Details best viewed in Original Size.

 

I photographed this African Elephant at the Little Vumbura Camp, a concession run by Wilderness Safaris, in the Okavango Delta of Botswana. I shot about a dozen frames and this was one of three keepers.

According to Wikipedia, both African Savanna Elephant (a.k.a. African Bush Elephant) and the African Forest Elephant have in the past been classified as a single species, known simply as the African elephant, but recent preliminary evidence has seen the forest elephant classified as a distinct species (although this status is not conclusively accepted due to concerns over conservation strategies until the reclassification is formalized). The African bush elephant is the largest and heaviest land animal on earth, being up to 13 feet (3.96m) tall at the shoulder and 23,000 lbs. (10.4 tonnes) in weight (a male shot in 1974, near Mucusso, southern Angola). On average, males are 10 feet (3.2m) tall at the shoulder and 13,200 lbs. (6 tonnes) in weight, while females are much smaller at 8.5 feet (2.6 m) tall at the shoulder and 6,600 lbs. (3 tonnes) in weight. The most characteristic features of African elephants are their very large ears, which they use to radiate excess heat and intimidate, and their trunk, a nose and an extension of the upper lip with two opposing extensions, or "fingers" at the end of it (in contrast to the Asian elephant, which only has one). The trunk is used for communication and handling objects and food. African elephants also have bigger tusks, large modified incisors that grow throughout an elephant's life. They occur in both males and females and are used in fights and for marking, feeding, and digging.

www.saranvaid.com

 

The Sloth Bear (Melursus ursinus), also known as the Lip Bear, is a mammal of the family which is native to the lowland forests of India, Nepal,Pakistan, Bhutan, Bangladesh, and Sri Lanka. The Sloth Bear is the only bear species classified in genus Melursus. Though originally classed as a "bear sloth" due to the shape of its claws and its arboreal habits, it was appropriately reclassified as a bear in the 1800s.

 

The body is 150–190 cm long, covered in long, shaggy fur, ranging from auburn to black, with a distinctive;V-shaped white mark on the chest, a whitish snout and black nose. The snout is long with bare lips and a lack of upper incisors, adaptations for its insect-based diet. The front feet are turned inwards and have 4 inch long, non-retractable, curved claws that are adapted for digging and climbing. The males, weighing 80–140 kg, are larger than the females, which weigh only 55–95 kg. Its pugmarks are very similar to a human footprint. The tail is 15-18 cm (6-7 inches) long, the longest in the bear family. Their natural lifespan is unknown, however, a captive specimen was recorded to have lived 40 years.

 

The age of sexual maturity in sloth bears is 3 years. Female Sloth Bears typically give birth to two cubs in December-early January after a gestation of 210 days. The cubs are typically born in a cave or under large boulders, where they remain for two to three months, and continue to accompany the mother for at least a further two years.

 

Sloth bears are the most nocturnal of bears, though sows with cubs will often move in daylight. They do not hibernate. Sloth bears are excellent climbers, and will stay in trees to feed and rest, though not to escape danger. They are capable of jumping from distances of 10 feet, and can hang upside-down in a sloth-like manner. They mark their territories either by rubbing their flanks against trees, or scraping bark with their claws.

 

The Sloth Bear primarily eats ants and termites, breaking into termite mounds with large powerful claws and eating the occupants. It uses its long tubular snout to blow away dirt and suck up the insects, the sounds of which can be heard from 100 meters away. It may also eat honey, eggs, birds, flowers, tubers, fruits, grains and meat. The animal's fondness for honey has caused it to be nicknamed the Honey bear (a nickname also given to the sun bear); it has been known to scale the occasional tree to knock down a bee honeycomb, which it will then enjoy on the ground below.

 

Though sloth bears kill fewer livestock than Asiatic black bears, in some areas of India and Burma, sloth bears are more feared than tigers, due to their unpredictable temperament. In Madhya Pradesh, sloth bear attacks accounted for the deaths of 48 people and the injuring of 686 others between the years 1989 and 1994, probably due in part to the density of population and competition for food sources. One specimen, known as the Sloth bear of Mysore, was singlehandedly responsible for the deaths of 12 people and the mutilation of 2 dozen others before being shot by Kenneth Anderson. Sloth bears defend themselves when surprised, with the majority of confrontations occurring at night. They typically charge on all fours with their head held low, before rearing on their hind legs and striking at their attackers with their claws and teeth.

            

African bush elephant

 

Afrikanischer Elefant

 

Kruger National Park is one of the largest game reserves in Africa. It covers an area of 19,485 km2 (7,523 sq mi) in the provinces of Limpopo and Mpumalanga in northeastern South Africa, and extends 360 km (220 mi) from north to south and 65 km (40 mi) from east to west. The administrative headquarters are in Skukuza. Areas of the park were first protected by the government of the South African Republic in 1898, and it became South Africa's first national park in 1926.

 

To the west and south of the Kruger National Park are the two South African provinces of Limpopo and Mpumalanga. In the north is Zimbabwe, and to the east is Mozambique. It is now part of the Great Limpopo Transfrontier Park, a peace park that links Kruger National Park with the Gonarezhou National Park in Zimbabwe, and with the Limpopo National Park in Mozambique.

 

The park is part of the Kruger to Canyons Biosphere an area designated by the United Nations Educational, Scientific and Cultural Organization (UNESCO) as an International Man and Biosphere Reserve (the "Biosphere").

 

The park has nine main gates allowing entrance to the different camps.

 

(Wikipedia)

 

Name

 

African Elephant or African Bush Elephant [Loxodonta africana]

 

Introduction

 

The Elephant is the world's largest land mammal, and weighs up to 7 tonnes and reaches heights of 3.3 m at the shoulder. Elephants can live to a potential age of 70 years. The massive tusks of older bulls can weigh up to 50 or 60 kilograms, but tusks weighing up to 90 kilograms have been recorded.

 

Appearance

 

What is the trunk and what is it used for?

The Elephant's trunk is a modified nose which is very sensitive and can even detect water under ground. There are as many as 50 000 muscles in an Elephant trunk. The sensitive finger-like appendages at the tip of the trunk enables them to pick the smallest twig or flower, pull the toughest reed of grass or even pick out a thorn from their feet.

 

Do elephants have knees or elbows?

 

The joints that are perceived as 'knees', are in fact wrists. This is a common misunderstanding due to the belief that a leg joint that bends between the foot and the body must be a knee. The main difference between us and the elephants is that our foot bones and hand bones are separate, whereas those of the elephant are one in the same, and have evolved to suit this four-legged mammal.

 

Why do elephants have tusks?

 

The tusks are used for obtaining food, fighting (amongst males) and for self defence. They are actually their upper incisors, and grow continuously until they die at around 60 years old. Although their skin is up to 3cm (1 inch) thick, it is quite sensitive.

 

Diet

 

Elephants are voracious feeders which in a day consume up to 272 kg (600 pounds) of grass, tender shoots and bark from trees. An adult Elephant can drink up to 200 litres of water in a single session. A single Elephant deposits up to 150kg (330 pounds) of dung every day - about one dollop every 15 minutes!

 

Breeding

 

African Elephant are not seasonal breeders. Generally they produce one calf every 3 to 4 years. The gestation period is about 22 months. At birth calves weigh about 100 kg (220 pounds) and are fully weaned between 18-24 months. An orphaned calf will usually be adopted by one of the family's lactating females or suckled by various females. Elephants are very attentive mothers, and because most Elephant behavior has to be learned, they keep their offspring with them for many years. Tusks erupt at 16 months but do not show externally until 30 months. Once weaned, usually at age 4 or 5, the calf still remains in the maternal group. Females mature at about 11 years and stay in the group, while the males, which mature between 12 and 15, are usually expelled from the maternal herd. Even though these young males are sexually mature, they do not breed until they are in their mid, or late 20s or even older and have moved up in the social hierarchy.

 

Behaviour

 

Mature males form bachelor groups and become solitary bulls. Elephant form strong family units of cows, calves and young offspring. Such herds are always led by an old female. Apart from drinking large quantities of water they also love wading or swimming in it. Elephants clearly relish mud baths.

It was once thought that family groups were led by old bull elephants, but these males are most often solitary. The female family groups are often visited by mature males checking for females in oestrus. Several interrelated family groups may inhabit an area and know each other well.

 

How do you tell an elephant's mock charge from a serious one?

 

It is imperative to keep in mind that Elephant are extremely intelligent, and each individual has a distinct character. Although there will be exceptions to the rules, the common signs of a mock charge are bush-bashing, dust-throwing, trumpeting and other vocalizations, open ears and an intimidating presence, can be considered a mock-display. Aggressive or startled elephants usually make sudden headshakes and flap their large ears against their head. Serious charges usually occur after all attempts to intimidate have failed, and the Elephant feels threatened. The ears are pinned back and head and trunk are lowered. Ultimately, the key lies in the intelligence of the animal and how they will react to the 'target' and unfamiliar actions, and a conscious decision is made.

 

Why do elephants rhythmically flap their ears?

 

Contrary to common belief, it is not an expression of anger. Being an animal of such a large size, with no sweat glands and a dark body colour, elephants flap their ears to cool the body and rid themselves of irritating insects.

 

Where are they found?

 

Once ranging across most of Africa the Elephant population has declined dramatically across the continent. In South Africa the Addo Elephant and Kruger National Park protect large herds. Due to rigorous conservation measures the Elephant population in South Africa has grown from a estimated 120 in 1920 in 4 locations, to 10 000 at 40 locations to date.

 

Notes

 

The African Elephant has recently been classified into two separate species, the more common African Bush Elephant [Loxodonta Africana] and the smaller African Forest Elephant [Loxodonta cyclotis] of the rainforest of Central Africa.

 

(krugerpark.co.za)

 

Der Kruger-Nationalpark (deutsch häufig falsch Krüger-Nationalpark) ist das größte Wildschutzgebiet Südafrikas. Er liegt im Nordosten des Landes in der Landschaft des Lowveld auf dem Gebiet der Provinz Limpopo sowie des östlichen Abschnitts von Mpumalanga. Seine Fläche erstreckt sich vom Crocodile-River im Süden bis zum Limpopo, dem Grenzfluss zu Simbabwe, im Norden. Die Nord-Süd-Ausdehnung beträgt etwa 350 km, in Ost-West-Richtung ist der Park durchschnittlich 54 km breit und umfasst eine Fläche von rund 20.000 Quadratkilometern. Damit gehört er zu den größten Nationalparks in Afrika.

 

Das Schutzgebiet wurde am 26. März 1898 unter dem Präsidenten Paul Kruger als Sabie Game Reserve zum Schutz der Wildnis gegründet. 1926 erhielt das Gebiet den Status Nationalpark und wurde in seinen heutigen Namen umbenannt. Im Park leben 147 Säugetierarten inklusive der „Big Five“, außerdem etwa 507 Vogelarten und 114 Reptilienarten, 49 Fischarten und 34 Amphibienarten.

 

(Wikipedia)

 

Der Afrikanische Elefant (Loxodonta africana), auch Afrikanischer Steppenelefant oder Afrikanischer Buschelefant, ist eine Art aus der Familie der Elefanten. Er ist das größte gegenwärtig lebende Landsäugetier und gleichzeitig das größte rezente landbewohnende Tier der Erde. Herausragende Kennzeichen sind neben den Stoßzähnen und dem markanten Rüssel die großen Ohren und die säulenförmigen Beine. In zahlreichen morphologischen und anatomischen Merkmalen unterscheidet sich der Afrikanische Elefant von seinen etwas kleineren Verwandten, dem Waldelefanten und dem Asiatischen Elefanten. Das Verbreitungsgebiet umfasst heute große Teile von Afrika südlich der Sahara. Die Tiere haben sich dort an zahlreiche unterschiedliche Lebensräume angepasst, die von geschlossenen Wäldern über offene Savannenlandschaften bis hin zu Sumpfgebieten und wüstenartigen Regionen reichen. Insgesamt ist das Vorkommen aber stark fragmentiert.

 

Die Lebensweise des Afrikanischen Elefanten ist durch intensive Studien gut erforscht. Sie wird durch einen stark sozialen Charakter geprägt. Weibliche Tiere und ihr Nachwuchs leben in Familienverbänden (Herden). Diese formieren sich wiederum zu einem enger verwandten Clan. Die einzelnen Herden treffen sich zu bestimmten Gelegenheiten und trennen sich danach wieder. Die männlichen Tiere bilden Junggesellengruppen. Die verschiedenen Verbände nutzen Aktionsräume, in denen sie teils im Jahreszyklus herumwandern. Für die Kommunikation untereinander nutzen die Tiere verschiedene Töne im niedrigen Frequenzbereich. Anhand der Lautgebung, aber auch durch bestimmte chemische Signale können sich die einzelnen Individuen untereinander erkennen. Darüber hinaus besteht ein umfangreiches Repertoire an Gesten. Hervorzuheben sind auch die kognitiven Fähigkeiten des Afrikanischen Elefanten.

 

Die Nahrung besteht sowohl aus weicher wie auch harter Pflanzenkost. Die genaue Zusammensetzung variiert dabei regional und jahreszeitlich. Generell verbringt der Afrikanische Elefant einen großen Teil seiner Tagesaktivitäten mit der Nahrungsaufnahme. Die Fortpflanzung erfolgt ganzjährig, regional gibt es Tendenzen zu einer stärkeren Saisonalisierung. Bullen kommen einmal jährlich in die Musth, während deren sie auf Wanderung zur Suche nach fortpflanzungswilligen Kühen gehen. Während der Musth ist die Aggressivität gesteigert, es finden dann auch Rivalenkämpfe statt. Der Sexualzyklus der Kühe dauert vergleichsweise lange und weist einen für Säugetiere untypischen Verlauf auf. Nach erfolgter Geburt setzt er in der Regel mehrere Jahre aus. Zumeist wird nach fast zweijähriger Tragzeit ein Jungtier geboren, das in der mütterlichen Herde aufwächst. Junge weibliche Tiere verbleiben später in der Herde, die jungen männlichen verlassen diese.

 

Die wissenschaftliche Erstbeschreibung des Afrikanischen Elefanten erfolgte im Jahr 1797 mit einer formalen artlichen Trennung des Afrikanischen vom Asiatischen Elefanten. Der heute gebräuchliche Gattungsname Loxodonta wurde offiziell erst dreißig Jahre später eingeführt. Die Bezeichnung bezieht sich auf markante Zahnunterschiede zwischen den asiatischen und den afrikanischen Elefanten. Im Verlauf des 20. Jahrhunderts wurden mehrere Unterarten unterschieden, darunter auch der Waldelefant des zentralen Afrikas. Letzterer gilt heute genetischen Untersuchungen zufolge als eigenständige Art, die weiteren Unterarten sind nicht anerkannt. Stammesgeschichtlich lässt sich der Afrikanische Elefant erstmals im beginnenden Mittleren Pleistozän belegen. Der Gesamtbestand gilt als gefährdet. Ursachen hierfür sind hauptsächlich die Jagd nach Elfenbein und Lebensraumverlust durch die zunehmend wachsende menschliche Bevölkerung. Der Afrikanische Elefant zählt zu den sogenannten „Big Five“ von Großwildjagd und Safari.

 

(Wikipedia)

Hippopotamus

 

Flusspferd

 

Kruger National Park is one of the largest game reserves in Africa. It covers an area of 19,485 km2 (7,523 sq mi) in the provinces of Limpopo and Mpumalanga in northeastern South Africa, and extends 360 km (220 mi) from north to south and 65 km (40 mi) from east to west. The administrative headquarters are in Skukuza. Areas of the park were first protected by the government of the South African Republic in 1898, and it became South Africa's first national park in 1926.

 

To the west and south of the Kruger National Park are the two South African provinces of Limpopo and Mpumalanga. In the north is Zimbabwe, and to the east is Mozambique. It is now part of the Great Limpopo Transfrontier Park, a peace park that links Kruger National Park with the Gonarezhou National Park in Zimbabwe, and with the Limpopo National Park in Mozambique.

 

The park is part of the Kruger to Canyons Biosphere an area designated by the United Nations Educational, Scientific and Cultural Organization (UNESCO) as an International Man and Biosphere Reserve (the "Biosphere").

 

The park has nine main gates allowing entrance to the different camps.

 

(Wikipedia)

 

The common hippopotamus (/ˌhɪpəˈpɒtəməs/ HIP-ə-POT-ə-məs; Hippopotamus amphibius), or hippo, is a large, mostly herbivorous, semiaquatic mammal and ungulate native to sub-Saharan Africa. It is one of only two extant species in the family Hippopotamidae, the other being the pygmy hippopotamus (Choeropsis liberiensis or Hexaprotodon liberiensis). The name comes from the ancient Greek for "river horse" (ἱπποπόταμος). After the elephant and rhinoceros, the common hippopotamus is the third-largest type of land mammal and the heaviest extant artiodactyl. Despite their physical resemblance to pigs and other terrestrial even-toed ungulates, the closest living relatives of the Hippopotamidae are cetaceans (whales, dolphins, porpoises, etc.) from which they diverged about 55 million years ago.

 

Common hippos are recognisable by their barrel-shaped torsos, wide-opening mouths revealing large canine tusks, nearly hairless bodies, columnar legs and large size; adults average 1,500 kg (3,310 lb) and 1,300 kg (2,870 lb) for males and females respectively. Despite its stocky shape and short legs, it is capable of running 30 km/h (19 mph) over short distances.

 

The common hippopotamus inhabits rivers, lakes and mangrove swamps, where territorial bulls preside over a stretch of river and groups of five to thirty females and young. During the day, they remain cool by staying in the water or mud; reproduction and childbirth both occur in water. They emerge at dusk to graze on grasses. While hippopotamuses rest near each other in the water, grazing is a solitary activity and hippos are not territorial on land. The hippopotamus is among the most dangerous animals in the world as it is highly aggressive and unpredictable. They are threatened by habitat loss and poaching for their meat and ivory canine teeth.

 

The Latin word "hippopotamus" is derived from the ancient Greek ἱπποπόταμος, hippopotamos, from ἵππος, hippos, "horse", and ποταμός, potamos, "river", meaning "horse of the river". In English, the plural is "hippopotamuses", but "hippopotami" is also used; "hippos" can be used as a short plural. Hippopotamuses are gregarious, living in groups of up to thirty animals. A group is called a pod, herd, dale, or bloat.

 

Hippopotami are among the largest living land mammals, being only smaller than elephants and some rhinoceroses. Amongst the extant African megafauna, behind the two African elephant species, they average smaller than the white rhinoceros but are larger by body mass than the black rhinoceros and the giraffe. Mean adult weight is around 1,500 kg (3,310 lb) and 1,300 kg (2,870 lb) for males and females respectively, very large males can reach 2,000 kg (4,410 lb) and exceptional males weighing 2,660 kg (5,860 lb) and 3,200 kg (7,050 lb) have been reported. Male hippos appear to continue growing throughout their lives while females reach maximum weight at around age 25.

 

Hippopotami have barrel-shaped bodies with short legs and long muzzles. Their skeletal structures are graviportal; adapted to carrying their enormous weight, and their specific gravity allows them to sink and move along the bottom of a river. Hippopotamuses have small legs (relative to other megafauna) because the water in which they live reduces the weight burden. Though they are bulky animals, hippopotamuses can gallop at 30 km/h (19 mph) on land but normally trot. They are incapable of jumping but do climb up steep banks. Despite being semiaquatic and having webbed feet, an adult hippo is not a particularly good swimmer nor can it float. It is rarely found in deep water; when it is, the animal moves by porpoise-like leaps from the bottom. The eyes, ears, and nostrils of hippos are placed high on the roof of their skulls. This allows these organs to remain above the surface while the rest of the body submerges. The testes of the males descend only partially and a scrotum is not present. In addition, the penis retracts into the body when not erect. The genitals of the female are unusual in that the vagina is ridged and two large diverticula protrude from the vulval vestibule. The function of these is unknown.

 

The hippo's jaw is powered by a large masseter and a well-developed digastric; the latter loops up behind the former to the hyoid. The jaw hinge is located far back enough to allow the animal to open its mouth at almost 180°. A moderate folding of the orbicularis oris muscle allows the hippo to achieve such a gape without tearing any tissue. The bite force of an adult female has been measured as 8,100 newtons (1,800 lbf). Hippopotamus teeth sharpen themselves as they grind together. The lower canines and lower incisors are enlarged, especially in males, and grow continuously. The incisors can reach 40 cm (1 ft 4 in), while the canines reach up to 50 cm (1 ft 8 in). The canines and incisors are used for combat and play no role in feeding. Hippos rely on their broad horny lips to grasp and pull grasses which are then ground by the molars. The hippo is considered to be a pseudoruminant; it has a complex three-chambered stomach but does not "chew cud".

 

Unlike most other semiaquatic animals, the hippopotamus has very little hair. The skin is 6 cm (2 in) thick, providing it great protection against conspecifics and predators. By contrast, its subcutaneous fat layer is thin. The animals' upper parts are purplish-grey to blue-black, while the under parts and areas around the eyes and ears can be brownish-pink. Their skin secretes a natural sunscreen substance which is red-coloured. The secretion is sometimes referred to as "blood sweat", but is neither blood nor sweat. This secretion is initially colourless and turns red-orange within minutes, eventually becoming brown. Two distinct pigments have been identified in the secretions, one red (hipposudoric acid) and one orange (norhipposudoric acid). The two pigments are highly acidic compounds. They inhibit the growth of disease-causing bacteria, and their light absorption peaks in the ultraviolet range, creating a sunscreen effect. All hippos, even those with different diets, secrete the pigments, so it does not appear that food is the source of the pigments. Instead, the animals may synthesise the pigments from precursors such as the amino acid tyrosine. Nevertheless, this natural sunscreen cannot prevent the animal's skin from cracking if it stays out of water too long.

 

A hippo's lifespan is typically 40–50 years. Donna the Hippo was the oldest living hippo in captivity. She lived at the Mesker Park Zoo in Evansville, Indiana in the US until her death in 2012 at the age of 61.

 

Different from all other large land mammals, hippos are of semiaquatic habits, spending the day in lakes and rivers. They can be found in both savannah and forest areas. Proper habitat requires enough water to submerge in and grass nearby. Larger densities of the animals inhabit quiet waters with mostly firm, smooth sloping beaches. Males may be found in very small numbers in rapid waters in rocky gorges. Hippo mostly live in freshwater habitats, however populations in West Africa mostly inhabit estuarine waters and may even be found at sea. With the exception of eating, most of hippopotamuses' lives occurs in the water. Hippos leave the water at dusk and travel inland, sometimes up to 10 km (6 mi), to graze on short grasses, their main source of food. They spend four to five hours grazing and can consume 68 kg (150 lb) of grass each night.

 

Like almost any herbivore, they consume other plants if presented with them, but their diet in nature consists almost entirely of grass, with only minimal consumption of aquatic plants. Hippos are born with sterile intestines, and require bacteria obtained from their mothers' feces to digest vegetation. Hippos have (albeit rarely) been filmed eating carrion, usually close to the water. There are other reports of meat-eating, and even cannibalism and predation. The stomach anatomy of a hippo is not suited to carnivory, and meat-eating is likely caused by aberrant behaviour or nutritional stress.

 

Hippo defecation creates allochthonous deposits of organic matter along the river beds. These deposits have an unclear ecological function. A 2015 study concluded that hippo dung provides nutrients from terrestrial material for fish and aquatic invertebrates, while a 2018 study found that their dung can be toxic to aquatic life in large quantities, due to absorption of dissolved oxygen in water bodies. Because of their size and their habit of taking the same paths to feed, hippos can have a significant impact on the land across which they walk, both by keeping the land clear of vegetation and depressing the ground. Over prolonged periods, hippos can divert the paths of swamps and channels.

A hippopotamus walking on the grass land in Serengeti National Park in the morning

 

Adult hippos move at speeds up to 8 km/h (5 mph) in water; typically resurfacing to breathe every three to five minutes. The young have to breathe every two to three minutes. The process of surfacing and breathing is subconscious: a hippo sleeping underwater will rise and breathe without waking up. A hippo closes its nostrils when it submerges into the water. As with fish and turtles on a coral reef, hippos occasionally visit cleaning stations and signal, by opening their mouths wide, their readiness for being cleaned of parasites by certain species of fishes. This is an example of mutualism, in which the hippo benefits from the cleaning while the fish receive food.

 

Hippopotamus coexist with a variety of formidable predators. Nile crocodiles, lions and spotted hyenas are known to prey on young hippos. However, due to their aggression and size, adult hippopotamus are not usually preyed upon by other animals. Cases where large lion prides have successfully preyed on adult hippopotamus have been reported; however, this predation is generally rare. Lions occasionally prey on adults at Gorongosa National Park and calves are taken at Virunga. Crocodiles are frequent targets of hippo aggression, probably because they often inhabit the same riparian habitats; crocodiles may be either aggressively displaced or killed by hippopotamuses. In turn, beyond cases of killing the seldom unguarded hippo calf, very large Nile crocodiles have been verified to occasionally prey on "half-grown" hippopotamuses and anecdotally perhaps adult female hippos. Aggregations of crocodiles have also been seen to dispatch still-living bull hippopotamuses that have been previously injured in mating battles with other bulls.

 

(Wikipedia)

 

Der Kruger-Nationalpark (deutsch häufig falsch Krüger-Nationalpark) ist das größte Wildschutzgebiet Südafrikas. Er liegt im Nordosten des Landes in der Landschaft des Lowveld auf dem Gebiet der Provinz Limpopo sowie des östlichen Abschnitts von Mpumalanga. Seine Fläche erstreckt sich vom Crocodile-River im Süden bis zum Limpopo, dem Grenzfluss zu Simbabwe, im Norden. Die Nord-Süd-Ausdehnung beträgt etwa 350 km, in Ost-West-Richtung ist der Park durchschnittlich 54 km breit und umfasst eine Fläche von rund 20.000 Quadratkilometern. Damit gehört er zu den größten Nationalparks in Afrika.

 

Das Schutzgebiet wurde am 26. März 1898 unter dem Präsidenten Paul Kruger als Sabie Game Reserve zum Schutz der Wildnis gegründet. 1926 erhielt das Gebiet den Status Nationalpark und wurde in seinen heutigen Namen umbenannt. Im Park leben 147 Säugetierarten inklusive der „Big Five“, außerdem etwa 507 Vogelarten und 114 Reptilienarten, 49 Fischarten und 34 Amphibienarten.

 

(Wikipedia)

 

Das Flusspferd (Hippopotamus amphibius), auch Nilpferd, Großflusspferd oder Hippopotamus genannt, ist ein großes, pflanzenfressendes Säugetier. Es lebt in Gewässernähe im mittleren und südlichen Afrika. Zusammen mit dem Breitmaulnashorn zählt es zu den schwersten landbewohnenden Säugetieren nach den Elefanten. Mit einem Gesamtbestand von rund 125.000 bis 150.000 Tieren und einem erwarteten weiteren Rückgang der Population ist die Art gefährdet.

 

Zusammen mit dem Zwergflusspferd und zahlreichen ausgestorbenen Arten bildet das Flusspferd die Familie der Flusspferde (Hippopotamidae). Obwohl sie „-pferde“ genannt werden, sind Flusspferde mit den Pferden nicht näher verwandt. Traditionell werden sie in die Ordnung der Paarhufer gestellt. Nach derzeitiger Lehrmeinung sind indessen die Wale die nächsten Verwandten der Flusspferde, die zusammen mit den paraphyletischen Paarhufern das Taxon der Cetartiodactyla bilden.

 

Die Bezeichnung Flusspferd ist eine Lehnübersetzung des griechischen Wortes ἱπποπόταμος hippopótamos (gebildet aus hippos „Pferd“ und potamos „Fluss“). Hippopotamus, der wissenschaftliche Name der Gattung, ist die latinisierte Form des griechischen Wortes. Der Namensbestandteil amphibius bezieht sich auf die amphibische Lebensweise im Wasser und an Land. Hiob Ludolf erklärte den griechischen Namen des Tieres mit der Ähnlichkeit des aus dem Wasser ragenden Kopfes mit einem Pferdekopf.

 

Der Name Nilpferd rührt daher, dass in der Literatur zunächst Flusspferde am Nil beschrieben wurden.

 

Flusspferde sind schwere Tiere mit einem fassförmigen Körper, einem wuchtigen Kopf und kurzen Gliedmaßen. Die kräftigen Beine enden in jeweils vier nach vorne ragenden Zehen, die mit Schwimmhäuten verbunden sind.

 

Flusspferde erreichen eine Kopf-Rumpf-Länge von 2,9 bis 5,1 Metern, wozu noch ein 40 bis 56 Zentimeter langer Schwanz kommt. Die Schulterhöhe beträgt 150 bis 165 Zentimeter. Das Gewicht schwankt zwischen 1000 und 4500 Kilogramm. Männchen sind generell größer und schwerer als Weibchen.

 

Die Haut der Tiere ist so spärlich mit kurzen, feinen Haaren bedeckt, dass sie nackt erscheint. Die Haut ist bräunlich, annähernd kupferfarben gefärbt, an der Oberseite ist sie dunkler und am Bauch purpurn. Auch im Gesicht, speziell um die Augen, Ohren und an den Wangen, können purpurne oder rosafarbene Flecken vorhanden sein.

 

Der Kopf des Flusspferds ist groß und wuchtig. Die Schnauze ist vorne sehr breit, was durch die Eckzahnfächer bedingt ist. Auf Höhe der Prämolaren ist sie stark eingeschnürt. Die Nasenöffnungen liegen erhöht und sind verschließbar. Auch die leicht hervorstehenden Augen und die bis zu 10 Zentimeter langen Ohren sitzen hoch am Kopf, so dass nur sie aus dem Wasser herausragen, wenn das Tier unter der Wasseroberfläche schwimmt.

 

Pro Kieferhälfte haben die Flusspferde zwei oder drei Schneidezähne, einen Eckzahn, vier Prämolaren und drei Molaren. Die Schneide- und Eckzähne wachsen das ganze Leben lang. Die Schneidezähne sind rundlich, glatt und weit voneinander entfernt. Die oberen Schneidezähne sind eher klein und nach unten gerichtet, die unteren sind länger (vor allem das innere Paar) und weisen nach vorne. Die Eckzähne sind hauerartig entwickelt, die unteren sind größer und können eine Gesamtlänge von 70 Zentimetern erreichen (von denen 30 Zentimeter aus dem Zahnfleisch ragen). Die Prämolaren haben in der Regel einen Höcker, die Molaren weisen zwei Paar Höcker auf, nur der hinterste drei. Flusspferde können ihre Kiefer bis zu ca. 150° aufklappen.

 

Lebensraum der Flusspferde sind Gebiete mit tieferen Seen und langsam fließenden Flüssen, die idealerweise mit Schilfgürteln umgeben sind. Zum Weiden benötigt es Grasgebiete in der näheren Umgebung der Gewässer. In Regenwäldern kommt es außer bei großen Flüssen in der Regel nicht vor.

 

Flusspferde verbringen praktisch den ganzen Tag schlafend oder ruhend, dazu halten sie sich im Wasser oder in Gewässernähe auf. Dabei tauchen sie oft bis auf die Augen, Ohren und Nasenlöcher unter. Obwohl Flusspferde gut an ein Leben im Wasser angepasst sind, sind sie schlechte Schwimmer. Meistens laufen sie auf dem Grund eines Gewässers entlang oder lassen sich vom Wasser tragen; ihre Fortbewegungsart wird manchmal als „Schwimmlaufen“ umschrieben. Wenn sie untertauchen, können sie ihre schlitzförmigen Nasenlöcher und Ohren verschließen. Tauchgänge sind in der Regel nicht länger als drei bis fünf Minuten, sie können aber länger unter Wasser bleiben, möglicherweise bis zu 30 Minuten. Im Schlaf erfolgt das Auftauchen ebenso automatisch wie das Luftholen.

 

Vorwiegend in der Nacht verlassen sie das schützende Wasser, um sich auf Nahrungssuche zu begeben. Dabei können sie sich mehrere Kilometer vom Wasser entfernen, um Grasflächen zu erreichen. Um zu ihren Weideflächen zu gelangen, bilden die Gruppen regelrechte Trampelpfade („Hippo Trails“). Trotz ihres behäbigen Äußeren können Flusspferde im Bedarfsfall schnell laufen, Schätzungen belaufen sich auf bis zu 50 Kilometer pro Stunde. Diese Geschwindigkeit halten sie aber nur wenige hundert Meter durch.

 

An Land muss ihre Haut feucht bleiben, sie wird rissig, wenn sie zu lange der Luft ausgesetzt ist. Spezielle Hautdrüsen sondern eine Flüssigkeit ab, die die Tiere vor der Austrocknung schützt. Diese zunächst farblose Flüssigkeit verfärbt sich innerhalb von ein paar Minuten rötlich und später bräunlich. Bestandteil dieser Flüssigkeit sind zwei saure Pigmente, hipposudoric acid und norhipposudoric acid genannt. Diese wirken sowohl als Sonnenschutz, indem sie UV-Strahlen absorbieren, als auch antibiotisch gegen verschiedene Krankheitserreger. Das rötliche Schimmern hat früher zu der Vermutung geführt, Flusspferde würden Blut schwitzen.

 

(Wikipedia)

#8 restoration

Sometimes, when the necklaces are big or too complicated to remove, some sleep with it arounf the neck!

The Pokhot live in the Baringo and Western Pokot districts of Kenya and in Uganda.

There are two main sub-groups depending of their location and way of life. The first group consist of the Hill Pokot who live in the rainy highlands in the west and in the central south, and are mainly farmers and pastoralists. The second group is made up of the Plains Pokot who live in dry and infertile plains, with their cattles. A homestead is composed of one or more buildings for a man, his wife and children; eventual co-wives live in separate houses. The role of the community in teaching children ethical rules. Most of the Pokot are nomadic and thus have interacted with different peoples, incorporating their social customs.The Pokot are very proud of their culture. The Songs, storytelling, and decorative arts, especially bodily decoration, are very appreciated. They adorn the body with beads, hairstyling, scarification, and the removal of the lower central incisors. Pokot girls wear a beaded necklace made of the stems of an asparagus tree. Most Pokot have some knowledge of herbal medicine, so they often use these treatments along with those of the hospitals. They belong to the Kenya's Nilotic-speaking peoples. .

For the Pokot, the universe has two realms: the above is the realm of the most powerful deities—Tororot, Asis (sun), and llat (rain); and the below is the one where live humans, animals, and plants. Humans are responsible for the realm that they inhabit, but they rely upon divinities to achieve and maintain peace and prosperity. They worship many deities like the sun, moon and believe in the spirit of death.The Pokot communicate with their deities through prayer and sacrifice. They perform it during ethnic festivals and dances. Oracles are responsible for maintaining the spiritual balance within the community. They are superstitious and believe in sorcery, so sometimes they call on shielding lucky sorcery. They have prophets, either male or female, who foresee advise, usually by the means of animal sacrifices. His or her ability is considered as a divine gift. Clan histories recount the changes of location, through poetry and song, emphasizing the vulnerability of humans and the importance of supernatural powers to help them overcome hunger, thirst, and even death. Ceremonies mark the transitions in the people's social lives. Among these are: the cleansing of a couple expecting their first child; the cleansing of newborn infants and their mothers; the cleansing of twins and other children who are born under unusual circumstances; male and female initiation; marriage; sapana, a coming-of-age ceremony for men; and summer-solstice, harvest, and healing ceremonies. The most important rite of passage for most Pokot is circumcision for boys and clitoridectomy for girls. These rites consist of a series of neighborhood-based ceremonies, emphasizing the importance of having a good behavior. When boys are circumcised, they acquire membership in one of eight age sets. Women do not have age-sets. After excisions, for several months, girls have a white painting on their face and wear a hood made of blackened leather with charcoal and oil. This means they are untouchable until the lepan ceremony, that marks the passage to womanhood. Unlike other tribes, the Pokot keep the affiliation to their clan throughout their lives, there is no disruption with marriage. Surprisingly, the agreement before marriage is made by gift giving, from the groom and his family to the bride and her family, often over a period of years (and not the contrary). It often implies the gift of a combination of livestock, goods, and cash to the bride's family, and the allotment of milk cows and rights to land to the bride. The bond between a husband and wife lasts for 3 generations, after what marriages can take place again between the two groups. Polygamy exists but is not prevalent among men before 40. The spirits of the elder anticipate reincarnation in their living descendants: when a child is said to resemble the elder, the same name is given. Disputes are resolved in neighborhood councils and in government courts. Some of the sanctions include shaming, cursing, and bewitching.

 

© Eric Lafforgue

www.ericlafforgue.com

 

A beaver was busy building a lodge at the edge of Reflections Lake. It looks to be a lot of hard work. The beaver had just woven a cottonwood branch into the pile.

 

He will keep piling up branches and small trees he has felled with his fabulous incisors. Then he will gnaw tunnels into the piles, creating a porch for drying off (accessed from under water, of course) and a sleeping den. In Alaska, the mud that he will line the den with will freeze hard as concrete in winter. A vent is kept open - an occupied beaver lodge in winter has a wispy curl of beaver breath coming from the top of the mound.

 

It is hard to overstate the importance of beavers in a healthy ecosystem. They raise water tables, mitigate effects of drought, purify streams of sediment and even bacteria, provide nesting sites for Trumpeter Swans (old lodges) and keep streams healthy and flowing for trout and salmon. In the 1930's, the CCC put 600 beavers to work repairing riparian environments in Oregon, Washington, Wyoming, and Utah. Each $5 beaver completed $300 worth of work.

 

They were once common from the subarctic in Alaska and Canada, all across North America coast to coast and even pretty far south on some rivers in Mexico. They have since been extirpated by humans from many regions. A lot of work has gone into trying to bring them back in places like the Wood River Refuge in Oregon.

 

And yet, some local oaf walking around the lake stopped to brag about shooting a beaver that was building a dam on 'his' land. Pretty sure the beaver was there first.

Traffic Block - African bush elephant

 

Verkehrshindernis - Afrikanischer Elefant

 

Kruger National Park is one of the largest game reserves in Africa. It covers an area of 19,485 km2 (7,523 sq mi) in the provinces of Limpopo and Mpumalanga in northeastern South Africa, and extends 360 km (220 mi) from north to south and 65 km (40 mi) from east to west. The administrative headquarters are in Skukuza. Areas of the park were first protected by the government of the South African Republic in 1898, and it became South Africa's first national park in 1926.

 

To the west and south of the Kruger National Park are the two South African provinces of Limpopo and Mpumalanga. In the north is Zimbabwe, and to the east is Mozambique. It is now part of the Great Limpopo Transfrontier Park, a peace park that links Kruger National Park with the Gonarezhou National Park in Zimbabwe, and with the Limpopo National Park in Mozambique.

 

The park is part of the Kruger to Canyons Biosphere an area designated by the United Nations Educational, Scientific and Cultural Organization (UNESCO) as an International Man and Biosphere Reserve (the "Biosphere").

 

The park has nine main gates allowing entrance to the different camps.

 

(Wikipedia)

 

Name

 

African Elephant or African Bush Elephant [Loxodonta africana]

 

Introduction

 

The Elephant is the world's largest land mammal, and weighs up to 7 tonnes and reaches heights of 3.3 m at the shoulder. Elephants can live to a potential age of 70 years. The massive tusks of older bulls can weigh up to 50 or 60 kilograms, but tusks weighing up to 90 kilograms have been recorded.

 

Appearance

 

What is the trunk and what is it used for?

The Elephant's trunk is a modified nose which is very sensitive and can even detect water under ground. There are as many as 50 000 muscles in an Elephant trunk. The sensitive finger-like appendages at the tip of the trunk enables them to pick the smallest twig or flower, pull the toughest reed of grass or even pick out a thorn from their feet.

 

Do elephants have knees or elbows?

 

The joints that are perceived as 'knees', are in fact wrists. This is a common misunderstanding due to the belief that a leg joint that bends between the foot and the body must be a knee. The main difference between us and the elephants is that our foot bones and hand bones are separate, whereas those of the elephant are one in the same, and have evolved to suit this four-legged mammal.

 

Why do elephants have tusks?

 

The tusks are used for obtaining food, fighting (amongst males) and for self defence. They are actually their upper incisors, and grow continuously until they die at around 60 years old. Although their skin is up to 3cm (1 inch) thick, it is quite sensitive.

 

Diet

 

Elephants are voracious feeders which in a day consume up to 272 kg (600 pounds) of grass, tender shoots and bark from trees. An adult Elephant can drink up to 200 litres of water in a single session. A single Elephant deposits up to 150kg (330 pounds) of dung every day - about one dollop every 15 minutes!

 

Breeding

 

African Elephant are not seasonal breeders. Generally they produce one calf every 3 to 4 years. The gestation period is about 22 months. At birth calves weigh about 100 kg (220 pounds) and are fully weaned between 18-24 months. An orphaned calf will usually be adopted by one of the family's lactating females or suckled by various females. Elephants are very attentive mothers, and because most Elephant behavior has to be learned, they keep their offspring with them for many years. Tusks erupt at 16 months but do not show externally until 30 months. Once weaned, usually at age 4 or 5, the calf still remains in the maternal group. Females mature at about 11 years and stay in the group, while the males, which mature between 12 and 15, are usually expelled from the maternal herd. Even though these young males are sexually mature, they do not breed until they are in their mid, or late 20s or even older and have moved up in the social hierarchy.

 

Behaviour

 

Mature males form bachelor groups and become solitary bulls. Elephant form strong family units of cows, calves and young offspring. Such herds are always led by an old female. Apart from drinking large quantities of water they also love wading or swimming in it. Elephants clearly relish mud baths.

It was once thought that family groups were led by old bull elephants, but these males are most often solitary. The female family groups are often visited by mature males checking for females in oestrus. Several interrelated family groups may inhabit an area and know each other well.

 

How do you tell an elephant's mock charge from a serious one?

 

It is imperative to keep in mind that Elephant are extremely intelligent, and each individual has a distinct character. Although there will be exceptions to the rules, the common signs of a mock charge are bush-bashing, dust-throwing, trumpeting and other vocalizations, open ears and an intimidating presence, can be considered a mock-display. Aggressive or startled elephants usually make sudden headshakes and flap their large ears against their head. Serious charges usually occur after all attempts to intimidate have failed, and the Elephant feels threatened. The ears are pinned back and head and trunk are lowered. Ultimately, the key lies in the intelligence of the animal and how they will react to the 'target' and unfamiliar actions, and a conscious decision is made.

 

Why do elephants rhythmically flap their ears?

 

Contrary to common belief, it is not an expression of anger. Being an animal of such a large size, with no sweat glands and a dark body colour, elephants flap their ears to cool the body and rid themselves of irritating insects.

 

Where are they found?

 

Once ranging across most of Africa the Elephant population has declined dramatically across the continent. In South Africa the Addo Elephant and Kruger National Park protect large herds. Due to rigorous conservation measures the Elephant population in South Africa has grown from a estimated 120 in 1920 in 4 locations, to 10 000 at 40 locations to date.

 

Notes

 

The African Elephant has recently been classified into two separate species, the more common African Bush Elephant [Loxodonta Africana] and the smaller African Forest Elephant [Loxodonta cyclotis] of the rainforest of Central Africa.

 

(krugerpark.co.za)

 

Der Kruger-Nationalpark (deutsch häufig falsch Krüger-Nationalpark) ist das größte Wildschutzgebiet Südafrikas. Er liegt im Nordosten des Landes in der Landschaft des Lowveld auf dem Gebiet der Provinz Limpopo sowie des östlichen Abschnitts von Mpumalanga. Seine Fläche erstreckt sich vom Crocodile-River im Süden bis zum Limpopo, dem Grenzfluss zu Simbabwe, im Norden. Die Nord-Süd-Ausdehnung beträgt etwa 350 km, in Ost-West-Richtung ist der Park durchschnittlich 54 km breit und umfasst eine Fläche von rund 20.000 Quadratkilometern. Damit gehört er zu den größten Nationalparks in Afrika.

 

Das Schutzgebiet wurde am 26. März 1898 unter dem Präsidenten Paul Kruger als Sabie Game Reserve zum Schutz der Wildnis gegründet. 1926 erhielt das Gebiet den Status Nationalpark und wurde in seinen heutigen Namen umbenannt. Im Park leben 147 Säugetierarten inklusive der „Big Five“, außerdem etwa 507 Vogelarten und 114 Reptilienarten, 49 Fischarten und 34 Amphibienarten.

 

(Wikipedia)

 

Der Afrikanische Elefant (Loxodonta africana), auch Afrikanischer Steppenelefant oder Afrikanischer Buschelefant, ist eine Art aus der Familie der Elefanten. Er ist das größte gegenwärtig lebende Landsäugetier und gleichzeitig das größte rezente landbewohnende Tier der Erde. Herausragende Kennzeichen sind neben den Stoßzähnen und dem markanten Rüssel die großen Ohren und die säulenförmigen Beine. In zahlreichen morphologischen und anatomischen Merkmalen unterscheidet sich der Afrikanische Elefant von seinen etwas kleineren Verwandten, dem Waldelefanten und dem Asiatischen Elefanten. Das Verbreitungsgebiet umfasst heute große Teile von Afrika südlich der Sahara. Die Tiere haben sich dort an zahlreiche unterschiedliche Lebensräume angepasst, die von geschlossenen Wäldern über offene Savannenlandschaften bis hin zu Sumpfgebieten und wüstenartigen Regionen reichen. Insgesamt ist das Vorkommen aber stark fragmentiert.

 

Die Lebensweise des Afrikanischen Elefanten ist durch intensive Studien gut erforscht. Sie wird durch einen stark sozialen Charakter geprägt. Weibliche Tiere und ihr Nachwuchs leben in Familienverbänden (Herden). Diese formieren sich wiederum zu einem enger verwandten Clan. Die einzelnen Herden treffen sich zu bestimmten Gelegenheiten und trennen sich danach wieder. Die männlichen Tiere bilden Junggesellengruppen. Die verschiedenen Verbände nutzen Aktionsräume, in denen sie teils im Jahreszyklus herumwandern. Für die Kommunikation untereinander nutzen die Tiere verschiedene Töne im niedrigen Frequenzbereich. Anhand der Lautgebung, aber auch durch bestimmte chemische Signale können sich die einzelnen Individuen untereinander erkennen. Darüber hinaus besteht ein umfangreiches Repertoire an Gesten. Hervorzuheben sind auch die kognitiven Fähigkeiten des Afrikanischen Elefanten.

 

Die Nahrung besteht sowohl aus weicher wie auch harter Pflanzenkost. Die genaue Zusammensetzung variiert dabei regional und jahreszeitlich. Generell verbringt der Afrikanische Elefant einen großen Teil seiner Tagesaktivitäten mit der Nahrungsaufnahme. Die Fortpflanzung erfolgt ganzjährig, regional gibt es Tendenzen zu einer stärkeren Saisonalisierung. Bullen kommen einmal jährlich in die Musth, während deren sie auf Wanderung zur Suche nach fortpflanzungswilligen Kühen gehen. Während der Musth ist die Aggressivität gesteigert, es finden dann auch Rivalenkämpfe statt. Der Sexualzyklus der Kühe dauert vergleichsweise lange und weist einen für Säugetiere untypischen Verlauf auf. Nach erfolgter Geburt setzt er in der Regel mehrere Jahre aus. Zumeist wird nach fast zweijähriger Tragzeit ein Jungtier geboren, das in der mütterlichen Herde aufwächst. Junge weibliche Tiere verbleiben später in der Herde, die jungen männlichen verlassen diese.

 

Die wissenschaftliche Erstbeschreibung des Afrikanischen Elefanten erfolgte im Jahr 1797 mit einer formalen artlichen Trennung des Afrikanischen vom Asiatischen Elefanten. Der heute gebräuchliche Gattungsname Loxodonta wurde offiziell erst dreißig Jahre später eingeführt. Die Bezeichnung bezieht sich auf markante Zahnunterschiede zwischen den asiatischen und den afrikanischen Elefanten. Im Verlauf des 20. Jahrhunderts wurden mehrere Unterarten unterschieden, darunter auch der Waldelefant des zentralen Afrikas. Letzterer gilt heute genetischen Untersuchungen zufolge als eigenständige Art, die weiteren Unterarten sind nicht anerkannt. Stammesgeschichtlich lässt sich der Afrikanische Elefant erstmals im beginnenden Mittleren Pleistozän belegen. Der Gesamtbestand gilt als gefährdet. Ursachen hierfür sind hauptsächlich die Jagd nach Elfenbein und Lebensraumverlust durch die zunehmend wachsende menschliche Bevölkerung. Der Afrikanische Elefant zählt zu den sogenannten „Big Five“ von Großwildjagd und Safari.

 

(Wikipedia)

Portrait of an Angolan Woman with filed teeth, jewllery and an uncomfortable rope sling.

Bonkers opening wide, practicing for the dentist. Or, just yawning. One of his top incisors was removed many years ago due to dental caries - since then, he doesn’t drool so much as chew in an awkward manner.

The incisor's crown has come off several times when the dentist is making an impression.

Mursi woman without her lip plate. In south Ethiopia, only Mursis and Surmas wear the lip plate. It's considered a sign of beauty by thoses tribes. It's made of wood or terracotta, and they have to remove the lower incisors to let some space for the disc. it's amazing to see them speak without any trouble, put it and remove it as a classic jewel. For some years, some Mursi women have removed the labret and have had surgical to close the hole (the lip has to be cut). Nobody knows really who does this, perhaps some christian missionaries, who try to convert the tribes in the area, but one thing is sure: i have seen women with a rebuilt lip. (pictures next)

The women are shaved, like the men, cos they hate hairiness!

 

© Eric Lafforgue

www.ericlafforgue.com

Surma or Suri are sedentary pastoral people living in south west of Ethiopia, on the western bank of the Omo River, in the Kibish and Tulgit area.

 

These breeders tribal groups have a cattle centred culture. They grow cabbage, beans, yams, tobacco and coffee and breed their cattle, mostly cows, on their traditional lands, located in the Omo Valley. Cows are tremendously important for the Suris. They do not see cattle simply as a material asset but as a life sustaining and meaningful companion. Suri even sing songs for them and make fires to warm them. These cows are not bred for their meat and are usually not killed unless they are needed for ceremonial purposes. They use their milk and their blood, which they both drink. Cows also have a social and symbolic meaning in Suri’s society. Suri men are judged on how much cattle they own. In desperate times, Suri men can risk their lives to steal cattle from other tribes. The average male in the Suri tribe owns 40 cows. Every young male is named after their cattle, which they have to look after since the age of 8. Cows are given to the bride’s family after the wedding ceremony. Usually 20 cows and they offer also a Kalashnikov as wedding gift.

 

This central role of the cow in their way of life accounts for the fierce independence they want to preserve and explains their warlike culture. Indeed, it’s quite common to see men and even women carrying Kalashnikovs, which are part of the daily life. Their remote homeland has always been a place of traditional rivalries with the neighbouring tribes such as the Bume (Nyangatom) or the Toposa from Sudan who regularly team up to raid the Suri’s cattle. These fights, and even sometimes battles, have become quite bloody since automatic firearms have become available from the parties in the Sudanese Civil War. This conflict has pushed neighbouring tribes into Suri’s land and is a constant competition to keep and protect their territory and owns.

 

Like their neighbours, the Surma also paint their bodies. They create a variety of designs on their necked bodies using their fingertips, which helps them to expose their dark skins. The painting could have both a beautifying and opponent frightening purpose. As one studies these body paintings whirls, stripes, flower and star designs are noticeable. Surma men who are generally believed to be expert artists also paint the girls.

 

A ritual chief in the villages known as the Komoru, dressed in colourful robes and wearing a crown of baboon fur leads the Suri. Village life is largely communal, sharing the produce of the cattle (milk and blood). The men in an assembly take decisions of the village.

Although their traditional remoteness and autarky is threatened, only few Surma are familiar with Amharic, the official language of Ethiopia, and their literacy level is very low. Lip plate and Donga stick fight are the two typical distinctive features of these people, which they share with the neighbouring Mursi people.

 

Suri women wear giant lip plate, a sign of beauty, like in Mursi tribe, and also a prime attraction for tourists which help to sustain a view of them, in guidebooks and travel articles, as an untouched people, living in one of the last wildernesses of Africa. When they are ready to marry, teenagers start to make a hole in the lower lip with a wood stick. It will be kept for one night, and is removed to put a bigger one. This is very painful at this time. Few months after, the lip plate has its full size, and the men see the girl as beautiful. The lip plate is made of wood or terracotta. They have to remove the lower incisors to let some space for the disc. Sometimes the pressure of the plate breaks the lip. This is a big problem for the girl because men will consider her as ugly, she won't be able to marry anyone in the tribe apart the old men or the sick people.

 

The Sagenai, called also donga ritual is a combat that brings both wounds and honor to both the winner and loser. The men bodies are decorated with ritual drawings and their heads are protected by a sort of helmet. For the boys participating to the donga, this challenge is a true moment of glory. The combat is taking place in a middle of a circle made by the crowd. The rules are simple and can be summed up as follows: the person who manages to stay on his feet is the winner, and one must absolutely not kill his opponent. The winner will be honored by the entire tribe and can choose girls to date.

The lands of the Suri are stolen by the Ethiopian government to be rented to foreign companies. A Malaysian company, Lim Slow Jin, runs the Koka plantation near Kibish on the east side of the Omo. The lands are confiscated and rented out for 1 euro per hectare for a year.

 

© Eric Lafforgue

www.ericlafforgue.com

Marmots are large rodents with characteristically short but robust legs, enlarged claws well adapted to digging, stout bodies and large heads and incisors to quickly process a variety of vegetation. Marmots are the largest members of the squirrel family.

Taken in the Camargue France. Note the huge webbed rear foot.

 

It is not a native species in France and was introduced from South America in the 19th century for its fur, (as were American Mink and Musk rats from North America). Following escapes and deliberate releases in the 1930’s when demand fell in the global depression they have spread to most regions of France.

 

Resembling a large rat or perhaps a Beaver with a rat like tail they grow to 40–60 cm in body length and have a 30–45 cm tail. They have a coarse, darkish brown outer fur with a soft dense grey under-fur, also called the nutria. They have webbed rear feet, large bright orange-yellow incisors and a whitish patch on the muzzle. Adults weigh 5–9 kg

 

Breeding takes place at any time of the year and with a gestation of 130 days it’s normal for a female to produce two or three broods in a year. They can give birth to as many as 12 young with an average of 6 or 7 many of which die in the first year. The young are born fully furred with open eyes and they can eat vegetation with their parents within hours of birth. Tunnels of about 20cm diameter and up to 10 metres long are dug in the bank sides, frequently with the entrance part submerged. The nest, made of dry grass and sedges, is usually subterranean in France and rarely constructed above ground in reed beds. With no natural predators in France the only population restraints are severe prolonged cold spells in winter and destruction by humans.

 

There is no doubt that they cause serious damage to the quality of the environment they inhabit with serious consequences for native species and the wider environment. Water quality can be seriously degraded as the natural filters provided by reeds, sedges and other bank side plants are removed and bank sides undermined. The destruction of these plants will also remove valuable sites for the birds and insects that depend upon them either for nesting, important forage or cover, and of course there is the considerable loss of some plant species. There is also a certain amount of economic damage caused to crops, particularly maize, carrots and other roots.

 

As an invasive introduced species eradication would be the ideal solution but this is clearly not possible and all that can be done is to strictly manage populations especially in habitats that are of great importance for other threatened or fragile native species. Cage trapping followed by humane killing would seem to be the best approach provided the traps are regularly visited as per the regulations. This allows for the release of any non target species that are captured.

Uno dei più importanti artisti del Novecento, ARTURO MARTINI (Treviso 1889- Milano 1947), che fu incisore, pittore, e soprattutto scultore, riteneva che la scultura debba essere non «una vistosa virtù, ma un oscuro grembo». Le intuizioni che egli colse in quel grembo profondo seppe esprimere nella pietra e nel marmo, nel bronzo e nel legno, nella ceramica e nel gesso, ma soprattutto nella terracotta. Nei primi anni ’30 l’Ilva Refrattari di Vado Ligure gli permise di avere un forno tutto suo, così che poté modellare le statue nella fornace stessa, ed evitare il rischio che si rompessero nel trasporto, prima di essere cotte. L’artista poté così cimentarsi con opere di grandi dimensioni, realizzate in esemplari unici.

Memore di voci antiche, classiche e persino etrusche, seppe riviverle con assoluta originalità, superando il suo innegabile virtuosismo (la «vistosa virtù»!) grazie a un lirismo affascinante, in cui trovano voce le moderne inquietudini, le tensioni, le emozioni della sua vita.

 

Sono opere che emanano un intenso fascino, come questa del 1931-32, «Chiaro di luna»: due giovani donne su un balcone, in abiti leggeri su cui gioca la luce, i volti protesi verso il cielo, guardano la luna.

È in terra refrattaria, alta cm.180x133x54.

Oggi si trova al Middelheim Museum di Anversa.

La foto è stata scattata a Bologna, in occasione della mostra

"Creature. Il sogno della terracotta", 2013-2014.

 

Dable Dam

 

African bush elephant

 

Afrikanischer Elefant

 

Kruger National Park is one of the largest game reserves in Africa. It covers an area of 19,485 km2 (7,523 sq mi) in the provinces of Limpopo and Mpumalanga in northeastern South Africa, and extends 360 km (220 mi) from north to south and 65 km (40 mi) from east to west. The administrative headquarters are in Skukuza. Areas of the park were first protected by the government of the South African Republic in 1898, and it became South Africa's first national park in 1926.

 

To the west and south of the Kruger National Park are the two South African provinces of Limpopo and Mpumalanga. In the north is Zimbabwe, and to the east is Mozambique. It is now part of the Great Limpopo Transfrontier Park, a peace park that links Kruger National Park with the Gonarezhou National Park in Zimbabwe, and with the Limpopo National Park in Mozambique.

 

The park is part of the Kruger to Canyons Biosphere an area designated by the United Nations Educational, Scientific and Cultural Organization (UNESCO) as an International Man and Biosphere Reserve (the "Biosphere").

 

The park has nine main gates allowing entrance to the different camps.

 

(Wikipedia)

 

Name

 

African Elephant or African Bush Elephant [Loxodonta africana]

 

Introduction

 

The Elephant is the world's largest land mammal, and weighs up to 7 tonnes and reaches heights of 3.3 m at the shoulder. Elephants can live to a potential age of 70 years. The massive tusks of older bulls can weigh up to 50 or 60 kilograms, but tusks weighing up to 90 kilograms have been recorded.

 

Appearance

 

What is the trunk and what is it used for?

The Elephant's trunk is a modified nose which is very sensitive and can even detect water under ground. There are as many as 50 000 muscles in an Elephant trunk. The sensitive finger-like appendages at the tip of the trunk enables them to pick the smallest twig or flower, pull the toughest reed of grass or even pick out a thorn from their feet.

 

Do elephants have knees or elbows?

 

The joints that are perceived as 'knees', are in fact wrists. This is a common misunderstanding due to the belief that a leg joint that bends between the foot and the body must be a knee. The main difference between us and the elephants is that our foot bones and hand bones are separate, whereas those of the elephant are one in the same, and have evolved to suit this four-legged mammal.

 

Why do elephants have tusks?

 

The tusks are used for obtaining food, fighting (amongst males) and for self defence. They are actually their upper incisors, and grow continuously until they die at around 60 years old. Although their skin is up to 3cm (1 inch) thick, it is quite sensitive.

 

Diet

 

Elephants are voracious feeders which in a day consume up to 272 kg (600 pounds) of grass, tender shoots and bark from trees. An adult Elephant can drink up to 200 litres of water in a single session. A single Elephant deposits up to 150kg (330 pounds) of dung every day - about one dollop every 15 minutes!

 

Breeding

 

African Elephant are not seasonal breeders. Generally they produce one calf every 3 to 4 years. The gestation period is about 22 months. At birth calves weigh about 100 kg (220 pounds) and are fully weaned between 18-24 months. An orphaned calf will usually be adopted by one of the family's lactating females or suckled by various females. Elephants are very attentive mothers, and because most Elephant behavior has to be learned, they keep their offspring with them for many years. Tusks erupt at 16 months but do not show externally until 30 months. Once weaned, usually at age 4 or 5, the calf still remains in the maternal group. Females mature at about 11 years and stay in the group, while the males, which mature between 12 and 15, are usually expelled from the maternal herd. Even though these young males are sexually mature, they do not breed until they are in their mid, or late 20s or even older and have moved up in the social hierarchy.

 

Behaviour

 

Mature males form bachelor groups and become solitary bulls. Elephant form strong family units of cows, calves and young offspring. Such herds are always led by an old female. Apart from drinking large quantities of water they also love wading or swimming in it. Elephants clearly relish mud baths.

It was once thought that family groups were led by old bull elephants, but these males are most often solitary. The female family groups are often visited by mature males checking for females in oestrus. Several interrelated family groups may inhabit an area and know each other well.

 

How do you tell an elephant's mock charge from a serious one?

 

It is imperative to keep in mind that Elephant are extremely intelligent, and each individual has a distinct character. Although there will be exceptions to the rules, the common signs of a mock charge are bush-bashing, dust-throwing, trumpeting and other vocalizations, open ears and an intimidating presence, can be considered a mock-display. Aggressive or startled elephants usually make sudden headshakes and flap their large ears against their head. Serious charges usually occur after all attempts to intimidate have failed, and the Elephant feels threatened. The ears are pinned back and head and trunk are lowered. Ultimately, the key lies in the intelligence of the animal and how they will react to the 'target' and unfamiliar actions, and a conscious decision is made.

 

Why do elephants rhythmically flap their ears?

 

Contrary to common belief, it is not an expression of anger. Being an animal of such a large size, with no sweat glands and a dark body colour, elephants flap their ears to cool the body and rid themselves of irritating insects.

 

Where are they found?

 

Once ranging across most of Africa the Elephant population has declined dramatically across the continent. In South Africa the Addo Elephant and Kruger National Park protect large herds. Due to rigorous conservation measures the Elephant population in South Africa has grown from a estimated 120 in 1920 in 4 locations, to 10 000 at 40 locations to date.

 

Notes

 

The African Elephant has recently been classified into two separate species, the more common African Bush Elephant [Loxodonta Africana] and the smaller African Forest Elephant [Loxodonta cyclotis] of the rainforest of Central Africa.

 

(krugerpark.co.za)

 

(Wikipedia)

 

Der Kruger-Nationalpark (deutsch häufig falsch Krüger-Nationalpark) ist das größte Wildschutzgebiet Südafrikas. Er liegt im Nordosten des Landes in der Landschaft des Lowveld auf dem Gebiet der Provinz Limpopo sowie des östlichen Abschnitts von Mpumalanga. Seine Fläche erstreckt sich vom Crocodile-River im Süden bis zum Limpopo, dem Grenzfluss zu Simbabwe, im Norden. Die Nord-Süd-Ausdehnung beträgt etwa 350 km, in Ost-West-Richtung ist der Park durchschnittlich 54 km breit und umfasst eine Fläche von rund 20.000 Quadratkilometern. Damit gehört er zu den größten Nationalparks in Afrika.

 

Das Schutzgebiet wurde am 26. März 1898 unter dem Präsidenten Paul Kruger als Sabie Game Reserve zum Schutz der Wildnis gegründet. 1926 erhielt das Gebiet den Status Nationalpark und wurde in seinen heutigen Namen umbenannt. Im Park leben 147 Säugetierarten inklusive der „Big Five“, außerdem etwa 507 Vogelarten und 114 Reptilienarten, 49 Fischarten und 34 Amphibienarten.

 

(Wikipedia)

 

Der Afrikanische Elefant (Loxodonta africana), auch Afrikanischer Steppenelefant oder Afrikanischer Buschelefant, ist eine Art aus der Familie der Elefanten. Er ist das größte gegenwärtig lebende Landsäugetier und gleichzeitig das größte rezente landbewohnende Tier der Erde. Herausragende Kennzeichen sind neben den Stoßzähnen und dem markanten Rüssel die großen Ohren und die säulenförmigen Beine. In zahlreichen morphologischen und anatomischen Merkmalen unterscheidet sich der Afrikanische Elefant von seinen etwas kleineren Verwandten, dem Waldelefanten und dem Asiatischen Elefanten. Das Verbreitungsgebiet umfasst heute große Teile von Afrika südlich der Sahara. Die Tiere haben sich dort an zahlreiche unterschiedliche Lebensräume angepasst, die von geschlossenen Wäldern über offene Savannenlandschaften bis hin zu Sumpfgebieten und wüstenartigen Regionen reichen. Insgesamt ist das Vorkommen aber stark fragmentiert.

 

Die Lebensweise des Afrikanischen Elefanten ist durch intensive Studien gut erforscht. Sie wird durch einen stark sozialen Charakter geprägt. Weibliche Tiere und ihr Nachwuchs leben in Familienverbänden (Herden). Diese formieren sich wiederum zu einem enger verwandten Clan. Die einzelnen Herden treffen sich zu bestimmten Gelegenheiten und trennen sich danach wieder. Die männlichen Tiere bilden Junggesellengruppen. Die verschiedenen Verbände nutzen Aktionsräume, in denen sie teils im Jahreszyklus herumwandern. Für die Kommunikation untereinander nutzen die Tiere verschiedene Töne im niedrigen Frequenzbereich. Anhand der Lautgebung, aber auch durch bestimmte chemische Signale können sich die einzelnen Individuen untereinander erkennen. Darüber hinaus besteht ein umfangreiches Repertoire an Gesten. Hervorzuheben sind auch die kognitiven Fähigkeiten des Afrikanischen Elefanten.

 

Die Nahrung besteht sowohl aus weicher wie auch harter Pflanzenkost. Die genaue Zusammensetzung variiert dabei regional und jahreszeitlich. Generell verbringt der Afrikanische Elefant einen großen Teil seiner Tagesaktivitäten mit der Nahrungsaufnahme. Die Fortpflanzung erfolgt ganzjährig, regional gibt es Tendenzen zu einer stärkeren Saisonalisierung. Bullen kommen einmal jährlich in die Musth, während deren sie auf Wanderung zur Suche nach fortpflanzungswilligen Kühen gehen. Während der Musth ist die Aggressivität gesteigert, es finden dann auch Rivalenkämpfe statt. Der Sexualzyklus der Kühe dauert vergleichsweise lange und weist einen für Säugetiere untypischen Verlauf auf. Nach erfolgter Geburt setzt er in der Regel mehrere Jahre aus. Zumeist wird nach fast zweijähriger Tragzeit ein Jungtier geboren, das in der mütterlichen Herde aufwächst. Junge weibliche Tiere verbleiben später in der Herde, die jungen männlichen verlassen diese.

 

Die wissenschaftliche Erstbeschreibung des Afrikanischen Elefanten erfolgte im Jahr 1797 mit einer formalen artlichen Trennung des Afrikanischen vom Asiatischen Elefanten. Der heute gebräuchliche Gattungsname Loxodonta wurde offiziell erst dreißig Jahre später eingeführt. Die Bezeichnung bezieht sich auf markante Zahnunterschiede zwischen den asiatischen und den afrikanischen Elefanten. Im Verlauf des 20. Jahrhunderts wurden mehrere Unterarten unterschieden, darunter auch der Waldelefant des zentralen Afrikas. Letzterer gilt heute genetischen Untersuchungen zufolge als eigenständige Art, die weiteren Unterarten sind nicht anerkannt. Stammesgeschichtlich lässt sich der Afrikanische Elefant erstmals im beginnenden Mittleren Pleistozän belegen. Der Gesamtbestand gilt als gefährdet. Ursachen hierfür sind hauptsächlich die Jagd nach Elfenbein und Lebensraumverlust durch die zunehmend wachsende menschliche Bevölkerung. Der Afrikanische Elefant zählt zu den sogenannten „Big Five“ von Großwildjagd und Safari.

 

(Wikipedia)

This male gelada baboon lives at Colchester Zoo in Essex.

 

These primates are called gelada baboons, however, they aren't baboons, instead, geladas form a separate genus of their own.

Geladas, which spend up to 99% of their time on the ground, are easily recognizable due to the hairless patches of skin on their chests. During the mating season, these chest patches acquire bright crimson coloration in females, leading to their alternate name of bleeding-heart baboon.

These brown and grey primates have a dark face and pale eyelids. Males are twice the size of females, averaging 41 lb. (18.5 kg) to 24 lb. (11 kg) for females. A males head-body length is 27 to 30 in (69 to 74 cm) plus a 17 to 20 in. (45 to 50 cm) tail. A females head-body length is 19 to 21 in (50 to 65 cm) plus a 11 to 16 in. (30 to 41 cm) tail.

They are the last surviving member of a grass-grazing primate group, which were abundant and widespread in the past. They generally feed upon blades of grasses though they will opportunistically eat fruits, invertebrates, and even cereal crops where agriculture abuts their habitat. Their small, powerful fingers are designed for pulling grass, while small incisors allow them to chew it. When eating, geladas move around with characteristic shuffle gait. When walking, they use all of their four limbs and slide their feet without changing their body posture.

The natural range of this species is restricted to Ethiopia Highlands, where these animals mainly occur in in the Semien Mountains National Park. During the night, they typically sleep on rocky cliffs and outcrops. In the morning, geladas typically look for food in nearby grasslands, at heights of 6,550 to 16,400 ft. (2,000 to 5,000 m) above sea level.

Geladas are highly social animals, forming so called 'one male units' (OMU). These are female led groups that consist of a single male and multiple females with their young. When a male from the outside challenges the male of the OMU in order to displace it, females of the group may support or oppose both of them, accepting the winning male and fiercely driving away the defeated one. Various OMU's occasionally share the same area, thus forming larger units called bands. As these animals are non-territorial, they may be observed grazing in separate bands in areas with abundant food without any conflicts. Males and females can often be observed grooming each other. In general, all members of the community participate in grooming, which enhances social bonds within the OMU.

Geladas are polygynous, meaning that one male gets an exclusive right to mating with multiple females. Although geladas can mate at any time of the year, births appear to peak during the rainy season. Gestation period lasts for 5 to 6 months, yielding a single baby, which feeds upon its mother's milk for 12 to 18 months. The infant is mainly cared for by its mother, who will carry, groom, nurse and protect the baby, until it reaches the age of independence. Meanwhile, the father will take little part in rearing its offspring. Average life span in the wild is between 14 to 20 years and up to 30 years in captivity.

Predators of geladas are leopards, hyenas, feral dogs, jackals, foxes, servals and bearded vultures.

One of the biggest threats to the population of this species is habitat reduction due to development of agriculture. Additionally, geladas are considered pests and thus shot because of their destruction of crops. According to the IUCN Red List, the total population of Geladas is around 200,000 animals. Currently, this species conservation status is classified as 'Least Concern' but its numbers are decreasing.

Two Guinea baboons (Papio papio) are eating fruits in their natural habitat in Niokolo-Koba National Park, Senegal.

 

The one in the front, a male with long canines and big fur on his shoulders, has a broken finger. I had to capture this humorous moment when he seemed to give me the finger while impudently biting his fruit.

Interestingly, although males have large and sharp canines, they still use their incisors to open fruit.

 

This picture was taken during a visit to the Centre de recherche de Primatologie Simenti, founded by the Cognitive Ethology Lab, Germany, in 2007. Since then, primatologists have been conducting long-term behavioural and ecological research there.

 

More information: www.dpz.eu/en/cognitive-ethology

en.wikipedia.org/wiki/Agama_(genus)

 

An agama is any one of the various small, long-tailed, insect-eating lizards of the genus Agama. The agamid genus is composed of at least 37 species across Africa, where they are the most common lizard. They can be found in many sizes, from 12.5 to 30 cm (5 in. to 1 ft.) in length and a wide variety of colours. One of the best known species is the Agama agama), widespread in sub-Saharan Africa. For Eurasian agamas, see the genus Laudakia.

 

Agamas originally lived in forest and bush across Africa, but have since adapted to live in villages and compounds where their habitat has been cleared. They live inside the thatch of huts and other small spaces, emerging only to feed. If caught out in the open, agamas are able to run quickly on their hind legs to reach shelter. The desert agama can still be found in the dry areas of North Africa. Despite their name, they avoid bare sand.[1]

 

Agamas are active during the day and are often found scampering around to snatch up their favorite foods. They can tolerate greater temperatures than most reptiles, but in the afternoon when temperatures reach around 38°C (100°F) they will settle into the shade and wait for it to cool. Frequent fighting breaks out between males; such fighting involves a lot of bobbing and weaving in an attempt to scare the opponent. If it comes to blows, they lash out with their tails and threaten each other with open jaws. Many older males have broken tails as a result of such fights. Females may sometimes chase and fight one another, while hatchlings mimic the adults in preparation for their future.[1]

 

Agamas are mainly insectivores. Their incisor-like front teeth are designed for quick cutting and chewing of their prey. They may also eat grass, berries, seeds and even the eggs of smaller lizards.

 

Most agamas are polygamous. Males may hold six or more females in their territory for breeding. During courtship, the male bobs his head to impress the female. Occasionally, females initiate courtship by offering their hindquarters to the male and then running until he is able to catch up. The breeding season is typically March-May with eggs being laid in June-September during the season after the rains. Eggs are laid in clutches of up to twelve.[

  

SpeciesListed alphabetically.[2]

  

Ground Agama (Agama aculeata), Namibia

Ground Agama (Agama aculeata), female, TanzaniaAgama aculeata Merrem, 1820 – Ground Agama

Agama agama (Linnaeus, 1758) – Red-headed Rock Agama, Common Agama, Rainbow Agama

Agama anchietae Bocage, 1896 – Western Rock Agama, Anchieta's Agama

Agama armata Peters, 1855 – Tropical Spiny Agama

Agama atra Daudin, 1802 – Southern Rock Agama

Agama bocourti Rochebrune, 1884 – Bocourt's Agama

Agama bottegi Boulenger, 1897 – Somali Agama

Agama boueti Chabanaud, 1917 – Mali Agama

Agama boulengeri Lataste, 1886 – Boulenger's Agama

Agama castroviejoi Padial, 2005 – Mauritania Agama

Agama caudospinosa Meek, 1910 – Elmenteita Rock Agama

Agama cornii Scortecci, 1928 – Scortecci's Agama

Agama doriae Boulenger, 1885 – Nigeria Agama

 

Kenyan Rock Agama (Agama lionotus), male on a wall in Samburu National Reserve, Kenya

Mwanza Flat-headed Rock Agama (Agama mwanzae), male, Serengeti, TanzaniaAgama etoshae McLachan, 1981 – Etosha Agama

Agama finchi Böhme, Wagner, Malonza, Lötters & Köhler, 2005 – Finch's Agama or Malaba Rock Agama

Agama gracilimembris Chabanaud, 1918 – Benin Agama

Agama hartmanni Peters, 1869 – Hartmann's Agama

Agama hispida (Kaup, 1827) – Common Spiny Agama

Agama impalearis Boettger, 1874 – Bibron's Agama

Agama insularis Chabanaud, 1918 – Insular Agama

Agama kaimosae Loveridge, 1935

Agama kirkii Boulenger, 1885 – Kirk's Rock Agama

Agama lionotus Boulenger, 1896 – Kenyan Rock Agama

Agama mehelyi Tornier, 1902 – Mehely's Agama

Agama montana Barbour & Loveridge, 1828 – Montane (Rock) Agama

Agama mossambica Peters, 1854 – Mozambique Agama

Agama mwanzae Loveridge, 1923 – Mwanza Flat-headed Agama

Agama paragama Grandison, 1968 – False Agama

Agama persimilis Parker 1942 – Somali Painted Agama, Similar Agama

Agama planiceps Peters, 1862 – Namib Rock Agama

Agama robecchii Boulenger, 1892 – Robecchi's Agama

Agama rueppelli or Agama ruppelli Vaillant, 1882 – Rüppell's Agama, Arboreal Agama

Agama sankaranica (Chabanaud, 1918 – Senegal Agama

Agama spinosa Gray 1831 – Lanza's Spiny Agama

Agama sylvana Macdonald, 1981

Agama turuensis Loveridge, 1896

Agama weidholzi Wettstein, 1932 – Gambia Agama

 

By the river Sabie

African Buffalos or Cape Buffalos (Syncerus caffer)

 

Am Fluß Sabi

Kaffernbüffel

 

Press 'L' for large Picture!

"L" drücken für großes Bild!

 

Kruger National Park is one of the largest game reserves in Africa. It covers an area of 19,485 km2 (7,523 sq mi) in the provinces of Limpopo and Mpumalanga in northeastern South Africa, and extends 360 km (220 mi) from north to south and 65 km (40 mi) from east to west. The administrative headquarters are in Skukuza. Areas of the park were first protected by the government of the South African Republic in 1898, and it became South Africa's first national park in 1926.

 

To the west and south of the Kruger National Park are the two South African provinces of Limpopo and Mpumalanga. In the north is Zimbabwe, and to the east is Mozambique. It is now part of the Great Limpopo Transfrontier Park, a peace park that links Kruger National Park with the Gonarezhou National Park in Zimbabwe, and with the Limpopo National Park in Mozambique.

 

The park is part of the Kruger to Canyons Biosphere an area designated by the United Nations Educational, Scientific and Cultural Organization (UNESCO) as an International Man and Biosphere Reserve (the "Biosphere").

 

The park has nine main gates allowing entrance to the different camps.

 

(Wikipedia)

 

The African buffalo or Cape buffalo (Syncerus caffer) is a large Sub-Saharan African bovine. Syncerus caffer caffer, the Cape buffalo, is the typical subspecies, and the largest one, found in Southern and East Africa. S. c. nanus (African forest buffalo) is the smallest subspecies, common in forest areas of Central and West Africa, while S. c. brachyceros is in West Africa and S. c. aequinoctialis is in the savannas of East Africa. The adult buffalo's horns are its characteristic feature: they have fused bases, forming a continuous bone shield across the top of the head referred to as a "boss". They are widely regarded as among the most dangerous animals on the African continent, and according to some estimates they gore, trample, and kill over 200 people every year.

 

The African buffalo is not an ancestor of domestic cattle and is only distantly related to other larger bovines. Its unpredictable temperament means that the African buffalo has never been domesticated, unlike its Asian counterpart, the water buffalo. African Cape buffaloes have few predators aside from lions and large crocodiles. As a member of the big five game, the Cape buffalo is a sought-after trophy in hunting.

 

The African buffalo is a very robust species. Its shoulder height can range from 1.0 to 1.7 m (3.3 to 5.6 ft) and its head-and-body length can range from 1.7 to 3.4 m (5.6 to 11.2 ft). Compared with other large bovids, it has a long but stocky body (the body length can exceed the wild water buffalo, which is heavier and taller) and short but thickset legs, resulting in a relatively short standing height. The tail can range from 70 to 110 cm (28 to 43 in) long. Savannah-type buffaloes weigh 500 to 1,000 kg (1,100 to 2,200 lb), with males normally larger than females, reaching the upper weight range. In comparison, forest-type buffaloes, at 250 to 450 kg (600 to 1,000 lb), are only half that size. Its head is carried low; its top is located below the backline. The front hooves of the buffalo are wider than the rear, which is associated with the need to support the weight of the front part of the body, which is heavier and more powerful than the back.

 

Savannah-type buffaloes have black or dark brown coats with age. Old bulls often have whitish circles around their eyes and on their face. Females tend to have more-reddish coats. Forest-type buffaloes are 30-40% smaller, reddish brown in colour, with much more hair growth around the ears and with horns that curve back and slightly up. Calves of both types have red coats.

 

A characteristic feature of the horns of adult male African buffalo (Southern and Eastern populations) is that the bases come very close together, forming a shield referred to as a "boss". From the base, the horns diverge downwards, then smoothly curve upwards and outwards and in some cases inwards and or backwards. In large bulls, the distance between the ends of the horns can reach upwards of one metre (the record being 64.5 inches 164 cm). The horns form fully when the animal reaches the age of five or six years but the bosses do not become "hard" till 8 to 9 years old. In cows, the horns are, on average, 10–20% smaller, and they do not have a boss. Forest buffalo horns are smaller than those of the savanna buffalo from Southern and Eastern Africa, usually measuring less than 40 centimetres (16 in), and are almost never fused.

 

The African buffalo is one of the most successful grazers in Africa. It lives in swamps and floodplains, as well as mopane grasslands, and the forests of the major mountains of Africa. This buffalo prefers a habitat with dense cover, such as reeds and thickets, but can also be found in open woodland. While not particularly demanding in regard to habitat, they require water daily, and so they depend on perennial sources of water. Like the plains zebra, the buffalo can live on tall, coarse grasses. Herds of buffalo mow down grasses and make way for more selective grazers. When feeding, the buffalo makes use of its tongue and wide incisor row to eat grass more quickly than most other African herbivores. Buffaloes do not stay on trampled or depleted areas for long.

 

Other than humans, African buffaloes have few predators and are capable of defending themselves against (and killing) lions. Lions do kill and eat buffalo regularly, and in some regions, the buffaloes are the lions' primary prey. It typically takes quite a few lions to bring down a single adult buffalo. Usually, the entire pride joins the hunt; however, several incidents have been reported in which lone adult male lions have been able to successfully bring down adult animals. The average-sized crocodile typically attacks only old solitary animals and young calves, though they can kill healthy adults, and exceptionally large, old male Nile crocodiles may become semi-habitual predators of buffalo. Also, this crocodilian is the only animal that typically takes down an adult buffalo alone, whereas a pride attack is the preferred method of lions when taking down such large prey. The cheetah, leopard, and spotted hyena are normally a threat only to newborn calves, though very large clans of spotted hyenas have been recorded killing cows (mainly pregnant ones) and, on very rare occasions, full-grown bulls.

 

The African buffalo is susceptible to many diseases, including bovine tuberculosis, corridor disease, and foot and mouth disease. As with many diseases, these problems remain dormant within a population as long as the health of the animals is good. These diseases do, however, restrict the legal movements of the animals and fencing infected areas from unaffected areas is enforced. Some wardens and game managers have managed to protect and breed "disease-free" herds which become very valuable because they can be transported. Most well-known are Lindsay Hunt's efforts to source uninfected animals from the Kruger National Park in South Africa. Some disease-free buffaloes in South Africa have been sold to breeders for close to US$130,000.

 

Herd size is highly variable. The core of the herds is made up of related females, and their offspring, in an almost linear dominance hierarchy. The basic herds are surrounded by subherds of subordinate males, high-ranking males and females, and old or invalid animals. The young males keep their distance from the dominant bull, which is recognizable by the thickness of his horns. During the dry season, males split from the herd and form bachelor groups. Two types of bachelor herds occur: ones made of males aged four to seven years and those of males 12 years or older. During the wet season, the younger bulls rejoin a herd to mate with the females. They stay with them throughout the season to protect the calves. Some older bulls cease to rejoin the herd, as they can no longer compete with the younger, more aggressive males. Males have a linear dominance hierarchy based on age and size. Since a buffalo is safer when a herd is larger, dominant bulls may rely on subordinate bulls and sometimes tolerate their copulation.

 

Adult bulls spar in play, dominance interactions, or actual fights. A bull approaches another, lowing, with his horns down, and waits for the other bull to do the same thing. When sparring, the bulls twist their horns from side to side.[27] If the sparring is for play, the bull may rub his opponent's face and body during the sparring session. Actual fights are violent but rare and brief. Calves may also spar in play, but adult females rarely spar at all.

 

African buffaloes are notable for their apparent altruism. Females appear to exhibit some sort of "voting behavior". During resting time, the females stand up, shuffle around, and sit back down again. They sit in the direction they think they should move. After an hour of more shuffling, the females travel in the direction they decide. This decision is communal and not based on hierarchy or dominance. When chased by predators, a herd sticks close together and makes it hard for the predators to pick off one member. Calves are gathered in the middle. A buffalo herd responds to the distress call of a captured member and tries to rescue it. A calf's distress call gets the attention of not only the mother, but also the herd. Buffaloes engage in mobbing behavior when fighting off predators. They have been recorded killing a lion and chasing lions up trees and keeping them there for two hours, after the lions have killed a member of their group. Lion cubs can get trampled and killed. In one videotaped instance, known as the Battle at Kruger, a calf survived an attack by both lions and a crocodile after intervention of the herd.

 

African buffaloes make various vocalizations. Many calls are lower-pitched versions of those emitted by domestic cattle. They emit low-pitched, two- to four-second calls intermittently at three- to six-second intervals to signal the herd to move. To signal to the herd to change direction, leaders emit "gritty", "creaking gate" sounds. When moving to drinking places, some individuals make long "maaa" calls up to 20 times a minute. When being aggressive, they make explosive grunts that may last long or turn into a rumbling growl. Cows produce croaking calls when looking for their calves. Calves make a similar call of a higher pitch when in distress. When threatened by predators, they make drawn-out "waaaa" calls. Dominant individuals make calls to announce their presence and location. A version of the same call, but more intense, is emitted as a warning to an encroaching inferior. When grazing, they make various sounds, such as brief bellows, grunts, honks, and croaks.

 

African buffaloes mate and give birth only during the rainy seasons. Birth peak takes place early in the season, while mating peaks later. A bull closely guards a cow that comes into heat, while keeping other bulls at bay. This is difficult, as cows are quite evasive and attract many males to the scene. By the time a cow is in full estrus, only the most dominant bull in the herd/subherd is there.

 

Cows first calve at five years of age, after a gestation period of 11.5 months. Newborn calves remain hidden in vegetation for the first few weeks while being nursed occasionally by the mother before joining the main herd. Older calves are held in the centre of the herd for safety.

 

The maternal bond between mother and calf lasts longer than in most bovids. However, when a new calf is born, the bonding ends and the mother keeps her previous offspring at bay with horn jabs. Nevertheless, the yearling follows its mother for another year or so. Males leave their mothers when they are two years old and join the bachelor groups. Young calves, unusually for bovids, suckle from behind their mothers, pushing their heads between the mothers' legs.

 

The current status of the African buffalo is dependent on the animal's value to both trophy hunters and tourists, paving the way for conservation efforts through anti-poaching patrols, village crop damage payouts, and CAMPFIRE payback programs to local areas.

 

The African buffalo was listed as least concern by the IUCN "as the species had a global population estimated at nearly 900,000 animals, of which more than three-quarters are in protected areas. However, in 2019 the African buffalo was listed as a near threatened species, with only 400,000 individuals left. While some populations (subspecies) are decreasing, others will remain unchanged in the long term if large, healthy populations continue to persist in a substantial number of national parks, equivalent reserves and hunting zones in southern and eastern Africa."

 

In the most recent and available census data at continental scale, the total estimated numbers of the three African buffalo savanna subspecies (S. c. caffer, S. c. brachyceros, S. c. aequinoctialis) are at 513,000 individuals.

 

In the past, numbers of African buffaloes suffered their most severe collapse during the great rinderpest epidemic of the 1890s, which, coupled with pleuro-pneumonia, caused mortalities as high as 95% among livestock and wild ungulates.

 

Being a member of the big five game group, a term originally used to describe the five most dangerous animals to hunt, the Cape buffalo is a sought-after trophy, with some hunters paying over $10,000 for the opportunity to hunt one. The larger bulls are targeted for their trophy value, although in some areas, buffaloes are still hunted for meat.

 

One of the "big five" African game, it is known as "the Black Death" or "the widowmaker", and is widely regarded as a very dangerous animal. According to some estimates, it gores and kills over 200 people every year. African buffaloes are sometimes reported to kill more people in Africa than any other animal, although the same claim is also made of hippopotamuses and crocodiles. These numbers may be somewhat overestimated; for example, in the country of Mozambique, attacks, especially fatal ones, were much less frequent on humans than those by hippos and, especially, Nile crocodiles. In Uganda, on the other hand, large herbivores were found to attack more people on average than lions or leopards and have a higher rate of inflicting fatalities during attacks than the predators (the African buffalo, in particular, killing humans in 49.5% of attacks on them), but hippos and even elephants may still kill more people per annum here than buffaloes. African buffaloes are notorious among big-game hunters as very dangerous animals, with wounded animals reported to ambush and attack pursuers.

 

(Wikipedia)

 

Der Kruger-Nationalpark (deutsch häufig falsch Krüger-Nationalpark) ist das größte Wildschutzgebiet Südafrikas. Er liegt im Nordosten des Landes in der Landschaft des Lowveld auf dem Gebiet der Provinz Limpopo sowie des östlichen Abschnitts von Mpumalanga. Seine Fläche erstreckt sich vom Crocodile-River im Süden bis zum Limpopo, dem Grenzfluss zu Simbabwe, im Norden. Die Nord-Süd-Ausdehnung beträgt etwa 350 km, in Ost-West-Richtung ist der Park durchschnittlich 54 km breit und umfasst eine Fläche von rund 20.000 Quadratkilometern. Damit gehört er zu den größten Nationalparks in Afrika.

 

Das Schutzgebiet wurde am 26. März 1898 unter dem Präsidenten Paul Kruger als Sabie Game Reserve zum Schutz der Wildnis gegründet. 1926 erhielt das Gebiet den Status Nationalpark und wurde in seinen heutigen Namen umbenannt. Im Park leben 147 Säugetierarten inklusive der „Big Five“, außerdem etwa 507 Vogelarten und 114 Reptilienarten, 49 Fischarten und 34 Amphibienarten.

 

(Wikipedia)

 

Der Kaffernbüffel (Syncerus caffer), auch Schwarzbüffel, Afrikanischer Büffel oder Steppenbüffel genannt, ist ein sehr großer Vertreter aus der Familie der Hornträger, welcher in weiten Teilen des östlichen und südlichen Afrikas vorkommt. Dort bewohnt er offene Savannenlandschaften und bewaldete Flussgebiete des Tief- und Hochlands. Charakteristisch ist der robuste Körperbau und die großen, abwärts geschwungenen Hörner, die auf großen Hornbasen auf der Stirn aufsitzen. Das Sozialsystem des Kaffernbüffels ist komplex und umfasst Herden aus verwandten Kühen mit Jungtieren und gelegentlich Bullen, Junggesellengruppen aus männlichen Tieren und einzelne männliche Individuen. Vor allem die jeweiligen Herden zeigen ein dynamisches Verhalten durch permanentes Aufspalten und Wiedervereinen. Sie ziehen in räumlich begrenzten Aktionsgebieten in einem jährlich von Witterung und Nahrungsangebot vorgegebenen Rhythmus umher. Die sie begleitenden oder flankierenden Bullen sind regelmäßig in Dominanzkämpfe um das Paarungsvorrecht eingebunden, die mit Kopf- oder Körperrammen durchgeführt werden. Kühe bringen in der Regel ein einzelnes Jungtier zur Welt, welches rund ein bis anderthalb Jahre gesäugt wird. Der weibliche Nachwuchs verbleibt zumeist in der Herde, der männliche verlässt sie später. Die Nahrung umfasst überwiegend harte Gräser, in Trockenzeiten kann aber auch der Anteil an weicheren Pflanzen deutlich ansteigen. Dadurch haben die Tiere saisonal und regional bedingt unterschiedliche Ernährungsstrategien. Die Bestände sind im Übergang vom 19. zum 20. Jahrhundert in weiten Teilen des Verbreitungsgebietes in Folge einer Rinderpest-Epidemie zusammengebrochen. Heute ist der Kaffernbüffel weitgehend auf Schutzgebiete beschränkt, gilt aber als häufig und nicht bedroht. Die Art wurde 1779 beschrieben.

 

Der Kaffernbüffel stellt den größten Vertreter der afrikanischen Rinder (Bovini) dar und erreicht die Ausmaße seiner asiatischen, wildlebenden Verwandten. Er besitzt eine Kopf-Rumpf-Länge von 240 bis 340 cm (zuzüglich eines 50 bis 110 cm langen Schwanzes), eine Schulterhöhe von 148 bis 175 cm und ein Gewicht von 350 bis 900 kg. Ein Sexualdimorphismus ist deutlich ausgeprägt, die Bullen sind größer und können bis zu doppelt so schwer wie die Weibchen werden. Untersuchungen von Tieren aus Tansania ergaben ein Gewicht für männliche Tiere von 661 bis 849 kg, für weibliche von 426 bis 468 kg. Entsprechende Angaben für Tiere aus Sambia liegen bei 472 bis 723 kg beziehungsweise 386 bis 536 kg. Insgesamt ist der Kaffernbüffel sehr kräftig gebaut und besitzt verhältnismäßig kurze Gliedmaßen und einen Buckel oberhalb der Schultern. Das Fell hat in beiden Geschlechtern eine rabenschwarze Färbung, bei alten Bullen kann es ausdünnen, so dass Flecken von dunkler Haut am Kopf sichtbar werden. Der Schwanz endet in einer auffälligen, ebenfalls dunklen Quaste. Der massige Kopf ist mit einem breiten Maul ausgestattet, die Nase ist feucht und nackt. Die großen, hängenden Ohren sind gefranst. Besonders auffällig zeigen sich die großen, am Ende spitzen Hörner, die quer über den vorderen Kopf verlaufend ansetzen und seitlich nach unten biegen. Sie spannen 73 bis 134 cm weit auseinander, die Länge der einzelnen Hörner über die Krümmung gemessen ist mit 66 bis 116 cm nahezu identisch zur Spannweite. Das längste jemals gemessene Horn erreichte 163 cm und stammt von einem Bullen aus dem Lake-Manyara-Nationalpark in Tansania. Die Hörner sind nicht geriffelt und vor allem bei Männchen besonders massiv, ihre Basen dehnen sich deutlich über den Vorderkopf und treffen sich auf der Mittellinie des Schädels. Es gibt aber keine Übereinstimmung mit dem Alter der Tiere und der Größe der Hornbasis. Im Gegensatz zu weiblichen Tieren mit ihren grazileren Hörnern wachsen bei den männlichen zwischen den Hornansätzen keine Haare. Die Hufe haben eine große, gerundete Form, zudem treten Afterklauen auf. Die Hinterfußlänge beträgt 56 bis 61 cm. Bestimmte Drüsen zum Absetzen von Sekreten sind nicht ausgebildet. Kühe haben einen kleinen, gerundeten Euter mit zwei Zitzenpaaren.

 

Der Schädel wird 44,8 bis 57,5 cm lang und am Warzenfortsatz des Schläfenbeins 24,1 bis 31,6 cm breit. Allgemein ist er kurz und breit und im Profil deutlich gerundet. Eine Voraugengrube (Fossa praeorbitalis) und eine Siebgrube (Fossa ethmoidalis) sind nicht ausgebildet. Im oberen Gebiss ist typisch für die Wiederkäuer anstatt der Schneidezähne eine Hornplatte ausgebildet. Die Backenzähne sind stark hochkronig (hypsodont) mit einer Kronenhöhe von 5 cm bei jungadulten Tieren, auf der Kaufläche sind scharfe Zahnschmelzleisten ausgebildet. Das Durchbruchsmuster der Zähne kann zur Altersbestimmung verwendet werden, der Eckzahn des Unterkiefers ist der letzte durchbrechende Zahn des Dauergebisses. Er erscheint mit viereinhalb bis fünfeinhalb Jahren.

 

Der Kaffernbüffel kommt in weiten Teilen des östlichen und südlichen Afrikas vor. Das Verbreitungsgebiet reicht vom südlichen Äthiopien und Somalia im Nordosten südwärts über Kenia, Tansania, Uganda und dem äußersten Osten und Südosten der Demokratischen Republik Kongo in Ostafrika sowie über Sambia, Simbabwe, Mosambik bis nach Südafrika, dem Nordosten von Namibia und Süden von Botswana. Die Art bewohnt eine Vielzahl von verschiedenen Landschaftstypen, die gut durchfeuchtete Savannen, Sümpfe und Überflutungsebenen umfassen. Daneben tritt sie aber auch in trockeneren Savannen und Auwäldern trockener Habitate auf, sofern Wasser zur Verfügung steht. Die Tiere sind auch in montanen Waldgebieten bis auf 3000 m Höhe anzutreffen, extrem hohe Nachweise liegen bei 4700 m am Mount Kenya. In besonders zuträglichen Gebieten, etwa im Lake-Manyara-Nationalpark in Tansania, kann der Kaffernbüffel mit rund 20 Individuen je Quadratkilometer in sehr hoher Anzahl auftreten, im Ngorongoro-Krater liegt die Populationsdichte entsprechend bei etwa 11, in der Serengeti bei 8 Individuen auf einer vergleichbar großen Fläche. In Trockenlandschaften wie etwa dem Lowveld in Südafrika geht sie auf bis zu 2 Individuen je Quadratkilometer zurück.

 

Der Kaffernbüffel besitzt ein komplexes Sozialsystem. Typisch ist der Verband aus miteinander verwandten Weibchen und ihren Jungtieren, der als Herde bezeichnet wird und in dem sich auch Bullen unterschiedlichen Alters aufhalten können. Die Größe der Herde variiert von einigen Dutzend bis mehreren Tausend Individuen. Die extrem großen Gruppen stellen aber vermutlich Herdenverbände dar, die nur temporär bestehen und keine innere Hierarchie besitzen. Ihre Bildungen sind abhängig von der Jahreszeit und dem Angebot an Nahrung und Wasser. Es gibt jedoch eine Variation der Herdengröße über das Jahr, die zu größeren Gruppen in der feuchten und kleineren in der trockenen Jahreszeit führen, wobei die Gruppen sich regelmäßig teilen und wieder vereinen.[5] Solche Trennungen und Neuformierungen sind aber nicht nur an äußere Bedingungen geknüpft, sondern teilweise auch an die Größe der Herde und ihre soziale Struktur, wie Untersuchungen am Lake Manyara ergaben. Die hierarchische Struktur einer Herde besteht aus dominanten Weibchen, die die Herde anführen, gefolgt von Kühen mit sehr jungen Kälbern und Jungtieren im mittleren Teil. Den Abschluss der Herde bilden Tiere mit untergeordnetem Status; die Position eines Tiers in der Herde gibt somit Aufschluss über dessen soziale Stellung. Sie bestimmt darüber hinaus auch die Qualität der Nahrung und die allgemeine körperliche Kondition der Kuh. Häufig führt die Geburt eines Kalbes zu einer höheren sozialen Stellung innerhalb der Herde. Es wird meist angenommen, dass die einzelnen Herden (ohne temporäre Aufsplittung) stabile Verbände darstellen und erwachsene Kühe nur selten ihre angestammte Gruppe verlassen, einige Herden wurden über einen Zeitraum von bis zu fünf Dekaden dokumentiert. Beobachtungen aus dem Chobe-Nationalpark in Botswana zeigen aber, dass gelegentlich weibliche Tiere ihre Herde verlassen. Eine Kuh legte während eines solchen Herdenwechsels bis zu 133 km zurück. Ein Wiederaufsuchen der alten Herde wurde dabei nicht festgestellt, möglicherweise handelt es sich bei wechselnden Tieren um solche mit einem eher geringen Status. Bullen leben einzeln oder schließen sich zu Junggesellengruppen mit einer Größe von 5 bis 10 Individuen zusammen, die zeitlich begrenzt bestehen. Innerhalb dieser Junggesellengruppen kommt es permanent zu Dominanzkämpfen um das Paarungsvorrecht. Kurzfristig können Bullen auch von Junggesellengruppen zu Herden und umgekehrt wechseln, innerhalb der Herde sind sie aber nicht in das dortige Sozialgefüge eingebunden. Bemerkenswert ist, dass Bullen, wenn sie sich über längere Zeit in Herden aufhalten, regelmäßig an Gewicht verlieren, da sie durchschnittlich weniger fressen. Vermutlich sind die hohen energetischen Kosten, die für die Fortpflanzung aufgebracht werden müssen, ursächlich dafür verantwortlich, weswegen Bullen die Herden nach kürzerer Zeit wieder verlassen.

 

Die Herden des Kaffernbüffels sind nicht migratorisch und legen somit keine großen Entfernungen zurück. Sie halten sich in mehr oder weniger begrenzten Aktionsräumen von 100 bis über 1000 km² auf. Innerhalb dieser Aktionsräume folgen sie einem regelmäßigen Wanderungsmuster über das Jahr hinweg, das durch den Zyklus von Niederschlägen und Pflanzenwachstum bestimmt wird. Diese einem bestimmten Rhythmus folgenden Wanderungen auf festgelegten Pfaden und Wegen führen dazu, dass der Boden aufgearbeitet und dadurch neues Pflanzenwachstum angeregt wird, was das wiederholte Weiden in der gleichen Region ermöglicht. Typische Wanderungsbewegungen führen zu Wasser- und Weidestellen, sie sind täglich umfangreicher, je weiter zugängliches Wasser verstreut liegt. Im Krüger-Nationalpark in Südafrika werden so Distanzen von durchschnittlich 3,35 km täglich überwunden, in der ostafrikanischen Serengeti sind es dazu im Vergleich bis zu 30 km. Einzelne Bullen und Junggesellengruppen haben wesentlich kleinere Territorien von 3 bis 4 km² Größe. Sie flankieren den Wanderungsverlauf der Herde. Der Tagesablauf folgt ähnlichen Mustern wie bei zahlreichen anderen Wiederkäuern und ist geprägt von der Nahrungsaufnahme und der daraus folgenden Ruhe und dem Wiederkäuen. Insgesamt ist die Dauer der jeweiligen Aktivität abhängig von der Qualität und Quantität des lokalen und saisonalen Nahrungsangebots. Die hauptsächlichen Aktivitäten erfolgen tagsüber, in Gebieten mit großem Jagddruck aber auch nachts. Am Lake Manyara fressen die Tiere zwischen 10:00 Uhr und 14:00 Uhr, wobei die Dauer je nach Intensität zwischen 1,5 und 4,5 Stunden betragen kann, was etwa 22 bis 56 % der Tagesaktivität entspricht (durchschnittlich 37 %). Die Nahrungsaufnahme trennt hier zwei Ruheperioden, die meist mit Wiederkäuen verbracht werden, häufig mit dem Körper im Wasser liegend, und wiederum mehrere Stunden andauern können. Sie nehmen zwischen 11 und 49 % des täglich verfügbaren Zeitbudgets in Anspruch (durchschnittlich 31 %). Somit verbringt der Kaffernbüffel am Lake Manyara jährlich etwa zwei Drittel seiner Tageszeit mit Fressen und dem zugehörigen Verdauen. Dabei ist die Dauer der Verdauung gegenläufig zur Dauer der Nahrungsaufnahme, da ein Tier in der Regenzeit durch das üppigere Angebot qualitativ hochwertiger Nahrung in kürzerer Zeit mehr fressen kann, aber längere Zeit zum Wiederkäuen benötigt. In der Trockenzeit mit einem qualitativ minderwertigerem Nahrungsangebot frisst er länger, verdaut aber kürzer. Im Chebera-Churchura-Nationalpark in Äthiopien weiden die Tiere in den frühen Morgen- und Abendstunden und verbrauchen dafür bis zur Hälfte ihres täglichen Zeitbudgets, ein weiteres gutes Drittel nimmt die Ruhe in Anspruch. Ähnlich lang verbringt der Kaffernbüffel im Krüger-Nationalpark mit dem Fressen, dies erfolgt hier aber teilweise auch nachts.

 

Zum Komfortverhalten gehören unter anderem Schlammbäder, um eventuell Insektenbisse zu vermeiden und Ruhephasen an erhöhten Landmarken. Letzteres dient wahrscheinlich dazu, kühle Brisen zu erhaschen, da der Kaffernbüffel allgemein schlecht an Hitze angepasst ist. Generell ist der Kaffernbüffel eher still. Bekannte Lautäußerungen sind ein Grunzen und Schnaufen, letzteres wird ausgestoßen, wenn ein Tier alarmiert ist oder zu Laufen anfängt. Ein Bellen äußert der Kaffernbüffel bei starken Verletzungen, beispielsweise durch einen Beutegreifer, das andere Artgenossen häufig animiert, zur Unterstützung zu kommen. Schwer verletzte Tiere suchen zum Schutz andere Kaffernbüffel auf. Ein Warnruf bei Gefahr besteht nicht, aber vor allem in der Nacht kann eine Herde sehr still werden, so dass ein Aufspüren sehr schwierig ist. Zwischen dominanten und unterwürfigen Tieren besteht ein ritualisiertes Verhaltensmuster. Erstere halten den Kopf und die Schultern nach oben, während die Nase nach unten gerichtet ist, so dass die Hörner optimal präsentiert werden. Bei letzteren liegt der Kopf tief und parallel zum Boden, teilweise führt das unterlegene Tier auch den Kopf zwischen die Hinterbeine des dominanten und stößt ein Bellen aus. Der Sehsinn ist recht gut entwickelt, ebenso wie das Gehör. Aufgrund der fehlenden Drüsen spielen Sekrete für die innerartliche Kommunikation nur eine untergeordnete Rolle.

 

Als weitgehende Offenlandart besteht die Hauptnahrung des Kaffernbüffels aus Gräsern. Hierbei bevorzugt er langhalmige Gräser vor breitblättrigen und toleriert auch solche mit einem hohen Faseranteil. Er meidet aber solche mit niedrigem Nährstoffgehalt oder Pflanzen, die ätherische Öle enthalten. Zu den am häufigsten verzehrten Pflanzen gehören Hundszahngräser, Fingerhirsen und Rispenhirsen sowie Vertreter der Gattungen Sporobolus und Heteropogon. Bei hoher Verfügbarkeit an Gräsern – etwa zur Regenzeit – frisst ein Tier große Mengen und ist wenig wählerisch, bei geringerer Verfügbarkeit oder schlechterer Qualität geht es eher selektiv vor. In der Serengeti bevorzugt der Kaffernbüffel in der Regenzeit weitgehend Grashalme, in der Trockenzeit, wenn die Gräser verschwinden, zieht er sich in Flussniederungen zurück, wo das Nahrungsangebot umfangreicher ist. In trockenen Landschaften wie der Sukkulent-Karoo des südlichen Afrikas, wo Gräser weitaus seltener sind, stellt weicheres Pflanzenmaterial einen größeren Teil der vertilgten Menge. Hier ergaben Untersuchungen, dass während der trockenen Jahresabschnitte der Anteil an weichen Pflanzen bis zu 33 % erreichen kann, während er in der feuchteren auf etwa 28 % absinkt. Die Tiere fressen dann unter anderem auch Akaziengewächse, Sternbüsche, Rautengewächse oder den Kap-Bleiwurz. In der grasreicheren Nama-Karoo konnten dagegen derartige Unterschiede nicht festgestellt werden, da Gräser hier 95 % der aufgenommenen Nahrung ausmachen. Allerdings konnte hier ein Wechsel an bevorzugten Pflanzen innerhalb der verschiedenen Jahreszeiten ausgemacht werden. So dominierten Sporobolus-Gräser in den feuchten Jahresabschnitten, Themada-Gräser dagegen in den trockenen. Täglich muss ein Tier rund 2,2 % seiner Körpermasse vertilgen, was in etwa 14 bis 15 kg entspricht. Da der Kaffernbüffel von Wasser abhängig ist, entfernt er sich selten weiter als 8 bis 20 km von offenen Wasserstellen. Bedeutend sind darüber hinaus Salzlecken, unter anderem auch von schwitzenden Artgenossen, oder das Fressen eisenhaltiger Substrate vor allem in großen Höhenlagen wie am Mount Kenya.

 

Die Paarung kann beim Kaffernbüffel prinzipiell das ganze Jahr über erfolgen. In Regionen mit ausgeprägten Jahreszeiten und somit einem qualitativ und quantitativ schwankenden Nahrungsangebot ist sie jedoch saisonal beschränkt. In der Serengeti werden Paarungen von November und zunehmend bis Juli beobachtet, in der südafrikanischen Provinz Limpopo sind sie von Regenfällen und dem jeweiligen Nahrungsangebot abhängig. Letzteres lässt darauf schließen, dass die körperliche Kondition der weiblichen Tiere ausschlaggebend für die Paarung und Geburt ist. Kühe gebären ihr erstes Kalb mit 4 bis 5 Jahren, Bullen sind mit 8 bis 9 Jahren voll geschlechtsreif. Die männlichen Tiere sind in ein hierarchisches System eingebunden, welches die Fortpflanzung regelt. Dazu führen die männlichen Tiere regelmäßig Paarungs- und Dominanzwettkämpfe durch, die unter anderem aus Kopf- und Körperrammen bestehen; ersteres findet häufig zwischen dominanten und unterlegenen, letzteres zwischen gleichrangigen Tieren statt. Sie beginnen damit, dass ein Tier mit erhobenem Kopf und nach unten weisender Nase stillsteht, vergleichbar dem Dominanzverhalten. Oftmals wird der Kopf dabei auch ab- und aufwärts geführt. Die ganze Prozedur kann schon in 30 m Entfernung voneinander starten und ist teilweise mit einem tiefen Grummeln verbunden. Danach kommt es zum Angriff, wobei das Tier den Kopf so weit absenkt, dass beim Aufeinandertreffen die Energie auf die verstärkte Basis der Hörner gelenkt wird. Der Gewinner wird durch die Kraft und Geschwindigkeit bestimmt, der Verlierer sucht im Abschluss zumeist das Weite, wird manchmal aber vom Sieger bis zu 100 m verfolgt.

 

Bullen testen die Empfangsbereitschaft der Kühe beständig durch Beschnuppern der Geschlechtsorgane oder des Urins und zeigen dabei ein Flehmen. Der Sexualzyklus der Kühe dauert etwa 18 bis 22 Tage, der Östrus selbst einen Tag. Paarungswillige Bullen legen ihr Kinn auf den Rumpf der Kuh und signalisieren so Bereitschaft. Die Kuh wiederum erwidert ihre Bereitschaft durch Stillstehen und Schwanzbewegung. Die Kopulation währt eher kurz und wird innerhalb von 30 Minuten mehrfach wiederholt. Die Tragezeit der Kühe dauert bis zu 11,5 Monate (durchschnittlich 340 Tage), das Geburtsintervall beträgt 15 bis 24 Monate. Meist kommt nur ein Kalb zur Welt, das 39 bis 41 kg schwer ist; Zwillingsgeburten sind sehr selten. Das Geburtsgewicht schwankt leicht mit der Jahreszeit. Das Fell der Jungtiere ist schwarz oder dunkelgrau und ändert teilweise während des Wachstums die Farbe. Das Kalb benötigt mehrere Stunden nach der Geburt, um sicher zu stehen und bewegt sich in den folgenden Wochen eher langsam und behäbig fort. Mutter und Jungtier sind nach der Geburt etwas getrennt von der Herde, in dieser Zeit achtet das Muttertier sorgfältig auf das Junge. Insgesamt ist die Herde sehr aufmerksam auf Jungtiere, so dass eine Erbeutung durch Beutegreifer selten erfolgt. Das Junge saugt unregelmäßig, anfänglich bis zu 10 Minuten lang. Nach rund vier Wochen wird es etwa 5 Minuten lang gestillt. Während der Wachstumsphase spielen oder tummeln Jungtiere kaum, jungadulte Bullen führen aber Sparringskämpfe durch. Die Stillzeit endet unterschiedlich, abhängig von der Dichte der Population. Am Lake Manyara mit einer hohen Individuenzahl werden Jungtiere nach anderthalb Jahren entwöhnt, in der weniger dicht besiedelten Serengeti schon nach zehn Monaten, wenn das Muttertier im siebten Monat trächtig mit neuem Nachwuchs ist. Männliche Tiere verlassen die mütterliche Herde, weibliche verbleiben dort. Durchschnittlich liegt die Lebenserwartung bei 18 bis 20 Jahren in freier Wildbahn, einzelne Tiere in menschlicher Gefangenschaft wurden über 30 Jahre alt.

 

Den bedeutendsten Fressfeind stellt der Löwe dar. Aufgrund des guten Augenlichts kann der Kaffernbüffel Löwen schon aus bis zu 1,5 km Entfernung wahrnehmen. Einzelne Tiere verteidigen sich gegen Löwenangriffe, indem sie sich mit dem Rücken zu einem Hindernis positionieren oder im Gebüsch verstecken, so dass die Beutegreifer nicht die verletzlichen Hinterbeine erreichen können. Auch Flucht ist möglich, da individuelle Tiere Geschwindigkeiten von bis zu 55 km/h über kurze Distanzen erreichen. Gruppen stellen sich häufig im Kreis auf und präsentieren eine Phalanx von Hörnern. Unter Umständen greift der Kaffernbüffel auch Löwen direkt an, vertreibt sie, wirbelt sie mit den Hörnern durch die Luft oder trampelt sie mitunter zu Tode. Die Raubkatzen können einen ausgewachsenen Büffel nur durch einen Kehlbiss töten, da sie nicht die Kraft aufbringen, das Genick zu brechen. Außerdem wirkt die Haut des Kaffernbüffels mit 2 bis 3 cm Dicke schützend. Der Einfluss des Löwen auf die lokalen Bestände des Kaffernbüffels ist unterschiedlich. In der Serengeti verursacht er nur 25 % der Todesfälle beim Kaffernbüffel, am Lake Manyara sind es bis zu 85 %. In der Regel fallen dem Beutegreifer ältere, einzeln wandernde Bullen zum Opfer, selten in Junggesellengruppen lebende Bullen oder Kühe und Jungtiere in Herden. Bemerkenswert in diesem Zusammenhang ist, dass in Gebieten mit größerer Löwenpopulation auch größere Herden des Kaffernbüffels zu beobachten sind. Jungtiere werden gelegentlich auch durch die Tüpfelhyäne und den Leoparden gerissen.

 

Zu den wichtigsten Krankheiten, die den Kaffernbüffel befallen, gehören die Rinderpest, die Lungenseuche und die Tuberkulose, die überwiegend von Hausrindern auf die Wildtiere übertragen werden. Untergeordnet spielen auch der Milzbrand und die Maul- und Klauenseuche eine Rolle. Zudem sind die Tiere als Reservoir des Küstenfiebers bekannt. Gegen einige originär tropische oder afrikanische Erkrankungen wie die Schlafkrankheit, die durch die Tsetse-Fliege übertragen wird, ist der Kaffernbüffel immun. An äußeren Parasiten sind zahlreiche Zecken belegt, etwa die Gattungen Amblyomma, Rhipicephalus, Hyalomma und Boophilus. Einige dieser Ectoparasiten gelten auch als Überträger der bereits erwähnten Krankheiten. In der Regel leiden ältere Bullen stärker unter dem Befall als jüngere Tiere. Zur Linderung werden Vögel wie der Madenhacker geduldet, der die äußeren Parasiten entfernt.

 

Der Bestand des Kaffernbüffels (und anderer Huftiere) war in der Wende vom 19. zum 20. Jahrhundert stark reduziert worden. Eine Hauptursache stellte eine Rinderpest-Epidemie um 1890 dar, die Krankheit hatte sich ab dem Beginn des 19. Jahrhunderts im Zuge der zunehmenden Besiedlung des südlichen Afrikas durch europäische Einwanderer mit ihrem Hausvieh über den Kontinent ausgebreitet. In Verbindung mit der Lungenseuche führte dies zu einer Mortalitätsrate innerhalb der einzelnen Populationen von etwa 95 %. Auch im Verlauf des 20. Jahrhunderts waren die Rinderpest und zusätzlich der Milzbrand für den Rückgang lokaler Bestände verantwortlich. Hinzu kommt ein hoher Jagddruck, da der Kaffernbüffel in zahlreichen Ländern als Nahrungsressource gilt. Die Fleischnutzung erfolgt aber nicht nur zur Eigenversorgung in der ländlichen Bevölkerung, sondern wird auch im industriellen Maßstab getätigt. Darüber hinaus gilt der Kaffernbüffel als Bestandteil der Big Five und hat den Ruf als eines der gefährlichsten Tiere der afrikanischen Savannen, womit die Trophäenjagd eine hohe Bedeutung erlangt. Nicht zuletzt hat die Zerstörung des Lebensraums einen großen Einfluss auf die einzelnen Bestände, vor allem im Randbereich von Schutzgebieten, wo es zur Konkurrenz mit Nutzvieh kommt. Heute sind die Populationen und Herden des Kaffernbüffels zum Großteil auf Schutzgebiete beschränkt. Da die Art gegenwärtig weit verbreitet ist, wird sie von der IUCN als „ungefährdet“ (Least Concern) eingestuft. Ihr Gesamtbestand wird auf etwa 670.000 Individuen geschätzt. Zu den bedeutendsten Schutzgebieten, in denen der Kaffernbüffel auftritt, gehören die Serengeti und der Lake-Manyara-Nationalpark in Tansania, der Chobe-Nationalpark in Botswana und der Hluhluwe-iMfolozi-Nationalpark sowie der Krüger-Nationalpark in Südafrika.

 

(Wikipedia)

Wanted to do a series about untold adventures of the Kamen Riders in Ventara.

 

Left to right: Incisor, Black Knight, Dragon Knight, Strike, Torque.

 

Figma Kamen Rider Dragon Knight action figures by Good Smile Company.

Nature is camouflaging within itself - The Caucasian squirrel's body reaches a maximum of 25 cm, their total weight is no more than 430 grams, they are small and the colors match the centuries - old olive bark. It is impossible to notice on an olive tree from a distance of 15 meters.

  

Caucasian squirrel is a tree squirrel, they sleep in their nests are usually found in the tree hollows in the centuries-old olive trees. When the temperature drops, they go out to feed again and when the sun goes down they go back to their nests. When I talked to the local people, I learned that Caucasian squirrel kittens are showing up a lot this year. Caucasian squirrel are now an endangered species. I hope tomorrow or after tomorrow I will come across this year's Caucasian squirrel kittens and share their photos with you.

 

The Caucasian squirrel - Sciurus anomalus ; The Caucasian squirrel or Persian squirrel, is a tree squirrel in the genus Sciurus found in temperate broadleaf and mixed forests in south-western Asia.

The species is usually said to have first been described in 1778 by Johann Friedrich Gmelin in the 13th edition of Systema Naturae,and named Sciurus anomalus. However, some authors argue that this work was actually published in 1788, and that the true first description was made by Johann Anton Güldenstädt in 1785.

Description - Caucasian squirrels are small tree squirrels, with a total length of 32 to 36 cm (13 to 14 in), including the 13 to 18 cm (5.1 to 7.1 in) tail, and weighing 250 to 410 g (8.8 to 14.5 oz). The color of the upper body fur ranges from greyish brown to pale grey, depending on the subspecies, while that of the underparts is rusty brown to yellowish, and that of the tail, yellow brown to deep red. The claws are relatively short, compared with those of other tree squirrels, and females have either eight or ten teats.

Samuel Griswold Goodrich described the Caucasian squirrel in 1885 as "Its color is grayish-brown above, and yellowish-brown below".

 

Physical Description - Caucasian squirrels have a dental formula of incisors 1/1, canines 0/0, premolars 1/1, and molars 3/3, totaling 20. They have four fingered fore feet and five fingered hind feet. Sex differences in body length or mass are not evident.

Distribution and habitat -

 

Caucasian squirrels are native to south-western Asia, where they are found from Turkey, and the islands of Gökçeada and Lesbos in the west, Iran in the southeast, and as far as Israel and Jordan in the south.It is one of only two species of the genus Sciurus to be found on Mediterranean islands,and, although Eurasian red squirrels have been recently introduced to some areas, is the only species of Sciurus native to the wider region.

The species mainly lives in forested areas dominated by oak, pine, and pistachio, up to altitudes of 2,000 metres (6,600 ft).

 

Biology and behavior -

The squirrels are diurnal, and solitary, although temporary groups may forage where food is plentiful. Their diet includes nuts, seeds, tree shoots, and buds,with the seeds of oak and pine being particularly favored. Like many other squirrels, they cache their food within tree cavities or loose soil, with some larders containing up to 6 kg (13 lb) of seeds. They live in trees, where they make their dens, but frequently forage on the ground, and are considered less arboreal than Eurasian red squirrels. They commonly nest in tree hollows lined with moss and leaves, and located 5 to 14 m (16 to 46 ft) above the ground, but nests are also sometimes found under rocks or tree roots. Their alarm call is high-pitched, and said to resemble the call of the European green woodpecker, and they mark their territories with urine and dung.

Breeding occurs throughout the year, but is more common in spring or autumn. Litters range from two to seven, with three or four being typical, and the young are fully mature by five or six months of age.

 

Conservation -

A survey in 2008 found that the species remained abundant within Turkey, however declines are noted in population within the Levant region. The guides for a survey in 1993 in Israel stated that they considered the species to be nearly extinct within the area studied. Whilst the Caucasian squirrel is threatened by poaching and deforestation, the declines recorded are not sufficient to qualify them as anything other than "Least Concern" by the International Union for Conservation of Nature.[1] Hunting of the species is banned by the Central Hunting Commission, and the Caucasian squirrel is protected by the Bern Convention and the EU Habitats Directive.

 

This information is sourced from "Wikipedia".

  

Thank you so much for visiting my stream, whether you comments , favorites or just have a look.

I appreciate it very much, wishing the best of luck and good light.

  

© All rights reserved R.Ertug Please do not use this image without my explicit written permission. Contact me by Flickr mail if you want to buy or use. Your comments and critiques are very well appreciated.

 

Lens - hand held or Monopod and definitely SPORT VR on. Aperture is f5.6 and full length. All my images have been converted from RAW to JPEG.

 

I started using Nikon Cross-Body Strap or Monopod on long walks. Here is my Carbon Monopod details : Gitzo GM2542 Series 2 4S Carbon Monopod - Really Right Stuff MH-01 Monopod Head with Standard Lever - Really Right Stuff LCF-11 Replacement Foot for Nikon AF-S 500mm /5.6E PF Lense -

 

Thanks for stopping and looking :)

I bit gruesome but looking better than it did when both incisors were over grown. I didn’t think this squirrel survived the winter then it showed up again this week looking plump, eating well and running like the wind.

Hippopotamus

 

Flusspferd

 

Kruger National Park is one of the largest game reserves in Africa. It covers an area of 19,485 km2 (7,523 sq mi) in the provinces of Limpopo and Mpumalanga in northeastern South Africa, and extends 360 km (220 mi) from north to south and 65 km (40 mi) from east to west. The administrative headquarters are in Skukuza. Areas of the park were first protected by the government of the South African Republic in 1898, and it became South Africa's first national park in 1926.

 

To the west and south of the Kruger National Park are the two South African provinces of Limpopo and Mpumalanga. In the north is Zimbabwe, and to the east is Mozambique. It is now part of the Great Limpopo Transfrontier Park, a peace park that links Kruger National Park with the Gonarezhou National Park in Zimbabwe, and with the Limpopo National Park in Mozambique.

 

The park is part of the Kruger to Canyons Biosphere an area designated by the United Nations Educational, Scientific and Cultural Organization (UNESCO) as an International Man and Biosphere Reserve (the "Biosphere").

 

The park has nine main gates allowing entrance to the different camps.

 

(Wikipedia)

 

The common hippopotamus (/ˌhɪpəˈpɒtəməs/ HIP-ə-POT-ə-məs; Hippopotamus amphibius), or hippo, is a large, mostly herbivorous, semiaquatic mammal and ungulate native to sub-Saharan Africa. It is one of only two extant species in the family Hippopotamidae, the other being the pygmy hippopotamus (Choeropsis liberiensis or Hexaprotodon liberiensis). The name comes from the ancient Greek for "river horse" (ἱπποπόταμος). After the elephant and rhinoceros, the common hippopotamus is the third-largest type of land mammal and the heaviest extant artiodactyl. Despite their physical resemblance to pigs and other terrestrial even-toed ungulates, the closest living relatives of the Hippopotamidae are cetaceans (whales, dolphins, porpoises, etc.) from which they diverged about 55 million years ago.

 

Common hippos are recognisable by their barrel-shaped torsos, wide-opening mouths revealing large canine tusks, nearly hairless bodies, columnar legs and large size; adults average 1,500 kg (3,310 lb) and 1,300 kg (2,870 lb) for males and females respectively. Despite its stocky shape and short legs, it is capable of running 30 km/h (19 mph) over short distances.

 

The common hippopotamus inhabits rivers, lakes and mangrove swamps, where territorial bulls preside over a stretch of river and groups of five to thirty females and young. During the day, they remain cool by staying in the water or mud; reproduction and childbirth both occur in water. They emerge at dusk to graze on grasses. While hippopotamuses rest near each other in the water, grazing is a solitary activity and hippos are not territorial on land. The hippopotamus is among the most dangerous animals in the world as it is highly aggressive and unpredictable. They are threatened by habitat loss and poaching for their meat and ivory canine teeth.

 

The Latin word "hippopotamus" is derived from the ancient Greek ἱπποπόταμος, hippopotamos, from ἵππος, hippos, "horse", and ποταμός, potamos, "river", meaning "horse of the river". In English, the plural is "hippopotamuses", but "hippopotami" is also used; "hippos" can be used as a short plural. Hippopotamuses are gregarious, living in groups of up to thirty animals. A group is called a pod, herd, dale, or bloat.

 

Hippopotami are among the largest living land mammals, being only smaller than elephants and some rhinoceroses. Amongst the extant African megafauna, behind the two African elephant species, they average smaller than the white rhinoceros but are larger by body mass than the black rhinoceros and the giraffe. Mean adult weight is around 1,500 kg (3,310 lb) and 1,300 kg (2,870 lb) for males and females respectively, very large males can reach 2,000 kg (4,410 lb) and exceptional males weighing 2,660 kg (5,860 lb) and 3,200 kg (7,050 lb) have been reported. Male hippos appear to continue growing throughout their lives while females reach maximum weight at around age 25.

 

Hippopotami have barrel-shaped bodies with short legs and long muzzles. Their skeletal structures are graviportal; adapted to carrying their enormous weight, and their specific gravity allows them to sink and move along the bottom of a river. Hippopotamuses have small legs (relative to other megafauna) because the water in which they live reduces the weight burden. Though they are bulky animals, hippopotamuses can gallop at 30 km/h (19 mph) on land but normally trot. They are incapable of jumping but do climb up steep banks. Despite being semiaquatic and having webbed feet, an adult hippo is not a particularly good swimmer nor can it float. It is rarely found in deep water; when it is, the animal moves by porpoise-like leaps from the bottom. The eyes, ears, and nostrils of hippos are placed high on the roof of their skulls. This allows these organs to remain above the surface while the rest of the body submerges. The testes of the males descend only partially and a scrotum is not present. In addition, the penis retracts into the body when not erect. The genitals of the female are unusual in that the vagina is ridged and two large diverticula protrude from the vulval vestibule. The function of these is unknown.

 

The hippo's jaw is powered by a large masseter and a well-developed digastric; the latter loops up behind the former to the hyoid. The jaw hinge is located far back enough to allow the animal to open its mouth at almost 180°. A moderate folding of the orbicularis oris muscle allows the hippo to achieve such a gape without tearing any tissue. The bite force of an adult female has been measured as 8,100 newtons (1,800 lbf). Hippopotamus teeth sharpen themselves as they grind together. The lower canines and lower incisors are enlarged, especially in males, and grow continuously. The incisors can reach 40 cm (1 ft 4 in), while the canines reach up to 50 cm (1 ft 8 in). The canines and incisors are used for combat and play no role in feeding. Hippos rely on their broad horny lips to grasp and pull grasses which are then ground by the molars. The hippo is considered to be a pseudoruminant; it has a complex three-chambered stomach but does not "chew cud".

 

Unlike most other semiaquatic animals, the hippopotamus has very little hair. The skin is 6 cm (2 in) thick, providing it great protection against conspecifics and predators. By contrast, its subcutaneous fat layer is thin. The animals' upper parts are purplish-grey to blue-black, while the under parts and areas around the eyes and ears can be brownish-pink. Their skin secretes a natural sunscreen substance which is red-coloured. The secretion is sometimes referred to as "blood sweat", but is neither blood nor sweat. This secretion is initially colourless and turns red-orange within minutes, eventually becoming brown. Two distinct pigments have been identified in the secretions, one red (hipposudoric acid) and one orange (norhipposudoric acid). The two pigments are highly acidic compounds. They inhibit the growth of disease-causing bacteria, and their light absorption peaks in the ultraviolet range, creating a sunscreen effect. All hippos, even those with different diets, secrete the pigments, so it does not appear that food is the source of the pigments. Instead, the animals may synthesise the pigments from precursors such as the amino acid tyrosine. Nevertheless, this natural sunscreen cannot prevent the animal's skin from cracking if it stays out of water too long.

 

A hippo's lifespan is typically 40–50 years. Donna the Hippo was the oldest living hippo in captivity. She lived at the Mesker Park Zoo in Evansville, Indiana in the US until her death in 2012 at the age of 61.

 

Different from all other large land mammals, hippos are of semiaquatic habits, spending the day in lakes and rivers. They can be found in both savannah and forest areas. Proper habitat requires enough water to submerge in and grass nearby. Larger densities of the animals inhabit quiet waters with mostly firm, smooth sloping beaches. Males may be found in very small numbers in rapid waters in rocky gorges. Hippo mostly live in freshwater habitats, however populations in West Africa mostly inhabit estuarine waters and may even be found at sea. With the exception of eating, most of hippopotamuses' lives occurs in the water. Hippos leave the water at dusk and travel inland, sometimes up to 10 km (6 mi), to graze on short grasses, their main source of food. They spend four to five hours grazing and can consume 68 kg (150 lb) of grass each night.

 

Like almost any herbivore, they consume other plants if presented with them, but their diet in nature consists almost entirely of grass, with only minimal consumption of aquatic plants. Hippos are born with sterile intestines, and require bacteria obtained from their mothers' feces to digest vegetation. Hippos have (albeit rarely) been filmed eating carrion, usually close to the water. There are other reports of meat-eating, and even cannibalism and predation. The stomach anatomy of a hippo is not suited to carnivory, and meat-eating is likely caused by aberrant behaviour or nutritional stress.

 

Hippo defecation creates allochthonous deposits of organic matter along the river beds. These deposits have an unclear ecological function. A 2015 study concluded that hippo dung provides nutrients from terrestrial material for fish and aquatic invertebrates, while a 2018 study found that their dung can be toxic to aquatic life in large quantities, due to absorption of dissolved oxygen in water bodies. Because of their size and their habit of taking the same paths to feed, hippos can have a significant impact on the land across which they walk, both by keeping the land clear of vegetation and depressing the ground. Over prolonged periods, hippos can divert the paths of swamps and channels.

A hippopotamus walking on the grass land in Serengeti National Park in the morning

 

Adult hippos move at speeds up to 8 km/h (5 mph) in water; typically resurfacing to breathe every three to five minutes. The young have to breathe every two to three minutes. The process of surfacing and breathing is subconscious: a hippo sleeping underwater will rise and breathe without waking up. A hippo closes its nostrils when it submerges into the water. As with fish and turtles on a coral reef, hippos occasionally visit cleaning stations and signal, by opening their mouths wide, their readiness for being cleaned of parasites by certain species of fishes. This is an example of mutualism, in which the hippo benefits from the cleaning while the fish receive food.

 

Hippopotamus coexist with a variety of formidable predators. Nile crocodiles, lions and spotted hyenas are known to prey on young hippos. However, due to their aggression and size, adult hippopotamus are not usually preyed upon by other animals. Cases where large lion prides have successfully preyed on adult hippopotamus have been reported; however, this predation is generally rare. Lions occasionally prey on adults at Gorongosa National Park and calves are taken at Virunga. Crocodiles are frequent targets of hippo aggression, probably because they often inhabit the same riparian habitats; crocodiles may be either aggressively displaced or killed by hippopotamuses. In turn, beyond cases of killing the seldom unguarded hippo calf, very large Nile crocodiles have been verified to occasionally prey on "half-grown" hippopotamuses and anecdotally perhaps adult female hippos. Aggregations of crocodiles have also been seen to dispatch still-living bull hippopotamuses that have been previously injured in mating battles with other bulls.

 

(Wikipedia)

 

Der Kruger-Nationalpark (deutsch häufig falsch Krüger-Nationalpark) ist das größte Wildschutzgebiet Südafrikas. Er liegt im Nordosten des Landes in der Landschaft des Lowveld auf dem Gebiet der Provinz Limpopo sowie des östlichen Abschnitts von Mpumalanga. Seine Fläche erstreckt sich vom Crocodile-River im Süden bis zum Limpopo, dem Grenzfluss zu Simbabwe, im Norden. Die Nord-Süd-Ausdehnung beträgt etwa 350 km, in Ost-West-Richtung ist der Park durchschnittlich 54 km breit und umfasst eine Fläche von rund 20.000 Quadratkilometern. Damit gehört er zu den größten Nationalparks in Afrika.

 

Das Schutzgebiet wurde am 26. März 1898 unter dem Präsidenten Paul Kruger als Sabie Game Reserve zum Schutz der Wildnis gegründet. 1926 erhielt das Gebiet den Status Nationalpark und wurde in seinen heutigen Namen umbenannt. Im Park leben 147 Säugetierarten inklusive der „Big Five“, außerdem etwa 507 Vogelarten und 114 Reptilienarten, 49 Fischarten und 34 Amphibienarten.

 

(Wikipedia)

 

Das Flusspferd (Hippopotamus amphibius), auch Nilpferd, Großflusspferd oder Hippopotamus genannt, ist ein großes, pflanzenfressendes Säugetier. Es lebt in Gewässernähe im mittleren und südlichen Afrika. Zusammen mit dem Breitmaulnashorn zählt es zu den schwersten landbewohnenden Säugetieren nach den Elefanten. Mit einem Gesamtbestand von rund 125.000 bis 150.000 Tieren und einem erwarteten weiteren Rückgang der Population ist die Art gefährdet.

 

Zusammen mit dem Zwergflusspferd und zahlreichen ausgestorbenen Arten bildet das Flusspferd die Familie der Flusspferde (Hippopotamidae). Obwohl sie „-pferde“ genannt werden, sind Flusspferde mit den Pferden nicht näher verwandt. Traditionell werden sie in die Ordnung der Paarhufer gestellt. Nach derzeitiger Lehrmeinung sind indessen die Wale die nächsten Verwandten der Flusspferde, die zusammen mit den paraphyletischen Paarhufern das Taxon der Cetartiodactyla bilden.

 

Die Bezeichnung Flusspferd ist eine Lehnübersetzung des griechischen Wortes ἱπποπόταμος hippopótamos (gebildet aus hippos „Pferd“ und potamos „Fluss“). Hippopotamus, der wissenschaftliche Name der Gattung, ist die latinisierte Form des griechischen Wortes. Der Namensbestandteil amphibius bezieht sich auf die amphibische Lebensweise im Wasser und an Land. Hiob Ludolf erklärte den griechischen Namen des Tieres mit der Ähnlichkeit des aus dem Wasser ragenden Kopfes mit einem Pferdekopf.

 

Der Name Nilpferd rührt daher, dass in der Literatur zunächst Flusspferde am Nil beschrieben wurden.

 

Flusspferde sind schwere Tiere mit einem fassförmigen Körper, einem wuchtigen Kopf und kurzen Gliedmaßen. Die kräftigen Beine enden in jeweils vier nach vorne ragenden Zehen, die mit Schwimmhäuten verbunden sind.

 

Flusspferde erreichen eine Kopf-Rumpf-Länge von 2,9 bis 5,1 Metern, wozu noch ein 40 bis 56 Zentimeter langer Schwanz kommt. Die Schulterhöhe beträgt 150 bis 165 Zentimeter. Das Gewicht schwankt zwischen 1000 und 4500 Kilogramm. Männchen sind generell größer und schwerer als Weibchen.

 

Die Haut der Tiere ist so spärlich mit kurzen, feinen Haaren bedeckt, dass sie nackt erscheint. Die Haut ist bräunlich, annähernd kupferfarben gefärbt, an der Oberseite ist sie dunkler und am Bauch purpurn. Auch im Gesicht, speziell um die Augen, Ohren und an den Wangen, können purpurne oder rosafarbene Flecken vorhanden sein.

 

Der Kopf des Flusspferds ist groß und wuchtig. Die Schnauze ist vorne sehr breit, was durch die Eckzahnfächer bedingt ist. Auf Höhe der Prämolaren ist sie stark eingeschnürt. Die Nasenöffnungen liegen erhöht und sind verschließbar. Auch die leicht hervorstehenden Augen und die bis zu 10 Zentimeter langen Ohren sitzen hoch am Kopf, so dass nur sie aus dem Wasser herausragen, wenn das Tier unter der Wasseroberfläche schwimmt.

 

Pro Kieferhälfte haben die Flusspferde zwei oder drei Schneidezähne, einen Eckzahn, vier Prämolaren und drei Molaren. Die Schneide- und Eckzähne wachsen das ganze Leben lang. Die Schneidezähne sind rundlich, glatt und weit voneinander entfernt. Die oberen Schneidezähne sind eher klein und nach unten gerichtet, die unteren sind länger (vor allem das innere Paar) und weisen nach vorne. Die Eckzähne sind hauerartig entwickelt, die unteren sind größer und können eine Gesamtlänge von 70 Zentimetern erreichen (von denen 30 Zentimeter aus dem Zahnfleisch ragen). Die Prämolaren haben in der Regel einen Höcker, die Molaren weisen zwei Paar Höcker auf, nur der hinterste drei. Flusspferde können ihre Kiefer bis zu ca. 150° aufklappen.

 

Lebensraum der Flusspferde sind Gebiete mit tieferen Seen und langsam fließenden Flüssen, die idealerweise mit Schilfgürteln umgeben sind. Zum Weiden benötigt es Grasgebiete in der näheren Umgebung der Gewässer. In Regenwäldern kommt es außer bei großen Flüssen in der Regel nicht vor.

 

Flusspferde verbringen praktisch den ganzen Tag schlafend oder ruhend, dazu halten sie sich im Wasser oder in Gewässernähe auf. Dabei tauchen sie oft bis auf die Augen, Ohren und Nasenlöcher unter. Obwohl Flusspferde gut an ein Leben im Wasser angepasst sind, sind sie schlechte Schwimmer. Meistens laufen sie auf dem Grund eines Gewässers entlang oder lassen sich vom Wasser tragen; ihre Fortbewegungsart wird manchmal als „Schwimmlaufen“ umschrieben. Wenn sie untertauchen, können sie ihre schlitzförmigen Nasenlöcher und Ohren verschließen. Tauchgänge sind in der Regel nicht länger als drei bis fünf Minuten, sie können aber länger unter Wasser bleiben, möglicherweise bis zu 30 Minuten. Im Schlaf erfolgt das Auftauchen ebenso automatisch wie das Luftholen.

 

Vorwiegend in der Nacht verlassen sie das schützende Wasser, um sich auf Nahrungssuche zu begeben. Dabei können sie sich mehrere Kilometer vom Wasser entfernen, um Grasflächen zu erreichen. Um zu ihren Weideflächen zu gelangen, bilden die Gruppen regelrechte Trampelpfade („Hippo Trails“). Trotz ihres behäbigen Äußeren können Flusspferde im Bedarfsfall schnell laufen, Schätzungen belaufen sich auf bis zu 50 Kilometer pro Stunde. Diese Geschwindigkeit halten sie aber nur wenige hundert Meter durch.

 

An Land muss ihre Haut feucht bleiben, sie wird rissig, wenn sie zu lange der Luft ausgesetzt ist. Spezielle Hautdrüsen sondern eine Flüssigkeit ab, die die Tiere vor der Austrocknung schützt. Diese zunächst farblose Flüssigkeit verfärbt sich innerhalb von ein paar Minuten rötlich und später bräunlich. Bestandteil dieser Flüssigkeit sind zwei saure Pigmente, hipposudoric acid und norhipposudoric acid genannt. Diese wirken sowohl als Sonnenschutz, indem sie UV-Strahlen absorbieren, als auch antibiotisch gegen verschiedene Krankheitserreger. Das rötliche Schimmern hat früher zu der Vermutung geführt, Flusspferde würden Blut schwitzen.

 

(Wikipedia)

Rabbits are small mammals in the family Leporidae of the order Lagomorpha (along with the hare and the pika). Oryctolagus cuniculus includes the European rabbit species and its descendants, the world's 305 breeds[1] of domestic rabbit. Sylvilagus includes 13 wild rabbit species, among them the seven types of cottontail. The European rabbit, which has been introduced on every continent except Antarctica, is familiar throughout the world as a wild prey animal and as a domesticated form of livestock and pet. With its widespread effect on ecologies and cultures, the rabbit (or bunny) is, in many areas of the world, a part of daily life—as food, clothing, a companion, and as a source of artistic inspiration.

 

Although once considered rodents, lagomorphs like rabbits have been placed in their own, separate family because of a number of traits their rodent cousins lack, like two extra incisors.

---

Het stikt in het Diemerpark van de konijnen, dat hadden we allemaal al gemerkt. Maar waarom zitten ze daar eigenlijk en hoeveel zijn het er? Is er sprake van een plaag? We vragen het aan stadsecoloog Els Corporaal van stadsdeel Oost.

 

Waar komen de konijnen vandaan?

“Ze zaten al vóór de sanering in het park. Tijdens de werkzaamheden hebben ze zich teruggetrokken op de Diemerzeedijk, die niet werd gesaneerd.”

 

Hoeveel zijn het er?

“Dat is moeilijk te zeggen, want ze planten zich enorm snel voort. In een jaar tijd kunnen er honderden bij komen. Maar bij een uitbraak van de konijnenziekte myxomatose vallen ze bij bosjes om.”

 

Wat is dat voor ziekte?

“Het gaat om een virus dat wordt overgedragen door muggen en vlooien, maar ook van konijn op konijn. Als die ziekte oprukt, legt 90 procent het loodje. Alleen de sterkste overleven. De konijnen krijgen gezwellen rond hun ogen en kunnen niet meer goed zien, waardoor ze al snel een prooi vormen voor roofdieren. Als ze blind zijn geworden komt het voor dat ze gewoon ergens dood neervallen, maar over het algemeen zoeken ze een bosje op om rustig te sterven. Gelukkig herstellen de populaties zich snel, konijnen reproduceren zich als een razende.”

 

Wanneer was de laatste uitbraak?

“Dat was vorig jaar, in de herfst. Vaak gebeurt het in het najaar of in het voorjaar. Sinds ik hier als stadsecoloog werk, heb ik al een paar uitbraken meegemaakt. Niet alleen in het Diemerpark, maar zo’n virus spreidt zich als een olievlek uit over de hele stad.”

 

Zie je ze dan ook echt minder?

“Eventjes ja, maar vanaf het voorjaar groeit de populatie weer snel aan: konijnen kunnen achter elkaar nieuwe jongen werpen. En die jongen werpen vervolgens zelf binnen een paar weken weer jongen. Er is dan bijna geen limiet aan. Natuurlijke vijanden heeft het konijn wel in het Diemerpark: roofvogels, de wezel, de hermelijn en de vos. Maar het gaat om zoveel konijnen, daar valt niet tegenop te jagen.”

 

Wat voor soorten zien we eigenlijk? Is het één soort of zijn het er meerdere? Soms zit er bijvoorbeeld een zwarte tussen.

“Bij die zwarte kun je er eigenlijk vanuit gaan dat het een gefokt exemplaar is dat door mensen het huis uitgezet is. Omdat de kinderen er niet meer naar omkeken, of omdat het verzorgen toch te veel moeite bleek. Dat gebeurt best veel bij konijnen. Net als bijvoorbeeld bij slangen en schildpadden trouwens. Het mag niet, maar het gebeurt wel. Oorspronkelijk zijn wilde konijnen lichtbruin of grijs, maar tegenwoordig is het een mengelmoes van allerlei soorten geworden.

De gefokte soorten doen het redelijk in het park, al zou het wel kunnen dat ze minder lang leven doordat ze bevattelijker zijn voor ziektes, of minder snel schuilen bij gevaar. Dan worden ze er eerder uitgepikt door roofdieren.”

 

Hoe komt het dat de konijnen in het Diemerpark zo weinig schuw zijn?

“Er komen zoveel mensen in het park dat de konijntjes eraan gewend zijn geraakt. In rustiger gebieden zoals de duinen rennen konijnen veel eerder weg. Van nature zijn konijnen bang voor mensen, maar in het Diemerpark weten ze inmiddels dat er niet zoveel te vrezen valt. Behalve af en toe van honden: die gaan wel eens achter een konijn aan. Het grappige is dat juist op het hondenuitrengebied ontzettend veel konijnen zitten. Daar omheen liggen stapels takken waarin ze zich kunnen verschuilen. Vaak blijven ze er dichtbij zitten: twee sprongen en ze zijn weer veilig.”

 

Is er sprake van een konijnenplaag?

“Ikzelf zie ze niet als een plaag. Het is nog niet zo dat je over de konijnen struikelt. Wij zien iets pas als een plaag als je er echt problemen mee krijgt. Als je niet meer in het Diemerpark kunt rondlopen zonder door een konijn in je benen gebeten te worden, bij wijze van spreken, maar dat doen konijnen niet. Of als het een gevaar is voor de volksgezondheid. Konijnen die holen graven onder een spoorlijn, daar heb je last van. Of op een begraafplaats is het ook pijnlijk als ze holletjes maken in de graven. Dan zou je maatregelen moeten nemen.”

 

Graven ze in het Diemerpark niet tot de ingepakte vuilstort die onder de leeflaag ligt?

“Nee, de holen gaan niet heel erg diep. Je hebt eerst de leeflaag en dan een afdeklaag van trisoplast. Die is zo stevig, daar kom je niet doorheen.”

 

Lees meer over de natuur in Amsterdam Oost en omstreken

en over de konijnenziekte

Door Kirsten Dorrestijn16082012

Rubriek NatuurTags Diemerpark, stadsdeel Oost, natuur Amsterdam Oost, konijnen diemerpark, els corporaal, myxomatose konijnen

 

Elephants on the run

 

Elefanten auf der Flucht

 

Kruger National Park is one of the largest game reserves in Africa. It covers an area of 19,485 km2 (7,523 sq mi) in the provinces of Limpopo and Mpumalanga in northeastern South Africa, and extends 360 km (220 mi) from north to south and 65 km (40 mi) from east to west. The administrative headquarters are in Skukuza. Areas of the park were first protected by the government of the South African Republic in 1898, and it became South Africa's first national park in 1926.

 

To the west and south of the Kruger National Park are the two South African provinces of Limpopo and Mpumalanga. In the north is Zimbabwe, and to the east is Mozambique. It is now part of the Great Limpopo Transfrontier Park, a peace park that links Kruger National Park with the Gonarezhou National Park in Zimbabwe, and with the Limpopo National Park in Mozambique.

 

The park is part of the Kruger to Canyons Biosphere an area designated by the United Nations Educational, Scientific and Cultural Organization (UNESCO) as an International Man and Biosphere Reserve (the "Biosphere").

 

The park has nine main gates allowing entrance to the different camps.

 

(Wikipedia)

 

Name

 

African Elephant or African Bush Elephant [Loxodonta africana]

 

Introduction

 

The Elephant is the world's largest land mammal, and weighs up to 7 tonnes and reaches heights of 3.3 m at the shoulder. Elephants can live to a potential age of 70 years. The massive tusks of older bulls can weigh up to 50 or 60 kilograms, but tusks weighing up to 90 kilograms have been recorded.

 

Appearance

 

What is the trunk and what is it used for?

The Elephant's trunk is a modified nose which is very sensitive and can even detect water under ground. There are as many as 50 000 muscles in an Elephant trunk. The sensitive finger-like appendages at the tip of the trunk enables them to pick the smallest twig or flower, pull the toughest reed of grass or even pick out a thorn from their feet.

 

Do elephants have knees or elbows?

 

The joints that are perceived as 'knees', are in fact wrists. This is a common misunderstanding due to the belief that a leg joint that bends between the foot and the body must be a knee. The main difference between us and the elephants is that our foot bones and hand bones are separate, whereas those of the elephant are one in the same, and have evolved to suit this four-legged mammal.

 

Why do elephants have tusks?

 

The tusks are used for obtaining food, fighting (amongst males) and for self defence. They are actually their upper incisors, and grow continuously until they die at around 60 years old. Although their skin is up to 3cm (1 inch) thick, it is quite sensitive.

 

Diet

 

Elephants are voracious feeders which in a day consume up to 272 kg (600 pounds) of grass, tender shoots and bark from trees. An adult Elephant can drink up to 200 litres of water in a single session. A single Elephant deposits up to 150kg (330 pounds) of dung every day - about one dollop every 15 minutes!

 

Breeding

 

African Elephant are not seasonal breeders. Generally they produce one calf every 3 to 4 years. The gestation period is about 22 months. At birth calves weigh about 100 kg (220 pounds) and are fully weaned between 18-24 months. An orphaned calf will usually be adopted by one of the family's lactating females or suckled by various females. Elephants are very attentive mothers, and because most Elephant behavior has to be learned, they keep their offspring with them for many years. Tusks erupt at 16 months but do not show externally until 30 months. Once weaned, usually at age 4 or 5, the calf still remains in the maternal group. Females mature at about 11 years and stay in the group, while the males, which mature between 12 and 15, are usually expelled from the maternal herd. Even though these young males are sexually mature, they do not breed until they are in their mid, or late 20s or even older and have moved up in the social hierarchy.

 

Behaviour

 

Mature males form bachelor groups and become solitary bulls. Elephant form strong family units of cows, calves and young offspring. Such herds are always led by an old female. Apart from drinking large quantities of water they also love wading or swimming in it. Elephants clearly relish mud baths.

It was once thought that family groups were led by old bull elephants, but these males are most often solitary. The female family groups are often visited by mature males checking for females in oestrus. Several interrelated family groups may inhabit an area and know each other well.

 

How do you tell an elephant's mock charge from a serious one?

 

It is imperative to keep in mind that Elephant are extremely intelligent, and each individual has a distinct character. Although there will be exceptions to the rules, the common signs of a mock charge are bush-bashing, dust-throwing, trumpeting and other vocalizations, open ears and an intimidating presence, can be considered a mock-display. Aggressive or startled elephants usually make sudden headshakes and flap their large ears against their head. Serious charges usually occur after all attempts to intimidate have failed, and the Elephant feels threatened. The ears are pinned back and head and trunk are lowered. Ultimately, the key lies in the intelligence of the animal and how they will react to the 'target' and unfamiliar actions, and a conscious decision is made.

 

Why do elephants rhythmically flap their ears?

 

Contrary to common belief, it is not an expression of anger. Being an animal of such a large size, with no sweat glands and a dark body colour, elephants flap their ears to cool the body and rid themselves of irritating insects.

 

Where are they found?

 

Once ranging across most of Africa the Elephant population has declined dramatically across the continent. In South Africa the Addo Elephant and Kruger National Park protect large herds. Due to rigorous conservation measures the Elephant population in South Africa has grown from a estimated 120 in 1920 in 4 locations, to 10 000 at 40 locations to date.

 

Notes

 

The African Elephant has recently been classified into two separate species, the more common African Bush Elephant [Loxodonta Africana] and the smaller African Forest Elephant [Loxodonta cyclotis] of the rainforest of Central Africa.

 

(krugerpark.co.za)

 

Der Kruger-Nationalpark (deutsch häufig falsch Krüger-Nationalpark) ist das größte Wildschutzgebiet Südafrikas. Er liegt im Nordosten des Landes in der Landschaft des Lowveld auf dem Gebiet der Provinz Limpopo sowie des östlichen Abschnitts von Mpumalanga. Seine Fläche erstreckt sich vom Crocodile-River im Süden bis zum Limpopo, dem Grenzfluss zu Simbabwe, im Norden. Die Nord-Süd-Ausdehnung beträgt etwa 350 km, in Ost-West-Richtung ist der Park durchschnittlich 54 km breit und umfasst eine Fläche von rund 20.000 Quadratkilometern. Damit gehört er zu den größten Nationalparks in Afrika.

 

Das Schutzgebiet wurde am 26. März 1898 unter dem Präsidenten Paul Kruger als Sabie Game Reserve zum Schutz der Wildnis gegründet. 1926 erhielt das Gebiet den Status Nationalpark und wurde in seinen heutigen Namen umbenannt. Im Park leben 147 Säugetierarten inklusive der „Big Five“, außerdem etwa 507 Vogelarten und 114 Reptilienarten, 49 Fischarten und 34 Amphibienarten.

 

(Wikipedia)

 

Der Afrikanische Elefant (Loxodonta africana), auch Afrikanischer Steppenelefant oder Afrikanischer Buschelefant, ist eine Art aus der Familie der Elefanten. Er ist das größte gegenwärtig lebende Landsäugetier und gleichzeitig das größte rezente landbewohnende Tier der Erde. Herausragende Kennzeichen sind neben den Stoßzähnen und dem markanten Rüssel die großen Ohren und die säulenförmigen Beine. In zahlreichen morphologischen und anatomischen Merkmalen unterscheidet sich der Afrikanische Elefant von seinen etwas kleineren Verwandten, dem Waldelefanten und dem Asiatischen Elefanten. Das Verbreitungsgebiet umfasst heute große Teile von Afrika südlich der Sahara. Die Tiere haben sich dort an zahlreiche unterschiedliche Lebensräume angepasst, die von geschlossenen Wäldern über offene Savannenlandschaften bis hin zu Sumpfgebieten und wüstenartigen Regionen reichen. Insgesamt ist das Vorkommen aber stark fragmentiert.

 

Die Lebensweise des Afrikanischen Elefanten ist durch intensive Studien gut erforscht. Sie wird durch einen stark sozialen Charakter geprägt. Weibliche Tiere und ihr Nachwuchs leben in Familienverbänden (Herden). Diese formieren sich wiederum zu einem enger verwandten Clan. Die einzelnen Herden treffen sich zu bestimmten Gelegenheiten und trennen sich danach wieder. Die männlichen Tiere bilden Junggesellengruppen. Die verschiedenen Verbände nutzen Aktionsräume, in denen sie teils im Jahreszyklus herumwandern. Für die Kommunikation untereinander nutzen die Tiere verschiedene Töne im niedrigen Frequenzbereich. Anhand der Lautgebung, aber auch durch bestimmte chemische Signale können sich die einzelnen Individuen untereinander erkennen. Darüber hinaus besteht ein umfangreiches Repertoire an Gesten. Hervorzuheben sind auch die kognitiven Fähigkeiten des Afrikanischen Elefanten.

 

Die Nahrung besteht sowohl aus weicher wie auch harter Pflanzenkost. Die genaue Zusammensetzung variiert dabei regional und jahreszeitlich. Generell verbringt der Afrikanische Elefant einen großen Teil seiner Tagesaktivitäten mit der Nahrungsaufnahme. Die Fortpflanzung erfolgt ganzjährig, regional gibt es Tendenzen zu einer stärkeren Saisonalisierung. Bullen kommen einmal jährlich in die Musth, während deren sie auf Wanderung zur Suche nach fortpflanzungswilligen Kühen gehen. Während der Musth ist die Aggressivität gesteigert, es finden dann auch Rivalenkämpfe statt. Der Sexualzyklus der Kühe dauert vergleichsweise lange und weist einen für Säugetiere untypischen Verlauf auf. Nach erfolgter Geburt setzt er in der Regel mehrere Jahre aus. Zumeist wird nach fast zweijähriger Tragzeit ein Jungtier geboren, das in der mütterlichen Herde aufwächst. Junge weibliche Tiere verbleiben später in der Herde, die jungen männlichen verlassen diese.

 

Die wissenschaftliche Erstbeschreibung des Afrikanischen Elefanten erfolgte im Jahr 1797 mit einer formalen artlichen Trennung des Afrikanischen vom Asiatischen Elefanten. Der heute gebräuchliche Gattungsname Loxodonta wurde offiziell erst dreißig Jahre später eingeführt. Die Bezeichnung bezieht sich auf markante Zahnunterschiede zwischen den asiatischen und den afrikanischen Elefanten. Im Verlauf des 20. Jahrhunderts wurden mehrere Unterarten unterschieden, darunter auch der Waldelefant des zentralen Afrikas. Letzterer gilt heute genetischen Untersuchungen zufolge als eigenständige Art, die weiteren Unterarten sind nicht anerkannt. Stammesgeschichtlich lässt sich der Afrikanische Elefant erstmals im beginnenden Mittleren Pleistozän belegen. Der Gesamtbestand gilt als gefährdet. Ursachen hierfür sind hauptsächlich die Jagd nach Elfenbein und Lebensraumverlust durch die zunehmend wachsende menschliche Bevölkerung. Der Afrikanische Elefant zählt zu den sogenannten „Big Five“ von Großwildjagd und Safari.

 

(Wikipedia)

i luv diz dude i think hez hot hez fine && sexii && i luv hez songz here they r

 

herez him rap battlein

 

www.youtube.com/watch?v=tLzgC0NL11k

 

www.youtube.com/watch?v=Szou7F6mYQ0&feature=related

  

"Don't Fuck With Us"

 

[John Cena]

We keep it hoppin like the cars with the shocks

We spittin heat on your block

We new to the game, but runnin the spot

Numbin your knot, with basslines that'll make ya neck break

This rook'll take your queen and put ya king in checkmate

Open your mind without makin ya meditate

We real champs; y'all just featherweight

Time to get it straight, I push your wig back

Crew loaded up with extra bread like a Big Mac

Beefin with us? We're leavin you face down

Stompin bitch rappers like I'm straight outta A-Town

Runnin the playground like it was a track meet

Shoes on the whip that be bigger than Shaq's feet

We into big things, bank account's overgrown

All types of cheese - swiss, cheddar, provolone

Guaranteed to burn wax like candles

Track hittin hard to the head like shots of Jack Daniels

 

[Chorus 2X: John Cena]

Y'all, bitch, crews, don't wanna fuck with us

Y'all bound, to, lose, another one bites the dust

 

[Tha Trademarc]

It's Trademarc the truth, laid back, aloof

I'm God, as if you needed some proof

You ain't hard I can see it on you, I need a roof

Fuck a droptop, crop if I'm creepin on you

Click-clack nickelback knickknacks if you got heaters on you

Spittin back live rounders, with five pounders

If we meetin on two, I put a beatin on you

Your sound's tired buddy, that's why I'm sleepin on you

We lean back in the ride, with cream stackin the rawhide

The sound of God slide with a raw vibe

Straight military camel clothes ash brown boots

So sick, I've been handlin flows, since enamel was gold tooth

And branded by low

You cold fuck like eskimo hoes at 7 below

You slow, you be the last to think

My hands seen more fuckin dirt than bathroom sinks

 

[Chorus]

 

[John Cena]

I got punks, dumps and switches, dump chumpses bitches

We feed you to the sharks, you can sleep with the fishes

Clean you like dishes but I ain't no busboy

You ain't family, you ain't earnin my trust boy

Seen too many bitches that'll double cross ya

We bring more drama than the Laker roster

Get the click pissed, ain't nobody can save ya

Throw heat without lookin like Fernando Valenzuela

 

[Tha Trademarc]

Marc Predka's the name, the rest of you lame

I'm ego drivin, seen with different women, every size and frame

I refine my game by fuckin famous bitches

But it's all the same, it's just ex to the next

for sex or brain, misses or Mrs.

Married or not, my game don't stop

It's cars bars bonds and stocks you ain't see my flow

Y'all are small-time suckers like a knee-high hoe

 

[Chorus]

  

"Bad, Bad Man"

 

[John Cena]

Aww, you done did it now

Chaos you shoulda put this one in the vault man!

They not ready - they don't know what's comin man!

Oh we gonna drop this on 'em right here

Y'all ain't ready for this, Y'ALL AIN'T READY FOR NONE OF THIS!

 

Your boy's a bad man, and we invadin the streets

Make unclever rappers scurred, they be droppin the heat

Shocked the world, now I'm standin alone

I flip fools like them clamshell cellular phones

You can't help but nod your head to the track

Fuck the watered down rap, we be takin it back

Give it to me straight - ain't no chasin it

Check yourself in the mirror - ain't no facin it

Cause you, playin the role and you plannin to fold

This the masterplan, we got the planet on hold

We all over the streets like your favorite sneaker

Breakin up your sound like a drive-through speaker

Everything that I be spittin is strong

After I rock, fast forward through the rest of the song

We the monkeywrench, that's gonna ruin your plan

And don't fuck with John Cena - I'm a BAD, BAD MAN

 

[Chorus 2X: Bumpy Knuckles]

With the mic in my hands I'm a bad man

Even in a fight with the hands I'm a bad man

Livin in the streets all my life I'm a bad man

I'm a bad man, I'm a bad man

 

[Tha Trademarc]

We devils - rockin ambient levels

We set loose among hot tunes to instrumentals

And cats got one-liners, I drop several

And I think it's funny you choose, losin progress

or runnin in place; we makin moves, and y'all settle

I rip rappers and take responsibility

for makin future hall-of-famers look third rate

Y'all are lost for words like conversation on your worst first date

and ride beats, creep through side streets

Looseleaf notepads that's where rhymes leak

Punchlines - man, don't even beg

I got knee-slappin tracks, y'all brusin your leg

You a rhyme writer - funny man, that's a joke

You ain't worthy of bein my secretary man that's a quote

I flood tracks like cracks in boats

And pussy rappers choked up with they own lines in they throat

 

[Chorus]

 

[Bumpy Knuckles a.k.a. Freddie Foxxx]

TURN UP THE MICROPHONE and feed me I'm a beast

MC's and they beats is what I eat, 16 I'll leave you in the street

My rhymes are sicker than gangrene in both feet

It's spreadin up the leg, and headed for the head

Your rhymes are whack your style is proof that the brain corrosion

is fuckin with your chosen flows, I'm nice with mics

My hands'll break your nose like Mikey Tyson

Fightin in his prime, one rhyme

And I shake up the room one time, BOOM! To the jaw

Your face is a coat type raw

And the blood and snot they mix, jelly on the floor

My love is cop them bricks, belly on the floor

I rob you, you soft and you really ain't a problem

I solve you, 357 long nose revolve you

Acid in your face, bad look, dissolve you

I'm a bad, bad man

 

Yeah, check it out

It's Bumpy Knuckles baby

And I want you to say hello to the BAD, BAD, MAN - C'MON!

 

[Chorus]

  

"Basic Thugonomics"

  

"So... you think you're untouchable?"

 

[Chorus: John Cena - scratched by DJ Chaos]

Word life! This is basic thugonomics

This is ba-basic thugomoics

Word life! {*scratching*}

"I'm untouchable, but I'm forcin you to feel me" - Esoteric

Word life! This is bas-{*scratch*}

Basic thugo-{*scratch*}-thugo-{*scratch*}-thugonomics

Word life! {*scratching*}

"I'm untouchable, but I'm forcin you to feel me" - Esoteric

 

[Verse One: John Cena]

Whether fightin, or spittin, my discipline is unforgiven

Got you backin up, in a defensive position

An ass-kickin anthem, heavyweight or bantam

Holdin camps for ransom, the microphone phantom

Teams hit the floor, this the new fight joint

Like a broken needle kid, you missin the point!

We dominate your conference with offense that's no nonsense

My theme song hits, get your reinforcements!

We strike quick with hard kicks, duckin ice picks

Bare-knuckle men through fight pits, beat you lifeless

Never survive this! Get forget like Alzheimer's

Two-face rappers, walk away with four shiners

The raw rhymer, turnin legends to old-timers

My incisor's like a viper, bitin through your one-liners!

New Deadman Inc. - and we about to make you famous

Takin over Earth and still kickin in Uranus!

 

[Chorus]

 

[Verse Two: Esoteric]

You ain't advanced enough to process potential phonetical concepts

The objects are foreign, like blot tests

Sponsored sex, a complex, regardless of your finesse

or your fitness, it's the condition of business

Your lame vision of a underground, physical image

You're underneath to undermine your whole, typical image

With the precision of percentages, and the collision of sedatives

Poetry, beats, and mics - we untouchable

like righteous sluts with no crevices

Streets unite, we rock right over dumber beats

Yo' cats couldn't come this hot if they {jerked} off in the summer heat

Forget two takes, kill y'all birds the first time

Yo' best {shit} ain't, worthy of my filler or worst rhymes

I'm better than nice, check the veteran stripes

Leave you beside yourself with fear, I kill you, and bury you twice

Despite the cover of night, trackin' your flight

Like guerilla warfare, where the grass is dense

Approachin me is a quick way to get referred to in the past tense

Dead that! When the light to mic is on

The crowd is dead like the intermission when you on the Titantron

 

[Chorus]

  

"Make It Loud"

 

[John Cena at a live show]

It's the joint baby, GOTTA MAKE IT LOUD [crowd cheers]

SO LET ME HEAR SOME NOISE FROM THE CROWD [more cheering]

 

[Tazz] That's noise!

 

[Chorus 2X: John Cena]

It's the joint baby, gotta make it loud

Get the point yo you gotat make it loud

Everybody in the club make it loud

SO LET ME HEAR SOME NOISE FROM THE CROWD

 

[John Cena]

Yeah, yeah

We came to kick the door down, it's time to hit the floor now

Yo... we got some shit in store now

So; clap your hands while we let the sax blow

Not quite Krispy Kreme, but we came to stack dough

We ain't maxed yo, we just try and get this money right

Bills made of Spandex, I still keep my money tight

Never stoppin, all I see is the money like

the kid on the mic is too +Raw+ for your Monday night

If you got in free, or your fuckin cover's paid

Bounce to this motherfucker like you was some Rubbermaid

This ain't that Cristal sippin type shit

It's that bottle breakin, startin riot type shit

So jump up and down 'til ya break the floor

Yo we keep it underground like a basement tour

East coast reppin, stretchin out to L.A.

Not double oh seven but we +Die Another Day+, what

 

[Chorus]

 

[Tha Trademarc]

I tear up any track, front to back

Like Roy Jones takin on fifty year-old cats

makin comebacks, where you at, cats spit soft shit

like whispers and gloves, I'm not hearin that

It's all love maybe if you wanna rub baby

Anything but that, step back lady

Trademarc, John Cena, clubbin it up

We got Chaos on the one and two, cuttin it up

I'm all about laid back, don't jock, I hate that

I see through haters games, don't mistake that

I still got love if you buyin our shit

If you claim you hatin us, but you ridin our dicks

Everybody hear the name, Marc Predka

It's gonna ring like an echo for years, I never left ya

All y'all raise your glass to this shit

Cause Trademarc's the head of the class of misfits

 

[Chorus]

 

[John Cena]

We steal your top spot, and you not gettin your number back

Chop down competition like I was a lumberjack

Clear out the club floor, we keep 'em comin back

Tough to bring down like an overweight runningback

Yeah - and we blaze 'em baby

Trademarc, John Cena, we amazin baby

Yo we tear up any crew, leave a motherfucker worn

Y'all are just soft like some Cinemax porn

 

[Tha Trademarc]

I move a crowd like a bomb scare

Grab the mic when we hittin it right, if you want fear

Some say Trademarc, he ain't all there

We old school like when Sonny, was on Cher

Take it back like a Richard Pyror 8-track

And grab a chunk of your change like a state tax

Man please, we want platinum plaques

I want cream, green, cheddar cheese, to grab in stacks

 

[Chorus]

 

[scratching Trademarc to fade]

"Chaos on the one and two, cuttin it up"

"That's that shit!"

 

[crowd chanting] "Ce-na, Ce-na, Ce-na" [at the end]

 

"my time is now"

 

[Chorus: John Cena]

Your time is up, my time is now

You can't see me, my time is now

It's the franchise, boy I'm shinin now

You can't see me, my time is now!

 

[John Cena]

In case you forgot or fell off I'm still hot - knock your shell off

My money stack fat plus I can't turn the swell off

The franchise, doin big bid'ness, I live this

It's automatic I win this - oh you hear those horns, you finished

A soldier, and I stay under you fightin

Plus I'm stormin on you chumps like I'm thunder and lightning

Ain't no way you breakin me kid, I'm harder than nails

Plus I keep it on lock, like I'm part of the jail

I'm slaughterin stale, competition, I got the whole block wishin

they could run with my division but they gone fishin -

- with no bait, kid your boy hold weight

I got my soul straight, I brush your mouth like Colgate

In any weather I'm never better your boy's so hot

you'll never catch me in the next man's sweater

If they hate, let 'em hate, I drop ya whole clan

Lay yo' ass DOWN for the three second TAN

 

[Chorus]

 

[Tha Trademarc]

Yeah, uhh

It's gon' be what it's gon' be

Five pounds of courage buddy, bass tint pants with a gold T

Uhh - it's a war dance and victory step

A raw stance is a gift, when you insist it's my rep

John Cena, Trademarc, y'all are so-so

And talk about the bread you make but don't know the recipe for dough though

Aimin guns in all your photos, that's a no-no

When this pop, you'll liplock, your big talk's a blatant no-show

See what happens when the ice age melt

You see monetary status is not what matters, but it helps

I rock a timepiece by Benny if any

The same reason y'all could love me is the same reason y'all condemn me

A man's measured by the way that he thinks

Not clothing lines, ice links, leather and minks

I spent 20 plus years seekin knowledge of self

So for now Marc Predka's livin live for wealth

 

[Chorus - repeat 2X]

  

(theirz more but these i luv da mozt)

 

(here da videoz)

 

(make it loud) www.youtube.com/watch?v=V02Sz51ySV0

 

(basic thugonomics) www.youtube.com/watch?v=n8Guq9KRQgA

 

(dont fuck with us) www.youtube.com/watch?v=oAs4-nEH0rE

 

(bad bad man) www.youtube.com/watch?v=_lKuoF3qz-w&feature=related

 

(my time is now) www.youtube.com/watch?v=_JAa3NvP6f4

  

(&& hez a wwe wrestler 2 =])

BMW's Hommage to Lagomorphic Cross Dressing

 

Left:

 

BMW’s faithful application of Bugs Bunny’s delicate feminine eye socket.

 

BMW logo -to-grille proportions the same as bunny nose-to-incisors.

 

BMW grille honors that famous gap and buck teeth.

 

BMW respectfully evokes the color of Bug's to-die-for

yellow halter.

  

Right:

Lower airfoil pays loving deference to Bug's sensuous, pouting, full lower lip

 

Oh, those bedroom eyes!

 

BMW ‘cheeky cheeks’ allude to Bug’s devilish smile.

  

"Goodbye cruel world...I grow tired of your monkeyshines"

 

I've a tooth-surgery in 3 hours to uproot/remove my Impacted 3-rd Molar. It has generally been a 'love-hate' acquaintance between the two of us. In the last couple of months, the relationship has largely fallen out, with gaping holes being unearthed. I've ceased to care about my 3-rd molar. My attention these days is largely devoted to the jazzy incisors. Those hussies!

 

I'm really really bad at making puns :(

 

...and that tooth looks ridiculously, fake, doesn't it?

Quality prints, greeting cards and many products can be purchased at >> kaye-menner.pixels.com/featured/beach-dog-more-play-by-ka...

 

This wonderful and friendly dog, whose name is Rambo, just loves the beach and swimming in the surf. Also, (like most dogs), he loves to play ball with his favorite orange ball.

THE FINE ART AMERICA LOGO WILL NOT APPEAR ON PURCHASED PRINTS OR PRODUCTS.

 

[From Wikipedia]

The Staffordshire Bull Terrier is a medium-sized, stocky, and very muscular dog, with a similar appearance to the much larger American Staffordshire terrier and American pit bull terrier. It has a broad head (male considerably more so than female), defined occipital muscles, a relatively short fore-face, dark round eyes and a wide mouth with a clean scissor-like bite (the top incisors slightly overlap the bottom incisors). The ears are small. The cheek muscles are very pronounced. The lips show no looseness. From above, the head loosely resembles a triangle. The head tapers down to a strong well-muscled neck and shoulders placed on squarely spaced forelimbs. They are tucked up in their loins and the last 1-2 ribs of the rib-cage are usually visible. The tail resembles an old fashioned pump handle. The hind quarters are well-muscled and are what give the Stafford drive when baiting. They are coloured brindle, black, red, fawn, blue, white, or any blending of these colors with white. White with any other colour broken up over the body is known as pied. Liver-colored, black and tan dogs can occur but are rare and it is advised not to breed from either as well as those with light eyes. The exception to the light eye rule are Blue staffies; all others should have dark brown eyes even if fawn coat. The coat is smooth and clings tightly to the body giving the dog a streamlined appearance.

 

The dogs stand 36 to 41 cm (14 to 16 in) at the withers and weigh 13 to 17 kg (29 to 37 lb) for males; females are 11 to 15.4 kg (24.3 to 34.0 lb).

 

Island of Madagascar

Off The East Coast Of Africa

Palmarium Reserve

 

Click On Image To Enlarge.

 

Red-ruffed lemur in a tree in the rainforest with sunlight on it’s face.

 

Wikipedia- The red ruffed lemur (Varecia rubra) is one of two species in the genus Varecia, the ruffed lemurs; the other is the black-and-white ruffed lemur (Varecia variegata). Like all lemurs, it is native to Madagascar and occurs only in the rainforests of Masoala, in the northeast of the island.

 

It is one of the largest primates of Madagascar with a body length of 53 cm, a tail length of 60 cm and a weight of 3.3–3.6 kg. Its soft, thick fur is red and black in colour and sports a buff or cream colored spot at the nape, but a few are known to have a white or pink patch on the back of the limbs or digits and a ring on the base of the tail in a similar color.

 

The red ruffed lemur is a very clean animal and spends a lot of time grooming itself and in social grooming. The lower incisors (front teeth) and the claw on the second toe of the hind foot are specially adapted for this behavior. The lower incisors grow forward in line with each other and are slightly spaced. This creates a toothcomb which can be used to groom its long, soft fur. The claw is also used for grooming.

 

The red ruffed lemur lives 15–20 years in the wild. In captivity, 25 years is not uncommon, and one lived to be about 33 years old. It is a diurnal animal, and most active in the morning and evening.

 

The red ruffed lemur is mainly a fruit-eater, though it is known to eat leaves and shoots. They especially like figs.

  

Fish, any of approximately 34,000 species of vertebrate animals (phylum Chordata) found in the fresh and salt waters of the world. Living species range from the primitive jawless lampreys and hagfishes through the cartilaginous sharks, skates, and rays to the abundant and diverse bony fishes. Most fish species are cold-blooded; however, one species, the opah (Lampris guttatus), is warm-blooded.

 

The term fish is applied to a variety of vertebrates of several evolutionary lines. It describes a life-form rather than a taxonomic group. As members of the phylum Chordata, fish share certain features with other vertebrates. These features are gill slits at some point in the life cycle, a notochord, or skeletal supporting rod, a dorsal hollow nerve cord, and a tail. Living fishes represent some five classes, which are as distinct from one another as are the four classes of familiar air-breathing animals—amphibians, reptiles, birds, and mammals. For example, the jawless fishes (Agnatha) have gills in pouches and lack limb girdles. Extant agnathans are the lampreys and the hagfishes. As the name implies, the skeletons of fishes of the class Chondrichthyes (from chondr, “cartilage,” and ichthyes, “fish”) are made entirely of cartilage. Modern fish of this class lack a swim bladder, and their scales and teeth are made up of the same placoid material. Sharks, skates, and rays are examples of cartilaginous fishes. The bony fishes are by far the largest class. Examples range from the tiny seahorse to the 450-kg (1,000-pound) blue marlin, from the flattened soles and flounders to the boxy puffers and ocean sunfishes. Unlike the scales of the cartilaginous fishes, those of bony fishes, when present, grow throughout life and are made up of thin overlapping plates of bone. Bony fishes also have an operculum that covers the gill slits.

 

The study of fishes, the science of ichthyology, is of broad importance. Fishes are of interest to humans for many reasons, the most important being their relationship with and dependence on the environment. A more obvious reason for interest in fishes is their role as a moderate but important part of the world’s food supply. This resource, once thought unlimited, is now realized to be finite and in delicate balance with the biological, chemical, and physical factors of the aquatic environment. Overfishing, pollution, and alteration of the environment are the chief enemies of proper fisheries management, both in fresh waters and in the ocean. (For a detailed discussion of the technology and economics of fisheries, see commercial fishing.) Another practical reason for studying fishes is their use in disease control. As predators on mosquito larvae, they help curb malaria and other mosquito-borne diseases.

 

Fishes are valuable laboratory animals in many aspects of medical and biological research. For example, the readiness of many fishes to acclimate to captivity has allowed biologists to study behaviour, physiology, and even ecology under relatively natural conditions. Fishes have been especially important in the study of animal behaviour, where research on fishes has provided a broad base for the understanding of the more flexible behaviour of the higher vertebrates. The zebra fish is used as a model in studies of gene expression.

 

There are aesthetic and recreational reasons for an interest in fishes. Millions of people keep live fishes in home aquariums for the simple pleasure of observing the beauty and behaviour of animals otherwise unfamiliar to them. Aquarium fishes provide a personal challenge to many aquarists, allowing them to test their ability to keep a small section of the natural environment in their homes. Sportfishing is another way of enjoying the natural environment, also indulged in by millions of people every year. Interest in aquarium fishes and sportfishing supports multimillion-dollar industries throughout the world.

 

Fishes have been in existence for more than 450 million years, during which time they have evolved repeatedly to fit into almost every conceivable type of aquatic habitat. In a sense, land vertebrates are simply highly modified fishes: when fishes colonized the land habitat, they became tetrapod (four-legged) land vertebrates. The popular conception of a fish as a slippery, streamlined aquatic animal that possesses fins and breathes by gills applies to many fishes, but far more fishes deviate from that conception than conform to it. For example, the body is elongate in many forms and greatly shortened in others; the body is flattened in some (principally in bottom-dwelling fishes) and laterally compressed in many others; the fins may be elaborately extended, forming intricate shapes, or they may be reduced or even lost; and the positions of the mouth, eyes, nostrils, and gill openings vary widely. Air breathers have appeared in several evolutionary lines.

 

Many fishes are cryptically coloured and shaped, closely matching their respective environments; others are among the most brilliantly coloured of all organisms, with a wide range of hues, often of striking intensity, on a single individual. The brilliance of pigments may be enhanced by the surface structure of the fish, so that it almost seems to glow. A number of unrelated fishes have actual light-producing organs. Many fishes are able to alter their coloration—some for the purpose of camouflage, others for the enhancement of behavioral signals.

 

Fishes range in adult length from less than 10 mm (0.4 inch) to more than 20 metres (60 feet) and in weight from about 1.5 grams (less than 0.06 ounce) to many thousands of kilograms. Some live in shallow thermal springs at temperatures slightly above 42 °C (100 °F), others in cold Arctic seas a few degrees below 0 °C (32 °F) or in cold deep waters more than 4,000 metres (13,100 feet) beneath the ocean surface. The structural and, especially, the physiological adaptations for life at such extremes are relatively poorly known and provide the scientifically curious with great incentive for study.

 

Almost all natural bodies of water bear fish life, the exceptions being very hot thermal ponds and extremely salt-alkaline lakes, such as the Dead Sea in Asia and the Great Salt Lake in North America. The present distribution of fishes is a result of the geological history and development of Earth as well as the ability of fishes to undergo evolutionary change and to adapt to the available habitats. Fishes may be seen to be distributed according to habitat and according to geographical area. Major habitat differences are marine and freshwater. For the most part, the fishes in a marine habitat differ from those in a freshwater habitat, even in adjacent areas, but some, such as the salmon, migrate from one to the other. The freshwater habitats may be seen to be of many kinds. Fishes found in mountain torrents, Arctic lakes, tropical lakes, temperate streams, and tropical rivers will all differ from each other, both in obvious gross structure and in physiological attributes. Even in closely adjacent habitats where, for example, a tropical mountain torrent enters a lowland stream, the fish fauna will differ. The marine habitats can be divided into deep ocean floors (benthic), mid-water oceanic (bathypelagic), surface oceanic (pelagic), rocky coast, sandy coast, muddy shores, bays, estuaries, and others. Also, for example, rocky coastal shores in tropical and temperate regions will have different fish faunas, even when such habitats occur along the same coastline.

 

Although much is known about the present geographical distribution of fishes, far less is known about how that distribution came about. Many parts of the fish fauna of the fresh waters of North America and Eurasia are related and undoubtedly have a common origin. The faunas of Africa and South America are related, extremely old, and probably an expression of the drifting apart of the two continents. The fauna of southern Asia is related to that of Central Asia, and some of it appears to have entered Africa. The extremely large shore-fish faunas of the Indian and tropical Pacific oceans comprise a related complex, but the tropical shore fauna of the Atlantic, although containing Indo-Pacific components, is relatively limited and probably younger. The Arctic and Antarctic marine faunas are quite different from each other. The shore fauna of the North Pacific is quite distinct, and that of the North Atlantic more limited and probably younger. Pelagic oceanic fishes, especially those in deep waters, are similar the world over, showing little geographical isolation in terms of family groups. The deep oceanic habitat is very much the same throughout the world, but species differences do exist, showing geographical areas determined by oceanic currents and water masses.

 

All aspects of the life of a fish are closely correlated with adaptation to the total environment, physical, chemical, and biological. In studies, all the interdependent aspects of fish, such as behaviour, locomotion, reproduction, and physical and physiological characteristics, must be taken into account.

 

Correlated with their adaptation to an extremely wide variety of habitats is the extremely wide variety of life cycles that fishes display. The great majority hatch from relatively small eggs a few days to several weeks or more after the eggs are scattered in the water. Newly hatched young are still partially undeveloped and are called larvae until body structures such as fins, skeleton, and some organs are fully formed. Larval life is often very short, usually less than a few weeks, but it can be very long, some lampreys continuing as larvae for at least five years. Young and larval fishes, before reaching sexual maturity, must grow considerably, and their small size and other factors often dictate that they live in a habitat different than that of the adults. For example, most tropical marine shore fishes have pelagic larvae. Larval food also is different, and larval fishes often live in shallow waters, where they may be less exposed to predators.

 

After a fish reaches adult size, the length of its life is subject to many factors, such as innate rates of aging, predation pressure, and the nature of the local climate. The longevity of a species in the protected environment of an aquarium may have nothing to do with how long members of that species live in the wild. Many small fishes live only one to three years at the most. In some species, however, individuals may live as long as 10 or 20 or even 100 years.

 

Fish behaviour is a complicated and varied subject. As in almost all animals with a central nervous system, the nature of a response of an individual fish to stimuli from its environment depends upon the inherited characteristics of its nervous system, on what it has learned from past experience, and on the nature of the stimuli. Compared with the variety of human responses, however, that of a fish is stereotyped, not subject to much modification by “thought” or learning, and investigators must guard against anthropomorphic interpretations of fish behaviour.

 

Fishes perceive the world around them by the usual senses of sight, smell, hearing, touch, and taste and by special lateral line water-current detectors. In the few fishes that generate electric fields, a process that might best be called electrolocation aids in perception. One or another of these senses often is emphasized at the expense of others, depending upon the fish’s other adaptations. In fishes with large eyes, the sense of smell may be reduced; others, with small eyes, hunt and feed primarily by smell (such as some eels).

 

Specialized behaviour is primarily concerned with the three most important activities in the fish’s life: feeding, reproduction, and escape from enemies. Schooling behaviour of sardines on the high seas, for instance, is largely a protective device to avoid enemies, but it is also associated with and modified by their breeding and feeding requirements. Predatory fishes are often solitary, lying in wait to dart suddenly after their prey, a kind of locomotion impossible for beaked parrot fishes, which feed on coral, swimming in small groups from one coral head to the next. In addition, some predatory fishes that inhabit pelagic environments, such as tunas, often school.

 

Sleep in fishes, all of which lack true eyelids, consists of a seemingly listless state in which the fish maintains its balance but moves slowly. If attacked or disturbed, most can dart away. A few kinds of fishes lie on the bottom to sleep. Most catfishes, some loaches, and some eels and electric fishes are strictly nocturnal, being active and hunting for food during the night and retiring during the day to holes, thick vegetation, or other protective parts of the environment.

 

Communication between members of a species or between members of two or more species often is extremely important, especially in breeding behaviour (see below Reproduction). The mode of communication may be visual, as between the small so-called cleaner fish and a large fish of a very different species. The larger fish often allows the cleaner to enter its mouth to remove gill parasites. The cleaner is recognized by its distinctive colour and actions and therefore is not eaten, even if the larger fish is normally a predator. Communication is often chemical, signals being sent by specific chemicals called pheromones.

 

Many fishes have a streamlined body and swim freely in open water. Fish locomotion is closely correlated with habitat and ecological niche (the general position of the animal to its environment).

 

Many fishes in both marine and fresh waters swim at the surface and have mouths adapted to feed best (and sometimes only) at the surface. Often such fishes are long and slender, able to dart at surface insects or at other surface fishes and in turn to dart away from predators; needlefishes, halfbeaks, and topminnows (such as killifish and mosquito fish) are good examples. Oceanic flying fishes escape their predators by gathering speed above the water surface, with the lower lobe of the tail providing thrust in the water. They then glide hundreds of yards on enlarged, winglike pectoral and pelvic fins. South American freshwater flying fishes escape their enemies by jumping and propelling their strongly keeled bodies out of the water.

 

So-called mid-water swimmers, the most common type of fish, are of many kinds and live in many habitats. The powerful fusiform tunas and the trouts, for example, are adapted for strong, fast swimming, the tunas to capture prey speedily in the open ocean and the trouts to cope with the swift currents of streams and rivers. The trout body form is well adapted to many habitats. Fishes that live in relatively quiet waters such as bays or lake shores or slow rivers usually are not strong, fast swimmers but are capable of short, quick bursts of speed to escape a predator. Many of these fishes have their sides flattened, examples being the sunfish and the freshwater angelfish of aquarists. Fish associated with the bottom or substrate usually are slow swimmers. Open-water plankton-feeding fishes almost always remain fusiform and are capable of rapid, strong movement (for example, sardines and herrings of the open ocean and also many small minnows of streams and lakes).

 

Bottom-living fishes are of many kinds and have undergone many types of modification of their body shape and swimming habits. Rays, which evolved from strong-swimming mid-water sharks, usually stay close to the bottom and move by undulating their large pectoral fins. Flounders live in a similar habitat and move over the bottom by undulating the entire body. Many bottom fishes dart from place to place, resting on the bottom between movements, a motion common in gobies. One goby relative, the mudskipper, has taken to living at the edge of pools along the shore of muddy mangrove swamps. It escapes its enemies by flipping rapidly over the mud, out of the water. Some catfishes, synbranchid eels, the so-called climbing perch, and a few other fishes venture out over damp ground to find more promising waters than those that they left. They move by wriggling their bodies, sometimes using strong pectoral fins; most have accessory air-breathing organs. Many bottom-dwelling fishes live in mud holes or rocky crevices. Marine eels and gobies commonly are found in such habitats and for the most part venture far beyond their cavelike homes. Some bottom dwellers, such as the clingfishes (Gobiesocidae), have developed powerful adhesive disks that enable them to remain in place on the substrate in areas such as rocky coasts, where the action of the waves is great.

 

The methods of reproduction in fishes are varied, but most fishes lay a large number of small eggs, fertilized and scattered outside of the body. The eggs of pelagic fishes usually remain suspended in the open water. Many shore and freshwater fishes lay eggs on the bottom or among plants. Some have adhesive eggs. The mortality of the young and especially of the eggs is very high, and often only a few individuals grow to maturity out of hundreds, thousands, and in some cases millions of eggs laid.

 

Males produce sperm, usually as a milky white substance called milt, in two (sometimes one) testes within the body cavity. In bony fishes a sperm duct leads from each testis to a urogenital opening behind the vent or anus. In sharks and rays and in cyclostomes the duct leads to a cloaca. Sometimes the pelvic fins are modified to help transmit the milt to the eggs at the female’s vent or on the substrate where the female has placed them. Sometimes accessory organs are used to fertilize females internally—for example, the claspers of many sharks and rays.

 

In the females the eggs are formed in two ovaries (sometimes only one) and pass through the ovaries to the urogenital opening and to the outside. In some fishes the eggs are fertilized internally but are shed before development takes place. Members of about a dozen families each of bony fishes (teleosts) and sharks bear live young. Many skates and rays also bear live young. In some bony fishes the eggs simply develop within the female, the young emerging when the eggs hatch (ovoviviparous). Others develop within the ovary and are nourished by ovarian tissues after hatching (viviparous). There are also other methods utilized by fishes to nourish young within the female. In all live-bearers the young are born at a relatively large size and are few in number. In one family of primarily marine fishes, the surfperches from the Pacific coast of North America, Japan, and Korea, the males of at least one species are born sexually mature, although they are not fully grown.

 

Some fishes are hermaphroditic—an individual producing both sperm and eggs, usually at different stages of its life. Self-fertilization, however, is probably rare.

 

Successful reproduction and, in many cases, defense of the eggs and the young are assured by rather stereotypical but often elaborate courtship and parental behaviour, either by the male or the female or both. Some fishes prepare nests by hollowing out depressions in the sand bottom (cichlids, for example), build nests with plant materials and sticky threads excreted by the kidneys (sticklebacks), or blow a cluster of mucus-covered bubbles at the water surface (gouramis). The eggs are laid in these structures. Some varieties of cichlids and catfishes incubate eggs in their mouths.

 

Some fishes, such as salmon, undergo long migrations from the ocean and up large rivers to spawn in the gravel beds where they themselves hatched (anadromous fishes). Some, such as the freshwater eels (family Anguillidae), live and grow to maturity in fresh water and migrate to the sea to spawn (catadromous fishes). Other fishes undertake shorter migrations from lakes into streams, within the ocean, or enter spawning habitats that they do not ordinarily occupy in other ways.

 

The basic structure and function of the fish body are similar to those of all other vertebrates. The usual four types of tissues are present: surface or epithelial, connective (bone, cartilage, and fibrous tissues, as well as their derivative, blood), nerve, and muscle tissues. In addition, the fish’s organs and organ systems parallel those of other vertebrates.

 

The typical fish body is streamlined and spindle-shaped, with an anterior head, a gill apparatus, and a heart, the latter lying in the midline just below the gill chamber. The body cavity, containing the vital organs, is situated behind the head in the lower anterior part of the body. The anus usually marks the posterior termination of the body cavity and most often occurs just in front of the base of the anal fin. The spinal cord and vertebral column continue from the posterior part of the head to the base of the tail fin, passing dorsal to the body cavity and through the caudal (tail) region behind the body cavity. Most of the body is of muscular tissue, a high proportion of which is necessitated by swimming. In the course of evolution this basic body plan has been modified repeatedly into the many varieties of fish shapes that exist today.

 

The skeleton forms an integral part of the fish’s locomotion system, as well as serving to protect vital parts. The internal skeleton consists of the skull bones (except for the roofing bones of the head, which are really part of the external skeleton), the vertebral column, and the fin supports (fin rays). The fin supports are derived from the external skeleton but will be treated here because of their close functional relationship to the internal skeleton. The internal skeleton of cyclostomes, sharks, and rays is of cartilage; that of many fossil groups and some primitive living fishes is mostly of cartilage but may include some bone. In place of the vertebral column, the earliest vertebrates had a fully developed notochord, a flexible stiff rod of viscous cells surrounded by a strong fibrous sheath. During the evolution of modern fishes the rod was replaced in part by cartilage and then by ossified cartilage. Sharks and rays retain a cartilaginous vertebral column; bony fishes have spool-shaped vertebrae that in the more primitive living forms only partially replace the notochord. The skull, including the gill arches and jaws of bony fishes, is fully, or at least partially, ossified. That of sharks and rays remains cartilaginous, at times partially replaced by calcium deposits but never by true bone.

 

The supportive elements of the fins (basal or radial bones or both) have changed greatly during fish evolution. Some of these changes are described in the section below (Evolution and paleontology). Most fishes possess a single dorsal fin on the midline of the back. Many have two and a few have three dorsal fins. The other fins are the single tail and anal fins and paired pelvic and pectoral fins. A small fin, the adipose fin, with hairlike fin rays, occurs in many of the relatively primitive teleosts (such as trout) on the back near the base of the caudal fin.

 

The skin of a fish must serve many functions. It aids in maintaining the osmotic balance, provides physical protection for the body, is the site of coloration, contains sensory receptors, and, in some fishes, functions in respiration. Mucous glands, which aid in maintaining the water balance and offer protection from bacteria, are extremely numerous in fish skin, especially in cyclostomes and teleosts. Since mucous glands are present in the modern lampreys, it is reasonable to assume that they were present in primitive fishes, such as the ancient Silurian and Devonian agnathans. Protection from abrasion and predation is another function of the fish skin, and dermal (skin) bone arose early in fish evolution in response to this need. It is thought that bone first evolved in skin and only later invaded the cartilaginous areas of the fish’s body, to provide additional support and protection. There is some argument as to which came first, cartilage or bone, and fossil evidence does not settle the question. In any event, dermal bone has played an important part in fish evolution and has different characteristics in different groups of fishes. Several groups are characterized at least in part by the kind of bony scales they possess.

 

Scales have played an important part in the evolution of fishes. Primitive fishes usually had thick bony plates or thick scales in several layers of bone, enamel, and related substances. Modern teleost fishes have scales of bone, which, while still protective, allow much more freedom of motion in the body. A few modern teleosts (some catfishes, sticklebacks, and others) have secondarily acquired bony plates in the skin. Modern and early sharks possessed placoid scales, a relatively primitive type of scale with a toothlike structure, consisting of an outside layer of enamel-like substance (vitrodentine), an inner layer of dentine, and a pulp cavity containing nerves and blood vessels. Primitive bony fishes had thick scales of either the ganoid or the cosmoid type. Cosmoid scales have a hard, enamel-like outer layer, an inner layer of cosmine (a form of dentine), and then a layer of vascular bone (isopedine). In ganoid scales the hard outer layer is different chemically and is called ganoin. Under this is a cosminelike layer and then a vascular bony layer. The thin, translucent bony scales of modern fishes, called cycloid and ctenoid (the latter distinguished by serrations at the edges), lack enameloid and dentine layers.

 

Skin has several other functions in fishes. It is well supplied with nerve endings and presumably receives tactile, thermal, and pain stimuli. Skin is also well supplied with blood vessels. Some fishes breathe in part through the skin, by the exchange of oxygen and carbon dioxide between the surrounding water and numerous small blood vessels near the skin surface.

 

Skin serves as protection through the control of coloration. Fishes exhibit an almost limitless range of colours. The colours often blend closely with the surroundings, effectively hiding the animal. Many fishes use bright colours for territorial advertisement or as recognition marks for other members of their own species, or sometimes for members of other species. Many fishes can change their colour to a greater or lesser degree, by movement of pigment within the pigment cells (chromatophores). Black pigment cells (melanophores), of almost universal occurrence in fishes, are often juxtaposed with other pigment cells. When placed beneath iridocytes or leucophores (bearing the silvery or white pigment guanine), melanophores produce structural colours of blue and green. These colours are often extremely intense, because they are formed by refraction of light through the needlelike crystals of guanine. The blue and green refracted colours are often relatively pure, lacking the red and yellow rays, which have been absorbed by the black pigment (melanin) of the melanophores. Yellow, orange, and red colours are produced by erythrophores, cells containing the appropriate carotenoid pigments. Other colours are produced by combinations of melanophores, erythrophores, and iridocytes.

 

The major portion of the body of most fishes consists of muscles. Most of the mass is trunk musculature, the fin muscles usually being relatively small. The caudal fin is usually the most powerful fin, being moved by the trunk musculature. The body musculature is usually arranged in rows of chevron-shaped segments on each side. Contractions of these segments, each attached to adjacent vertebrae and vertebral processes, bends the body on the vertebral joint, producing successive undulations of the body, passing from the head to the tail, and producing driving strokes of the tail. It is the latter that provides the strong forward movement for most fishes.

 

The digestive system, in a functional sense, starts at the mouth, with the teeth used to capture prey or collect plant foods. Mouth shape and tooth structure vary greatly in fishes, depending on the kind of food normally eaten. Most fishes are predacious, feeding on small invertebrates or other fishes and have simple conical teeth on the jaws, on at least some of the bones of the roof of the mouth, and on special gill arch structures just in front of the esophagus. The latter are throat teeth. Most predacious fishes swallow their prey whole, and the teeth are used for grasping and holding prey, for orienting prey to be swallowed (head first) and for working the prey toward the esophagus. There are a variety of tooth types in fishes. Some fishes, such as sharks and piranhas, have cutting teeth for biting chunks out of their victims. A shark’s tooth, although superficially like that of a piranha, appears in many respects to be a modified scale, while that of the piranha is like that of other bony fishes, consisting of dentine and enamel. Parrot fishes have beaklike mouths with short incisor-like teeth for breaking off coral and have heavy pavementlike throat teeth for crushing the coral. Some catfishes have small brushlike teeth, arranged in rows on the jaws, for scraping plant and animal growth from rocks. Many fishes (such as the Cyprinidae or minnows) have no jaw teeth at all but have very strong throat teeth.

 

Some fishes gather planktonic food by straining it from their gill cavities with numerous elongate stiff rods (gill rakers) anchored by one end to the gill bars. The food collected on these rods is passed to the throat, where it is swallowed. Most fishes have only short gill rakers that help keep food particles from escaping out the mouth cavity into the gill chamber.

 

Once reaching the throat, food enters a short, often greatly distensible esophagus, a simple tube with a muscular wall leading into a stomach. The stomach varies greatly in fishes, depending upon the diet. In most predacious fishes it is a simple straight or curved tube or pouch with a muscular wall and a glandular lining. Food is largely digested there and leaves the stomach in liquid form.

 

Between the stomach and the intestine, ducts enter the digestive tube from the liver and pancreas. The liver is a large, clearly defined organ. The pancreas may be embedded in it, diffused through it, or broken into small parts spread along some of the intestine. The junction between the stomach and the intestine is marked by a muscular valve. Pyloric ceca (blind sacs) occur in some fishes at this junction and have a digestive or absorptive function or both.

 

The intestine itself is quite variable in length, depending upon the fish’s diet. It is short in predacious forms, sometimes no longer than the body cavity, but long in herbivorous forms, being coiled and several times longer than the entire length of the fish in some species of South American catfishes. The intestine is primarily an organ for absorbing nutrients into the bloodstream. The larger its internal surface, the greater its absorptive efficiency, and a spiral valve is one method of increasing its absorption surface.

 

Sharks, rays, chimaeras, lungfishes, surviving chondrosteans, holosteans, and even a few of the more primitive teleosts have a spiral valve or at least traces of it in the intestine. Most modern teleosts have increased the area of the intestinal walls by having numerous folds and villi (fingerlike projections) somewhat like those in humans. Undigested substances are passed to the exterior through the anus in most teleost fishes. In lungfishes, sharks, and rays, it is first passed through the cloaca, a common cavity receiving the intestinal opening and the ducts from the urogenital system.

 

Oxygen and carbon dioxide dissolve in water, and most fishes exchange dissolved oxygen and carbon dioxide in water by means of the gills. The gills lie behind and to the side of the mouth cavity and consist of fleshy filaments supported by the gill arches and filled with blood vessels, which give gills a bright red colour. Water taken in continuously through the mouth passes backward between the gill bars and over the gill filaments, where the exchange of gases takes place. The gills are protected by a gill cover in teleosts and many other fishes but by flaps of skin in sharks, rays, and some of the older fossil fish groups. The blood capillaries in the gill filaments are close to the gill surface to take up oxygen from the water and to give up excess carbon dioxide to the water.

 

Most modern fishes have a hydrostatic (ballast) organ, called the swim bladder, that lies in the body cavity just below the kidney and above the stomach and intestine. It originated as a diverticulum of the digestive canal. In advanced teleosts, especially the acanthopterygians, the bladder has lost its connection with the digestive tract, a condition called physoclistic. The connection has been retained (physostomous) by many relatively primitive teleosts. In several unrelated lines of fishes, the bladder has become specialized as a lung or, at least, as a highly vascularized accessory breathing organ. Some fishes with such accessory organs are obligate air breathers and will drown if denied access to the surface, even in well-oxygenated water. Fishes with a hydrostatic form of swim bladder can control their depth by regulating the amount of gas in the bladder. The gas, mostly oxygen, is secreted into the bladder by special glands, rendering the fish more buoyant; the gas is absorbed into the bloodstream by another special organ, reducing the overall buoyancy and allowing the fish to sink. Some deep-sea fishes may have oils, rather than gas, in the bladder. Other deep-sea and some bottom-living forms have much-reduced swim bladders or have lost the organ entirely.

 

The swim bladder of fishes follows the same developmental pattern as the lungs of land vertebrates. There is no doubt that the two structures have the same historical origin in primitive fishes. More or less intermediate forms still survive among the more primitive types of fishes, such as the lungfishes Lepidosiren and Protopterus.

 

The circulatory, or blood vascular, system consists of the heart, the arteries, the capillaries, and the veins. It is in the capillaries that the interchange of oxygen, carbon dioxide, nutrients, and other substances such as hormones and waste products takes place. The capillaries lead to the veins, which return the venous blood with its waste products to the heart, kidneys, and gills. There are two kinds of capillary beds: those in the gills and those in the rest of the body. The heart, a folded continuous muscular tube with three or four saclike enlargements, undergoes rhythmic contractions and receives venous blood in a sinus venosus. It passes the blood to an auricle and then into a thick muscular pump, the ventricle. From the ventricle the blood goes to a bulbous structure at the base of a ventral aorta just below the gills. The blood passes to the afferent (receiving) arteries of the gill arches and then to the gill capillaries. There waste gases are given off to the environment, and oxygen is absorbed. The oxygenated blood enters efferent (exuant) arteries of the gill arches and then flows into the dorsal aorta. From there blood is distributed to the tissues and organs of the body. One-way valves prevent backflow. The circulation of fishes thus differs from that of the reptiles, birds, and mammals in that oxygenated blood is not returned to the heart prior to distribution to the other parts of the body.

 

The primary excretory organ in fishes, as in other vertebrates, is the kidney. In fishes some excretion also takes place in the digestive tract, skin, and especially the gills (where ammonia is given off). Compared with land vertebrates, fishes have a special problem in maintaining their internal environment at a constant concentration of water and dissolved substances, such as salts. Proper balance of the internal environment (homeostasis) of a fish is in a great part maintained by the excretory system, especially the kidney.

 

The kidney, gills, and skin play an important role in maintaining a fish’s internal environment and checking the effects of osmosis. Marine fishes live in an environment in which the water around them has a greater concentration of salts than they can have inside their body and still maintain life. Freshwater fishes, on the other hand, live in water with a much lower concentration of salts than they require inside their bodies. Osmosis tends to promote the loss of water from the body of a marine fish and absorption of water by that of a freshwater fish. Mucus in the skin tends to slow the process but is not a sufficient barrier to prevent the movement of fluids through the permeable skin. When solutions on two sides of a permeable membrane have different concentrations of dissolved substances, water will pass through the membrane into the more concentrated solution, while the dissolved chemicals move into the area of lower concentration (diffusion).

 

The kidney of freshwater fishes is often larger in relation to body weight than that of marine fishes. In both groups the kidney excretes wastes from the body, but the kidney of freshwater fishes also excretes large amounts of water, counteracting the water absorbed through the skin. Freshwater fishes tend to lose salt to the environment and must replace it. They get some salt from their food, but the gills and skin inside the mouth actively absorb salt from water passed through the mouth. This absorption is performed by special cells capable of moving salts against the diffusion gradient. Freshwater fishes drink very little water and take in little water with their food.

 

Marine fishes must conserve water, and therefore their kidneys excrete little water. To maintain their water balance, marine fishes drink large quantities of seawater, retaining most of the water and excreting the salt. Most nitrogenous waste in marine fishes appears to be secreted by the gills as ammonia. Marine fishes can excrete salt by clusters of special cells (chloride cells) in the gills.

 

There are several teleosts—for example, the salmon—that travel between fresh water and seawater and must adjust to the reversal of osmotic gradients. They adjust their physiological processes by spending time (often surprisingly little time) in the intermediate brackish environment.

 

Marine hagfishes, sharks, and rays have osmotic concentrations in their blood about equal to that of seawater and so do not have to drink water nor perform much physiological work to maintain their osmotic balance. In sharks and rays the osmotic concentration is kept high by retention of urea in the blood. Freshwater sharks have a lowered concentration of urea in the blood.

 

Endocrine glands secrete their products into the bloodstream and body tissues and, along with the central nervous system, control and regulate many kinds of body functions. Cyclostomes have a well-developed endocrine system, and presumably it was well developed in the early Agnatha, ancestral to modern fishes. Although the endocrine system in fishes is similar to that of higher vertebrates, there are numerous differences in detail. The pituitary, the thyroid, the suprarenals, the adrenals, the pancreatic islets, the sex glands (ovaries and testes), the inner wall of the intestine, and the bodies of the ultimobranchial gland make up the endocrine system in fishes. There are some others whose function is not well understood. These organs regulate sexual activity and reproduction, growth, osmotic pressure, general metabolic activities such as the storage of fat and the utilization of foodstuffs, blood pressure, and certain aspects of skin colour. Many of these activities are also controlled in part by the central nervous system, which works with the endocrine system in maintaining the life of a fish. Some parts of the endocrine system are developmentally, and undoubtedly evolutionarily, derived from the nervous system.

 

As in all vertebrates, the nervous system of fishes is the primary mechanism coordinating body activities, as well as integrating these activities in the appropriate manner with stimuli from the environment. The central nervous system, consisting of the brain and spinal cord, is the primary integrating mechanism. The peripheral nervous system, consisting of nerves that connect the brain and spinal cord to various body organs, carries sensory information from special receptor organs such as the eyes, internal ears, nares (sense of smell), taste glands, and others to the integrating centres of the brain and spinal cord. The peripheral nervous system also carries information via different nerve cells from the integrating centres of the brain and spinal cord. This coded information is carried to the various organs and body systems, such as the skeletal muscular system, for appropriate action in response to the original external or internal stimulus. Another branch of the nervous system, the autonomic nervous system, helps to coordinate the activities of many glands and organs and is itself closely connected to the integrating centres of the brain.

 

The brain of the fish is divided into several anatomical and functional parts, all closely interconnected but each serving as the primary centre of integrating particular kinds of responses and activities. Several of these centres or parts are primarily associated with one type of sensory perception, such as sight, hearing, or smell (olfaction).

 

The sense of smell is important in almost all fishes. Certain eels with tiny eyes depend mostly on smell for location of food. The olfactory, or nasal, organ of fishes is located on the dorsal surface of the snout. The lining of the nasal organ has special sensory cells that perceive chemicals dissolved in the water, such as substances from food material, and send sensory information to the brain by way of the first cranial nerve. Odour also serves as an alarm system. Many fishes, especially various species of freshwater minnows, react with alarm to a chemical released from the skin of an injured member of their own species.

 

Many fishes have a well-developed sense of taste, and tiny pitlike taste buds or organs are located not only within their mouth cavities but also over their heads and parts of their body. Catfishes, which often have poor vision, have barbels (“whiskers”) that serve as supplementary taste organs, those around the mouth being actively used to search out food on the bottom. Some species of naturally blind cave fishes are especially well supplied with taste buds, which often cover most of their body surface.

 

Sight is extremely important in most fishes. The eye of a fish is basically like that of all other vertebrates, but the eyes of fishes are extremely varied in structure and adaptation. In general, fishes living in dark and dim water habitats have large eyes, unless they have specialized in some compensatory way so that another sense (such as smell) is dominant, in which case the eyes will often be reduced. Fishes living in brightly lighted shallow waters often will have relatively small but efficient eyes. Cyclostomes have somewhat less elaborate eyes than other fishes, with skin stretched over the eyeball perhaps making their vision somewhat less effective. Most fishes have a spherical lens and accommodate their vision to far or near subjects by moving the lens within the eyeball. A few sharks accommodate by changing the shape of the lens, as in land vertebrates. Those fishes that are heavily dependent upon the eyes have especially strong muscles for accommodation. Most fishes see well, despite the restrictions imposed by frequent turbidity of the water and by light refraction.

 

Fossil evidence suggests that colour vision evolved in fishes more than 300 million years ago, but not all living fishes have retained this ability. Experimental evidence indicates that many shallow-water fishes, if not all, have colour vision and see some colours especially well, but some bottom-dwelling shore fishes live in areas where the water is sufficiently deep to filter out most if not all colours, and these fishes apparently never see colours. When tested in shallow water, they apparently are unable to respond to colour differences.

 

Sound perception and balance are intimately associated senses in a fish. The organs of hearing are entirely internal, located within the skull, on each side of the brain and somewhat behind the eyes. Sound waves, especially those of low frequencies, travel readily through water and impinge directly upon the bones and fluids of the head and body, to be transmitted to the hearing organs. Fishes readily respond to sound; for example, a trout conditioned to escape by the approach of fishermen will take flight upon perceiving footsteps on a stream bank even if it cannot see a fisherman. Compared with humans, however, the range of sound frequencies heard by fishes is greatly restricted. Many fishes communicate with each other by producing sounds in their swim bladders, in their throats by rasping their teeth, and in other ways.

 

A fish or other vertebrate seldom has to rely on a single type of sensory information to determine the nature of the environment around it. A catfish uses taste and touch when examining a food object with its oral barbels. Like most other animals, fishes have many touch receptors over their body surface. Pain and temperature receptors also are present in fishes and presumably produce the same kind of information to a fish as to humans. Fishes react in a negative fashion to stimuli that would be painful to human beings, suggesting that they feel a sensation of pain.

 

An important sensory system in fishes that is absent in other vertebrates (except some amphibians) is the lateral line system. This consists of a series of heavily innervated small canals located in the skin and bone around the eyes, along the lower jaw, over the head, and down the mid-side of the body, where it is associated with the scales. Intermittently along these canals are located tiny sensory organs (pit organs) that apparently detect changes in pressure. The system allows a fish to sense changes in water currents and pressure, thereby helping the fish to orient itself to the various changes that occur in the physical environment.

 

Although a great many fossil fishes have been found and described, they represent a tiny portion of the long and complex evolution of fishes, and knowledge of fish evolution remains relatively fragmentary. In the classification presented in this article, fishlike vertebrates are divided into seven categories, the members of each having a different basic structural organization and different physical and physiological adaptations for the problems presented by the environment. The broad basic pattern has been one of successive replacement of older groups by newer, better-adapted groups. One or a few members of a group evolved a basically more efficient means of feeding, breathing, or swimming or several better ways of living. These better-adapted groups then forced the extinction of members of the older group with which they competed for available food, breeding places, or other necessities of life. As the new fishes became well established, some of them evolved further and adapted to other habitats, where they continued to replace members of the old group already there. The process was repeated until all or almost all members of the old group in a variety of habitats had been replaced by members of the newer evolutionary line.

 

The earliest vertebrate fossils of certain relationships are fragments of dermal armour of jawless fishes (superclass Agnatha, order Heterostraci) from the Upper Ordovician Period in North America, about 450 million years in age. Early Ordovician toothlike fragments from the former Soviet Union are less certainly remains of agnathans. It is uncertain whether the North American jawless fishes inhabited shallow coastal marine waters, where their remains became fossilized, or were freshwater vertebrates washed into coastal deposits by stream action.

 

Jawless fishes probably arose from ancient, small, soft-bodied filter-feeding organisms much like and probably also ancestral to the modern sand-dwelling filter feeders, the Cephalochordata (Amphioxus and its relatives). The body in the ancestral animals was probably stiffened by a notochord. Although a vertebrate origin in fresh water is much debated by paleontologists, it is possible that mobility of the body and protection provided by dermal armour arose in response to streamflow in the freshwater environment and to the need to escape from and resist the clawed invertebrate eurypterids that lived in the same waters. Because of the marine distribution of the surviving primitive chordates, however, many paleontologists doubt that the vertebrates arose in fresh water.

 

Heterostracan remains are next found in what appear to be delta deposits in two North American localities of Silurian age. By the close of the Silurian, about 416 million years ago, European heterostracan remains are found in what appear to be delta or coastal deposits. In the Late Silurian of the Baltic area, lagoon or freshwater deposits yield jawless fishes of the order Osteostraci. Somewhat later in the Silurian from the same region, layers contain fragments of jawed acanthodians, the earliest group of jawed vertebrates, and of jawless fishes. These layers lie between marine beds but appear to be washed out from fresh waters of a coastal region.

 

It is evident, therefore, that by the end of the Silurian both jawed and jawless vertebrates were well established and already must have had a long history of development. Yet paleontologists have remains only of specialized forms that cannot have been the ancestors of the placoderms and bony fishes that appear in the next period, the Devonian. No fossils are known of the more primitive ancestors of the agnathans and acanthodians. The extensive marine beds of the Silurian and those of the Ordovician are essentially void of vertebrate history. It is believed that the ancestors of fishlike vertebrates evolved in upland fresh waters, where whatever few and relatively small fossil beds were made probably have been long since eroded away. Remains of the earliest vertebrates may never be found.

 

By the close of the Silurian, all known orders of jawless vertebrates had evolved, except perhaps the modern cyclostomes, which are without the hard parts that ordinarily are preserved as fossils. Cyclostomes were unknown as fossils until 1968, when a lamprey of modern body structure was reported from the Middle Pennsylvanian of Illinois, in deposits more than 300 million years old. Fossil evidence of the four orders of armoured jawless vertebrates is absent from deposits later than the Devonian. Presumably, these vertebrates became extinct at that time, being replaced by the more efficient and probably more aggressive placoderms, acanthodians, selachians (sharks and relatives), and by early bony fishes. Cyclostomes survived probably because early on they evolved from anaspid agnathans and developed a rasping tonguelike structure and a sucking mouth, enabling them to prey on other fishes. With this way of life they apparently had no competition from other fish groups. Cyclostomes, the hagfishes and lampreys, were once thought to be closely related because of the similarity in their suctorial mouths, but it is now understood that the hagfishes, order Myxiniformes, are the most primitive living chordates, and they are classified separately from the lampreys, order Petromyzontiformes.

 

Early jawless vertebrates probably fed on tiny organisms by filter feeding, as do the larvae of their descendants, the modern lampreys. The gill cavity of the early agnathans was large. It is thought that small organisms taken from the bottom by a nibbling action of the mouth, or more certainly by a sucking action through the mouth, were passed into the gill cavity along with water for breathing. Small organisms then were strained out by the gill apparatus and directed to the food canal. The gill apparatus thus evolved as a feeding, as well as a breathing, structure. The head and gills in the agnathans were protected by a heavy dermal armour; the tail region was free, allowing motion for swimming.

 

Most important for the evolution of fishes and vertebrates in general was the early appearance of bone, cartilage, and enamel-like substance. These materials became modified in later fishes, enabling them to adapt to many aquatic environments and finally even to land. Other basic organs and tissues of the vertebrates—such as the central nervous system, heart, liver, digestive tract, kidney, and circulatory system— undoubtedly were present in the ancestors of the agnathans. In many ways, bone, both external and internal, was the key to vertebrate evolution.

 

The next class of fishes to appear was the Acanthodii, containing the earliest known jawed vertebrates, which arose in the Late Silurian, more than 416 million years ago. The acanthodians declined after the Devonian but lasted into the Early Permian, a little less than 280 million years ago. The first complete specimens appear in Lower Devonian freshwater deposits, but later in the Devonian and Permian some members appear to have been marine. Most were small fishes, not more than 75 cm (approximately 30 inches) in length.

 

We know nothing of the ancestors of the acanthodians. They must have arisen from some jawless vertebrate, probably in fresh water. They appear to have been active swimmers with almost no head armour but with large eyes, indicating that they depended heavily on vision. Perhaps they preyed on invertebrates. The rows of spines and spinelike fins between the pectoral and pelvic fins give some credence to the idea that paired fins arose from “fin folds” along the body sides.

 

The relationships of the acanthodians to other jawed vertebrates are obscure. They possess features found in both sharks and bony fishes. They are like early bony fishes in possessing ganoidlike scales and a partially ossified internal skeleton. Certain aspects of the jaw appear to be more like those of bony fishes than sharks, but the bony fin spines and certain aspects of the gill apparatus would seem to favour relationships with early sharks. Acanthodians do not seem particularly close to the Placodermi, although, like the placoderms, they apparently possessed less efficient tooth replacement and tooth structure than the sharks and the bony fishes, possibly one reason for their subsequent extinction.

Sheep (pl.: sheep) or domestic sheep (Ovis aries) are a domesticated, ruminant mammal typically kept as livestock. Although the term sheep can apply to other species in the genus Ovis, in everyday usage it almost always refers to domesticated sheep. Like all ruminants, sheep are members of the order Artiodactyla, the even-toed ungulates. Numbering a little over one billion, domestic sheep are also the most numerous species of sheep. An adult female is referred to as a ewe (/juː/ yoo), an intact male as a ram, occasionally a tup, a castrated male as a wether, and a young sheep as a lamb.

 

Sheep are most likely descended from the wild mouflon of Europe and Asia, with Iran being a geographic envelope of the domestication center. One of the earliest animals to be domesticated for agricultural purposes, sheep are raised for fleeces, meat (lamb, hogget or mutton) and milk. A sheep's wool is the most widely used animal fiber, and is usually harvested by shearing. In Commonwealth countries, ovine meat is called lamb when from younger animals and mutton when from older ones; in the United States, meat from both older and younger animals is usually called lamb. Sheep continue to be important for wool and meat today, and are also occasionally raised for pelts, as dairy animals, or as model organisms for science.

 

Sheep husbandry is practised throughout the majority of the inhabited world, and has been fundamental to many civilizations. In the modern era, Australia, New Zealand, the southern and central South American nations, and the British Isles are most closely associated with sheep production.

 

There is a large lexicon of unique terms for sheep husbandry which vary considerably by region and dialect. Use of the word sheep began in Middle English as a derivation of the Old English word scēap. A group of sheep is called a flock. Many other specific terms for the various life stages of sheep exist, generally related to lambing, shearing, and age.

 

Being a key animal in the history of farming, sheep have a deeply entrenched place in human culture, and are represented in much modern language and symbolism. As livestock, sheep are most often associated with pastoral, Arcadian imagery. Sheep figure in many mythologies—such as the Golden Fleece—and major religions, especially the Abrahamic traditions. In both ancient and modern religious ritual, sheep are used as sacrificial animals.

 

History

Main article: History of the domestic sheep

The exact line of descent from wild ancestors to domestic sheep is unclear. The most common hypothesis states that Ovis aries is descended from the Asiatic (O. gmelini) species of mouflon; the European mouflon (Ovis aries musimon) is a direct descendant of this population. Sheep were among the first animals to be domesticated by humankind (although the domestication of dogs probably took place 10 to 20 thousand years earlier); the domestication date is estimated to fall between 11,000 and 9000 B.C in Mesopotamia and possibly around 7000 BC in Mehrgarh in the Indus Valley. The rearing of sheep for secondary products, and the resulting breed development, began in either southwest Asia or western Europe. Initially, sheep were kept solely for meat, milk and skins. Archaeological evidence from statuary found at sites in Iran suggests that selection for woolly sheep may have begun around 6000 BC, and the earliest woven wool garments have been dated to two to three thousand years later.

 

Sheep husbandry spread quickly in Europe. Excavations show that in about 6000 BC, during the Neolithic period of prehistory, the Castelnovien people, living around Châteauneuf-les-Martigues near present-day Marseille in the south of France, were among the first in Europe to keep domestic sheep. Practically from its inception, ancient Greek civilization relied on sheep as primary livestock, and were even said to name individual animals. Ancient Romans kept sheep on a wide scale, and were an important agent in the spread of sheep raising. Pliny the Elder, in his Natural History (Naturalis Historia), speaks at length about sheep and wool. European colonists spread the practice to the New World from 1493 onwards.

 

Characteristics

Domestic sheep are relatively small ruminants, usually with a crimped hair called wool and often with horns forming a lateral spiral. They differ from their wild relatives and ancestors in several respects, having become uniquely neotenic as a result of selective breeding by humans. A few primitive breeds of sheep retain some of the characteristics of their wild cousins, such as short tails. Depending on breed, domestic sheep may have no horns at all (i.e. polled), or horns in both sexes, or in males only. Most horned breeds have a single pair, but a few breeds may have several.

 

Sheep in Turkmenistan

Another trait unique to domestic sheep as compared to wild ovines is their wide variation in color. Wild sheep are largely variations of brown hues, and variation within species is extremely limited. Colors of domestic sheep range from pure white to dark chocolate brown, and even spotted or piebald. Sheep keepers also sometimes artificially paint "smit marks" onto their sheep in any pattern or color for identification. Selection for easily dyeable white fleeces began early in sheep domestication, and as white wool is a dominant trait it spread quickly. However, colored sheep do appear in many modern breeds, and may even appear as a recessive trait in white flocks. While white wool is desirable for large commercial markets, there is a niche market for colored fleeces, mostly for handspinning. The nature of the fleece varies widely among the breeds, from dense and highly crimped, to long and hairlike. There is variation of wool type and quality even among members of the same flock, so wool classing is a step in the commercial processing of the fibre.

  

Suffolks are a medium wool, black-faced breed of meat sheep that make up 60% of the sheep population in the U.S.

Depending on breed, sheep show a range of heights and weights. Their rate of growth and mature weight is a heritable trait that is often selected for in breeding. Ewes typically weigh between 45 and 100 kilograms (100 and 220 lb), and rams between 45 and 160 kilograms (100 and 350 lb). When all deciduous teeth have erupted, the sheep has 20 teeth. Mature sheep have 32 teeth. As with other ruminants, the front teeth in the lower jaw bite against a hard, toothless pad in the upper jaw. These are used to pick off vegetation, then the rear teeth grind it before it is swallowed. There are eight lower front teeth in ruminants, but there is some disagreement as to whether these are eight incisors, or six incisors and two incisor-shaped canines. This means that the dental formula for sheep is either

0.0.3.3

4.0.3.3

or

0.0.3.3

3.1.3.3

There is a large diastema between the incisors and the molars.

 

In the first few years of life one can calculate the age of sheep from their front teeth, as a pair of milk teeth is replaced by larger adult teeth each year, the full set of eight adult front teeth being complete at about four years of age. The front teeth are then gradually lost as sheep age, making it harder for them to feed and hindering the health and productivity of the animal. For this reason, domestic sheep on normal pasture begin to slowly decline from four years on, and the life expectancy of a sheep is 10 to 12 years, though some sheep may live as long as 20 years.

 

Skull

Sheep have good hearing, and are sensitive to noise when being handled. Sheep have horizontal slit-shaped pupils, with excellent peripheral vision; with visual fields of about 270° to 320°, sheep can see behind themselves without turning their heads. Many breeds have only short hair on the face, and some have facial wool (if any) confined to the poll and or the area of the mandibular angle; the wide angles of peripheral vision apply to these breeds. A few breeds tend to have considerable wool on the face; for some individuals of these breeds, peripheral vision may be greatly reduced by "wool blindness", unless recently shorn about the face. Sheep have poor depth perception; shadows and dips in the ground may cause sheep to baulk. In general, sheep have a tendency to move out of the dark and into well-lit areas, and prefer to move uphill when disturbed. Sheep also have an excellent sense of smell, and, like all species of their genus, have scent glands just in front of the eyes, and interdigitally on the feet. The purpose of these glands is uncertain, but those on the face may be used in breeding behaviors. The foot glands might also be related to reproduction, but alternative functions, such as secretion of a waste product or a scent marker to help lost sheep find their flock, have also been proposed.

 

Comparison with goats

Sheep and goats are closely related: both are in the subfamily Caprinae. However, they are separate species, so hybrids rarely occur and are always infertile. A hybrid of a ewe and a buck (a male goat) is called a sheep-goat hybrid, known as geep. Visual differences between sheep and goats include the beard of goats and divided upper lip of sheep. Sheep tails also hang down, even when short or docked, while the short tails of goats are held upwards. Also, sheep breeds are often naturally polled (either in both sexes or just in the female), while naturally polled goats are rare (though many are polled artificially). Males of the two species differ in that buck goats acquire a unique and strong odor during the rut, whereas rams do not.

 

Breeds

The domestic sheep is a multi-purpose animal, and the more than 200 breeds now in existence were created to serve these diverse purposes. Some sources give a count of a thousand or more breeds, but these numbers cannot be verified, according to some sources. However, several hundred breeds of sheep have been identified by the Food and Agriculture Organization of the UN (FAO), with the estimated number varying somewhat from time to time: e.g. 863 breeds as of 1993, 1314 breeds as of 1995 and 1229 breeds as of 2006. (These numbers exclude extinct breeds, which are also tallied by the FAO.) For the purpose of such tallies, the FAO definition of a breed is "either a subspecific group of domestic livestock with definable and identifiable external characteristics that enable it to be separated by visual appraisal from other similarly defined groups within the same species or a group for which geographical and/or cultural separation from phenotypically similar groups has led to acceptance of its separate identity." Almost all sheep are classified as being best suited to furnishing a certain product: wool, meat, milk, hides, or a combination in a dual-purpose breed. Other features used when classifying sheep include face color (generally white or black), tail length, presence or lack of horns, and the topography for which the breed has been developed. This last point is especially stressed in the UK, where breeds are described as either upland (hill or mountain) or lowland breeds. A sheep may also be of a fat-tailed type, which is a dual-purpose sheep common in Africa and Asia with larger deposits of fat within and around its tail.

 

Breeds are often categorized by the type of their wool. Fine wool breeds are those that have wool of great crimp and density, which are preferred for textiles. Most of these were derived from Merino sheep, and the breed continues to dominate the world sheep industry. Downs breeds have wool between the extremes, and are typically fast-growing meat and ram breeds with dark faces. Some major medium wool breeds, such as the Corriedale, are dual-purpose crosses of long and fine-wooled breeds and were created for high-production commercial flocks. Long wool breeds are the largest of sheep, with long wool and a slow rate of growth. Long wool sheep are most valued for crossbreeding to improve the attributes of other sheep types. For example: the American Columbia breed was developed by crossing Lincoln rams (a long wool breed) with fine-wooled Rambouillet ewes.

 

Coarse or carpet wool sheep are those with a medium to long length wool of characteristic coarseness. Breeds traditionally used for carpet wool show great variability, but the chief requirement is a wool that will not break down under heavy use (as would that of the finer breeds). As the demand for carpet-quality wool declines, some breeders of this type of sheep are attempting to use a few of these traditional breeds for alternative purposes. Others have always been primarily meat-class sheep.

 

A minor class of sheep are the dairy breeds. Dual-purpose breeds that may primarily be meat or wool sheep are often used secondarily as milking animals, but there are a few breeds that are predominantly used for milking. These sheep produce a higher quantity of milk and have slightly longer lactation curves. In the quality of their milk, the fat and protein content percentages of dairy sheep vary from non-dairy breeds, but lactose content does not.

 

A last group of sheep breeds is that of fur or hair sheep, which do not grow wool at all. Hair sheep are similar to the early domesticated sheep kept before woolly breeds were developed, and are raised for meat and pelts. Some modern breeds of hair sheep, such as the Dorper, result from crosses between wool and hair breeds. For meat and hide producers, hair sheep are cheaper to keep, as they do not need shearing. Hair sheep are also more resistant to parasites and hot weather.

 

With the modern rise of corporate agribusiness and the decline of localized family farms, many breeds of sheep are in danger of extinction. The Rare Breeds Survival Trust of the UK lists 22 native breeds as having only 3,000 registered animals (each), and The Livestock Conservancy lists 14 as either "critical" or "threatened". Preferences for breeds with uniform characteristics and fast growth have pushed heritage (or heirloom) breeds to the margins of the sheep industry. Those that remain are maintained through the efforts of conservation organizations, breed registries, and individual farmers dedicated to their preservation.

 

Diet

Sheep are herbivorous mammals. Most breeds prefer to graze on grass and other short roughage, avoiding the taller woody parts of plants that goats readily consume. Both sheep and goats use their lips and tongues to select parts of the plant that are easier to digest or higher in nutrition. Sheep, however, graze well in monoculture pastures where most goats fare poorly.

 

Ruminant system of a sheep

Like all ruminants, sheep have a complex digestive system composed of four chambers, allowing them to break down cellulose from stems, leaves, and seed hulls into simpler carbohydrates. When sheep graze, vegetation is chewed into a mass called a bolus, which is then passed into the rumen, via the reticulum. The rumen is a 19- to 38-liter (5 to 10 gallon) organ in which feed is fermented. The fermenting organisms include bacteria, fungi, and protozoa. (Other important rumen organisms include some archaea, which produce methane from carbon dioxide.) The bolus is periodically regurgitated back to the mouth as cud for additional chewing and salivation. After fermentation in the rumen, feed passes into the reticulum and the omasum; special feeds such as grains may bypass the rumen altogether. After the first three chambers, food moves into the abomasum for final digestion before processing by the intestines. The abomasum is the only one of the four chambers analogous to the human stomach, and is sometimes called the "true stomach".

 

Other than forage, the other staple feed for sheep is hay, often during the winter months. The ability to thrive solely on pasture (even without hay) varies with breed, but all sheep can survive on this diet. Also included in some sheep's diets are minerals, either in a trace mix or in licks. Feed provided to sheep must be specially formulated, as most cattle, poultry, pig, and even some goat feeds contain levels of copper that are lethal to sheep. The same danger applies to mineral supplements such as salt licks.

 

Grazing behavior

Sheep follow a diurnal pattern of activity, feeding from dawn to dusk, stopping sporadically to rest and chew their cud. Ideal pasture for sheep is not lawnlike grass, but an array of grasses, legumes and forbs. Types of land where sheep are raised vary widely, from pastures that are seeded and improved intentionally to rough, native lands. Common plants toxic to sheep are present in most of the world, and include (but are not limited to) cherry, some oaks and acorns, tomato, yew, rhubarb, potato, and rhododendron.

 

Effects on pasture

Sheep are largely grazing herbivores, unlike browsing animals such as goats and deer that prefer taller foliage. With a much narrower face, sheep crop plants very close to the ground and can overgraze a pasture much faster than cattle. For this reason, many shepherds use managed intensive rotational grazing, where a flock is rotated through multiple pastures, giving plants time to recover. Paradoxically, sheep can both cause and solve the spread of invasive plant species. By disturbing the natural state of pasture, sheep and other livestock can pave the way for invasive plants. However, sheep also prefer to eat invasives such as cheatgrass, leafy spurge, kudzu and spotted knapweed over native species such as sagebrush, making grazing sheep effective for conservation grazing. Research conducted in Imperial County, California compared lamb grazing with herbicides for weed control in seedling alfalfa fields. Three trials demonstrated that grazing lambs were just as effective as herbicides in controlling winter weeds. Entomologists also compared grazing lambs to insecticides for insect control in winter alfalfa. In this trial, lambs provided insect control as effectively as insecticides.

 

Behavior

Sheep are flock animals and strongly gregarious; much sheep behavior can be understood on the basis of these tendencies. The dominance hierarchy of sheep and their natural inclination to follow a leader to new pastures were the pivotal factors in sheep being one of the first domesticated livestock species. Furthermore, in contrast to the red deer and gazelle (two other ungulates of primary importance to meat production in prehistoric times), sheep do not defend territories although they do form home ranges. All sheep have a tendency to congregate close to other members of a flock, although this behavior varies with breed, and sheep can become stressed when separated from their flock members. During flocking, sheep have a strong tendency to follow, and a leader may simply be the first individual to move. Relationships in flocks tend to be closest among related sheep: in mixed-breed flocks, subgroups of the same breed tend to form, and a ewe and her direct descendants often move as a unit within large flocks. Sheep can become hefted to one particular local pasture (heft) so they do not roam freely in unfenced landscapes. Lambs learn the heft from ewes and if whole flocks are culled it must be retaught to the replacement animals.

 

Flock behaviour in sheep is generally only exhibited in groups of four or more sheep; fewer sheep may not react as expected when alone or with few other sheep. Being a prey species, the primary defense mechanism of sheep is to flee from danger when their flight zone is entered. Cornered sheep may charge and butt, or threaten by hoof stamping and adopting an aggressive posture. This is particularly true for ewes with newborn lambs.

 

In regions where sheep have no natural predators, none of the native breeds of sheep exhibit a strong flocking behavior.

 

Herding

Farmers exploit flocking behavior to keep sheep together on unfenced pastures such as hill farming, and to move them more easily. For this purpose shepherds may use herding dogs in this effort, with a highly bred herding ability. Sheep are food-oriented, and association of humans with regular feeding often results in sheep soliciting people for food. Those who are moving sheep may exploit this behavior by leading sheep with buckets of feed.

 

Dominance hierarchy

Sheep establish a dominance hierarchy through fighting, threats and competitiveness. Dominant animals are inclined to be more aggressive with other sheep, and usually feed first at troughs. Primarily among rams, horn size is a factor in the flock hierarchy. Rams with different size horns may be less inclined to fight to establish the dominance order, while rams with similarly sized horns are more so. Merinos have an almost linear hierarchy whereas there is a less rigid structure in Border Leicesters when a competitive feeding situation arises.

 

In sheep, position in a moving flock is highly correlated with social dominance, but there is no definitive study to show consistent voluntary leadership by an individual sheep.

 

Intelligence and learning ability

Sheep are frequently thought of as unintelligent animals. Their flocking behavior and quickness to flee and panic can make shepherding a difficult endeavor for the uninitiated. Despite these perceptions, a University of Illinois monograph on sheep reported their intelligence to be just below that of pigs and on par with that of cattle. Sheep can recognize individual human and ovine faces and remember them for years; they can remember 50 other different sheep faces for over two years; they can recognize and are attracted to individual sheep and humans by their faces, as they possess similar specialized neural systems in the temporal and frontal lobes of their brains to humans and have a greater involvement of the right brain hemisphere. In addition to long-term facial recognition of individuals, sheep can also differentiate emotional states through facial characteristics.[68][69] If worked with patiently, sheep may learn their names, and many sheep are trained to be led by halter for showing and other purposes. Sheep have also responded well to clicker training. Sheep have been used as pack animals; Tibetan nomads distribute baggage equally throughout a flock as it is herded between living sites.

 

It has been reported that some sheep have apparently shown problem-solving abilities; a flock in West Yorkshire, England allegedly found a way to get over cattle grids by rolling on their backs, although documentation of this has relied on anecdotal accounts.

 

Vocalisations

Sounds made by domestic sheep include bleats, grunts, rumbles and snorts. Bleating ("baaing") is used mostly for contact communication, especially between dam and lambs, but also at times between other flock members. The bleats of individual sheep are distinctive, enabling the ewe and her lambs to recognize each other's vocalizations. Vocal communication between lambs and their dam declines to a very low level within several weeks after parturition. A variety of bleats may be heard, depending on sheep age and circumstances. Apart from contact communication, bleating may signal distress, frustration or impatience; however, sheep are usually silent when in pain. Isolation commonly prompts bleating by sheep. Pregnant ewes may grunt when in labor. Rumbling sounds are made by the ram during courting; somewhat similar rumbling sounds may be made by the ewe, especially when with her neonate lambs. A snort (explosive exhalation through the nostrils) may signal aggression or a warning, and is often elicited from startled sheep.

 

Lamb

In sheep breeds lacking facial wool, the visual field is wide. In 10 sheep (Cambridge, Lleyn and Welsh Mountain breeds, which lack facial wool), the visual field ranged from 298° to 325°, averaging 313.1°, with binocular overlap ranging from 44.5° to 74°, averaging 61.7°. In some breeds, unshorn facial wool can limit the visual field; in some individuals, this may be enough to cause "wool blindness". In 60 Merinos, visual fields ranged from 219.1° to 303.0°, averaging 269.9°, and the binocular field ranged from 8.9° to 77.7°, averaging 47.5°; 36% of the measurements were limited by wool, although photographs of the experiments indicate that only limited facial wool regrowth had occurred since shearing. In addition to facial wool (in some breeds), visual field limitations can include ears and (in some breeds) horns, so the visual field can be extended by tilting the head. Sheep eyes exhibit very low hyperopia and little astigmatism. Such visual characteristics are likely to produce a well-focused retinal image of objects in both the middle and long distance. Because sheep eyes have no accommodation, one might expect the image of very near objects to be blurred, but a rather clear near image could be provided by the tapetum and large retinal image of the sheep's eye, and adequate close vision may occur at muzzle length. Good depth perception, inferred from the sheep's sure-footedness, was confirmed in "visual cliff" experiments; behavioral responses indicating depth perception are seen in lambs at one day old. Sheep are thought to have colour vision, and can distinguish between a variety of colours: black, red, brown, green, yellow and white. Sight is a vital part of sheep communication, and when grazing, they maintain visual contact with each other. Each sheep lifts its head upwards to check the position of other sheep in the flock. This constant monitoring is probably what keeps the sheep in a flock as they move along grazing. Sheep become stressed when isolated; this stress is reduced if they are provided with a mirror, indicating that the sight of other sheep reduces stress.

 

Taste is the most important sense in sheep, establishing forage preferences, with sweet and sour plants being preferred and bitter plants being more commonly rejected. Touch and sight are also important in relation to specific plant characteristics, such as succulence and growth form.

 

The ram uses his vomeronasal organ (sometimes called the Jacobson's organ) to sense the pheromones of ewes and detect when they are in estrus. The ewe uses her vomeronasal organ for early recognition of her neonate lamb.

 

Reproduction

Sheep follow a similar reproductive strategy to other herd animals. A group of ewes is generally mated by a single ram, who has either been chosen by a breeder or (in feral populations) has established dominance through physical contest with other rams. Most sheep are seasonal breeders, although some are able to breed year-round. Ewes generally reach sexual maturity at six to eight months old, and rams generally at four to six months. However, there are exceptions. For example, Finnsheep ewe lambs may reach puberty as early as 3 to 4 months, and Merino ewes sometimes reach puberty at 18 to 20 months. Ewes have estrus cycles about every 17 days, during which they emit a scent and indicate readiness through physical displays towards rams.

 

In feral sheep, rams may fight during the rut to determine which individuals may mate with ewes. Rams, especially unfamiliar ones, will also fight outside the breeding period to establish dominance; rams can kill one another if allowed to mix freely. During the rut, even usually friendly rams may become aggressive towards humans due to increases in their hormone levels.

 

After mating, sheep have a gestation period of about five months, and normal labor takes one to three hours. Although some breeds regularly throw larger litters of lambs, most produce single or twin lambs. During or soon after labor, ewes and lambs may be confined to small lambing jugs, small pens designed to aid both careful observation of ewes and to cement the bond between them and their lambs.

  

A lamb's first steps

Ovine obstetrics can be problematic. By selectively breeding ewes that produce multiple offspring with higher birth weights for generations, sheep producers have inadvertently caused some domestic sheep to have difficulty lambing; balancing ease of lambing with high productivity is one of the dilemmas of sheep breeding. In the case of any such problems, those present at lambing may assist the ewe by extracting or repositioning lambs. After the birth, ewes ideally break the amniotic sac (if it is not broken during labor), and begin licking clean the lamb. Most lambs will begin standing within an hour of birth. In normal situations, lambs nurse after standing, receiving vital colostrum milk. Lambs that either fail to nurse or are rejected by the ewe require help to survive, such as bottle-feeding or fostering by another ewe.

 

Most lambs begin life being born outdoors. After lambs are several weeks old, lamb marking (ear tagging, docking, mulesing, and castrating) is carried out. Vaccinations are usually carried out at this point as well. Ear tags with numbers are attached, or ear marks are applied, for ease of later identification of sheep. Docking and castration are commonly done after 24 hours (to avoid interference with maternal bonding and consumption of colostrum) and are often done not later than one week after birth, to minimize pain, stress, recovery time and complications. The first course of vaccinations (commonly anti-clostridial) is commonly given at an age of about 10 to 12 weeks; i.e. when the concentration of maternal antibodies passively acquired via colostrum is expected to have fallen low enough to permit development of active immunity. Ewes are often revaccinated annually about 3 weeks before lambing, to provide high antibody concentrations in colostrum during the first several hours after lambing. Ram lambs that will either be slaughtered or separated from ewes before sexual maturity are not usually castrated. Objections to all these procedures have been raised by animal rights groups, but farmers defend them by saying they save money, and inflict only temporary pain.

 

Homosexuality

Sheep are the only species of mammal except for humans which exhibits exclusive homosexual behavior. About 10% of rams refuse to mate with ewes but readily mate with other rams, and thirty percent of all rams demonstrate at least some homosexual behavior. Additionally, a small number of females that were accompanied by a male fetus in utero (i.e. as fraternal twins) are freemartins (female animals that are behaviorally masculine and lack functioning ovaries).

 

Health

Sheep may fall victim to poisons, infectious diseases, and physical injuries. As a prey species, a sheep's system is adapted to hide the obvious signs of illness, to prevent being targeted by predators. However, some signs of ill health are obvious, with sick sheep eating little, vocalizing excessively, and being generally listless. Throughout history, much of the money and labor of sheep husbandry has aimed to prevent sheep ailments. Historically, shepherds often created remedies by experimentation on the farm. In some developed countries, including the United States, sheep lack the economic importance for drug companies to perform expensive clinical trials required to approve more than a relatively limited number of drugs for ovine use. However, extra-label drug use in sheep production is permitted in many jurisdictions, subject to certain restrictions. In the US, for example, regulations governing extra-label drug use in animals are found in 21 CFR (Code of Federal Regulations) Part 530. In the 20th and 21st centuries, a minority of sheep owners have turned to alternative treatments such as homeopathy, herbalism and even traditional Chinese medicine to treat sheep veterinary problems. Despite some favorable anecdotal evidence, the effectiveness of alternative veterinary medicine has been met with skepticism in scientific journals. The need for traditional anti-parasite drugs and antibiotics is widespread, and is the main impediment to certified organic farming with sheep.

 

Many breeders take a variety of preventive measures to ward off problems. The first is to ensure all sheep are healthy when purchased. Many buyers avoid outlets known to be clearing houses for animals culled from healthy flocks as either sick or simply inferior. This can also mean maintaining a closed flock, and quarantining new sheep for a month. Two fundamental preventive programs are maintaining good nutrition and reducing stress in the sheep. Restraint, isolation, loud noises, novel situations, pain, heat, extreme cold, fatigue and other stressors can lead to secretion of cortisol, a stress hormone, in amounts that may indicate welfare problems. Excessive stress can compromise the immune system. "Shipping fever" (pneumonic mannheimiosis, formerly called pasteurellosis) is a disease of particular concern, that can occur as a result of stress, notably during transport and (or) handling. Pain, fear and several other stressors can cause secretion of epinephrine (adrenaline). Considerable epinephrine secretion in the final days before slaughter can adversely affect meat quality (by causing glycogenolysis, removing the substrate for normal post-slaughter acidification of meat) and result in meat becoming more susceptible to colonization by spoilage bacteria. Because of such issues, low-stress handling is essential in sheep management. Avoiding poisoning is also important; common poisons are pesticide sprays, inorganic fertilizer, motor oil, as well as radiator coolant containing ethylene glycol.

 

Common forms of preventive medication for sheep are vaccinations and treatments for parasites. Both external and internal parasites are the most prevalent malady in sheep, and are either fatal, or reduce the productivity of flocks. Worms are the most common internal parasites. They are ingested during grazing, incubate within the sheep, and are expelled through the digestive system (beginning the cycle again). Oral anti-parasitic medicines, known as drenches, are given to a flock to treat worms, sometimes after worm eggs in the feces has been counted to assess infestation levels. Afterwards, sheep may be moved to a new pasture to avoid ingesting the same parasites. External sheep parasites include: lice (for different parts of the body), sheep keds, nose bots, sheep itch mites, and maggots. Keds are blood-sucking parasites that cause general malnutrition and decreased productivity, but are not fatal. Maggots are those of the bot fly and the blow-fly, commonly Lucilia sericata or its relative L. cuprina. Fly maggots cause the extremely destructive condition of flystrike. Flies lay their eggs in wounds or wet, manure-soiled wool; when the maggots hatch they burrow into a sheep's flesh, eventually causing death if untreated. In addition to other treatments, crutching (shearing wool from a sheep's rump) is a common preventive method. Some countries allow mulesing, a practice that involves stripping away the skin on the rump to prevent fly-strike, normally performed when the sheep is a lamb. Nose bots are fly larvae that inhabit a sheep's sinuses, causing breathing difficulties and discomfort. Common signs are a discharge from the nasal passage, sneezing, and frantic movement such as head shaking. External parasites may be controlled through the use of backliners, sprays or immersive sheep dips.

 

A wide array of bacterial and viral diseases affect sheep. Diseases of the hoof, such as foot rot and foot scald may occur, and are treated with footbaths and other remedies. Foot rot is present in over 97% of flocks in the UK. These painful conditions cause lameness and hinder feeding. Ovine Johne's disease is a wasting disease that affects young sheep. Bluetongue disease is an insect-borne illness causing fever and inflammation of the mucous membranes. Ovine rinderpest (or peste des petits ruminants) is a highly contagious and often fatal viral disease affecting sheep and goats. Sheep may also be affected by primary or secondary photosensitization. Tetanus can also afflict sheep through wounds from shearing, docking, castration, or vaccination. The organism also can be introduced into the reproductive tract by unsanitary humans who assist ewes during lambing.

 

A few sheep conditions are transmissible to humans. Orf (also known as scabby mouth, contagious ecthyma or soremouth) is a skin disease leaving lesions that is transmitted through skin-to-skin contact. Cutaneous anthrax is also called woolsorter's disease, as the spores can be transmitted in unwashed wool. More seriously, the organisms that can cause spontaneous enzootic abortion in sheep are easily transmitted to pregnant women. Also of concern are the prion disease scrapie and the virus that causes foot-and-mouth disease (FMD), as both can devastate flocks. The latter poses a slight risk to humans. During the 2001 FMD pandemic in the UK, hundreds of sheep were culled and some rare British breeds were at risk of extinction due to this.

 

Of the 600,300 sheep lost to the US economy in 2004, 37.3% were lost to predators, while 26.5% were lost to some form of disease. Poisoning accounted for 1.7% of non-productive deaths.

 

Predators

A lamb being attacked by coyotes with a bite to the throat

Other than parasites and disease, predation is a threat to sheep and the profitability of sheep raising. Sheep have little ability to defend themselves, compared with other species kept as livestock. Even if sheep survive an attack, they may die from their injuries or simply from panic. However, the impact of predation varies dramatically with region. In Africa, Australia, the Americas, and parts of Europe and Asia predators are a serious problem. In the United States, for instance, over one third of sheep deaths in 2004 were caused by predation. In contrast, other nations are virtually devoid of sheep predators, particularly islands known for extensive sheep husbandry. Worldwide, canids—including the domestic dog—are responsible for most sheep deaths. Other animals that occasionally prey on sheep include: felines, bears, birds of prey, ravens and feral hogs.

 

Sheep producers have used a wide variety of measures to combat predation. Pre-modern shepherds used their own presence, livestock guardian dogs, and protective structures such as barns and fencing. Fencing (both regular and electric), penning sheep at night and lambing indoors all continue to be widely used. More modern shepherds used guns, traps, and poisons to kill predators, causing significant decreases in predator populations. In the wake of the environmental and conservation movements, the use of these methods now usually falls under the purview of specially designated government agencies in most developed countries.

 

The 1970s saw a resurgence in the use of livestock guardian dogs and the development of new methods of predator control by sheep producers, many of them non-lethal. Donkeys and guard llamas have been used since the 1980s in sheep operations, using the same basic principle as livestock guardian dogs. Interspecific pasturing, usually with larger livestock such as cattle or horses, may help to deter predators, even if such species do not actively guard sheep. In addition to animal guardians, contemporary sheep operations may use non-lethal predator deterrents such as motion-activated lights and noisy alarms.

 

Economic importance

Main article: Agricultural economics

Global sheep stock

in 2019

Number in millions

1. China163.5 (13.19%)

2. India74.3 (5.99%)

3. Australia65.8 (5.31%)

4. Nigeria46.9 (3.78%)

5. Iran41.3 (3.33%)

6. Sudan40.9 (3.3%)

7. Chad35.9 (2.9%)

8. Turkey35.2 (2.84%)

9. United Kingdom33.6 (2.71%)

10. Mongolia32.3 (2.61%)

World total1,239.8

 

Source: UN Food and Agriculture Organization

Sheep are an important part of the global agricultural economy. However, their once vital status has been largely replaced by other livestock species, especially the pig, chicken, and cow. China, Australia, India, and Iran have the largest modern flocks, and serve both local and exportation needs for wool and mutton. Other countries such as New Zealand have smaller flocks but retain a large international economic impact due to their export of sheep products. Sheep also play a major role in many local economies, which may be niche markets focused on organic or sustainable agriculture and local food customers. Especially in developing countries, such flocks may be a part of subsistence agriculture rather than a system of trade. Sheep themselves may be a medium of trade in barter economies.

 

Domestic sheep provide a wide array of raw materials. Wool was one of the first textiles, although in the late 20th century wool prices began to fall dramatically as the result of the popularity and cheap prices for synthetic fabrics. For many sheep owners, the cost of shearing is greater than the possible profit from the fleece, making subsisting on wool production alone practically impossible without farm subsidies. Fleeces are used as material in making alternative products such as wool insulation. In the 21st century, the sale of meat is the most profitable enterprise in the sheep industry, even though far less sheep meat is consumed than chicken, pork or beef.

 

Sheepskin is likewise used for making clothes, footwear, rugs, and other products. Byproducts from the slaughter of sheep are also of value: sheep tallow can be used in candle and soap making, sheep bone and cartilage has been used to furnish carved items such as dice and buttons as well as rendered glue and gelatin. Sheep intestine can be formed into sausage casings, and lamb intestine has been formed into surgical sutures, as well as strings for musical instruments and tennis rackets. Sheep droppings, which are high in cellulose, have even been sterilized and mixed with traditional pulp materials to make paper. Of all sheep byproducts, perhaps the most valuable is lanolin: the waterproof, fatty substance found naturally in sheep's wool and used as a base for innumerable cosmetics and other products.

 

Some farmers who keep sheep also make a profit from live sheep. Providing lambs for youth programs such as 4-H and competition at agricultural shows is often a dependable avenue for the sale of sheep. Farmers may also choose to focus on a particular breed of sheep in order to sell registered purebred animals, as well as provide a ram rental service for breeding. A new option for deriving profit from live sheep is the rental of flocks for grazing; these "mowing services" are hired in order to keep unwanted vegetation down in public spaces and to lessen fire hazard.

 

Despite the falling demand and price for sheep products in many markets, sheep have distinct economic advantages when compared with other livestock. They do not require expensive housing, such as that used in the intensive farming of chickens or pigs. They are an efficient use of land; roughly six sheep can be kept on the amount that would suffice for a single cow or horse. Sheep can also consume plants, such as noxious weeds, that most other animals will not touch, and produce more young at a faster rate. Also, in contrast to most livestock species, the cost of raising sheep is not necessarily tied to the price of feed crops such as grain, soybeans and corn. Combined with the lower cost of quality sheep, all these factors combine to equal a lower overhead for sheep producers, thus entailing a higher profitability potential for the small farmer. Sheep are especially beneficial for independent producers, including family farms with limited resources, as the sheep industry is one of the few types of animal agriculture that has not been vertically integrated by agribusiness. However, small flocks, from 10 to 50 ewes, often are not profitable because they tend to be poorly managed. The primary reason is that mechanization is not feasible, so return per hour of labor is not maximized. Small farm flocks generally are used simply to control weeds on irrigation ditches or maintained as a hobby.

 

Shoulder of lamb

Sheep meat and milk were one of the earliest staple proteins consumed by human civilization after the transition from hunting and gathering to agriculture. Sheep meat prepared for food is known as either mutton or lamb, and approximately 540 million sheep are slaughtered each year for meat worldwide. "Mutton" is derived from the Old French moton, which was the word for sheep used by the Anglo-Norman rulers of much of the British Isles in the Middle Ages. This became the name for sheep meat in English, while the Old English word sceap was kept for the live animal. Throughout modern history, "mutton" has been limited to the meat of mature sheep usually at least two years of age; "lamb" is used for that of immature sheep less than a year.

 

In the 21st century, the nations with the highest consumption of sheep meat are the Arab states of the Persian Gulf, New Zealand, Australia, Greece, Uruguay, the United Kingdom and Ireland. These countries eat 14–40 lbs (3–18 kg) of sheep meat per capita, per annum. Sheep meat is also popular in France, Africa (especially the Arab world), the Caribbean, the rest of the Middle East, India, and parts of China. This often reflects a history of sheep production. In these countries in particular, dishes comprising alternative cuts and offal may be popular or traditional. Sheep testicles—called animelles or lamb fries—are considered a delicacy in many parts of the world. Perhaps the most unusual dish of sheep meat is the Scottish haggis, composed of various sheep innards cooked along with oatmeal and chopped onions inside its stomach. In comparison, countries such as the U.S. consume only a pound or less (under 0.5 kg), with Americans eating 50 pounds (22 kg) of pork and 65 pounds (29 kg) of beef. In addition, such countries rarely eat mutton, and may favor the more expensive cuts of lamb: mostly lamb chops and leg of lamb.

 

Though sheep's milk may be drunk rarely in fresh form, today it is used predominantly in cheese and yogurt making. Sheep have only two teats, and produce a far smaller volume of milk than cows. However, as sheep's milk contains far more fat, solids, and minerals than cow's milk, it is ideal for the cheese-making process. It also resists contamination during cooling better because of its much higher calcium content. Well-known cheeses made from sheep milk include the feta of Bulgaria and Greece, Roquefort of France, Manchego from Spain, the pecorino romano (the Italian word for "sheep" is pecore) and ricotta of Italy. Yogurts, especially some forms of strained yogurt, may also be made from sheep milk. Many of these products are now often made with cow's milk, especially when produced outside their country of origin. Sheep milk contains 4.8% lactose, which may affect those who are intolerant.

 

As with other domestic animals, the meat of uncastrated males is inferior in quality, especially as they grow. A "bucky" lamb is a lamb which was not castrated early enough, or which was castrated improperly (resulting in one testicle being retained). These lambs are worth less at market.

 

In science

Sheep are generally too large and reproduce too slowly to make ideal research subjects, and thus are not a common model organism. They have, however, played an influential role in some fields of science. In particular, the Roslin Institute of Edinburgh, Scotland used sheep for genetics research that produced groundbreaking results. In 1995, two ewes named Megan and Morag were the first mammals cloned from differentiated cells, also referred to as gynomerogony. A year later, a Finnish Dorset sheep named Dolly, dubbed "the world's most famous sheep" in Scientific American, was the first mammal to be cloned from an adult somatic cell. Following this, Polly and Molly were the first mammals to be simultaneously cloned and transgenic.

 

As of 2008, the sheep genome has not been fully sequenced, although a detailed genetic map has been published, and a draft version of the complete genome produced by assembling sheep DNA sequences using information given by the genomes of other mammals. In 2012, a transgenic sheep named "Peng Peng" was cloned by Chinese scientists, who spliced his genes with that of a roundworm (C. elegans) in order to increase production of fats healthier for human consumption.

 

In the study of natural selection, the population of Soay sheep that remain on the island of Hirta have been used to explore the relation of body size and coloration to reproductive success. Soay sheep come in several colors, and researchers investigated why the larger, darker sheep were in decline; this occurrence contradicted the rule of thumb that larger members of a population tend to be more successful reproductively. The feral Soays on Hirta are especially useful subjects because they are isolated.

 

Domestic sheep are sometimes used in medical research, particularly for researching cardiovascular physiology, in areas such as hypertension and heart failure. Pregnant sheep are also a useful model for human pregnancy, and have been used to investigate the effects on fetal development of malnutrition and hypoxia. In behavioral sciences, sheep have been used in isolated cases for the study of facial recognition, as their mental process of recognition is qualitatively similar to humans.

 

Cultural impact

Sheep have had a strong presence in many cultures, especially in areas where they form the most common type of livestock. In the English language, to call someone a sheep or ovine may allude that they are timid and easily led. In contradiction to this image, male sheep are often used as symbols of virility and power; the logos of the Los Angeles Rams football team and the Dodge Ram pickup truck allude to males of the bighorn sheep, Ovis canadensis.

 

Counting sheep is popularly said to be an aid to sleep, and some ancient systems of counting sheep persist today. Sheep also enter in colloquial sayings and idiom frequently with such phrases as "black sheep". To call an individual a black sheep implies that they are an odd or disreputable member of a group. This usage derives from the recessive trait that causes an occasional black lamb to be born into an entirely white flock. These black sheep were considered undesirable by shepherds, as black wool is not as commercially viable as white wool. Citizens who accept overbearing governments have been referred to by the Portmanteau neologism of sheeple. Somewhat differently, the adjective "sheepish" is also used to describe embarrassment.

 

In heraldry

In British heraldry, sheep appear in the form of rams, sheep proper and lambs. These are distinguished by the ram being depicted with horns and a tail, the sheep with neither and the lamb with its tail only. A further variant of the lamb, termed the Paschal lamb, is depicted as carrying a Christian cross and with a halo over its head. Rams' heads, portrayed without a neck and facing the viewer, are also found in British armories. The fleece, depicted as an entire sheepskin carried by a ring around its midsection, originally became known through its use in the arms of the Order of the Golden Fleece and was later adopted by towns and individuals with connections to the wool industry. A sheep on a blue field is depicted on the greater/royal arms of the king of Denmark to represent the Faroe Islands. In 2004 a modernized arms has been adopted by the Faroe Islands, which based on a 15th century coat of arms.

 

Religion and folklore

In antiquity, symbolism involving sheep cropped up in religions in the ancient Near East, the Mideast, and the Mediterranean area: Çatalhöyük, ancient Egyptian religion, the Cana'anite and Phoenician tradition, Judaism, Greek religion, and others. Religious symbolism and ritual involving sheep began with some of the first known faiths: Skulls of rams (along with bulls) occupied central placement in shrines at the Çatalhöyük settlement in 8,000 BCE. In Ancient Egyptian religion, the ram was the symbol of several gods: Khnum, Heryshaf and Amun (in his incarnation as a god of fertility). Other deities occasionally shown with ram features include the goddess Ishtar, the Phoenician god Baal-Hamon, and the Babylonian god Ea-Oannes. In Madagascar, sheep were not eaten as they were believed to be incarnations of the souls of ancestors.

 

There are many ancient Greek references to sheep: that of Chrysomallos, the golden-fleeced ram, continuing to be told through into the modern era. Astrologically, Aries, the ram, is the first sign of the classical Greek zodiac, and the sheep is the eighth of the twelve animals associated with the 12-year cycle of in the Chinese zodiac, related to the Chinese calendar. It is said in Chinese traditions that Hou ji sacrificed sheep. Mongolia, shagai are an ancient form of dice made from the cuboid bones of sheep that are often used for fortunetelling purposes.

 

Sheep play an important role in all the Abrahamic faiths; Abraham, Isaac, Jacob, Moses, and King David were all shepherds. According to the Biblical story of the Binding of Isaac, a ram is sacrificed as a substitute for Isaac after an angel stays Abraham's hand (in the Islamic tradition, Abraham was about to sacrifice Ishmael). Eid al-Adha is a major annual festival in Islam in which sheep (or other animals) are sacrificed in remembrance of this act. Sheep are occasionally sacrificed to commemorate important secular events in Islamic cultures. Greeks and Romans sacrificed sheep regularly in religious practice, and Judaism once sacrificed sheep as a Korban (sacrifice), such as the Passover lamb. Ovine symbols—such as the ceremonial blowing of a shofar—still find a presence in modern Judaic traditions.

 

Collectively, followers of Christianity are often referred to as a flock, with Christ as the Good Shepherd, and sheep are an element in the Christian iconography of the birth of Jesus. Some Christian saints are considered patrons of shepherds, and even of sheep themselves. Christ is also portrayed as the Sacrificial lamb of God (Agnus Dei) and Easter celebrations in Greece and Romania traditionally feature a meal of Paschal lamb. A church leader is often called the pastor, which is derived from the Latin word for shepherd. In many western Christian traditions bishops carry a staff, which also serves as a symbol of the episcopal office, known as a crosier, which is modeled on the shepherd's crook.

 

Sheep are key symbols in fables and nursery rhymes like The Wolf in Sheep's Clothing, Little Bo Peep, Baa, Baa, Black Sheep, and Mary Had a Little Lamb; novels such as George Orwell's Animal Farm and Haruki Murakami's A Wild Sheep Chase; songs such as Bach's Sheep may safely graze (Schafe können sicher weiden) and Pink Floyd's "Sheep", and poems like William Blake's "The Lamb".

This Pokot girl wears a necklace of beads cut from the stem of an asparagus tree.

The Pokhot live in the Baringo and Western Pokot districts of Kenya and in Uganda.

There are two main sub-groups depending of their location and way of life. The first group consist of the Hill Pokot who live in the rainy highlands in the west and in the central south, and are mainly farmers and pastoralists. The second group is made up of the Plains Pokot who live in dry and infertile plains, with their cattles. A homestead is composed of one or more buildings for a man, his wife and children; eventual co-wives live in separate houses. The role of the community in teaching children ethical rules. Most of the Pokot are nomadic and thus have interacted with different peoples, incorporating their social customs.The Pokot are very proud of their culture. The Songs, storytelling, and decorative arts, especially bodily decoration, are very appreciated. They adorn the body with beads, hairstyling, scarification, and the removal of the lower central incisors. Pokot girls wear a beaded necklace made of the stems of an asparagus tree. Most Pokot have some knowledge of herbal medicine, so they often use these treatments along with those of the hospitals. They belong to the Kenya's Nilotic-speaking peoples. .

For the Pokot, the universe has two realms: the above is the realm of the most powerful deities—Tororot, Asis (sun), and llat (rain); and the below is the one where live humans, animals, and plants. Humans are responsible for the realm that they inhabit, but they rely upon divinities to achieve and maintain peace and prosperity. They worship many deities like the sun, moon and believe in the spirit of death.The Pokot communicate with their deities through prayer and sacrifice. They perform it during ethnic festivals and dances. Oracles are responsible for maintaining the spiritual balance within the community. They are superstitious and believe in sorcery, so sometimes they call on shielding lucky sorcery. They have prophets, either male or female, who foresee advise, usually by the means of animal sacrifices. His or her ability is considered as a divine gift. Clan histories recount the changes of location, through poetry and song, emphasizing the vulnerability of humans and the importance of supernatural powers to help them overcome hunger, thirst, and even death. Ceremonies mark the transitions in the people's social lives. Among these are: the cleansing of a couple expecting their first child; the cleansing of newborn infants and their mothers; the cleansing of twins and other children who are born under unusual circumstances; male and female initiation; marriage; sapana, a coming-of-age ceremony for men; and summer-solstice, harvest, and healing ceremonies. The most important rite of passage for most Pokot is circumcision for boys and clitoridectomy for girls. These rites consist of a series of neighborhood-based ceremonies, emphasizing the importance of having a good behavior. When boys are circumcised, they acquire membership in one of eight age sets. Women do not have age-sets. After excisions, for several months, girls have a white painting on their face and wear a hood made of blackened leather with charcoal and oil. This means they are untouchable until the lepan ceremony, that marks the passage to womanhood. Unlike other tribes, the Pokot keep the affiliation to their clan throughout their lives, there is no disruption with marriage. Surprisingly, the agreement before marriage is made by gift giving, from the groom and his family to the bride and her family, often over a period of years (and not the contrary). It often implies the gift of a combination of livestock, goods, and cash to the bride's family, and the allotment of milk cows and rights to land to the bride. The bond between a husband and wife lasts for 3 generations, after what marriages can take place again between the two groups. Polygamy exists but is not prevalent among men before 40. The spirits of the elder anticipate reincarnation in their living descendants: when a child is said to resemble the elder, the same name is given. Disputes are resolved in neighborhood councils and in government courts. Some of the sanctions include shaming, cursing, and bewitching.

 

© Eric Lafforgue

www.ericlafforgue.com

     

French postcard by Imp. Georges Lang, Paris, offered by Chocolats Tobler. Image: Walt Disney.

 

Si and Am are Aunt Sarah's twin Siamese cats in Disney's animation film Lady and the Tramp (Clyde Geronimi, Wilfred Jackson, Hamilton Luske, 1955).

 

Si and Am are a sinister pair of cats who use the love they gain from Aunt Sarah to get others into a large amount of trouble. The cats are sneaky and love to cause trouble and make messes for their own amusement. It is indicated that they care for nothing but each other and take advantage of Aunt Sarah, seeing her as a protector and not as the beloved owner she is. Si and Am are identical seal-point Siamese cats. They are sleek and elegant cream-colored felines with angular, wedge-shaped faces, and large, blue almond eyes, which are oval-shaped and narrow out to points near the ends. They have a deep brown facial point with a darker, almost black muzzle, and dark triangular ears. Their noses are black and small, and they both have two middle incisors that hang over their lower lip; these buck teeth are a commonly used Asian stereotype during the late 1940s and 1950s. Si and Am also have dark brown 'stockings' on their front legs and shorter 'socks' on their hind legs and have dark, long, winding tails.

 

In Lady and the Tramp (1955), Si and Am's owner Aunt Sarah is in charge of Lady and the newborn child. The two cats are commonly seen toted in a picnic basket owned by Aunt Sarah, from which they emerge after her brief departure. While Aunt Sarah is upstairs, Si and Am perform their trademark musical number, 'The Siamese Cat Song'. During their song, they wreak havoc throughout the household, claiming it as their own, and even attempt to eat both the family fish and family bird. During the closure of their song, Lady ferociously chases after them upon hearing of their intentions to sample some of the newborn baby's milk. Collateral damage ensues, and although the cats caused all the trouble, Si and Am feign injury in order to get away with it and lay all the blame on Lady. Si and Am are carried upstairs by Aunt Sarah as they deviously shake their tails. As a result, Lady is muzzled by Aunt Sarah.

 

Source: Disney Wiki and IMDb.

 

And, please check out our blog European Film Star Postcards.

African buffalos

 

Kaffernbüffel

 

Kruger National Park is one of the largest game reserves in Africa. It covers an area of 19,485 km2 (7,523 sq mi) in the provinces of Limpopo and Mpumalanga in northeastern South Africa, and extends 360 km (220 mi) from north to south and 65 km (40 mi) from east to west. The administrative headquarters are in Skukuza. Areas of the park were first protected by the government of the South African Republic in 1898, and it became South Africa's first national park in 1926.

 

To the west and south of the Kruger National Park are the two South African provinces of Limpopo and Mpumalanga. In the north is Zimbabwe, and to the east is Mozambique. It is now part of the Great Limpopo Transfrontier Park, a peace park that links Kruger National Park with the Gonarezhou National Park in Zimbabwe, and with the Limpopo National Park in Mozambique.

 

The park is part of the Kruger to Canyons Biosphere an area designated by the United Nations Educational, Scientific and Cultural Organization (UNESCO) as an International Man and Biosphere Reserve (the "Biosphere").

 

The park has nine main gates allowing entrance to the different camps.

 

(Wikipedia)

 

The African buffalo or Cape buffalo (Syncerus caffer) is a large Sub-Saharan African bovine. Syncerus caffer caffer, the Cape buffalo, is the typical subspecies, and the largest one, found in Southern and East Africa. S. c. nanus (African forest buffalo) is the smallest subspecies, common in forest areas of Central and West Africa, while S. c. brachyceros is in West Africa and S. c. aequinoctialis is in the savannas of East Africa. The adult buffalo's horns are its characteristic feature: they have fused bases, forming a continuous bone shield across the top of the head referred to as a "boss". They are widely regarded as among the most dangerous animals on the African continent, and according to some estimates they gore, trample, and kill over 200 people every year.

 

The African buffalo is not an ancestor of domestic cattle and is only distantly related to other larger bovines. Its unpredictable temperament means that the African buffalo has never been domesticated, unlike its Asian counterpart, the water buffalo. African Cape buffaloes have few predators aside from lions and large crocodiles. As a member of the big five game, the Cape buffalo is a sought-after trophy in hunting.

 

The African buffalo is a very robust species. Its shoulder height can range from 1.0 to 1.7 m (3.3 to 5.6 ft) and its head-and-body length can range from 1.7 to 3.4 m (5.6 to 11.2 ft). Compared with other large bovids, it has a long but stocky body (the body length can exceed the wild water buffalo, which is heavier and taller) and short but thickset legs, resulting in a relatively short standing height. The tail can range from 70 to 110 cm (28 to 43 in) long. Savannah-type buffaloes weigh 500 to 1,000 kg (1,100 to 2,200 lb), with males normally larger than females, reaching the upper weight range. In comparison, forest-type buffaloes, at 250 to 450 kg (600 to 1,000 lb), are only half that size. Its head is carried low; its top is located below the backline. The front hooves of the buffalo are wider than the rear, which is associated with the need to support the weight of the front part of the body, which is heavier and more powerful than the back.

 

Savannah-type buffaloes have black or dark brown coats with age. Old bulls often have whitish circles around their eyes and on their face. Females tend to have more-reddish coats. Forest-type buffaloes are 30-40% smaller, reddish brown in colour, with much more hair growth around the ears and with horns that curve back and slightly up. Calves of both types have red coats.

 

A characteristic feature of the horns of adult male African buffalo (Southern and Eastern populations) is that the bases come very close together, forming a shield referred to as a "boss". From the base, the horns diverge downwards, then smoothly curve upwards and outwards and in some cases inwards and or backwards. In large bulls, the distance between the ends of the horns can reach upwards of one metre (the record being 64.5 inches 164 cm). The horns form fully when the animal reaches the age of five or six years but the bosses do not become "hard" till 8 to 9 years old. In cows, the horns are, on average, 10–20% smaller, and they do not have a boss. Forest buffalo horns are smaller than those of the savanna buffalo from Southern and Eastern Africa, usually measuring less than 40 centimetres (16 in), and are almost never fused.

 

The African buffalo is one of the most successful grazers in Africa. It lives in swamps and floodplains, as well as mopane grasslands, and the forests of the major mountains of Africa. This buffalo prefers a habitat with dense cover, such as reeds and thickets, but can also be found in open woodland. While not particularly demanding in regard to habitat, they require water daily, and so they depend on perennial sources of water. Like the plains zebra, the buffalo can live on tall, coarse grasses. Herds of buffalo mow down grasses and make way for more selective grazers. When feeding, the buffalo makes use of its tongue and wide incisor row to eat grass more quickly than most other African herbivores. Buffaloes do not stay on trampled or depleted areas for long.

 

Other than humans, African buffaloes have few predators and are capable of defending themselves against (and killing) lions. Lions do kill and eat buffalo regularly, and in some regions, the buffaloes are the lions' primary prey. It typically takes quite a few lions to bring down a single adult buffalo. Usually, the entire pride joins the hunt; however, several incidents have been reported in which lone adult male lions have been able to successfully bring down adult animals. The average-sized crocodile typically attacks only old solitary animals and young calves, though they can kill healthy adults, and exceptionally large, old male Nile crocodiles may become semi-habitual predators of buffalo. Also, this crocodilian is the only animal that typically takes down an adult buffalo alone, whereas a pride attack is the preferred method of lions when taking down such large prey. The cheetah, leopard, and spotted hyena are normally a threat only to newborn calves, though very large clans of spotted hyenas have been recorded killing cows (mainly pregnant ones) and, on very rare occasions, full-grown bulls.

 

The African buffalo is susceptible to many diseases, including bovine tuberculosis, corridor disease, and foot and mouth disease. As with many diseases, these problems remain dormant within a population as long as the health of the animals is good. These diseases do, however, restrict the legal movements of the animals and fencing infected areas from unaffected areas is enforced. Some wardens and game managers have managed to protect and breed "disease-free" herds which become very valuable because they can be transported. Most well-known are Lindsay Hunt's efforts to source uninfected animals from the Kruger National Park in South Africa. Some disease-free buffaloes in South Africa have been sold to breeders for close to US$130,000.

 

Herd size is highly variable. The core of the herds is made up of related females, and their offspring, in an almost linear dominance hierarchy. The basic herds are surrounded by subherds of subordinate males, high-ranking males and females, and old or invalid animals. The young males keep their distance from the dominant bull, which is recognizable by the thickness of his horns. During the dry season, males split from the herd and form bachelor groups. Two types of bachelor herds occur: ones made of males aged four to seven years and those of males 12 years or older. During the wet season, the younger bulls rejoin a herd to mate with the females. They stay with them throughout the season to protect the calves. Some older bulls cease to rejoin the herd, as they can no longer compete with the younger, more aggressive males. Males have a linear dominance hierarchy based on age and size. Since a buffalo is safer when a herd is larger, dominant bulls may rely on subordinate bulls and sometimes tolerate their copulation.

 

Adult bulls spar in play, dominance interactions, or actual fights. A bull approaches another, lowing, with his horns down, and waits for the other bull to do the same thing. When sparring, the bulls twist their horns from side to side.[27] If the sparring is for play, the bull may rub his opponent's face and body during the sparring session. Actual fights are violent but rare and brief. Calves may also spar in play, but adult females rarely spar at all.

 

African buffaloes are notable for their apparent altruism. Females appear to exhibit some sort of "voting behavior". During resting time, the females stand up, shuffle around, and sit back down again. They sit in the direction they think they should move. After an hour of more shuffling, the females travel in the direction they decide. This decision is communal and not based on hierarchy or dominance. When chased by predators, a herd sticks close together and makes it hard for the predators to pick off one member. Calves are gathered in the middle. A buffalo herd responds to the distress call of a captured member and tries to rescue it. A calf's distress call gets the attention of not only the mother, but also the herd. Buffaloes engage in mobbing behavior when fighting off predators. They have been recorded killing a lion and chasing lions up trees and keeping them there for two hours, after the lions have killed a member of their group. Lion cubs can get trampled and killed. In one videotaped instance, known as the Battle at Kruger, a calf survived an attack by both lions and a crocodile after intervention of the herd.

 

African buffaloes make various vocalizations. Many calls are lower-pitched versions of those emitted by domestic cattle. They emit low-pitched, two- to four-second calls intermittently at three- to six-second intervals to signal the herd to move. To signal to the herd to change direction, leaders emit "gritty", "creaking gate" sounds. When moving to drinking places, some individuals make long "maaa" calls up to 20 times a minute. When being aggressive, they make explosive grunts that may last long or turn into a rumbling growl. Cows produce croaking calls when looking for their calves. Calves make a similar call of a higher pitch when in distress. When threatened by predators, they make drawn-out "waaaa" calls. Dominant individuals make calls to announce their presence and location. A version of the same call, but more intense, is emitted as a warning to an encroaching inferior. When grazing, they make various sounds, such as brief bellows, grunts, honks, and croaks.

 

African buffaloes mate and give birth only during the rainy seasons. Birth peak takes place early in the season, while mating peaks later. A bull closely guards a cow that comes into heat, while keeping other bulls at bay. This is difficult, as cows are quite evasive and attract many males to the scene. By the time a cow is in full estrus, only the most dominant bull in the herd/subherd is there.

 

Cows first calve at five years of age, after a gestation period of 11.5 months. Newborn calves remain hidden in vegetation for the first few weeks while being nursed occasionally by the mother before joining the main herd. Older calves are held in the centre of the herd for safety.

 

The maternal bond between mother and calf lasts longer than in most bovids. However, when a new calf is born, the bonding ends and the mother keeps her previous offspring at bay with horn jabs. Nevertheless, the yearling follows its mother for another year or so. Males leave their mothers when they are two years old and join the bachelor groups. Young calves, unusually for bovids, suckle from behind their mothers, pushing their heads between the mothers' legs.

 

The current status of the African buffalo is dependent on the animal's value to both trophy hunters and tourists, paving the way for conservation efforts through anti-poaching patrols, village crop damage payouts, and CAMPFIRE payback programs to local areas.

 

The African buffalo was listed as least concern by the IUCN "as the species had a global population estimated at nearly 900,000 animals, of which more than three-quarters are in protected areas. However, in 2019 the African buffalo was listed as a near threatened species, with only 400,000 individuals left. While some populations (subspecies) are decreasing, others will remain unchanged in the long term if large, healthy populations continue to persist in a substantial number of national parks, equivalent reserves and hunting zones in southern and eastern Africa."

 

In the most recent and available census data at continental scale, the total estimated numbers of the three African buffalo savanna subspecies (S. c. caffer, S. c. brachyceros, S. c. aequinoctialis) are at 513,000 individuals.

 

In the past, numbers of African buffaloes suffered their most severe collapse during the great rinderpest epidemic of the 1890s, which, coupled with pleuro-pneumonia, caused mortalities as high as 95% among livestock and wild ungulates.

 

Being a member of the big five game group, a term originally used to describe the five most dangerous animals to hunt, the Cape buffalo is a sought-after trophy, with some hunters paying over $10,000 for the opportunity to hunt one. The larger bulls are targeted for their trophy value, although in some areas, buffaloes are still hunted for meat.

 

One of the "big five" African game, it is known as "the Black Death" or "the widowmaker", and is widely regarded as a very dangerous animal. According to some estimates, it gores and kills over 200 people every year. African buffaloes are sometimes reported to kill more people in Africa than any other animal, although the same claim is also made of hippopotamuses and crocodiles. These numbers may be somewhat overestimated; for example, in the country of Mozambique, attacks, especially fatal ones, were much less frequent on humans than those by hippos and, especially, Nile crocodiles. In Uganda, on the other hand, large herbivores were found to attack more people on average than lions or leopards and have a higher rate of inflicting fatalities during attacks than the predators (the African buffalo, in particular, killing humans in 49.5% of attacks on them), but hippos and even elephants may still kill more people per annum here than buffaloes. African buffaloes are notorious among big-game hunters as very dangerous animals, with wounded animals reported to ambush and attack pursuers.

 

(Wikipedia)

 

Der Kruger-Nationalpark (deutsch häufig falsch Krüger-Nationalpark) ist das größte Wildschutzgebiet Südafrikas. Er liegt im Nordosten des Landes in der Landschaft des Lowveld auf dem Gebiet der Provinz Limpopo sowie des östlichen Abschnitts von Mpumalanga. Seine Fläche erstreckt sich vom Crocodile-River im Süden bis zum Limpopo, dem Grenzfluss zu Simbabwe, im Norden. Die Nord-Süd-Ausdehnung beträgt etwa 350 km, in Ost-West-Richtung ist der Park durchschnittlich 54 km breit und umfasst eine Fläche von rund 20.000 Quadratkilometern. Damit gehört er zu den größten Nationalparks in Afrika.

 

Das Schutzgebiet wurde am 26. März 1898 unter dem Präsidenten Paul Kruger als Sabie Game Reserve zum Schutz der Wildnis gegründet. 1926 erhielt das Gebiet den Status Nationalpark und wurde in seinen heutigen Namen umbenannt. Im Park leben 147 Säugetierarten inklusive der „Big Five“, außerdem etwa 507 Vogelarten und 114 Reptilienarten, 49 Fischarten und 34 Amphibienarten.

 

(Wikipedia)

 

Der Kaffernbüffel (Syncerus caffer), auch Schwarzbüffel, Afrikanischer Büffel oder Steppenbüffel genannt, ist ein sehr großer Vertreter aus der Familie der Hornträger, welcher in weiten Teilen des östlichen und südlichen Afrikas vorkommt. Dort bewohnt er offene Savannenlandschaften und bewaldete Flussgebiete des Tief- und Hochlands. Charakteristisch ist der robuste Körperbau und die großen, abwärts geschwungenen Hörner, die auf großen Hornbasen auf der Stirn aufsitzen. Das Sozialsystem des Kaffernbüffels ist komplex und umfasst Herden aus verwandten Kühen mit Jungtieren und gelegentlich Bullen, Junggesellengruppen aus männlichen Tieren und einzelne männliche Individuen. Vor allem die jeweiligen Herden zeigen ein dynamisches Verhalten durch permanentes Aufspalten und Wiedervereinen. Sie ziehen in räumlich begrenzten Aktionsgebieten in einem jährlich von Witterung und Nahrungsangebot vorgegebenen Rhythmus umher. Die sie begleitenden oder flankierenden Bullen sind regelmäßig in Dominanzkämpfe um das Paarungsvorrecht eingebunden, die mit Kopf- oder Körperrammen durchgeführt werden. Kühe bringen in der Regel ein einzelnes Jungtier zur Welt, welches rund ein bis anderthalb Jahre gesäugt wird. Der weibliche Nachwuchs verbleibt zumeist in der Herde, der männliche verlässt sie später. Die Nahrung umfasst überwiegend harte Gräser, in Trockenzeiten kann aber auch der Anteil an weicheren Pflanzen deutlich ansteigen. Dadurch haben die Tiere saisonal und regional bedingt unterschiedliche Ernährungsstrategien. Die Bestände sind im Übergang vom 19. zum 20. Jahrhundert in weiten Teilen des Verbreitungsgebietes in Folge einer Rinderpest-Epidemie zusammengebrochen. Heute ist der Kaffernbüffel weitgehend auf Schutzgebiete beschränkt, gilt aber als häufig und nicht bedroht. Die Art wurde 1779 beschrieben.

 

Der Kaffernbüffel stellt den größten Vertreter der afrikanischen Rinder (Bovini) dar und erreicht die Ausmaße seiner asiatischen, wildlebenden Verwandten. Er besitzt eine Kopf-Rumpf-Länge von 240 bis 340 cm (zuzüglich eines 50 bis 110 cm langen Schwanzes), eine Schulterhöhe von 148 bis 175 cm und ein Gewicht von 350 bis 900 kg. Ein Sexualdimorphismus ist deutlich ausgeprägt, die Bullen sind größer und können bis zu doppelt so schwer wie die Weibchen werden. Untersuchungen von Tieren aus Tansania ergaben ein Gewicht für männliche Tiere von 661 bis 849 kg, für weibliche von 426 bis 468 kg. Entsprechende Angaben für Tiere aus Sambia liegen bei 472 bis 723 kg beziehungsweise 386 bis 536 kg. Insgesamt ist der Kaffernbüffel sehr kräftig gebaut und besitzt verhältnismäßig kurze Gliedmaßen und einen Buckel oberhalb der Schultern. Das Fell hat in beiden Geschlechtern eine rabenschwarze Färbung, bei alten Bullen kann es ausdünnen, so dass Flecken von dunkler Haut am Kopf sichtbar werden. Der Schwanz endet in einer auffälligen, ebenfalls dunklen Quaste. Der massige Kopf ist mit einem breiten Maul ausgestattet, die Nase ist feucht und nackt. Die großen, hängenden Ohren sind gefranst. Besonders auffällig zeigen sich die großen, am Ende spitzen Hörner, die quer über den vorderen Kopf verlaufend ansetzen und seitlich nach unten biegen. Sie spannen 73 bis 134 cm weit auseinander, die Länge der einzelnen Hörner über die Krümmung gemessen ist mit 66 bis 116 cm nahezu identisch zur Spannweite. Das längste jemals gemessene Horn erreichte 163 cm und stammt von einem Bullen aus dem Lake-Manyara-Nationalpark in Tansania. Die Hörner sind nicht geriffelt und vor allem bei Männchen besonders massiv, ihre Basen dehnen sich deutlich über den Vorderkopf und treffen sich auf der Mittellinie des Schädels. Es gibt aber keine Übereinstimmung mit dem Alter der Tiere und der Größe der Hornbasis. Im Gegensatz zu weiblichen Tieren mit ihren grazileren Hörnern wachsen bei den männlichen zwischen den Hornansätzen keine Haare. Die Hufe haben eine große, gerundete Form, zudem treten Afterklauen auf. Die Hinterfußlänge beträgt 56 bis 61 cm. Bestimmte Drüsen zum Absetzen von Sekreten sind nicht ausgebildet. Kühe haben einen kleinen, gerundeten Euter mit zwei Zitzenpaaren.

 

Der Schädel wird 44,8 bis 57,5 cm lang und am Warzenfortsatz des Schläfenbeins 24,1 bis 31,6 cm breit. Allgemein ist er kurz und breit und im Profil deutlich gerundet. Eine Voraugengrube (Fossa praeorbitalis) und eine Siebgrube (Fossa ethmoidalis) sind nicht ausgebildet. Im oberen Gebiss ist typisch für die Wiederkäuer anstatt der Schneidezähne eine Hornplatte ausgebildet. Die Backenzähne sind stark hochkronig (hypsodont) mit einer Kronenhöhe von 5 cm bei jungadulten Tieren, auf der Kaufläche sind scharfe Zahnschmelzleisten ausgebildet. Das Durchbruchsmuster der Zähne kann zur Altersbestimmung verwendet werden, der Eckzahn des Unterkiefers ist der letzte durchbrechende Zahn des Dauergebisses. Er erscheint mit viereinhalb bis fünfeinhalb Jahren.

 

Der Kaffernbüffel kommt in weiten Teilen des östlichen und südlichen Afrikas vor. Das Verbreitungsgebiet reicht vom südlichen Äthiopien und Somalia im Nordosten südwärts über Kenia, Tansania, Uganda und dem äußersten Osten und Südosten der Demokratischen Republik Kongo in Ostafrika sowie über Sambia, Simbabwe, Mosambik bis nach Südafrika, dem Nordosten von Namibia und Süden von Botswana. Die Art bewohnt eine Vielzahl von verschiedenen Landschaftstypen, die gut durchfeuchtete Savannen, Sümpfe und Überflutungsebenen umfassen. Daneben tritt sie aber auch in trockeneren Savannen und Auwäldern trockener Habitate auf, sofern Wasser zur Verfügung steht. Die Tiere sind auch in montanen Waldgebieten bis auf 3000 m Höhe anzutreffen, extrem hohe Nachweise liegen bei 4700 m am Mount Kenya. In besonders zuträglichen Gebieten, etwa im Lake-Manyara-Nationalpark in Tansania, kann der Kaffernbüffel mit rund 20 Individuen je Quadratkilometer in sehr hoher Anzahl auftreten, im Ngorongoro-Krater liegt die Populationsdichte entsprechend bei etwa 11, in der Serengeti bei 8 Individuen auf einer vergleichbar großen Fläche. In Trockenlandschaften wie etwa dem Lowveld in Südafrika geht sie auf bis zu 2 Individuen je Quadratkilometer zurück.

 

Der Kaffernbüffel besitzt ein komplexes Sozialsystem. Typisch ist der Verband aus miteinander verwandten Weibchen und ihren Jungtieren, der als Herde bezeichnet wird und in dem sich auch Bullen unterschiedlichen Alters aufhalten können. Die Größe der Herde variiert von einigen Dutzend bis mehreren Tausend Individuen. Die extrem großen Gruppen stellen aber vermutlich Herdenverbände dar, die nur temporär bestehen und keine innere Hierarchie besitzen. Ihre Bildungen sind abhängig von der Jahreszeit und dem Angebot an Nahrung und Wasser. Es gibt jedoch eine Variation der Herdengröße über das Jahr, die zu größeren Gruppen in der feuchten und kleineren in der trockenen Jahreszeit führen, wobei die Gruppen sich regelmäßig teilen und wieder vereinen.[5] Solche Trennungen und Neuformierungen sind aber nicht nur an äußere Bedingungen geknüpft, sondern teilweise auch an die Größe der Herde und ihre soziale Struktur, wie Untersuchungen am Lake Manyara ergaben. Die hierarchische Struktur einer Herde besteht aus dominanten Weibchen, die die Herde anführen, gefolgt von Kühen mit sehr jungen Kälbern und Jungtieren im mittleren Teil. Den Abschluss der Herde bilden Tiere mit untergeordnetem Status; die Position eines Tiers in der Herde gibt somit Aufschluss über dessen soziale Stellung. Sie bestimmt darüber hinaus auch die Qualität der Nahrung und die allgemeine körperliche Kondition der Kuh. Häufig führt die Geburt eines Kalbes zu einer höheren sozialen Stellung innerhalb der Herde. Es wird meist angenommen, dass die einzelnen Herden (ohne temporäre Aufsplittung) stabile Verbände darstellen und erwachsene Kühe nur selten ihre angestammte Gruppe verlassen, einige Herden wurden über einen Zeitraum von bis zu fünf Dekaden dokumentiert. Beobachtungen aus dem Chobe-Nationalpark in Botswana zeigen aber, dass gelegentlich weibliche Tiere ihre Herde verlassen. Eine Kuh legte während eines solchen Herdenwechsels bis zu 133 km zurück. Ein Wiederaufsuchen der alten Herde wurde dabei nicht festgestellt, möglicherweise handelt es sich bei wechselnden Tieren um solche mit einem eher geringen Status. Bullen leben einzeln oder schließen sich zu Junggesellengruppen mit einer Größe von 5 bis 10 Individuen zusammen, die zeitlich begrenzt bestehen. Innerhalb dieser Junggesellengruppen kommt es permanent zu Dominanzkämpfen um das Paarungsvorrecht. Kurzfristig können Bullen auch von Junggesellengruppen zu Herden und umgekehrt wechseln, innerhalb der Herde sind sie aber nicht in das dortige Sozialgefüge eingebunden. Bemerkenswert ist, dass Bullen, wenn sie sich über längere Zeit in Herden aufhalten, regelmäßig an Gewicht verlieren, da sie durchschnittlich weniger fressen. Vermutlich sind die hohen energetischen Kosten, die für die Fortpflanzung aufgebracht werden müssen, ursächlich dafür verantwortlich, weswegen Bullen die Herden nach kürzerer Zeit wieder verlassen.

 

Die Herden des Kaffernbüffels sind nicht migratorisch und legen somit keine großen Entfernungen zurück. Sie halten sich in mehr oder weniger begrenzten Aktionsräumen von 100 bis über 1000 km² auf. Innerhalb dieser Aktionsräume folgen sie einem regelmäßigen Wanderungsmuster über das Jahr hinweg, das durch den Zyklus von Niederschlägen und Pflanzenwachstum bestimmt wird. Diese einem bestimmten Rhythmus folgenden Wanderungen auf festgelegten Pfaden und Wegen führen dazu, dass der Boden aufgearbeitet und dadurch neues Pflanzenwachstum angeregt wird, was das wiederholte Weiden in der gleichen Region ermöglicht. Typische Wanderungsbewegungen führen zu Wasser- und Weidestellen, sie sind täglich umfangreicher, je weiter zugängliches Wasser verstreut liegt. Im Krüger-Nationalpark in Südafrika werden so Distanzen von durchschnittlich 3,35 km täglich überwunden, in der ostafrikanischen Serengeti sind es dazu im Vergleich bis zu 30 km. Einzelne Bullen und Junggesellengruppen haben wesentlich kleinere Territorien von 3 bis 4 km² Größe. Sie flankieren den Wanderungsverlauf der Herde. Der Tagesablauf folgt ähnlichen Mustern wie bei zahlreichen anderen Wiederkäuern und ist geprägt von der Nahrungsaufnahme und der daraus folgenden Ruhe und dem Wiederkäuen. Insgesamt ist die Dauer der jeweiligen Aktivität abhängig von der Qualität und Quantität des lokalen und saisonalen Nahrungsangebots. Die hauptsächlichen Aktivitäten erfolgen tagsüber, in Gebieten mit großem Jagddruck aber auch nachts. Am Lake Manyara fressen die Tiere zwischen 10:00 Uhr und 14:00 Uhr, wobei die Dauer je nach Intensität zwischen 1,5 und 4,5 Stunden betragen kann, was etwa 22 bis 56 % der Tagesaktivität entspricht (durchschnittlich 37 %). Die Nahrungsaufnahme trennt hier zwei Ruheperioden, die meist mit Wiederkäuen verbracht werden, häufig mit dem Körper im Wasser liegend, und wiederum mehrere Stunden andauern können. Sie nehmen zwischen 11 und 49 % des täglich verfügbaren Zeitbudgets in Anspruch (durchschnittlich 31 %). Somit verbringt der Kaffernbüffel am Lake Manyara jährlich etwa zwei Drittel seiner Tageszeit mit Fressen und dem zugehörigen Verdauen. Dabei ist die Dauer der Verdauung gegenläufig zur Dauer der Nahrungsaufnahme, da ein Tier in der Regenzeit durch das üppigere Angebot qualitativ hochwertiger Nahrung in kürzerer Zeit mehr fressen kann, aber längere Zeit zum Wiederkäuen benötigt. In der Trockenzeit mit einem qualitativ minderwertigerem Nahrungsangebot frisst er länger, verdaut aber kürzer. Im Chebera-Churchura-Nationalpark in Äthiopien weiden die Tiere in den frühen Morgen- und Abendstunden und verbrauchen dafür bis zur Hälfte ihres täglichen Zeitbudgets, ein weiteres gutes Drittel nimmt die Ruhe in Anspruch. Ähnlich lang verbringt der Kaffernbüffel im Krüger-Nationalpark mit dem Fressen, dies erfolgt hier aber teilweise auch nachts.

 

Zum Komfortverhalten gehören unter anderem Schlammbäder, um eventuell Insektenbisse zu vermeiden und Ruhephasen an erhöhten Landmarken. Letzteres dient wahrscheinlich dazu, kühle Brisen zu erhaschen, da der Kaffernbüffel allgemein schlecht an Hitze angepasst ist. Generell ist der Kaffernbüffel eher still. Bekannte Lautäußerungen sind ein Grunzen und Schnaufen, letzteres wird ausgestoßen, wenn ein Tier alarmiert ist oder zu Laufen anfängt. Ein Bellen äußert der Kaffernbüffel bei starken Verletzungen, beispielsweise durch einen Beutegreifer, das andere Artgenossen häufig animiert, zur Unterstützung zu kommen. Schwer verletzte Tiere suchen zum Schutz andere Kaffernbüffel auf. Ein Warnruf bei Gefahr besteht nicht, aber vor allem in der Nacht kann eine Herde sehr still werden, so dass ein Aufspüren sehr schwierig ist. Zwischen dominanten und unterwürfigen Tieren besteht ein ritualisiertes Verhaltensmuster. Erstere halten den Kopf und die Schultern nach oben, während die Nase nach unten gerichtet ist, so dass die Hörner optimal präsentiert werden. Bei letzteren liegt der Kopf tief und parallel zum Boden, teilweise führt das unterlegene Tier auch den Kopf zwischen die Hinterbeine des dominanten und stößt ein Bellen aus. Der Sehsinn ist recht gut entwickelt, ebenso wie das Gehör. Aufgrund der fehlenden Drüsen spielen Sekrete für die innerartliche Kommunikation nur eine untergeordnete Rolle.

 

Als weitgehende Offenlandart besteht die Hauptnahrung des Kaffernbüffels aus Gräsern. Hierbei bevorzugt er langhalmige Gräser vor breitblättrigen und toleriert auch solche mit einem hohen Faseranteil. Er meidet aber solche mit niedrigem Nährstoffgehalt oder Pflanzen, die ätherische Öle enthalten. Zu den am häufigsten verzehrten Pflanzen gehören Hundszahngräser, Fingerhirsen und Rispenhirsen sowie Vertreter der Gattungen Sporobolus und Heteropogon. Bei hoher Verfügbarkeit an Gräsern – etwa zur Regenzeit – frisst ein Tier große Mengen und ist wenig wählerisch, bei geringerer Verfügbarkeit oder schlechterer Qualität geht es eher selektiv vor. In der Serengeti bevorzugt der Kaffernbüffel in der Regenzeit weitgehend Grashalme, in der Trockenzeit, wenn die Gräser verschwinden, zieht er sich in Flussniederungen zurück, wo das Nahrungsangebot umfangreicher ist. In trockenen Landschaften wie der Sukkulent-Karoo des südlichen Afrikas, wo Gräser weitaus seltener sind, stellt weicheres Pflanzenmaterial einen größeren Teil der vertilgten Menge. Hier ergaben Untersuchungen, dass während der trockenen Jahresabschnitte der Anteil an weichen Pflanzen bis zu 33 % erreichen kann, während er in der feuchteren auf etwa 28 % absinkt. Die Tiere fressen dann unter anderem auch Akaziengewächse, Sternbüsche, Rautengewächse oder den Kap-Bleiwurz. In der grasreicheren Nama-Karoo konnten dagegen derartige Unterschiede nicht festgestellt werden, da Gräser hier 95 % der aufgenommenen Nahrung ausmachen. Allerdings konnte hier ein Wechsel an bevorzugten Pflanzen innerhalb der verschiedenen Jahreszeiten ausgemacht werden. So dominierten Sporobolus-Gräser in den feuchten Jahresabschnitten, Themada-Gräser dagegen in den trockenen. Täglich muss ein Tier rund 2,2 % seiner Körpermasse vertilgen, was in etwa 14 bis 15 kg entspricht. Da der Kaffernbüffel von Wasser abhängig ist, entfernt er sich selten weiter als 8 bis 20 km von offenen Wasserstellen. Bedeutend sind darüber hinaus Salzlecken, unter anderem auch von schwitzenden Artgenossen, oder das Fressen eisenhaltiger Substrate vor allem in großen Höhenlagen wie am Mount Kenya.

 

Die Paarung kann beim Kaffernbüffel prinzipiell das ganze Jahr über erfolgen. In Regionen mit ausgeprägten Jahreszeiten und somit einem qualitativ und quantitativ schwankenden Nahrungsangebot ist sie jedoch saisonal beschränkt. In der Serengeti werden Paarungen von November und zunehmend bis Juli beobachtet, in der südafrikanischen Provinz Limpopo sind sie von Regenfällen und dem jeweiligen Nahrungsangebot abhängig. Letzteres lässt darauf schließen, dass die körperliche Kondition der weiblichen Tiere ausschlaggebend für die Paarung und Geburt ist. Kühe gebären ihr erstes Kalb mit 4 bis 5 Jahren, Bullen sind mit 8 bis 9 Jahren voll geschlechtsreif. Die männlichen Tiere sind in ein hierarchisches System eingebunden, welches die Fortpflanzung regelt. Dazu führen die männlichen Tiere regelmäßig Paarungs- und Dominanzwettkämpfe durch, die unter anderem aus Kopf- und Körperrammen bestehen; ersteres findet häufig zwischen dominanten und unterlegenen, letzteres zwischen gleichrangigen Tieren statt. Sie beginnen damit, dass ein Tier mit erhobenem Kopf und nach unten weisender Nase stillsteht, vergleichbar dem Dominanzverhalten. Oftmals wird der Kopf dabei auch ab- und aufwärts geführt. Die ganze Prozedur kann schon in 30 m Entfernung voneinander starten und ist teilweise mit einem tiefen Grummeln verbunden. Danach kommt es zum Angriff, wobei das Tier den Kopf so weit absenkt, dass beim Aufeinandertreffen die Energie auf die verstärkte Basis der Hörner gelenkt wird. Der Gewinner wird durch die Kraft und Geschwindigkeit bestimmt, der Verlierer sucht im Abschluss zumeist das Weite, wird manchmal aber vom Sieger bis zu 100 m verfolgt.

 

Bullen testen die Empfangsbereitschaft der Kühe beständig durch Beschnuppern der Geschlechtsorgane oder des Urins und zeigen dabei ein Flehmen. Der Sexualzyklus der Kühe dauert etwa 18 bis 22 Tage, der Östrus selbst einen Tag. Paarungswillige Bullen legen ihr Kinn auf den Rumpf der Kuh und signalisieren so Bereitschaft. Die Kuh wiederum erwidert ihre Bereitschaft durch Stillstehen und Schwanzbewegung. Die Kopulation währt eher kurz und wird innerhalb von 30 Minuten mehrfach wiederholt. Die Tragezeit der Kühe dauert bis zu 11,5 Monate (durchschnittlich 340 Tage), das Geburtsintervall beträgt 15 bis 24 Monate. Meist kommt nur ein Kalb zur Welt, das 39 bis 41 kg schwer ist; Zwillingsgeburten sind sehr selten. Das Geburtsgewicht schwankt leicht mit der Jahreszeit. Das Fell der Jungtiere ist schwarz oder dunkelgrau und ändert teilweise während des Wachstums die Farbe. Das Kalb benötigt mehrere Stunden nach der Geburt, um sicher zu stehen und bewegt sich in den folgenden Wochen eher langsam und behäbig fort. Mutter und Jungtier sind nach der Geburt etwas getrennt von der Herde, in dieser Zeit achtet das Muttertier sorgfältig auf das Junge. Insgesamt ist die Herde sehr aufmerksam auf Jungtiere, so dass eine Erbeutung durch Beutegreifer selten erfolgt. Das Junge saugt unregelmäßig, anfänglich bis zu 10 Minuten lang. Nach rund vier Wochen wird es etwa 5 Minuten lang gestillt. Während der Wachstumsphase spielen oder tummeln Jungtiere kaum, jungadulte Bullen führen aber Sparringskämpfe durch. Die Stillzeit endet unterschiedlich, abhängig von der Dichte der Population. Am Lake Manyara mit einer hohen Individuenzahl werden Jungtiere nach anderthalb Jahren entwöhnt, in der weniger dicht besiedelten Serengeti schon nach zehn Monaten, wenn das Muttertier im siebten Monat trächtig mit neuem Nachwuchs ist. Männliche Tiere verlassen die mütterliche Herde, weibliche verbleiben dort. Durchschnittlich liegt die Lebenserwartung bei 18 bis 20 Jahren in freier Wildbahn, einzelne Tiere in menschlicher Gefangenschaft wurden über 30 Jahre alt.

 

Den bedeutendsten Fressfeind stellt der Löwe dar. Aufgrund des guten Augenlichts kann der Kaffernbüffel Löwen schon aus bis zu 1,5 km Entfernung wahrnehmen. Einzelne Tiere verteidigen sich gegen Löwenangriffe, indem sie sich mit dem Rücken zu einem Hindernis positionieren oder im Gebüsch verstecken, so dass die Beutegreifer nicht die verletzlichen Hinterbeine erreichen können. Auch Flucht ist möglich, da individuelle Tiere Geschwindigkeiten von bis zu 55 km/h über kurze Distanzen erreichen. Gruppen stellen sich häufig im Kreis auf und präsentieren eine Phalanx von Hörnern. Unter Umständen greift der Kaffernbüffel auch Löwen direkt an, vertreibt sie, wirbelt sie mit den Hörnern durch die Luft oder trampelt sie mitunter zu Tode. Die Raubkatzen können einen ausgewachsenen Büffel nur durch einen Kehlbiss töten, da sie nicht die Kraft aufbringen, das Genick zu brechen. Außerdem wirkt die Haut des Kaffernbüffels mit 2 bis 3 cm Dicke schützend. Der Einfluss des Löwen auf die lokalen Bestände des Kaffernbüffels ist unterschiedlich. In der Serengeti verursacht er nur 25 % der Todesfälle beim Kaffernbüffel, am Lake Manyara sind es bis zu 85 %. In der Regel fallen dem Beutegreifer ältere, einzeln wandernde Bullen zum Opfer, selten in Junggesellengruppen lebende Bullen oder Kühe und Jungtiere in Herden. Bemerkenswert in diesem Zusammenhang ist, dass in Gebieten mit größerer Löwenpopulation auch größere Herden des Kaffernbüffels zu beobachten sind. Jungtiere werden gelegentlich auch durch die Tüpfelhyäne und den Leoparden gerissen.

 

Zu den wichtigsten Krankheiten, die den Kaffernbüffel befallen, gehören die Rinderpest, die Lungenseuche und die Tuberkulose, die überwiegend von Hausrindern auf die Wildtiere übertragen werden. Untergeordnet spielen auch der Milzbrand und die Maul- und Klauenseuche eine Rolle. Zudem sind die Tiere als Reservoir des Küstenfiebers bekannt. Gegen einige originär tropische oder afrikanische Erkrankungen wie die Schlafkrankheit, die durch die Tsetse-Fliege übertragen wird, ist der Kaffernbüffel immun. An äußeren Parasiten sind zahlreiche Zecken belegt, etwa die Gattungen Amblyomma, Rhipicephalus, Hyalomma und Boophilus. Einige dieser Ectoparasiten gelten auch als Überträger der bereits erwähnten Krankheiten. In der Regel leiden ältere Bullen stärker unter dem Befall als jüngere Tiere. Zur Linderung werden Vögel wie der Madenhacker geduldet, der die äußeren Parasiten entfernt.

 

Der Bestand des Kaffernbüffels (und anderer Huftiere) war in der Wende vom 19. zum 20. Jahrhundert stark reduziert worden. Eine Hauptursache stellte eine Rinderpest-Epidemie um 1890 dar, die Krankheit hatte sich ab dem Beginn des 19. Jahrhunderts im Zuge der zunehmenden Besiedlung des südlichen Afrikas durch europäische Einwanderer mit ihrem Hausvieh über den Kontinent ausgebreitet. In Verbindung mit der Lungenseuche führte dies zu einer Mortalitätsrate innerhalb der einzelnen Populationen von etwa 95 %. Auch im Verlauf des 20. Jahrhunderts waren die Rinderpest und zusätzlich der Milzbrand für den Rückgang lokaler Bestände verantwortlich. Hinzu kommt ein hoher Jagddruck, da der Kaffernbüffel in zahlreichen Ländern als Nahrungsressource gilt. Die Fleischnutzung erfolgt aber nicht nur zur Eigenversorgung in der ländlichen Bevölkerung, sondern wird auch im industriellen Maßstab getätigt. Darüber hinaus gilt der Kaffernbüffel als Bestandteil der Big Five und hat den Ruf als eines der gefährlichsten Tiere der afrikanischen Savannen, womit die Trophäenjagd eine hohe Bedeutung erlangt. Nicht zuletzt hat die Zerstörung des Lebensraums einen großen Einfluss auf die einzelnen Bestände, vor allem im Randbereich von Schutzgebieten, wo es zur Konkurrenz mit Nutzvieh kommt. Heute sind die Populationen und Herden des Kaffernbüffels zum Großteil auf Schutzgebiete beschränkt. Da die Art gegenwärtig weit verbreitet ist, wird sie von der IUCN als „ungefährdet“ (Least Concern) eingestuft. Ihr Gesamtbestand wird auf etwa 670.000 Individuen geschätzt. Zu den bedeutendsten Schutzgebieten, in denen der Kaffernbüffel auftritt, gehören die Serengeti und der Lake-Manyara-Nationalpark in Tansania, der Chobe-Nationalpark in Botswana und der Hluhluwe-iMfolozi-Nationalpark sowie der Krüger-Nationalpark in Südafrika.

 

(Wikipedia)

Porcupines are amongst the largest of the rodents weighing between 24 to 40 lb. (11 to 18 kg) and measuring from nose to tail between 31 to 39 in. (78 to 100 cm). As with all members of this family their incisor teeth must be in regular use to keep them in trim and prevent them from over growing.

The most distinctive feature of this mammal is the converted body hair, most of which is seen as a protective covering of sharp ended spines. These may be raised or lowered at will, both for body protection and as a means of communication. An angry or disturbed porcupine will raise it's spines and even rattle them as a means of warning to a would be aggressor. Predators of the Indian crested porcupine include large cats, caracals, wolves, striped hyenas, Asian wild dogs, saltwater crocodiles and humans.

Indian crested porcupines are found throughout southwest and central Asia. Due to their flexible environmental tolerances, Indian crested porcupines occupy a broad range of habitats. They prefer rocky hillsides, but are also common in tropical and temperate shrublands, grasslands, forests, plantations, and gardens. Their range seems to be limited by seasonal densities of forage and the availability of suitable substrates for digging burrows. They do not occur above latitudes where minimum night duration is less than 7 hours, presumably because of the amount of foraging time required to find food. They consume a variety of natural and agricultural plant material, including roots, bulbs, fruits, grains, drupe and tubers, along with insects and small vertebrates.

The Indian crested porcupine's conservation status is LC (least concern).

The Pokhot live in the Baringo and Western Pokot districts of Kenya and in Uganda.

There are two main sub-groups depending of their location and way of life. The first group consist of the Hill Pokot who live in the rainy highlands in the west and in the central south, and are mainly farmers and pastoralists. The second group is made up of the Plains Pokot who live in dry and infertile plains, with their cattles. A homestead is composed of one or more buildings for a man, his wife and children; eventual co-wives live in separate houses. The role of the community in teaching children ethical rules. Most of the Pokot are nomadic and thus have interacted with different peoples, incorporating their social customs.The Pokot are very proud of their culture. The Songs, storytelling, and decorative arts, especially bodily decoration, are very appreciated. They adorn the body with beads, hairstyling, scarification, and the removal of the lower central incisors. Pokot girls wear a beaded necklace made of the stems of an asparagus tree. Most Pokot have some knowledge of herbal medicine, so they often use these treatments along with those of the hospitals. They belong to the Kenya's Nilotic-speaking peoples. .

For the Pokot, the universe has two realms: the above is the realm of the most powerful deities—Tororot, Asis (sun), and llat (rain); and the below is the one where live humans, animals, and plants. Humans are responsible for the realm that they inhabit, but they rely upon divinities to achieve and maintain peace and prosperity. They worship many deities like the sun, moon and believe in the spirit of death.The Pokot communicate with their deities through prayer and sacrifice. They perform it during ethnic festivals and dances. Oracles are responsible for maintaining the spiritual balance within the community. They are superstitious and believe in sorcery, so sometimes they call on shielding lucky sorcery. They have prophets, either male or female, who foresee advise, usually by the means of animal sacrifices. His or her ability is considered as a divine gift. Clan histories recount the changes of location, through poetry and song, emphasizing the vulnerability of humans and the importance of supernatural powers to help them overcome hunger, thirst, and even death. Ceremonies mark the transitions in the people's social lives. Among these are: the cleansing of a couple expecting their first child; the cleansing of newborn infants and their mothers; the cleansing of twins and other children who are born under unusual circumstances; male and female initiation; marriage; sapana, a coming-of-age ceremony for men; and summer-solstice, harvest, and healing ceremonies. The most important rite of passage for most Pokot is circumcision for boys and clitoridectomy for girls. These rites consist of a series of neighborhood-based ceremonies, emphasizing the importance of having a good behavior. When boys are circumcised, they acquire membership in one of eight age sets. Women do not have age-sets. After excisions, for several months, girls have a white painting on their face and wear a hood made of blackened leather with charcoal and oil. This means they are untouchable until the lepan ceremony, that marks the passage to womanhood. Unlike other tribes, the Pokot keep the affiliation to their clan throughout their lives, there is no disruption with marriage. Surprisingly, the agreement before marriage is made by gift giving, from the groom and his family to the bride and her family, often over a period of years (and not the contrary). It often implies the gift of a combination of livestock, goods, and cash to the bride's family, and the allotment of milk cows and rights to land to the bride. The bond between a husband and wife lasts for 3 generations, after what marriages can take place again between the two groups. Polygamy exists but is not prevalent among men before 40. The spirits of the elder anticipate reincarnation in their living descendants: when a child is said to resemble the elder, the same name is given. Disputes are resolved in neighborhood councils and in government courts. Some of the sanctions include shaming, cursing, and bewitching.

 

© Eric Lafforgue

www.ericlafforgue.com

     

Fish, any of approximately 34,000 species of vertebrate animals (phylum Chordata) found in the fresh and salt waters of the world. Living species range from the primitive jawless lampreys and hagfishes through the cartilaginous sharks, skates, and rays to the abundant and diverse bony fishes. Most fish species are cold-blooded; however, one species, the opah (Lampris guttatus), is warm-blooded.

 

The term fish is applied to a variety of vertebrates of several evolutionary lines. It describes a life-form rather than a taxonomic group. As members of the phylum Chordata, fish share certain features with other vertebrates. These features are gill slits at some point in the life cycle, a notochord, or skeletal supporting rod, a dorsal hollow nerve cord, and a tail. Living fishes represent some five classes, which are as distinct from one another as are the four classes of familiar air-breathing animals—amphibians, reptiles, birds, and mammals. For example, the jawless fishes (Agnatha) have gills in pouches and lack limb girdles. Extant agnathans are the lampreys and the hagfishes. As the name implies, the skeletons of fishes of the class Chondrichthyes (from chondr, “cartilage,” and ichthyes, “fish”) are made entirely of cartilage. Modern fish of this class lack a swim bladder, and their scales and teeth are made up of the same placoid material. Sharks, skates, and rays are examples of cartilaginous fishes. The bony fishes are by far the largest class. Examples range from the tiny seahorse to the 450-kg (1,000-pound) blue marlin, from the flattened soles and flounders to the boxy puffers and ocean sunfishes. Unlike the scales of the cartilaginous fishes, those of bony fishes, when present, grow throughout life and are made up of thin overlapping plates of bone. Bony fishes also have an operculum that covers the gill slits.

 

The study of fishes, the science of ichthyology, is of broad importance. Fishes are of interest to humans for many reasons, the most important being their relationship with and dependence on the environment. A more obvious reason for interest in fishes is their role as a moderate but important part of the world’s food supply. This resource, once thought unlimited, is now realized to be finite and in delicate balance with the biological, chemical, and physical factors of the aquatic environment. Overfishing, pollution, and alteration of the environment are the chief enemies of proper fisheries management, both in fresh waters and in the ocean. (For a detailed discussion of the technology and economics of fisheries, see commercial fishing.) Another practical reason for studying fishes is their use in disease control. As predators on mosquito larvae, they help curb malaria and other mosquito-borne diseases.

 

Fishes are valuable laboratory animals in many aspects of medical and biological research. For example, the readiness of many fishes to acclimate to captivity has allowed biologists to study behaviour, physiology, and even ecology under relatively natural conditions. Fishes have been especially important in the study of animal behaviour, where research on fishes has provided a broad base for the understanding of the more flexible behaviour of the higher vertebrates. The zebra fish is used as a model in studies of gene expression.

 

There are aesthetic and recreational reasons for an interest in fishes. Millions of people keep live fishes in home aquariums for the simple pleasure of observing the beauty and behaviour of animals otherwise unfamiliar to them. Aquarium fishes provide a personal challenge to many aquarists, allowing them to test their ability to keep a small section of the natural environment in their homes. Sportfishing is another way of enjoying the natural environment, also indulged in by millions of people every year. Interest in aquarium fishes and sportfishing supports multimillion-dollar industries throughout the world.

 

Fishes have been in existence for more than 450 million years, during which time they have evolved repeatedly to fit into almost every conceivable type of aquatic habitat. In a sense, land vertebrates are simply highly modified fishes: when fishes colonized the land habitat, they became tetrapod (four-legged) land vertebrates. The popular conception of a fish as a slippery, streamlined aquatic animal that possesses fins and breathes by gills applies to many fishes, but far more fishes deviate from that conception than conform to it. For example, the body is elongate in many forms and greatly shortened in others; the body is flattened in some (principally in bottom-dwelling fishes) and laterally compressed in many others; the fins may be elaborately extended, forming intricate shapes, or they may be reduced or even lost; and the positions of the mouth, eyes, nostrils, and gill openings vary widely. Air breathers have appeared in several evolutionary lines.

 

Many fishes are cryptically coloured and shaped, closely matching their respective environments; others are among the most brilliantly coloured of all organisms, with a wide range of hues, often of striking intensity, on a single individual. The brilliance of pigments may be enhanced by the surface structure of the fish, so that it almost seems to glow. A number of unrelated fishes have actual light-producing organs. Many fishes are able to alter their coloration—some for the purpose of camouflage, others for the enhancement of behavioral signals.

 

Fishes range in adult length from less than 10 mm (0.4 inch) to more than 20 metres (60 feet) and in weight from about 1.5 grams (less than 0.06 ounce) to many thousands of kilograms. Some live in shallow thermal springs at temperatures slightly above 42 °C (100 °F), others in cold Arctic seas a few degrees below 0 °C (32 °F) or in cold deep waters more than 4,000 metres (13,100 feet) beneath the ocean surface. The structural and, especially, the physiological adaptations for life at such extremes are relatively poorly known and provide the scientifically curious with great incentive for study.

 

Almost all natural bodies of water bear fish life, the exceptions being very hot thermal ponds and extremely salt-alkaline lakes, such as the Dead Sea in Asia and the Great Salt Lake in North America. The present distribution of fishes is a result of the geological history and development of Earth as well as the ability of fishes to undergo evolutionary change and to adapt to the available habitats. Fishes may be seen to be distributed according to habitat and according to geographical area. Major habitat differences are marine and freshwater. For the most part, the fishes in a marine habitat differ from those in a freshwater habitat, even in adjacent areas, but some, such as the salmon, migrate from one to the other. The freshwater habitats may be seen to be of many kinds. Fishes found in mountain torrents, Arctic lakes, tropical lakes, temperate streams, and tropical rivers will all differ from each other, both in obvious gross structure and in physiological attributes. Even in closely adjacent habitats where, for example, a tropical mountain torrent enters a lowland stream, the fish fauna will differ. The marine habitats can be divided into deep ocean floors (benthic), mid-water oceanic (bathypelagic), surface oceanic (pelagic), rocky coast, sandy coast, muddy shores, bays, estuaries, and others. Also, for example, rocky coastal shores in tropical and temperate regions will have different fish faunas, even when such habitats occur along the same coastline.

 

Although much is known about the present geographical distribution of fishes, far less is known about how that distribution came about. Many parts of the fish fauna of the fresh waters of North America and Eurasia are related and undoubtedly have a common origin. The faunas of Africa and South America are related, extremely old, and probably an expression of the drifting apart of the two continents. The fauna of southern Asia is related to that of Central Asia, and some of it appears to have entered Africa. The extremely large shore-fish faunas of the Indian and tropical Pacific oceans comprise a related complex, but the tropical shore fauna of the Atlantic, although containing Indo-Pacific components, is relatively limited and probably younger. The Arctic and Antarctic marine faunas are quite different from each other. The shore fauna of the North Pacific is quite distinct, and that of the North Atlantic more limited and probably younger. Pelagic oceanic fishes, especially those in deep waters, are similar the world over, showing little geographical isolation in terms of family groups. The deep oceanic habitat is very much the same throughout the world, but species differences do exist, showing geographical areas determined by oceanic currents and water masses.

 

All aspects of the life of a fish are closely correlated with adaptation to the total environment, physical, chemical, and biological. In studies, all the interdependent aspects of fish, such as behaviour, locomotion, reproduction, and physical and physiological characteristics, must be taken into account.

 

Correlated with their adaptation to an extremely wide variety of habitats is the extremely wide variety of life cycles that fishes display. The great majority hatch from relatively small eggs a few days to several weeks or more after the eggs are scattered in the water. Newly hatched young are still partially undeveloped and are called larvae until body structures such as fins, skeleton, and some organs are fully formed. Larval life is often very short, usually less than a few weeks, but it can be very long, some lampreys continuing as larvae for at least five years. Young and larval fishes, before reaching sexual maturity, must grow considerably, and their small size and other factors often dictate that they live in a habitat different than that of the adults. For example, most tropical marine shore fishes have pelagic larvae. Larval food also is different, and larval fishes often live in shallow waters, where they may be less exposed to predators.

 

After a fish reaches adult size, the length of its life is subject to many factors, such as innate rates of aging, predation pressure, and the nature of the local climate. The longevity of a species in the protected environment of an aquarium may have nothing to do with how long members of that species live in the wild. Many small fishes live only one to three years at the most. In some species, however, individuals may live as long as 10 or 20 or even 100 years.

 

Fish behaviour is a complicated and varied subject. As in almost all animals with a central nervous system, the nature of a response of an individual fish to stimuli from its environment depends upon the inherited characteristics of its nervous system, on what it has learned from past experience, and on the nature of the stimuli. Compared with the variety of human responses, however, that of a fish is stereotyped, not subject to much modification by “thought” or learning, and investigators must guard against anthropomorphic interpretations of fish behaviour.

 

Fishes perceive the world around them by the usual senses of sight, smell, hearing, touch, and taste and by special lateral line water-current detectors. In the few fishes that generate electric fields, a process that might best be called electrolocation aids in perception. One or another of these senses often is emphasized at the expense of others, depending upon the fish’s other adaptations. In fishes with large eyes, the sense of smell may be reduced; others, with small eyes, hunt and feed primarily by smell (such as some eels).

 

Specialized behaviour is primarily concerned with the three most important activities in the fish’s life: feeding, reproduction, and escape from enemies. Schooling behaviour of sardines on the high seas, for instance, is largely a protective device to avoid enemies, but it is also associated with and modified by their breeding and feeding requirements. Predatory fishes are often solitary, lying in wait to dart suddenly after their prey, a kind of locomotion impossible for beaked parrot fishes, which feed on coral, swimming in small groups from one coral head to the next. In addition, some predatory fishes that inhabit pelagic environments, such as tunas, often school.

 

Sleep in fishes, all of which lack true eyelids, consists of a seemingly listless state in which the fish maintains its balance but moves slowly. If attacked or disturbed, most can dart away. A few kinds of fishes lie on the bottom to sleep. Most catfishes, some loaches, and some eels and electric fishes are strictly nocturnal, being active and hunting for food during the night and retiring during the day to holes, thick vegetation, or other protective parts of the environment.

 

Communication between members of a species or between members of two or more species often is extremely important, especially in breeding behaviour (see below Reproduction). The mode of communication may be visual, as between the small so-called cleaner fish and a large fish of a very different species. The larger fish often allows the cleaner to enter its mouth to remove gill parasites. The cleaner is recognized by its distinctive colour and actions and therefore is not eaten, even if the larger fish is normally a predator. Communication is often chemical, signals being sent by specific chemicals called pheromones.

 

Many fishes have a streamlined body and swim freely in open water. Fish locomotion is closely correlated with habitat and ecological niche (the general position of the animal to its environment).

 

Many fishes in both marine and fresh waters swim at the surface and have mouths adapted to feed best (and sometimes only) at the surface. Often such fishes are long and slender, able to dart at surface insects or at other surface fishes and in turn to dart away from predators; needlefishes, halfbeaks, and topminnows (such as killifish and mosquito fish) are good examples. Oceanic flying fishes escape their predators by gathering speed above the water surface, with the lower lobe of the tail providing thrust in the water. They then glide hundreds of yards on enlarged, winglike pectoral and pelvic fins. South American freshwater flying fishes escape their enemies by jumping and propelling their strongly keeled bodies out of the water.

 

So-called mid-water swimmers, the most common type of fish, are of many kinds and live in many habitats. The powerful fusiform tunas and the trouts, for example, are adapted for strong, fast swimming, the tunas to capture prey speedily in the open ocean and the trouts to cope with the swift currents of streams and rivers. The trout body form is well adapted to many habitats. Fishes that live in relatively quiet waters such as bays or lake shores or slow rivers usually are not strong, fast swimmers but are capable of short, quick bursts of speed to escape a predator. Many of these fishes have their sides flattened, examples being the sunfish and the freshwater angelfish of aquarists. Fish associated with the bottom or substrate usually are slow swimmers. Open-water plankton-feeding fishes almost always remain fusiform and are capable of rapid, strong movement (for example, sardines and herrings of the open ocean and also many small minnows of streams and lakes).

 

Bottom-living fishes are of many kinds and have undergone many types of modification of their body shape and swimming habits. Rays, which evolved from strong-swimming mid-water sharks, usually stay close to the bottom and move by undulating their large pectoral fins. Flounders live in a similar habitat and move over the bottom by undulating the entire body. Many bottom fishes dart from place to place, resting on the bottom between movements, a motion common in gobies. One goby relative, the mudskipper, has taken to living at the edge of pools along the shore of muddy mangrove swamps. It escapes its enemies by flipping rapidly over the mud, out of the water. Some catfishes, synbranchid eels, the so-called climbing perch, and a few other fishes venture out over damp ground to find more promising waters than those that they left. They move by wriggling their bodies, sometimes using strong pectoral fins; most have accessory air-breathing organs. Many bottom-dwelling fishes live in mud holes or rocky crevices. Marine eels and gobies commonly are found in such habitats and for the most part venture far beyond their cavelike homes. Some bottom dwellers, such as the clingfishes (Gobiesocidae), have developed powerful adhesive disks that enable them to remain in place on the substrate in areas such as rocky coasts, where the action of the waves is great.

 

The methods of reproduction in fishes are varied, but most fishes lay a large number of small eggs, fertilized and scattered outside of the body. The eggs of pelagic fishes usually remain suspended in the open water. Many shore and freshwater fishes lay eggs on the bottom or among plants. Some have adhesive eggs. The mortality of the young and especially of the eggs is very high, and often only a few individuals grow to maturity out of hundreds, thousands, and in some cases millions of eggs laid.

 

Males produce sperm, usually as a milky white substance called milt, in two (sometimes one) testes within the body cavity. In bony fishes a sperm duct leads from each testis to a urogenital opening behind the vent or anus. In sharks and rays and in cyclostomes the duct leads to a cloaca. Sometimes the pelvic fins are modified to help transmit the milt to the eggs at the female’s vent or on the substrate where the female has placed them. Sometimes accessory organs are used to fertilize females internally—for example, the claspers of many sharks and rays.

 

In the females the eggs are formed in two ovaries (sometimes only one) and pass through the ovaries to the urogenital opening and to the outside. In some fishes the eggs are fertilized internally but are shed before development takes place. Members of about a dozen families each of bony fishes (teleosts) and sharks bear live young. Many skates and rays also bear live young. In some bony fishes the eggs simply develop within the female, the young emerging when the eggs hatch (ovoviviparous). Others develop within the ovary and are nourished by ovarian tissues after hatching (viviparous). There are also other methods utilized by fishes to nourish young within the female. In all live-bearers the young are born at a relatively large size and are few in number. In one family of primarily marine fishes, the surfperches from the Pacific coast of North America, Japan, and Korea, the males of at least one species are born sexually mature, although they are not fully grown.

 

Some fishes are hermaphroditic—an individual producing both sperm and eggs, usually at different stages of its life. Self-fertilization, however, is probably rare.

 

Successful reproduction and, in many cases, defense of the eggs and the young are assured by rather stereotypical but often elaborate courtship and parental behaviour, either by the male or the female or both. Some fishes prepare nests by hollowing out depressions in the sand bottom (cichlids, for example), build nests with plant materials and sticky threads excreted by the kidneys (sticklebacks), or blow a cluster of mucus-covered bubbles at the water surface (gouramis). The eggs are laid in these structures. Some varieties of cichlids and catfishes incubate eggs in their mouths.

 

Some fishes, such as salmon, undergo long migrations from the ocean and up large rivers to spawn in the gravel beds where they themselves hatched (anadromous fishes). Some, such as the freshwater eels (family Anguillidae), live and grow to maturity in fresh water and migrate to the sea to spawn (catadromous fishes). Other fishes undertake shorter migrations from lakes into streams, within the ocean, or enter spawning habitats that they do not ordinarily occupy in other ways.

 

The basic structure and function of the fish body are similar to those of all other vertebrates. The usual four types of tissues are present: surface or epithelial, connective (bone, cartilage, and fibrous tissues, as well as their derivative, blood), nerve, and muscle tissues. In addition, the fish’s organs and organ systems parallel those of other vertebrates.

 

The typical fish body is streamlined and spindle-shaped, with an anterior head, a gill apparatus, and a heart, the latter lying in the midline just below the gill chamber. The body cavity, containing the vital organs, is situated behind the head in the lower anterior part of the body. The anus usually marks the posterior termination of the body cavity and most often occurs just in front of the base of the anal fin. The spinal cord and vertebral column continue from the posterior part of the head to the base of the tail fin, passing dorsal to the body cavity and through the caudal (tail) region behind the body cavity. Most of the body is of muscular tissue, a high proportion of which is necessitated by swimming. In the course of evolution this basic body plan has been modified repeatedly into the many varieties of fish shapes that exist today.

 

The skeleton forms an integral part of the fish’s locomotion system, as well as serving to protect vital parts. The internal skeleton consists of the skull bones (except for the roofing bones of the head, which are really part of the external skeleton), the vertebral column, and the fin supports (fin rays). The fin supports are derived from the external skeleton but will be treated here because of their close functional relationship to the internal skeleton. The internal skeleton of cyclostomes, sharks, and rays is of cartilage; that of many fossil groups and some primitive living fishes is mostly of cartilage but may include some bone. In place of the vertebral column, the earliest vertebrates had a fully developed notochord, a flexible stiff rod of viscous cells surrounded by a strong fibrous sheath. During the evolution of modern fishes the rod was replaced in part by cartilage and then by ossified cartilage. Sharks and rays retain a cartilaginous vertebral column; bony fishes have spool-shaped vertebrae that in the more primitive living forms only partially replace the notochord. The skull, including the gill arches and jaws of bony fishes, is fully, or at least partially, ossified. That of sharks and rays remains cartilaginous, at times partially replaced by calcium deposits but never by true bone.

 

The supportive elements of the fins (basal or radial bones or both) have changed greatly during fish evolution. Some of these changes are described in the section below (Evolution and paleontology). Most fishes possess a single dorsal fin on the midline of the back. Many have two and a few have three dorsal fins. The other fins are the single tail and anal fins and paired pelvic and pectoral fins. A small fin, the adipose fin, with hairlike fin rays, occurs in many of the relatively primitive teleosts (such as trout) on the back near the base of the caudal fin.

 

The skin of a fish must serve many functions. It aids in maintaining the osmotic balance, provides physical protection for the body, is the site of coloration, contains sensory receptors, and, in some fishes, functions in respiration. Mucous glands, which aid in maintaining the water balance and offer protection from bacteria, are extremely numerous in fish skin, especially in cyclostomes and teleosts. Since mucous glands are present in the modern lampreys, it is reasonable to assume that they were present in primitive fishes, such as the ancient Silurian and Devonian agnathans. Protection from abrasion and predation is another function of the fish skin, and dermal (skin) bone arose early in fish evolution in response to this need. It is thought that bone first evolved in skin and only later invaded the cartilaginous areas of the fish’s body, to provide additional support and protection. There is some argument as to which came first, cartilage or bone, and fossil evidence does not settle the question. In any event, dermal bone has played an important part in fish evolution and has different characteristics in different groups of fishes. Several groups are characterized at least in part by the kind of bony scales they possess.

 

Scales have played an important part in the evolution of fishes. Primitive fishes usually had thick bony plates or thick scales in several layers of bone, enamel, and related substances. Modern teleost fishes have scales of bone, which, while still protective, allow much more freedom of motion in the body. A few modern teleosts (some catfishes, sticklebacks, and others) have secondarily acquired bony plates in the skin. Modern and early sharks possessed placoid scales, a relatively primitive type of scale with a toothlike structure, consisting of an outside layer of enamel-like substance (vitrodentine), an inner layer of dentine, and a pulp cavity containing nerves and blood vessels. Primitive bony fishes had thick scales of either the ganoid or the cosmoid type. Cosmoid scales have a hard, enamel-like outer layer, an inner layer of cosmine (a form of dentine), and then a layer of vascular bone (isopedine). In ganoid scales the hard outer layer is different chemically and is called ganoin. Under this is a cosminelike layer and then a vascular bony layer. The thin, translucent bony scales of modern fishes, called cycloid and ctenoid (the latter distinguished by serrations at the edges), lack enameloid and dentine layers.

 

Skin has several other functions in fishes. It is well supplied with nerve endings and presumably receives tactile, thermal, and pain stimuli. Skin is also well supplied with blood vessels. Some fishes breathe in part through the skin, by the exchange of oxygen and carbon dioxide between the surrounding water and numerous small blood vessels near the skin surface.

 

Skin serves as protection through the control of coloration. Fishes exhibit an almost limitless range of colours. The colours often blend closely with the surroundings, effectively hiding the animal. Many fishes use bright colours for territorial advertisement or as recognition marks for other members of their own species, or sometimes for members of other species. Many fishes can change their colour to a greater or lesser degree, by movement of pigment within the pigment cells (chromatophores). Black pigment cells (melanophores), of almost universal occurrence in fishes, are often juxtaposed with other pigment cells. When placed beneath iridocytes or leucophores (bearing the silvery or white pigment guanine), melanophores produce structural colours of blue and green. These colours are often extremely intense, because they are formed by refraction of light through the needlelike crystals of guanine. The blue and green refracted colours are often relatively pure, lacking the red and yellow rays, which have been absorbed by the black pigment (melanin) of the melanophores. Yellow, orange, and red colours are produced by erythrophores, cells containing the appropriate carotenoid pigments. Other colours are produced by combinations of melanophores, erythrophores, and iridocytes.

 

The major portion of the body of most fishes consists of muscles. Most of the mass is trunk musculature, the fin muscles usually being relatively small. The caudal fin is usually the most powerful fin, being moved by the trunk musculature. The body musculature is usually arranged in rows of chevron-shaped segments on each side. Contractions of these segments, each attached to adjacent vertebrae and vertebral processes, bends the body on the vertebral joint, producing successive undulations of the body, passing from the head to the tail, and producing driving strokes of the tail. It is the latter that provides the strong forward movement for most fishes.

 

The digestive system, in a functional sense, starts at the mouth, with the teeth used to capture prey or collect plant foods. Mouth shape and tooth structure vary greatly in fishes, depending on the kind of food normally eaten. Most fishes are predacious, feeding on small invertebrates or other fishes and have simple conical teeth on the jaws, on at least some of the bones of the roof of the mouth, and on special gill arch structures just in front of the esophagus. The latter are throat teeth. Most predacious fishes swallow their prey whole, and the teeth are used for grasping and holding prey, for orienting prey to be swallowed (head first) and for working the prey toward the esophagus. There are a variety of tooth types in fishes. Some fishes, such as sharks and piranhas, have cutting teeth for biting chunks out of their victims. A shark’s tooth, although superficially like that of a piranha, appears in many respects to be a modified scale, while that of the piranha is like that of other bony fishes, consisting of dentine and enamel. Parrot fishes have beaklike mouths with short incisor-like teeth for breaking off coral and have heavy pavementlike throat teeth for crushing the coral. Some catfishes have small brushlike teeth, arranged in rows on the jaws, for scraping plant and animal growth from rocks. Many fishes (such as the Cyprinidae or minnows) have no jaw teeth at all but have very strong throat teeth.

 

Some fishes gather planktonic food by straining it from their gill cavities with numerous elongate stiff rods (gill rakers) anchored by one end to the gill bars. The food collected on these rods is passed to the throat, where it is swallowed. Most fishes have only short gill rakers that help keep food particles from escaping out the mouth cavity into the gill chamber.

 

Once reaching the throat, food enters a short, often greatly distensible esophagus, a simple tube with a muscular wall leading into a stomach. The stomach varies greatly in fishes, depending upon the diet. In most predacious fishes it is a simple straight or curved tube or pouch with a muscular wall and a glandular lining. Food is largely digested there and leaves the stomach in liquid form.

 

Between the stomach and the intestine, ducts enter the digestive tube from the liver and pancreas. The liver is a large, clearly defined organ. The pancreas may be embedded in it, diffused through it, or broken into small parts spread along some of the intestine. The junction between the stomach and the intestine is marked by a muscular valve. Pyloric ceca (blind sacs) occur in some fishes at this junction and have a digestive or absorptive function or both.

 

The intestine itself is quite variable in length, depending upon the fish’s diet. It is short in predacious forms, sometimes no longer than the body cavity, but long in herbivorous forms, being coiled and several times longer than the entire length of the fish in some species of South American catfishes. The intestine is primarily an organ for absorbing nutrients into the bloodstream. The larger its internal surface, the greater its absorptive efficiency, and a spiral valve is one method of increasing its absorption surface.

 

Sharks, rays, chimaeras, lungfishes, surviving chondrosteans, holosteans, and even a few of the more primitive teleosts have a spiral valve or at least traces of it in the intestine. Most modern teleosts have increased the area of the intestinal walls by having numerous folds and villi (fingerlike projections) somewhat like those in humans. Undigested substances are passed to the exterior through the anus in most teleost fishes. In lungfishes, sharks, and rays, it is first passed through the cloaca, a common cavity receiving the intestinal opening and the ducts from the urogenital system.

 

Oxygen and carbon dioxide dissolve in water, and most fishes exchange dissolved oxygen and carbon dioxide in water by means of the gills. The gills lie behind and to the side of the mouth cavity and consist of fleshy filaments supported by the gill arches and filled with blood vessels, which give gills a bright red colour. Water taken in continuously through the mouth passes backward between the gill bars and over the gill filaments, where the exchange of gases takes place. The gills are protected by a gill cover in teleosts and many other fishes but by flaps of skin in sharks, rays, and some of the older fossil fish groups. The blood capillaries in the gill filaments are close to the gill surface to take up oxygen from the water and to give up excess carbon dioxide to the water.

 

Most modern fishes have a hydrostatic (ballast) organ, called the swim bladder, that lies in the body cavity just below the kidney and above the stomach and intestine. It originated as a diverticulum of the digestive canal. In advanced teleosts, especially the acanthopterygians, the bladder has lost its connection with the digestive tract, a condition called physoclistic. The connection has been retained (physostomous) by many relatively primitive teleosts. In several unrelated lines of fishes, the bladder has become specialized as a lung or, at least, as a highly vascularized accessory breathing organ. Some fishes with such accessory organs are obligate air breathers and will drown if denied access to the surface, even in well-oxygenated water. Fishes with a hydrostatic form of swim bladder can control their depth by regulating the amount of gas in the bladder. The gas, mostly oxygen, is secreted into the bladder by special glands, rendering the fish more buoyant; the gas is absorbed into the bloodstream by another special organ, reducing the overall buoyancy and allowing the fish to sink. Some deep-sea fishes may have oils, rather than gas, in the bladder. Other deep-sea and some bottom-living forms have much-reduced swim bladders or have lost the organ entirely.

 

The swim bladder of fishes follows the same developmental pattern as the lungs of land vertebrates. There is no doubt that the two structures have the same historical origin in primitive fishes. More or less intermediate forms still survive among the more primitive types of fishes, such as the lungfishes Lepidosiren and Protopterus.

 

The circulatory, or blood vascular, system consists of the heart, the arteries, the capillaries, and the veins. It is in the capillaries that the interchange of oxygen, carbon dioxide, nutrients, and other substances such as hormones and waste products takes place. The capillaries lead to the veins, which return the venous blood with its waste products to the heart, kidneys, and gills. There are two kinds of capillary beds: those in the gills and those in the rest of the body. The heart, a folded continuous muscular tube with three or four saclike enlargements, undergoes rhythmic contractions and receives venous blood in a sinus venosus. It passes the blood to an auricle and then into a thick muscular pump, the ventricle. From the ventricle the blood goes to a bulbous structure at the base of a ventral aorta just below the gills. The blood passes to the afferent (receiving) arteries of the gill arches and then to the gill capillaries. There waste gases are given off to the environment, and oxygen is absorbed. The oxygenated blood enters efferent (exuant) arteries of the gill arches and then flows into the dorsal aorta. From there blood is distributed to the tissues and organs of the body. One-way valves prevent backflow. The circulation of fishes thus differs from that of the reptiles, birds, and mammals in that oxygenated blood is not returned to the heart prior to distribution to the other parts of the body.

 

The primary excretory organ in fishes, as in other vertebrates, is the kidney. In fishes some excretion also takes place in the digestive tract, skin, and especially the gills (where ammonia is given off). Compared with land vertebrates, fishes have a special problem in maintaining their internal environment at a constant concentration of water and dissolved substances, such as salts. Proper balance of the internal environment (homeostasis) of a fish is in a great part maintained by the excretory system, especially the kidney.

 

The kidney, gills, and skin play an important role in maintaining a fish’s internal environment and checking the effects of osmosis. Marine fishes live in an environment in which the water around them has a greater concentration of salts than they can have inside their body and still maintain life. Freshwater fishes, on the other hand, live in water with a much lower concentration of salts than they require inside their bodies. Osmosis tends to promote the loss of water from the body of a marine fish and absorption of water by that of a freshwater fish. Mucus in the skin tends to slow the process but is not a sufficient barrier to prevent the movement of fluids through the permeable skin. When solutions on two sides of a permeable membrane have different concentrations of dissolved substances, water will pass through the membrane into the more concentrated solution, while the dissolved chemicals move into the area of lower concentration (diffusion).

 

The kidney of freshwater fishes is often larger in relation to body weight than that of marine fishes. In both groups the kidney excretes wastes from the body, but the kidney of freshwater fishes also excretes large amounts of water, counteracting the water absorbed through the skin. Freshwater fishes tend to lose salt to the environment and must replace it. They get some salt from their food, but the gills and skin inside the mouth actively absorb salt from water passed through the mouth. This absorption is performed by special cells capable of moving salts against the diffusion gradient. Freshwater fishes drink very little water and take in little water with their food.

 

Marine fishes must conserve water, and therefore their kidneys excrete little water. To maintain their water balance, marine fishes drink large quantities of seawater, retaining most of the water and excreting the salt. Most nitrogenous waste in marine fishes appears to be secreted by the gills as ammonia. Marine fishes can excrete salt by clusters of special cells (chloride cells) in the gills.

 

There are several teleosts—for example, the salmon—that travel between fresh water and seawater and must adjust to the reversal of osmotic gradients. They adjust their physiological processes by spending time (often surprisingly little time) in the intermediate brackish environment.

 

Marine hagfishes, sharks, and rays have osmotic concentrations in their blood about equal to that of seawater and so do not have to drink water nor perform much physiological work to maintain their osmotic balance. In sharks and rays the osmotic concentration is kept high by retention of urea in the blood. Freshwater sharks have a lowered concentration of urea in the blood.

 

Endocrine glands secrete their products into the bloodstream and body tissues and, along with the central nervous system, control and regulate many kinds of body functions. Cyclostomes have a well-developed endocrine system, and presumably it was well developed in the early Agnatha, ancestral to modern fishes. Although the endocrine system in fishes is similar to that of higher vertebrates, there are numerous differences in detail. The pituitary, the thyroid, the suprarenals, the adrenals, the pancreatic islets, the sex glands (ovaries and testes), the inner wall of the intestine, and the bodies of the ultimobranchial gland make up the endocrine system in fishes. There are some others whose function is not well understood. These organs regulate sexual activity and reproduction, growth, osmotic pressure, general metabolic activities such as the storage of fat and the utilization of foodstuffs, blood pressure, and certain aspects of skin colour. Many of these activities are also controlled in part by the central nervous system, which works with the endocrine system in maintaining the life of a fish. Some parts of the endocrine system are developmentally, and undoubtedly evolutionarily, derived from the nervous system.

 

As in all vertebrates, the nervous system of fishes is the primary mechanism coordinating body activities, as well as integrating these activities in the appropriate manner with stimuli from the environment. The central nervous system, consisting of the brain and spinal cord, is the primary integrating mechanism. The peripheral nervous system, consisting of nerves that connect the brain and spinal cord to various body organs, carries sensory information from special receptor organs such as the eyes, internal ears, nares (sense of smell), taste glands, and others to the integrating centres of the brain and spinal cord. The peripheral nervous system also carries information via different nerve cells from the integrating centres of the brain and spinal cord. This coded information is carried to the various organs and body systems, such as the skeletal muscular system, for appropriate action in response to the original external or internal stimulus. Another branch of the nervous system, the autonomic nervous system, helps to coordinate the activities of many glands and organs and is itself closely connected to the integrating centres of the brain.

 

The brain of the fish is divided into several anatomical and functional parts, all closely interconnected but each serving as the primary centre of integrating particular kinds of responses and activities. Several of these centres or parts are primarily associated with one type of sensory perception, such as sight, hearing, or smell (olfaction).

 

The sense of smell is important in almost all fishes. Certain eels with tiny eyes depend mostly on smell for location of food. The olfactory, or nasal, organ of fishes is located on the dorsal surface of the snout. The lining of the nasal organ has special sensory cells that perceive chemicals dissolved in the water, such as substances from food material, and send sensory information to the brain by way of the first cranial nerve. Odour also serves as an alarm system. Many fishes, especially various species of freshwater minnows, react with alarm to a chemical released from the skin of an injured member of their own species.

 

Many fishes have a well-developed sense of taste, and tiny pitlike taste buds or organs are located not only within their mouth cavities but also over their heads and parts of their body. Catfishes, which often have poor vision, have barbels (“whiskers”) that serve as supplementary taste organs, those around the mouth being actively used to search out food on the bottom. Some species of naturally blind cave fishes are especially well supplied with taste buds, which often cover most of their body surface.

 

Sight is extremely important in most fishes. The eye of a fish is basically like that of all other vertebrates, but the eyes of fishes are extremely varied in structure and adaptation. In general, fishes living in dark and dim water habitats have large eyes, unless they have specialized in some compensatory way so that another sense (such as smell) is dominant, in which case the eyes will often be reduced. Fishes living in brightly lighted shallow waters often will have relatively small but efficient eyes. Cyclostomes have somewhat less elaborate eyes than other fishes, with skin stretched over the eyeball perhaps making their vision somewhat less effective. Most fishes have a spherical lens and accommodate their vision to far or near subjects by moving the lens within the eyeball. A few sharks accommodate by changing the shape of the lens, as in land vertebrates. Those fishes that are heavily dependent upon the eyes have especially strong muscles for accommodation. Most fishes see well, despite the restrictions imposed by frequent turbidity of the water and by light refraction.

 

Fossil evidence suggests that colour vision evolved in fishes more than 300 million years ago, but not all living fishes have retained this ability. Experimental evidence indicates that many shallow-water fishes, if not all, have colour vision and see some colours especially well, but some bottom-dwelling shore fishes live in areas where the water is sufficiently deep to filter out most if not all colours, and these fishes apparently never see colours. When tested in shallow water, they apparently are unable to respond to colour differences.

 

Sound perception and balance are intimately associated senses in a fish. The organs of hearing are entirely internal, located within the skull, on each side of the brain and somewhat behind the eyes. Sound waves, especially those of low frequencies, travel readily through water and impinge directly upon the bones and fluids of the head and body, to be transmitted to the hearing organs. Fishes readily respond to sound; for example, a trout conditioned to escape by the approach of fishermen will take flight upon perceiving footsteps on a stream bank even if it cannot see a fisherman. Compared with humans, however, the range of sound frequencies heard by fishes is greatly restricted. Many fishes communicate with each other by producing sounds in their swim bladders, in their throats by rasping their teeth, and in other ways.

 

A fish or other vertebrate seldom has to rely on a single type of sensory information to determine the nature of the environment around it. A catfish uses taste and touch when examining a food object with its oral barbels. Like most other animals, fishes have many touch receptors over their body surface. Pain and temperature receptors also are present in fishes and presumably produce the same kind of information to a fish as to humans. Fishes react in a negative fashion to stimuli that would be painful to human beings, suggesting that they feel a sensation of pain.

 

An important sensory system in fishes that is absent in other vertebrates (except some amphibians) is the lateral line system. This consists of a series of heavily innervated small canals located in the skin and bone around the eyes, along the lower jaw, over the head, and down the mid-side of the body, where it is associated with the scales. Intermittently along these canals are located tiny sensory organs (pit organs) that apparently detect changes in pressure. The system allows a fish to sense changes in water currents and pressure, thereby helping the fish to orient itself to the various changes that occur in the physical environment.

  

Due to its remarkable adaptability to almost any environment, the mouse is one of the most successful mammalian genera living on Earth today.

All rodents share the characteristic of dentition highly specialised for gnawing. This specialisation gives rodents their name from the Latin, rodere, to gnaw.

Rodent incisors grow continuously and must be kept worn down by gnawing.

he wood mouse (Apodemus sylvaticus) is a common rodent from Europe.

This one was displaced when I constructed a new fence between my neighbor, as part of the job the heavy growth along the old fnce was cleared. this little mouse must have nested in the undergrowth.

Geoffrey’s marmoset is an exceptionally distinctive monkey, most readily recognised for its conspicuous white cheeks, forehead and throat, which contrast starkly against its elongate black ear-tufts, tan to black face, and dark coat. The body is greyish-black mottled with yellow-orange on the upperparts, brown on the underparts, and the long black tail is lightly ringed. Like all marmosets, Geoffroy’s marmoset has incisor teeth that are specially adapted to carving out small holes in the trunks of trees, through which they drink the sap and gum that oozes out ...

We like the Hindi name Bhalu, it is much more attractive for such an endearing bear than Sloth Bear, the name which was derived from the thought that they were related to the South American Sloths (wooly with long sickle claws and no upper middle incisors), which of course has persisted.

 

They have rather a depressing history of exploitation by man. Listed as Vulnerable in the IUCN Red List they now have legal protection: the government of India have banned their use for entertainment and international trade is banned.

 

This one sighted at Ranthambore, Rajastan, India

Addo Elephant National Park is a diverse wildlife conservation park situated close to Port Elizabeth in South Africa and is one of the country's 19 national parks. It currently ranks third in size after Kruger National Park and the Kgalagadi Transfrontier Park.

 

The original section of the park was founded in 1931, in part due to the efforts of Sydney Skaife, in order to provide a sanctuary for the eleven remaining elephants in the area. The park has proved to be very successful and currently houses more than 600 elephants and a large number of other mammals.

 

The original park has subsequently been expanded to include the Woody Cape Nature Reserve that extends from the Sundays River mouth towards Alexandria and a marine reserve, which includes St. Croix Island and Bird Island, both important breeding habitat for gannets and penguins, not to mention a large variety of other marine life. Bird Island is home to the world's largest breeding colony of gannets - about 120,000 birds - and also hosts the second largest breeding colony of African penguins, the largest breeding colony being St. Croix island. These marine assets form part of the plan to expand the 1,640 km² Addo National Elephant Park into the 3,600 km² Greater Addo Elephant National Park.

 

The expansion will mean not only that the park contains five of South Africa's seven major vegetation zones (biomes), but also that it will be the only park in the world to house Africa's "Big 7" (elephant, rhinoceros, lion, buffalo, leopard, whale and great white shark) in their natural habitat.

 

More than 600 elephants, 400 Cape buffaloes, over 48 endangered black rhinos as well as a variety of antelope species. Transvaal lion and spotted hyena have also recently been re-introduced to the area. The largest remaining population of the flightless dung beetle (Circellium bacchus) is located within the park. The flora within the AENP is quite varied, and like all plant life, is a central factor to the ecological system in place. Several species of rare and endemic plants, particularly succulent shrubs and geophytes are native to the South African region within the AENP. Many species are under environmental pressure, however, and are facing possible extinction.

 

(Wikipedia)

 

Name

 

African Elephant or African Bush Elephant [Loxodonta africana]

 

Introduction

 

The Elephant is the world's largest land mammal, and weighs up to 7 tonnes and reaches heights of 3.3 m at the shoulder. Elephants can live to a potential age of 70 years. The massive tusks of older bulls can weigh up to 50 or 60 kilograms, but tusks weighing up to 90 kilograms have been recorded.

 

Appearance

 

What is the trunk and what is it used for?

The Elephant's trunk is a modified nose which is very sensitive and can even detect water under ground. There are as many as 50 000 muscles in an Elephant trunk. The sensitive finger-like appendages at the tip of the trunk enables them to pick the smallest twig or flower, pull the toughest reed of grass or even pick out a thorn from their feet.

 

Do elephants have knees or elbows?

 

The joints that are perceived as 'knees', are in fact wrists. This is a common misunderstanding due to the belief that a leg joint that bends between the foot and the body must be a knee. The main difference between us and the elephants is that our foot bones and hand bones are separate, whereas those of the elephant are one in the same, and have evolved to suit this four-legged mammal.

 

Why do elephants have tusks?

 

The tusks are used for obtaining food, fighting (amongst males) and for self defence. They are actually their upper incisors, and grow continuously until they die at around 60 years old. Although their skin is up to 3cm (1 inch) thick, it is quite sensitive.

 

Diet

 

Elephants are voracious feeders which in a day consume up to 272 kg (600 pounds) of grass, tender shoots and bark from trees. An adult Elephant can drink up to 200 litres of water in a single session. A single Elephant deposits up to 150kg (330 pounds) of dung every day - about one dollop every 15 minutes!

 

Breeding

 

African Elephant are not seasonal breeders. Generally they produce one calf every 3 to 4 years. The gestation period is about 22 months. At birth calves weigh about 100 kg (220 pounds) and are fully weaned between 18-24 months. An orphaned calf will usually be adopted by one of the family's lactating females or suckled by various females. Elephants are very attentive mothers, and because most Elephant behavior has to be learned, they keep their offspring with them for many years. Tusks erupt at 16 months but do not show externally until 30 months. Once weaned, usually at age 4 or 5, the calf still remains in the maternal group. Females mature at about 11 years and stay in the group, while the males, which mature between 12 and 15, are usually expelled from the maternal herd. Even though these young males are sexually mature, they do not breed until they are in their mid, or late 20s or even older and have moved up in the social hierarchy.

 

Behaviour

 

Mature males form bachelor groups and become solitary bulls. Elephant form strong family units of cows, calves and young offspring. Such herds are always led by an old female. Apart from drinking large quantities of water they also love wading or swimming in it. Elephants clearly relish mud baths.

It was once thought that family groups were led by old bull elephants, but these males are most often solitary. The female family groups are often visited by mature males checking for females in oestrus. Several interrelated family groups may inhabit an area and know each other well.

 

How do you tell an elephant's mock charge from a serious one?

 

It is imperative to keep in mind that Elephant are extremely intelligent, and each individual has a distinct character. Although there will be exceptions to the rules, the common signs of a mock charge are bush-bashing, dust-throwing, trumpeting and other vocalizations, open ears and an intimidating presence, can be considered a mock-display. Aggressive or startled elephants usually make sudden headshakes and flap their large ears against their head. Serious charges usually occur after all attempts to intimidate have failed, and the Elephant feels threatened. The ears are pinned back and head and trunk are lowered. Ultimately, the key lies in the intelligence of the animal and how they will react to the 'target' and unfamiliar actions, and a conscious decision is made.

 

Why do elephants rhythmically flap their ears?

 

Contrary to common belief, it is not an expression of anger. Being an animal of such a large size, with no sweat glands and a dark body colour, elephants flap their ears to cool the body and rid themselves of irritating insects.

 

Where are they found?

 

Once ranging across most of Africa the Elephant population has declined dramatically across the continent. In South Africa the Addo Elephant and Kruger National Park protect large herds. Due to rigorous conservation measures the Elephant population in South Africa has grown from a estimated 120 in 1920 in 4 locations, to 10 000 at 40 locations to date.

 

Notes

 

The African Elephant has recently been classified into two separate species, the more common African Bush Elephant [Loxodonta Africana] and the smaller African Forest Elephant [Loxodonta cyclotis] of the rainforest of Central Africa.

 

(krugerpark.co.za)

 

Der Addo-Elefanten-Nationalpark (afrikaans: Addo Olifant Nasionale Park, englisch: Addo Elephant National Park) liegt im Distrikt Cacadu, im westlichen Teil der Provinz Ostkap in Südafrika, 70 Kilometer nordöstlich von Port Elizabeth im Sundays River Valley. Der Elefanten-Nationalpark ist mit 1640 km² der größte Nationalpark im Ostkap.

 

Der Nationalpark wurde 1931 zum Schutz der elf letzten überlebenden Elefanten der Region eingerichtet, die bis zu diesem Zeitpunkt noch nicht zum Opfer von Elfenbeinjägern oder Farmern geworden waren. Der Gründung vorausgegangen war eine von Seiten der Regierung initiierte Jagd auf die hier heimischen Kap-Elefanten, die auf der Suche nach Nahrung immer wieder die Felder und Gärten der hier ansässigen Farmer verwüsteten. Nachdem es zu öffentlichen Protesten gekommen war, als der „letzte große weiße Jäger“ Major P. J. Pretorius in einem Jahr 130 Elefanten erlegte, wurde im Addo-Busch das seitdem mehrfach erweiterte Wildreservat eingerichtet. Im Jahr 1954, als es 22 Elefanten gab, ließ der damalige Parkmanager Graham Armstrong eine Fläche von 2270 Hektar mit Elefantenzäunen umgeben. Dieser Zaun wird noch heute vom Park genutzt und wird nach seinem Erfinder Armstrong-Zaun genannt.

 

Im Jahr 2004 lebten im Park etwa 350 Elefanten; 2006 wurden bereits knapp über 400 Elefanten gezählt. Damit erreichte der Nationalpark das ökologisch vertretbare Maximum an Elefanten.

 

Langfristig soll der Addo Elephant Park der drittgrößte Park Südafrikas werden. Der Park soll auf eine Größe von 3600 km² anwachsen.

 

Neben Elefanten leben im Addo-Elefanten-Nationalpark Kudus, Afrikanischer Büffel, Elenantilopen, Südafrikanische Kuhantilopen, Buschböcke, Warzenschweine, Steppenzebras, Spitzmaulnashörner, Hyänen und Leoparden. In einigen Randgebieten, etwa im Bereich des Darlington-Dammes beziehungsweise in der Nähe der Zuurberg Mountains, leben auch einige für das Kapgebiet typische Huftierarten, wie Bergzebras, Weißschwanzgnus, Oryxantilopen und Springböcke. Am Sundays River leben Flusspferde. 2003 wurden Löwen im Park angesiedelt, so dass man seither die sogenannten Big Five im Park antreffen kann. Fleckenhyänen wurden ebenfalls angesiedelt, und auch die Wiederansiedlung von Wildhunden und Geparden ist geplant.

 

Der Nationalpark beherbergt mehr als 500 verschiedene Pflanzenarten aus rund 70 Familien. Man findet hier vorrangig kleine Pflanzenarten sowie verschiedene Buscharten wie Schotia afra und Portulacaria afra.

 

(Wikipedia)

 

Der Afrikanische Elefant (Loxodonta africana), auch Afrikanischer Steppenelefant oder Afrikanischer Buschelefant, ist eine Art aus der Familie der Elefanten. Er ist das größte gegenwärtig lebende Landsäugetier und gleichzeitig das größte rezente landbewohnende Tier der Erde. Herausragende Kennzeichen sind neben den Stoßzähnen und dem markanten Rüssel die großen Ohren und die säulenförmigen Beine. In zahlreichen morphologischen und anatomischen Merkmalen unterscheidet sich der Afrikanische Elefant von seinen etwas kleineren Verwandten, dem Waldelefanten und dem Asiatischen Elefanten. Das Verbreitungsgebiet umfasst heute große Teile von Afrika südlich der Sahara. Die Tiere haben sich dort an zahlreiche unterschiedliche Lebensräume angepasst, die von geschlossenen Wäldern über offene Savannenlandschaften bis hin zu Sumpfgebieten und wüstenartigen Regionen reichen. Insgesamt ist das Vorkommen aber stark fragmentiert.

 

Die Lebensweise des Afrikanischen Elefanten ist durch intensive Studien gut erforscht. Sie wird durch einen stark sozialen Charakter geprägt. Weibliche Tiere und ihr Nachwuchs leben in Familienverbänden (Herden). Diese formieren sich wiederum zu einem enger verwandten Clan. Die einzelnen Herden treffen sich zu bestimmten Gelegenheiten und trennen sich danach wieder. Die männlichen Tiere bilden Junggesellengruppen. Die verschiedenen Verbände nutzen Aktionsräume, in denen sie teils im Jahreszyklus herumwandern. Für die Kommunikation untereinander nutzen die Tiere verschiedene Töne im niedrigen Frequenzbereich. Anhand der Lautgebung, aber auch durch bestimmte chemische Signale können sich die einzelnen Individuen untereinander erkennen. Darüber hinaus besteht ein umfangreiches Repertoire an Gesten. Hervorzuheben sind auch die kognitiven Fähigkeiten des Afrikanischen Elefanten.

 

Die Nahrung besteht sowohl aus weicher wie auch harter Pflanzenkost. Die genaue Zusammensetzung variiert dabei regional und jahreszeitlich. Generell verbringt der Afrikanische Elefant einen großen Teil seiner Tagesaktivitäten mit der Nahrungsaufnahme. Die Fortpflanzung erfolgt ganzjährig, regional gibt es Tendenzen zu einer stärkeren Saisonalisierung. Bullen kommen einmal jährlich in die Musth, während deren sie auf Wanderung zur Suche nach fortpflanzungswilligen Kühen gehen. Während der Musth ist die Aggressivität gesteigert, es finden dann auch Rivalenkämpfe statt. Der Sexualzyklus der Kühe dauert vergleichsweise lange und weist einen für Säugetiere untypischen Verlauf auf. Nach erfolgter Geburt setzt er in der Regel mehrere Jahre aus. Zumeist wird nach fast zweijähriger Tragzeit ein Jungtier geboren, das in der mütterlichen Herde aufwächst. Junge weibliche Tiere verbleiben später in der Herde, die jungen männlichen verlassen diese.

 

Die wissenschaftliche Erstbeschreibung des Afrikanischen Elefanten erfolgte im Jahr 1797 mit einer formalen artlichen Trennung des Afrikanischen vom Asiatischen Elefanten. Der heute gebräuchliche Gattungsname Loxodonta wurde offiziell erst dreißig Jahre später eingeführt. Die Bezeichnung bezieht sich auf markante Zahnunterschiede zwischen den asiatischen und den afrikanischen Elefanten. Im Verlauf des 20. Jahrhunderts wurden mehrere Unterarten unterschieden, darunter auch der Waldelefant des zentralen Afrikas. Letzterer gilt heute genetischen Untersuchungen zufolge als eigenständige Art, die weiteren Unterarten sind nicht anerkannt. Stammesgeschichtlich lässt sich der Afrikanische Elefant erstmals im beginnenden Mittleren Pleistozän belegen. Der Gesamtbestand gilt als gefährdet. Ursachen hierfür sind hauptsächlich die Jagd nach Elfenbein und Lebensraumverlust durch die zunehmend wachsende menschliche Bevölkerung. Der Afrikanische Elefant zählt zu den sogenannten „Big Five“ von Großwildjagd und Safari.

 

(Wikipedia)

Antoon van Dyck (Antwerp, 1599 - London, 1641) - Paulus Pontius (engraver) - Gaspar De Crayer - Van Dyck exhibition Court painter - Turin, Royal Museums - Palatine Hall of the Sabauda Gallery

Piazza Santa Maria Novella, Firenze

Known for their huge bodies, unique trunks, and long lives, elephants are the largest land animals in the world. They live in social groups called herds which are composed of related females, called cows, and their offspring. The leader of the herd is a female, usually the oldest, most experienced member of the herd. She decides the herd’s activities. The females stay with their female relatives for life, but young males leave the herd when they reach puberty, either traveling alone or in bachelor groups. Females will practice motherhood in the herd by being “aunts” to the calves.

 

Elephants will display dominance by raising their head, trunk and ears.They will also shake their heads and make trumpeting noises, followed by loud rumbles. They use their huge ears as fans, flapping them to cool themselves off, and their trunks perform a variety of jobs from spraying water and dust to picking up tiny objects to heavy lifting. Their tusks are actually their incisor teeth and they use them to dig, lift things, and protect themselves.

 

DIET

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