around food resources. Adult males, in particular, are more tolerant of each other in land environments and outside the breeding season. They have been recorded forming stable "alliances", travelling, resting and playing together. A dominant hierarchy exists among polar bears with the largest mature males ranking at the top. Adult females outrank subadults and adolescents and younger males outrank females of the same age. In addition, cubs with their mothers outrank those on their own. Females with dependent offspring tend to stay away from males, but are sometimes associated with other female–offspring units, creating "composite families".
Polar bears are generally quiet but can produce various sounds. Chuffing, a soft pulsing call, is made by mother bears presumably to keep in contact with their young. During the breeding season, adult males will chuff at potential mates. Unlike other animals where chuffing is passed through the nostrils, in polar bears it is emitted through a partially open mouth. Cubs will cry for attention and produce humming noises while nursing. Teeth chops, jaw pops, blows, huffs, moans, growls and roars are heard in more hostile encounters. A polar bear visually communicates with its eyes, ears, nose and lips. Chemical communication can also be important: bears secrete their scent from their foot pads into their tracks, allowing individuals to keep track of one another.
Diet and hunting
The polar bear is a hypercarnivore, and the most carnivorous species of bear. It is an apex predator of the Arctic, preying on ice-living seals and consuming their energy-rich blubber. The most commonly taken species is the ringed seal, but they also prey on bearded seals and harp seals. Ringed seals are ideal prey as they are abundant and small enough to be overpowered by even small bears. Bearded seal adults are larger and are more likely to break free from an attacking bear, hence adult male bears are more successful in hunting them. Less common prey are hooded seals, spotted seals, ribbon seals and the more temperate-living harbour seals. Polar bears, mostly adult males, will occasionally hunt walruses, both on land and ice, though they mainly target the young, as adults are too large and formidable, with their thick skin and long tusks.
Bear feeding on a bearded seal
Besides seals, bears will prey on cetacean species such as beluga whales and narwhals, as well as reindeer, birds and their eggs, fish and marine invertebrates. They rarely eat plant material as their digestive system is too specialized for animal matter, though they have been recorded eating berries, moss, grass and seaweed. In their southern range, especially near Hudson Bay and James Bay, polar bears endure all summer without sea ice to hunt from and must subsist more on terrestrial foods. Fat reserves allow polar bears to survive for months without eating. Cannibalism is known to occur in the species.
Polar bears hunt their prey in several different ways. When a bear spots a seal hauling out on the sea ice, it slowly stalks it with the head and neck lowered, possibly to make its dark nose and eyes less noticeable. As it gets closer, the bear crouches more and eventually charges at a high speed, attempting to catch the seal before it can escape into its ice hole. Some stalking bears need to move through water; traversing through water cavities in the ice when approaching the seal or swimming towards a seal on an ice floe. The polar bear can stay underwater with its nose exposed. When it gets close enough, the animal lunges from the water to attack.
During a limited time in spring, polar bears will search for ringed seal pups in their birth lairs underneath the ice. Once a bear catches the scent of a hiding pup and pinpoints its location, it approaches the den quietly to not alert it. It uses its front feet to smash through the ice and then pokes its head in to catch the pup before it can escape. A ringed seal's lair can be more than 1 m (3.3 ft) below the surface of the ice and thus more massive bears are better equipped for breaking in. Some bears may simply stay still near a breathing hole or other spot near the water and wait for prey to come by. This can last hours and when a seal surfaces the bear will try to pull it out with its paws and claws. This tactic is the primary hunting method from winter to early spring.
Bear with whale carcass
Bears hunt walrus groups by provoking them into stampeding and then look for young that have been crushed or separated from their mothers during the turmoil. There are reports of bears trying to kill or injure walruses by throwing rocks and pieces of ice on them. Belugas and narwhals are vulnerable to bear attacks when they are stranded in shallow water or stuck in isolated breathing holes in the ice. When stalking reindeer, polar bears will hide in vegetation before an ambush. On some occasions, bears may try to catch prey in open water, swimming underneath a seal or aquatic bird. Seals in particular, however, are more agile than bears in the water. Polar bears rely on raw power when trying to kill their prey, and will employ bites and paw swipes. They have the strength to pull a mid-sized seal out of the water or haul a beluga carcass for quite some distance. Polar bears only occasionally store food for later—burying it under snow—and only in the short term.
Arctic foxes routinely follow polar bears and scavenge scraps from their kills. The bears usually tolerate them but will charge a fox that gets too close when they are feeding. Polar bears themselves will scavenge. Subadult bears will eat remains left behind by others. Females with cubs often abandon a carcass when they see an adult male approaching, though are less likely to if they have not eaten in a long time. Whale carcasses are a valuable food source, particularly on land and after the sea ice melts, and attract several bears. In one area in northeastern Alaska, polar bears have been recorded competing with grizzly bears for whale carcasses. Despite their smaller size, grizzlies are more aggressive and polar bears are likely to yield to them in confrontations. Polar bears will also scavenge at garbage dumps during ice-free periods.
Reproduction and development
Polar bear mating takes place on the sea ice and during spring, mostly between March and May. Males search for females in estrus and often travel in twisting paths which reduces the chances of them encountering other males while still allowing them to find females. The movements of females remain linear and they travel more widely. The mating system can be labelled as female-defence polygyny, serial monogamy or promiscuity.
Upon finding a female, a male will try to isolate and guard her. Courtship can be somewhat aggressive, and a male will pursue a female if she tries to run away. It can take days for the male to mate with the female which induces ovulation. After their first copulation, the couple bond. Undisturbed polar bear pairings typically last around two weeks during which they will sleep together and mate multiple times. Competition for mates can be intense and this has led to sexual selection for bigger males. Polar bear males often have scars from fighting. A male and female that have already bonded will flee together when another male arrives. A female mates with multiple males in a season and a single litter can have more than one father.
Polar bear cubs
When the mating season ends, the female will build up more fat reserves to sustain both herself and her young. Sometime between August and October, the female constructs and enters a maternity den for winter. Depending on the area, maternity dens can be found in sea ice just off the coastline or further inland and may be dug underneath snow, earth or a combination of both. The inside of these shelters can be around 1.5 m (4.9 ft) wide with a ceiling height of 1.2 m (3.9 ft) while the entrance may be 2.1 m (6.9 ft) long and 1.2 m (3.9 ft) wide. The temperature of a den can be much higher than the outside. Females hibernate and give birth to their cubs in the dens. Hibernating bears fast and internally recycle bodily waste. Polar bears experience delayed implantation and the fertilized embryo does not start development until the fall, between mid-September and mid-October. With delayed implantation, gestation in the species lasts seven to nine months but actual pregnancy is only two months.
Mother polar bears typically give birth to two cubs per litter. As with other bear species, newborn polar bears are tiny and altricial. The newborns have woolly hair and pink skin, with a weight of around 600 g (21 oz). Their eyes remain closed for a month. The mother's fatty milk fuels their growth, and the cubs are kept warm both by the mother's body heat and the den. The mother emerges from the den between late February and early April, and her cubs are well-developed and capable of walking with her. At this time they weigh 10–15 kilograms (22–33 lb). A polar bear family stays near the den for roughly two weeks; during this time the cubs will move and play around while the mother mostly rests. They eventually head out on the sea ice.
Cubs under a year old stay close to their mother. When she hunts, they stay still and watch until she calls them back. Observing and imitating the mother helps the cubs hone their hunting skills. After their first year they become more independent and explore. At around two years old, they are capable of hunting on their own. The young suckle their mother as she is lying on her side or sitting on her rump. A lactating female cannot conceive and give birth, and cubs are weaned between two and two-and-a-half years. She may simply leave her weaned young or they may be chased away by a courting male. Polar bears reach sexual maturity at around four years for females and six years for males. Females reach their adult size at 4 or 5 years of age while males are fully grown at twice that age.
Mortality
Polar bears can live up to 30 years. The bear's long lifespan and ability to consistently produce young offsets cub deaths in a population. Some cubs die in the dens or the womb if the female is not in good condition. Nevertheless, the female has a chance to produce a surviving litter the next spring if she can eat better in the coming year. Cubs will eventually starve if their mothers cannot kill enough prey. Cubs also face threats from wolves and adult male bears. Males kill cubs to bring their mother back into estrus but also kill young outside the breeding season for food. A female and her cubs can flee from the slower male. If the male can get close to a cub, the mother may try to fight him off, sometimes at the cost of her life.
Subadult bears, who are independent but not quite mature, have a particularly rough time as they are not as successful hunters as adults. Even when they do succeed, their kill will likely be stolen by a larger bear. Hence subadults have to scavenge and are often underweight and at risk of starvation. At adulthood, polar bears have a high survival rate, though adult males suffer injuries from fights over mates. Polar bears are especially susceptible to Trichinella, a parasitic roundworm they contract through cannibalism.
Conservation status
In 2015, the IUCN Red List categorized the polar bear as vulnerable due to a "decline in area of occupancy, extent of occurrence and/or quality of habitat". It estimated the total population to be between 22,000 and 31,000, and the current population trend is unknown. Threats to polar bear populations include climate change, pollution and energy development.
In 2021, the IUCN/SSC Polar Bear Specialist Group labelled four subpopulations (Barents and Chukchi Sea, Foxe Basin and Gulf of Boothia) as "likely stable", two (Kane Basin and M'Clintock Channel) as "likely increased" and three (Southern Beaufort Sea, Southern and Western Hudson Bay) as "likely decreased" over specific periods between the 1980s and 2010s. The remaining ten did not have enough data. A 2008 study predicted two-thirds of the world's polar bears may disappear by 2050, based on the reduction of sea ice, and only one population would likely survive in 50 years. A 2016 study projected a likely decline in polar bear numbers of more than 30 percent over three generations. The study concluded that declines of more than 50 percent are much less likely. A 2012 review suggested that polar bears may become regionally extinct in southern areas by 2050 if trends continue, leaving the Canadian Archipelago and northern Greenland as strongholds.
The key danger from climate change is malnutrition or starvation due to habitat loss. Polar bears hunt seals on the sea ice, and rising temperatures cause the ice to melt earlier in the year, driving the bears to shore before they have built sufficient fat reserves to survive the period of scarce food in the late summer and early fall. Thinner sea ice tends to break more easily, which makes it more difficult for polar bears to access seals. Insufficient nourishment leads to lower reproductive rates in adult females and lower survival rates in cubs and juvenile bears. Lack of access to seals also causes bears to find food on land which increases the risk of conflict with humans.
Reduction in sea ice cover also forces bears to swim longer distances, which further depletes their energy stores and occasionally leads to drowning. Increased ice mobility may result in less stable sites for dens or longer distances for mothers travelling to and from dens on land. Thawing of permafrost would lead to more fire-prone roofs for bears denning underground. Less snow may affect insulation while more rain could cause more cave-ins. The maximum corticosteroid-binding capacity of corticosteroid-binding globulin in polar bear serum correlates with stress in polar bears, and this has increased with climate warming. Disease-causing bacteria and parasites would flourish more readily in a warmer climate.
Oil and gas development also affects polar bear habitat. The Chukchi Sea Planning Area of northwestern Alaska, which has had many drilling leases, was found to be an important site for non-denning female bears. Oil spills are also a risk. A 2018 study found that ten percent or less of prime bear habitat in the Chukchi Sea is vulnerable to a potential spill, but a spill at full reach could impact nearly 40 percent of the polar bear population. Polar bears accumulate high levels of persistent organic pollutants such as polychlorinated biphenyl (PCBs) and chlorinated pesticides, due to their position at the top of the ecological pyramid. Many of these chemicals have been internationally banned due to the recognition of their harm to the environment. Traces of them have slowly dwindled in polar bears but persist and have even increased in some populations.
Polar bears receive some legal protection in all the countries they inhabit. The species has been labelled as threatened under the US Endangered Species Act since 2008, while the Committee on the Status of Endangered Wildlife in Canada listed it as of 'Special concern' since 1991. In 1973, the Agreement on the Conservation of Polar Bears was signed by all five nations with polar bear populations, Canada, Denmark (of which Greenland is an autonomous territory), Russia (then USSR), Norway and the US. This banned most harvesting of polar bears, allowed indigenous hunting using traditional methods, and promoted the preservation of bear habitat. The Convention on International Trade in Endangered Species of Wild Fauna lists the species under Appendix II, which allows regulated trade.
Relationship with humans
Polar bears have coexisted and interacted with circumpolar peoples for millennia. "White bears" are mentioned as commercial items in the Japanese book Nihon Shoki in the seventh century. It is not clear if these were polar bears or white-coloured brown bears. During the Middle Ages, Europeans considered white bears to be a novelty and were more familiar with brown- and black-coloured bears. The first known written account of the polar bear in its natural environment is found in the 13th-century anonymous Norwegian text Konungs skuggsjá, which mentions that "the white bear of Greenland wanders most of the time on the ice of the sea, hunting seals and whales and feeding on them" and says the bear is "as skillful a swimmer as any seal or whale".
Over the next centuries, several European explorers would mention polar bears and describe their habits. Such accounts became more accurate after the Enlightenment, and both living and dead specimens were brought back. Nevertheless, some fanciful reports continued, including the idea that polar bears cover their noses during hunts. A relatively accurate drawing of a polar bear is found in Henry Ellis's work A Voyage to Hudson's Bay (1748). Polar bears were formally classified as a species by Constantine Phipps after his 1773 voyage to the Arctic. Accompanying him was a young Horatio Nelson, who was said to have wanted to get a polar bear coat for his father but failed in his hunt. In his 1785 edition of Histoire Naturelle, Comte de Buffon mentions and depicts a "sea bear", clearly a polar bear, and "land bears", likely brown and black bears. This helped promote ideas about speciation. Buffon also mentioned a "white bear of the forest", possibly a Kermode bear.
Exploitation
Polar bears were hunted as early as 8,000 years ago, as indicated by archaeological remains at Zhokhov Island in the East Siberian Sea. The oldest graphic depiction of a polar bear shows it being hunted by a man with three dogs. This rock art was among several petroglyphs found at Pegtymel in Siberia and dates from the fifth to eighth centuries. Before access to firearms, native people used lances, bows and arrows and hunted in groups accompanied by dogs. Though hunting typically took place on foot, some people killed swimming bears from boats with a harpoon. Polar bears were sometimes killed in their dens. Killing a polar bear was considered a rite of passage for boys in some cultures. Native people respected the animal and hunts were subject to strict rituals. Bears were harvested for the fur, meat, fat, tendons, bones and teeth. The fur was worn and slept on, while the bones and teeth were made into tools. For the Netsilik, the individual who finally killed the bear had the right to its fur while the meat was passed to all in the party. Some people kept the cubs of slain bears.
Norsemen in Greenland traded polar bear furs in the Middle Ages. Russia traded polar bear products as early as 1556, with Novaya Zemlya and Franz Josef Land being important commercial centres. Large-scale hunting of bears at Svalbard occurred since at least the 18th century, when no less than 150 bears were killed each year by Russian explorers. In the next century, more Norwegians were harvesting the bears on the island. From the 1870s to the 1970s, around 22,000 of the animals were hunted in total. Over 150,000 polar bears in total were either killed or captured in Russia and Svalbard, from the 18th to the 20th century. In the Canadian Arctic, bears were harvested by commercial whalers especially if they could not get enough whales. The Hudson's Bay Company is estimated to have sold 15,000 polar bear coats between the late 19th century and early 20th century. In the mid-20th century, countries began to regulate polar bear harvesting, culminating in the 1973 agreement.
Polar bear meat was commonly eaten as rations by explorers and sailors in the Arctic. Its taste and texture have been described both positively and negatively. Some have called it too coarse with a powerful smell, while others praised it as a "royal dish". The liver was known for being too toxic to eat. This is due to the accumulation of vitamin A from their prey. Polar bear fat was also used in lamps when other fuel was unavailable. Polar bear rugs were almost ubiquitous on the floors of Norwegian churches by the 13th and 14th centuries. In more modern times, classical Hollywood actors would pose on bearskin rugs, notably Marilyn Monroe. Such images often had sexual connotations.
Conflicts
Road sign warning about the presence of polar bears. The Norwegian text translates into "Applies to all of Svalbard".
When the sea ice melts, polar bears, particularly subadults, conflict with humans over resources on land. They are attracted to the smell of human-made foods, particularly at garbage dumps and may be shot when they encroach on private property. In Churchill, Manitoba, local authorities maintain a "polar bear jail" where nuisance bears are held until the sea ice freezes again. Climate change has increased conflicts between the two species. Over 50 polar bears swarmed a town in Novaya Zemlya in February 2019, leading local authorities to declare a state of emergency.
From 1870 to 2014, there were an estimated 73 polar bear attacks on humans, which led to 20 deaths. The majority of attacks were by hungry males, typically subadults, while female attacks were usually in defence of the young. In comparison to brown and American black bears, attacks by polar bears were more often near and around where humans lived. This may be due to the bears getting desperate for food and thus more likely to seek out human settlements. As with the other two bear species, polar bears are unlikely to target more than two people at once. Though popularly thought of as the most dangerous bear, the polar bear is no more aggressive to humans than other species.
Captivity
The polar bear was a particularly sought-after species for exotic animal collectors due to being relatively rare and remote living, and its reputation as a ferocious beast. It is one of the few marine mammals that can reproduce well in captivity. They were originally kept only by royals and elites. The Tower of London got a polar bear as early as 1252 under King Henry III. In 1609, James VI and I of Scotland, England and Ireland were given two polar bear cubs by the sailor Jonas Poole, who got them during a trip to Svalbard. At the end of the 17th century, Frederick I of Prussia housed polar bears in menageries with other wild animals. He had their claws and canines removed to perform mock fights. Around 1726, Catherine I of Russia gifted two polar bears to Augustus II the Strong of Poland, who desired them for his animal collection. Later, polar bears were displayed to the public in zoos and circuses. In early 19th century, the species was exhibited at the Exeter Exchange in London, as well as menageries in Vienna and Paris. The first zoo in North America to exhibit a polar bear was the Philadelphia Zoo in 1859.
Polar bear exhibits were innovated by Carl Hagenbeck, who replaced cages and pits with settings that mimicked the animal's natural environment. In 1907, he revealed a complex panoramic structure at the Tierpark Hagenbeck Zoo in Hamburg consisting of exhibits made of artificial snow and ice separated by moats. Different polar animals were displayed on each platform, giving the illusion of them living together. Starting in 1975, Hellabrunn Zoo in Munich housed its polar bears in an exhibit which consisted of a glass barrier, a house, concrete platforms mimicking ice floes and a large pool. Inside the house were maternity dens, and rooms for the staff to prepare and store the food. The exhibit was connected to an outdoor yard for extra room. Similar naturalistic and "immersive" exhibits were opened in the early 21st century, such as the "Arctic Ring of Life" at the Detroit Zoo and Ontario's Cochrane Polar Bear Habitat. Many zoos in Europe and North America have stopped keeping polar bears due to the size and costs of their complex exhibits. In North America, the population of polar bears in zoos reached its zenith in 1975 with 229 animals and declined in the 21st century.
Polar bears have been trained to perform in circuses. Bears in general, being large, powerful, easy to train and human-like in form, were widespread in circuses, and the white coat of polar bears made them particularly attractive. Circuses helped change the polar bear's image from a fearsome monster to something more comical. Performing polar bears were used in 1888 by Circus Krone in Germany and later in 1904 by the Bostock and Wombwell Menagerie in England. Circus director Wilhelm Hagenbeck trained up to 75 polar bears to slide into a large tank through a chute. He began performing with them in 1908 and they had a particularly well-received show at the Hippodrome in London. Other circus tricks performed by polar bears involved tightropes, balls, roller skates and motorcycles. One of the most famous polar bear trainers in the second half of the twentieth century was the East German Ursula Böttcher, whose small stature contrasted with that of the large bears. Starting in the late 20th century, most polar bear acts were retired and the use of these bears for the circus is now prohibited in the US.
Several captive polar bears gained celebrity status in the late 20th and early 21st century, notably Knut of the Berlin Zoological Garden, who was rejected by his mother and had to be hand-reared by zookeepers. Another bear, Binky of the Alaska Zoo in Anchorage, became famous for attacking two visitors who got too close. Captive polar bears may pace back and forth, a stereotypical behaviour. In one study, they were recorded to have spent 14 percent of their days pacing. Gus of the Central Park Zoo was prescribed Prozac by a therapist for constantly swimming in his pool. To reduce stereotypical behaviours, zookeepers provide the bears with enrichment items to trigger their play behaviour. Zoo polar bears may appear green due to algae concentrations.
Cultural significance
Polar bears have prominent roles in Inuit culture and religion. The deity Torngarsuk is sometimes imagined as a giant polar bear. He resides underneath the sea floor in an underworld of the dead and has power over sea creatures. Kalaallit shamans would worship him through singing and dancing and were expected to be taken by him to the sea and consumed if he considered them worthy. Polar bears were also associated with the goddess Nuliajuk who was responsible for their creation, along with other sea creatures. It is believed that shamans could reach the Moon or the bottom of the ocean by riding on a guardian spirit in the form of a polar bear. Some folklore involves people turning into or disguising themselves as polar bears by donning their skins or the reverse, with polar bears removing their skins. In Inuit astronomy, the Pleiades star cluster is conceived of as a polar bear trapped by dogs while Orion's Belt, the Hyades and Aldebaran represent hunters, dogs and a wounded bear respectively.
Nordic folklore and literature have also featured polar bears. In The Tale of Auðun of the West Fjords, written around 1275, a poor man named Auðun spends all his money on a polar bear in Greenland, but ends up wealthy after giving the bear to the king of Denmark. In the 14th-century manuscript Hauksbók, a man named Odd kills and eats a polar bear that killed his father and brother. In the story of The Grimsey Man and the Bear, a mother bear nurses and rescues a farmer stuck on an ice floe and is repaid with sheep meat. 18th-century Icelandic writings mention the legend of a "polar bear king" known as the bjarndýrakóngur. This beast was depicted as a polar bear with "ruddy cheeks" and a unicorn-like horn, which glows in the dark. The king could understand when humans talk and was considered to be very astute. Two Norwegian fairy tales, "East of the Sun and West of the Moon" and "White-Bear-King-Valemon", involve white bears turning into men and seducing women.
Drawings of polar bears have been featured on maps of the northern regions. Possibly the earliest depictions of a polar bear on a map is the Swedish Carta marina of 1539, which has a white bear on Iceland or "Islandia". A 1544 map of North America includes two polar bears near Quebec. Notable paintings featuring polar bears include François-Auguste Biard's Fighting Polar Bears (1839) and Edwin Landseer's Man Proposes, God Disposes (1864). Polar bears have also been filmed for cinema. An Inuit polar bear hunt was shot for the 1932 documentary Igloo, while the 1974 film The White Dawn filmed a simulated stabbing of a trained bear for a scene. In the film The Big Show (1961), two characters are killed by a circus polar bear. The scenes were shot using animal trainers instead of the actors. In modern literature, polar bears have been characters in both children's fiction, like Hans Beer's Little Polar Bear and the Whales and Sakiasi Qaunaq's The Orphan and the Polar Bear, and fantasy novels, like Philip Pullman's His Dark Materials series. In radio, Mel Blanc provided the vocals for Jack Benny's pet polar bear Carmichael on The Jack Benny Program. The polar bear is featured on flags and coats of arms, like the coat of arms of Greenland, and in many advertisements, notably for Coca-Cola since 1922.
As charismatic megafauna, polar bears have been used to raise awareness of the dangers of climate change. Aurora the polar bear is a giant marionette created by Greenpeace for climate protests. The World Wide Fund for Nature has sold plush polar bears as part of its "Arctic Home" campaign. Photographs of polar bears have been featured in National Geographic and Time magazines, including ones of them standing on ice floes, while the climate change documentary and advocacy film An Inconvenient Truth (2006) includes an animated bear swimming. Automobile manufacturer Nissan used a polar bear in one of its commercials, hugging a man for using an electric car. To make a statement about global warming, in 2009 a Copenhagen ice statue of a polar bear with a bronze skeleton was purposely left to melt in the sun.
The raccoon (/rəˈkuːn/ or US: /ræˈkuːn/ ⓘ, Procyon lotor), also spelled racoon[3] and sometimes called the common raccoon to distinguish it from the other species, is a mammal native to North America. It is the largest of the procyonid family, having a body length of 40 to 70 cm (16 to 28 in), and a body weight of 5 to 26 kg (11 to 57 lb). Its grayish coat mostly consists of dense underfur, which insulates it against cold weather. Three of the raccoon's most distinctive features are its extremely dexterous front paws, its facial mask, and its ringed tail, which are themes in the mythologies of the indigenous peoples of the Americas relating to the animal. The raccoon is noted for its intelligence, as studies show that it is able to remember the solution to tasks for at least three years. It is usually nocturnal and omnivorous, eating about 40% invertebrates, 33% plants, and 27% vertebrates.
The original habitats of the raccoon are deciduous and mixed forests, but due to their adaptability, they have extended their range to mountainous areas, coastal marshes, and urban areas, where some homeowners consider them to be pests. As a result of escapes and deliberate introductions in the mid-20th century, raccoons are now also distributed across central Europe, the Caucasus, and Japan.
In Europe, the raccoon is included since 2016 in the list of Invasive Alien Species of Union concern (the Union list).[4] This implies that this species cannot be imported, bred, transported, commercialized, or intentionally released into the environment in the whole of the European Union.[5]
Though previously thought to be generally solitary, there is now evidence that raccoons engage in sex-specific social behavior. Related females often share a common area, while unrelated males live together in groups of up to four raccoons in order to maintain their positions against foreign males during the mating season and against other potential invaders. Home range sizes vary anywhere from 3 ha (7.4 acres) for females in cities, to 5,000 ha (12,000 acres) for males in prairies. After a gestation period of about 65 days, two to five young known as "kits" are born in spring. The kits are subsequently raised by their mother until dispersal in late fall. Although captive raccoons have been known to live over 20 years, their life expectancy in the wild is only 1.8 to 3.1 years. In many areas, hunting and vehicular injury are the two most common causes of death.
Etymology
The mask of a raccoon is often interrupted by a brown-black streak that extends from forehead to nose.[6]
Names for the species include the common raccoon,[7] North American raccoon,[8] and northern raccoon.[9] In various North American native languages, the reference to the animal's manual dexterity, or use of its hands is the source for the names.[10] The word raccoon was adopted into English from the native Powhatan term meaning 'animal that scratches with its hands', as used in the Colony of Virginia. It was recorded on John Smith's list of Powhatan words as aroughcun, and on that of William Strachey as arathkone.[11] It has also been identified as a reflex of a Proto-Algonquian root *ahrah-koon-em, meaning '[the] one who rubs, scrubs and scratches with its hands'.[12] The word is sometimes spelled as racoon.[13]
In Spanish, the raccoon is called mapache, derived from the Nahuatl mapachtli of the Aztecs, meaning '[the] one who takes everything in its hands'.[14]
Its Latin name literally means 'before-dog washer'.[15] The genus Procyon was named by Gottlieb Conrad Christian Storr.[10] The animal's observed habit of "washing" or "dowsing" (see below) is the source of its name in other languages.[16][17] For example, the French "raton laveur" means "washing rat".
The colloquial abbreviation coon is used in words like coonskin for fur clothing and in phrases like old coon as a self-designation of trappers.[18][19] In the 1830s, the United States Whig Party used the raccoon as an emblem, causing them to be pejoratively known as "coons" by their political opponents, who saw them as too sympathetic to African-Americans. Soon after that the term became an ethnic slur,[20] especially in use between 1880 and 1920 (see coon song), and the term is still considered offensive.[21] Dogs bred to hunt raccoons are called coonhound and coon dog.[22]
Taxonomy
Skins of P. lotor and P. cancrivorus
Skulls of P. lotor and P. cancrivorus
In the first decades after its discovery by the members of the expedition of Christopher Columbus, who were the first Europeans to leave a written record about the species, taxonomists thought the raccoon was related to many different species, including dogs, cats, badgers and particularly bears.[23] Carl Linnaeus, the father of modern taxonomy, placed the raccoon in the genus Ursus, first as Ursus cauda elongata ('long-tailed bear') in the second edition of his Systema Naturae (1740), then as Ursus Lotor ('washer bear') in the tenth edition (1758–59).[24][25] In 1780, Gottlieb Conrad Christian Storr placed the raccoon in its own genus Procyon, which can be translated as either 'before the dog' or 'doglike'.[26][27] It is also possible that Storr had its nocturnal lifestyle in mind and chose the star Procyon as eponym for the species.[28][29]
Evolution
Based on fossil evidence from Russia and Bulgaria, the first known members of the family Procyonidae lived in Europe in the late Oligocene about 25 million years ago.[30] Similar tooth and skull structures suggest procyonids and weasels share a common ancestor, but molecular analysis indicates a closer relationship between raccoons and bears.[31] After the then-existing species crossed the Bering Strait at least six million years later in the early Miocene, the center of its distribution was probably in Central America.[32] Coatis (Nasua and Nasuella) and raccoons (Procyon) have been considered to share common descent from a species in the genus Paranasua present between 5.2 and 6.0 million years ago.[33] This assumption, based on morphological comparisons of fossils, conflicts with a 2006 genetic analysis which indicates raccoons are more closely related to ringtails.[34] Unlike other procyonids, such as the crab-eating raccoon (Procyon cancrivorus), the ancestors of the common raccoon left tropical and subtropical areas and migrated farther north about 2.5 million years ago, in a migration that has been confirmed by the discovery of fossils in the Great Plains dating back to the middle of the Pliocene.[35][33] Its most recent ancestor was likely Procyon rexroadensis, a large Blancan raccoon from the Rexroad Formation characterized by its narrow back teeth and large lower jaw.[36]
Subspecies
A Torch Key raccoon (P. l. incautus) in Cudjoe Key, Florida. Subspecies inhabiting the Florida Keys are characterized by their small size and very pale fur.
Female raccoon of the Vancouver Island subspecies at Sidney, British Columbia, with characteristic dark fur
As of 2005, Mammal Species of the World recognizes 22 subspecies of raccoons.[37] Four of these subspecies living only on small Central American and Caribbean islands were often regarded as distinct species after their discovery. These are the Bahamian raccoon and Guadeloupe raccoon, which are very similar to each other; the Tres Marias raccoon, which is larger than average and has an angular skull; and the extinct Barbados raccoon. Studies of their morphological and genetic traits in 1999, 2003 and 2005 led all these island raccoons to be listed as subspecies of the common raccoon in Mammal Species of the World's third edition. A fifth island raccoon population, the Cozumel raccoon, which weighs only 3 to 4 kg (6.6 to 8.8 lb) and has notably small teeth, is still regarded as a separate species.[38][39][40][41]
The four smallest raccoon subspecies, with a typical weight of 1.8 to 2.7 kg (4.0 to 6.0 lb), live along the southern coast of Florida and on the adjacent islands; an example is the Ten Thousand Islands raccoon (Procyon lotor marinus).[42] Most of the other 15 subspecies differ only slightly from each other in coat color, size and other physical characteristics.[43][44] The two most widespread subspecies are the eastern raccoon (Procyon lotor lotor) and the Upper Mississippi Valley raccoon (Procyon lotor hirtus). Both share a comparatively dark coat with long hairs, but the Upper Mississippi Valley raccoon is larger than the eastern raccoon. The eastern raccoon occurs in all U.S. states and Canadian provinces to the north of South Carolina and Tennessee. The adjacent range of the Upper Mississippi Valley raccoon covers all U.S. states and Canadian provinces to the north of Louisiana, Texas and New Mexico.[45]
The taxonomic identity of feral raccoons inhabiting Central Europe, Causasia and Japan is unknown, as the founding populations consisted of uncategorized specimens from zoos and fur farms.
Description
Physical characteristics
Lower side of front paw with visible vibrissae on the tips of the digits
Skeleton
Skull with dentition: 2/2 molars, 4/4 premolars, 1/1 canines, 3/3 incisors
Baculum or penis bone
Female genitourinary system
Male genitourinary system
Head to hindquarters, raccoons measure between 40 and 70 cm (16 and 28 in), not including the bushy tail which can measure between 20 and 40 cm (7.9 and 15.7 in), but is usually not much longer than 25 cm (9.8 in).[71][72][73] The shoulder height is between 23 and 30 cm (9.1 and 11.8 in).[74] The body weight of an adult raccoon varies considerably with habitat, making the raccoon one of the most variably sized mammals. It can range from 2 to 26 kg (4.4 to 57.3 lb), but is usually between 5 and 12 kg (11 and 26 lb). The smallest specimens live in southern Florida, while those near the northern limits of the raccoon's range tend to be the largest (see Bergmann's rule).[75] Males are usually 15 to 20% heavier than females.[76] At the beginning of winter, a raccoon can weigh twice as much as in spring because of fat storage.[77][78][79] The largest recorded wild raccoon weighed 28.4 kg (63 lb) and measured 140 cm (55 in) in total length, by far the largest size recorded for a procyonid.[80][81]
The most characteristic physical feature of the raccoon is the area of black fur around the eyes, which contrasts sharply with the surrounding white face coloring. This is reminiscent of a "bandit's mask" and has thus enhanced the animal's reputation for mischief.[82][83] The slightly rounded ears are also bordered by white fur. Raccoons are assumed to recognize the facial expression and posture of other members of their species more quickly because of the conspicuous facial coloration and the alternating light and dark rings on the tail.[84][85][86] The dark mask may also reduce glare and thus enhance night vision.[85][86] On other parts of the body, the long and stiff guard hairs, which shed moisture, are usually colored in shades of gray and, to a lesser extent, brown.[87] Raccoons with a very dark coat are more common in the German population because individuals with such coloring were among those initially released to the wild.[88] The dense underfur, which accounts for almost 90% of the coat, insulates against cold weather and is composed of 2 to 3 cm (0.79 to 1.18 in) long hairs.[87]
The raccoon, whose method of locomotion is usually considered to be plantigrade, can stand on its hind legs to examine objects with its front paws.[89][90] As raccoons have short legs compared to their compact torso, they are usually not able either to run quickly or jump great distances.[91][92] Their top speed over short distances is 16 to 24 km/h (9.9 to 14.9 mph).[93][94] Raccoons can swim with an average speed of about 5 km/h (3.1 mph) and can stay in the water for several hours.[95][92] For climbing down a tree headfirst—an unusual ability for a mammal of its size—a raccoon rotates its hind feet so they are pointing backwards.[96][92] Raccoons have a dual cooling system to regulate their temperature; that is, they are able to both sweat and pant for heat dissipation.[97][98]
Raccoon skulls have a short and wide facial region and a voluminous braincase. The facial length of the skull is less than the cranial, and their nasal bones are short and quite broad. The auditory bullae are inflated in form, and the sagittal crest is weakly developed.[99] The dentition—40 teeth with the dental formula:
3.1.4.2
3.1.4.2
—is adapted to their omnivorous diet: the carnassials are not as sharp and pointed as those of a full-time carnivore, but the molars are not as wide as those of a herbivore.[100] The penis bone of males is about 10 cm (3.9 in) long and strongly bent at the front end,[101][102] and its shape can be used to distinguish juvenile males from mature males.[103][104][105] Seven of the thirteen identified vocal calls are used in communication between the mother and her kits, one of these being the birdlike twittering of newborns.[106][107][98]
Senses
The most important sense for the raccoon is its sense of touch.[108][109][110] The "hyper sensitive"[109] front paws are protected by a thin horny layer that becomes pliable when wet.[111][112] The five digits of the paws have no webbing between them, which is unusual for a carnivoran.[113] Almost two-thirds of the area responsible for sensory perception in the raccoon's cerebral cortex is specialized for the interpretation of tactile impulses, more than in any other studied animal.[114] They are able to identify objects before touching them with vibrissae located above their sharp, nonretractable claws.[89][110] The raccoon's paws lack an opposable thumb; thus, it does not have the agility of the hands of primates.[110][112] There is no observed negative effect on tactile perception when a raccoon stands in water below 10 °C (50 °F) for hours.[115]
Raccoons are thought to be color blind or at least poorly able to distinguish color, though their eyes are well-adapted for sensing green light.[116][117][118] Although their accommodation of 11 dioptre is comparable to that of humans and they see well in twilight because of the tapetum lucidum behind the retina, visual perception is of subordinate importance to raccoons because of their poor long-distance vision.[119][120][121] In addition to being useful for orientation in the dark, their sense of smell is important for intraspecific communication. Glandular secretions (usually from their anal glands), urine and feces are used for marking.[122][123][124] With their broad auditory range, they can perceive tones up to 50–85 kHz as well as quiet noises, like those produced by earthworms underground.[125][126]
Intelligence
Zoologist Clinton Hart Merriam described raccoons as "clever beasts", and that "in certain directions their cunning surpasses that of the fox". The animal's intelligence gave rise to the epithet "sly coon".[127] Only a few studies have been undertaken to determine the mental abilities of raccoons, most of them based on the animal's sense of touch. In a study by the ethologist H. B. Davis in 1908, raccoons were able to open 11 of 13 complex locks in fewer than 10 tries and had no problems repeating the action when the locks were rearranged or turned upside down. Davis concluded that they understood the abstract principles of the locking mechanisms and their learning speed was equivalent to that of rhesus macaques.[128]
Studies in 1963, 1973, 1975 and 1992 concentrated on raccoon memory showed that they can remember the solutions to tasks for at least three years.[129] In a study by B. Pohl in 1992, raccoons were able to instantly differentiate between identical and different symbols three years after the short initial learning phase.[129] Stanislas Dehaene reports in his book The Number Sense that raccoons can distinguish boxes containing two or four grapes from those containing three.[130] In research by Suzana Herculano-Houzel and other neuroscientists, raccoons have been found to be comparable to primates in density of neurons in the cerebral cortex, which they have proposed to be a neuroanatomical indicator of intelligence.[131][132]
Behavior
Social behavior
Eastern raccoons (P. l. lotor) in a tree: The raccoon's social structure is grouped into what Ulf Hohmann calls a "three-class society".
California raccoon (P. l. psora) climbing a tree in Lower Klamath National Wildlife Refuge
Baby raccoon chatter
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Studies in the 1990s by the ethologists Stanley D. Gehrt and Ulf Hohmann suggest that raccoons engage in sex-specific social behaviors and are not typically solitary, as was previously thought.[133][134] Related females often live in a so-called "fission-fusion society"; that is, they share a common area and occasionally meet at feeding or resting grounds.[135][136] Unrelated males often form loose male social groups to maintain their position against foreign males during the mating season—or against other potential invaders.[137] Such a group does not usually consist of more than four individuals.[138][139] Since some males show aggressive behavior towards unrelated kits, mothers will isolate themselves from other raccoons until their kits are big enough to defend themselves.[140]
With respect to these three different modes of life prevalent among raccoons, Hohmann called their social structure a "three-class society".[141] Samuel I. Zeveloff, professor of zoology at Weber State University and author of the book Raccoons: A Natural History, is more cautious in his interpretation and concludes at least the females are solitary most of the time and, according to Erik K. Fritzell's study in North Dakota in 1978, males in areas with low population densities are solitary as well.[142]
The shape and size of a raccoon's home range varies depending on age, sex, and habitat, with adults claiming areas more than twice as large as juveniles.[143] While the size of home ranges in the habitat of North Dakota's prairies lie between 7 and 50 km2 (3 and 20 sq mi) for males and between 2 and 16 km2 (1 and 6 sq mi) for females, the average size in a marsh at Lake Erie was 0.5 km2 (0.19 sq mi).[144] Irrespective of whether the home ranges of adjacent groups overlap, they are most likely not actively defended outside the mating season if food supplies are sufficient.[145] Odor marks on prominent spots are assumed to establish home ranges and identify individuals.[124] Urine and feces left at shared raccoon latrines may provide additional information about feeding grounds, since raccoons were observed to meet there later for collective eating, sleeping and playing.[146]
Concerning the general behavior patterns of raccoons, Gehrt points out that "typically you'll find 10 to 15 percent that will do the opposite" of what is expected.[147]
Diet
Though usually nocturnal, the raccoon is sometimes active in daylight to take advantage of available food sources.[148][149] Its diet consists of about 40% invertebrates, 33% plant material and 27% vertebrates.[150] Since its diet consists of such a variety of different foods, Zeveloff argues the raccoon "may well be one of the world's most omnivorous animals".[151] While its diet in spring and early summer consists mostly of insects, worms, and other animals already available early in the year, it prefers fruits and nuts, such as acorns and walnuts, which emerge in late summer and autumn, and represent a rich calorie source for building up fat needed for winter.[152][153]
Contrary to popular belief, raccoons only occasionally eat active or large prey, such as birds and mammals. They prefer prey that is easier to catch, specifically crayfish, insects,[154] fish, amphibians and bird eggs.[155] Raccoons are virulent predators of eggs and hatchlings in both birds and reptile nests, to such a degree that, for threatened prey species, raccoons may need to be removed from the area or nests may need to be relocated to mitigate the effect of their predations (i.e. in the case of some globally threatened turtles).[156][157][158][159][160] When food is plentiful, raccoons can develop strong individual preferences for specific foods.[78] In the northern parts of their range, raccoons go into a winter rest, reducing their activity drastically as long as a permanent snow cover makes searching for food difficult.[161]
Dousing
Captive raccoons often douse their food before eating.
One aspect of raccoon behavior is so well known that it gives the animal part of its scientific name, Procyon lotor; lotor is Latin for 'washer'. In the wild, raccoons often dabble for underwater food near the shore-line. They then often pick up the food item with their front paws to examine it and rub the item, sometimes to remove unwanted parts. This gives the appearance of the raccoon "washing" the food. The tactile sensitivity of raccoons' paws is increased if this rubbing action is performed underwater, since the water softens the hard layer covering the paws.[109][162] However, the behavior observed in captive raccoons in which they carry their food to water to "wash" or douse it before eating has not been observed in the wild.[163][164] Naturalist Georges-Louis Leclerc, Comte de Buffon, believed that raccoons do not have adequate saliva production to moisten food thereby necessitating dousing, but this hypothesis is now considered to be incorrect.[162][163][165][166] Captive raccoons douse their food more frequently when a watering hole with a layout similar to a stream is not farther away than 3 m (10 ft).[166] The widely accepted theory is that dousing in captive raccoons is a fixed action pattern from the dabbling behavior performed when foraging at shores for aquatic foods.[162][166][167][168] This is supported by the observation that aquatic foods are doused more frequently. Cleaning dirty food does not seem to be a reason for "washing".[166]
Reproduction
Raccoons usually mate in a period triggered by increasing daylight between late January and mid-March.[169][170][171] However, there are large regional differences which are not completely explicable by solar conditions. For example, while raccoons in southern states typically mate later than average, the mating season in Manitoba also peaks later than usual in March and extends until June.[171] During the mating season, males restlessly roam their home ranges in search of females in an attempt to court them during the three- to four-day period when conception is possible. These encounters will often occur at central meeting places.[172][173][174] Copulation, including foreplay, can last over an hour and is repeated over several nights.[175] The weaker members of a male social group also are assumed to get the opportunity to mate, since the stronger ones cannot mate with all available females.[176] In a study in southern Texas during the mating seasons from 1990 to 1992, about one third of all females mated with more than one male.[177] If a female does not become pregnant or if she loses her kits early, she will sometimes become fertile again 80 to 140 days later.[178][179][180]
An eastern raccoon (P. l. lotor) kit
After usually 63 to 65 days of gestation (although anywhere from 54 to 70 days is possible), a litter of typically two to five young is born.[181][182] The average litter size varies widely with habitat, ranging from 2.5 in Alabama to 4.8 in North Dakota.[183][184] Larger litters are more common in areas with a high mortality rate, due, for example, to hunting or severe winters.[185][184] While male yearlings usually reach their sexual maturity only after the main mating season, female yearlings can compensate for high mortality rates and may be responsible for about 50% of all young born in a year.[186][187][188] Males have no part in raising young.[138][189][190] The kits (also called "cubs") are blind and deaf at birth, but their mask is already visible against their light fur.[191][192] The birth weight of the about 10 cm (4 in)-long kits is between 60 and 75 g (2.1 and 2.6 oz).[192] Their ear canals open after around 18 to 23 days, a few days before their eyes open for the first time.[193] Once the kits weigh about 1 kg (2 lb), they begin to explore outside the den, consuming solid food for the first time after six to nine weeks.[194][195] After this point, their mother suckles them with decreasing frequency; they are usually weaned by 16 weeks.[196] In the fall, after their mother has shown them dens and feeding grounds, the juvenile group splits up.[197] [198] While many females will stay close to the home range of their mother, males can sometimes move more than 20 km (12 mi) away. This is considered an instinctive behavior, preventing inbreeding. However, mother and offspring may share a den during the first winter in cold areas.
Life expectancy
Captive raccoons have been known to live for more than 20 years. However, the species' life expectancy in the wild is only 1.8 to 3.1 years, depending on the local conditions such as traffic volume, hunting, and weather severity. It is not unusual for only half of the young born in one year to survive a full year. After this point, the annual mortality rate drops to between 10% and 30%. Young raccoons are vulnerable to losing their mother and to starvation, particularly in long and cold winters. The most frequent natural cause of death in the North American raccoon population is distemper, which can reach epidemic proportions and kill most of a local raccoon population. In areas with heavy vehicular traffic and extensive hunting, these factors can account for up to 90% of all deaths of adult raccoons. The most important natural predators of the raccoon are bobcats, coyotes, and great horned owls, the latter mainly preying on young raccoons but capable of killing adults in some cases. In Florida, they have been reported to fall victim to larger carnivores like American black bear and cougars and these species may also be a threat on occasion in other areas. Where still present, gray wolves may still occasionally take raccoons as a supplemental prey item. Also in the southeast, they are among the favored prey for adult American alligators. On occasion, both bald and golden eagles will prey on raccoons. In the tropics, raccoons are known to fall prey to smaller eagles such as ornate hawk-eagles and black hawk-eagles, although it is not clear whether adults or merely juvenile raccoons are taken by these. In rare cases of overlap, they may fall victim from carnivores ranging from species averaging smaller than themselves such as fishers to those as large and formidable as jaguars in Mexico. In their introduced range in the former Soviet Union, their main predators are wolves, lynxes and Eurasian eagle-owls. However, predation is not a significant cause of death, especially because larger predators have been exterminated in many areas inhabited by raccoons.
Range
Although they have thrived in sparsely wooded areas in the last decades, raccoons depend on vertical structures to climb when they feel threatened. Therefore, they avoid open terrain and areas with high concentrations of beech trees, as beech bark is too smooth to climb. Tree hollows in old oaks or other trees and rock crevices are preferred by raccoons as sleeping, winter and litter dens. If such dens are unavailable or accessing them is inconvenient, raccoons use burrows dug by other mammals, dense undergrowth or tree crotches. In a study in the Solling range of hills in Germany, more than 60% of all sleeping places were used only once, but those used at least ten times accounted for about 70% of all uses. Since amphibians, crustaceans, and other animals around the shore of lakes and rivers are an important part of the raccoon's diet, lowland deciduous or mixed forests abundant with water and marshes sustain the highest population densities. While population densities range from 0.5 to 3.2 animals per square kilometer (1.3 to 8.3 animals per square mile) in prairies and do not usually exceed 6 animals per square kilometer (15.5 animals per square mile) in upland hardwood forests, more than 20 raccoons per square kilometer (51.8 animals per square mile) can live in lowland forests and marshes.
Distribution in North America
Raccoons are common throughout North America from Canada to Panama, where the subspecies Procyon lotor pumilus coexists with the crab-eating raccoon (Procyon cancrivorus). The population on Hispaniola was exterminated as early as 1513 by Spanish colonists who hunted them for their meat. Raccoons were also exterminated in Cuba and Jamaica, where the last sightings were reported in 1687. The Barbados raccoon became extinct relatively recently, in 1964. When they were still considered separate species, the Bahamas raccoon, Guadeloupe raccoon and Tres Marias raccoon were classified as endangered by the IUCN in 1996.
A Raccoon sleeping on a tree in High Park, Toronto
There is archeological evidence that in pre-Columbian times raccoons were numerous only along rivers and in the woodlands of the Southeastern United States. As raccoons were not mentioned in earlier reports of pioneers exploring the central and north-central parts of the United States, their initial spread may have begun a few decades before the 20th century. Since the 1950s, raccoons have expanded their range from Vancouver Island—formerly the northernmost limit of their range—far into the northern portions of the four south-central Canadian provinces. New habitats which have recently been occupied by raccoons (aside from urban areas) include mountain ranges, such as the Western Rocky Mountains, prairies and coastal marshes. After a population explosion starting in the 1940s, the estimated number of raccoons in North America in the late 1980s was 15 to 20 times higher than in the 1930s, when raccoons were comparatively rare. Urbanization, the expansion of agriculture, deliberate introductions, and the extermination of natural predators of the raccoon have probably caused this increase in abundance and distribution.
Distribution outside North America
s a result of escapes and deliberate introductions in the mid-20th century, the raccoon is now distributed in several European and Asian countries. Sightings have occurred in all the countries bordering Germany, which hosts the largest population outside of North America. Another stable population exists in northern France, where several pet raccoons were released by members of the U.S. Air Force near the Laon-Couvron Air Base in 1966. Furthermore, raccoons have been known to be in the area around Madrid since the early 1970s. In 2013, the city authorized "the capture and death of any specimen". It is also present in Italy, with one self-sustaining population in Lombardy.
About 1,240 animals were released in nine regions of the former Soviet Union between 1936 and 1958 for the purpose of establishing a population to be hunted for their fur. Two of these introductions were successful – one in the south of Belarus between 1954 and 1958, and another in Azerbaijan between 1941 and 1957. With a seasonal harvest of between 1,000~1,500 animals, in 1974 the estimated size of the population distributed in the Caucasus region was around 20,000 animals and the density was four animals per square kilometer (10 animals per square mile).
Distribution in Japan
In Japan, up to 1,500 raccoons were imported as pets each year after the success of the anime series Rascal the Raccoon (1977). In 2004, the descendants of discarded or escaped animals lived in 42 of 47 prefectures. The range of raccoons in the wild in Japan grew from 17 prefectures in 2000 to all 47 prefectures in 2008. It is estimated that raccoons cause thirty million yen (~$275,000) of agricultural damage on Hokkaido alone.
Distribution in Germany
In Germany – where the raccoon is called the Waschbär (literally, 'wash-bear' or 'washing bear') due to its habit of "dousing" food in water – two pairs of pet raccoons were released into the German countryside at the Edersee reservoir in the north of Hesse in April 1934 by a forester upon request of their owner, a poultry farmer. He released them two weeks before receiving permission from the Prussian hunting office to "enrich the fauna". Several prior attempts to introduce raccoons in Germany had been unsuccessful. A second population was established in eastern Germany in 1945 when 25 raccoons escaped from a fur farm at Wolfshagen (today district of Altlandsberg), east of Berlin, after an air strike. The two populations are parasitologically distinguishable: 70% of the raccoons of the Hessian population are infected with the roundworm Baylisascaris procyonis, but none of the Brandenburgian population is known to have the parasite. In the Hessian region, there were an estimated 285 raccoons in 1956, which increased to over 20,000 in 1970; in 2008 there were between 200,000 and 400,000 raccoons in the whole of Germany. By 2012 it was estimated that Germany now had more than a million raccoons.
The raccoon was once a protected species in Germany, but has been declared a game animal in 14 of the 16 German states since 1954. Hunters and environmentalists argue the raccoon spreads uncontrollably, threatens protected bird species, and supersedes indigenous competitors. This view is opposed by the zoologist Frank-Uwe Michler, who finds no evidence that a high population density of raccoons leads to negative effects on the biodiversity of an area. Hohmann holds that extensive hunting cannot be justified by the absence of natural predators, because predation is not a significant cause of death in the North American raccoon population.
Graphs are unavailable due to technical issues. There is more info on Phabricator and on MediaWiki.org.
The raccoon is extensively hunted in Germany as it is seen as an invasive species and pest. In the 1990s, only about 400 raccoons were hunted yearly. This increased dramatically over the next quarter-century: during the 2015–2016 hunting season, 128,100 raccoons were hunted, 60 percent of them in the state of Hesse.
Distribution in the former Soviet Union
Experiments in acclimatising raccoons into the Soviet Union began in 1936, and were repeated a further 25 times until 1962. Overall, 1,222 individuals were released, 64 of which came from zoos and fur farms (38 of them having been imports from western Europe). The remainder originated from a population previously established in Transcaucasia. The range of Soviet raccoons was never single or continuous, as they were often introduced to different locations far from each other. All introductions into the Russian Far East failed; melanistic raccoons were released on Petrov Island near Vladivostok and some areas of southern Primorsky Krai, but died. In Middle Asia, raccoons were released in Kyrgyzstan's Jalal-Abad Province, though they were later recorded as "practically absent" there in January 1963. A large and stable raccoon population (yielding 1,000~1,500 catches a year) was established in Azerbaijan after an introduction to the area in 1937. Raccoons apparently survived an introduction near Terek, along the Sulak River into the Dagestani lowlands. Attempts to settle raccoons on the Kuban River's left tributary and Kabardino-Balkaria were unsuccessful. A successful acclimatization occurred in Belarus, where three introductions (consisting of 52, 37, and 38 individuals in 1954 and 1958) took place. By January 1963, 700 individuals were recorded in the country.
Urban raccoons
Due to its adaptability, the raccoon has been able to use urban areas as a habitat. The first sightings were recorded in a suburb of Cincinnati in the 1920s. Since the 1950s, raccoons have been present in metropolitan areas like Washington, DC, Chicago, Toronto, and New York City. Since the 1960s, Kassel has hosted Europe's first and densest population in a large urban area, with about 50 to 150 animals per square kilometer (130 to 390 animals per square mile), a figure comparable to those of urban habitats in North America. Home range sizes of urban raccoons are only 3 to 40 hectares (7.5 to 100 acres) for females and 8 to 80 hectares (20 to 200 acres) for males. In small towns and suburbs, many raccoons sleep in a nearby forest after foraging in the settlement area. Fruit and insects in gardens and leftovers in municipal waste are easily available food sources. Furthermore, a large number of additional sleeping areas exist in these areas, such as hollows in old garden trees, cottages, garages, abandoned houses, and attics. The percentage of urban raccoons sleeping in abandoned or occupied houses varies from 15% in Washington, DC (1991) to 43% in Kassel (2003).
Health
Raccoons can carry rabies, a lethal disease caused by the neurotropic rabies virus carried in the saliva and transmitted by bites. Its spread began in Florida and Georgia in the 1950s and was facilitated by the introduction of infected individuals to Virginia and North Dakota in the late 1970s. Of the 6,940 documented rabies cases reported in the United States in 2006, 2,615 (37.7%) were in raccoons. The U.S. Department of Agriculture, as well as local authorities in several U.S. states and Canadian provinces, has developed oral vaccination programs to fight the spread of the disease in endangered populations. Only one human fatality has been reported after transmission of the rabies virus strain commonly known as "raccoon rabies". Among the main symptoms for rabies in raccoons are a generally sickly appearance, impaired mobility, abnormal vocalization, and aggressiveness. There may be no visible signs at all, however, and most individuals do not show the aggressive behavior seen in infected canids; rabid raccoons will often retire to their dens instead. Organizations like the U.S. Forest Service encourage people to stay away from animals with unusual behavior or appearance, and to notify the proper authorities, such as an animal control officer from the local health department. Since healthy animals, especially nursing mothers, will occasionally forage during the day, daylight activity is not a reliable indicator of illness in raccoons.
Unlike rabies and at least a dozen other pathogens carried by raccoons, distemper, an epizootic virus, does not affect humans. This disease is the most frequent natural cause of death in the North American raccoon population and affects individuals of all age groups. For example, 94 of 145 raccoons died during an outbreak in Clifton, Ohio, in 1968. It may occur along with a following inflammation of the brain (encephalitis), causing the animal to display rabies-like symptoms. In Germany, the first eight cases of distemper were reported in 2007.
Some of the most important bacterial diseases which affect raccoons are leptospirosis, listeriosis, tetanus, and tularemia. Although internal parasites weaken their immune systems, well-fed individuals can carry a great many roundworms in their digestive tracts without showing symptoms. The larvae of the roundworm Baylisascaris procyonis, which can be contained in the feces and seldom causes a severe illness in humans, can be ingested when cleaning raccoon latrines without wearing breathing protection.
While not endemic, the worm Trichinella does infect raccoons, and undercooked raccoon meat has caused trichinosis in humans.
Trematode Metorchis conjunctus can also infect raccoons.
Relationship with humans
Raccoons have become notorious in urban areas for consuming food waste. They possess impressive problem-solving abilities and can break into all but the most secure food waste bins, which has earned them the derisive nickname trash panda. The presence of raccoons in close proximity to humans may be undesirable, as raccoon droppings (like most wild animals) contain parasites and other disease vectors. Raccoon roundworm is of particular concern to public health. It can be contracted in humans by accidental ingestion or inhalation of the eggs, which are present in the feces of infected raccoons. While usually harmless to the host, it causes progressive neurological damage in humans, and is eventually fatal if untreated. It is found in about 60% of adult raccoons. The general presence of raccoons in an area is not typically of concern, but nests or droppings found within or near structures should be destroyed. Roundworm eggs are very robust and bleach alone is insufficient; burning or treatment with hot solutions of sodium hydroxide is required. The keeping of raccoons as pets is illegal in some jurisdictions due to these risks.
The increasing number of raccoons in urban areas has resulted in diverse reactions in humans, ranging from outrage at their presence to deliberate feeding. Some wildlife experts and most public authorities caution against feeding wild animals because they might become increasingly obtrusive and dependent on humans as a food source. Other experts challenge such arguments and give advice on feeding raccoons and other wildlife in their books. Raccoons without a fear of humans are a concern to those who attribute this trait to rabies, but scientists point out this behavior is much more likely to be a behavioral adjustment to living in habitats with regular contact to humans for many generations. Raccoons usually do not prey on domestic cats and dogs, but isolated cases of killings have been reported. Attacks on pets may also target their owners.
A Florida raccoon (P. l. elucus) in the Everglades approaches a group of humans, hoping to be fed
While overturned waste containers and raided fruit trees are just a nuisance to homeowners, it can cost several thousand dollars to repair damage caused by the use of attic space as dens. Relocating or killing raccoons without a permit is forbidden in many urban areas on grounds of animal welfare. These methods usually only solve problems with particularly wild or aggressive individuals, since adequate dens are either known to several raccoons or will quickly be rediscovered. Loud noises, flashing lights, and unpleasant odors have proven particularly effective in driving away a mother and her kits before they would normally leave the nesting place (when the kits are about eight weeks old). Typically, though, only precautionary measures to restrict access to food waste and den sites are effective in the long term.
Among all fruits and crops cultivated in agricultural areas, sweet corn in its milk stage is particularly popular among raccoons.[309][310] In a two-year study by Purdue University researchers, published in 2004, raccoons were responsible for 87% of the damage to corn plants. Like other predators, raccoons searching for food can break into poultry houses to feed on chickens, ducks, their eggs, or food.
Mythology, arts, and entertainment
In the mythology of the Indigenous peoples of the Americas, the raccoon is the subject of folk tales. Stories such as "How raccoons catch so many crayfish" from the Tuscarora centered on its skills at foraging. In other tales, the raccoon played the role of the trickster which outsmarts other animals, like coyotes and wolves. Among others, the Dakota believe the raccoon has natural spirit powers, since its mask resembles the facial paintings, two-fingered swashes of black and white, used during rituals to connect to spirit beings. The Aztecs linked supernatural abilities especially to females, whose commitment to their young was associated with the role of wise women in their society.
The raccoon also appears in Native American art across a wide geographic range. Petroglyphs with engraved raccoon tracks were found in Lewis Canyon, Texas; at the Crow Hollow petroglyph site in Grayson County, Kentucky; and in river drainages near Tularosa, the San Francisco River of New Mexico and Arizona. The meaning and significance of the Raccoon Priests Gorget, which features a stylized carving of a raccoon and was found at the Spiro Mounds, Oklahoma, remains unknown.
Hunting and fur trade
The fur of raccoons is used for clothing, especially for coats and coonskin caps. At present, it is the material used for the inaccurately named "sealskin" cap worn by the Royal Fusiliers of Great Britain. Sporrans made of raccoon pelt and hide have sometimes been used as part of traditional Scottish highland men's apparel since the 18th century, especially in North America. Such sporrans may or may not be of the "full-mask" type. Historically, Native American tribes not only used the fur for winter clothing, but also used the tails for ornament. The famous Sioux leader Spotted Tail took his name from a raccoon skin hat with the tail attached he acquired from a fur trader. Since the late 18th century, various types of scent hounds, called coonhounds, which are able to tree animals have been bred in the United States. In the 19th century, when coonskins occasionally even served as means of payment, several thousand raccoons were killed each year in the United States. This number rose quickly when automobile coats became popular after the turn of the 20th century. In the 1920s, wearing a raccoon coat was regarded as status symbol among college students. Attempts to breed raccoons in fur farms in the 1920s and 1930s in North America and Europe turned out not to be profitable, and farming was abandoned after prices for long-haired pelts dropped in the 1940s. Although raccoons had become rare in the 1930s, at least 388,000 were killed during the hunting season of 1934–1935.
After persistent population increases began in the 1940s, the seasonal coon hunting harvest reached about one million animals in 1946–1947 and two million in 1962–1963. The broadcast of three television episodes about the frontiersman Davy Crockett and the film Davy Crockett, King of the Wild Frontier in 1954 and 1955 led to a high demand for coonskin caps in the United States, although it is unlikely either Crockett or the actor who played him, Fess Parker, actually wore a cap made from raccoon fur. The seasonal hunt reached an all-time high with 5.2 million animals in 1976–1977 and ranged between 3.2 and 4.7 million for most of the 1980s. In 1982, the average pelt price was $20. As of 1987, the raccoon was identified as the most important wild furbearer in North America in terms of revenue. In the first half of the 1990s, the seasonal hunt dropped to 0.9 from 1.9 million due to decreasing pelt prices.
Food
While primarily hunted for their fur, raccoons were also a source of food for Native Americans and early American settlers. According to Ernest Thompson Seton, young specimens killed without a fight are palatable, whereas old raccoons caught after a lengthy battle are inedible. Raccoon meat was extensively eaten during the early years of California, where it was sold in the San Francisco market for $1–3 apiece. American slaves occasionally ate raccoon at Christmas, but it was not necessarily a dish of the poor or rural. The first edition of The Joy of Cooking, released in 1931, contained a recipe for preparing raccoon, and US President Calvin Coolidge's pet raccoon Rebecca was originally sent to be served at the White House Thanksgiving Dinner.
Although the idea of eating raccoons may seem repulsive to most mainstream consumers, who see them as endearing, cute, or vermin, several thousand raccoons are still eaten each year in the United States, primarily in the Southern United States. Some people tout the taste of the meat.
Other uses
In addition to the fur and meat, the raccoon baculum (penis bone) have had numerous traditional uses in the Southern United States and beyond. Indigenous people used the bones as a pipe cleaning tool. The bones were used by moonshine distillers to guide the flow of whiskey from the drip tube to the bottle. With their tips filed down, the bones were used as toothpicks under the moniker "coon rods". In hoodoo, the folk magic of the American South, the baculum is sometimes worn as an amulet for love or luck. The bones also have decorative uses (e.g. on the trademark hat of stock car racer Richard Petty or as earrings by actresses Sarah Jessica Parker and Vanessa Williams).
Pet raccoons
Raccoons are sometimes kept as pets, which is discouraged by many experts because the raccoon is not a domesticated species. Raccoons may act unpredictably and aggressively and it is extremely difficult to teach them to obey commands. In places where keeping raccoons as pets is not forbidden, such as in Wisconsin and other U.S. states, an exotic pet permit may be required. One notable pet raccoon was Rebecca, kept by US president Calvin Coolidge.
Their propensity for unruly behavior exceeds that of captive skunks, and they are even less trustworthy when allowed to roam freely. Because of their intelligence and nimble forelimbs, even inexperienced raccoons are easily capable of unscrewing jars, uncorking bottles and opening door latches, with more experienced specimens having been recorded to open door knobs. Sexually mature raccoons often show aggressive natural behaviors such as biting during the mating season. Neutering them at around five or six months of age decreases the chances of aggressive behavior developing. Raccoons can become obese and suffer from other disorders due to poor diet and lack of exercise. When fed with cat food over a long time period, raccoons can develop gout. With respect to the research results regarding their social behavior, it is now required by law in Austria and Germany to keep at least two individuals to prevent loneliness. Raccoons are usually kept in a pen (indoor or outdoor), also a legal requirement in Austria and Germany, rather than in the apartment where their natural curiosity may result in damage to property.
When orphaned, it is possible for kits to be rehabilitated and reintroduced to the wild. However, it is uncertain whether they readapt well to life in the wild. Feeding unweaned kits with cow's milk rather than a kitten replacement milk or a similar product can be dangerous to their health.
Hidden
The coat of the klipspringer, yellowish grey to reddish brown, acts as an efficient camouflage in its rocky habitat; the underbelly is white. Unlike most other antelopes, the klipspringer has a thick and coarse coat with hollow, brittle hairs. The incisors might even get damaged by the hairs while grooming. However, the coat is a significant adaptation that saves the animal during steep falls and provides effective insulation in the extreme climates characteristic of its mountain habitat. A study showed that ticks occur in larger numbers on the underbelly, where the hair is less coarse. The hair often turns erect, especially if the animal is ill or if its temperature increases.
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The photo was taken during January 2014 at Mabula Private Game Reserve, Bela-Bela, Limpopo, South Africa.
Info source: en.wikipedia.org/wiki/Klipspringer
Sunda Colugo (Galeopterus variegatus). The Sunda or Malayan Colugo is the most widespread of the genus Galeopterus; it occurs in parts of southern Myanmar and southern Thailand, localised areas of Laos, Cambodia and Vietnam, and has an extensive range in Peninsular Malaysia, Singapore and Sumatra where suitable forest still exists.
A divergent lineage of Galeopterus occurs in Java (Janečka et al, 2008), which some consider as a separate species, and the island of Borneo supports the widely-recognised Bornean Colugo Galeopterus borneanus. Mason (2016) concluded that there may actually be 6 species of Galeopterus.
Colugos have large eyes with excellent night vision, small ears and a pointed snout. Some of its teeth are unusual, for example the lower incisors point outwards and are comb-like in structure.
Their diet includes leaves and young shoots. During the day colugos rest mainly on tree trunks, generally high up, but sometimes just three metres or so from the ground; they may also make use of tree holes. At dusk they become active, gliding from tree to tree to feed.
Females carry a single young (rarely twins) which clings underneath; when roosting, the young often poke their head out from beneath the female. In flight, the young are carried clinging to the flight membrane. Males are smaller than females.
The Sunda Colugo is generally mottled grey or greenish-grey but many individuals are reddish to yellowish-orange; males are predominantly reddish.
Colugos are rarely heard vocalising; Lim (2007) describes calls from sparring Sunda Colugo males as "a cracking or ripping sound, like the ripping of a thick piece of cardboard".
In Singapore this species has proven itself to be adaptable to fragmented forests in semi-urban settings; it makes use of tree plantings next to major roads and other linear plantings to navigate between fragments of habitat. It has been observed gliding across 6-lane highways, if the median divider supports mature trees.
Photo by Nick Dobbs, Berjaya Resort, Langkawi 18-04-2025
Sheep (pl.: sheep) or domestic sheep (Ovis aries) are a domesticated, ruminant mammal typically kept as livestock. Although the term sheep can apply to other species in the genus Ovis, in everyday usage it almost always refers to domesticated sheep. Like all ruminants, sheep are members of the order Artiodactyla, the even-toed ungulates. Numbering a little over one billion, domestic sheep are also the most numerous species of sheep. An adult female is referred to as a ewe (/juː/ yoo), an intact male as a ram, occasionally a tup, a castrated male as a wether, and a young sheep as a lamb.
Sheep are most likely descended from the wild mouflon of Europe and Asia, with Iran being a geographic envelope of the domestication center. One of the earliest animals to be domesticated for agricultural purposes, sheep are raised for fleeces, meat (lamb, hogget or mutton) and milk. A sheep's wool is the most widely used animal fiber, and is usually harvested by shearing. In Commonwealth countries, ovine meat is called lamb when from younger animals and mutton when from older ones; in the United States, meat from both older and younger animals is usually called lamb. Sheep continue to be important for wool and meat today, and are also occasionally raised for pelts, as dairy animals, or as model organisms for science.
Sheep husbandry is practised throughout the majority of the inhabited world, and has been fundamental to many civilizations. In the modern era, Australia, New Zealand, the southern and central South American nations, and the British Isles are most closely associated with sheep production.
There is a large lexicon of unique terms for sheep husbandry which vary considerably by region and dialect. Use of the word sheep began in Middle English as a derivation of the Old English word scēap. A group of sheep is called a flock. Many other specific terms for the various life stages of sheep exist, generally related to lambing, shearing, and age.
Being a key animal in the history of farming, sheep have a deeply entrenched place in human culture, and are represented in much modern language and symbolism. As livestock, sheep are most often associated with pastoral, Arcadian imagery. Sheep figure in many mythologies—such as the Golden Fleece—and major religions, especially the Abrahamic traditions. In both ancient and modern religious ritual, sheep are used as sacrificial animals.
History
Main article: History of the domestic sheep
The exact line of descent from wild ancestors to domestic sheep is unclear. The most common hypothesis states that Ovis aries is descended from the Asiatic (O. gmelini) species of mouflon; the European mouflon (Ovis aries musimon) is a direct descendant of this population. Sheep were among the first animals to be domesticated by humankind (although the domestication of dogs probably took place 10 to 20 thousand years earlier); the domestication date is estimated to fall between 11,000 and 9000 B.C in Mesopotamia and possibly around 7000 BC in Mehrgarh in the Indus Valley. The rearing of sheep for secondary products, and the resulting breed development, began in either southwest Asia or western Europe. Initially, sheep were kept solely for meat, milk and skins. Archaeological evidence from statuary found at sites in Iran suggests that selection for woolly sheep may have begun around 6000 BC, and the earliest woven wool garments have been dated to two to three thousand years later.
Sheep husbandry spread quickly in Europe. Excavations show that in about 6000 BC, during the Neolithic period of prehistory, the Castelnovien people, living around Châteauneuf-les-Martigues near present-day Marseille in the south of France, were among the first in Europe to keep domestic sheep. Practically from its inception, ancient Greek civilization relied on sheep as primary livestock, and were even said to name individual animals. Ancient Romans kept sheep on a wide scale, and were an important agent in the spread of sheep raising. Pliny the Elder, in his Natural History (Naturalis Historia), speaks at length about sheep and wool. European colonists spread the practice to the New World from 1493 onwards.
Characteristics
Domestic sheep are relatively small ruminants, usually with a crimped hair called wool and often with horns forming a lateral spiral. They differ from their wild relatives and ancestors in several respects, having become uniquely neotenic as a result of selective breeding by humans. A few primitive breeds of sheep retain some of the characteristics of their wild cousins, such as short tails. Depending on breed, domestic sheep may have no horns at all (i.e. polled), or horns in both sexes, or in males only. Most horned breeds have a single pair, but a few breeds may have several.
Sheep in Turkmenistan
Another trait unique to domestic sheep as compared to wild ovines is their wide variation in color. Wild sheep are largely variations of brown hues, and variation within species is extremely limited. Colors of domestic sheep range from pure white to dark chocolate brown, and even spotted or piebald. Sheep keepers also sometimes artificially paint "smit marks" onto their sheep in any pattern or color for identification. Selection for easily dyeable white fleeces began early in sheep domestication, and as white wool is a dominant trait it spread quickly. However, colored sheep do appear in many modern breeds, and may even appear as a recessive trait in white flocks. While white wool is desirable for large commercial markets, there is a niche market for colored fleeces, mostly for handspinning. The nature of the fleece varies widely among the breeds, from dense and highly crimped, to long and hairlike. There is variation of wool type and quality even among members of the same flock, so wool classing is a step in the commercial processing of the fibre.
Suffolks are a medium wool, black-faced breed of meat sheep that make up 60% of the sheep population in the U.S.
Depending on breed, sheep show a range of heights and weights. Their rate of growth and mature weight is a heritable trait that is often selected for in breeding. Ewes typically weigh between 45 and 100 kilograms (100 and 220 lb), and rams between 45 and 160 kilograms (100 and 350 lb). When all deciduous teeth have erupted, the sheep has 20 teeth. Mature sheep have 32 teeth. As with other ruminants, the front teeth in the lower jaw bite against a hard, toothless pad in the upper jaw. These are used to pick off vegetation, then the rear teeth grind it before it is swallowed. There are eight lower front teeth in ruminants, but there is some disagreement as to whether these are eight incisors, or six incisors and two incisor-shaped canines. This means that the dental formula for sheep is either
0.0.3.3
4.0.3.3
or
0.0.3.3
3.1.3.3
There is a large diastema between the incisors and the molars.
In the first few years of life one can calculate the age of sheep from their front teeth, as a pair of milk teeth is replaced by larger adult teeth each year, the full set of eight adult front teeth being complete at about four years of age. The front teeth are then gradually lost as sheep age, making it harder for them to feed and hindering the health and productivity of the animal. For this reason, domestic sheep on normal pasture begin to slowly decline from four years on, and the life expectancy of a sheep is 10 to 12 years, though some sheep may live as long as 20 years.
Skull
Sheep have good hearing, and are sensitive to noise when being handled. Sheep have horizontal slit-shaped pupils, with excellent peripheral vision; with visual fields of about 270° to 320°, sheep can see behind themselves without turning their heads. Many breeds have only short hair on the face, and some have facial wool (if any) confined to the poll and or the area of the mandibular angle; the wide angles of peripheral vision apply to these breeds. A few breeds tend to have considerable wool on the face; for some individuals of these breeds, peripheral vision may be greatly reduced by "wool blindness", unless recently shorn about the face. Sheep have poor depth perception; shadows and dips in the ground may cause sheep to baulk. In general, sheep have a tendency to move out of the dark and into well-lit areas, and prefer to move uphill when disturbed. Sheep also have an excellent sense of smell, and, like all species of their genus, have scent glands just in front of the eyes, and interdigitally on the feet. The purpose of these glands is uncertain, but those on the face may be used in breeding behaviors. The foot glands might also be related to reproduction, but alternative functions, such as secretion of a waste product or a scent marker to help lost sheep find their flock, have also been proposed.
Comparison with goats
Sheep and goats are closely related: both are in the subfamily Caprinae. However, they are separate species, so hybrids rarely occur and are always infertile. A hybrid of a ewe and a buck (a male goat) is called a sheep-goat hybrid, known as geep. Visual differences between sheep and goats include the beard of goats and divided upper lip of sheep. Sheep tails also hang down, even when short or docked, while the short tails of goats are held upwards. Also, sheep breeds are often naturally polled (either in both sexes or just in the female), while naturally polled goats are rare (though many are polled artificially). Males of the two species differ in that buck goats acquire a unique and strong odor during the rut, whereas rams do not.
Breeds
The domestic sheep is a multi-purpose animal, and the more than 200 breeds now in existence were created to serve these diverse purposes. Some sources give a count of a thousand or more breeds, but these numbers cannot be verified, according to some sources. However, several hundred breeds of sheep have been identified by the Food and Agriculture Organization of the UN (FAO), with the estimated number varying somewhat from time to time: e.g. 863 breeds as of 1993, 1314 breeds as of 1995 and 1229 breeds as of 2006. (These numbers exclude extinct breeds, which are also tallied by the FAO.) For the purpose of such tallies, the FAO definition of a breed is "either a subspecific group of domestic livestock with definable and identifiable external characteristics that enable it to be separated by visual appraisal from other similarly defined groups within the same species or a group for which geographical and/or cultural separation from phenotypically similar groups has led to acceptance of its separate identity." Almost all sheep are classified as being best suited to furnishing a certain product: wool, meat, milk, hides, or a combination in a dual-purpose breed. Other features used when classifying sheep include face color (generally white or black), tail length, presence or lack of horns, and the topography for which the breed has been developed. This last point is especially stressed in the UK, where breeds are described as either upland (hill or mountain) or lowland breeds. A sheep may also be of a fat-tailed type, which is a dual-purpose sheep common in Africa and Asia with larger deposits of fat within and around its tail.
Breeds are often categorized by the type of their wool. Fine wool breeds are those that have wool of great crimp and density, which are preferred for textiles. Most of these were derived from Merino sheep, and the breed continues to dominate the world sheep industry. Downs breeds have wool between the extremes, and are typically fast-growing meat and ram breeds with dark faces. Some major medium wool breeds, such as the Corriedale, are dual-purpose crosses of long and fine-wooled breeds and were created for high-production commercial flocks. Long wool breeds are the largest of sheep, with long wool and a slow rate of growth. Long wool sheep are most valued for crossbreeding to improve the attributes of other sheep types. For example: the American Columbia breed was developed by crossing Lincoln rams (a long wool breed) with fine-wooled Rambouillet ewes.
Coarse or carpet wool sheep are those with a medium to long length wool of characteristic coarseness. Breeds traditionally used for carpet wool show great variability, but the chief requirement is a wool that will not break down under heavy use (as would that of the finer breeds). As the demand for carpet-quality wool declines, some breeders of this type of sheep are attempting to use a few of these traditional breeds for alternative purposes. Others have always been primarily meat-class sheep.
A minor class of sheep are the dairy breeds. Dual-purpose breeds that may primarily be meat or wool sheep are often used secondarily as milking animals, but there are a few breeds that are predominantly used for milking. These sheep produce a higher quantity of milk and have slightly longer lactation curves. In the quality of their milk, the fat and protein content percentages of dairy sheep vary from non-dairy breeds, but lactose content does not.
A last group of sheep breeds is that of fur or hair sheep, which do not grow wool at all. Hair sheep are similar to the early domesticated sheep kept before woolly breeds were developed, and are raised for meat and pelts. Some modern breeds of hair sheep, such as the Dorper, result from crosses between wool and hair breeds. For meat and hide producers, hair sheep are cheaper to keep, as they do not need shearing. Hair sheep are also more resistant to parasites and hot weather.
With the modern rise of corporate agribusiness and the decline of localized family farms, many breeds of sheep are in danger of extinction. The Rare Breeds Survival Trust of the UK lists 22 native breeds as having only 3,000 registered animals (each), and The Livestock Conservancy lists 14 as either "critical" or "threatened". Preferences for breeds with uniform characteristics and fast growth have pushed heritage (or heirloom) breeds to the margins of the sheep industry. Those that remain are maintained through the efforts of conservation organizations, breed registries, and individual farmers dedicated to their preservation.
Diet
Sheep are herbivorous mammals. Most breeds prefer to graze on grass and other short roughage, avoiding the taller woody parts of plants that goats readily consume. Both sheep and goats use their lips and tongues to select parts of the plant that are easier to digest or higher in nutrition. Sheep, however, graze well in monoculture pastures where most goats fare poorly.
Ruminant system of a sheep
Like all ruminants, sheep have a complex digestive system composed of four chambers, allowing them to break down cellulose from stems, leaves, and seed hulls into simpler carbohydrates. When sheep graze, vegetation is chewed into a mass called a bolus, which is then passed into the rumen, via the reticulum. The rumen is a 19- to 38-liter (5 to 10 gallon) organ in which feed is fermented. The fermenting organisms include bacteria, fungi, and protozoa. (Other important rumen organisms include some archaea, which produce methane from carbon dioxide.) The bolus is periodically regurgitated back to the mouth as cud for additional chewing and salivation. After fermentation in the rumen, feed passes into the reticulum and the omasum; special feeds such as grains may bypass the rumen altogether. After the first three chambers, food moves into the abomasum for final digestion before processing by the intestines. The abomasum is the only one of the four chambers analogous to the human stomach, and is sometimes called the "true stomach".
Other than forage, the other staple feed for sheep is hay, often during the winter months. The ability to thrive solely on pasture (even without hay) varies with breed, but all sheep can survive on this diet. Also included in some sheep's diets are minerals, either in a trace mix or in licks. Feed provided to sheep must be specially formulated, as most cattle, poultry, pig, and even some goat feeds contain levels of copper that are lethal to sheep. The same danger applies to mineral supplements such as salt licks.
Grazing behavior
Sheep follow a diurnal pattern of activity, feeding from dawn to dusk, stopping sporadically to rest and chew their cud. Ideal pasture for sheep is not lawnlike grass, but an array of grasses, legumes and forbs. Types of land where sheep are raised vary widely, from pastures that are seeded and improved intentionally to rough, native lands. Common plants toxic to sheep are present in most of the world, and include (but are not limited to) cherry, some oaks and acorns, tomato, yew, rhubarb, potato, and rhododendron.
Effects on pasture
Sheep are largely grazing herbivores, unlike browsing animals such as goats and deer that prefer taller foliage. With a much narrower face, sheep crop plants very close to the ground and can overgraze a pasture much faster than cattle. For this reason, many shepherds use managed intensive rotational grazing, where a flock is rotated through multiple pastures, giving plants time to recover. Paradoxically, sheep can both cause and solve the spread of invasive plant species. By disturbing the natural state of pasture, sheep and other livestock can pave the way for invasive plants. However, sheep also prefer to eat invasives such as cheatgrass, leafy spurge, kudzu and spotted knapweed over native species such as sagebrush, making grazing sheep effective for conservation grazing. Research conducted in Imperial County, California compared lamb grazing with herbicides for weed control in seedling alfalfa fields. Three trials demonstrated that grazing lambs were just as effective as herbicides in controlling winter weeds. Entomologists also compared grazing lambs to insecticides for insect control in winter alfalfa. In this trial, lambs provided insect control as effectively as insecticides.
Behavior
Sheep are flock animals and strongly gregarious; much sheep behavior can be understood on the basis of these tendencies. The dominance hierarchy of sheep and their natural inclination to follow a leader to new pastures were the pivotal factors in sheep being one of the first domesticated livestock species. Furthermore, in contrast to the red deer and gazelle (two other ungulates of primary importance to meat production in prehistoric times), sheep do not defend territories although they do form home ranges. All sheep have a tendency to congregate close to other members of a flock, although this behavior varies with breed, and sheep can become stressed when separated from their flock members. During flocking, sheep have a strong tendency to follow, and a leader may simply be the first individual to move. Relationships in flocks tend to be closest among related sheep: in mixed-breed flocks, subgroups of the same breed tend to form, and a ewe and her direct descendants often move as a unit within large flocks. Sheep can become hefted to one particular local pasture (heft) so they do not roam freely in unfenced landscapes. Lambs learn the heft from ewes and if whole flocks are culled it must be retaught to the replacement animals.
Flock behaviour in sheep is generally only exhibited in groups of four or more sheep; fewer sheep may not react as expected when alone or with few other sheep. Being a prey species, the primary defense mechanism of sheep is to flee from danger when their flight zone is entered. Cornered sheep may charge and butt, or threaten by hoof stamping and adopting an aggressive posture. This is particularly true for ewes with newborn lambs.
In regions where sheep have no natural predators, none of the native breeds of sheep exhibit a strong flocking behavior.
Herding
Farmers exploit flocking behavior to keep sheep together on unfenced pastures such as hill farming, and to move them more easily. For this purpose shepherds may use herding dogs in this effort, with a highly bred herding ability. Sheep are food-oriented, and association of humans with regular feeding often results in sheep soliciting people for food. Those who are moving sheep may exploit this behavior by leading sheep with buckets of feed.
Dominance hierarchy
Sheep establish a dominance hierarchy through fighting, threats and competitiveness. Dominant animals are inclined to be more aggressive with other sheep, and usually feed first at troughs. Primarily among rams, horn size is a factor in the flock hierarchy. Rams with different size horns may be less inclined to fight to establish the dominance order, while rams with similarly sized horns are more so. Merinos have an almost linear hierarchy whereas there is a less rigid structure in Border Leicesters when a competitive feeding situation arises.
In sheep, position in a moving flock is highly correlated with social dominance, but there is no definitive study to show consistent voluntary leadership by an individual sheep.
Intelligence and learning ability
Sheep are frequently thought of as unintelligent animals. Their flocking behavior and quickness to flee and panic can make shepherding a difficult endeavor for the uninitiated. Despite these perceptions, a University of Illinois monograph on sheep reported their intelligence to be just below that of pigs and on par with that of cattle. Sheep can recognize individual human and ovine faces and remember them for years; they can remember 50 other different sheep faces for over two years; they can recognize and are attracted to individual sheep and humans by their faces, as they possess similar specialized neural systems in the temporal and frontal lobes of their brains to humans and have a greater involvement of the right brain hemisphere. In addition to long-term facial recognition of individuals, sheep can also differentiate emotional states through facial characteristics.[68][69] If worked with patiently, sheep may learn their names, and many sheep are trained to be led by halter for showing and other purposes. Sheep have also responded well to clicker training. Sheep have been used as pack animals; Tibetan nomads distribute baggage equally throughout a flock as it is herded between living sites.
It has been reported that some sheep have apparently shown problem-solving abilities; a flock in West Yorkshire, England allegedly found a way to get over cattle grids by rolling on their backs, although documentation of this has relied on anecdotal accounts.
Vocalisations
Sounds made by domestic sheep include bleats, grunts, rumbles and snorts. Bleating ("baaing") is used mostly for contact communication, especially between dam and lambs, but also at times between other flock members. The bleats of individual sheep are distinctive, enabling the ewe and her lambs to recognize each other's vocalizations. Vocal communication between lambs and their dam declines to a very low level within several weeks after parturition. A variety of bleats may be heard, depending on sheep age and circumstances. Apart from contact communication, bleating may signal distress, frustration or impatience; however, sheep are usually silent when in pain. Isolation commonly prompts bleating by sheep. Pregnant ewes may grunt when in labor. Rumbling sounds are made by the ram during courting; somewhat similar rumbling sounds may be made by the ewe, especially when with her neonate lambs. A snort (explosive exhalation through the nostrils) may signal aggression or a warning, and is often elicited from startled sheep.
Lamb
In sheep breeds lacking facial wool, the visual field is wide. In 10 sheep (Cambridge, Lleyn and Welsh Mountain breeds, which lack facial wool), the visual field ranged from 298° to 325°, averaging 313.1°, with binocular overlap ranging from 44.5° to 74°, averaging 61.7°. In some breeds, unshorn facial wool can limit the visual field; in some individuals, this may be enough to cause "wool blindness". In 60 Merinos, visual fields ranged from 219.1° to 303.0°, averaging 269.9°, and the binocular field ranged from 8.9° to 77.7°, averaging 47.5°; 36% of the measurements were limited by wool, although photographs of the experiments indicate that only limited facial wool regrowth had occurred since shearing. In addition to facial wool (in some breeds), visual field limitations can include ears and (in some breeds) horns, so the visual field can be extended by tilting the head. Sheep eyes exhibit very low hyperopia and little astigmatism. Such visual characteristics are likely to produce a well-focused retinal image of objects in both the middle and long distance. Because sheep eyes have no accommodation, one might expect the image of very near objects to be blurred, but a rather clear near image could be provided by the tapetum and large retinal image of the sheep's eye, and adequate close vision may occur at muzzle length. Good depth perception, inferred from the sheep's sure-footedness, was confirmed in "visual cliff" experiments; behavioral responses indicating depth perception are seen in lambs at one day old. Sheep are thought to have colour vision, and can distinguish between a variety of colours: black, red, brown, green, yellow and white. Sight is a vital part of sheep communication, and when grazing, they maintain visual contact with each other. Each sheep lifts its head upwards to check the position of other sheep in the flock. This constant monitoring is probably what keeps the sheep in a flock as they move along grazing. Sheep become stressed when isolated; this stress is reduced if they are provided with a mirror, indicating that the sight of other sheep reduces stress.
Taste is the most important sense in sheep, establishing forage preferences, with sweet and sour plants being preferred and bitter plants being more commonly rejected. Touch and sight are also important in relation to specific plant characteristics, such as succulence and growth form.
The ram uses his vomeronasal organ (sometimes called the Jacobson's organ) to sense the pheromones of ewes and detect when they are in estrus. The ewe uses her vomeronasal organ for early recognition of her neonate lamb.
Reproduction
Sheep follow a similar reproductive strategy to other herd animals. A group of ewes is generally mated by a single ram, who has either been chosen by a breeder or (in feral populations) has established dominance through physical contest with other rams. Most sheep are seasonal breeders, although some are able to breed year-round. Ewes generally reach sexual maturity at six to eight months old, and rams generally at four to six months. However, there are exceptions. For example, Finnsheep ewe lambs may reach puberty as early as 3 to 4 months, and Merino ewes sometimes reach puberty at 18 to 20 months. Ewes have estrus cycles about every 17 days, during which they emit a scent and indicate readiness through physical displays towards rams.
In feral sheep, rams may fight during the rut to determine which individuals may mate with ewes. Rams, especially unfamiliar ones, will also fight outside the breeding period to establish dominance; rams can kill one another if allowed to mix freely. During the rut, even usually friendly rams may become aggressive towards humans due to increases in their hormone levels.
After mating, sheep have a gestation period of about five months, and normal labor takes one to three hours. Although some breeds regularly throw larger litters of lambs, most produce single or twin lambs. During or soon after labor, ewes and lambs may be confined to small lambing jugs, small pens designed to aid both careful observation of ewes and to cement the bond between them and their lambs.
A lamb's first steps
Ovine obstetrics can be problematic. By selectively breeding ewes that produce multiple offspring with higher birth weights for generations, sheep producers have inadvertently caused some domestic sheep to have difficulty lambing; balancing ease of lambing with high productivity is one of the dilemmas of sheep breeding. In the case of any such problems, those present at lambing may assist the ewe by extracting or repositioning lambs. After the birth, ewes ideally break the amniotic sac (if it is not broken during labor), and begin licking clean the lamb. Most lambs will begin standing within an hour of birth. In normal situations, lambs nurse after standing, receiving vital colostrum milk. Lambs that either fail to nurse or are rejected by the ewe require help to survive, such as bottle-feeding or fostering by another ewe.
Most lambs begin life being born outdoors. After lambs are several weeks old, lamb marking (ear tagging, docking, mulesing, and castrating) is carried out. Vaccinations are usually carried out at this point as well. Ear tags with numbers are attached, or ear marks are applied, for ease of later identification of sheep. Docking and castration are commonly done after 24 hours (to avoid interference with maternal bonding and consumption of colostrum) and are often done not later than one week after birth, to minimize pain, stress, recovery time and complications. The first course of vaccinations (commonly anti-clostridial) is commonly given at an age of about 10 to 12 weeks; i.e. when the concentration of maternal antibodies passively acquired via colostrum is expected to have fallen low enough to permit development of active immunity. Ewes are often revaccinated annually about 3 weeks before lambing, to provide high antibody concentrations in colostrum during the first several hours after lambing. Ram lambs that will either be slaughtered or separated from ewes before sexual maturity are not usually castrated. Objections to all these procedures have been raised by animal rights groups, but farmers defend them by saying they save money, and inflict only temporary pain.
Homosexuality
Sheep are the only species of mammal except for humans which exhibits exclusive homosexual behavior. About 10% of rams refuse to mate with ewes but readily mate with other rams, and thirty percent of all rams demonstrate at least some homosexual behavior. Additionally, a small number of females that were accompanied by a male fetus in utero (i.e. as fraternal twins) are freemartins (female animals that are behaviorally masculine and lack functioning ovaries).
Health
Sheep may fall victim to poisons, infectious diseases, and physical injuries. As a prey species, a sheep's system is adapted to hide the obvious signs of illness, to prevent being targeted by predators. However, some signs of ill health are obvious, with sick sheep eating little, vocalizing excessively, and being generally listless. Throughout history, much of the money and labor of sheep husbandry has aimed to prevent sheep ailments. Historically, shepherds often created remedies by experimentation on the farm. In some developed countries, including the United States, sheep lack the economic importance for drug companies to perform expensive clinical trials required to approve more than a relatively limited number of drugs for ovine use. However, extra-label drug use in sheep production is permitted in many jurisdictions, subject to certain restrictions. In the US, for example, regulations governing extra-label drug use in animals are found in 21 CFR (Code of Federal Regulations) Part 530. In the 20th and 21st centuries, a minority of sheep owners have turned to alternative treatments such as homeopathy, herbalism and even traditional Chinese medicine to treat sheep veterinary problems. Despite some favorable anecdotal evidence, the effectiveness of alternative veterinary medicine has been met with skepticism in scientific journals. The need for traditional anti-parasite drugs and antibiotics is widespread, and is the main impediment to certified organic farming with sheep.
Many breeders take a variety of preventive measures to ward off problems. The first is to ensure all sheep are healthy when purchased. Many buyers avoid outlets known to be clearing houses for animals culled from healthy flocks as either sick or simply inferior. This can also mean maintaining a closed flock, and quarantining new sheep for a month. Two fundamental preventive programs are maintaining good nutrition and reducing stress in the sheep. Restraint, isolation, loud noises, novel situations, pain, heat, extreme cold, fatigue and other stressors can lead to secretion of cortisol, a stress hormone, in amounts that may indicate welfare problems. Excessive stress can compromise the immune system. "Shipping fever" (pneumonic mannheimiosis, formerly called pasteurellosis) is a disease of particular concern, that can occur as a result of stress, notably during transport and (or) handling. Pain, fear and several other stressors can cause secretion of epinephrine (adrenaline). Considerable epinephrine secretion in the final days before slaughter can adversely affect meat quality (by causing glycogenolysis, removing the substrate for normal post-slaughter acidification of meat) and result in meat becoming more susceptible to colonization by spoilage bacteria. Because of such issues, low-stress handling is essential in sheep management. Avoiding poisoning is also important; common poisons are pesticide sprays, inorganic fertilizer, motor oil, as well as radiator coolant containing ethylene glycol.
Common forms of preventive medication for sheep are vaccinations and treatments for parasites. Both external and internal parasites are the most prevalent malady in sheep, and are either fatal, or reduce the productivity of flocks. Worms are the most common internal parasites. They are ingested during grazing, incubate within the sheep, and are expelled through the digestive system (beginning the cycle again). Oral anti-parasitic medicines, known as drenches, are given to a flock to treat worms, sometimes after worm eggs in the feces has been counted to assess infestation levels. Afterwards, sheep may be moved to a new pasture to avoid ingesting the same parasites. External sheep parasites include: lice (for different parts of the body), sheep keds, nose bots, sheep itch mites, and maggots. Keds are blood-sucking parasites that cause general malnutrition and decreased productivity, but are not fatal. Maggots are those of the bot fly and the blow-fly, commonly Lucilia sericata or its relative L. cuprina. Fly maggots cause the extremely destructive condition of flystrike. Flies lay their eggs in wounds or wet, manure-soiled wool; when the maggots hatch they burrow into a sheep's flesh, eventually causing death if untreated. In addition to other treatments, crutching (shearing wool from a sheep's rump) is a common preventive method. Some countries allow mulesing, a practice that involves stripping away the skin on the rump to prevent fly-strike, normally performed when the sheep is a lamb. Nose bots are fly larvae that inhabit a sheep's sinuses, causing breathing difficulties and discomfort. Common signs are a discharge from the nasal passage, sneezing, and frantic movement such as head shaking. External parasites may be controlled through the use of backliners, sprays or immersive sheep dips.
A wide array of bacterial and viral diseases affect sheep. Diseases of the hoof, such as foot rot and foot scald may occur, and are treated with footbaths and other remedies. Foot rot is present in over 97% of flocks in the UK. These painful conditions cause lameness and hinder feeding. Ovine Johne's disease is a wasting disease that affects young sheep. Bluetongue disease is an insect-borne illness causing fever and inflammation of the mucous membranes. Ovine rinderpest (or peste des petits ruminants) is a highly contagious and often fatal viral disease affecting sheep and goats. Sheep may also be affected by primary or secondary photosensitization. Tetanus can also afflict sheep through wounds from shearing, docking, castration, or vaccination. The organism also can be introduced into the reproductive tract by unsanitary humans who assist ewes during lambing.
A few sheep conditions are transmissible to humans. Orf (also known as scabby mouth, contagious ecthyma or soremouth) is a skin disease leaving lesions that is transmitted through skin-to-skin contact. Cutaneous anthrax is also called woolsorter's disease, as the spores can be transmitted in unwashed wool. More seriously, the organisms that can cause spontaneous enzootic abortion in sheep are easily transmitted to pregnant women. Also of concern are the prion disease scrapie and the virus that causes foot-and-mouth disease (FMD), as both can devastate flocks. The latter poses a slight risk to humans. During the 2001 FMD pandemic in the UK, hundreds of sheep were culled and some rare British breeds were at risk of extinction due to this.
Of the 600,300 sheep lost to the US economy in 2004, 37.3% were lost to predators, while 26.5% were lost to some form of disease. Poisoning accounted for 1.7% of non-productive deaths.
Predators
A lamb being attacked by coyotes with a bite to the throat
Other than parasites and disease, predation is a threat to sheep and the profitability of sheep raising. Sheep have little ability to defend themselves, compared with other species kept as livestock. Even if sheep survive an attack, they may die from their injuries or simply from panic. However, the impact of predation varies dramatically with region. In Africa, Australia, the Americas, and parts of Europe and Asia predators are a serious problem. In the United States, for instance, over one third of sheep deaths in 2004 were caused by predation. In contrast, other nations are virtually devoid of sheep predators, particularly islands known for extensive sheep husbandry. Worldwide, canids—including the domestic dog—are responsible for most sheep deaths. Other animals that occasionally prey on sheep include: felines, bears, birds of prey, ravens and feral hogs.
Sheep producers have used a wide variety of measures to combat predation. Pre-modern shepherds used their own presence, livestock guardian dogs, and protective structures such as barns and fencing. Fencing (both regular and electric), penning sheep at night and lambing indoors all continue to be widely used. More modern shepherds used guns, traps, and poisons to kill predators, causing significant decreases in predator populations. In the wake of the environmental and conservation movements, the use of these methods now usually falls under the purview of specially designated government agencies in most developed countries.
The 1970s saw a resurgence in the use of livestock guardian dogs and the development of new methods of predator control by sheep producers, many of them non-lethal. Donkeys and guard llamas have been used since the 1980s in sheep operations, using the same basic principle as livestock guardian dogs. Interspecific pasturing, usually with larger livestock such as cattle or horses, may help to deter predators, even if such species do not actively guard sheep. In addition to animal guardians, contemporary sheep operations may use non-lethal predator deterrents such as motion-activated lights and noisy alarms.
Economic importance
Main article: Agricultural economics
Global sheep stock
in 2019
Number in millions
1. China163.5 (13.19%)
2. India74.3 (5.99%)
3. Australia65.8 (5.31%)
4. Nigeria46.9 (3.78%)
5. Iran41.3 (3.33%)
6. Sudan40.9 (3.3%)
7. Chad35.9 (2.9%)
8. Turkey35.2 (2.84%)
9. United Kingdom33.6 (2.71%)
10. Mongolia32.3 (2.61%)
World total1,239.8
Source: UN Food and Agriculture Organization
Sheep are an important part of the global agricultural economy. However, their once vital status has been largely replaced by other livestock species, especially the pig, chicken, and cow. China, Australia, India, and Iran have the largest modern flocks, and serve both local and exportation needs for wool and mutton. Other countries such as New Zealand have smaller flocks but retain a large international economic impact due to their export of sheep products. Sheep also play a major role in many local economies, which may be niche markets focused on organic or sustainable agriculture and local food customers. Especially in developing countries, such flocks may be a part of subsistence agriculture rather than a system of trade. Sheep themselves may be a medium of trade in barter economies.
Domestic sheep provide a wide array of raw materials. Wool was one of the first textiles, although in the late 20th century wool prices began to fall dramatically as the result of the popularity and cheap prices for synthetic fabrics. For many sheep owners, the cost of shearing is greater than the possible profit from the fleece, making subsisting on wool production alone practically impossible without farm subsidies. Fleeces are used as material in making alternative products such as wool insulation. In the 21st century, the sale of meat is the most profitable enterprise in the sheep industry, even though far less sheep meat is consumed than chicken, pork or beef.
Sheepskin is likewise used for making clothes, footwear, rugs, and other products. Byproducts from the slaughter of sheep are also of value: sheep tallow can be used in candle and soap making, sheep bone and cartilage has been used to furnish carved items such as dice and buttons as well as rendered glue and gelatin. Sheep intestine can be formed into sausage casings, and lamb intestine has been formed into surgical sutures, as well as strings for musical instruments and tennis rackets. Sheep droppings, which are high in cellulose, have even been sterilized and mixed with traditional pulp materials to make paper. Of all sheep byproducts, perhaps the most valuable is lanolin: the waterproof, fatty substance found naturally in sheep's wool and used as a base for innumerable cosmetics and other products.
Some farmers who keep sheep also make a profit from live sheep. Providing lambs for youth programs such as 4-H and competition at agricultural shows is often a dependable avenue for the sale of sheep. Farmers may also choose to focus on a particular breed of sheep in order to sell registered purebred animals, as well as provide a ram rental service for breeding. A new option for deriving profit from live sheep is the rental of flocks for grazing; these "mowing services" are hired in order to keep unwanted vegetation down in public spaces and to lessen fire hazard.
Despite the falling demand and price for sheep products in many markets, sheep have distinct economic advantages when compared with other livestock. They do not require expensive housing, such as that used in the intensive farming of chickens or pigs. They are an efficient use of land; roughly six sheep can be kept on the amount that would suffice for a single cow or horse. Sheep can also consume plants, such as noxious weeds, that most other animals will not touch, and produce more young at a faster rate. Also, in contrast to most livestock species, the cost of raising sheep is not necessarily tied to the price of feed crops such as grain, soybeans and corn. Combined with the lower cost of quality sheep, all these factors combine to equal a lower overhead for sheep producers, thus entailing a higher profitability potential for the small farmer. Sheep are especially beneficial for independent producers, including family farms with limited resources, as the sheep industry is one of the few types of animal agriculture that has not been vertically integrated by agribusiness. However, small flocks, from 10 to 50 ewes, often are not profitable because they tend to be poorly managed. The primary reason is that mechanization is not feasible, so return per hour of labor is not maximized. Small farm flocks generally are used simply to control weeds on irrigation ditches or maintained as a hobby.
Shoulder of lamb
Sheep meat and milk were one of the earliest staple proteins consumed by human civilization after the transition from hunting and gathering to agriculture. Sheep meat prepared for food is known as either mutton or lamb, and approximately 540 million sheep are slaughtered each year for meat worldwide. "Mutton" is derived from the Old French moton, which was the word for sheep used by the Anglo-Norman rulers of much of the British Isles in the Middle Ages. This became the name for sheep meat in English, while the Old English word sceap was kept for the live animal. Throughout modern history, "mutton" has been limited to the meat of mature sheep usually at least two years of age; "lamb" is used for that of immature sheep less than a year.
In the 21st century, the nations with the highest consumption of sheep meat are the Arab states of the Persian Gulf, New Zealand, Australia, Greece, Uruguay, the United Kingdom and Ireland. These countries eat 14–40 lbs (3–18 kg) of sheep meat per capita, per annum. Sheep meat is also popular in France, Africa (especially the Arab world), the Caribbean, the rest of the Middle East, India, and parts of China. This often reflects a history of sheep production. In these countries in particular, dishes comprising alternative cuts and offal may be popular or traditional. Sheep testicles—called animelles or lamb fries—are considered a delicacy in many parts of the world. Perhaps the most unusual dish of sheep meat is the Scottish haggis, composed of various sheep innards cooked along with oatmeal and chopped onions inside its stomach. In comparison, countries such as the U.S. consume only a pound or less (under 0.5 kg), with Americans eating 50 pounds (22 kg) of pork and 65 pounds (29 kg) of beef. In addition, such countries rarely eat mutton, and may favor the more expensive cuts of lamb: mostly lamb chops and leg of lamb.
Though sheep's milk may be drunk rarely in fresh form, today it is used predominantly in cheese and yogurt making. Sheep have only two teats, and produce a far smaller volume of milk than cows. However, as sheep's milk contains far more fat, solids, and minerals than cow's milk, it is ideal for the cheese-making process. It also resists contamination during cooling better because of its much higher calcium content. Well-known cheeses made from sheep milk include the feta of Bulgaria and Greece, Roquefort of France, Manchego from Spain, the pecorino romano (the Italian word for "sheep" is pecore) and ricotta of Italy. Yogurts, especially some forms of strained yogurt, may also be made from sheep milk. Many of these products are now often made with cow's milk, especially when produced outside their country of origin. Sheep milk contains 4.8% lactose, which may affect those who are intolerant.
As with other domestic animals, the meat of uncastrated males is inferior in quality, especially as they grow. A "bucky" lamb is a lamb which was not castrated early enough, or which was castrated improperly (resulting in one testicle being retained). These lambs are worth less at market.
In science
Sheep are generally too large and reproduce too slowly to make ideal research subjects, and thus are not a common model organism. They have, however, played an influential role in some fields of science. In particular, the Roslin Institute of Edinburgh, Scotland used sheep for genetics research that produced groundbreaking results. In 1995, two ewes named Megan and Morag were the first mammals cloned from differentiated cells, also referred to as gynomerogony. A year later, a Finnish Dorset sheep named Dolly, dubbed "the world's most famous sheep" in Scientific American, was the first mammal to be cloned from an adult somatic cell. Following this, Polly and Molly were the first mammals to be simultaneously cloned and transgenic.
As of 2008, the sheep genome has not been fully sequenced, although a detailed genetic map has been published, and a draft version of the complete genome produced by assembling sheep DNA sequences using information given by the genomes of other mammals. In 2012, a transgenic sheep named "Peng Peng" was cloned by Chinese scientists, who spliced his genes with that of a roundworm (C. elegans) in order to increase production of fats healthier for human consumption.
In the study of natural selection, the population of Soay sheep that remain on the island of Hirta have been used to explore the relation of body size and coloration to reproductive success. Soay sheep come in several colors, and researchers investigated why the larger, darker sheep were in decline; this occurrence contradicted the rule of thumb that larger members of a population tend to be more successful reproductively. The feral Soays on Hirta are especially useful subjects because they are isolated.
Domestic sheep are sometimes used in medical research, particularly for researching cardiovascular physiology, in areas such as hypertension and heart failure. Pregnant sheep are also a useful model for human pregnancy, and have been used to investigate the effects on fetal development of malnutrition and hypoxia. In behavioral sciences, sheep have been used in isolated cases for the study of facial recognition, as their mental process of recognition is qualitatively similar to humans.
Cultural impact
Sheep have had a strong presence in many cultures, especially in areas where they form the most common type of livestock. In the English language, to call someone a sheep or ovine may allude that they are timid and easily led. In contradiction to this image, male sheep are often used as symbols of virility and power; the logos of the Los Angeles Rams football team and the Dodge Ram pickup truck allude to males of the bighorn sheep, Ovis canadensis.
Counting sheep is popularly said to be an aid to sleep, and some ancient systems of counting sheep persist today. Sheep also enter in colloquial sayings and idiom frequently with such phrases as "black sheep". To call an individual a black sheep implies that they are an odd or disreputable member of a group. This usage derives from the recessive trait that causes an occasional black lamb to be born into an entirely white flock. These black sheep were considered undesirable by shepherds, as black wool is not as commercially viable as white wool. Citizens who accept overbearing governments have been referred to by the Portmanteau neologism of sheeple. Somewhat differently, the adjective "sheepish" is also used to describe embarrassment.
In heraldry
In British heraldry, sheep appear in the form of rams, sheep proper and lambs. These are distinguished by the ram being depicted with horns and a tail, the sheep with neither and the lamb with its tail only. A further variant of the lamb, termed the Paschal lamb, is depicted as carrying a Christian cross and with a halo over its head. Rams' heads, portrayed without a neck and facing the viewer, are also found in British armories. The fleece, depicted as an entire sheepskin carried by a ring around its midsection, originally became known through its use in the arms of the Order of the Golden Fleece and was later adopted by towns and individuals with connections to the wool industry. A sheep on a blue field is depicted on the greater/royal arms of the king of Denmark to represent the Faroe Islands. In 2004 a modernized arms has been adopted by the Faroe Islands, which based on a 15th century coat of arms.
Religion and folklore
In antiquity, symbolism involving sheep cropped up in religions in the ancient Near East, the Mideast, and the Mediterranean area: Çatalhöyük, ancient Egyptian religion, the Cana'anite and Phoenician tradition, Judaism, Greek religion, and others. Religious symbolism and ritual involving sheep began with some of the first known faiths: Skulls of rams (along with bulls) occupied central placement in shrines at the Çatalhöyük settlement in 8,000 BCE. In Ancient Egyptian religion, the ram was the symbol of several gods: Khnum, Heryshaf and Amun (in his incarnation as a god of fertility). Other deities occasionally shown with ram features include the goddess Ishtar, the Phoenician god Baal-Hamon, and the Babylonian god Ea-Oannes. In Madagascar, sheep were not eaten as they were believed to be incarnations of the souls of ancestors.
There are many ancient Greek references to sheep: that of Chrysomallos, the golden-fleeced ram, continuing to be told through into the modern era. Astrologically, Aries, the ram, is the first sign of the classical Greek zodiac, and the sheep is the eighth of the twelve animals associated with the 12-year cycle of in the Chinese zodiac, related to the Chinese calendar. It is said in Chinese traditions that Hou ji sacrificed sheep. Mongolia, shagai are an ancient form of dice made from the cuboid bones of sheep that are often used for fortunetelling purposes.
Sheep play an important role in all the Abrahamic faiths; Abraham, Isaac, Jacob, Moses, and King David were all shepherds. According to the Biblical story of the Binding of Isaac, a ram is sacrificed as a substitute for Isaac after an angel stays Abraham's hand (in the Islamic tradition, Abraham was about to sacrifice Ishmael). Eid al-Adha is a major annual festival in Islam in which sheep (or other animals) are sacrificed in remembrance of this act. Sheep are occasionally sacrificed to commemorate important secular events in Islamic cultures. Greeks and Romans sacrificed sheep regularly in religious practice, and Judaism once sacrificed sheep as a Korban (sacrifice), such as the Passover lamb. Ovine symbols—such as the ceremonial blowing of a shofar—still find a presence in modern Judaic traditions.
Collectively, followers of Christianity are often referred to as a flock, with Christ as the Good Shepherd, and sheep are an element in the Christian iconography of the birth of Jesus. Some Christian saints are considered patrons of shepherds, and even of sheep themselves. Christ is also portrayed as the Sacrificial lamb of God (Agnus Dei) and Easter celebrations in Greece and Romania traditionally feature a meal of Paschal lamb. A church leader is often called the pastor, which is derived from the Latin word for shepherd. In many western Christian traditions bishops carry a staff, which also serves as a symbol of the episcopal office, known as a crosier, which is modeled on the shepherd's crook.
Sheep are key symbols in fables and nursery rhymes like The Wolf in Sheep's Clothing, Little Bo Peep, Baa, Baa, Black Sheep, and Mary Had a Little Lamb; novels such as George Orwell's Animal Farm and Haruki Murakami's A Wild Sheep Chase; songs such as Bach's Sheep may safely graze (Schafe können sicher weiden) and Pink Floyd's "Sheep", and poems like William Blake's "The Lamb".
Today, July 5, 2017, is the grand opening of Land Of Lemurs at the Calgary Zoo.
• Lemurs are prosimians, meaning they are “more primitive”
and more like the ancestor to all primates.
• This group also contains galagos, aye-ayes, pottos, lorises
and bushbabies.
• Prosimians have:
• claws instead of fingernails
• long, bony snouts
• little teeth differentiation, except for dental comb made
of lower front incisors that are used for grooming
• incomplete eye orbitals
• smaller braincases than monkeys or apes
• shorter forelimbs than hindlimbs
• most prosimians are nocturnal, but since there are no
monkey species on Madagascar to compete
with, lemurs are also diurnal
• many of the well-known lemur species (ie: ring-tailed,
ruffed, etc.) are diurnal, but some species
have retained their nocturnal lifestyle
• Lemurs are also the only prosimians to live in social
groups (this does not apply to all species)
• Lemurs are found on Madagascar and the neighboring
ComorosIslands (where they were introduced), and
are found nowhere else on earth.
• In 2016 Madagascar is home to 104 types of lemurs but new
species are often found.
• Lemurs are an important conservation focus for their ability
to regenerate forest habitat through seed dispersal
Here's "Vince" enjoying the remaining portion of the watermelon he was sharing with "Mulan". His lower incisors are working like a melon scoop, gouging out chunks of melon. He gradually worked his way completely around the edge, leaving only some melon in the deepest section in the center... a treat for the younger bucks who were waiting for him to leave. He somehow got a bit stuck on the tip of one brow tine.
100-6956
(Between a rock and a hard place)
Young Rock Hyrax (Dassie) trying to hide from the "Big Eye".
Best viewed full screen.
The rock hyrax (Procavia capensis), also called rock badger, rock rabbit, and Cape hyrax, is commonly referred to in South African English as the dassie. It is one of the four living species of the order Hyracoidea, and the only living species in the genus Procavia. Like all hyraxes, it is a medium-sized (~4 kg) terrestrial mammal, with short ears and tail.
The closest living relatives to hyraxes are the modern-day elephants and sirenians. The rock hyrax is found across Africa and the Middle East in habitats with rock crevices into which it escapes from predators. It is the only extant terrestrial afrotherian in the Middle East. Hyraxes typically live in groups of 10–80 animals, and forage as a group. They have been reported to use sentries: one or more animals take up position on a vantage point and issue alarm calls on the approach of predators.
The rock hyrax has incomplete thermoregulation and is most active in the morning and evening, although its activity pattern varies substantially with season and climate.
Over most of its range, the rock hyrax is not endangered, and in some areas is considered a minor pest. In Ethiopia, Israel and Jordan, it is a reservoir of the leishmaniasis parasite.
The rock hyrax is squat and heavily built, adults reaching a length of 50 cm (20 in) and weighing around 4 kg (8.8 lb), with a slight sexual dimorphism, males being approximately 10% heavier than females. Their fur is thick and grey-brown, although this varies strongly between different environments: from dark brown in wetter habitats, to light grey in desert living individuals. Hyrax size (as measured by skull length and humerus diameter) is correlated to precipitation, probably because of the effect on preferred hyrax forage.
Prominent in, and apparently unique to hyraxes, is the dorsal gland, which excretes an odour used for social communication and territorial marking. The gland is most clearly visible in dominant males.
The head of the rock hyrax is pointed, having a short neck with rounded ears. They have long black whiskers on their muzzles. The rock hyrax has a prominent pair of long, pointed tusk-like upper incisors which are reminiscent of the elephant, to which the hyrax is distantly related. The forefeet are plantigrade, and the hind feet semi-digitigrade. The soles of the feet have large, soft pads that are kept moist with sweat-like secretions. In males, the testes are permanently abdominal, another anatomical feature that hyraxes share with their relatives elephants and sirenians.
Thermoregulation in the rock hyrax has been subject to much research, as their body temperature varies with a diurnal rhythm. However, animals kept in constant environmental conditions also display such variation and this internal mechanism may be related to water balance regulation.
The rock hyrax occurs across sub-Saharan Africa, with the exception of the Congo basin and Madagascar. A larger, longer-haired subspecies is abundant in the glacial moraines in the alpine zone of Mount Kenya. The distribution continues into southern Algeria, Libya and Egypt, and the Middle East, with populations in Israel, Jordan, Lebanon and the Arabian peninsula.
A mammal of similar appearance by convergent evolution, but unrelated, is the rock cavy of Brazil.
Rock hyraxes build dwelling holes in any type of rock with suitable cavities such as sedimentary rocks and soil. In Mount Kenya, rock hyraxes live in colonies comprising an adult male, differing numbers of adult females and immatures. They are active during the day, and sometimes during moonlit nights. The dominant male defends and watches over the group. The male also marks its territory.
In Africa, hyraxes are preyed on by leopards, Egyptian cobras, puff adders, rock pythons, caracals, wild dogs, hawks, and owls. Verreaux's eagle in particular is a specialist hunter of hyrax. In Israel, the rock hyrax is reportedly rarely preyed upon by terrestrial predators, as their system of sentries and their reliable refuges provide considerable protection. Hyrax remains are almost absent from the droppings of wolves in the Judean Desert.
Hyraxes feed on a wide variety of different plants, including Lobelia and broad-leafed plants. They also have been reported to eat insects and grubs. The rock hyraxes forage for food up to about 50 metres from their refuge, usually feeding as a group and with one or more acting as sentries from a prominent lookout position. On the approach of danger, the sentries give an alarm call, and the animals quickly retreat to their refuge.
They are able to go for many days without water due to the moisture they obtain through their food, but will quickly dehydrate under direct sunlight. Despite their seemingly clumsy build, they are able to climb trees (although not as readily as Heterohyrax), and will readily enter residential gardens to feed on the leaves of citrus and other trees.
The rock hyrax also makes a loud grunting sound while moving its jaws as if chewing, and this behaviour may be a sign of aggression. Some authors have proposed that observation of this behaviour by ancient Israelites gave rise to the misconception given in Leviticus 11:4-8 that the hyrax chews the cud In fact, hyraxes are not ruminants.
Rock hyraxes give birth to between two and four young after a gestation period of 6–7 months (long, for their size). The young are well developed at birth with fully opened eyes and complete pelage. Young can ingest solid food after two weeks and are weaned at ten weeks. After 16 months, the rock hyraxes become sexually mature, they reach adult size at three years, and they typically live about ten years. During seasonal changes, the weight of the male reproductive organs (testis, seminal vesicles) changes due to sexual activity. A study showed that between May and January, the males were inactive sexually. From February onward, there was a dramatic increase to the weight of these organs, and the males are able to copulate.
Social behaviour
In a study of their social networks, it was found that hyraxes that live in more "egalitarian" groups, in which social associations are spread more evenly among group members, survive longer. In addition, hyraxes are the first non-human species in which structural balance was described. They follow "the friend of my friend is my friend" rule, and avoid unbalanced social configurations.
Captive rock hyraxes make more than 20 different noises and vocal signals. The most familiar signal is a high trill, given in response to perceived danger. Rock hyrax calls can provide important biological information such as size, age, social status, body weight, condition, and hormonal state of the caller, as determined by measuring their call length, patterns, complexity, and frequency. More recently, researchers have found rich syntactic structure and geographical variations in the calls of rock hyraxes, a first in the vocalization of mammalian taxa other than primates, cetaceans, and bats. Higher ranked males tend to sing more often, although the energetic cost of singing is relatively low.
The rock hyrax spends approximately 95% of its time resting. During this time, they can often be seen basking in the sun, which is thought to be an element of their complex thermoregulation.
Dispersal
Male hyraxes have been categorised into four classes: territorial, peripheral, early dispersers, and late dispersers. The territorial males are dominant. Peripheral males are more solitary and sometimes take over a group when the dominant male is missing. Early-dispersing males are juveniles that leave the birth site around 16 to 24 months of age. Late dispersers are also juvenile males, but they leave the birth site much later; around 30 or more months of age.
Names
They are known as dassies in South Africa, and sometimes rock rabbits. The Swahili names for them are pimbi, pelele and wibari, though the latter two names are nowadays reserved for the tree hyraxes. This species has many subspecies, many of which are also known as rock or Cape hyrax, although the former usually refers to African varieties.
In Arabic, the rock hyrax is called "wabr" or "tabsoun". In Hebrew, the rock hyrax is called "shafan sela", meaning rock "shafan", where the meaning of shafan is obscure, but is colloquially used as a synonym for rabbit in modern Hebrew. According to Gerald Durrell local people in Bafut, Cameroon, call the rock hyrax the n'eer.
Naturopathic use
Rock hyraxes produce large quantities of hyraceum—a sticky mass of dung and urine that has been employed as a South African folk remedy in the treatment of several medical disorders, including epilepsy and convulsions. Hyraceum is now being used by perfumers who tincture it in alcohol to yield a natural animal musk.
In culture
The rock hyrax is classified as non-kosher in the Jewish Torah. Nonetheless, it is also included in Proverbs 30:26 as one of a number of remarkable animals for being small, but exceedingly wise, in this case because "the rock badgers are a people not mighty, yet they make their homes in the cliffs".
In Joy Adamson's books and the movie Born Free, a rock hyrax called Pati-Pati was her companion for six years before Elsa and her siblings came along; Pati-Pati took the role of nanny and watched over them with great care.
The 2013 animated film Khumba features a number of rock hyraxes who sacrifice one of their own to a white black eagle.
The species was introduced to Jebel Hafeet, which is on the border of Oman and the United Arab Emirates.
Dikhololo
Rural Area
Brits
South Africa
I wasn’t sure when I pulled out the whisky-loaded bonbon. Half-popped caramel corn glared a challenge at my teeth. But I bit in gingerly with my incisors and the salty sweet tickled my tongue. The whisky ganache underneath smoooothed its way down. And the macadamia nuts gave in softly. The milk chocolate enrobing it all, with its wavy stripe of dark, finished the heavenly bite.
The common marmoset (Callithrix Jacchus) is classed as a New World monkey. It originally lived on the North-eastern coast of Brazil; however release of captive individuals has expanded its range to include Southeast Brazil. They are very small monkeys with relatively long tails. Males have an average height of 188 mm (7.40 in) and females have an average height of 185 mm (7.28 in). Males weigh 256 g (9.03 oz) on average and females weigh 236 g (8.32 oz) on average.
The common marmosets have claw-like nails on most of their fingers, with only their big toes having the flat nails that most other primates have. These nails, the shape of their incisors, and their specialist gut help their unique diet of gum, sap, latex and resins. They use their nails to cling to the side of a tree and, with their long lower incisors, chew a hole in the tree bark. They will then lick up the exudates.
Above information taken from Wikipedia.
You can see this typical feeding behaviour, as well as see them catching and eating insects and the occasional small bird, when visiting these marmosets at Longleat. They also enjoy the meal worms and sweet corn that the handlers use as treats to tempt them out into the viewing enclosure.
Picherali, Pompeo. - Architetto, pittore e incisore (Siracusa 1670 - ivi 1743), autore, tra l'altro, della Chiesa dello Spirito Santo (1727), del portico esterno della chiesa di S. Lucia e delle decorazioni della chiesa della Madonna del Carmelo, tutte situate a Siracusa.
Fish, any of approximately 34,000 species of vertebrate animals (phylum Chordata) found in the fresh and salt waters of the world. Living species range from the primitive jawless lampreys and hagfishes through the cartilaginous sharks, skates, and rays to the abundant and diverse bony fishes. Most fish species are cold-blooded; however, one species, the opah (Lampris guttatus), is warm-blooded.
The term fish is applied to a variety of vertebrates of several evolutionary lines. It describes a life-form rather than a taxonomic group. As members of the phylum Chordata, fish share certain features with other vertebrates. These features are gill slits at some point in the life cycle, a notochord, or skeletal supporting rod, a dorsal hollow nerve cord, and a tail. Living fishes represent some five classes, which are as distinct from one another as are the four classes of familiar air-breathing animals—amphibians, reptiles, birds, and mammals. For example, the jawless fishes (Agnatha) have gills in pouches and lack limb girdles. Extant agnathans are the lampreys and the hagfishes. As the name implies, the skeletons of fishes of the class Chondrichthyes (from chondr, “cartilage,” and ichthyes, “fish”) are made entirely of cartilage. Modern fish of this class lack a swim bladder, and their scales and teeth are made up of the same placoid material. Sharks, skates, and rays are examples of cartilaginous fishes. The bony fishes are by far the largest class. Examples range from the tiny seahorse to the 450-kg (1,000-pound) blue marlin, from the flattened soles and flounders to the boxy puffers and ocean sunfishes. Unlike the scales of the cartilaginous fishes, those of bony fishes, when present, grow throughout life and are made up of thin overlapping plates of bone. Bony fishes also have an operculum that covers the gill slits.
The study of fishes, the science of ichthyology, is of broad importance. Fishes are of interest to humans for many reasons, the most important being their relationship with and dependence on the environment. A more obvious reason for interest in fishes is their role as a moderate but important part of the world’s food supply. This resource, once thought unlimited, is now realized to be finite and in delicate balance with the biological, chemical, and physical factors of the aquatic environment. Overfishing, pollution, and alteration of the environment are the chief enemies of proper fisheries management, both in fresh waters and in the ocean. (For a detailed discussion of the technology and economics of fisheries, see commercial fishing.) Another practical reason for studying fishes is their use in disease control. As predators on mosquito larvae, they help curb malaria and other mosquito-borne diseases.
Fishes are valuable laboratory animals in many aspects of medical and biological research. For example, the readiness of many fishes to acclimate to captivity has allowed biologists to study behaviour, physiology, and even ecology under relatively natural conditions. Fishes have been especially important in the study of animal behaviour, where research on fishes has provided a broad base for the understanding of the more flexible behaviour of the higher vertebrates. The zebra fish is used as a model in studies of gene expression.
There are aesthetic and recreational reasons for an interest in fishes. Millions of people keep live fishes in home aquariums for the simple pleasure of observing the beauty and behaviour of animals otherwise unfamiliar to them. Aquarium fishes provide a personal challenge to many aquarists, allowing them to test their ability to keep a small section of the natural environment in their homes. Sportfishing is another way of enjoying the natural environment, also indulged in by millions of people every year. Interest in aquarium fishes and sportfishing supports multimillion-dollar industries throughout the world.
Fishes have been in existence for more than 450 million years, during which time they have evolved repeatedly to fit into almost every conceivable type of aquatic habitat. In a sense, land vertebrates are simply highly modified fishes: when fishes colonized the land habitat, they became tetrapod (four-legged) land vertebrates. The popular conception of a fish as a slippery, streamlined aquatic animal that possesses fins and breathes by gills applies to many fishes, but far more fishes deviate from that conception than conform to it. For example, the body is elongate in many forms and greatly shortened in others; the body is flattened in some (principally in bottom-dwelling fishes) and laterally compressed in many others; the fins may be elaborately extended, forming intricate shapes, or they may be reduced or even lost; and the positions of the mouth, eyes, nostrils, and gill openings vary widely. Air breathers have appeared in several evolutionary lines.
Many fishes are cryptically coloured and shaped, closely matching their respective environments; others are among the most brilliantly coloured of all organisms, with a wide range of hues, often of striking intensity, on a single individual. The brilliance of pigments may be enhanced by the surface structure of the fish, so that it almost seems to glow. A number of unrelated fishes have actual light-producing organs. Many fishes are able to alter their coloration—some for the purpose of camouflage, others for the enhancement of behavioral signals.
Fishes range in adult length from less than 10 mm (0.4 inch) to more than 20 metres (60 feet) and in weight from about 1.5 grams (less than 0.06 ounce) to many thousands of kilograms. Some live in shallow thermal springs at temperatures slightly above 42 °C (100 °F), others in cold Arctic seas a few degrees below 0 °C (32 °F) or in cold deep waters more than 4,000 metres (13,100 feet) beneath the ocean surface. The structural and, especially, the physiological adaptations for life at such extremes are relatively poorly known and provide the scientifically curious with great incentive for study.
Almost all natural bodies of water bear fish life, the exceptions being very hot thermal ponds and extremely salt-alkaline lakes, such as the Dead Sea in Asia and the Great Salt Lake in North America. The present distribution of fishes is a result of the geological history and development of Earth as well as the ability of fishes to undergo evolutionary change and to adapt to the available habitats. Fishes may be seen to be distributed according to habitat and according to geographical area. Major habitat differences are marine and freshwater. For the most part, the fishes in a marine habitat differ from those in a freshwater habitat, even in adjacent areas, but some, such as the salmon, migrate from one to the other. The freshwater habitats may be seen to be of many kinds. Fishes found in mountain torrents, Arctic lakes, tropical lakes, temperate streams, and tropical rivers will all differ from each other, both in obvious gross structure and in physiological attributes. Even in closely adjacent habitats where, for example, a tropical mountain torrent enters a lowland stream, the fish fauna will differ. The marine habitats can be divided into deep ocean floors (benthic), mid-water oceanic (bathypelagic), surface oceanic (pelagic), rocky coast, sandy coast, muddy shores, bays, estuaries, and others. Also, for example, rocky coastal shores in tropical and temperate regions will have different fish faunas, even when such habitats occur along the same coastline.
Although much is known about the present geographical distribution of fishes, far less is known about how that distribution came about. Many parts of the fish fauna of the fresh waters of North America and Eurasia are related and undoubtedly have a common origin. The faunas of Africa and South America are related, extremely old, and probably an expression of the drifting apart of the two continents. The fauna of southern Asia is related to that of Central Asia, and some of it appears to have entered Africa. The extremely large shore-fish faunas of the Indian and tropical Pacific oceans comprise a related complex, but the tropical shore fauna of the Atlantic, although containing Indo-Pacific components, is relatively limited and probably younger. The Arctic and Antarctic marine faunas are quite different from each other. The shore fauna of the North Pacific is quite distinct, and that of the North Atlantic more limited and probably younger. Pelagic oceanic fishes, especially those in deep waters, are similar the world over, showing little geographical isolation in terms of family groups. The deep oceanic habitat is very much the same throughout the world, but species differences do exist, showing geographical areas determined by oceanic currents and water masses.
All aspects of the life of a fish are closely correlated with adaptation to the total environment, physical, chemical, and biological. In studies, all the interdependent aspects of fish, such as behaviour, locomotion, reproduction, and physical and physiological characteristics, must be taken into account.
Correlated with their adaptation to an extremely wide variety of habitats is the extremely wide variety of life cycles that fishes display. The great majority hatch from relatively small eggs a few days to several weeks or more after the eggs are scattered in the water. Newly hatched young are still partially undeveloped and are called larvae until body structures such as fins, skeleton, and some organs are fully formed. Larval life is often very short, usually less than a few weeks, but it can be very long, some lampreys continuing as larvae for at least five years. Young and larval fishes, before reaching sexual maturity, must grow considerably, and their small size and other factors often dictate that they live in a habitat different than that of the adults. For example, most tropical marine shore fishes have pelagic larvae. Larval food also is different, and larval fishes often live in shallow waters, where they may be less exposed to predators.
After a fish reaches adult size, the length of its life is subject to many factors, such as innate rates of aging, predation pressure, and the nature of the local climate. The longevity of a species in the protected environment of an aquarium may have nothing to do with how long members of that species live in the wild. Many small fishes live only one to three years at the most. In some species, however, individuals may live as long as 10 or 20 or even 100 years.
Fish behaviour is a complicated and varied subject. As in almost all animals with a central nervous system, the nature of a response of an individual fish to stimuli from its environment depends upon the inherited characteristics of its nervous system, on what it has learned from past experience, and on the nature of the stimuli. Compared with the variety of human responses, however, that of a fish is stereotyped, not subject to much modification by “thought” or learning, and investigators must guard against anthropomorphic interpretations of fish behaviour.
Fishes perceive the world around them by the usual senses of sight, smell, hearing, touch, and taste and by special lateral line water-current detectors. In the few fishes that generate electric fields, a process that might best be called electrolocation aids in perception. One or another of these senses often is emphasized at the expense of others, depending upon the fish’s other adaptations. In fishes with large eyes, the sense of smell may be reduced; others, with small eyes, hunt and feed primarily by smell (such as some eels).
Specialized behaviour is primarily concerned with the three most important activities in the fish’s life: feeding, reproduction, and escape from enemies. Schooling behaviour of sardines on the high seas, for instance, is largely a protective device to avoid enemies, but it is also associated with and modified by their breeding and feeding requirements. Predatory fishes are often solitary, lying in wait to dart suddenly after their prey, a kind of locomotion impossible for beaked parrot fishes, which feed on coral, swimming in small groups from one coral head to the next. In addition, some predatory fishes that inhabit pelagic environments, such as tunas, often school.
Sleep in fishes, all of which lack true eyelids, consists of a seemingly listless state in which the fish maintains its balance but moves slowly. If attacked or disturbed, most can dart away. A few kinds of fishes lie on the bottom to sleep. Most catfishes, some loaches, and some eels and electric fishes are strictly nocturnal, being active and hunting for food during the night and retiring during the day to holes, thick vegetation, or other protective parts of the environment.
Communication between members of a species or between members of two or more species often is extremely important, especially in breeding behaviour (see below Reproduction). The mode of communication may be visual, as between the small so-called cleaner fish and a large fish of a very different species. The larger fish often allows the cleaner to enter its mouth to remove gill parasites. The cleaner is recognized by its distinctive colour and actions and therefore is not eaten, even if the larger fish is normally a predator. Communication is often chemical, signals being sent by specific chemicals called pheromones.
Many fishes have a streamlined body and swim freely in open water. Fish locomotion is closely correlated with habitat and ecological niche (the general position of the animal to its environment).
Many fishes in both marine and fresh waters swim at the surface and have mouths adapted to feed best (and sometimes only) at the surface. Often such fishes are long and slender, able to dart at surface insects or at other surface fishes and in turn to dart away from predators; needlefishes, halfbeaks, and topminnows (such as killifish and mosquito fish) are good examples. Oceanic flying fishes escape their predators by gathering speed above the water surface, with the lower lobe of the tail providing thrust in the water. They then glide hundreds of yards on enlarged, winglike pectoral and pelvic fins. South American freshwater flying fishes escape their enemies by jumping and propelling their strongly keeled bodies out of the water.
So-called mid-water swimmers, the most common type of fish, are of many kinds and live in many habitats. The powerful fusiform tunas and the trouts, for example, are adapted for strong, fast swimming, the tunas to capture prey speedily in the open ocean and the trouts to cope with the swift currents of streams and rivers. The trout body form is well adapted to many habitats. Fishes that live in relatively quiet waters such as bays or lake shores or slow rivers usually are not strong, fast swimmers but are capable of short, quick bursts of speed to escape a predator. Many of these fishes have their sides flattened, examples being the sunfish and the freshwater angelfish of aquarists. Fish associated with the bottom or substrate usually are slow swimmers. Open-water plankton-feeding fishes almost always remain fusiform and are capable of rapid, strong movement (for example, sardines and herrings of the open ocean and also many small minnows of streams and lakes).
Bottom-living fishes are of many kinds and have undergone many types of modification of their body shape and swimming habits. Rays, which evolved from strong-swimming mid-water sharks, usually stay close to the bottom and move by undulating their large pectoral fins. Flounders live in a similar habitat and move over the bottom by undulating the entire body. Many bottom fishes dart from place to place, resting on the bottom between movements, a motion common in gobies. One goby relative, the mudskipper, has taken to living at the edge of pools along the shore of muddy mangrove swamps. It escapes its enemies by flipping rapidly over the mud, out of the water. Some catfishes, synbranchid eels, the so-called climbing perch, and a few other fishes venture out over damp ground to find more promising waters than those that they left. They move by wriggling their bodies, sometimes using strong pectoral fins; most have accessory air-breathing organs. Many bottom-dwelling fishes live in mud holes or rocky crevices. Marine eels and gobies commonly are found in such habitats and for the most part venture far beyond their cavelike homes. Some bottom dwellers, such as the clingfishes (Gobiesocidae), have developed powerful adhesive disks that enable them to remain in place on the substrate in areas such as rocky coasts, where the action of the waves is great.
The methods of reproduction in fishes are varied, but most fishes lay a large number of small eggs, fertilized and scattered outside of the body. The eggs of pelagic fishes usually remain suspended in the open water. Many shore and freshwater fishes lay eggs on the bottom or among plants. Some have adhesive eggs. The mortality of the young and especially of the eggs is very high, and often only a few individuals grow to maturity out of hundreds, thousands, and in some cases millions of eggs laid.
Males produce sperm, usually as a milky white substance called milt, in two (sometimes one) testes within the body cavity. In bony fishes a sperm duct leads from each testis to a urogenital opening behind the vent or anus. In sharks and rays and in cyclostomes the duct leads to a cloaca. Sometimes the pelvic fins are modified to help transmit the milt to the eggs at the female’s vent or on the substrate where the female has placed them. Sometimes accessory organs are used to fertilize females internally—for example, the claspers of many sharks and rays.
In the females the eggs are formed in two ovaries (sometimes only one) and pass through the ovaries to the urogenital opening and to the outside. In some fishes the eggs are fertilized internally but are shed before development takes place. Members of about a dozen families each of bony fishes (teleosts) and sharks bear live young. Many skates and rays also bear live young. In some bony fishes the eggs simply develop within the female, the young emerging when the eggs hatch (ovoviviparous). Others develop within the ovary and are nourished by ovarian tissues after hatching (viviparous). There are also other methods utilized by fishes to nourish young within the female. In all live-bearers the young are born at a relatively large size and are few in number. In one family of primarily marine fishes, the surfperches from the Pacific coast of North America, Japan, and Korea, the males of at least one species are born sexually mature, although they are not fully grown.
Some fishes are hermaphroditic—an individual producing both sperm and eggs, usually at different stages of its life. Self-fertilization, however, is probably rare.
Successful reproduction and, in many cases, defense of the eggs and the young are assured by rather stereotypical but often elaborate courtship and parental behaviour, either by the male or the female or both. Some fishes prepare nests by hollowing out depressions in the sand bottom (cichlids, for example), build nests with plant materials and sticky threads excreted by the kidneys (sticklebacks), or blow a cluster of mucus-covered bubbles at the water surface (gouramis). The eggs are laid in these structures. Some varieties of cichlids and catfishes incubate eggs in their mouths.
Some fishes, such as salmon, undergo long migrations from the ocean and up large rivers to spawn in the gravel beds where they themselves hatched (anadromous fishes). Some, such as the freshwater eels (family Anguillidae), live and grow to maturity in fresh water and migrate to the sea to spawn (catadromous fishes). Other fishes undertake shorter migrations from lakes into streams, within the ocean, or enter spawning habitats that they do not ordinarily occupy in other ways.
The basic structure and function of the fish body are similar to those of all other vertebrates. The usual four types of tissues are present: surface or epithelial, connective (bone, cartilage, and fibrous tissues, as well as their derivative, blood), nerve, and muscle tissues. In addition, the fish’s organs and organ systems parallel those of other vertebrates.
The typical fish body is streamlined and spindle-shaped, with an anterior head, a gill apparatus, and a heart, the latter lying in the midline just below the gill chamber. The body cavity, containing the vital organs, is situated behind the head in the lower anterior part of the body. The anus usually marks the posterior termination of the body cavity and most often occurs just in front of the base of the anal fin. The spinal cord and vertebral column continue from the posterior part of the head to the base of the tail fin, passing dorsal to the body cavity and through the caudal (tail) region behind the body cavity. Most of the body is of muscular tissue, a high proportion of which is necessitated by swimming. In the course of evolution this basic body plan has been modified repeatedly into the many varieties of fish shapes that exist today.
The skeleton forms an integral part of the fish’s locomotion system, as well as serving to protect vital parts. The internal skeleton consists of the skull bones (except for the roofing bones of the head, which are really part of the external skeleton), the vertebral column, and the fin supports (fin rays). The fin supports are derived from the external skeleton but will be treated here because of their close functional relationship to the internal skeleton. The internal skeleton of cyclostomes, sharks, and rays is of cartilage; that of many fossil groups and some primitive living fishes is mostly of cartilage but may include some bone. In place of the vertebral column, the earliest vertebrates had a fully developed notochord, a flexible stiff rod of viscous cells surrounded by a strong fibrous sheath. During the evolution of modern fishes the rod was replaced in part by cartilage and then by ossified cartilage. Sharks and rays retain a cartilaginous vertebral column; bony fishes have spool-shaped vertebrae that in the more primitive living forms only partially replace the notochord. The skull, including the gill arches and jaws of bony fishes, is fully, or at least partially, ossified. That of sharks and rays remains cartilaginous, at times partially replaced by calcium deposits but never by true bone.
The supportive elements of the fins (basal or radial bones or both) have changed greatly during fish evolution. Some of these changes are described in the section below (Evolution and paleontology). Most fishes possess a single dorsal fin on the midline of the back. Many have two and a few have three dorsal fins. The other fins are the single tail and anal fins and paired pelvic and pectoral fins. A small fin, the adipose fin, with hairlike fin rays, occurs in many of the relatively primitive teleosts (such as trout) on the back near the base of the caudal fin.
The skin of a fish must serve many functions. It aids in maintaining the osmotic balance, provides physical protection for the body, is the site of coloration, contains sensory receptors, and, in some fishes, functions in respiration. Mucous glands, which aid in maintaining the water balance and offer protection from bacteria, are extremely numerous in fish skin, especially in cyclostomes and teleosts. Since mucous glands are present in the modern lampreys, it is reasonable to assume that they were present in primitive fishes, such as the ancient Silurian and Devonian agnathans. Protection from abrasion and predation is another function of the fish skin, and dermal (skin) bone arose early in fish evolution in response to this need. It is thought that bone first evolved in skin and only later invaded the cartilaginous areas of the fish’s body, to provide additional support and protection. There is some argument as to which came first, cartilage or bone, and fossil evidence does not settle the question. In any event, dermal bone has played an important part in fish evolution and has different characteristics in different groups of fishes. Several groups are characterized at least in part by the kind of bony scales they possess.
Scales have played an important part in the evolution of fishes. Primitive fishes usually had thick bony plates or thick scales in several layers of bone, enamel, and related substances. Modern teleost fishes have scales of bone, which, while still protective, allow much more freedom of motion in the body. A few modern teleosts (some catfishes, sticklebacks, and others) have secondarily acquired bony plates in the skin. Modern and early sharks possessed placoid scales, a relatively primitive type of scale with a toothlike structure, consisting of an outside layer of enamel-like substance (vitrodentine), an inner layer of dentine, and a pulp cavity containing nerves and blood vessels. Primitive bony fishes had thick scales of either the ganoid or the cosmoid type. Cosmoid scales have a hard, enamel-like outer layer, an inner layer of cosmine (a form of dentine), and then a layer of vascular bone (isopedine). In ganoid scales the hard outer layer is different chemically and is called ganoin. Under this is a cosminelike layer and then a vascular bony layer. The thin, translucent bony scales of modern fishes, called cycloid and ctenoid (the latter distinguished by serrations at the edges), lack enameloid and dentine layers.
Skin has several other functions in fishes. It is well supplied with nerve endings and presumably receives tactile, thermal, and pain stimuli. Skin is also well supplied with blood vessels. Some fishes breathe in part through the skin, by the exchange of oxygen and carbon dioxide between the surrounding water and numerous small blood vessels near the skin surface.
Skin serves as protection through the control of coloration. Fishes exhibit an almost limitless range of colours. The colours often blend closely with the surroundings, effectively hiding the animal. Many fishes use bright colours for territorial advertisement or as recognition marks for other members of their own species, or sometimes for members of other species. Many fishes can change their colour to a greater or lesser degree, by movement of pigment within the pigment cells (chromatophores). Black pigment cells (melanophores), of almost universal occurrence in fishes, are often juxtaposed with other pigment cells. When placed beneath iridocytes or leucophores (bearing the silvery or white pigment guanine), melanophores produce structural colours of blue and green. These colours are often extremely intense, because they are formed by refraction of light through the needlelike crystals of guanine. The blue and green refracted colours are often relatively pure, lacking the red and yellow rays, which have been absorbed by the black pigment (melanin) of the melanophores. Yellow, orange, and red colours are produced by erythrophores, cells containing the appropriate carotenoid pigments. Other colours are produced by combinations of melanophores, erythrophores, and iridocytes.
The major portion of the body of most fishes consists of muscles. Most of the mass is trunk musculature, the fin muscles usually being relatively small. The caudal fin is usually the most powerful fin, being moved by the trunk musculature. The body musculature is usually arranged in rows of chevron-shaped segments on each side. Contractions of these segments, each attached to adjacent vertebrae and vertebral processes, bends the body on the vertebral joint, producing successive undulations of the body, passing from the head to the tail, and producing driving strokes of the tail. It is the latter that provides the strong forward movement for most fishes.
The digestive system, in a functional sense, starts at the mouth, with the teeth used to capture prey or collect plant foods. Mouth shape and tooth structure vary greatly in fishes, depending on the kind of food normally eaten. Most fishes are predacious, feeding on small invertebrates or other fishes and have simple conical teeth on the jaws, on at least some of the bones of the roof of the mouth, and on special gill arch structures just in front of the esophagus. The latter are throat teeth. Most predacious fishes swallow their prey whole, and the teeth are used for grasping and holding prey, for orienting prey to be swallowed (head first) and for working the prey toward the esophagus. There are a variety of tooth types in fishes. Some fishes, such as sharks and piranhas, have cutting teeth for biting chunks out of their victims. A shark’s tooth, although superficially like that of a piranha, appears in many respects to be a modified scale, while that of the piranha is like that of other bony fishes, consisting of dentine and enamel. Parrot fishes have beaklike mouths with short incisor-like teeth for breaking off coral and have heavy pavementlike throat teeth for crushing the coral. Some catfishes have small brushlike teeth, arranged in rows on the jaws, for scraping plant and animal growth from rocks. Many fishes (such as the Cyprinidae or minnows) have no jaw teeth at all but have very strong throat teeth.
Some fishes gather planktonic food by straining it from their gill cavities with numerous elongate stiff rods (gill rakers) anchored by one end to the gill bars. The food collected on these rods is passed to the throat, where it is swallowed. Most fishes have only short gill rakers that help keep food particles from escaping out the mouth cavity into the gill chamber.
Once reaching the throat, food enters a short, often greatly distensible esophagus, a simple tube with a muscular wall leading into a stomach. The stomach varies greatly in fishes, depending upon the diet. In most predacious fishes it is a simple straight or curved tube or pouch with a muscular wall and a glandular lining. Food is largely digested there and leaves the stomach in liquid form.
Between the stomach and the intestine, ducts enter the digestive tube from the liver and pancreas. The liver is a large, clearly defined organ. The pancreas may be embedded in it, diffused through it, or broken into small parts spread along some of the intestine. The junction between the stomach and the intestine is marked by a muscular valve. Pyloric ceca (blind sacs) occur in some fishes at this junction and have a digestive or absorptive function or both.
The intestine itself is quite variable in length, depending upon the fish’s diet. It is short in predacious forms, sometimes no longer than the body cavity, but long in herbivorous forms, being coiled and several times longer than the entire length of the fish in some species of South American catfishes. The intestine is primarily an organ for absorbing nutrients into the bloodstream. The larger its internal surface, the greater its absorptive efficiency, and a spiral valve is one method of increasing its absorption surface.
Sharks, rays, chimaeras, lungfishes, surviving chondrosteans, holosteans, and even a few of the more primitive teleosts have a spiral valve or at least traces of it in the intestine. Most modern teleosts have increased the area of the intestinal walls by having numerous folds and villi (fingerlike projections) somewhat like those in humans. Undigested substances are passed to the exterior through the anus in most teleost fishes. In lungfishes, sharks, and rays, it is first passed through the cloaca, a common cavity receiving the intestinal opening and the ducts from the urogenital system.
Oxygen and carbon dioxide dissolve in water, and most fishes exchange dissolved oxygen and carbon dioxide in water by means of the gills. The gills lie behind and to the side of the mouth cavity and consist of fleshy filaments supported by the gill arches and filled with blood vessels, which give gills a bright red colour. Water taken in continuously through the mouth passes backward between the gill bars and over the gill filaments, where the exchange of gases takes place. The gills are protected by a gill cover in teleosts and many other fishes but by flaps of skin in sharks, rays, and some of the older fossil fish groups. The blood capillaries in the gill filaments are close to the gill surface to take up oxygen from the water and to give up excess carbon dioxide to the water.
Most modern fishes have a hydrostatic (ballast) organ, called the swim bladder, that lies in the body cavity just below the kidney and above the stomach and intestine. It originated as a diverticulum of the digestive canal. In advanced teleosts, especially the acanthopterygians, the bladder has lost its connection with the digestive tract, a condition called physoclistic. The connection has been retained (physostomous) by many relatively primitive teleosts. In several unrelated lines of fishes, the bladder has become specialized as a lung or, at least, as a highly vascularized accessory breathing organ. Some fishes with such accessory organs are obligate air breathers and will drown if denied access to the surface, even in well-oxygenated water. Fishes with a hydrostatic form of swim bladder can control their depth by regulating the amount of gas in the bladder. The gas, mostly oxygen, is secreted into the bladder by special glands, rendering the fish more buoyant; the gas is absorbed into the bloodstream by another special organ, reducing the overall buoyancy and allowing the fish to sink. Some deep-sea fishes may have oils, rather than gas, in the bladder. Other deep-sea and some bottom-living forms have much-reduced swim bladders or have lost the organ entirely.
The swim bladder of fishes follows the same developmental pattern as the lungs of land vertebrates. There is no doubt that the two structures have the same historical origin in primitive fishes. More or less intermediate forms still survive among the more primitive types of fishes, such as the lungfishes Lepidosiren and Protopterus.
The circulatory, or blood vascular, system consists of the heart, the arteries, the capillaries, and the veins. It is in the capillaries that the interchange of oxygen, carbon dioxide, nutrients, and other substances such as hormones and waste products takes place. The capillaries lead to the veins, which return the venous blood with its waste products to the heart, kidneys, and gills. There are two kinds of capillary beds: those in the gills and those in the rest of the body. The heart, a folded continuous muscular tube with three or four saclike enlargements, undergoes rhythmic contractions and receives venous blood in a sinus venosus. It passes the blood to an auricle and then into a thick muscular pump, the ventricle. From the ventricle the blood goes to a bulbous structure at the base of a ventral aorta just below the gills. The blood passes to the afferent (receiving) arteries of the gill arches and then to the gill capillaries. There waste gases are given off to the environment, and oxygen is absorbed. The oxygenated blood enters efferent (exuant) arteries of the gill arches and then flows into the dorsal aorta. From there blood is distributed to the tissues and organs of the body. One-way valves prevent backflow. The circulation of fishes thus differs from that of the reptiles, birds, and mammals in that oxygenated blood is not returned to the heart prior to distribution to the other parts of the body.
The primary excretory organ in fishes, as in other vertebrates, is the kidney. In fishes some excretion also takes place in the digestive tract, skin, and especially the gills (where ammonia is given off). Compared with land vertebrates, fishes have a special problem in maintaining their internal environment at a constant concentration of water and dissolved substances, such as salts. Proper balance of the internal environment (homeostasis) of a fish is in a great part maintained by the excretory system, especially the kidney.
The kidney, gills, and skin play an important role in maintaining a fish’s internal environment and checking the effects of osmosis. Marine fishes live in an environment in which the water around them has a greater concentration of salts than they can have inside their body and still maintain life. Freshwater fishes, on the other hand, live in water with a much lower concentration of salts than they require inside their bodies. Osmosis tends to promote the loss of water from the body of a marine fish and absorption of water by that of a freshwater fish. Mucus in the skin tends to slow the process but is not a sufficient barrier to prevent the movement of fluids through the permeable skin. When solutions on two sides of a permeable membrane have different concentrations of dissolved substances, water will pass through the membrane into the more concentrated solution, while the dissolved chemicals move into the area of lower concentration (diffusion).
The kidney of freshwater fishes is often larger in relation to body weight than that of marine fishes. In both groups the kidney excretes wastes from the body, but the kidney of freshwater fishes also excretes large amounts of water, counteracting the water absorbed through the skin. Freshwater fishes tend to lose salt to the environment and must replace it. They get some salt from their food, but the gills and skin inside the mouth actively absorb salt from water passed through the mouth. This absorption is performed by special cells capable of moving salts against the diffusion gradient. Freshwater fishes drink very little water and take in little water with their food.
Marine fishes must conserve water, and therefore their kidneys excrete little water. To maintain their water balance, marine fishes drink large quantities of seawater, retaining most of the water and excreting the salt. Most nitrogenous waste in marine fishes appears to be secreted by the gills as ammonia. Marine fishes can excrete salt by clusters of special cells (chloride cells) in the gills.
There are several teleosts—for example, the salmon—that travel between fresh water and seawater and must adjust to the reversal of osmotic gradients. They adjust their physiological processes by spending time (often surprisingly little time) in the intermediate brackish environment.
Marine hagfishes, sharks, and rays have osmotic concentrations in their blood about equal to that of seawater and so do not have to drink water nor perform much physiological work to maintain their osmotic balance. In sharks and rays the osmotic concentration is kept high by retention of urea in the blood. Freshwater sharks have a lowered concentration of urea in the blood.
Endocrine glands secrete their products into the bloodstream and body tissues and, along with the central nervous system, control and regulate many kinds of body functions. Cyclostomes have a well-developed endocrine system, and presumably it was well developed in the early Agnatha, ancestral to modern fishes. Although the endocrine system in fishes is similar to that of higher vertebrates, there are numerous differences in detail. The pituitary, the thyroid, the suprarenals, the adrenals, the pancreatic islets, the sex glands (ovaries and testes), the inner wall of the intestine, and the bodies of the ultimobranchial gland make up the endocrine system in fishes. There are some others whose function is not well understood. These organs regulate sexual activity and reproduction, growth, osmotic pressure, general metabolic activities such as the storage of fat and the utilization of foodstuffs, blood pressure, and certain aspects of skin colour. Many of these activities are also controlled in part by the central nervous system, which works with the endocrine system in maintaining the life of a fish. Some parts of the endocrine system are developmentally, and undoubtedly evolutionarily, derived from the nervous system.
As in all vertebrates, the nervous system of fishes is the primary mechanism coordinating body activities, as well as integrating these activities in the appropriate manner with stimuli from the environment. The central nervous system, consisting of the brain and spinal cord, is the primary integrating mechanism. The peripheral nervous system, consisting of nerves that connect the brain and spinal cord to various body organs, carries sensory information from special receptor organs such as the eyes, internal ears, nares (sense of smell), taste glands, and others to the integrating centres of the brain and spinal cord. The peripheral nervous system also carries information via different nerve cells from the integrating centres of the brain and spinal cord. This coded information is carried to the various organs and body systems, such as the skeletal muscular system, for appropriate action in response to the original external or internal stimulus. Another branch of the nervous system, the autonomic nervous system, helps to coordinate the activities of many glands and organs and is itself closely connected to the integrating centres of the brain.
The brain of the fish is divided into several anatomical and functional parts, all closely interconnected but each serving as the primary centre of integrating particular kinds of responses and activities. Several of these centres or parts are primarily associated with one type of sensory perception, such as sight, hearing, or smell (olfaction).
The sense of smell is important in almost all fishes. Certain eels with tiny eyes depend mostly on smell for location of food. The olfactory, or nasal, organ of fishes is located on the dorsal surface of the snout. The lining of the nasal organ has special sensory cells that perceive chemicals dissolved in the water, such as substances from food material, and send sensory information to the brain by way of the first cranial nerve. Odour also serves as an alarm system. Many fishes, especially various species of freshwater minnows, react with alarm to a chemical released from the skin of an injured member of their own species.
Many fishes have a well-developed sense of taste, and tiny pitlike taste buds or organs are located not only within their mouth cavities but also over their heads and parts of their body. Catfishes, which often have poor vision, have barbels (“whiskers”) that serve as supplementary taste organs, those around the mouth being actively used to search out food on the bottom. Some species of naturally blind cave fishes are especially well supplied with taste buds, which often cover most of their body surface.
Sight is extremely important in most fishes. The eye of a fish is basically like that of all other vertebrates, but the eyes of fishes are extremely varied in structure and adaptation. In general, fishes living in dark and dim water habitats have large eyes, unless they have specialized in some compensatory way so that another sense (such as smell) is dominant, in which case the eyes will often be reduced. Fishes living in brightly lighted shallow waters often will have relatively small but efficient eyes. Cyclostomes have somewhat less elaborate eyes than other fishes, with skin stretched over the eyeball perhaps making their vision somewhat less effective. Most fishes have a spherical lens and accommodate their vision to far or near subjects by moving the lens within the eyeball. A few sharks accommodate by changing the shape of the lens, as in land vertebrates. Those fishes that are heavily dependent upon the eyes have especially strong muscles for accommodation. Most fishes see well, despite the restrictions imposed by frequent turbidity of the water and by light refraction.
Fossil evidence suggests that colour vision evolved in fishes more than 300 million years ago, but not all living fishes have retained this ability. Experimental evidence indicates that many shallow-water fishes, if not all, have colour vision and see some colours especially well, but some bottom-dwelling shore fishes live in areas where the water is sufficiently deep to filter out most if not all colours, and these fishes apparently never see colours. When tested in shallow water, they apparently are unable to respond to colour differences.
Sound perception and balance are intimately associated senses in a fish. The organs of hearing are entirely internal, located within the skull, on each side of the brain and somewhat behind the eyes. Sound waves, especially those of low frequencies, travel readily through water and impinge directly upon the bones and fluids of the head and body, to be transmitted to the hearing organs. Fishes readily respond to sound; for example, a trout conditioned to escape by the approach of fishermen will take flight upon perceiving footsteps on a stream bank even if it cannot see a fisherman. Compared with humans, however, the range of sound frequencies heard by fishes is greatly restricted. Many fishes communicate with each other by producing sounds in their swim bladders, in their throats by rasping their teeth, and in other ways.
A fish or other vertebrate seldom has to rely on a single type of sensory information to determine the nature of the environment around it. A catfish uses taste and touch when examining a food object with its oral barbels. Like most other animals, fishes have many touch receptors over their body surface. Pain and temperature receptors also are present in fishes and presumably produce the same kind of information to a fish as to humans. Fishes react in a negative fashion to stimuli that would be painful to human beings, suggesting that they feel a sensation of pain.
An important sensory system in fishes that is absent in other vertebrates (except some amphibians) is the lateral line system. This consists of a series of heavily innervated small canals located in the skin and bone around the eyes, along the lower jaw, over the head, and down the mid-side of the body, where it is associated with the scales. Intermittently along these canals are located tiny sensory organs (pit organs) that apparently detect changes in pressure. The system allows a fish to sense changes in water currents and pressure, thereby helping the fish to orient itself to the various changes that occur in the physical environment.
This youngster, walking across the road to finally join his family who had crossed a few minutes earlier, just looked like a stroppy teenager to us. Before he reached the herd he proceded to dust himself down with more red dirt, not that he wasn't already red enough! LOL.
Elephants are large mammals of the family Elephantidae and the order Proboscidea. Two species are traditionally recognised, the African elephant (Loxodonta africana) and the Asian elephant (Elephas maximus), although some evidence suggests that African bush elephants and African forest elephants are separate species (L. africana and L. cyclotis respectively). Elephants are scattered throughout sub-Saharan Africa, South Asia, and Southeast Asia. Elephantidae is the only surviving family of the order Proboscidea; other, now extinct, members of the order include deinotheres, gomphotheres, mammoths, and mastodons. Male African elephants are the largest extant terrestrial animals and can reach a height of 4 m (13 ft) and weigh 7,000 kg (15,000 lb). All elephants have several distinctive features the most notable of which is a long trunk or proboscis, used for many purposes, particularly breathing, lifting water and grasping objects. Their incisors grow into tusks, which can serve as weapons and as tools for moving objects and digging. Elephants' large ear flaps help to control their body temperature. Their pillar-like legs can carry their great weight. African elephants have larger ears and concave backs while Asian elephants have smaller ears and convex or level backs.
Elephants are herbivorous and can be found in different habitats including savannahs, forests, deserts and marshes. They prefer to stay near water. They are considered to be keystone species due to their impact on their environments. Other animals tend to keep their distance where predators such as lions, tigers, hyenas, and wild dogs usually target only the young elephants (or "calves"). Females ("cows") tend to live in family groups, which can consist of one female with her calves or several related females with offspring. The groups are led by an individual known as the matriarch, often the oldest cow. Elephants have a fission–fusion society in which multiple family groups come together to socialise. Males ("bulls") leave their family groups when they reach puberty, and may live alone or with other males. Adult bulls mostly interact with family groups when looking for a mate and enter a state of increased testosterone and aggression known as musth, which helps them gain dominance and reproductive success. Calves are the centre of attention in their family groups and rely on their mothers for as long as three years. Elephants can live up to 70 years in the wild. They communicate by touch, sight, smell and sound; elephants use infrasound, and seismic communication over long distances. Elephant intelligence has been compared with that of primates and cetaceans. They appear to have self-awareness and show empathy for dying or dead individuals of their kind.
African bush elephant
Afrikanischer Elefant
Kruger National Park is one of the largest game reserves in Africa. It covers an area of 19,485 km2 (7,523 sq mi) in the provinces of Limpopo and Mpumalanga in northeastern South Africa, and extends 360 km (220 mi) from north to south and 65 km (40 mi) from east to west. The administrative headquarters are in Skukuza. Areas of the park were first protected by the government of the South African Republic in 1898, and it became South Africa's first national park in 1926.
To the west and south of the Kruger National Park are the two South African provinces of Limpopo and Mpumalanga. In the north is Zimbabwe, and to the east is Mozambique. It is now part of the Great Limpopo Transfrontier Park, a peace park that links Kruger National Park with the Gonarezhou National Park in Zimbabwe, and with the Limpopo National Park in Mozambique.
The park is part of the Kruger to Canyons Biosphere an area designated by the United Nations Educational, Scientific and Cultural Organization (UNESCO) as an International Man and Biosphere Reserve (the "Biosphere").
The park has nine main gates allowing entrance to the different camps.
(Wikipedia)
Name
African Elephant or African Bush Elephant [Loxodonta africana]
Introduction
The Elephant is the world's largest land mammal, and weighs up to 7 tonnes and reaches heights of 3.3 m at the shoulder. Elephants can live to a potential age of 70 years. The massive tusks of older bulls can weigh up to 50 or 60 kilograms, but tusks weighing up to 90 kilograms have been recorded.
Appearance
What is the trunk and what is it used for?
The Elephant's trunk is a modified nose which is very sensitive and can even detect water under ground. There are as many as 50 000 muscles in an Elephant trunk. The sensitive finger-like appendages at the tip of the trunk enables them to pick the smallest twig or flower, pull the toughest reed of grass or even pick out a thorn from their feet.
Do elephants have knees or elbows?
The joints that are perceived as 'knees', are in fact wrists. This is a common misunderstanding due to the belief that a leg joint that bends between the foot and the body must be a knee. The main difference between us and the elephants is that our foot bones and hand bones are separate, whereas those of the elephant are one in the same, and have evolved to suit this four-legged mammal.
Why do elephants have tusks?
The tusks are used for obtaining food, fighting (amongst males) and for self defence. They are actually their upper incisors, and grow continuously until they die at around 60 years old. Although their skin is up to 3cm (1 inch) thick, it is quite sensitive.
Diet
Elephants are voracious feeders which in a day consume up to 272 kg (600 pounds) of grass, tender shoots and bark from trees. An adult Elephant can drink up to 200 litres of water in a single session. A single Elephant deposits up to 150kg (330 pounds) of dung every day - about one dollop every 15 minutes!
Breeding
African Elephant are not seasonal breeders. Generally they produce one calf every 3 to 4 years. The gestation period is about 22 months. At birth calves weigh about 100 kg (220 pounds) and are fully weaned between 18-24 months. An orphaned calf will usually be adopted by one of the family's lactating females or suckled by various females. Elephants are very attentive mothers, and because most Elephant behavior has to be learned, they keep their offspring with them for many years. Tusks erupt at 16 months but do not show externally until 30 months. Once weaned, usually at age 4 or 5, the calf still remains in the maternal group. Females mature at about 11 years and stay in the group, while the males, which mature between 12 and 15, are usually expelled from the maternal herd. Even though these young males are sexually mature, they do not breed until they are in their mid, or late 20s or even older and have moved up in the social hierarchy.
Behaviour
Mature males form bachelor groups and become solitary bulls. Elephant form strong family units of cows, calves and young offspring. Such herds are always led by an old female. Apart from drinking large quantities of water they also love wading or swimming in it. Elephants clearly relish mud baths.
It was once thought that family groups were led by old bull elephants, but these males are most often solitary. The female family groups are often visited by mature males checking for females in oestrus. Several interrelated family groups may inhabit an area and know each other well.
How do you tell an elephant's mock charge from a serious one?
It is imperative to keep in mind that Elephant are extremely intelligent, and each individual has a distinct character. Although there will be exceptions to the rules, the common signs of a mock charge are bush-bashing, dust-throwing, trumpeting and other vocalizations, open ears and an intimidating presence, can be considered a mock-display. Aggressive or startled elephants usually make sudden headshakes and flap their large ears against their head. Serious charges usually occur after all attempts to intimidate have failed, and the Elephant feels threatened. The ears are pinned back and head and trunk are lowered. Ultimately, the key lies in the intelligence of the animal and how they will react to the 'target' and unfamiliar actions, and a conscious decision is made.
Why do elephants rhythmically flap their ears?
Contrary to common belief, it is not an expression of anger. Being an animal of such a large size, with no sweat glands and a dark body colour, elephants flap their ears to cool the body and rid themselves of irritating insects.
Where are they found?
Once ranging across most of Africa the Elephant population has declined dramatically across the continent. In South Africa the Addo Elephant and Kruger National Park protect large herds. Due to rigorous conservation measures the Elephant population in South Africa has grown from a estimated 120 in 1920 in 4 locations, to 10 000 at 40 locations to date.
Notes
The African Elephant has recently been classified into two separate species, the more common African Bush Elephant [Loxodonta Africana] and the smaller African Forest Elephant [Loxodonta cyclotis] of the rainforest of Central Africa.
(krugerpark.co.za)
Der Kruger-Nationalpark (deutsch häufig falsch Krüger-Nationalpark) ist das größte Wildschutzgebiet Südafrikas. Er liegt im Nordosten des Landes in der Landschaft des Lowveld auf dem Gebiet der Provinz Limpopo sowie des östlichen Abschnitts von Mpumalanga. Seine Fläche erstreckt sich vom Crocodile-River im Süden bis zum Limpopo, dem Grenzfluss zu Simbabwe, im Norden. Die Nord-Süd-Ausdehnung beträgt etwa 350 km, in Ost-West-Richtung ist der Park durchschnittlich 54 km breit und umfasst eine Fläche von rund 20.000 Quadratkilometern. Damit gehört er zu den größten Nationalparks in Afrika.
Das Schutzgebiet wurde am 26. März 1898 unter dem Präsidenten Paul Kruger als Sabie Game Reserve zum Schutz der Wildnis gegründet. 1926 erhielt das Gebiet den Status Nationalpark und wurde in seinen heutigen Namen umbenannt. Im Park leben 147 Säugetierarten inklusive der „Big Five“, außerdem etwa 507 Vogelarten und 114 Reptilienarten, 49 Fischarten und 34 Amphibienarten.
(Wikipedia)
Der Afrikanische Elefant (Loxodonta africana), auch Afrikanischer Steppenelefant oder Afrikanischer Buschelefant, ist eine Art aus der Familie der Elefanten. Er ist das größte gegenwärtig lebende Landsäugetier und gleichzeitig das größte rezente landbewohnende Tier der Erde. Herausragende Kennzeichen sind neben den Stoßzähnen und dem markanten Rüssel die großen Ohren und die säulenförmigen Beine. In zahlreichen morphologischen und anatomischen Merkmalen unterscheidet sich der Afrikanische Elefant von seinen etwas kleineren Verwandten, dem Waldelefanten und dem Asiatischen Elefanten. Das Verbreitungsgebiet umfasst heute große Teile von Afrika südlich der Sahara. Die Tiere haben sich dort an zahlreiche unterschiedliche Lebensräume angepasst, die von geschlossenen Wäldern über offene Savannenlandschaften bis hin zu Sumpfgebieten und wüstenartigen Regionen reichen. Insgesamt ist das Vorkommen aber stark fragmentiert.
Die Lebensweise des Afrikanischen Elefanten ist durch intensive Studien gut erforscht. Sie wird durch einen stark sozialen Charakter geprägt. Weibliche Tiere und ihr Nachwuchs leben in Familienverbänden (Herden). Diese formieren sich wiederum zu einem enger verwandten Clan. Die einzelnen Herden treffen sich zu bestimmten Gelegenheiten und trennen sich danach wieder. Die männlichen Tiere bilden Junggesellengruppen. Die verschiedenen Verbände nutzen Aktionsräume, in denen sie teils im Jahreszyklus herumwandern. Für die Kommunikation untereinander nutzen die Tiere verschiedene Töne im niedrigen Frequenzbereich. Anhand der Lautgebung, aber auch durch bestimmte chemische Signale können sich die einzelnen Individuen untereinander erkennen. Darüber hinaus besteht ein umfangreiches Repertoire an Gesten. Hervorzuheben sind auch die kognitiven Fähigkeiten des Afrikanischen Elefanten.
Die Nahrung besteht sowohl aus weicher wie auch harter Pflanzenkost. Die genaue Zusammensetzung variiert dabei regional und jahreszeitlich. Generell verbringt der Afrikanische Elefant einen großen Teil seiner Tagesaktivitäten mit der Nahrungsaufnahme. Die Fortpflanzung erfolgt ganzjährig, regional gibt es Tendenzen zu einer stärkeren Saisonalisierung. Bullen kommen einmal jährlich in die Musth, während deren sie auf Wanderung zur Suche nach fortpflanzungswilligen Kühen gehen. Während der Musth ist die Aggressivität gesteigert, es finden dann auch Rivalenkämpfe statt. Der Sexualzyklus der Kühe dauert vergleichsweise lange und weist einen für Säugetiere untypischen Verlauf auf. Nach erfolgter Geburt setzt er in der Regel mehrere Jahre aus. Zumeist wird nach fast zweijähriger Tragzeit ein Jungtier geboren, das in der mütterlichen Herde aufwächst. Junge weibliche Tiere verbleiben später in der Herde, die jungen männlichen verlassen diese.
Die wissenschaftliche Erstbeschreibung des Afrikanischen Elefanten erfolgte im Jahr 1797 mit einer formalen artlichen Trennung des Afrikanischen vom Asiatischen Elefanten. Der heute gebräuchliche Gattungsname Loxodonta wurde offiziell erst dreißig Jahre später eingeführt. Die Bezeichnung bezieht sich auf markante Zahnunterschiede zwischen den asiatischen und den afrikanischen Elefanten. Im Verlauf des 20. Jahrhunderts wurden mehrere Unterarten unterschieden, darunter auch der Waldelefant des zentralen Afrikas. Letzterer gilt heute genetischen Untersuchungen zufolge als eigenständige Art, die weiteren Unterarten sind nicht anerkannt. Stammesgeschichtlich lässt sich der Afrikanische Elefant erstmals im beginnenden Mittleren Pleistozän belegen. Der Gesamtbestand gilt als gefährdet. Ursachen hierfür sind hauptsächlich die Jagd nach Elfenbein und Lebensraumverlust durch die zunehmend wachsende menschliche Bevölkerung. Der Afrikanische Elefant zählt zu den sogenannten „Big Five“ von Großwildjagd und Safari.
(Wikipedia)
The Mursi (also called Murzu) is the most popular tribe in the southwestern Ethiopia lower Omo Valley, 100 km north of Kenyan. They are estimated to 10 000 people and live in the Mago National Park, established in 1979. Due to the climate, they move twice a year between the winter and summer months. They herd cattle and grow crops along the banks of the Omo River. The Mursi are sedentary rather than nomadic. Their language belongs to the Nilo-Saharan linguistic family.Very few Mursi people speak Amharic, the official Ethiopian language. Although a small percentage of the Mursi tribe are Christians, most still practice animism. Mursi women wear giant lip plate, a sign of beauty, like in Suri tribe, and also a prime attraction for tourists which help to sustain a view of them, in guidebooks and travel articles, as an untouched people, living in one of the last wildernesses of Africa. When they are ready to marry, teenagers start to make a hole in the lower lip with a wood stick.
It will be kept for one night, and is removed to put a bigger one. This is very painful at this time. Few months after, the lip plate has its full size, and the girl is seen as beautiful by the men. The lip plate is made of wood or terracotta. They have to remove the lower incisors to let some space for the disc. Sometimes the lip is broken by the pressure of the plate. This is a big problem for the girl because men will consider her as ugly, she won't be able to marry anyone in the tribe apart the old men or the sick people. Women and men are shaved because they hate hairiness. Both like to make scarifications on their bodies. Women as a beauty sign, men after killing animals or ennemies as competition for grazing land has led to tribal conflicts.
The Mursi men have a reputation for being aggressive and are famous for their stick fighting ceremony called donga. The winner of the donga will be able to select the girl of his choice to have relations with if she agrees. Similar to the Surma tribe, the Mursi tribe commonly drink a mixture of blood and milk. Over the past few decades they and their neighbours have faced growing threats to their livelihoods cause the Ethiopian government officials have been actively evicting Mursi people from the Omo National Park, without any compensation to rent their land to foreign investors. Drought has made it difficult for many families to feed themselves by means of their traditional mix of subsistence activities. The establishment of hunting concessions has added to the pressure on scarce ressources.
© Eric Lafforgue
Commentary.
Amazing Suilven changes in form
as we circumnavigate it.
From the west, a sugar-loaf dome, near vertical.
From others, a giant elephant.
West peak, its rump.
Central col, a dip in its backbone.
Easterly peak, a sharper point to the top of its skull.
From yet others it appears like an incisor tooth,
thrusting up from an undulating, rocky base
of “Knock and Lochan” or small hill and lake.
The mountains of Sutherland don’t reach 1,000 metres.
But because of their stark, isolated rise,
they seem double their actual height.
They arrest one’s attention.
They demand focus.
They bemuse, by constantly changing form, shape and character,
and none more so, than the captivating,
iconic, monolith known as Suilven!
The brown-throated sloth (Bradypus variegatus) is a species of three-toed sloth found in the Neotropical realm of Central and South America.
It is the most common of the four species of three-toed sloth, and is found in the forests of South and Central America.
The brown-throated sloth is of similar size and build to most other species of three-toed sloths, with both males and females being 42 to 80 cm (17 to 31 in) in total body length. The tail is relatively short, only 2.5 to 9 cm (1.0 to 3.5 in) long. Adults weigh from 2.25 to 6.3 kg (5.0 to 13.9 lb), with no significant size difference between males and females. Each foot has three fingers, ending in long, curved claws, which are 7 to 8 cm (2.8 to 3.1 in) long on the fore feet, and 5 to 5.5 cm (2.0 to 2.2 in) on the hind feet.
The head is rounded, with a blunt nose and inconspicuous ears. As with other sloths, the brown-throated sloth has no incisor or canine teeth, and the cheek teeth are simple and peg-like. They have no gall bladder, cecum, or appendix.
The brown-throated sloth has grayish-brown to beige-color fur over the body, with darker brown fur on the throat, the sides of the face, and the forehead. The face is generally paler in color, with a stripe of very dark fur running beneath the eyes.
This image was taken in the Animal and Bird Park, near the Cruise Terminal at Cartagena in Columbia.
A herd of Impalas looking back in the direction that they came from.
Impala - Fast food of the African bush!
The impala even has the distinctive MacDonald's "M" on its rump and are also called predator take-aways. There are thousands of them in the Kruger Park and we get so used to spotting them that eventually we just ignore them.
Impalas have evolved a number of unique adaptations that allow them to thrive in their dangerous environment.
Grooming and tick removal are high on the list of survival skills. The impala is the only hoofed animal that allogrooms - mutual grooming from one animal to another - as well as grooming itself.
It is also the smallest antelope that tolerates oxpecker birds to assist in removing ticks. Ticks can reduce blood reserves exposing the antelope to disease and malnutrition. To assist them in grooming, impalas possess an "antelope's toothcomb" comprised of canines and incisors adapted for removing ticks and other parasites. By allogrooming they get rid of ticks etc in the unreachable places, like around the ears, head and neck. When it comes to the area under the tails the two black stripes of the MacDonalds logo come into play. The constantly wagging tail of the impala brushes the ticks towards the warmer black hair where the impala can reach with its teeth.
Unlike most animals, the impalas graze and browse, thereby maximising the availability of food. For safety they move in herds as large as the available food will allow. They scatter in all directions if a predator should charge, leaping in strides of up to 12 meters. Impala ewes give birth away from the herd and rejoins with her calf within two days. With lambs in the herd, vigilance is sharpened, but the lambs still fall prey to wild dogs and other predators.
Even though they are considered predator "fast food", their numbers are increasing and these graceful antelope will not be on the endangered list for some time to come.
Source: Getaway Magazine - January 2004
Kruger National Park as seen from Ngwenya Lodge
Mpumalanga
South Africa
This Somali wild ass lives at Africa Alive at Kessingland in Suffolk.
Wild asses have been an important part of Africa's history. The Egyptians domesticated asses more than 6,000 years ago. Their images have been painted in caves by ancient peoples in North Africa, and, at one time, sultans of the area ordered a man's hand cut off if he killed a wild ass.
The Somali wild ass is found in Somalia, the Southern Red Sea region of Eritrea and the Afar Region of Ethiopia. They have a soft grey body, white belly, spiky upright black and grey mane and unique black- and white stripes on their legs that hint to their zebra family connections. Their small, narrow hooves allow the ass move quickly and safely through its stony habitat. Adults average 6ft 6in. (2 m) in length and stand between 4 to 5ft. (1.22 to 1.53 m) tall at the shoulder. Their average weight is 605 lb. (275 kg).
Native to rocky desert, the animal has to deal with extremely high temperatures. Their large ears not only help them pick up sounds, they assist with the dissipation of heat. A wild ass can lose almost one-third of its body weight in water and still survive. They rest under any shade they can find until the temperature cools down in the late afternoon and evening and graze in the early morning hours.
All wild asses are herbivores. They grazing on grasses, but they also eat scrub, bark, and tough desert plants. Their upper and lower incisor teeth clip grass and other vegetation, molars help to grind the rough food before it's swallowed.
Somali wild asses often live alone due to the lack of food to be found in their environment. Small herds do exist, though, usually comprised of mares and their offspring. Occasionally, larger groups form during the wet season when food and water are more plentiful. This is also when foals are born.
Usually 1 foal is born every other year. At birth it will weigh between 50 to 60 lb. (23 to 27 kg). Foals are able to follow their mother within a few hours after birth, and usually by the time they are one year old, they are half grown, weaned, and no longer need their mother. The foal can expect to live for up to 40 years.
Wild asses are intelligent creatures, with excellent vision and hearing. They'd rather run from predators than fight. If cornered, though, they can kick hard to protect themselves. One swift kick from a sharp hoof is enough to drive away most predators, such as wolves.
There are likely fewer than 1,000 animals in the wild and as of 2011, there were about 200 individuals in captivity around the world. Political unrest, encroachment of their land and water sources by domestic herds, poaching, and a general lack of concern for their welfare are all major threats to the wild asses. Another problem is Somali wild ass will mate with domestic donkeys and this is a serious threat to the gene pool of the wild ass species.
The conservation status of this animal is 'Critically Endangered'.
Fun fact: Adult sloth bears are missing their top two front teeth (incisors), which enables them to suck up termites and other insects with ease.
Read more about these unique bears here:
Beavers are large, semiaquatic rodents in the genus Castor native to the temperate Northern Hemisphere. There are two extant species: the North American beaver (Castor canadensis) and the Eurasian beaver (C. fiber). Beavers are the second-largest living rodents after the capybaras. They have stout bodies with large heads, long chisel-like incisors, brown or gray fur, hand-like front feet, webbed back feet and flat, scaly tails. The Eurasian beaver has a more elongated skull with a more triangular nasal bone opening, lighter fur color and a narrower tail. The animals can be found in a number of freshwater habitats, such as rivers, streams, lakes and ponds. They are herbivorous, consuming tree bark, aquatic plants, grasses and sedges.
Sheep (pl.: sheep) or domestic sheep (Ovis aries) are a domesticated, ruminant mammal typically kept as livestock. Although the term sheep can apply to other species in the genus Ovis, in everyday usage it almost always refers to domesticated sheep. Like all ruminants, sheep are members of the order Artiodactyla, the even-toed ungulates. Numbering a little over one billion, domestic sheep are also the most numerous species of sheep. An adult female is referred to as a ewe (/juː/ yoo), an intact male as a ram, occasionally a tup, a castrated male as a wether, and a young sheep as a lamb.
Sheep are most likely descended from the wild mouflon of Europe and Asia, with Iran being a geographic envelope of the domestication center. One of the earliest animals to be domesticated for agricultural purposes, sheep are raised for fleeces, meat (lamb, hogget or mutton) and milk. A sheep's wool is the most widely used animal fiber, and is usually harvested by shearing. In Commonwealth countries, ovine meat is called lamb when from younger animals and mutton when from older ones; in the United States, meat from both older and younger animals is usually called lamb. Sheep continue to be important for wool and meat today, and are also occasionally raised for pelts, as dairy animals, or as model organisms for science.
Sheep husbandry is practised throughout the majority of the inhabited world, and has been fundamental to many civilizations. In the modern era, Australia, New Zealand, the southern and central South American nations, and the British Isles are most closely associated with sheep production.
There is a large lexicon of unique terms for sheep husbandry which vary considerably by region and dialect. Use of the word sheep began in Middle English as a derivation of the Old English word scēap. A group of sheep is called a flock. Many other specific terms for the various life stages of sheep exist, generally related to lambing, shearing, and age.
Being a key animal in the history of farming, sheep have a deeply entrenched place in human culture, and are represented in much modern language and symbolism. As livestock, sheep are most often associated with pastoral, Arcadian imagery. Sheep figure in many mythologies—such as the Golden Fleece—and major religions, especially the Abrahamic traditions. In both ancient and modern religious ritual, sheep are used as sacrificial animals.
History
Main article: History of the domestic sheep
The exact line of descent from wild ancestors to domestic sheep is unclear. The most common hypothesis states that Ovis aries is descended from the Asiatic (O. gmelini) species of mouflon; the European mouflon (Ovis aries musimon) is a direct descendant of this population. Sheep were among the first animals to be domesticated by humankind (although the domestication of dogs probably took place 10 to 20 thousand years earlier); the domestication date is estimated to fall between 11,000 and 9000 B.C in Mesopotamia and possibly around 7000 BC in Mehrgarh in the Indus Valley. The rearing of sheep for secondary products, and the resulting breed development, began in either southwest Asia or western Europe. Initially, sheep were kept solely for meat, milk and skins. Archaeological evidence from statuary found at sites in Iran suggests that selection for woolly sheep may have begun around 6000 BC, and the earliest woven wool garments have been dated to two to three thousand years later.
Sheep husbandry spread quickly in Europe. Excavations show that in about 6000 BC, during the Neolithic period of prehistory, the Castelnovien people, living around Châteauneuf-les-Martigues near present-day Marseille in the south of France, were among the first in Europe to keep domestic sheep. Practically from its inception, ancient Greek civilization relied on sheep as primary livestock, and were even said to name individual animals. Ancient Romans kept sheep on a wide scale, and were an important agent in the spread of sheep raising. Pliny the Elder, in his Natural History (Naturalis Historia), speaks at length about sheep and wool. European colonists spread the practice to the New World from 1493 onwards.
Characteristics
Domestic sheep are relatively small ruminants, usually with a crimped hair called wool and often with horns forming a lateral spiral. They differ from their wild relatives and ancestors in several respects, having become uniquely neotenic as a result of selective breeding by humans. A few primitive breeds of sheep retain some of the characteristics of their wild cousins, such as short tails. Depending on breed, domestic sheep may have no horns at all (i.e. polled), or horns in both sexes, or in males only. Most horned breeds have a single pair, but a few breeds may have several.
Sheep in Turkmenistan
Another trait unique to domestic sheep as compared to wild ovines is their wide variation in color. Wild sheep are largely variations of brown hues, and variation within species is extremely limited. Colors of domestic sheep range from pure white to dark chocolate brown, and even spotted or piebald. Sheep keepers also sometimes artificially paint "smit marks" onto their sheep in any pattern or color for identification. Selection for easily dyeable white fleeces began early in sheep domestication, and as white wool is a dominant trait it spread quickly. However, colored sheep do appear in many modern breeds, and may even appear as a recessive trait in white flocks. While white wool is desirable for large commercial markets, there is a niche market for colored fleeces, mostly for handspinning. The nature of the fleece varies widely among the breeds, from dense and highly crimped, to long and hairlike. There is variation of wool type and quality even among members of the same flock, so wool classing is a step in the commercial processing of the fibre.
Suffolks are a medium wool, black-faced breed of meat sheep that make up 60% of the sheep population in the U.S.
Depending on breed, sheep show a range of heights and weights. Their rate of growth and mature weight is a heritable trait that is often selected for in breeding. Ewes typically weigh between 45 and 100 kilograms (100 and 220 lb), and rams between 45 and 160 kilograms (100 and 350 lb). When all deciduous teeth have erupted, the sheep has 20 teeth. Mature sheep have 32 teeth. As with other ruminants, the front teeth in the lower jaw bite against a hard, toothless pad in the upper jaw. These are used to pick off vegetation, then the rear teeth grind it before it is swallowed. There are eight lower front teeth in ruminants, but there is some disagreement as to whether these are eight incisors, or six incisors and two incisor-shaped canines. This means that the dental formula for sheep is either
0.0.3.3
4.0.3.3
or
0.0.3.3
3.1.3.3
There is a large diastema between the incisors and the molars.
In the first few years of life one can calculate the age of sheep from their front teeth, as a pair of milk teeth is replaced by larger adult teeth each year, the full set of eight adult front teeth being complete at about four years of age. The front teeth are then gradually lost as sheep age, making it harder for them to feed and hindering the health and productivity of the animal. For this reason, domestic sheep on normal pasture begin to slowly decline from four years on, and the life expectancy of a sheep is 10 to 12 years, though some sheep may live as long as 20 years.
Skull
Sheep have good hearing, and are sensitive to noise when being handled. Sheep have horizontal slit-shaped pupils, with excellent peripheral vision; with visual fields of about 270° to 320°, sheep can see behind themselves without turning their heads. Many breeds have only short hair on the face, and some have facial wool (if any) confined to the poll and or the area of the mandibular angle; the wide angles of peripheral vision apply to these breeds. A few breeds tend to have considerable wool on the face; for some individuals of these breeds, peripheral vision may be greatly reduced by "wool blindness", unless recently shorn about the face. Sheep have poor depth perception; shadows and dips in the ground may cause sheep to baulk. In general, sheep have a tendency to move out of the dark and into well-lit areas, and prefer to move uphill when disturbed. Sheep also have an excellent sense of smell, and, like all species of their genus, have scent glands just in front of the eyes, and interdigitally on the feet. The purpose of these glands is uncertain, but those on the face may be used in breeding behaviors. The foot glands might also be related to reproduction, but alternative functions, such as secretion of a waste product or a scent marker to help lost sheep find their flock, have also been proposed.
Comparison with goats
Sheep and goats are closely related: both are in the subfamily Caprinae. However, they are separate species, so hybrids rarely occur and are always infertile. A hybrid of a ewe and a buck (a male goat) is called a sheep-goat hybrid, known as geep. Visual differences between sheep and goats include the beard of goats and divided upper lip of sheep. Sheep tails also hang down, even when short or docked, while the short tails of goats are held upwards. Also, sheep breeds are often naturally polled (either in both sexes or just in the female), while naturally polled goats are rare (though many are polled artificially). Males of the two species differ in that buck goats acquire a unique and strong odor during the rut, whereas rams do not.
Breeds
The domestic sheep is a multi-purpose animal, and the more than 200 breeds now in existence were created to serve these diverse purposes. Some sources give a count of a thousand or more breeds, but these numbers cannot be verified, according to some sources. However, several hundred breeds of sheep have been identified by the Food and Agriculture Organization of the UN (FAO), with the estimated number varying somewhat from time to time: e.g. 863 breeds as of 1993, 1314 breeds as of 1995 and 1229 breeds as of 2006. (These numbers exclude extinct breeds, which are also tallied by the FAO.) For the purpose of such tallies, the FAO definition of a breed is "either a subspecific group of domestic livestock with definable and identifiable external characteristics that enable it to be separated by visual appraisal from other similarly defined groups within the same species or a group for which geographical and/or cultural separation from phenotypically similar groups has led to acceptance of its separate identity." Almost all sheep are classified as being best suited to furnishing a certain product: wool, meat, milk, hides, or a combination in a dual-purpose breed. Other features used when classifying sheep include face color (generally white or black), tail length, presence or lack of horns, and the topography for which the breed has been developed. This last point is especially stressed in the UK, where breeds are described as either upland (hill or mountain) or lowland breeds. A sheep may also be of a fat-tailed type, which is a dual-purpose sheep common in Africa and Asia with larger deposits of fat within and around its tail.
Breeds are often categorized by the type of their wool. Fine wool breeds are those that have wool of great crimp and density, which are preferred for textiles. Most of these were derived from Merino sheep, and the breed continues to dominate the world sheep industry. Downs breeds have wool between the extremes, and are typically fast-growing meat and ram breeds with dark faces. Some major medium wool breeds, such as the Corriedale, are dual-purpose crosses of long and fine-wooled breeds and were created for high-production commercial flocks. Long wool breeds are the largest of sheep, with long wool and a slow rate of growth. Long wool sheep are most valued for crossbreeding to improve the attributes of other sheep types. For example: the American Columbia breed was developed by crossing Lincoln rams (a long wool breed) with fine-wooled Rambouillet ewes.
Coarse or carpet wool sheep are those with a medium to long length wool of characteristic coarseness. Breeds traditionally used for carpet wool show great variability, but the chief requirement is a wool that will not break down under heavy use (as would that of the finer breeds). As the demand for carpet-quality wool declines, some breeders of this type of sheep are attempting to use a few of these traditional breeds for alternative purposes. Others have always been primarily meat-class sheep.
A minor class of sheep are the dairy breeds. Dual-purpose breeds that may primarily be meat or wool sheep are often used secondarily as milking animals, but there are a few breeds that are predominantly used for milking. These sheep produce a higher quantity of milk and have slightly longer lactation curves. In the quality of their milk, the fat and protein content percentages of dairy sheep vary from non-dairy breeds, but lactose content does not.
A last group of sheep breeds is that of fur or hair sheep, which do not grow wool at all. Hair sheep are similar to the early domesticated sheep kept before woolly breeds were developed, and are raised for meat and pelts. Some modern breeds of hair sheep, such as the Dorper, result from crosses between wool and hair breeds. For meat and hide producers, hair sheep are cheaper to keep, as they do not need shearing. Hair sheep are also more resistant to parasites and hot weather.
With the modern rise of corporate agribusiness and the decline of localized family farms, many breeds of sheep are in danger of extinction. The Rare Breeds Survival Trust of the UK lists 22 native breeds as having only 3,000 registered animals (each), and The Livestock Conservancy lists 14 as either "critical" or "threatened". Preferences for breeds with uniform characteristics and fast growth have pushed heritage (or heirloom) breeds to the margins of the sheep industry. Those that remain are maintained through the efforts of conservation organizations, breed registries, and individual farmers dedicated to their preservation.
Diet
Sheep are herbivorous mammals. Most breeds prefer to graze on grass and other short roughage, avoiding the taller woody parts of plants that goats readily consume. Both sheep and goats use their lips and tongues to select parts of the plant that are easier to digest or higher in nutrition. Sheep, however, graze well in monoculture pastures where most goats fare poorly.
Ruminant system of a sheep
Like all ruminants, sheep have a complex digestive system composed of four chambers, allowing them to break down cellulose from stems, leaves, and seed hulls into simpler carbohydrates. When sheep graze, vegetation is chewed into a mass called a bolus, which is then passed into the rumen, via the reticulum. The rumen is a 19- to 38-liter (5 to 10 gallon) organ in which feed is fermented. The fermenting organisms include bacteria, fungi, and protozoa. (Other important rumen organisms include some archaea, which produce methane from carbon dioxide.) The bolus is periodically regurgitated back to the mouth as cud for additional chewing and salivation. After fermentation in the rumen, feed passes into the reticulum and the omasum; special feeds such as grains may bypass the rumen altogether. After the first three chambers, food moves into the abomasum for final digestion before processing by the intestines. The abomasum is the only one of the four chambers analogous to the human stomach, and is sometimes called the "true stomach".
Other than forage, the other staple feed for sheep is hay, often during the winter months. The ability to thrive solely on pasture (even without hay) varies with breed, but all sheep can survive on this diet. Also included in some sheep's diets are minerals, either in a trace mix or in licks. Feed provided to sheep must be specially formulated, as most cattle, poultry, pig, and even some goat feeds contain levels of copper that are lethal to sheep. The same danger applies to mineral supplements such as salt licks.
Grazing behavior
Sheep follow a diurnal pattern of activity, feeding from dawn to dusk, stopping sporadically to rest and chew their cud. Ideal pasture for sheep is not lawnlike grass, but an array of grasses, legumes and forbs. Types of land where sheep are raised vary widely, from pastures that are seeded and improved intentionally to rough, native lands. Common plants toxic to sheep are present in most of the world, and include (but are not limited to) cherry, some oaks and acorns, tomato, yew, rhubarb, potato, and rhododendron.
Effects on pasture
Sheep are largely grazing herbivores, unlike browsing animals such as goats and deer that prefer taller foliage. With a much narrower face, sheep crop plants very close to the ground and can overgraze a pasture much faster than cattle. For this reason, many shepherds use managed intensive rotational grazing, where a flock is rotated through multiple pastures, giving plants time to recover. Paradoxically, sheep can both cause and solve the spread of invasive plant species. By disturbing the natural state of pasture, sheep and other livestock can pave the way for invasive plants. However, sheep also prefer to eat invasives such as cheatgrass, leafy spurge, kudzu and spotted knapweed over native species such as sagebrush, making grazing sheep effective for conservation grazing. Research conducted in Imperial County, California compared lamb grazing with herbicides for weed control in seedling alfalfa fields. Three trials demonstrated that grazing lambs were just as effective as herbicides in controlling winter weeds. Entomologists also compared grazing lambs to insecticides for insect control in winter alfalfa. In this trial, lambs provided insect control as effectively as insecticides.
Behavior
Sheep are flock animals and strongly gregarious; much sheep behavior can be understood on the basis of these tendencies. The dominance hierarchy of sheep and their natural inclination to follow a leader to new pastures were the pivotal factors in sheep being one of the first domesticated livestock species. Furthermore, in contrast to the red deer and gazelle (two other ungulates of primary importance to meat production in prehistoric times), sheep do not defend territories although they do form home ranges. All sheep have a tendency to congregate close to other members of a flock, although this behavior varies with breed, and sheep can become stressed when separated from their flock members. During flocking, sheep have a strong tendency to follow, and a leader may simply be the first individual to move. Relationships in flocks tend to be closest among related sheep: in mixed-breed flocks, subgroups of the same breed tend to form, and a ewe and her direct descendants often move as a unit within large flocks. Sheep can become hefted to one particular local pasture (heft) so they do not roam freely in unfenced landscapes. Lambs learn the heft from ewes and if whole flocks are culled it must be retaught to the replacement animals.
Flock behaviour in sheep is generally only exhibited in groups of four or more sheep; fewer sheep may not react as expected when alone or with few other sheep. Being a prey species, the primary defense mechanism of sheep is to flee from danger when their flight zone is entered. Cornered sheep may charge and butt, or threaten by hoof stamping and adopting an aggressive posture. This is particularly true for ewes with newborn lambs.
In regions where sheep have no natural predators, none of the native breeds of sheep exhibit a strong flocking behavior.
Herding
Farmers exploit flocking behavior to keep sheep together on unfenced pastures such as hill farming, and to move them more easily. For this purpose shepherds may use herding dogs in this effort, with a highly bred herding ability. Sheep are food-oriented, and association of humans with regular feeding often results in sheep soliciting people for food. Those who are moving sheep may exploit this behavior by leading sheep with buckets of feed.
Dominance hierarchy
Sheep establish a dominance hierarchy through fighting, threats and competitiveness. Dominant animals are inclined to be more aggressive with other sheep, and usually feed first at troughs. Primarily among rams, horn size is a factor in the flock hierarchy. Rams with different size horns may be less inclined to fight to establish the dominance order, while rams with similarly sized horns are more so. Merinos have an almost linear hierarchy whereas there is a less rigid structure in Border Leicesters when a competitive feeding situation arises.
In sheep, position in a moving flock is highly correlated with social dominance, but there is no definitive study to show consistent voluntary leadership by an individual sheep.
Intelligence and learning ability
Sheep are frequently thought of as unintelligent animals. Their flocking behavior and quickness to flee and panic can make shepherding a difficult endeavor for the uninitiated. Despite these perceptions, a University of Illinois monograph on sheep reported their intelligence to be just below that of pigs and on par with that of cattle. Sheep can recognize individual human and ovine faces and remember them for years; they can remember 50 other different sheep faces for over two years; they can recognize and are attracted to individual sheep and humans by their faces, as they possess similar specialized neural systems in the temporal and frontal lobes of their brains to humans and have a greater involvement of the right brain hemisphere. In addition to long-term facial recognition of individuals, sheep can also differentiate emotional states through facial characteristics.[68][69] If worked with patiently, sheep may learn their names, and many sheep are trained to be led by halter for showing and other purposes. Sheep have also responded well to clicker training. Sheep have been used as pack animals; Tibetan nomads distribute baggage equally throughout a flock as it is herded between living sites.
It has been reported that some sheep have apparently shown problem-solving abilities; a flock in West Yorkshire, England allegedly found a way to get over cattle grids by rolling on their backs, although documentation of this has relied on anecdotal accounts.
Vocalisations
Sounds made by domestic sheep include bleats, grunts, rumbles and snorts. Bleating ("baaing") is used mostly for contact communication, especially between dam and lambs, but also at times between other flock members. The bleats of individual sheep are distinctive, enabling the ewe and her lambs to recognize each other's vocalizations. Vocal communication between lambs and their dam declines to a very low level within several weeks after parturition. A variety of bleats may be heard, depending on sheep age and circumstances. Apart from contact communication, bleating may signal distress, frustration or impatience; however, sheep are usually silent when in pain. Isolation commonly prompts bleating by sheep. Pregnant ewes may grunt when in labor. Rumbling sounds are made by the ram during courting; somewhat similar rumbling sounds may be made by the ewe, especially when with her neonate lambs. A snort (explosive exhalation through the nostrils) may signal aggression or a warning, and is often elicited from startled sheep.
Lamb
In sheep breeds lacking facial wool, the visual field is wide. In 10 sheep (Cambridge, Lleyn and Welsh Mountain breeds, which lack facial wool), the visual field ranged from 298° to 325°, averaging 313.1°, with binocular overlap ranging from 44.5° to 74°, averaging 61.7°. In some breeds, unshorn facial wool can limit the visual field; in some individuals, this may be enough to cause "wool blindness". In 60 Merinos, visual fields ranged from 219.1° to 303.0°, averaging 269.9°, and the binocular field ranged from 8.9° to 77.7°, averaging 47.5°; 36% of the measurements were limited by wool, although photographs of the experiments indicate that only limited facial wool regrowth had occurred since shearing. In addition to facial wool (in some breeds), visual field limitations can include ears and (in some breeds) horns, so the visual field can be extended by tilting the head. Sheep eyes exhibit very low hyperopia and little astigmatism. Such visual characteristics are likely to produce a well-focused retinal image of objects in both the middle and long distance. Because sheep eyes have no accommodation, one might expect the image of very near objects to be blurred, but a rather clear near image could be provided by the tapetum and large retinal image of the sheep's eye, and adequate close vision may occur at muzzle length. Good depth perception, inferred from the sheep's sure-footedness, was confirmed in "visual cliff" experiments; behavioral responses indicating depth perception are seen in lambs at one day old. Sheep are thought to have colour vision, and can distinguish between a variety of colours: black, red, brown, green, yellow and white. Sight is a vital part of sheep communication, and when grazing, they maintain visual contact with each other. Each sheep lifts its head upwards to check the position of other sheep in the flock. This constant monitoring is probably what keeps the sheep in a flock as they move along grazing. Sheep become stressed when isolated; this stress is reduced if they are provided with a mirror, indicating that the sight of other sheep reduces stress.
Taste is the most important sense in sheep, establishing forage preferences, with sweet and sour plants being preferred and bitter plants being more commonly rejected. Touch and sight are also important in relation to specific plant characteristics, such as succulence and growth form.
The ram uses his vomeronasal organ (sometimes called the Jacobson's organ) to sense the pheromones of ewes and detect when they are in estrus. The ewe uses her vomeronasal organ for early recognition of her neonate lamb.
Reproduction
Sheep follow a similar reproductive strategy to other herd animals. A group of ewes is generally mated by a single ram, who has either been chosen by a breeder or (in feral populations) has established dominance through physical contest with other rams. Most sheep are seasonal breeders, although some are able to breed year-round. Ewes generally reach sexual maturity at six to eight months old, and rams generally at four to six months. However, there are exceptions. For example, Finnsheep ewe lambs may reach puberty as early as 3 to 4 months, and Merino ewes sometimes reach puberty at 18 to 20 months. Ewes have estrus cycles about every 17 days, during which they emit a scent and indicate readiness through physical displays towards rams.
In feral sheep, rams may fight during the rut to determine which individuals may mate with ewes. Rams, especially unfamiliar ones, will also fight outside the breeding period to establish dominance; rams can kill one another if allowed to mix freely. During the rut, even usually friendly rams may become aggressive towards humans due to increases in their hormone levels.
After mating, sheep have a gestation period of about five months, and normal labor takes one to three hours. Although some breeds regularly throw larger litters of lambs, most produce single or twin lambs. During or soon after labor, ewes and lambs may be confined to small lambing jugs, small pens designed to aid both careful observation of ewes and to cement the bond between them and their lambs.
A lamb's first steps
Ovine obstetrics can be problematic. By selectively breeding ewes that produce multiple offspring with higher birth weights for generations, sheep producers have inadvertently caused some domestic sheep to have difficulty lambing; balancing ease of lambing with high productivity is one of the dilemmas of sheep breeding. In the case of any such problems, those present at lambing may assist the ewe by extracting or repositioning lambs. After the birth, ewes ideally break the amniotic sac (if it is not broken during labor), and begin licking clean the lamb. Most lambs will begin standing within an hour of birth. In normal situations, lambs nurse after standing, receiving vital colostrum milk. Lambs that either fail to nurse or are rejected by the ewe require help to survive, such as bottle-feeding or fostering by another ewe.
Most lambs begin life being born outdoors. After lambs are several weeks old, lamb marking (ear tagging, docking, mulesing, and castrating) is carried out. Vaccinations are usually carried out at this point as well. Ear tags with numbers are attached, or ear marks are applied, for ease of later identification of sheep. Docking and castration are commonly done after 24 hours (to avoid interference with maternal bonding and consumption of colostrum) and are often done not later than one week after birth, to minimize pain, stress, recovery time and complications. The first course of vaccinations (commonly anti-clostridial) is commonly given at an age of about 10 to 12 weeks; i.e. when the concentration of maternal antibodies passively acquired via colostrum is expected to have fallen low enough to permit development of active immunity. Ewes are often revaccinated annually about 3 weeks before lambing, to provide high antibody concentrations in colostrum during the first several hours after lambing. Ram lambs that will either be slaughtered or separated from ewes before sexual maturity are not usually castrated. Objections to all these procedures have been raised by animal rights groups, but farmers defend them by saying they save money, and inflict only temporary pain.
Homosexuality
Sheep are the only species of mammal except for humans which exhibits exclusive homosexual behavior. About 10% of rams refuse to mate with ewes but readily mate with other rams, and thirty percent of all rams demonstrate at least some homosexual behavior. Additionally, a small number of females that were accompanied by a male fetus in utero (i.e. as fraternal twins) are freemartins (female animals that are behaviorally masculine and lack functioning ovaries).
Health
Sheep may fall victim to poisons, infectious diseases, and physical injuries. As a prey species, a sheep's system is adapted to hide the obvious signs of illness, to prevent being targeted by predators. However, some signs of ill health are obvious, with sick sheep eating little, vocalizing excessively, and being generally listless. Throughout history, much of the money and labor of sheep husbandry has aimed to prevent sheep ailments. Historically, shepherds often created remedies by experimentation on the farm. In some developed countries, including the United States, sheep lack the economic importance for drug companies to perform expensive clinical trials required to approve more than a relatively limited number of drugs for ovine use. However, extra-label drug use in sheep production is permitted in many jurisdictions, subject to certain restrictions. In the US, for example, regulations governing extra-label drug use in animals are found in 21 CFR (Code of Federal Regulations) Part 530. In the 20th and 21st centuries, a minority of sheep owners have turned to alternative treatments such as homeopathy, herbalism and even traditional Chinese medicine to treat sheep veterinary problems. Despite some favorable anecdotal evidence, the effectiveness of alternative veterinary medicine has been met with skepticism in scientific journals. The need for traditional anti-parasite drugs and antibiotics is widespread, and is the main impediment to certified organic farming with sheep.
Many breeders take a variety of preventive measures to ward off problems. The first is to ensure all sheep are healthy when purchased. Many buyers avoid outlets known to be clearing houses for animals culled from healthy flocks as either sick or simply inferior. This can also mean maintaining a closed flock, and quarantining new sheep for a month. Two fundamental preventive programs are maintaining good nutrition and reducing stress in the sheep. Restraint, isolation, loud noises, novel situations, pain, heat, extreme cold, fatigue and other stressors can lead to secretion of cortisol, a stress hormone, in amounts that may indicate welfare problems. Excessive stress can compromise the immune system. "Shipping fever" (pneumonic mannheimiosis, formerly called pasteurellosis) is a disease of particular concern, that can occur as a result of stress, notably during transport and (or) handling. Pain, fear and several other stressors can cause secretion of epinephrine (adrenaline). Considerable epinephrine secretion in the final days before slaughter can adversely affect meat quality (by causing glycogenolysis, removing the substrate for normal post-slaughter acidification of meat) and result in meat becoming more susceptible to colonization by spoilage bacteria. Because of such issues, low-stress handling is essential in sheep management. Avoiding poisoning is also important; common poisons are pesticide sprays, inorganic fertilizer, motor oil, as well as radiator coolant containing ethylene glycol.
Common forms of preventive medication for sheep are vaccinations and treatments for parasites. Both external and internal parasites are the most prevalent malady in sheep, and are either fatal, or reduce the productivity of flocks. Worms are the most common internal parasites. They are ingested during grazing, incubate within the sheep, and are expelled through the digestive system (beginning the cycle again). Oral anti-parasitic medicines, known as drenches, are given to a flock to treat worms, sometimes after worm eggs in the feces has been counted to assess infestation levels. Afterwards, sheep may be moved to a new pasture to avoid ingesting the same parasites. External sheep parasites include: lice (for different parts of the body), sheep keds, nose bots, sheep itch mites, and maggots. Keds are blood-sucking parasites that cause general malnutrition and decreased productivity, but are not fatal. Maggots are those of the bot fly and the blow-fly, commonly Lucilia sericata or its relative L. cuprina. Fly maggots cause the extremely destructive condition of flystrike. Flies lay their eggs in wounds or wet, manure-soiled wool; when the maggots hatch they burrow into a sheep's flesh, eventually causing death if untreated. In addition to other treatments, crutching (shearing wool from a sheep's rump) is a common preventive method. Some countries allow mulesing, a practice that involves stripping away the skin on the rump to prevent fly-strike, normally performed when the sheep is a lamb. Nose bots are fly larvae that inhabit a sheep's sinuses, causing breathing difficulties and discomfort. Common signs are a discharge from the nasal passage, sneezing, and frantic movement such as head shaking. External parasites may be controlled through the use of backliners, sprays or immersive sheep dips.
A wide array of bacterial and viral diseases affect sheep. Diseases of the hoof, such as foot rot and foot scald may occur, and are treated with footbaths and other remedies. Foot rot is present in over 97% of flocks in the UK. These painful conditions cause lameness and hinder feeding. Ovine Johne's disease is a wasting disease that affects young sheep. Bluetongue disease is an insect-borne illness causing fever and inflammation of the mucous membranes. Ovine rinderpest (or peste des petits ruminants) is a highly contagious and often fatal viral disease affecting sheep and goats. Sheep may also be affected by primary or secondary photosensitization. Tetanus can also afflict sheep through wounds from shearing, docking, castration, or vaccination. The organism also can be introduced into the reproductive tract by unsanitary humans who assist ewes during lambing.
A few sheep conditions are transmissible to humans. Orf (also known as scabby mouth, contagious ecthyma or soremouth) is a skin disease leaving lesions that is transmitted through skin-to-skin contact. Cutaneous anthrax is also called woolsorter's disease, as the spores can be transmitted in unwashed wool. More seriously, the organisms that can cause spontaneous enzootic abortion in sheep are easily transmitted to pregnant women. Also of concern are the prion disease scrapie and the virus that causes foot-and-mouth disease (FMD), as both can devastate flocks. The latter poses a slight risk to humans. During the 2001 FMD pandemic in the UK, hundreds of sheep were culled and some rare British breeds were at risk of extinction due to this.
Of the 600,300 sheep lost to the US economy in 2004, 37.3% were lost to predators, while 26.5% were lost to some form of disease. Poisoning accounted for 1.7% of non-productive deaths.
Predators
A lamb being attacked by coyotes with a bite to the throat
Other than parasites and disease, predation is a threat to sheep and the profitability of sheep raising. Sheep have little ability to defend themselves, compared with other species kept as livestock. Even if sheep survive an attack, they may die from their injuries or simply from panic. However, the impact of predation varies dramatically with region. In Africa, Australia, the Americas, and parts of Europe and Asia predators are a serious problem. In the United States, for instance, over one third of sheep deaths in 2004 were caused by predation. In contrast, other nations are virtually devoid of sheep predators, particularly islands known for extensive sheep husbandry. Worldwide, canids—including the domestic dog—are responsible for most sheep deaths. Other animals that occasionally prey on sheep include: felines, bears, birds of prey, ravens and feral hogs.
Sheep producers have used a wide variety of measures to combat predation. Pre-modern shepherds used their own presence, livestock guardian dogs, and protective structures such as barns and fencing. Fencing (both regular and electric), penning sheep at night and lambing indoors all continue to be widely used. More modern shepherds used guns, traps, and poisons to kill predators, causing significant decreases in predator populations. In the wake of the environmental and conservation movements, the use of these methods now usually falls under the purview of specially designated government agencies in most developed countries.
The 1970s saw a resurgence in the use of livestock guardian dogs and the development of new methods of predator control by sheep producers, many of them non-lethal. Donkeys and guard llamas have been used since the 1980s in sheep operations, using the same basic principle as livestock guardian dogs. Interspecific pasturing, usually with larger livestock such as cattle or horses, may help to deter predators, even if such species do not actively guard sheep. In addition to animal guardians, contemporary sheep operations may use non-lethal predator deterrents such as motion-activated lights and noisy alarms.
Economic importance
Main article: Agricultural economics
Global sheep stock
in 2019
Number in millions
1. China163.5 (13.19%)
2. India74.3 (5.99%)
3. Australia65.8 (5.31%)
4. Nigeria46.9 (3.78%)
5. Iran41.3 (3.33%)
6. Sudan40.9 (3.3%)
7. Chad35.9 (2.9%)
8. Turkey35.2 (2.84%)
9. United Kingdom33.6 (2.71%)
10. Mongolia32.3 (2.61%)
World total1,239.8
Source: UN Food and Agriculture Organization
Sheep are an important part of the global agricultural economy. However, their once vital status has been largely replaced by other livestock species, especially the pig, chicken, and cow. China, Australia, India, and Iran have the largest modern flocks, and serve both local and exportation needs for wool and mutton. Other countries such as New Zealand have smaller flocks but retain a large international economic impact due to their export of sheep products. Sheep also play a major role in many local economies, which may be niche markets focused on organic or sustainable agriculture and local food customers. Especially in developing countries, such flocks may be a part of subsistence agriculture rather than a system of trade. Sheep themselves may be a medium of trade in barter economies.
Domestic sheep provide a wide array of raw materials. Wool was one of the first textiles, although in the late 20th century wool prices began to fall dramatically as the result of the popularity and cheap prices for synthetic fabrics. For many sheep owners, the cost of shearing is greater than the possible profit from the fleece, making subsisting on wool production alone practically impossible without farm subsidies. Fleeces are used as material in making alternative products such as wool insulation. In the 21st century, the sale of meat is the most profitable enterprise in the sheep industry, even though far less sheep meat is consumed than chicken, pork or beef.
Sheepskin is likewise used for making clothes, footwear, rugs, and other products. Byproducts from the slaughter of sheep are also of value: sheep tallow can be used in candle and soap making, sheep bone and cartilage has been used to furnish carved items such as dice and buttons as well as rendered glue and gelatin. Sheep intestine can be formed into sausage casings, and lamb intestine has been formed into surgical sutures, as well as strings for musical instruments and tennis rackets. Sheep droppings, which are high in cellulose, have even been sterilized and mixed with traditional pulp materials to make paper. Of all sheep byproducts, perhaps the most valuable is lanolin: the waterproof, fatty substance found naturally in sheep's wool and used as a base for innumerable cosmetics and other products.
Some farmers who keep sheep also make a profit from live sheep. Providing lambs for youth programs such as 4-H and competition at agricultural shows is often a dependable avenue for the sale of sheep. Farmers may also choose to focus on a particular breed of sheep in order to sell registered purebred animals, as well as provide a ram rental service for breeding. A new option for deriving profit from live sheep is the rental of flocks for grazing; these "mowing services" are hired in order to keep unwanted vegetation down in public spaces and to lessen fire hazard.
Despite the falling demand and price for sheep products in many markets, sheep have distinct economic advantages when compared with other livestock. They do not require expensive housing, such as that used in the intensive farming of chickens or pigs. They are an efficient use of land; roughly six sheep can be kept on the amount that would suffice for a single cow or horse. Sheep can also consume plants, such as noxious weeds, that most other animals will not touch, and produce more young at a faster rate. Also, in contrast to most livestock species, the cost of raising sheep is not necessarily tied to the price of feed crops such as grain, soybeans and corn. Combined with the lower cost of quality sheep, all these factors combine to equal a lower overhead for sheep producers, thus entailing a higher profitability potential for the small farmer. Sheep are especially beneficial for independent producers, including family farms with limited resources, as the sheep industry is one of the few types of animal agriculture that has not been vertically integrated by agribusiness. However, small flocks, from 10 to 50 ewes, often are not profitable because they tend to be poorly managed. The primary reason is that mechanization is not feasible, so return per hour of labor is not maximized. Small farm flocks generally are used simply to control weeds on irrigation ditches or maintained as a hobby.
Shoulder of lamb
Sheep meat and milk were one of the earliest staple proteins consumed by human civilization after the transition from hunting and gathering to agriculture. Sheep meat prepared for food is known as either mutton or lamb, and approximately 540 million sheep are slaughtered each year for meat worldwide. "Mutton" is derived from the Old French moton, which was the word for sheep used by the Anglo-Norman rulers of much of the British Isles in the Middle Ages. This became the name for sheep meat in English, while the Old English word sceap was kept for the live animal. Throughout modern history, "mutton" has been limited to the meat of mature sheep usually at least two years of age; "lamb" is used for that of immature sheep less than a year.
In the 21st century, the nations with the highest consumption of sheep meat are the Arab states of the Persian Gulf, New Zealand, Australia, Greece, Uruguay, the United Kingdom and Ireland. These countries eat 14–40 lbs (3–18 kg) of sheep meat per capita, per annum. Sheep meat is also popular in France, Africa (especially the Arab world), the Caribbean, the rest of the Middle East, India, and parts of China. This often reflects a history of sheep production. In these countries in particular, dishes comprising alternative cuts and offal may be popular or traditional. Sheep testicles—called animelles or lamb fries—are considered a delicacy in many parts of the world. Perhaps the most unusual dish of sheep meat is the Scottish haggis, composed of various sheep innards cooked along with oatmeal and chopped onions inside its stomach. In comparison, countries such as the U.S. consume only a pound or less (under 0.5 kg), with Americans eating 50 pounds (22 kg) of pork and 65 pounds (29 kg) of beef. In addition, such countries rarely eat mutton, and may favor the more expensive cuts of lamb: mostly lamb chops and leg of lamb.
Though sheep's milk may be drunk rarely in fresh form, today it is used predominantly in cheese and yogurt making. Sheep have only two teats, and produce a far smaller volume of milk than cows. However, as sheep's milk contains far more fat, solids, and minerals than cow's milk, it is ideal for the cheese-making process. It also resists contamination during cooling better because of its much higher calcium content. Well-known cheeses made from sheep milk include the feta of Bulgaria and Greece, Roquefort of France, Manchego from Spain, the pecorino romano (the Italian word for "sheep" is pecore) and ricotta of Italy. Yogurts, especially some forms of strained yogurt, may also be made from sheep milk. Many of these products are now often made with cow's milk, especially when produced outside their country of origin. Sheep milk contains 4.8% lactose, which may affect those who are intolerant.
As with other domestic animals, the meat of uncastrated males is inferior in quality, especially as they grow. A "bucky" lamb is a lamb which was not castrated early enough, or which was castrated improperly (resulting in one testicle being retained). These lambs are worth less at market.
In science
Sheep are generally too large and reproduce too slowly to make ideal research subjects, and thus are not a common model organism. They have, however, played an influential role in some fields of science. In particular, the Roslin Institute of Edinburgh, Scotland used sheep for genetics research that produced groundbreaking results. In 1995, two ewes named Megan and Morag were the first mammals cloned from differentiated cells, also referred to as gynomerogony. A year later, a Finnish Dorset sheep named Dolly, dubbed "the world's most famous sheep" in Scientific American, was the first mammal to be cloned from an adult somatic cell. Following this, Polly and Molly were the first mammals to be simultaneously cloned and transgenic.
As of 2008, the sheep genome has not been fully sequenced, although a detailed genetic map has been published, and a draft version of the complete genome produced by assembling sheep DNA sequences using information given by the genomes of other mammals. In 2012, a transgenic sheep named "Peng Peng" was cloned by Chinese scientists, who spliced his genes with that of a roundworm (C. elegans) in order to increase production of fats healthier for human consumption.
In the study of natural selection, the population of Soay sheep that remain on the island of Hirta have been used to explore the relation of body size and coloration to reproductive success. Soay sheep come in several colors, and researchers investigated why the larger, darker sheep were in decline; this occurrence contradicted the rule of thumb that larger members of a population tend to be more successful reproductively. The feral Soays on Hirta are especially useful subjects because they are isolated.
Domestic sheep are sometimes used in medical research, particularly for researching cardiovascular physiology, in areas such as hypertension and heart failure. Pregnant sheep are also a useful model for human pregnancy, and have been used to investigate the effects on fetal development of malnutrition and hypoxia. In behavioral sciences, sheep have been used in isolated cases for the study of facial recognition, as their mental process of recognition is qualitatively similar to humans.
Cultural impact
Sheep have had a strong presence in many cultures, especially in areas where they form the most common type of livestock. In the English language, to call someone a sheep or ovine may allude that they are timid and easily led. In contradiction to this image, male sheep are often used as symbols of virility and power; the logos of the Los Angeles Rams football team and the Dodge Ram pickup truck allude to males of the bighorn sheep, Ovis canadensis.
Counting sheep is popularly said to be an aid to sleep, and some ancient systems of counting sheep persist today. Sheep also enter in colloquial sayings and idiom frequently with such phrases as "black sheep". To call an individual a black sheep implies that they are an odd or disreputable member of a group. This usage derives from the recessive trait that causes an occasional black lamb to be born into an entirely white flock. These black sheep were considered undesirable by shepherds, as black wool is not as commercially viable as white wool. Citizens who accept overbearing governments have been referred to by the Portmanteau neologism of sheeple. Somewhat differently, the adjective "sheepish" is also used to describe embarrassment.
In heraldry
In British heraldry, sheep appear in the form of rams, sheep proper and lambs. These are distinguished by the ram being depicted with horns and a tail, the sheep with neither and the lamb with its tail only. A further variant of the lamb, termed the Paschal lamb, is depicted as carrying a Christian cross and with a halo over its head. Rams' heads, portrayed without a neck and facing the viewer, are also found in British armories. The fleece, depicted as an entire sheepskin carried by a ring around its midsection, originally became known through its use in the arms of the Order of the Golden Fleece and was later adopted by towns and individuals with connections to the wool industry. A sheep on a blue field is depicted on the greater/royal arms of the king of Denmark to represent the Faroe Islands. In 2004 a modernized arms has been adopted by the Faroe Islands, which based on a 15th century coat of arms.
Religion and folklore
In antiquity, symbolism involving sheep cropped up in religions in the ancient Near East, the Mideast, and the Mediterranean area: Çatalhöyük, ancient Egyptian religion, the Cana'anite and Phoenician tradition, Judaism, Greek religion, and others. Religious symbolism and ritual involving sheep began with some of the first known faiths: Skulls of rams (along with bulls) occupied central placement in shrines at the Çatalhöyük settlement in 8,000 BCE. In Ancient Egyptian religion, the ram was the symbol of several gods: Khnum, Heryshaf and Amun (in his incarnation as a god of fertility). Other deities occasionally shown with ram features include the goddess Ishtar, the Phoenician god Baal-Hamon, and the Babylonian god Ea-Oannes. In Madagascar, sheep were not eaten as they were believed to be incarnations of the souls of ancestors.
There are many ancient Greek references to sheep: that of Chrysomallos, the golden-fleeced ram, continuing to be told through into the modern era. Astrologically, Aries, the ram, is the first sign of the classical Greek zodiac, and the sheep is the eighth of the twelve animals associated with the 12-year cycle of in the Chinese zodiac, related to the Chinese calendar. It is said in Chinese traditions that Hou ji sacrificed sheep. Mongolia, shagai are an ancient form of dice made from the cuboid bones of sheep that are often used for fortunetelling purposes.
Sheep play an important role in all the Abrahamic faiths; Abraham, Isaac, Jacob, Moses, and King David were all shepherds. According to the Biblical story of the Binding of Isaac, a ram is sacrificed as a substitute for Isaac after an angel stays Abraham's hand (in the Islamic tradition, Abraham was about to sacrifice Ishmael). Eid al-Adha is a major annual festival in Islam in which sheep (or other animals) are sacrificed in remembrance of this act. Sheep are occasionally sacrificed to commemorate important secular events in Islamic cultures. Greeks and Romans sacrificed sheep regularly in religious practice, and Judaism once sacrificed sheep as a Korban (sacrifice), such as the Passover lamb. Ovine symbols—such as the ceremonial blowing of a shofar—still find a presence in modern Judaic traditions.
Collectively, followers of Christianity are often referred to as a flock, with Christ as the Good Shepherd, and sheep are an element in the Christian iconography of the birth of Jesus. Some Christian saints are considered patrons of shepherds, and even of sheep themselves. Christ is also portrayed as the Sacrificial lamb of God (Agnus Dei) and Easter celebrations in Greece and Romania traditionally feature a meal of Paschal lamb. A church leader is often called the pastor, which is derived from the Latin word for shepherd. In many western Christian traditions bishops carry a staff, which also serves as a symbol of the episcopal office, known as a crosier, which is modeled on the shepherd's crook.
Sheep are key symbols in fables and nursery rhymes like The Wolf in Sheep's Clothing, Little Bo Peep, Baa, Baa, Black Sheep, and Mary Had a Little Lamb; novels such as George Orwell's Animal Farm and Haruki Murakami's A Wild Sheep Chase; songs such as Bach's Sheep may safely graze (Schafe können sicher weiden) and Pink Floyd's "Sheep", and poems like William Blake's "The Lamb".
The Mursi (also called Murzu) is the most popular tribe in the southwestern Ethiopia lower Omo Valley, 100 km north of Kenyan. They are estimated to 10 000 people and live in the Mago National Park, established in 1979. Due to the climate, they move twice a year between the winter and summer months. They herd cattle and grow crops along the banks of the Omo River. The Mursi are sedentary rather than nomadic. Their language belongs to the Nilo-Saharan linguistic family.Very few Mursi people speak Amharic, the official Ethiopian language. Although a small percentage of the Mursi tribe are Christians, most still practice animism. Mursi women wear giant lip plate, a sign of beauty, like in Suri tribe, and also a prime attraction for tourists which help to sustain a view of them, in guidebooks and travel articles, as an untouched people, living in one of the last wildernesses of Africa. When they are ready to marry, teenagers start to make a hole in the lower lip with a wood stick.
It will be kept for one night, and is removed to put a bigger one. This is very painful at this time. Few months after, the lip plate has its full size, and the girl is seen as beautiful by the men. The lip plate is made of wood or terracotta. They have to remove the lower incisors to let some space for the disc. Sometimes the lip is broken by the pressure of the plate. This is a big problem for the girl because men will consider her as ugly, she won't be able to marry anyone in the tribe apart the old men or the sick people. Women and men are shaved because they hate hairiness. Both like to make scarifications on their bodies. Women as a beauty sign, men after killing animals or ennemies as competition for grazing land has led to tribal conflicts.
The Mursi men have a reputation for being aggressive and are famous for their stick fighting ceremony called donga. The winner of the donga will be able to select the girl of his choice to have relations with if she agrees. Similar to the Surma tribe, the Mursi tribe commonly drink a mixture of blood and milk. Over the past few decades they and their neighbours have faced growing threats to their livelihoods cause the Ethiopian government officials have been actively evicting Mursi people from the Omo National Park, without any compensation to rent their land to foreign investors. Drought has made it difficult for many families to feed themselves by means of their traditional mix of subsistence activities. The establishment of hunting concessions has added to the pressure on scarce ressources.
© Eric Lafforgue
Sheep (pl.: sheep) or domestic sheep (Ovis aries) are a domesticated, ruminant mammal typically kept as livestock. Although the term sheep can apply to other species in the genus Ovis, in everyday usage it almost always refers to domesticated sheep. Like all ruminants, sheep are members of the order Artiodactyla, the even-toed ungulates. Numbering a little over one billion, domestic sheep are also the most numerous species of sheep. An adult female is referred to as a ewe (/juː/ yoo), an intact male as a ram, occasionally a tup, a castrated male as a wether, and a young sheep as a lamb.
Sheep are most likely descended from the wild mouflon of Europe and Asia, with Iran being a geographic envelope of the domestication center. One of the earliest animals to be domesticated for agricultural purposes, sheep are raised for fleeces, meat (lamb, hogget or mutton) and milk. A sheep's wool is the most widely used animal fiber, and is usually harvested by shearing. In Commonwealth countries, ovine meat is called lamb when from younger animals and mutton when from older ones; in the United States, meat from both older and younger animals is usually called lamb. Sheep continue to be important for wool and meat today, and are also occasionally raised for pelts, as dairy animals, or as model organisms for science.
Sheep husbandry is practised throughout the majority of the inhabited world, and has been fundamental to many civilizations. In the modern era, Australia, New Zealand, the southern and central South American nations, and the British Isles are most closely associated with sheep production.
There is a large lexicon of unique terms for sheep husbandry which vary considerably by region and dialect. Use of the word sheep began in Middle English as a derivation of the Old English word scēap. A group of sheep is called a flock. Many other specific terms for the various life stages of sheep exist, generally related to lambing, shearing, and age.
Being a key animal in the history of farming, sheep have a deeply entrenched place in human culture, and are represented in much modern language and symbolism. As livestock, sheep are most often associated with pastoral, Arcadian imagery. Sheep figure in many mythologies—such as the Golden Fleece—and major religions, especially the Abrahamic traditions. In both ancient and modern religious ritual, sheep are used as sacrificial animals.
History
Main article: History of the domestic sheep
The exact line of descent from wild ancestors to domestic sheep is unclear. The most common hypothesis states that Ovis aries is descended from the Asiatic (O. gmelini) species of mouflon; the European mouflon (Ovis aries musimon) is a direct descendant of this population. Sheep were among the first animals to be domesticated by humankind (although the domestication of dogs probably took place 10 to 20 thousand years earlier); the domestication date is estimated to fall between 11,000 and 9000 B.C in Mesopotamia and possibly around 7000 BC in Mehrgarh in the Indus Valley. The rearing of sheep for secondary products, and the resulting breed development, began in either southwest Asia or western Europe. Initially, sheep were kept solely for meat, milk and skins. Archaeological evidence from statuary found at sites in Iran suggests that selection for woolly sheep may have begun around 6000 BC, and the earliest woven wool garments have been dated to two to three thousand years later.
Sheep husbandry spread quickly in Europe. Excavations show that in about 6000 BC, during the Neolithic period of prehistory, the Castelnovien people, living around Châteauneuf-les-Martigues near present-day Marseille in the south of France, were among the first in Europe to keep domestic sheep. Practically from its inception, ancient Greek civilization relied on sheep as primary livestock, and were even said to name individual animals. Ancient Romans kept sheep on a wide scale, and were an important agent in the spread of sheep raising. Pliny the Elder, in his Natural History (Naturalis Historia), speaks at length about sheep and wool. European colonists spread the practice to the New World from 1493 onwards.
Characteristics
Domestic sheep are relatively small ruminants, usually with a crimped hair called wool and often with horns forming a lateral spiral. They differ from their wild relatives and ancestors in several respects, having become uniquely neotenic as a result of selective breeding by humans. A few primitive breeds of sheep retain some of the characteristics of their wild cousins, such as short tails. Depending on breed, domestic sheep may have no horns at all (i.e. polled), or horns in both sexes, or in males only. Most horned breeds have a single pair, but a few breeds may have several.
Sheep in Turkmenistan
Another trait unique to domestic sheep as compared to wild ovines is their wide variation in color. Wild sheep are largely variations of brown hues, and variation within species is extremely limited. Colors of domestic sheep range from pure white to dark chocolate brown, and even spotted or piebald. Sheep keepers also sometimes artificially paint "smit marks" onto their sheep in any pattern or color for identification. Selection for easily dyeable white fleeces began early in sheep domestication, and as white wool is a dominant trait it spread quickly. However, colored sheep do appear in many modern breeds, and may even appear as a recessive trait in white flocks. While white wool is desirable for large commercial markets, there is a niche market for colored fleeces, mostly for handspinning. The nature of the fleece varies widely among the breeds, from dense and highly crimped, to long and hairlike. There is variation of wool type and quality even among members of the same flock, so wool classing is a step in the commercial processing of the fibre.
Suffolks are a medium wool, black-faced breed of meat sheep that make up 60% of the sheep population in the U.S.
Depending on breed, sheep show a range of heights and weights. Their rate of growth and mature weight is a heritable trait that is often selected for in breeding. Ewes typically weigh between 45 and 100 kilograms (100 and 220 lb), and rams between 45 and 160 kilograms (100 and 350 lb). When all deciduous teeth have erupted, the sheep has 20 teeth. Mature sheep have 32 teeth. As with other ruminants, the front teeth in the lower jaw bite against a hard, toothless pad in the upper jaw. These are used to pick off vegetation, then the rear teeth grind it before it is swallowed. There are eight lower front teeth in ruminants, but there is some disagreement as to whether these are eight incisors, or six incisors and two incisor-shaped canines. This means that the dental formula for sheep is either
0.0.3.3
4.0.3.3
or
0.0.3.3
3.1.3.3
There is a large diastema between the incisors and the molars.
In the first few years of life one can calculate the age of sheep from their front teeth, as a pair of milk teeth is replaced by larger adult teeth each year, the full set of eight adult front teeth being complete at about four years of age. The front teeth are then gradually lost as sheep age, making it harder for them to feed and hindering the health and productivity of the animal. For this reason, domestic sheep on normal pasture begin to slowly decline from four years on, and the life expectancy of a sheep is 10 to 12 years, though some sheep may live as long as 20 years.
Skull
Sheep have good hearing, and are sensitive to noise when being handled. Sheep have horizontal slit-shaped pupils, with excellent peripheral vision; with visual fields of about 270° to 320°, sheep can see behind themselves without turning their heads. Many breeds have only short hair on the face, and some have facial wool (if any) confined to the poll and or the area of the mandibular angle; the wide angles of peripheral vision apply to these breeds. A few breeds tend to have considerable wool on the face; for some individuals of these breeds, peripheral vision may be greatly reduced by "wool blindness", unless recently shorn about the face. Sheep have poor depth perception; shadows and dips in the ground may cause sheep to baulk. In general, sheep have a tendency to move out of the dark and into well-lit areas, and prefer to move uphill when disturbed. Sheep also have an excellent sense of smell, and, like all species of their genus, have scent glands just in front of the eyes, and interdigitally on the feet. The purpose of these glands is uncertain, but those on the face may be used in breeding behaviors. The foot glands might also be related to reproduction, but alternative functions, such as secretion of a waste product or a scent marker to help lost sheep find their flock, have also been proposed.
Comparison with goats
Sheep and goats are closely related: both are in the subfamily Caprinae. However, they are separate species, so hybrids rarely occur and are always infertile. A hybrid of a ewe and a buck (a male goat) is called a sheep-goat hybrid, known as geep. Visual differences between sheep and goats include the beard of goats and divided upper lip of sheep. Sheep tails also hang down, even when short or docked, while the short tails of goats are held upwards. Also, sheep breeds are often naturally polled (either in both sexes or just in the female), while naturally polled goats are rare (though many are polled artificially). Males of the two species differ in that buck goats acquire a unique and strong odor during the rut, whereas rams do not.
Breeds
The domestic sheep is a multi-purpose animal, and the more than 200 breeds now in existence were created to serve these diverse purposes. Some sources give a count of a thousand or more breeds, but these numbers cannot be verified, according to some sources. However, several hundred breeds of sheep have been identified by the Food and Agriculture Organization of the UN (FAO), with the estimated number varying somewhat from time to time: e.g. 863 breeds as of 1993, 1314 breeds as of 1995 and 1229 breeds as of 2006. (These numbers exclude extinct breeds, which are also tallied by the FAO.) For the purpose of such tallies, the FAO definition of a breed is "either a subspecific group of domestic livestock with definable and identifiable external characteristics that enable it to be separated by visual appraisal from other similarly defined groups within the same species or a group for which geographical and/or cultural separation from phenotypically similar groups has led to acceptance of its separate identity." Almost all sheep are classified as being best suited to furnishing a certain product: wool, meat, milk, hides, or a combination in a dual-purpose breed. Other features used when classifying sheep include face color (generally white or black), tail length, presence or lack of horns, and the topography for which the breed has been developed. This last point is especially stressed in the UK, where breeds are described as either upland (hill or mountain) or lowland breeds. A sheep may also be of a fat-tailed type, which is a dual-purpose sheep common in Africa and Asia with larger deposits of fat within and around its tail.
Breeds are often categorized by the type of their wool. Fine wool breeds are those that have wool of great crimp and density, which are preferred for textiles. Most of these were derived from Merino sheep, and the breed continues to dominate the world sheep industry. Downs breeds have wool between the extremes, and are typically fast-growing meat and ram breeds with dark faces. Some major medium wool breeds, such as the Corriedale, are dual-purpose crosses of long and fine-wooled breeds and were created for high-production commercial flocks. Long wool breeds are the largest of sheep, with long wool and a slow rate of growth. Long wool sheep are most valued for crossbreeding to improve the attributes of other sheep types. For example: the American Columbia breed was developed by crossing Lincoln rams (a long wool breed) with fine-wooled Rambouillet ewes.
Coarse or carpet wool sheep are those with a medium to long length wool of characteristic coarseness. Breeds traditionally used for carpet wool show great variability, but the chief requirement is a wool that will not break down under heavy use (as would that of the finer breeds). As the demand for carpet-quality wool declines, some breeders of this type of sheep are attempting to use a few of these traditional breeds for alternative purposes. Others have always been primarily meat-class sheep.
A minor class of sheep are the dairy breeds. Dual-purpose breeds that may primarily be meat or wool sheep are often used secondarily as milking animals, but there are a few breeds that are predominantly used for milking. These sheep produce a higher quantity of milk and have slightly longer lactation curves. In the quality of their milk, the fat and protein content percentages of dairy sheep vary from non-dairy breeds, but lactose content does not.
A last group of sheep breeds is that of fur or hair sheep, which do not grow wool at all. Hair sheep are similar to the early domesticated sheep kept before woolly breeds were developed, and are raised for meat and pelts. Some modern breeds of hair sheep, such as the Dorper, result from crosses between wool and hair breeds. For meat and hide producers, hair sheep are cheaper to keep, as they do not need shearing. Hair sheep are also more resistant to parasites and hot weather.
With the modern rise of corporate agribusiness and the decline of localized family farms, many breeds of sheep are in danger of extinction. The Rare Breeds Survival Trust of the UK lists 22 native breeds as having only 3,000 registered animals (each), and The Livestock Conservancy lists 14 as either "critical" or "threatened". Preferences for breeds with uniform characteristics and fast growth have pushed heritage (or heirloom) breeds to the margins of the sheep industry. Those that remain are maintained through the efforts of conservation organizations, breed registries, and individual farmers dedicated to their preservation.
Diet
Sheep are herbivorous mammals. Most breeds prefer to graze on grass and other short roughage, avoiding the taller woody parts of plants that goats readily consume. Both sheep and goats use their lips and tongues to select parts of the plant that are easier to digest or higher in nutrition. Sheep, however, graze well in monoculture pastures where most goats fare poorly.
Ruminant system of a sheep
Like all ruminants, sheep have a complex digestive system composed of four chambers, allowing them to break down cellulose from stems, leaves, and seed hulls into simpler carbohydrates. When sheep graze, vegetation is chewed into a mass called a bolus, which is then passed into the rumen, via the reticulum. The rumen is a 19- to 38-liter (5 to 10 gallon) organ in which feed is fermented. The fermenting organisms include bacteria, fungi, and protozoa. (Other important rumen organisms include some archaea, which produce methane from carbon dioxide.) The bolus is periodically regurgitated back to the mouth as cud for additional chewing and salivation. After fermentation in the rumen, feed passes into the reticulum and the omasum; special feeds such as grains may bypass the rumen altogether. After the first three chambers, food moves into the abomasum for final digestion before processing by the intestines. The abomasum is the only one of the four chambers analogous to the human stomach, and is sometimes called the "true stomach".
Other than forage, the other staple feed for sheep is hay, often during the winter months. The ability to thrive solely on pasture (even without hay) varies with breed, but all sheep can survive on this diet. Also included in some sheep's diets are minerals, either in a trace mix or in licks. Feed provided to sheep must be specially formulated, as most cattle, poultry, pig, and even some goat feeds contain levels of copper that are lethal to sheep. The same danger applies to mineral supplements such as salt licks.
Grazing behavior
Sheep follow a diurnal pattern of activity, feeding from dawn to dusk, stopping sporadically to rest and chew their cud. Ideal pasture for sheep is not lawnlike grass, but an array of grasses, legumes and forbs. Types of land where sheep are raised vary widely, from pastures that are seeded and improved intentionally to rough, native lands. Common plants toxic to sheep are present in most of the world, and include (but are not limited to) cherry, some oaks and acorns, tomato, yew, rhubarb, potato, and rhododendron.
Effects on pasture
Sheep are largely grazing herbivores, unlike browsing animals such as goats and deer that prefer taller foliage. With a much narrower face, sheep crop plants very close to the ground and can overgraze a pasture much faster than cattle. For this reason, many shepherds use managed intensive rotational grazing, where a flock is rotated through multiple pastures, giving plants time to recover. Paradoxically, sheep can both cause and solve the spread of invasive plant species. By disturbing the natural state of pasture, sheep and other livestock can pave the way for invasive plants. However, sheep also prefer to eat invasives such as cheatgrass, leafy spurge, kudzu and spotted knapweed over native species such as sagebrush, making grazing sheep effective for conservation grazing. Research conducted in Imperial County, California compared lamb grazing with herbicides for weed control in seedling alfalfa fields. Three trials demonstrated that grazing lambs were just as effective as herbicides in controlling winter weeds. Entomologists also compared grazing lambs to insecticides for insect control in winter alfalfa. In this trial, lambs provided insect control as effectively as insecticides.
Behavior
Sheep are flock animals and strongly gregarious; much sheep behavior can be understood on the basis of these tendencies. The dominance hierarchy of sheep and their natural inclination to follow a leader to new pastures were the pivotal factors in sheep being one of the first domesticated livestock species. Furthermore, in contrast to the red deer and gazelle (two other ungulates of primary importance to meat production in prehistoric times), sheep do not defend territories although they do form home ranges. All sheep have a tendency to congregate close to other members of a flock, although this behavior varies with breed, and sheep can become stressed when separated from their flock members. During flocking, sheep have a strong tendency to follow, and a leader may simply be the first individual to move. Relationships in flocks tend to be closest among related sheep: in mixed-breed flocks, subgroups of the same breed tend to form, and a ewe and her direct descendants often move as a unit within large flocks. Sheep can become hefted to one particular local pasture (heft) so they do not roam freely in unfenced landscapes. Lambs learn the heft from ewes and if whole flocks are culled it must be retaught to the replacement animals.
Flock behaviour in sheep is generally only exhibited in groups of four or more sheep; fewer sheep may not react as expected when alone or with few other sheep. Being a prey species, the primary defense mechanism of sheep is to flee from danger when their flight zone is entered. Cornered sheep may charge and butt, or threaten by hoof stamping and adopting an aggressive posture. This is particularly true for ewes with newborn lambs.
In regions where sheep have no natural predators, none of the native breeds of sheep exhibit a strong flocking behavior.
Herding
Farmers exploit flocking behavior to keep sheep together on unfenced pastures such as hill farming, and to move them more easily. For this purpose shepherds may use herding dogs in this effort, with a highly bred herding ability. Sheep are food-oriented, and association of humans with regular feeding often results in sheep soliciting people for food. Those who are moving sheep may exploit this behavior by leading sheep with buckets of feed.
Dominance hierarchy
Sheep establish a dominance hierarchy through fighting, threats and competitiveness. Dominant animals are inclined to be more aggressive with other sheep, and usually feed first at troughs. Primarily among rams, horn size is a factor in the flock hierarchy. Rams with different size horns may be less inclined to fight to establish the dominance order, while rams with similarly sized horns are more so. Merinos have an almost linear hierarchy whereas there is a less rigid structure in Border Leicesters when a competitive feeding situation arises.
In sheep, position in a moving flock is highly correlated with social dominance, but there is no definitive study to show consistent voluntary leadership by an individual sheep.
Intelligence and learning ability
Sheep are frequently thought of as unintelligent animals. Their flocking behavior and quickness to flee and panic can make shepherding a difficult endeavor for the uninitiated. Despite these perceptions, a University of Illinois monograph on sheep reported their intelligence to be just below that of pigs and on par with that of cattle. Sheep can recognize individual human and ovine faces and remember them for years; they can remember 50 other different sheep faces for over two years; they can recognize and are attracted to individual sheep and humans by their faces, as they possess similar specialized neural systems in the temporal and frontal lobes of their brains to humans and have a greater involvement of the right brain hemisphere. In addition to long-term facial recognition of individuals, sheep can also differentiate emotional states through facial characteristics.[68][69] If worked with patiently, sheep may learn their names, and many sheep are trained to be led by halter for showing and other purposes. Sheep have also responded well to clicker training. Sheep have been used as pack animals; Tibetan nomads distribute baggage equally throughout a flock as it is herded between living sites.
It has been reported that some sheep have apparently shown problem-solving abilities; a flock in West Yorkshire, England allegedly found a way to get over cattle grids by rolling on their backs, although documentation of this has relied on anecdotal accounts.
Vocalisations
Sounds made by domestic sheep include bleats, grunts, rumbles and snorts. Bleating ("baaing") is used mostly for contact communication, especially between dam and lambs, but also at times between other flock members. The bleats of individual sheep are distinctive, enabling the ewe and her lambs to recognize each other's vocalizations. Vocal communication between lambs and their dam declines to a very low level within several weeks after parturition. A variety of bleats may be heard, depending on sheep age and circumstances. Apart from contact communication, bleating may signal distress, frustration or impatience; however, sheep are usually silent when in pain. Isolation commonly prompts bleating by sheep. Pregnant ewes may grunt when in labor. Rumbling sounds are made by the ram during courting; somewhat similar rumbling sounds may be made by the ewe, especially when with her neonate lambs. A snort (explosive exhalation through the nostrils) may signal aggression or a warning, and is often elicited from startled sheep.
Lamb
In sheep breeds lacking facial wool, the visual field is wide. In 10 sheep (Cambridge, Lleyn and Welsh Mountain breeds, which lack facial wool), the visual field ranged from 298° to 325°, averaging 313.1°, with binocular overlap ranging from 44.5° to 74°, averaging 61.7°. In some breeds, unshorn facial wool can limit the visual field; in some individuals, this may be enough to cause "wool blindness". In 60 Merinos, visual fields ranged from 219.1° to 303.0°, averaging 269.9°, and the binocular field ranged from 8.9° to 77.7°, averaging 47.5°; 36% of the measurements were limited by wool, although photographs of the experiments indicate that only limited facial wool regrowth had occurred since shearing. In addition to facial wool (in some breeds), visual field limitations can include ears and (in some breeds) horns, so the visual field can be extended by tilting the head. Sheep eyes exhibit very low hyperopia and little astigmatism. Such visual characteristics are likely to produce a well-focused retinal image of objects in both the middle and long distance. Because sheep eyes have no accommodation, one might expect the image of very near objects to be blurred, but a rather clear near image could be provided by the tapetum and large retinal image of the sheep's eye, and adequate close vision may occur at muzzle length. Good depth perception, inferred from the sheep's sure-footedness, was confirmed in "visual cliff" experiments; behavioral responses indicating depth perception are seen in lambs at one day old. Sheep are thought to have colour vision, and can distinguish between a variety of colours: black, red, brown, green, yellow and white. Sight is a vital part of sheep communication, and when grazing, they maintain visual contact with each other. Each sheep lifts its head upwards to check the position of other sheep in the flock. This constant monitoring is probably what keeps the sheep in a flock as they move along grazing. Sheep become stressed when isolated; this stress is reduced if they are provided with a mirror, indicating that the sight of other sheep reduces stress.
Taste is the most important sense in sheep, establishing forage preferences, with sweet and sour plants being preferred and bitter plants being more commonly rejected. Touch and sight are also important in relation to specific plant characteristics, such as succulence and growth form.
The ram uses his vomeronasal organ (sometimes called the Jacobson's organ) to sense the pheromones of ewes and detect when they are in estrus. The ewe uses her vomeronasal organ for early recognition of her neonate lamb.
Reproduction
Sheep follow a similar reproductive strategy to other herd animals. A group of ewes is generally mated by a single ram, who has either been chosen by a breeder or (in feral populations) has established dominance through physical contest with other rams. Most sheep are seasonal breeders, although some are able to breed year-round. Ewes generally reach sexual maturity at six to eight months old, and rams generally at four to six months. However, there are exceptions. For example, Finnsheep ewe lambs may reach puberty as early as 3 to 4 months, and Merino ewes sometimes reach puberty at 18 to 20 months. Ewes have estrus cycles about every 17 days, during which they emit a scent and indicate readiness through physical displays towards rams.
In feral sheep, rams may fight during the rut to determine which individuals may mate with ewes. Rams, especially unfamiliar ones, will also fight outside the breeding period to establish dominance; rams can kill one another if allowed to mix freely. During the rut, even usually friendly rams may become aggressive towards humans due to increases in their hormone levels.
After mating, sheep have a gestation period of about five months, and normal labor takes one to three hours. Although some breeds regularly throw larger litters of lambs, most produce single or twin lambs. During or soon after labor, ewes and lambs may be confined to small lambing jugs, small pens designed to aid both careful observation of ewes and to cement the bond between them and their lambs.
A lamb's first steps
Ovine obstetrics can be problematic. By selectively breeding ewes that produce multiple offspring with higher birth weights for generations, sheep producers have inadvertently caused some domestic sheep to have difficulty lambing; balancing ease of lambing with high productivity is one of the dilemmas of sheep breeding. In the case of any such problems, those present at lambing may assist the ewe by extracting or repositioning lambs. After the birth, ewes ideally break the amniotic sac (if it is not broken during labor), and begin licking clean the lamb. Most lambs will begin standing within an hour of birth. In normal situations, lambs nurse after standing, receiving vital colostrum milk. Lambs that either fail to nurse or are rejected by the ewe require help to survive, such as bottle-feeding or fostering by another ewe.
Most lambs begin life being born outdoors. After lambs are several weeks old, lamb marking (ear tagging, docking, mulesing, and castrating) is carried out. Vaccinations are usually carried out at this point as well. Ear tags with numbers are attached, or ear marks are applied, for ease of later identification of sheep. Docking and castration are commonly done after 24 hours (to avoid interference with maternal bonding and consumption of colostrum) and are often done not later than one week after birth, to minimize pain, stress, recovery time and complications. The first course of vaccinations (commonly anti-clostridial) is commonly given at an age of about 10 to 12 weeks; i.e. when the concentration of maternal antibodies passively acquired via colostrum is expected to have fallen low enough to permit development of active immunity. Ewes are often revaccinated annually about 3 weeks before lambing, to provide high antibody concentrations in colostrum during the first several hours after lambing. Ram lambs that will either be slaughtered or separated from ewes before sexual maturity are not usually castrated. Objections to all these procedures have been raised by animal rights groups, but farmers defend them by saying they save money, and inflict only temporary pain.
Homosexuality
Sheep are the only species of mammal except for humans which exhibits exclusive homosexual behavior. About 10% of rams refuse to mate with ewes but readily mate with other rams, and thirty percent of all rams demonstrate at least some homosexual behavior. Additionally, a small number of females that were accompanied by a male fetus in utero (i.e. as fraternal twins) are freemartins (female animals that are behaviorally masculine and lack functioning ovaries).
Health
Sheep may fall victim to poisons, infectious diseases, and physical injuries. As a prey species, a sheep's system is adapted to hide the obvious signs of illness, to prevent being targeted by predators. However, some signs of ill health are obvious, with sick sheep eating little, vocalizing excessively, and being generally listless. Throughout history, much of the money and labor of sheep husbandry has aimed to prevent sheep ailments. Historically, shepherds often created remedies by experimentation on the farm. In some developed countries, including the United States, sheep lack the economic importance for drug companies to perform expensive clinical trials required to approve more than a relatively limited number of drugs for ovine use. However, extra-label drug use in sheep production is permitted in many jurisdictions, subject to certain restrictions. In the US, for example, regulations governing extra-label drug use in animals are found in 21 CFR (Code of Federal Regulations) Part 530. In the 20th and 21st centuries, a minority of sheep owners have turned to alternative treatments such as homeopathy, herbalism and even traditional Chinese medicine to treat sheep veterinary problems. Despite some favorable anecdotal evidence, the effectiveness of alternative veterinary medicine has been met with skepticism in scientific journals. The need for traditional anti-parasite drugs and antibiotics is widespread, and is the main impediment to certified organic farming with sheep.
Many breeders take a variety of preventive measures to ward off problems. The first is to ensure all sheep are healthy when purchased. Many buyers avoid outlets known to be clearing houses for animals culled from healthy flocks as either sick or simply inferior. This can also mean maintaining a closed flock, and quarantining new sheep for a month. Two fundamental preventive programs are maintaining good nutrition and reducing stress in the sheep. Restraint, isolation, loud noises, novel situations, pain, heat, extreme cold, fatigue and other stressors can lead to secretion of cortisol, a stress hormone, in amounts that may indicate welfare problems. Excessive stress can compromise the immune system. "Shipping fever" (pneumonic mannheimiosis, formerly called pasteurellosis) is a disease of particular concern, that can occur as a result of stress, notably during transport and (or) handling. Pain, fear and several other stressors can cause secretion of epinephrine (adrenaline). Considerable epinephrine secretion in the final days before slaughter can adversely affect meat quality (by causing glycogenolysis, removing the substrate for normal post-slaughter acidification of meat) and result in meat becoming more susceptible to colonization by spoilage bacteria. Because of such issues, low-stress handling is essential in sheep management. Avoiding poisoning is also important; common poisons are pesticide sprays, inorganic fertilizer, motor oil, as well as radiator coolant containing ethylene glycol.
Common forms of preventive medication for sheep are vaccinations and treatments for parasites. Both external and internal parasites are the most prevalent malady in sheep, and are either fatal, or reduce the productivity of flocks. Worms are the most common internal parasites. They are ingested during grazing, incubate within the sheep, and are expelled through the digestive system (beginning the cycle again). Oral anti-parasitic medicines, known as drenches, are given to a flock to treat worms, sometimes after worm eggs in the feces has been counted to assess infestation levels. Afterwards, sheep may be moved to a new pasture to avoid ingesting the same parasites. External sheep parasites include: lice (for different parts of the body), sheep keds, nose bots, sheep itch mites, and maggots. Keds are blood-sucking parasites that cause general malnutrition and decreased productivity, but are not fatal. Maggots are those of the bot fly and the blow-fly, commonly Lucilia sericata or its relative L. cuprina. Fly maggots cause the extremely destructive condition of flystrike. Flies lay their eggs in wounds or wet, manure-soiled wool; when the maggots hatch they burrow into a sheep's flesh, eventually causing death if untreated. In addition to other treatments, crutching (shearing wool from a sheep's rump) is a common preventive method. Some countries allow mulesing, a practice that involves stripping away the skin on the rump to prevent fly-strike, normally performed when the sheep is a lamb. Nose bots are fly larvae that inhabit a sheep's sinuses, causing breathing difficulties and discomfort. Common signs are a discharge from the nasal passage, sneezing, and frantic movement such as head shaking. External parasites may be controlled through the use of backliners, sprays or immersive sheep dips.
A wide array of bacterial and viral diseases affect sheep. Diseases of the hoof, such as foot rot and foot scald may occur, and are treated with footbaths and other remedies. Foot rot is present in over 97% of flocks in the UK. These painful conditions cause lameness and hinder feeding. Ovine Johne's disease is a wasting disease that affects young sheep. Bluetongue disease is an insect-borne illness causing fever and inflammation of the mucous membranes. Ovine rinderpest (or peste des petits ruminants) is a highly contagious and often fatal viral disease affecting sheep and goats. Sheep may also be affected by primary or secondary photosensitization. Tetanus can also afflict sheep through wounds from shearing, docking, castration, or vaccination. The organism also can be introduced into the reproductive tract by unsanitary humans who assist ewes during lambing.
A few sheep conditions are transmissible to humans. Orf (also known as scabby mouth, contagious ecthyma or soremouth) is a skin disease leaving lesions that is transmitted through skin-to-skin contact. Cutaneous anthrax is also called woolsorter's disease, as the spores can be transmitted in unwashed wool. More seriously, the organisms that can cause spontaneous enzootic abortion in sheep are easily transmitted to pregnant women. Also of concern are the prion disease scrapie and the virus that causes foot-and-mouth disease (FMD), as both can devastate flocks. The latter poses a slight risk to humans. During the 2001 FMD pandemic in the UK, hundreds of sheep were culled and some rare British breeds were at risk of extinction due to this.
Of the 600,300 sheep lost to the US economy in 2004, 37.3% were lost to predators, while 26.5% were lost to some form of disease. Poisoning accounted for 1.7% of non-productive deaths.
Predators
A lamb being attacked by coyotes with a bite to the throat
Other than parasites and disease, predation is a threat to sheep and the profitability of sheep raising. Sheep have little ability to defend themselves, compared with other species kept as livestock. Even if sheep survive an attack, they may die from their injuries or simply from panic. However, the impact of predation varies dramatically with region. In Africa, Australia, the Americas, and parts of Europe and Asia predators are a serious problem. In the United States, for instance, over one third of sheep deaths in 2004 were caused by predation. In contrast, other nations are virtually devoid of sheep predators, particularly islands known for extensive sheep husbandry. Worldwide, canids—including the domestic dog—are responsible for most sheep deaths. Other animals that occasionally prey on sheep include: felines, bears, birds of prey, ravens and feral hogs.
Sheep producers have used a wide variety of measures to combat predation. Pre-modern shepherds used their own presence, livestock guardian dogs, and protective structures such as barns and fencing. Fencing (both regular and electric), penning sheep at night and lambing indoors all continue to be widely used. More modern shepherds used guns, traps, and poisons to kill predators, causing significant decreases in predator populations. In the wake of the environmental and conservation movements, the use of these methods now usually falls under the purview of specially designated government agencies in most developed countries.
The 1970s saw a resurgence in the use of livestock guardian dogs and the development of new methods of predator control by sheep producers, many of them non-lethal. Donkeys and guard llamas have been used since the 1980s in sheep operations, using the same basic principle as livestock guardian dogs. Interspecific pasturing, usually with larger livestock such as cattle or horses, may help to deter predators, even if such species do not actively guard sheep. In addition to animal guardians, contemporary sheep operations may use non-lethal predator deterrents such as motion-activated lights and noisy alarms.
Economic importance
Main article: Agricultural economics
Global sheep stock
in 2019
Number in millions
1. China163.5 (13.19%)
2. India74.3 (5.99%)
3. Australia65.8 (5.31%)
4. Nigeria46.9 (3.78%)
5. Iran41.3 (3.33%)
6. Sudan40.9 (3.3%)
7. Chad35.9 (2.9%)
8. Turkey35.2 (2.84%)
9. United Kingdom33.6 (2.71%)
10. Mongolia32.3 (2.61%)
World total1,239.8
Source: UN Food and Agriculture Organization
Sheep are an important part of the global agricultural economy. However, their once vital status has been largely replaced by other livestock species, especially the pig, chicken, and cow. China, Australia, India, and Iran have the largest modern flocks, and serve both local and exportation needs for wool and mutton. Other countries such as New Zealand have smaller flocks but retain a large international economic impact due to their export of sheep products. Sheep also play a major role in many local economies, which may be niche markets focused on organic or sustainable agriculture and local food customers. Especially in developing countries, such flocks may be a part of subsistence agriculture rather than a system of trade. Sheep themselves may be a medium of trade in barter economies.
Domestic sheep provide a wide array of raw materials. Wool was one of the first textiles, although in the late 20th century wool prices began to fall dramatically as the result of the popularity and cheap prices for synthetic fabrics. For many sheep owners, the cost of shearing is greater than the possible profit from the fleece, making subsisting on wool production alone practically impossible without farm subsidies. Fleeces are used as material in making alternative products such as wool insulation. In the 21st century, the sale of meat is the most profitable enterprise in the sheep industry, even though far less sheep meat is consumed than chicken, pork or beef.
Sheepskin is likewise used for making clothes, footwear, rugs, and other products. Byproducts from the slaughter of sheep are also of value: sheep tallow can be used in candle and soap making, sheep bone and cartilage has been used to furnish carved items such as dice and buttons as well as rendered glue and gelatin. Sheep intestine can be formed into sausage casings, and lamb intestine has been formed into surgical sutures, as well as strings for musical instruments and tennis rackets. Sheep droppings, which are high in cellulose, have even been sterilized and mixed with traditional pulp materials to make paper. Of all sheep byproducts, perhaps the most valuable is lanolin: the waterproof, fatty substance found naturally in sheep's wool and used as a base for innumerable cosmetics and other products.
Some farmers who keep sheep also make a profit from live sheep. Providing lambs for youth programs such as 4-H and competition at agricultural shows is often a dependable avenue for the sale of sheep. Farmers may also choose to focus on a particular breed of sheep in order to sell registered purebred animals, as well as provide a ram rental service for breeding. A new option for deriving profit from live sheep is the rental of flocks for grazing; these "mowing services" are hired in order to keep unwanted vegetation down in public spaces and to lessen fire hazard.
Despite the falling demand and price for sheep products in many markets, sheep have distinct economic advantages when compared with other livestock. They do not require expensive housing, such as that used in the intensive farming of chickens or pigs. They are an efficient use of land; roughly six sheep can be kept on the amount that would suffice for a single cow or horse. Sheep can also consume plants, such as noxious weeds, that most other animals will not touch, and produce more young at a faster rate. Also, in contrast to most livestock species, the cost of raising sheep is not necessarily tied to the price of feed crops such as grain, soybeans and corn. Combined with the lower cost of quality sheep, all these factors combine to equal a lower overhead for sheep producers, thus entailing a higher profitability potential for the small farmer. Sheep are especially beneficial for independent producers, including family farms with limited resources, as the sheep industry is one of the few types of animal agriculture that has not been vertically integrated by agribusiness. However, small flocks, from 10 to 50 ewes, often are not profitable because they tend to be poorly managed. The primary reason is that mechanization is not feasible, so return per hour of labor is not maximized. Small farm flocks generally are used simply to control weeds on irrigation ditches or maintained as a hobby.
Shoulder of lamb
Sheep meat and milk were one of the earliest staple proteins consumed by human civilization after the transition from hunting and gathering to agriculture. Sheep meat prepared for food is known as either mutton or lamb, and approximately 540 million sheep are slaughtered each year for meat worldwide. "Mutton" is derived from the Old French moton, which was the word for sheep used by the Anglo-Norman rulers of much of the British Isles in the Middle Ages. This became the name for sheep meat in English, while the Old English word sceap was kept for the live animal. Throughout modern history, "mutton" has been limited to the meat of mature sheep usually at least two years of age; "lamb" is used for that of immature sheep less than a year.
In the 21st century, the nations with the highest consumption of sheep meat are the Arab states of the Persian Gulf, New Zealand, Australia, Greece, Uruguay, the United Kingdom and Ireland. These countries eat 14–40 lbs (3–18 kg) of sheep meat per capita, per annum. Sheep meat is also popular in France, Africa (especially the Arab world), the Caribbean, the rest of the Middle East, India, and parts of China. This often reflects a history of sheep production. In these countries in particular, dishes comprising alternative cuts and offal may be popular or traditional. Sheep testicles—called animelles or lamb fries—are considered a delicacy in many parts of the world. Perhaps the most unusual dish of sheep meat is the Scottish haggis, composed of various sheep innards cooked along with oatmeal and chopped onions inside its stomach. In comparison, countries such as the U.S. consume only a pound or less (under 0.5 kg), with Americans eating 50 pounds (22 kg) of pork and 65 pounds (29 kg) of beef. In addition, such countries rarely eat mutton, and may favor the more expensive cuts of lamb: mostly lamb chops and leg of lamb.
Though sheep's milk may be drunk rarely in fresh form, today it is used predominantly in cheese and yogurt making. Sheep have only two teats, and produce a far smaller volume of milk than cows. However, as sheep's milk contains far more fat, solids, and minerals than cow's milk, it is ideal for the cheese-making process. It also resists contamination during cooling better because of its much higher calcium content. Well-known cheeses made from sheep milk include the feta of Bulgaria and Greece, Roquefort of France, Manchego from Spain, the pecorino romano (the Italian word for "sheep" is pecore) and ricotta of Italy. Yogurts, especially some forms of strained yogurt, may also be made from sheep milk. Many of these products are now often made with cow's milk, especially when produced outside their country of origin. Sheep milk contains 4.8% lactose, which may affect those who are intolerant.
As with other domestic animals, the meat of uncastrated males is inferior in quality, especially as they grow. A "bucky" lamb is a lamb which was not castrated early enough, or which was castrated improperly (resulting in one testicle being retained). These lambs are worth less at market.
In science
Sheep are generally too large and reproduce too slowly to make ideal research subjects, and thus are not a common model organism. They have, however, played an influential role in some fields of science. In particular, the Roslin Institute of Edinburgh, Scotland used sheep for genetics research that produced groundbreaking results. In 1995, two ewes named Megan and Morag were the first mammals cloned from differentiated cells, also referred to as gynomerogony. A year later, a Finnish Dorset sheep named Dolly, dubbed "the world's most famous sheep" in Scientific American, was the first mammal to be cloned from an adult somatic cell. Following this, Polly and Molly were the first mammals to be simultaneously cloned and transgenic.
As of 2008, the sheep genome has not been fully sequenced, although a detailed genetic map has been published, and a draft version of the complete genome produced by assembling sheep DNA sequences using information given by the genomes of other mammals. In 2012, a transgenic sheep named "Peng Peng" was cloned by Chinese scientists, who spliced his genes with that of a roundworm (C. elegans) in order to increase production of fats healthier for human consumption.
In the study of natural selection, the population of Soay sheep that remain on the island of Hirta have been used to explore the relation of body size and coloration to reproductive success. Soay sheep come in several colors, and researchers investigated why the larger, darker sheep were in decline; this occurrence contradicted the rule of thumb that larger members of a population tend to be more successful reproductively. The feral Soays on Hirta are especially useful subjects because they are isolated.
Domestic sheep are sometimes used in medical research, particularly for researching cardiovascular physiology, in areas such as hypertension and heart failure. Pregnant sheep are also a useful model for human pregnancy, and have been used to investigate the effects on fetal development of malnutrition and hypoxia. In behavioral sciences, sheep have been used in isolated cases for the study of facial recognition, as their mental process of recognition is qualitatively similar to humans.
Cultural impact
Sheep have had a strong presence in many cultures, especially in areas where they form the most common type of livestock. In the English language, to call someone a sheep or ovine may allude that they are timid and easily led. In contradiction to this image, male sheep are often used as symbols of virility and power; the logos of the Los Angeles Rams football team and the Dodge Ram pickup truck allude to males of the bighorn sheep, Ovis canadensis.
Counting sheep is popularly said to be an aid to sleep, and some ancient systems of counting sheep persist today. Sheep also enter in colloquial sayings and idiom frequently with such phrases as "black sheep". To call an individual a black sheep implies that they are an odd or disreputable member of a group. This usage derives from the recessive trait that causes an occasional black lamb to be born into an entirely white flock. These black sheep were considered undesirable by shepherds, as black wool is not as commercially viable as white wool. Citizens who accept overbearing governments have been referred to by the Portmanteau neologism of sheeple. Somewhat differently, the adjective "sheepish" is also used to describe embarrassment.
In heraldry
In British heraldry, sheep appear in the form of rams, sheep proper and lambs. These are distinguished by the ram being depicted with horns and a tail, the sheep with neither and the lamb with its tail only. A further variant of the lamb, termed the Paschal lamb, is depicted as carrying a Christian cross and with a halo over its head. Rams' heads, portrayed without a neck and facing the viewer, are also found in British armories. The fleece, depicted as an entire sheepskin carried by a ring around its midsection, originally became known through its use in the arms of the Order of the Golden Fleece and was later adopted by towns and individuals with connections to the wool industry. A sheep on a blue field is depicted on the greater/royal arms of the king of Denmark to represent the Faroe Islands. In 2004 a modernized arms has been adopted by the Faroe Islands, which based on a 15th century coat of arms.
Religion and folklore
In antiquity, symbolism involving sheep cropped up in religions in the ancient Near East, the Mideast, and the Mediterranean area: Çatalhöyük, ancient Egyptian religion, the Cana'anite and Phoenician tradition, Judaism, Greek religion, and others. Religious symbolism and ritual involving sheep began with some of the first known faiths: Skulls of rams (along with bulls) occupied central placement in shrines at the Çatalhöyük settlement in 8,000 BCE. In Ancient Egyptian religion, the ram was the symbol of several gods: Khnum, Heryshaf and Amun (in his incarnation as a god of fertility). Other deities occasionally shown with ram features include the goddess Ishtar, the Phoenician god Baal-Hamon, and the Babylonian god Ea-Oannes. In Madagascar, sheep were not eaten as they were believed to be incarnations of the souls of ancestors.
There are many ancient Greek references to sheep: that of Chrysomallos, the golden-fleeced ram, continuing to be told through into the modern era. Astrologically, Aries, the ram, is the first sign of the classical Greek zodiac, and the sheep is the eighth of the twelve animals associated with the 12-year cycle of in the Chinese zodiac, related to the Chinese calendar. It is said in Chinese traditions that Hou ji sacrificed sheep. Mongolia, shagai are an ancient form of dice made from the cuboid bones of sheep that are often used for fortunetelling purposes.
Sheep play an important role in all the Abrahamic faiths; Abraham, Isaac, Jacob, Moses, and King David were all shepherds. According to the Biblical story of the Binding of Isaac, a ram is sacrificed as a substitute for Isaac after an angel stays Abraham's hand (in the Islamic tradition, Abraham was about to sacrifice Ishmael). Eid al-Adha is a major annual festival in Islam in which sheep (or other animals) are sacrificed in remembrance of this act. Sheep are occasionally sacrificed to commemorate important secular events in Islamic cultures. Greeks and Romans sacrificed sheep regularly in religious practice, and Judaism once sacrificed sheep as a Korban (sacrifice), such as the Passover lamb. Ovine symbols—such as the ceremonial blowing of a shofar—still find a presence in modern Judaic traditions.
Collectively, followers of Christianity are often referred to as a flock, with Christ as the Good Shepherd, and sheep are an element in the Christian iconography of the birth of Jesus. Some Christian saints are considered patrons of shepherds, and even of sheep themselves. Christ is also portrayed as the Sacrificial lamb of God (Agnus Dei) and Easter celebrations in Greece and Romania traditionally feature a meal of Paschal lamb. A church leader is often called the pastor, which is derived from the Latin word for shepherd. In many western Christian traditions bishops carry a staff, which also serves as a symbol of the episcopal office, known as a crosier, which is modeled on the shepherd's crook.
Sheep are key symbols in fables and nursery rhymes like The Wolf in Sheep's Clothing, Little Bo Peep, Baa, Baa, Black Sheep, and Mary Had a Little Lamb; novels such as George Orwell's Animal Farm and Haruki Murakami's A Wild Sheep Chase; songs such as Bach's Sheep may safely graze (Schafe können sicher weiden) and Pink Floyd's "Sheep", and poems like William Blake's "The Lamb".
The label in the image describes this as the Lesser Vampire (Vampyrus auritus). Today this mammal is actually known to be the big-eared woolly bat or (Peter's) woolly false vampire bat (Chrotopterus auritus). The label goes on to say that the specimen was recovered from Brazil and presented to the Natural History Museum in London by Dr C Peters in 1870. It is preserved in alcohol in a glass case. I love that the detail of the wing membranes remain so well defined.
The name Chrotopterus is derived from Greek roots chariots (skin, colour), and pteron (wing). The epithet auritus refers to the large ears. Big-eared woolly bats are very large predatory bats, the second largest bat species in the neotropics. Their body mass typically ranges from 75 to 96 grams. The length of the forearm ranges from 78.7 to 83.1 mm. There are only three New World phyllostomid bats of comparable size. The dorsal hair is about 12mm long, which is longer than most of species in the Phyllostomids. They also possess two lower incisors, a trait typically shared with smaller bats.
This bat species lives in warm sub-tropical forests, usually roosting in caves and hollow logs, where prey is consumed. Geographically, it is found between southern Mexico and Bolivia.
Big-eared woolly bats have diverse feeding habits. They have been shown to feed primarily on small arthropods and small vertebrates, including beetles, moths, small mammals (including opossums and rodents), birds (including passerine birds) and even other bat species. Though primarily carnivorous or insectivorous, it has also been known to eat fruit. It is NOT a vampire!
Londolozi Game Reserve
South Africa
Click On Image To enlarge.
You can see part of the elephant's trunk in the foliage. Below are some quick fact about elephants.
The elephant is the largest of all land mammals.
Life Span - elephants can live for up to 70 years.
Elephants normally walk about 4 mph.
Elephants are able to swim for long distances.
Elephants spend about 16 hours a day eating.
They consume as much as 495 pounds of food per day.
They live in tight social units led by an older matriarch.
Males leave the herd between the ages of 12 and 15.
Their tusks are of ivory and are actually enormously enlarged incisors.
The elephant's eyes are small and its eyesight is poor.
They have the largest brains in the animal kingdom.
Sheep (pl.: sheep) or domestic sheep (Ovis aries) are a domesticated, ruminant mammal typically kept as livestock. Although the term sheep can apply to other species in the genus Ovis, in everyday usage it almost always refers to domesticated sheep. Like all ruminants, sheep are members of the order Artiodactyla, the even-toed ungulates. Numbering a little over one billion, domestic sheep are also the most numerous species of sheep. An adult female is referred to as a ewe (/juː/ yoo), an intact male as a ram, occasionally a tup, a castrated male as a wether, and a young sheep as a lamb.
Sheep are most likely descended from the wild mouflon of Europe and Asia, with Iran being a geographic envelope of the domestication center. One of the earliest animals to be domesticated for agricultural purposes, sheep are raised for fleeces, meat (lamb, hogget or mutton) and milk. A sheep's wool is the most widely used animal fiber, and is usually harvested by shearing. In Commonwealth countries, ovine meat is called lamb when from younger animals and mutton when from older ones; in the United States, meat from both older and younger animals is usually called lamb. Sheep continue to be important for wool and meat today, and are also occasionally raised for pelts, as dairy animals, or as model organisms for science.
Sheep husbandry is practised throughout the majority of the inhabited world, and has been fundamental to many civilizations. In the modern era, Australia, New Zealand, the southern and central South American nations, and the British Isles are most closely associated with sheep production.
There is a large lexicon of unique terms for sheep husbandry which vary considerably by region and dialect. Use of the word sheep began in Middle English as a derivation of the Old English word scēap. A group of sheep is called a flock. Many other specific terms for the various life stages of sheep exist, generally related to lambing, shearing, and age.
Being a key animal in the history of farming, sheep have a deeply entrenched place in human culture, and are represented in much modern language and symbolism. As livestock, sheep are most often associated with pastoral, Arcadian imagery. Sheep figure in many mythologies—such as the Golden Fleece—and major religions, especially the Abrahamic traditions. In both ancient and modern religious ritual, sheep are used as sacrificial animals.
History
Main article: History of the domestic sheep
The exact line of descent from wild ancestors to domestic sheep is unclear. The most common hypothesis states that Ovis aries is descended from the Asiatic (O. gmelini) species of mouflon; the European mouflon (Ovis aries musimon) is a direct descendant of this population. Sheep were among the first animals to be domesticated by humankind (although the domestication of dogs probably took place 10 to 20 thousand years earlier); the domestication date is estimated to fall between 11,000 and 9000 B.C in Mesopotamia and possibly around 7000 BC in Mehrgarh in the Indus Valley. The rearing of sheep for secondary products, and the resulting breed development, began in either southwest Asia or western Europe. Initially, sheep were kept solely for meat, milk and skins. Archaeological evidence from statuary found at sites in Iran suggests that selection for woolly sheep may have begun around 6000 BC, and the earliest woven wool garments have been dated to two to three thousand years later.
Sheep husbandry spread quickly in Europe. Excavations show that in about 6000 BC, during the Neolithic period of prehistory, the Castelnovien people, living around Châteauneuf-les-Martigues near present-day Marseille in the south of France, were among the first in Europe to keep domestic sheep. Practically from its inception, ancient Greek civilization relied on sheep as primary livestock, and were even said to name individual animals. Ancient Romans kept sheep on a wide scale, and were an important agent in the spread of sheep raising. Pliny the Elder, in his Natural History (Naturalis Historia), speaks at length about sheep and wool. European colonists spread the practice to the New World from 1493 onwards.
Characteristics
Domestic sheep are relatively small ruminants, usually with a crimped hair called wool and often with horns forming a lateral spiral. They differ from their wild relatives and ancestors in several respects, having become uniquely neotenic as a result of selective breeding by humans. A few primitive breeds of sheep retain some of the characteristics of their wild cousins, such as short tails. Depending on breed, domestic sheep may have no horns at all (i.e. polled), or horns in both sexes, or in males only. Most horned breeds have a single pair, but a few breeds may have several.
Sheep in Turkmenistan
Another trait unique to domestic sheep as compared to wild ovines is their wide variation in color. Wild sheep are largely variations of brown hues, and variation within species is extremely limited. Colors of domestic sheep range from pure white to dark chocolate brown, and even spotted or piebald. Sheep keepers also sometimes artificially paint "smit marks" onto their sheep in any pattern or color for identification. Selection for easily dyeable white fleeces began early in sheep domestication, and as white wool is a dominant trait it spread quickly. However, colored sheep do appear in many modern breeds, and may even appear as a recessive trait in white flocks. While white wool is desirable for large commercial markets, there is a niche market for colored fleeces, mostly for handspinning. The nature of the fleece varies widely among the breeds, from dense and highly crimped, to long and hairlike. There is variation of wool type and quality even among members of the same flock, so wool classing is a step in the commercial processing of the fibre.
Suffolks are a medium wool, black-faced breed of meat sheep that make up 60% of the sheep population in the U.S.
Depending on breed, sheep show a range of heights and weights. Their rate of growth and mature weight is a heritable trait that is often selected for in breeding. Ewes typically weigh between 45 and 100 kilograms (100 and 220 lb), and rams between 45 and 160 kilograms (100 and 350 lb). When all deciduous teeth have erupted, the sheep has 20 teeth. Mature sheep have 32 teeth. As with other ruminants, the front teeth in the lower jaw bite against a hard, toothless pad in the upper jaw. These are used to pick off vegetation, then the rear teeth grind it before it is swallowed. There are eight lower front teeth in ruminants, but there is some disagreement as to whether these are eight incisors, or six incisors and two incisor-shaped canines. This means that the dental formula for sheep is either
0.0.3.3
4.0.3.3
or
0.0.3.3
3.1.3.3
There is a large diastema between the incisors and the molars.
In the first few years of life one can calculate the age of sheep from their front teeth, as a pair of milk teeth is replaced by larger adult teeth each year, the full set of eight adult front teeth being complete at about four years of age. The front teeth are then gradually lost as sheep age, making it harder for them to feed and hindering the health and productivity of the animal. For this reason, domestic sheep on normal pasture begin to slowly decline from four years on, and the life expectancy of a sheep is 10 to 12 years, though some sheep may live as long as 20 years.
Skull
Sheep have good hearing, and are sensitive to noise when being handled. Sheep have horizontal slit-shaped pupils, with excellent peripheral vision; with visual fields of about 270° to 320°, sheep can see behind themselves without turning their heads. Many breeds have only short hair on the face, and some have facial wool (if any) confined to the poll and or the area of the mandibular angle; the wide angles of peripheral vision apply to these breeds. A few breeds tend to have considerable wool on the face; for some individuals of these breeds, peripheral vision may be greatly reduced by "wool blindness", unless recently shorn about the face. Sheep have poor depth perception; shadows and dips in the ground may cause sheep to baulk. In general, sheep have a tendency to move out of the dark and into well-lit areas, and prefer to move uphill when disturbed. Sheep also have an excellent sense of smell, and, like all species of their genus, have scent glands just in front of the eyes, and interdigitally on the feet. The purpose of these glands is uncertain, but those on the face may be used in breeding behaviors. The foot glands might also be related to reproduction, but alternative functions, such as secretion of a waste product or a scent marker to help lost sheep find their flock, have also been proposed.
Comparison with goats
Sheep and goats are closely related: both are in the subfamily Caprinae. However, they are separate species, so hybrids rarely occur and are always infertile. A hybrid of a ewe and a buck (a male goat) is called a sheep-goat hybrid, known as geep. Visual differences between sheep and goats include the beard of goats and divided upper lip of sheep. Sheep tails also hang down, even when short or docked, while the short tails of goats are held upwards. Also, sheep breeds are often naturally polled (either in both sexes or just in the female), while naturally polled goats are rare (though many are polled artificially). Males of the two species differ in that buck goats acquire a unique and strong odor during the rut, whereas rams do not.
Breeds
The domestic sheep is a multi-purpose animal, and the more than 200 breeds now in existence were created to serve these diverse purposes. Some sources give a count of a thousand or more breeds, but these numbers cannot be verified, according to some sources. However, several hundred breeds of sheep have been identified by the Food and Agriculture Organization of the UN (FAO), with the estimated number varying somewhat from time to time: e.g. 863 breeds as of 1993, 1314 breeds as of 1995 and 1229 breeds as of 2006. (These numbers exclude extinct breeds, which are also tallied by the FAO.) For the purpose of such tallies, the FAO definition of a breed is "either a subspecific group of domestic livestock with definable and identifiable external characteristics that enable it to be separated by visual appraisal from other similarly defined groups within the same species or a group for which geographical and/or cultural separation from phenotypically similar groups has led to acceptance of its separate identity." Almost all sheep are classified as being best suited to furnishing a certain product: wool, meat, milk, hides, or a combination in a dual-purpose breed. Other features used when classifying sheep include face color (generally white or black), tail length, presence or lack of horns, and the topography for which the breed has been developed. This last point is especially stressed in the UK, where breeds are described as either upland (hill or mountain) or lowland breeds. A sheep may also be of a fat-tailed type, which is a dual-purpose sheep common in Africa and Asia with larger deposits of fat within and around its tail.
Breeds are often categorized by the type of their wool. Fine wool breeds are those that have wool of great crimp and density, which are preferred for textiles. Most of these were derived from Merino sheep, and the breed continues to dominate the world sheep industry. Downs breeds have wool between the extremes, and are typically fast-growing meat and ram breeds with dark faces. Some major medium wool breeds, such as the Corriedale, are dual-purpose crosses of long and fine-wooled breeds and were created for high-production commercial flocks. Long wool breeds are the largest of sheep, with long wool and a slow rate of growth. Long wool sheep are most valued for crossbreeding to improve the attributes of other sheep types. For example: the American Columbia breed was developed by crossing Lincoln rams (a long wool breed) with fine-wooled Rambouillet ewes.
Coarse or carpet wool sheep are those with a medium to long length wool of characteristic coarseness. Breeds traditionally used for carpet wool show great variability, but the chief requirement is a wool that will not break down under heavy use (as would that of the finer breeds). As the demand for carpet-quality wool declines, some breeders of this type of sheep are attempting to use a few of these traditional breeds for alternative purposes. Others have always been primarily meat-class sheep.
A minor class of sheep are the dairy breeds. Dual-purpose breeds that may primarily be meat or wool sheep are often used secondarily as milking animals, but there are a few breeds that are predominantly used for milking. These sheep produce a higher quantity of milk and have slightly longer lactation curves. In the quality of their milk, the fat and protein content percentages of dairy sheep vary from non-dairy breeds, but lactose content does not.
A last group of sheep breeds is that of fur or hair sheep, which do not grow wool at all. Hair sheep are similar to the early domesticated sheep kept before woolly breeds were developed, and are raised for meat and pelts. Some modern breeds of hair sheep, such as the Dorper, result from crosses between wool and hair breeds. For meat and hide producers, hair sheep are cheaper to keep, as they do not need shearing. Hair sheep are also more resistant to parasites and hot weather.
With the modern rise of corporate agribusiness and the decline of localized family farms, many breeds of sheep are in danger of extinction. The Rare Breeds Survival Trust of the UK lists 22 native breeds as having only 3,000 registered animals (each), and The Livestock Conservancy lists 14 as either "critical" or "threatened". Preferences for breeds with uniform characteristics and fast growth have pushed heritage (or heirloom) breeds to the margins of the sheep industry. Those that remain are maintained through the efforts of conservation organizations, breed registries, and individual farmers dedicated to their preservation.
Diet
Sheep are herbivorous mammals. Most breeds prefer to graze on grass and other short roughage, avoiding the taller woody parts of plants that goats readily consume. Both sheep and goats use their lips and tongues to select parts of the plant that are easier to digest or higher in nutrition. Sheep, however, graze well in monoculture pastures where most goats fare poorly.
Ruminant system of a sheep
Like all ruminants, sheep have a complex digestive system composed of four chambers, allowing them to break down cellulose from stems, leaves, and seed hulls into simpler carbohydrates. When sheep graze, vegetation is chewed into a mass called a bolus, which is then passed into the rumen, via the reticulum. The rumen is a 19- to 38-liter (5 to 10 gallon) organ in which feed is fermented. The fermenting organisms include bacteria, fungi, and protozoa. (Other important rumen organisms include some archaea, which produce methane from carbon dioxide.) The bolus is periodically regurgitated back to the mouth as cud for additional chewing and salivation. After fermentation in the rumen, feed passes into the reticulum and the omasum; special feeds such as grains may bypass the rumen altogether. After the first three chambers, food moves into the abomasum for final digestion before processing by the intestines. The abomasum is the only one of the four chambers analogous to the human stomach, and is sometimes called the "true stomach".
Other than forage, the other staple feed for sheep is hay, often during the winter months. The ability to thrive solely on pasture (even without hay) varies with breed, but all sheep can survive on this diet. Also included in some sheep's diets are minerals, either in a trace mix or in licks. Feed provided to sheep must be specially formulated, as most cattle, poultry, pig, and even some goat feeds contain levels of copper that are lethal to sheep. The same danger applies to mineral supplements such as salt licks.
Grazing behavior
Sheep follow a diurnal pattern of activity, feeding from dawn to dusk, stopping sporadically to rest and chew their cud. Ideal pasture for sheep is not lawnlike grass, but an array of grasses, legumes and forbs. Types of land where sheep are raised vary widely, from pastures that are seeded and improved intentionally to rough, native lands. Common plants toxic to sheep are present in most of the world, and include (but are not limited to) cherry, some oaks and acorns, tomato, yew, rhubarb, potato, and rhododendron.
Effects on pasture
Sheep are largely grazing herbivores, unlike browsing animals such as goats and deer that prefer taller foliage. With a much narrower face, sheep crop plants very close to the ground and can overgraze a pasture much faster than cattle. For this reason, many shepherds use managed intensive rotational grazing, where a flock is rotated through multiple pastures, giving plants time to recover. Paradoxically, sheep can both cause and solve the spread of invasive plant species. By disturbing the natural state of pasture, sheep and other livestock can pave the way for invasive plants. However, sheep also prefer to eat invasives such as cheatgrass, leafy spurge, kudzu and spotted knapweed over native species such as sagebrush, making grazing sheep effective for conservation grazing. Research conducted in Imperial County, California compared lamb grazing with herbicides for weed control in seedling alfalfa fields. Three trials demonstrated that grazing lambs were just as effective as herbicides in controlling winter weeds. Entomologists also compared grazing lambs to insecticides for insect control in winter alfalfa. In this trial, lambs provided insect control as effectively as insecticides.
Behavior
Sheep are flock animals and strongly gregarious; much sheep behavior can be understood on the basis of these tendencies. The dominance hierarchy of sheep and their natural inclination to follow a leader to new pastures were the pivotal factors in sheep being one of the first domesticated livestock species. Furthermore, in contrast to the red deer and gazelle (two other ungulates of primary importance to meat production in prehistoric times), sheep do not defend territories although they do form home ranges. All sheep have a tendency to congregate close to other members of a flock, although this behavior varies with breed, and sheep can become stressed when separated from their flock members. During flocking, sheep have a strong tendency to follow, and a leader may simply be the first individual to move. Relationships in flocks tend to be closest among related sheep: in mixed-breed flocks, subgroups of the same breed tend to form, and a ewe and her direct descendants often move as a unit within large flocks. Sheep can become hefted to one particular local pasture (heft) so they do not roam freely in unfenced landscapes. Lambs learn the heft from ewes and if whole flocks are culled it must be retaught to the replacement animals.
Flock behaviour in sheep is generally only exhibited in groups of four or more sheep; fewer sheep may not react as expected when alone or with few other sheep. Being a prey species, the primary defense mechanism of sheep is to flee from danger when their flight zone is entered. Cornered sheep may charge and butt, or threaten by hoof stamping and adopting an aggressive posture. This is particularly true for ewes with newborn lambs.
In regions where sheep have no natural predators, none of the native breeds of sheep exhibit a strong flocking behavior.
Herding
Farmers exploit flocking behavior to keep sheep together on unfenced pastures such as hill farming, and to move them more easily. For this purpose shepherds may use herding dogs in this effort, with a highly bred herding ability. Sheep are food-oriented, and association of humans with regular feeding often results in sheep soliciting people for food. Those who are moving sheep may exploit this behavior by leading sheep with buckets of feed.
Dominance hierarchy
Sheep establish a dominance hierarchy through fighting, threats and competitiveness. Dominant animals are inclined to be more aggressive with other sheep, and usually feed first at troughs. Primarily among rams, horn size is a factor in the flock hierarchy. Rams with different size horns may be less inclined to fight to establish the dominance order, while rams with similarly sized horns are more so. Merinos have an almost linear hierarchy whereas there is a less rigid structure in Border Leicesters when a competitive feeding situation arises.
In sheep, position in a moving flock is highly correlated with social dominance, but there is no definitive study to show consistent voluntary leadership by an individual sheep.
Intelligence and learning ability
Sheep are frequently thought of as unintelligent animals. Their flocking behavior and quickness to flee and panic can make shepherding a difficult endeavor for the uninitiated. Despite these perceptions, a University of Illinois monograph on sheep reported their intelligence to be just below that of pigs and on par with that of cattle. Sheep can recognize individual human and ovine faces and remember them for years; they can remember 50 other different sheep faces for over two years; they can recognize and are attracted to individual sheep and humans by their faces, as they possess similar specialized neural systems in the temporal and frontal lobes of their brains to humans and have a greater involvement of the right brain hemisphere. In addition to long-term facial recognition of individuals, sheep can also differentiate emotional states through facial characteristics.[68][69] If worked with patiently, sheep may learn their names, and many sheep are trained to be led by halter for showing and other purposes. Sheep have also responded well to clicker training. Sheep have been used as pack animals; Tibetan nomads distribute baggage equally throughout a flock as it is herded between living sites.
It has been reported that some sheep have apparently shown problem-solving abilities; a flock in West Yorkshire, England allegedly found a way to get over cattle grids by rolling on their backs, although documentation of this has relied on anecdotal accounts.
Vocalisations
Sounds made by domestic sheep include bleats, grunts, rumbles and snorts. Bleating ("baaing") is used mostly for contact communication, especially between dam and lambs, but also at times between other flock members. The bleats of individual sheep are distinctive, enabling the ewe and her lambs to recognize each other's vocalizations. Vocal communication between lambs and their dam declines to a very low level within several weeks after parturition. A variety of bleats may be heard, depending on sheep age and circumstances. Apart from contact communication, bleating may signal distress, frustration or impatience; however, sheep are usually silent when in pain. Isolation commonly prompts bleating by sheep. Pregnant ewes may grunt when in labor. Rumbling sounds are made by the ram during courting; somewhat similar rumbling sounds may be made by the ewe, especially when with her neonate lambs. A snort (explosive exhalation through the nostrils) may signal aggression or a warning, and is often elicited from startled sheep.
Lamb
In sheep breeds lacking facial wool, the visual field is wide. In 10 sheep (Cambridge, Lleyn and Welsh Mountain breeds, which lack facial wool), the visual field ranged from 298° to 325°, averaging 313.1°, with binocular overlap ranging from 44.5° to 74°, averaging 61.7°. In some breeds, unshorn facial wool can limit the visual field; in some individuals, this may be enough to cause "wool blindness". In 60 Merinos, visual fields ranged from 219.1° to 303.0°, averaging 269.9°, and the binocular field ranged from 8.9° to 77.7°, averaging 47.5°; 36% of the measurements were limited by wool, although photographs of the experiments indicate that only limited facial wool regrowth had occurred since shearing. In addition to facial wool (in some breeds), visual field limitations can include ears and (in some breeds) horns, so the visual field can be extended by tilting the head. Sheep eyes exhibit very low hyperopia and little astigmatism. Such visual characteristics are likely to produce a well-focused retinal image of objects in both the middle and long distance. Because sheep eyes have no accommodation, one might expect the image of very near objects to be blurred, but a rather clear near image could be provided by the tapetum and large retinal image of the sheep's eye, and adequate close vision may occur at muzzle length. Good depth perception, inferred from the sheep's sure-footedness, was confirmed in "visual cliff" experiments; behavioral responses indicating depth perception are seen in lambs at one day old. Sheep are thought to have colour vision, and can distinguish between a variety of colours: black, red, brown, green, yellow and white. Sight is a vital part of sheep communication, and when grazing, they maintain visual contact with each other. Each sheep lifts its head upwards to check the position of other sheep in the flock. This constant monitoring is probably what keeps the sheep in a flock as they move along grazing. Sheep become stressed when isolated; this stress is reduced if they are provided with a mirror, indicating that the sight of other sheep reduces stress.
Taste is the most important sense in sheep, establishing forage preferences, with sweet and sour plants being preferred and bitter plants being more commonly rejected. Touch and sight are also important in relation to specific plant characteristics, such as succulence and growth form.
The ram uses his vomeronasal organ (sometimes called the Jacobson's organ) to sense the pheromones of ewes and detect when they are in estrus. The ewe uses her vomeronasal organ for early recognition of her neonate lamb.
Reproduction
Sheep follow a similar reproductive strategy to other herd animals. A group of ewes is generally mated by a single ram, who has either been chosen by a breeder or (in feral populations) has established dominance through physical contest with other rams. Most sheep are seasonal breeders, although some are able to breed year-round. Ewes generally reach sexual maturity at six to eight months old, and rams generally at four to six months. However, there are exceptions. For example, Finnsheep ewe lambs may reach puberty as early as 3 to 4 months, and Merino ewes sometimes reach puberty at 18 to 20 months. Ewes have estrus cycles about every 17 days, during which they emit a scent and indicate readiness through physical displays towards rams.
In feral sheep, rams may fight during the rut to determine which individuals may mate with ewes. Rams, especially unfamiliar ones, will also fight outside the breeding period to establish dominance; rams can kill one another if allowed to mix freely. During the rut, even usually friendly rams may become aggressive towards humans due to increases in their hormone levels.
After mating, sheep have a gestation period of about five months, and normal labor takes one to three hours. Although some breeds regularly throw larger litters of lambs, most produce single or twin lambs. During or soon after labor, ewes and lambs may be confined to small lambing jugs, small pens designed to aid both careful observation of ewes and to cement the bond between them and their lambs.
A lamb's first steps
Ovine obstetrics can be problematic. By selectively breeding ewes that produce multiple offspring with higher birth weights for generations, sheep producers have inadvertently caused some domestic sheep to have difficulty lambing; balancing ease of lambing with high productivity is one of the dilemmas of sheep breeding. In the case of any such problems, those present at lambing may assist the ewe by extracting or repositioning lambs. After the birth, ewes ideally break the amniotic sac (if it is not broken during labor), and begin licking clean the lamb. Most lambs will begin standing within an hour of birth. In normal situations, lambs nurse after standing, receiving vital colostrum milk. Lambs that either fail to nurse or are rejected by the ewe require help to survive, such as bottle-feeding or fostering by another ewe.
Most lambs begin life being born outdoors. After lambs are several weeks old, lamb marking (ear tagging, docking, mulesing, and castrating) is carried out. Vaccinations are usually carried out at this point as well. Ear tags with numbers are attached, or ear marks are applied, for ease of later identification of sheep. Docking and castration are commonly done after 24 hours (to avoid interference with maternal bonding and consumption of colostrum) and are often done not later than one week after birth, to minimize pain, stress, recovery time and complications. The first course of vaccinations (commonly anti-clostridial) is commonly given at an age of about 10 to 12 weeks; i.e. when the concentration of maternal antibodies passively acquired via colostrum is expected to have fallen low enough to permit development of active immunity. Ewes are often revaccinated annually about 3 weeks before lambing, to provide high antibody concentrations in colostrum during the first several hours after lambing. Ram lambs that will either be slaughtered or separated from ewes before sexual maturity are not usually castrated. Objections to all these procedures have been raised by animal rights groups, but farmers defend them by saying they save money, and inflict only temporary pain.
Homosexuality
Sheep are the only species of mammal except for humans which exhibits exclusive homosexual behavior. About 10% of rams refuse to mate with ewes but readily mate with other rams, and thirty percent of all rams demonstrate at least some homosexual behavior. Additionally, a small number of females that were accompanied by a male fetus in utero (i.e. as fraternal twins) are freemartins (female animals that are behaviorally masculine and lack functioning ovaries).
Health
Sheep may fall victim to poisons, infectious diseases, and physical injuries. As a prey species, a sheep's system is adapted to hide the obvious signs of illness, to prevent being targeted by predators. However, some signs of ill health are obvious, with sick sheep eating little, vocalizing excessively, and being generally listless. Throughout history, much of the money and labor of sheep husbandry has aimed to prevent sheep ailments. Historically, shepherds often created remedies by experimentation on the farm. In some developed countries, including the United States, sheep lack the economic importance for drug companies to perform expensive clinical trials required to approve more than a relatively limited number of drugs for ovine use. However, extra-label drug use in sheep production is permitted in many jurisdictions, subject to certain restrictions. In the US, for example, regulations governing extra-label drug use in animals are found in 21 CFR (Code of Federal Regulations) Part 530. In the 20th and 21st centuries, a minority of sheep owners have turned to alternative treatments such as homeopathy, herbalism and even traditional Chinese medicine to treat sheep veterinary problems. Despite some favorable anecdotal evidence, the effectiveness of alternative veterinary medicine has been met with skepticism in scientific journals. The need for traditional anti-parasite drugs and antibiotics is widespread, and is the main impediment to certified organic farming with sheep.
Many breeders take a variety of preventive measures to ward off problems. The first is to ensure all sheep are healthy when purchased. Many buyers avoid outlets known to be clearing houses for animals culled from healthy flocks as either sick or simply inferior. This can also mean maintaining a closed flock, and quarantining new sheep for a month. Two fundamental preventive programs are maintaining good nutrition and reducing stress in the sheep. Restraint, isolation, loud noises, novel situations, pain, heat, extreme cold, fatigue and other stressors can lead to secretion of cortisol, a stress hormone, in amounts that may indicate welfare problems. Excessive stress can compromise the immune system. "Shipping fever" (pneumonic mannheimiosis, formerly called pasteurellosis) is a disease of particular concern, that can occur as a result of stress, notably during transport and (or) handling. Pain, fear and several other stressors can cause secretion of epinephrine (adrenaline). Considerable epinephrine secretion in the final days before slaughter can adversely affect meat quality (by causing glycogenolysis, removing the substrate for normal post-slaughter acidification of meat) and result in meat becoming more susceptible to colonization by spoilage bacteria. Because of such issues, low-stress handling is essential in sheep management. Avoiding poisoning is also important; common poisons are pesticide sprays, inorganic fertilizer, motor oil, as well as radiator coolant containing ethylene glycol.
Common forms of preventive medication for sheep are vaccinations and treatments for parasites. Both external and internal parasites are the most prevalent malady in sheep, and are either fatal, or reduce the productivity of flocks. Worms are the most common internal parasites. They are ingested during grazing, incubate within the sheep, and are expelled through the digestive system (beginning the cycle again). Oral anti-parasitic medicines, known as drenches, are given to a flock to treat worms, sometimes after worm eggs in the feces has been counted to assess infestation levels. Afterwards, sheep may be moved to a new pasture to avoid ingesting the same parasites. External sheep parasites include: lice (for different parts of the body), sheep keds, nose bots, sheep itch mites, and maggots. Keds are blood-sucking parasites that cause general malnutrition and decreased productivity, but are not fatal. Maggots are those of the bot fly and the blow-fly, commonly Lucilia sericata or its relative L. cuprina. Fly maggots cause the extremely destructive condition of flystrike. Flies lay their eggs in wounds or wet, manure-soiled wool; when the maggots hatch they burrow into a sheep's flesh, eventually causing death if untreated. In addition to other treatments, crutching (shearing wool from a sheep's rump) is a common preventive method. Some countries allow mulesing, a practice that involves stripping away the skin on the rump to prevent fly-strike, normally performed when the sheep is a lamb. Nose bots are fly larvae that inhabit a sheep's sinuses, causing breathing difficulties and discomfort. Common signs are a discharge from the nasal passage, sneezing, and frantic movement such as head shaking. External parasites may be controlled through the use of backliners, sprays or immersive sheep dips.
A wide array of bacterial and viral diseases affect sheep. Diseases of the hoof, such as foot rot and foot scald may occur, and are treated with footbaths and other remedies. Foot rot is present in over 97% of flocks in the UK. These painful conditions cause lameness and hinder feeding. Ovine Johne's disease is a wasting disease that affects young sheep. Bluetongue disease is an insect-borne illness causing fever and inflammation of the mucous membranes. Ovine rinderpest (or peste des petits ruminants) is a highly contagious and often fatal viral disease affecting sheep and goats. Sheep may also be affected by primary or secondary photosensitization. Tetanus can also afflict sheep through wounds from shearing, docking, castration, or vaccination. The organism also can be introduced into the reproductive tract by unsanitary humans who assist ewes during lambing.
A few sheep conditions are transmissible to humans. Orf (also known as scabby mouth, contagious ecthyma or soremouth) is a skin disease leaving lesions that is transmitted through skin-to-skin contact. Cutaneous anthrax is also called woolsorter's disease, as the spores can be transmitted in unwashed wool. More seriously, the organisms that can cause spontaneous enzootic abortion in sheep are easily transmitted to pregnant women. Also of concern are the prion disease scrapie and the virus that causes foot-and-mouth disease (FMD), as both can devastate flocks. The latter poses a slight risk to humans. During the 2001 FMD pandemic in the UK, hundreds of sheep were culled and some rare British breeds were at risk of extinction due to this.
Of the 600,300 sheep lost to the US economy in 2004, 37.3% were lost to predators, while 26.5% were lost to some form of disease. Poisoning accounted for 1.7% of non-productive deaths.
Predators
A lamb being attacked by coyotes with a bite to the throat
Other than parasites and disease, predation is a threat to sheep and the profitability of sheep raising. Sheep have little ability to defend themselves, compared with other species kept as livestock. Even if sheep survive an attack, they may die from their injuries or simply from panic. However, the impact of predation varies dramatically with region. In Africa, Australia, the Americas, and parts of Europe and Asia predators are a serious problem. In the United States, for instance, over one third of sheep deaths in 2004 were caused by predation. In contrast, other nations are virtually devoid of sheep predators, particularly islands known for extensive sheep husbandry. Worldwide, canids—including the domestic dog—are responsible for most sheep deaths. Other animals that occasionally prey on sheep include: felines, bears, birds of prey, ravens and feral hogs.
Sheep producers have used a wide variety of measures to combat predation. Pre-modern shepherds used their own presence, livestock guardian dogs, and protective structures such as barns and fencing. Fencing (both regular and electric), penning sheep at night and lambing indoors all continue to be widely used. More modern shepherds used guns, traps, and poisons to kill predators, causing significant decreases in predator populations. In the wake of the environmental and conservation movements, the use of these methods now usually falls under the purview of specially designated government agencies in most developed countries.
The 1970s saw a resurgence in the use of livestock guardian dogs and the development of new methods of predator control by sheep producers, many of them non-lethal. Donkeys and guard llamas have been used since the 1980s in sheep operations, using the same basic principle as livestock guardian dogs. Interspecific pasturing, usually with larger livestock such as cattle or horses, may help to deter predators, even if such species do not actively guard sheep. In addition to animal guardians, contemporary sheep operations may use non-lethal predator deterrents such as motion-activated lights and noisy alarms.
Economic importance
Main article: Agricultural economics
Global sheep stock
in 2019
Number in millions
1. China163.5 (13.19%)
2. India74.3 (5.99%)
3. Australia65.8 (5.31%)
4. Nigeria46.9 (3.78%)
5. Iran41.3 (3.33%)
6. Sudan40.9 (3.3%)
7. Chad35.9 (2.9%)
8. Turkey35.2 (2.84%)
9. United Kingdom33.6 (2.71%)
10. Mongolia32.3 (2.61%)
World total1,239.8
Source: UN Food and Agriculture Organization
Sheep are an important part of the global agricultural economy. However, their once vital status has been largely replaced by other livestock species, especially the pig, chicken, and cow. China, Australia, India, and Iran have the largest modern flocks, and serve both local and exportation needs for wool and mutton. Other countries such as New Zealand have smaller flocks but retain a large international economic impact due to their export of sheep products. Sheep also play a major role in many local economies, which may be niche markets focused on organic or sustainable agriculture and local food customers. Especially in developing countries, such flocks may be a part of subsistence agriculture rather than a system of trade. Sheep themselves may be a medium of trade in barter economies.
Domestic sheep provide a wide array of raw materials. Wool was one of the first textiles, although in the late 20th century wool prices began to fall dramatically as the result of the popularity and cheap prices for synthetic fabrics. For many sheep owners, the cost of shearing is greater than the possible profit from the fleece, making subsisting on wool production alone practically impossible without farm subsidies. Fleeces are used as material in making alternative products such as wool insulation. In the 21st century, the sale of meat is the most profitable enterprise in the sheep industry, even though far less sheep meat is consumed than chicken, pork or beef.
Sheepskin is likewise used for making clothes, footwear, rugs, and other products. Byproducts from the slaughter of sheep are also of value: sheep tallow can be used in candle and soap making, sheep bone and cartilage has been used to furnish carved items such as dice and buttons as well as rendered glue and gelatin. Sheep intestine can be formed into sausage casings, and lamb intestine has been formed into surgical sutures, as well as strings for musical instruments and tennis rackets. Sheep droppings, which are high in cellulose, have even been sterilized and mixed with traditional pulp materials to make paper. Of all sheep byproducts, perhaps the most valuable is lanolin: the waterproof, fatty substance found naturally in sheep's wool and used as a base for innumerable cosmetics and other products.
Some farmers who keep sheep also make a profit from live sheep. Providing lambs for youth programs such as 4-H and competition at agricultural shows is often a dependable avenue for the sale of sheep. Farmers may also choose to focus on a particular breed of sheep in order to sell registered purebred animals, as well as provide a ram rental service for breeding. A new option for deriving profit from live sheep is the rental of flocks for grazing; these "mowing services" are hired in order to keep unwanted vegetation down in public spaces and to lessen fire hazard.
Despite the falling demand and price for sheep products in many markets, sheep have distinct economic advantages when compared with other livestock. They do not require expensive housing, such as that used in the intensive farming of chickens or pigs. They are an efficient use of land; roughly six sheep can be kept on the amount that would suffice for a single cow or horse. Sheep can also consume plants, such as noxious weeds, that most other animals will not touch, and produce more young at a faster rate. Also, in contrast to most livestock species, the cost of raising sheep is not necessarily tied to the price of feed crops such as grain, soybeans and corn. Combined with the lower cost of quality sheep, all these factors combine to equal a lower overhead for sheep producers, thus entailing a higher profitability potential for the small farmer. Sheep are especially beneficial for independent producers, including family farms with limited resources, as the sheep industry is one of the few types of animal agriculture that has not been vertically integrated by agribusiness. However, small flocks, from 10 to 50 ewes, often are not profitable because they tend to be poorly managed. The primary reason is that mechanization is not feasible, so return per hour of labor is not maximized. Small farm flocks generally are used simply to control weeds on irrigation ditches or maintained as a hobby.
Shoulder of lamb
Sheep meat and milk were one of the earliest staple proteins consumed by human civilization after the transition from hunting and gathering to agriculture. Sheep meat prepared for food is known as either mutton or lamb, and approximately 540 million sheep are slaughtered each year for meat worldwide. "Mutton" is derived from the Old French moton, which was the word for sheep used by the Anglo-Norman rulers of much of the British Isles in the Middle Ages. This became the name for sheep meat in English, while the Old English word sceap was kept for the live animal. Throughout modern history, "mutton" has been limited to the meat of mature sheep usually at least two years of age; "lamb" is used for that of immature sheep less than a year.
In the 21st century, the nations with the highest consumption of sheep meat are the Arab states of the Persian Gulf, New Zealand, Australia, Greece, Uruguay, the United Kingdom and Ireland. These countries eat 14–40 lbs (3–18 kg) of sheep meat per capita, per annum. Sheep meat is also popular in France, Africa (especially the Arab world), the Caribbean, the rest of the Middle East, India, and parts of China. This often reflects a history of sheep production. In these countries in particular, dishes comprising alternative cuts and offal may be popular or traditional. Sheep testicles—called animelles or lamb fries—are considered a delicacy in many parts of the world. Perhaps the most unusual dish of sheep meat is the Scottish haggis, composed of various sheep innards cooked along with oatmeal and chopped onions inside its stomach. In comparison, countries such as the U.S. consume only a pound or less (under 0.5 kg), with Americans eating 50 pounds (22 kg) of pork and 65 pounds (29 kg) of beef. In addition, such countries rarely eat mutton, and may favor the more expensive cuts of lamb: mostly lamb chops and leg of lamb.
Though sheep's milk may be drunk rarely in fresh form, today it is used predominantly in cheese and yogurt making. Sheep have only two teats, and produce a far smaller volume of milk than cows. However, as sheep's milk contains far more fat, solids, and minerals than cow's milk, it is ideal for the cheese-making process. It also resists contamination during cooling better because of its much higher calcium content. Well-known cheeses made from sheep milk include the feta of Bulgaria and Greece, Roquefort of France, Manchego from Spain, the pecorino romano (the Italian word for "sheep" is pecore) and ricotta of Italy. Yogurts, especially some forms of strained yogurt, may also be made from sheep milk. Many of these products are now often made with cow's milk, especially when produced outside their country of origin. Sheep milk contains 4.8% lactose, which may affect those who are intolerant.
As with other domestic animals, the meat of uncastrated males is inferior in quality, especially as they grow. A "bucky" lamb is a lamb which was not castrated early enough, or which was castrated improperly (resulting in one testicle being retained). These lambs are worth less at market.
In science
Sheep are generally too large and reproduce too slowly to make ideal research subjects, and thus are not a common model organism. They have, however, played an influential role in some fields of science. In particular, the Roslin Institute of Edinburgh, Scotland used sheep for genetics research that produced groundbreaking results. In 1995, two ewes named Megan and Morag were the first mammals cloned from differentiated cells, also referred to as gynomerogony. A year later, a Finnish Dorset sheep named Dolly, dubbed "the world's most famous sheep" in Scientific American, was the first mammal to be cloned from an adult somatic cell. Following this, Polly and Molly were the first mammals to be simultaneously cloned and transgenic.
As of 2008, the sheep genome has not been fully sequenced, although a detailed genetic map has been published, and a draft version of the complete genome produced by assembling sheep DNA sequences using information given by the genomes of other mammals. In 2012, a transgenic sheep named "Peng Peng" was cloned by Chinese scientists, who spliced his genes with that of a roundworm (C. elegans) in order to increase production of fats healthier for human consumption.
In the study of natural selection, the population of Soay sheep that remain on the island of Hirta have been used to explore the relation of body size and coloration to reproductive success. Soay sheep come in several colors, and researchers investigated why the larger, darker sheep were in decline; this occurrence contradicted the rule of thumb that larger members of a population tend to be more successful reproductively. The feral Soays on Hirta are especially useful subjects because they are isolated.
Domestic sheep are sometimes used in medical research, particularly for researching cardiovascular physiology, in areas such as hypertension and heart failure. Pregnant sheep are also a useful model for human pregnancy, and have been used to investigate the effects on fetal development of malnutrition and hypoxia. In behavioral sciences, sheep have been used in isolated cases for the study of facial recognition, as their mental process of recognition is qualitatively similar to humans.
Cultural impact
Sheep have had a strong presence in many cultures, especially in areas where they form the most common type of livestock. In the English language, to call someone a sheep or ovine may allude that they are timid and easily led. In contradiction to this image, male sheep are often used as symbols of virility and power; the logos of the Los Angeles Rams football team and the Dodge Ram pickup truck allude to males of the bighorn sheep, Ovis canadensis.
Counting sheep is popularly said to be an aid to sleep, and some ancient systems of counting sheep persist today. Sheep also enter in colloquial sayings and idiom frequently with such phrases as "black sheep". To call an individual a black sheep implies that they are an odd or disreputable member of a group. This usage derives from the recessive trait that causes an occasional black lamb to be born into an entirely white flock. These black sheep were considered undesirable by shepherds, as black wool is not as commercially viable as white wool. Citizens who accept overbearing governments have been referred to by the Portmanteau neologism of sheeple. Somewhat differently, the adjective "sheepish" is also used to describe embarrassment.
In heraldry
In British heraldry, sheep appear in the form of rams, sheep proper and lambs. These are distinguished by the ram being depicted with horns and a tail, the sheep with neither and the lamb with its tail only. A further variant of the lamb, termed the Paschal lamb, is depicted as carrying a Christian cross and with a halo over its head. Rams' heads, portrayed without a neck and facing the viewer, are also found in British armories. The fleece, depicted as an entire sheepskin carried by a ring around its midsection, originally became known through its use in the arms of the Order of the Golden Fleece and was later adopted by towns and individuals with connections to the wool industry. A sheep on a blue field is depicted on the greater/royal arms of the king of Denmark to represent the Faroe Islands. In 2004 a modernized arms has been adopted by the Faroe Islands, which based on a 15th century coat of arms.
Religion and folklore
In antiquity, symbolism involving sheep cropped up in religions in the ancient Near East, the Mideast, and the Mediterranean area: Çatalhöyük, ancient Egyptian religion, the Cana'anite and Phoenician tradition, Judaism, Greek religion, and others. Religious symbolism and ritual involving sheep began with some of the first known faiths: Skulls of rams (along with bulls) occupied central placement in shrines at the Çatalhöyük settlement in 8,000 BCE. In Ancient Egyptian religion, the ram was the symbol of several gods: Khnum, Heryshaf and Amun (in his incarnation as a god of fertility). Other deities occasionally shown with ram features include the goddess Ishtar, the Phoenician god Baal-Hamon, and the Babylonian god Ea-Oannes. In Madagascar, sheep were not eaten as they were believed to be incarnations of the souls of ancestors.
There are many ancient Greek references to sheep: that of Chrysomallos, the golden-fleeced ram, continuing to be told through into the modern era. Astrologically, Aries, the ram, is the first sign of the classical Greek zodiac, and the sheep is the eighth of the twelve animals associated with the 12-year cycle of in the Chinese zodiac, related to the Chinese calendar. It is said in Chinese traditions that Hou ji sacrificed sheep. Mongolia, shagai are an ancient form of dice made from the cuboid bones of sheep that are often used for fortunetelling purposes.
Sheep play an important role in all the Abrahamic faiths; Abraham, Isaac, Jacob, Moses, and King David were all shepherds. According to the Biblical story of the Binding of Isaac, a ram is sacrificed as a substitute for Isaac after an angel stays Abraham's hand (in the Islamic tradition, Abraham was about to sacrifice Ishmael). Eid al-Adha is a major annual festival in Islam in which sheep (or other animals) are sacrificed in remembrance of this act. Sheep are occasionally sacrificed to commemorate important secular events in Islamic cultures. Greeks and Romans sacrificed sheep regularly in religious practice, and Judaism once sacrificed sheep as a Korban (sacrifice), such as the Passover lamb. Ovine symbols—such as the ceremonial blowing of a shofar—still find a presence in modern Judaic traditions.
Collectively, followers of Christianity are often referred to as a flock, with Christ as the Good Shepherd, and sheep are an element in the Christian iconography of the birth of Jesus. Some Christian saints are considered patrons of shepherds, and even of sheep themselves. Christ is also portrayed as the Sacrificial lamb of God (Agnus Dei) and Easter celebrations in Greece and Romania traditionally feature a meal of Paschal lamb. A church leader is often called the pastor, which is derived from the Latin word for shepherd. In many western Christian traditions bishops carry a staff, which also serves as a symbol of the episcopal office, known as a crosier, which is modeled on the shepherd's crook.
Sheep are key symbols in fables and nursery rhymes like The Wolf in Sheep's Clothing, Little Bo Peep, Baa, Baa, Black Sheep, and Mary Had a Little Lamb; novels such as George Orwell's Animal Farm and Haruki Murakami's A Wild Sheep Chase; songs such as Bach's Sheep may safely graze (Schafe können sicher weiden) and Pink Floyd's "Sheep", and poems like William Blake's "The Lamb".
Nkumbe - Sentinels
Nkumbe - Wächter
Klipspringer
Klippspringer
Kruger National Park is one of the largest game reserves in Africa. It covers an area of 19,485 km2 (7,523 sq mi) in the provinces of Limpopo and Mpumalanga in northeastern South Africa, and extends 360 km (220 mi) from north to south and 65 km (40 mi) from east to west. The administrative headquarters are in Skukuza. Areas of the park were first protected by the government of the South African Republic in 1898, and it became South Africa's first national park in 1926.
To the west and south of the Kruger National Park are the two South African provinces of Limpopo and Mpumalanga. In the north is Zimbabwe, and to the east is Mozambique. It is now part of the Great Limpopo Transfrontier Park, a peace park that links Kruger National Park with the Gonarezhou National Park in Zimbabwe, and with the Limpopo National Park in Mozambique.
The park is part of the Kruger to Canyons Biosphere an area designated by the United Nations Educational, Scientific and Cultural Organization (UNESCO) as an International Man and Biosphere Reserve (the "Biosphere").
The park has nine main gates allowing entrance to the different camps.
(Wikipedia)
The klipspringer (/ˈklɪpˌsprɪŋər/; Oreotragus oreotragus) is a small antelope found in eastern and southern Africa. The sole member of its genus, the klipspringer was first described by German zoologist Eberhard August Wilhelm von Zimmermann in 1783. The klipspringer is a small, sturdy antelope; it reaches 43–60 centimetres (17–24 in) at the shoulder and weighs from 8 to 18 kilograms (18 to 40 lb). The coat of the klipspringer, yellowish gray to reddish brown, acts as an efficient camouflage in its rocky habitat. Unlike most other antelopes, the klipspringer has a thick and coarse coat with hollow, brittle hairs. The horns, short and spiky, typically measure 7.5–9 centimetres (3.0–3.5 in).
Typically nocturnal, the klipspringer rests during the middle of the day and late at night. A gregarious animal, the klipspringer is monogamous to a much greater extent than other antelopes; individuals of opposite sexes exhibit long-term to lifelong pair bonding. The mates tend to stay as close as within 5 metres (16 ft) of each other at most times. Males form territories, 7.5–49 hectares (19–121 acres), in which they stay with their partners and offspring. Primarily a browser, the klipspringer prefers young plants, fruits and flowers. Gestation lasts around six months, following which a single calf is born; births peak from spring to early summer. The calf leaves its mother when it turns a year old.
The klipspringer inhabits places characterised by rocky terrain and sparse vegetation. Its range extends from northeastern Sudan, Eritrea, northern Somalia and Ethiopia in the east to South Africa in the south, and along coastal Angola and Namibia. The International Union for the Conservation of Nature and Natural Resources (IUCN) classifies the klipspringer as Least Concern. There are no major threats to the survival of the klipspringer, as its habitat is inaccessible and unfavourable for hunting. Significant numbers occur on private farmlands. As of 2008, nearly 25% of the populations occur in protected areas throughout its range.
The klipspringer is a small, sturdy antelope reaching 43–60 centimetres (17–24 in) at the shoulder. The head-and-body length is typically between 75 and 115 centimetres (30 and 45 in). It weighs from 8 to 18 kilograms (18 to 40 lb). The klipspringer is sexually dimorphic; females are slightly larger and heavier than the males. The tail measures 6.5–10.5 centimetres (2.6–4.1 in). Prominent facial features include the brown forehead, short ears marked with black, prominent preorbital glands near the eyes, and white lips and chin. The horns, short and spiky, present only on males, typically measure 7.5–9 centimetres (3.0–3.5 in); the maximum recorded horn length is 15.9 centimetres (6.3 in).
The coat of the klipspringer, yellowish gray to reddish brown, acts as an efficient camouflage in its rocky habitat; the underbelly is white. Unlike most other antelopes, the klipspringer has a thick and coarse coat with hollow, brittle hairs. The incisors might even get damaged by the hairs while grooming. However, the coat is a significant adaptation that saves the animal during steep falls and provides effective insulation in the extreme climates characteristic of its mountain habitat. A study showed that ticks occur in larger numbers on the underbelly, where the hair is less coarse. The hair often turns erect, especially if the animal is ill or if its temperature increases. Another feature unique to the klipspringer is its gait; it walks on the tips of its cylindrical, blunt hooves. This enhances the grip on the ground, enabling the animal to deftly climb and jump over rocky surfaces.
The subspecies vary in coat colour – from golden yellow in the Cape klipspringer, Ethiopian klipspringer, golden klipspringer and Transvaal klipspringer to ochre or rufous in the Maasai klipspringer, Stevenson's klipspringer and Zambian klipspringer. Cape klipspringer populations tend to have the largest males, while Maasai klipspringer exhibit the largest females.
Typically nocturnal (active mainly at night), the klipspringer rests during the midday and at late night; the animal tends to be more active on moonlit nights. It basks in the morning sunlight to warm itself. A gregarious animal, the klipspringer, like the dik-diks and the oribi, exhibits monogamy to a much greater extent than other antelopes; individuals of opposite sexes form pairs that might last until one dies.[ The mates tend to stay as close as within 5 metres (16 ft) of each other at most times; for instance, they take turns at keeping a lookout for predators while the other feeds, and face any danger together. The klipspringer will hop a few metres away from the danger. Other social groups include small family herds of 8 or more members or solitary individuals. Klipspringer greet one another by rubbing cheeks at social meetings.
Males form territories, 7.5–49 hectares (19–121 acres) large (the size depends on rainfall patterns), in which they stay with their partners and offspring. Males are generally more vigilant than females. Klipspringer form large dung heaps, nearly 1 metre (3.3 ft) across and 10 centimetres (3.9 in) deep, at the borders of territories; another form of marking is the secretion of a thick, black substance, measuring 5 millimetres (0.20 in) across, from the preorbital glands onto vegetation and rocks in the territories. A study revealed that the tick Ixodes neitzi detects and aggregates on twigs marked by the klipspringer. Another study showed that plants near the borders with neighbouring territories are particularly preferred for marking. The main vocalisation is a shrill whistle, given out by the klipspringer pair in a duet, as a means of communication or anti-predator response. Predators include the baboon, black-backed jackal, caracal, eagle, leopard, martial eagle, serval, spotted hyaena and Verreaux's eagle. Birds such as familiar chats, pale-winged starlings, red-winged starlings and yellow-bellied bulbuls have been observed feeding on ectoparasites of klipspringer.
Primarily a browser, the klipspringer prefers young plants, fruits and flowers. Grasses, eaten mainly in the wet season, form a minor portion of the diet. Some plants, such as Vellozia, may be preferred seasonally. Klipspringer depend mainly on succulent plants, and not on water bodies, to meet their water requirement. They can stand on their hindlegs to reach tall branches up to 1.2 metres (3.9 ft) above the ground; some individuals in Namibia were observed climbing Faidherbia albida trees up to a height of 5.4 metres (18 ft).
The klipspringer is a seasonal breeder; the time when mating occurs varies geographically. Females become sexually mature by the time they are a year old; males take slightly longer to mature. Mating behaviour has not been extensively observed. Gestation lasts around six months, following which a single calf, weighing slightly more than 1 kilogram (2.2 lb), is born; births peak from spring to early summer. Births take place in dense vegetation. The newborn is carefully hidden for up to three months to protect it from the view of predators; the mother suckles it three to four times a day, the visits gradually lengthen as the offspring grows. Males are protective of their offspring, keeping a watch for other males and predators. The calf is weaned at four to five months, and leaves its mother when it turns a year old. The klipspringer lives for around 15 years.
The klipspringer inhabits places characterised by rocky terrain and sparse vegetation. It migrates to lowlands at times of food scarcity. The klipspringer occurs at altitudes as high as 4,500 metres (14,800 ft) on Mount Kilimanjaro. The klipspringer can occur at high population densities in favourable habitats extending over a large area; 10 to 14 individuals occur per square kilometre in the Simien Mountains National Park, Ethiopia. However, the habitat is typically rocky over long stretches and grassy terrain is discontinuous; consequently the population density is typically between 0.01 and 0.1 individual per square kilometre.
The antelope occurs in significant numbers across eastern and southern Africa; its range extends from northeastern Sudan, Eritrea, northern Somalia and Ethiopia in the east to South Africa in the south, and along coastal Angola and Namibia. Smaller populations occur in the northern and western highlands of Central African Republic, southeastern Democratic Republic of Congo, Jos Plateau and east of Gashaka Gumti National Park in Nigeria. It is feared to be extinct in Burundi.
The International Union for the Conservation of Nature and Natural Resources (IUCN) classifies the klipspringer as Least Concern. The klipspringer is hunted for its meat, leather and hair. However, there are no major threats to the survival of the klipspringer, as its habitat is inaccessible and unfavourable for hunting. Moreover, the antelope does not have to compete with livestock, that do not frequent montane areas. However, populations at lower altitudes are more vulnerable to elimination.
In 1999, Rod East of the IUCN SSC Antelope Specialist Group estimated the total population of klipspringer at 42,000. Significant numbers occur on private farmlands. As of 2008, nearly 25% of the populations occur in protected areas such as the Simien and Bale Mountains National Parks (Ethiopia); Tsavo East and West National Parks (Kenya); North and South Luangwa National Parks (Zambia); Nyika National Park (Malawi); Namib-Naukluft National Park (Namibia); and Matobo National Park (Zimbabwe).
(Wikipedia)
Der Kruger-Nationalpark (deutsch häufig falsch Krüger-Nationalpark) ist das größte Wildschutzgebiet Südafrikas. Er liegt im Nordosten des Landes in der Landschaft des Lowveld auf dem Gebiet der Provinz Limpopo sowie des östlichen Abschnitts von Mpumalanga. Seine Fläche erstreckt sich vom Crocodile-River im Süden bis zum Limpopo, dem Grenzfluss zu Simbabwe, im Norden. Die Nord-Süd-Ausdehnung beträgt etwa 350 km, in Ost-West-Richtung ist der Park durchschnittlich 54 km breit und umfasst eine Fläche von rund 20.000 Quadratkilometern. Damit gehört er zu den größten Nationalparks in Afrika.
Das Schutzgebiet wurde am 26. März 1898 unter dem Präsidenten Paul Kruger als Sabie Game Reserve zum Schutz der Wildnis gegründet. 1926 erhielt das Gebiet den Status Nationalpark und wurde in seinen heutigen Namen umbenannt. Im Park leben 147 Säugetierarten inklusive der „Big Five“, außerdem etwa 507 Vogelarten und 114 Reptilienarten, 49 Fischarten und 34 Amphibienarten.
(Wikipedia)
Die Klippspringer (Oreotragus) sind eine kleine afrikanische Antilopengattung und gleichzeitig eine monotypische Tribus. Der Name stammt aus dem Afrikaans – niederländisch klipspringer bedeutet „Felsenspringer“.
Die Klippspringer gehören zu den zierlichsten Vertretern der Antilopen und erreichen eine Schulterhöhe von knapp 60 cm und ein Körpergewicht von 10–13 kg. Die Weibchen sind dabei etwas größer und schwerer als die Männchen. Die Männchen tragen darüber hinaus kurze, gerade, weit auseinanderstehende Spieße, von einer Länge von ungefähr 20–25 cm. Bei den nördlichen Beständen tragen auch die Weibchen Hörner.
Mit ihrem groben, borstigen, dichten gelb bis braungrauen Fell, das eine schwarze Melierung aufweist und auf der Unterseite weiß ist, sind sie vor den „Kopjes“ (Felsblöcken) kaum zu entdecken. Der Bauch, das Kinn und die Lippen werden von weißen oder cremefarbenen Haar bedeckt.
Die Klippspringer weisen zwei Charakteristika auf, die eine Anpassung an das Leben in einem felsigen Habitat sind. Sie sind die einzigen Antilopen, die auf den Hufspitzen laufen. Diese werden so abgelaufen, dass zylindrische Hufe entstehen, die eine unverwechselbare, zweikreisige Spur hinterlassen. Durch diese spezifische Form ihrer Hufe haben Klippspringer einen besseren Halt auf glatten Felsoberflächen. Als zweite Anpassung ist ihr Fell fest und dick. Jedes einzelne Haar ist hohl und borstenähnlich. Das Fell bietet eine gute Isolation gegen die Temperaturextreme ihrer hochgelegenen Lebensräume. Es isoliert sowohl gegenüber extremer Hitze als auch gegen große Kälte.
Diese Antilopengattung ist vom südafrikanischen Kap der Guten Hoffnung über Ostafrika bis nach Äthiopien zu finden. Sie kommt unter anderem in folgenden Naturschutzgebieten vor: Bale-Mountains-Nationalpark, Karoo, Mountain Zebra, Giant’s Castle, Krügerpark, Namib-Naukluft, Hwange, Upemba, Tsavo, Manyarasee, Ngorongoro, Serengeti, Kilimandscharo und Samburu.
Klippspringer halten sich generell auf oder in der Nähe von felsigen Hügeln oder Felsformationen auf. Häufig kann man sie bewegungslos auf der Spitze hoher Felsen entdecken. Oft kann man den lauten, nasalen Warnpfiff der Tiere hören. Auch im unwegsamen Gelände sind sie sehr flink. Bei Gefahr flüchten sie in der Regel nicht weit, sondern bleiben nach kurzer Flucht stehen und beobachten den Prädator. Dabei ist häufig der Warnpfiff zu hören. Dieses Verhalten trägt allerdings auch dazu bei, dass sich Klippspringer durch den Menschen leicht erjagen lassen.
Man findet sowohl Pärchen als auch Familiengruppen; die Männchen verteidigen ihr Revier. Die Paarbeziehungen enden in der Regel erst mit dem Tod eines der beiden Partner. Zu den typischen Verhaltensmerkmalen der Klippspringer gehören die großen Kothaufen, die sich meist am Rand des Revieres festlegen. Weibchen reiben nach dem Koten an nahen Zweigen ihre Voraugendrüsen ab, wodurch eine dicke, schwarze Substanz auf die Zweige übertragen wird. Diese strömen einen starken süßen Duft aus, wenn sie feucht sind. Gewöhnlich setzt das Männchen seine Duftmarke auf die des Weibchens. Die Klippspringer ernähren sich vorwiegend von Laub.
Die Brunstzeit der Klippspringer dauert von September bis Januar. Die Tragzeit beträgt 214 Tage. Das einzelne Junge wiegt bei der Geburt etwa 1 kg. Ihre Lebenserwartung beträgt 10 bis 12 Jahre.
In verschiedenen Gegenden Afrikas wurden Klippspringer von jeher gejagt. Aus ihrer Haut lässt sich ein Leder gewinnen, das zur Herstellung von Taschen verwendet wird. Das Fleisch soll sehr wohlschmeckend sein. Durch die Bejagung ist der Klippspringer im westafrikanischen Teil seines Verbreitungsgebiets nahezu ausgestorben (Unterart O. o. porteousi). Im übrigen Verbreitungsgebiet gilt er nicht als bedroht.
Da in den letzten Jahren einige Zoos die Haltung der Klippspringer beendeten, gibt es keine europäische Zoopopulationen mit Zukunftschancen. Im Moment werden drei Tiere in den zoologischen Gärten in Frankfurt am Main und Valencia gepflegt. Importbemühungen seitens Frankfurt scheiterten vor einigen Jahren. Da es keine Nachzuchten gibt, ist damit zu rechnen, dass Klippspringer in den nächsten Jahren aus der europäischen Zoolandschaft verschwinden werden. Seit 1994 werden in Europa nur noch Äthiopien-Klippspringer gepflegt.
(Wikipedia)
Bush elephant
Afrikanischer Elefant
Kruger National Park is one of the largest game reserves in Africa. It covers an area of 19,485 km2 (7,523 sq mi) in the provinces of Limpopo and Mpumalanga in northeastern South Africa, and extends 360 km (220 mi) from north to south and 65 km (40 mi) from east to west. The administrative headquarters are in Skukuza. Areas of the park were first protected by the government of the South African Republic in 1898, and it became South Africa's first national park in 1926.
To the west and south of the Kruger National Park are the two South African provinces of Limpopo and Mpumalanga. In the north is Zimbabwe, and to the east is Mozambique. It is now part of the Great Limpopo Transfrontier Park, a peace park that links Kruger National Park with the Gonarezhou National Park in Zimbabwe, and with the Limpopo National Park in Mozambique.
The park is part of the Kruger to Canyons Biosphere an area designated by the United Nations Educational, Scientific and Cultural Organization (UNESCO) as an International Man and Biosphere Reserve (the "Biosphere").
The park has nine main gates allowing entrance to the different camps.
(Wikipedia)
Name
African Elephant or African Bush Elephant [Loxodonta africana]
Introduction
The Elephant is the world's largest land mammal, and weighs up to 7 tonnes and reaches heights of 3.3 m at the shoulder. Elephants can live to a potential age of 70 years. The massive tusks of older bulls can weigh up to 50 or 60 kilograms, but tusks weighing up to 90 kilograms have been recorded.
Appearance
What is the trunk and what is it used for?
The Elephant's trunk is a modified nose which is very sensitive and can even detect water under ground. There are as many as 50 000 muscles in an Elephant trunk. The sensitive finger-like appendages at the tip of the trunk enables them to pick the smallest twig or flower, pull the toughest reed of grass or even pick out a thorn from their feet.
Do elephants have knees or elbows?
The joints that are perceived as 'knees', are in fact wrists. This is a common misunderstanding due to the belief that a leg joint that bends between the foot and the body must be a knee. The main difference between us and the elephants is that our foot bones and hand bones are separate, whereas those of the elephant are one in the same, and have evolved to suit this four-legged mammal.
Why do elephants have tusks?
The tusks are used for obtaining food, fighting (amongst males) and for self defence. They are actually their upper incisors, and grow continuously until they die at around 60 years old. Although their skin is up to 3cm (1 inch) thick, it is quite sensitive.
Diet
Elephants are voracious feeders which in a day consume up to 272 kg (600 pounds) of grass, tender shoots and bark from trees. An adult Elephant can drink up to 200 litres of water in a single session. A single Elephant deposits up to 150kg (330 pounds) of dung every day - about one dollop every 15 minutes!
Breeding
African Elephant are not seasonal breeders. Generally they produce one calf every 3 to 4 years. The gestation period is about 22 months. At birth calves weigh about 100 kg (220 pounds) and are fully weaned between 18-24 months. An orphaned calf will usually be adopted by one of the family's lactating females or suckled by various females. Elephants are very attentive mothers, and because most Elephant behavior has to be learned, they keep their offspring with them for many years. Tusks erupt at 16 months but do not show externally until 30 months. Once weaned, usually at age 4 or 5, the calf still remains in the maternal group. Females mature at about 11 years and stay in the group, while the males, which mature between 12 and 15, are usually expelled from the maternal herd. Even though these young males are sexually mature, they do not breed until they are in their mid, or late 20s or even older and have moved up in the social hierarchy.
Behaviour
Mature males form bachelor groups and become solitary bulls. Elephant form strong family units of cows, calves and young offspring. Such herds are always led by an old female. Apart from drinking large quantities of water they also love wading or swimming in it. Elephants clearly relish mud baths.
It was once thought that family groups were led by old bull elephants, but these males are most often solitary. The female family groups are often visited by mature males checking for females in oestrus. Several interrelated family groups may inhabit an area and know each other well.
How do you tell an elephant's mock charge from a serious one?
It is imperative to keep in mind that Elephant are extremely intelligent, and each individual has a distinct character. Although there will be exceptions to the rules, the common signs of a mock charge are bush-bashing, dust-throwing, trumpeting and other vocalizations, open ears and an intimidating presence, can be considered a mock-display. Aggressive or startled elephants usually make sudden headshakes and flap their large ears against their head. Serious charges usually occur after all attempts to intimidate have failed, and the Elephant feels threatened. The ears are pinned back and head and trunk are lowered. Ultimately, the key lies in the intelligence of the animal and how they will react to the 'target' and unfamiliar actions, and a conscious decision is made.
Why do elephants rhythmically flap their ears?
Contrary to common belief, it is not an expression of anger. Being an animal of such a large size, with no sweat glands and a dark body colour, elephants flap their ears to cool the body and rid themselves of irritating insects.
Where are they found?
Once ranging across most of Africa the Elephant population has declined dramatically across the continent. In South Africa the Addo Elephant and Kruger National Park protect large herds. Due to rigorous conservation measures the Elephant population in South Africa has grown from a estimated 120 in 1920 in 4 locations, to 10 000 at 40 locations to date.
Notes
The African Elephant has recently been classified into two separate species, the more common African Bush Elephant [Loxodonta Africana] and the smaller African Forest Elephant [Loxodonta cyclotis] of the rainforest of Central Africa.
(krugerpark.co.za)
Der Kruger-Nationalpark (deutsch häufig falsch Krüger-Nationalpark) ist das größte Wildschutzgebiet Südafrikas. Er liegt im Nordosten des Landes in der Landschaft des Lowveld auf dem Gebiet der Provinz Limpopo sowie des östlichen Abschnitts von Mpumalanga. Seine Fläche erstreckt sich vom Crocodile-River im Süden bis zum Limpopo, dem Grenzfluss zu Simbabwe, im Norden. Die Nord-Süd-Ausdehnung beträgt etwa 350 km, in Ost-West-Richtung ist der Park durchschnittlich 54 km breit und umfasst eine Fläche von rund 20.000 Quadratkilometern. Damit gehört er zu den größten Nationalparks in Afrika.
Das Schutzgebiet wurde am 26. März 1898 unter dem Präsidenten Paul Kruger als Sabie Game Reserve zum Schutz der Wildnis gegründet. 1926 erhielt das Gebiet den Status Nationalpark und wurde in seinen heutigen Namen umbenannt. Im Park leben 147 Säugetierarten inklusive der „Big Five“, außerdem etwa 507 Vogelarten und 114 Reptilienarten, 49 Fischarten und 34 Amphibienarten.
(Wikipedia)
Der Afrikanische Elefant (Loxodonta africana), auch Afrikanischer Steppenelefant oder Afrikanischer Buschelefant, ist eine Art aus der Familie der Elefanten. Er ist das größte gegenwärtig lebende Landsäugetier und gleichzeitig das größte rezente landbewohnende Tier der Erde. Herausragende Kennzeichen sind neben den Stoßzähnen und dem markanten Rüssel die großen Ohren und die säulenförmigen Beine. In zahlreichen morphologischen und anatomischen Merkmalen unterscheidet sich der Afrikanische Elefant von seinen etwas kleineren Verwandten, dem Waldelefanten und dem Asiatischen Elefanten. Das Verbreitungsgebiet umfasst heute große Teile von Afrika südlich der Sahara. Die Tiere haben sich dort an zahlreiche unterschiedliche Lebensräume angepasst, die von geschlossenen Wäldern über offene Savannenlandschaften bis hin zu Sumpfgebieten und wüstenartigen Regionen reichen. Insgesamt ist das Vorkommen aber stark fragmentiert.
Die Lebensweise des Afrikanischen Elefanten ist durch intensive Studien gut erforscht. Sie wird durch einen stark sozialen Charakter geprägt. Weibliche Tiere und ihr Nachwuchs leben in Familienverbänden (Herden). Diese formieren sich wiederum zu einem enger verwandten Clan. Die einzelnen Herden treffen sich zu bestimmten Gelegenheiten und trennen sich danach wieder. Die männlichen Tiere bilden Junggesellengruppen. Die verschiedenen Verbände nutzen Aktionsräume, in denen sie teils im Jahreszyklus herumwandern. Für die Kommunikation untereinander nutzen die Tiere verschiedene Töne im niedrigen Frequenzbereich. Anhand der Lautgebung, aber auch durch bestimmte chemische Signale können sich die einzelnen Individuen untereinander erkennen. Darüber hinaus besteht ein umfangreiches Repertoire an Gesten. Hervorzuheben sind auch die kognitiven Fähigkeiten des Afrikanischen Elefanten.
Die Nahrung besteht sowohl aus weicher wie auch harter Pflanzenkost. Die genaue Zusammensetzung variiert dabei regional und jahreszeitlich. Generell verbringt der Afrikanische Elefant einen großen Teil seiner Tagesaktivitäten mit der Nahrungsaufnahme. Die Fortpflanzung erfolgt ganzjährig, regional gibt es Tendenzen zu einer stärkeren Saisonalisierung. Bullen kommen einmal jährlich in die Musth, während deren sie auf Wanderung zur Suche nach fortpflanzungswilligen Kühen gehen. Während der Musth ist die Aggressivität gesteigert, es finden dann auch Rivalenkämpfe statt. Der Sexualzyklus der Kühe dauert vergleichsweise lange und weist einen für Säugetiere untypischen Verlauf auf. Nach erfolgter Geburt setzt er in der Regel mehrere Jahre aus. Zumeist wird nach fast zweijähriger Tragzeit ein Jungtier geboren, das in der mütterlichen Herde aufwächst. Junge weibliche Tiere verbleiben später in der Herde, die jungen männlichen verlassen diese.
Die wissenschaftliche Erstbeschreibung des Afrikanischen Elefanten erfolgte im Jahr 1797 mit einer formalen artlichen Trennung des Afrikanischen vom Asiatischen Elefanten. Der heute gebräuchliche Gattungsname Loxodonta wurde offiziell erst dreißig Jahre später eingeführt. Die Bezeichnung bezieht sich auf markante Zahnunterschiede zwischen den asiatischen und den afrikanischen Elefanten. Im Verlauf des 20. Jahrhunderts wurden mehrere Unterarten unterschieden, darunter auch der Waldelefant des zentralen Afrikas. Letzterer gilt heute genetischen Untersuchungen zufolge als eigenständige Art, die weiteren Unterarten sind nicht anerkannt. Stammesgeschichtlich lässt sich der Afrikanische Elefant erstmals im beginnenden Mittleren Pleistozän belegen. Der Gesamtbestand gilt als gefährdet. Ursachen hierfür sind hauptsächlich die Jagd nach Elfenbein und Lebensraumverlust durch die zunehmend wachsende menschliche Bevölkerung. Der Afrikanische Elefant zählt zu den sogenannten „Big Five“ von Großwildjagd und Safari.
(Wikipedia)
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Ickworth Park. With over 1,800 acres of parkland designed by Capability Brown, the house and its grounds were created as an homage to Italy, the country so beloved by Frederick Augustus Hervey, the 4th Earl of Bristol. The Earl-Bishop spent his life travelling the continent, gathering together a vast collection of paintings, sculpture and artefacts. Already possessed of several houses, he conceived Ickworth primarily as a museum for his treasures. At his death only the Rotunda - the giant circular structure at the centre of the two wings, described by Hervey's wife as 'a stupendous moment of Folly' - was nearing completion. The house was eventually finished by his son. Although Hervey's treasures, confiscated during the French invasion of Italy, were destined never to occupy Ickworth, his descendants made it their life's work to rebuild what has become an exceptional collection of art and silver. Paintings housed in the galleries include works by Velázquez, Titian and Poussin, while the collection of 18th-century portraits of the family is exceptionally fine, featuring canvases by Gainsborough, Reynolds, Vigée-Lebrun and Hogarth. In addition to one of the very best British collections of Georgian Huguenot silver, Ickworth is also home to an impressive array of Regency furniture, porcelain, and domestic objects. More made a career of producing idealised Italian landscapes. His Landscape with Classical Figures, Cicero at his Villa, painted in 1780 and funded in 1993, is a typical work, the misty soft-focus and pastel light adding to its appeal. Hugh Douglas Hamilton's The Earl Bishop of Bristol and Derry Seated before the Prospect of Rome shows Hervey seated at what is thought to be the southern tip of the Borghese Gardens.
Ickworth's parklands and gardens can provide a day's activity in their own right. The south gardens are modelled on the formal Italian style, while the gardens to the west of the house are more informal. Visitors can walk or cycle out into the park itself and up to the Fairy Lake. Bright and modern, The West Wing Restaurant overlooks the gardens and can be guaranteed to catch any sunlight on offer. It serves everything from hot meals to snacks, and at weekends the restaurant is open for breakfast. If you're after something rather more formal, try Frederick's restaurant at Ickworth Hotel in the grounds.
The sheep is a quadrupedal, ruminant mammal typically kept as livestock. Like all ruminants, sheep are members of the order Artiodactyla, the even-toed ungulates. Although the name "sheep" applies to many species in the genus Ovis, in everyday usage it almost always refers to Ovis aries. Numbering a little over one billion, domestic sheep are also the most numerous species of sheep. An adult female sheep is referred to as a ewe (/juː/), an intact male as a ram or occasionally a tup, a castrated male as a wether, and a younger sheep as a lamb. Sheep are most likely descended from the wild mouflon of Europe and Asia. One of the earliest animals to be domesticated for agricultural purposes, sheep are raised for fleece, meat (lamb, hogget or mutton) and milk. A sheep's wool is the most widely used animal fiber, and is usually harvested by shearing. Ovine meat is called lamb when from younger animals and mutton when from older ones. Sheep continue to be important for wool and meat today, and are also occasionally raised for pelts, as dairy animals, or as model organisms for science. Sheep husbandry is practised throughout the majority of the inhabited world, and has been fundamental to many civilizations. In the modern era, Australia, New Zealand, the southern and central South American nations, and the British Isles are most closely associated with sheep production. Sheepraising has a large lexicon of unique terms which vary considerably by region and dialect. Use of the word sheep began in Middle English as a derivation of the Old English word scēap; it is both the singular and plural name for the animal. A group of sheep is called a flock, herd or mob. Many other specific terms for the various life stages of sheep exist, generally related to lambing, shearing, and age. Being a key animal in the history of farming, sheep have a deeply entrenched place in human culture, and find representation in much modern language and symbology. As livestock, sheep are most often associated with pastoral, Arcadian imagery. Sheep figure in many mythologies—such as the Golden Fleece—and major religions, especially the Abrahamic traditions. In both ancient and modern religious ritual, sheep are used as sacrificial animals. Domestic sheep are relatively small ruminants, usually with a crimped hair called wool and often with horns forming a lateral spiral. Domestic sheep differ from their wild relatives and ancestors in several respects, having become uniquely neotenic as a result of selective breeding by humans. A few primitive breeds of sheep retain some of the characteristics of their wild cousins, such as short tails. Depending on breed, domestic sheep may have no horns at all, or horns in both sexes, or in males only. Most horned breeds have a single pair, but a few breeds may have several. Another trait unique to domestic sheep as compared to wild ovines is their wide variation in color. Wild sheep are largely variations of brown hues, and variation within species is extremely limited. Colors of domestic sheep range from pure white to dark chocolate brown and even spotted or piebald. Selection for easily dyeable white fleeces began early in sheep domestication, and as white wool is a dominant trait it spread quickly. However, colored sheep do appear in many modern breeds, and may even appear as a recessive trait in white flocks. While white wool is desirable for large commercial markets, there is a niche market for colored fleeces, mostly for handspinning. The nature of the fleece varies widely among the breeds, from dense and highly crimped, to long and hairlike. There is variation of wool type and quality even among members of the same flock, so wool classing is a step in the commercial processing of the fibre. Depending on breed, sheep show a range of heights and weights. Their rate of growth and mature weight is a heritable trait that is often selected for in breeding. Ewes typically weigh between 45 and 100 kilograms (99 and 220 lb), and rams between 45 and 160 kilograms (99 and 353 lb). When all deciduous teeth have erupted, the sheep has 20 teeth. Mature sheep have 32 teeth. As with other ruminants, the front teeth in the lower jaw bite against a hard, toothless pad in the upper jaw. These are used to pick off vegetation, then the rear teeth grind it before it is swallowed. There are eight lower front teeth in ruminants, but there is some disagreement as to whether these are eight incisors, or six incisors and two incisor-shaped canines. There is a large diastema between the incisors and the molars. For the first few years of life it is possible to calculate the age of sheep from their front teeth, as a pair of milk teeth is replaced by larger adult teeth each year, the full set of eight adult front teeth being complete at about four years of age. The front teeth are then gradually lost as sheep age, making it harder for them to feed and hindering the health and productivity of the animal. For this reason, domestic sheep on normal pasture begin to slowly decline from four years on, and the average life expectancy of a sheep is 10 to 12 years, though some sheep may live as long as 20 years. Sheep have good hearing, and are sensitive to noise when being handled. Sheep have horizontal slit-shaped pupils, possessing excellent peripheral vision; with visual fields of approximately 270° to 320°, sheep can see behind themselves without turning their heads. Many breeds have only short hair on the face, and some have facial wool (if any) confined to the poll and or the area of the mandibular angle; the wide angles of peripheral vision apply to these breeds. A few breeds tend to have considerable wool on the face; for some individuals of these breeds, peripheral vision may be greatly reduced by "wool blindness", unless recently shorn about the face. Sheep have poor depth perception; shadows and dips in the ground may cause sheep to baulk. In general, sheep have a tendency to move out of the dark and into well lit areas, and prefer to move uphill when disturbed. Sheep also have an excellent sense of smell, and, like all species of their genus, have scent glands just in front of the eyes, and interdigitally on the feet. The purpose of these glands is uncertain, but those on the face may be used in breeding behaviors. The foot glands might also be related to reproduction, but alternative reasons, such as secretion of a waste product or a scent marker to help lost sheep find their flock, have also been proposed.
From Wikipedia, the free encyclopedia
The Pokot live in the Baringo district and in the Western Pokot district in Kenya. There are two main sub-groups depending of their location and way of life : the Hill Pokot and the Plains Pokot. A homestead is composed of one or more buildings (if there are several wifes). Most of the Pokot are nomadic and thus have interacted with different peoples, incorporating their customs. The songs and decorative arts, in particular body decoration, are very appreciated among the Pokot. They adorn the body with beads and hairstyling, and proceed to scarifications and the removal of the lower central incisors. Pokot girls wear a beaded necklace made of the stems of an asparagus tree.For the Pokot, the universe has two realms: the above is the realm of the most powerful deities, and the below is the one where live humans, animals, and plants. Humans are responsible for the realm that they inhabit, but they rely upon divinities to achieve and maintain peace and prosperity. The Pokot communicate with their deities through prayer and sacrifice. They are superstitious and believe in sorcery. They have oracles and prophets. Ceremonies mark the transitions in the people's social lives. The most important rite of passage is circumcision for boys and clitoridectomy for girls. The Pokot keep the affiliation to their clan throughout their lives, there is no disruption with marriage. The bond between a husband and wife lasts for 3 generations. Polygamy exists but is not prevalent among men before 40.
Les Pokot vivent dans le district de Baringo et à l’ouest du district de Pokot au Kenya.Il existe deux principaux sous-groupes selon leur localisation et mode de vie : les Pokot des collines et les Pokot des plaines. Chaque propriété familiale est composée d’un ou plusieurs bâtiments (s’il y a plusieurs femmes). La plupart des Pokot sont nomades et ont donc interagi avec différents peuples, incorporant leurs coutumes. Les chants et arts décoratifs, en particulier la décoration du corps, sont très appréciés chez les Pokot. Ils parent leur corps de perles et coiffures originales, et procèdent à des scarifications et au retrait des incisives centrales inférieures. Les filles Pokot portent un collier de perles fait de tiges d’asparagus.Pour les Pokot, l’univers a deux royaumes : celui d’en haut est le royaume des déités les plus puissantes, celui d’en bas est celui où vivent les humains, animaux et plantes. Les humains sont responsables du royaume qu’ils habitent, mais ils reposent sur les divinités pour atteindre et maintenir la paix et la prospérité. Les Pokot communiquent avec leurs déités par la prière et le sacrifice. Ils sont superstitieux et croient aux sortilèges. Ils ont des oracles et prophètes. Les cérémonies marquent les transitions dans la vie sociale des individus. Le rite de passage le plus important est la circoncision pour les garçons et la clitorectomie pour les filles. Les Pokot gardent l’affiliation à leur clan toute leur vie, il n’y a aucune rupture lors du mariage. Le lien entre le mari et la femme dure pendant 3 générations. La polygamie existe mais ne prévaut pas chez les hommes de moins de 40 ans.
© Eric Lafforgue