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Ipê Amarelo, Tabebuia [chrysotricha or ochracea].
Ipê-amarelo em Brasília, Brasil.
This tree is in Brasília, Capital of Brazil.
Text, in english, from Wikipedia, the free encyclopedia
"Trumpet tree" redirects here. This term is occasionally used for the Shield-leaved Pumpwood (Cecropia peltata).
Tabebuia
Flowering Araguaney or ipê-amarelo (Tabebuia chrysantha) in central Brazil
Scientific classification
Kingdom: Plantae
(unranked): Angiosperms
(unranked): Eudicots
(unranked): Asterids
Order: Lamiales
Family: Bignoniaceae
Tribe: Tecomeae
Genus: Tabebuia
Gomez
Species
Nearly 100.
Tabebuia is a neotropical genus of about 100 species in the tribe Tecomeae of the family Bignoniaceae. The species range from northern Mexico and the Antilles south to northern Argentina and central Venezuela, including the Caribbean islands of Hispaniola (Dominican Republic and Haiti) and Cuba. Well-known common names include Ipê, Poui, trumpet trees and pau d'arco.
They are large shrubs and trees growing to 5 to 50 m (16 to 160 ft.) tall depending on the species; many species are dry-season deciduous but some are evergreen. The leaves are opposite pairs, complex or palmately compound with 3–7 leaflets.
Tabebuia is a notable flowering tree. The flowers are 3 to 11 cm (1 to 4 in.) wide and are produced in dense clusters. They present a cupular calyx campanulate to tubular, truncate, bilabiate or 5-lobed. Corolla colors vary between species ranging from white, light pink, yellow, lavender, magenta, or red. The outside texture of the flower tube is either glabrous or pubescentThe fruit is a dehiscent pod, 10 to 50 cm (4 to 20 in.) long, containing numerous—in some species winged—seeds. These pods often remain on the tree through dry season until the beginning of the rainy.
Species in this genus are important as timber trees. The wood is used for furniture, decking, and other outdoor uses. It is increasingly popular as a decking material due to its insect resistance and durability. By 2007, FSC-certified ipê wood had become readily available on the market, although certificates are occasionally forged.
Tabebuia is widely used as ornamental tree in the tropics in landscaping gardens, public squares, and boulevards due to its impressive and colorful flowering. Many flowers appear on still leafless stems at the end of the dry season, making the floral display more conspicuous. They are useful as honey plants for bees, and are popular with certain hummingbirds. Naturalist Madhaviah Krishnan on the other hand once famously took offense at ipé grown in India, where it is not native.
Lapacho teaThe bark of several species has medical properties. The bark is dried, shredded, and then boiled making a bitter or sour-tasting brownish-colored tea. Tea from the inner bark of Pink Ipê (T. impetiginosa) is known as Lapacho or Taheebo. Its main active principles are lapachol, quercetin, and other flavonoids. It is also available in pill form. The herbal remedy is typically used during flu and cold season and for easing smoker's cough. It apparently works as expectorant, by promoting the lungs to cough up and free deeply embedded mucus and contaminants. However, lapachol is rather toxic and therefore a more topical use e.g. as antibiotic or pesticide may be advisable. Other species with significant folk medical use are T. alba and Yellow Lapacho (T. serratifolia)
Tabebuia heteropoda, T. incana, and other species are occasionally used as an additive to the entheogenic drink Ayahuasca.
Mycosphaerella tabebuiae, a plant pathogenic sac fungus, was first discovered on an ipê tree.
Tabebuia alba
Tabebuia anafensis
Tabebuia arimaoensis
Tabebuia aurea – Caribbean Trumpet Tree
Tabebuia bilbergii
Tabebuia bibracteolata
Tabebuia cassinoides
Tabebuia chrysantha – Araguaney, Yellow Ipê, tajibo (Bolivia), ipê-amarelo (Brazil), cañaguate (N Colombia)
Tabebuia chrysotricha – Golden Trumpet Tree
Tabebuia donnell-smithii Rose – Gold Tree, "Prima Vera", Cortez blanco (El Salvador), San Juan (Honduras), palo blanco (Guatemala),duranga (Mexico)
A native of Mexico and Central Americas, considered one of the most colorful of all Central American trees. The leaves are deciduous. Masses of golden-yellow flowers cover the crown after the leaves are shed.
Tabebuia dubia
Tabebuia ecuadorensis
Tabebuia elongata
Tabebuia furfuracea
Tabebuia geminiflora Rizz. & Mattos
Tabebuia guayacan (Seem.) Hemsl.
Tabebuia haemantha
Tabebuia heptaphylla (Vell.) Toledo – tajy
Tabebuia heterophylla – roble prieto
Tabebuia heteropoda
Tabebuia hypoleuca
Tabebuia impetiginosa – Pink Ipê, Pink Lapacho, ipê-cavatã, ipê-comum, ipê-reto, ipê-rosa, ipê-roxo-damata, pau d'arco-roxo, peúva, piúva (Brazil), lapacho negro (Spanish); not "brazilwood"
Tabebuia incana
Tabebuia jackiana
Tabebuia lapacho – lapacho amarillo
Tabebuia orinocensis A.H. Gentry[verification needed]
Tabebuia ochracea
Tabebuia oligolepis
Tabebuia pallida – Cuban Pink Trumpet Tree
Tabebuia platyantha
Tabebuia polymorpha
Tabebuia rosea (Bertol.) DC.[verification needed] (= T. pentaphylla (L.) Hemsley) – Pink Poui, Pink Tecoma, apama, apamate, matilisguate
A popular street tree in tropical cities because of its multi-annular masses of light pink to purple flowers and modest size. The roots are not especially destructive for roads and sidewalks. It is the national tree of El Salvador and the state tree of Cojedes, Venezuela
Tabebuia roseo-alba – White Ipê, ipê-branco (Brazil), lapacho blanco
Tabebuia serratifolia – Yellow Lapacho, Yellow Poui, ipê-roxo (Brazil)
Tabebuia shaferi
Tabebuia striata
Tabebuia subtilis Sprague & Sandwith
Tabebuia umbellata
Tabebuia vellosoi Toledo
Ipê-do-cerrado
Texto, em português, da Wikipédia, a enciclopédia livre.
Ipê-do-cerrado
Classificação científica
Reino: Plantae
Divisão: Magnoliophyta
Classe: Magnoliopsida
Subclasse: Asteridae
Ordem: Lamiales
Família: Bignoniaceae
Género: Tabebuia
Espécie: T. ochracea
Nome binomial
Tabebuia ochracea
(Cham.) Standl. 1832
Sinónimos
Bignonia tomentosa Pav. ex DC.
Handroanthus ochraceus (Cham.) Mattos
Tabebuia chrysantha (Jacq.) G. Nicholson
Tabebuia hypodictyon A. DC.) Standl.
Tabebuia neochrysantha A.H. Gentry
Tabebuia ochracea subsp. heteropoda (A. DC.) A.H. Gentry
Tabebuia ochracea subsp. neochrysantha (A.H. Gentry) A.H. Gentry
Tecoma campinae Kraenzl.
ecoma grandiceps Kraenzl.
Tecoma hassleri Sprague
Tecoma hemmendorffiana Kraenzl.
Tecoma heteropoda A. DC.
Tecoma hypodictyon A. DC.
Tecoma ochracea Cham.
Ipê-do-cerrado é um dos nomes populares da Tabebuia ochracea (Cham.) Standl. 1832, nativa do cerrado brasileiro, no estados de Amazonas, Pará, Maranhão, Piauí, Ceará, Pernambuco, Bahia, Espírito Santo, Goiás, Mato Grosso, Mato Grosso do Sul, Minas Gerais, Rio de Janeiro, São Paulo e Paraná.
Está na lista de espécies ameaçadas do estado de São Paulo, onde é encontrda também no domínio da Mata Atlântica[1].
Ocorre também na Argentina, Paraguai, Bolívia, Equador, Peru, Venezuela, Guiana, El Salvador, Guatemala e Panamá[2].
Há uma espécie homônima descrita por A.H. Gentry em 1992.
Outros nomes populares: ipê-amarelo, ipê-cascudo, ipê-do-campo, ipê-pardo, pau-d'arco-do-campo, piúva, tarumã.
Características
Altura de 6 a 14 m. Tronco tortuso com até 50 cm de diâmetro. Folhas pilosas em ambas as faces, mais na inferior, que é mais clara.
Planta decídua, heliófita, xerófita, nativa do cerrado em solos bem drenados.
Floresce de julho a setembro. Os frutos amadurecem de setembro a outubro.
FloresProduz grande quantidade de sementes leves, aladas com pequenas reservas, e que perdem a viabilidade em menos de 90 dias após coleta. A sua conservação vem sendo estudada em termos de determinação da condição ideal de armazenamento, e tem demonstrado a importância de se conhecer o comportamento da espécie quando armazenada com diferentes teores de umidade inicial, e a umidade de equilíbrio crítica para a espécie (KANO; MÁRQUEZ & KAGEYAMA, 1978). As levíssimas sementes aladas da espécie não necessitam de quebra de dormência. Podem apenas ser expostas ao sol por cerca de 6 horas e semeadas diretamente nos saquinhos. A germinação ocorre após 30 dias e de 80%. As sementes são ortodoxas e há aproximadamente 72 000 sementes em cada quilo.
O desenvolvimento da planta é rápido.
Como outros ipês, a madeira é usada em tacos, assoalhos, e em dormentes e postes. Presta-se também para peças torneadas e instrumento musicais.
Tabebuia alba (Ipê-Amarelo)
Texto, em português, produzido pela Acadêmica Giovana Beatriz Theodoro Marto
Supervisão e orientação do Prof. Luiz Ernesto George Barrichelo e do Eng. Paulo Henrique Müller
Atualizado em 10/07/2006
O ipê amarelo é a árvore brasileira mais conhecida, a mais cultivada e, sem dúvida nenhuma, a mais bela. É na verdade um complexo de nove ou dez espécies com características mais ou menos semelhantes, com flores brancas, amarelas ou roxas. Não há região do país onde não exista pelo menos uma espécie dele, porém a existência do ipê em habitat natural nos dias atuais é rara entre a maioria das espécies (LORENZI,2000).
A espécie Tabebuia alba, nativa do Brasil, é uma das espécies do gênero Tabebuia que possui “Ipê Amarelo” como nome popular. O nome alba provém de albus (branco em latim) e é devido ao tomento branco dos ramos e folhas novas.
As árvores desta espécie proporcionam um belo espetáculo com sua bela floração na arborização de ruas em algumas cidades brasileiras. São lindas árvores que embelezam e promovem um colorido no final do inverno. Existe uma crença popular de que quando o ipê-amarelo floresce não vão ocorrer mais geadas. Infelizmente, a espécie é considerada vulnerável quanto à ameaça de extinção.
A Tabebuia alba, natural do semi-árido alagoano está adaptada a todas as regiões fisiográficas, levando o governo, por meio do Decreto nº 6239, a transformar a espécie como a árvore símbolo do estado, estando, pois sob a sua tutela, não mais podendo ser suprimida de seus habitats naturais.
Taxonomia
Família: Bignoniaceae
Espécie: Tabebuia Alba (Chamiso) Sandwith
Sinonímia botânica: Handroanthus albus (Chamiso) Mattos; Tecoma alba Chamisso
Outros nomes vulgares: ipê-amarelo, ipê, aipê, ipê-branco, ipê-mamono, ipê-mandioca, ipê-ouro, ipê-pardo, ipê-vacariano, ipê-tabaco, ipê-do-cerrado, ipê-dourado, ipê-da-serra, ipezeiro, pau-d’arco-amarelo, taipoca.
Aspectos Ecológicos
O ipê-amarelo é uma espécie heliófita (Planta adaptada ao crescimento em ambiente aberto ou exposto à luz direta) e decídua (que perde as folhas em determinada época do ano). Pertence ao grupo das espécies secundárias iniciais (DURIGAN & NOGUEIRA, 1990).
Abrange a Floresta Pluvial da Mata Atlântica e da Floresta Latifoliada Semidecídua, ocorrendo principalmente no interior da Floresta Primária Densa. É característica de sub-bosques dos pinhais, onde há regeneração regular.
Informações Botânicas
Morfologia
As árvores de Tabebuia alba possuem cerca de 30 metros de altura. O tronco é reto ou levemente tortuoso, com fuste de 5 a 8 m de altura. A casca externa é grisáceo-grossa, possuindo fissuras longitudinais esparas e profundas. A coloração desta é cinza-rosa intenso, com camadas fibrosas, muito resistentes e finas, porém bem distintas.
Com ramos grossos, tortuosos e compridos, o ipê-amarelo possui copa alongada e alargada na base. As raízes de sustentação e absorção são vigorosas e profundas.
As folhas, deciduais, são opostas, digitadas e compostas. A face superior destas folhas é verde-escura, e, a face inferior, acinzentada, sendo ambas as faces tomentosas. Os pecíolos das folhas medem de 2,5 a 10 cm de comprimento. Os folíolos, geralmente, apresentam-se em número de 5 a 7, possuindo de 7 a 18 cm de comprimento por 2 a 6 cm de largura. Quando jovem estes folíolos são densamente pilosos em ambas as faces. O ápice destes é pontiagudo, com base arredondada e margem serreada.
As flores, grandes e lanceoladas, são de coloração amarelo-ouro. Possuem em média 8X15 cm.
Quanto aos frutos, estes possuem forma de cápsula bivalvar e são secos e deiscentes. Do tipo síliqua, lembram uma vagem. Medem de 15 a 30 cm de comprimento por 1,5 a 2,5 cm de largura. As valvas são finamente tomentosas com pêlos ramificados. Possuem grande quantidade de sementes.
As sementes são membranáceas brilhantes e esbranquiçadas, de coloração marrom. Possuem de 2 a 3 cm de comprimento por 7 a 9 mm de largura e são aladas.
Reprodução
A espécie é caducifólia e a queda das folhas coincide com o período de floração. A floração inicia-se no final de agosto, podendo ocorrer alguma variação devido a fenômenos climáticos. Como a espécie floresce no final do inverno é influenciada pela intensidade do mesmo. Quanto mais frio e seco for o inverno, maior será a intensidade da florada do ipê amarelo.
As flores por sua exuberância, atraem abelhas e pássaros, principalmente beija-flores que são importantes agentes polinizadores. Segundo CARVALHO (2003), a espécie possui como vetor de polinização a abelha mamangava (Bombus morio).
As sementes são dispersas pelo vento.
A planta é hermafrodita, e frutifica nos meses de setembro, outubro, novembro, dezembro, janeiro e fevereiro, dependendo da sua localização. Em cultivo, a espécie inicia o processo reprodutivo após o terceiro ano.
Ocorrência Natural
Ocorre naturalmente na Floresta Estaciobal Semidecicual, Floresta de Araucária e no Cerrado.
Segundo o IBGE, a Tabebuia alba (Cham.) Sandw. é uma árvore do Cerrado, Cerradão e Mata Seca. Apresentando-se nos campos secos (savana gramíneo-lenhosa), próximo às escarpas.
Clima
Segundo a classificação de Köppen, o ipê-amarelo abrange locais de clima tropical (Aw), subtropical úmido (Cfa), sutropical de altitude (Cwa e Cwb) e temperado.
A T.alba pode tolerar até 81 geadas em um ano. Ocorre em locais onde a temperatura média anual varia de 14,4ºC como mínimo e 22,4ºC como máximo.
Solo
A espécie prefere solos úmidos, com drenagem lenta e geralmente não muito ondulados (LONGHI, 1995).
Aparece em terras de boa à média fertilidade, em solos profundos ou rasos, nas matas e raramente cerradões (NOGUEIRA, 1977).
Pragas e Doenças
De acordo com CARVALHO (2003), possui como praga a espécie de coleópteros Cydianerus bohemani da família Curculionoideae e um outro coleóptero da família Chrysomellidae. Apesar da constatação de elevados índices populacionais do primeiro, os danos ocasionados até o momento são leves. Nas praças e ruas de Curitiba - PR, 31% das árvores foram atacadas pela Cochonilha Ceroplastes grandis.
ZIDKO (2002), ao estudar no município de Piracicaba a associação de coleópteros em espécies arbóreas, verificou a presença de insetos adultos da espécie Sitophilus linearis da família de coleópteros, Curculionidae, em estruturas reprodutivas. Os insetos adultos da espécie emergiram das vagens do ipê, danificando as sementes desta espécie nativa.
ANDRADE (1928) assinalou diversas espécies de Cerambycidae atacando essências florestais vivas, como ingazeiro, cinamomo, cangerana, cedro, caixeta, jacarandá, araribá, jatobá, entre outras como o ipê amarelo.
A Madeira
A Tabebuia alba produz madeira de grande durabilidade e resistência ao apodrecimento (LONGHI,1995).
MANIERI (1970) caracteriza o cerne desta espécie como de cor pardo-havana-claro, pardo-havan-escuro, ou pardo-acastanhado, com reflexos esverdeados. A superfície da madeira é irregularmente lustrosa, lisa ao tato, possuindo textura media e grã-direita.
Com densidade entre 0,90 e 1,15 grama por centímetro cúbico, a madeira é muito dura (LORENZI, 1992), apresentando grande dificuldade ao serrar.
A madeira possui cheiro e gosto distintos. Segundo LORENZI (1992), o cheiro característico é devido à presença da substância lapachol, ou ipeína.
Usos da Madeira
Sendo pesada, com cerne escuro, adquire grande valor comercial na marcenaria e carpintaria. Também é utilizada para fabricação de dormentes, moirões, pontes, postes, eixos de roda, varais de carroça, moendas de cana, etc.
Produtos Não-Madeireiros
A entrecasca do ipê-amarelo possui propriedades terapêuticas como adstringente, usada no tratamento de garganta e estomatites. É também usada como diurético.
O ipê-amarelo possui flores melíferas e que maduras podem ser utilizadas na alimentação humana.
Outros Usos
É comumente utilizada em paisagismo de parques e jardins pela beleza e porte. Além disso, é muito utilizada na arborização urbana.
Segundo MOREIRA & SOUZA (1987), o ipê-amarelo costuma povoar as beiras dos rios sendo, portanto, indicado para recomposição de matas ciliares. MARTINS (1986), também cita a espécie para recomposição de matas ciliares da Floresta Estacional Semidecidual, abrangendo alguns municípios das regiões Norte, Noroeste e parte do Oeste do Estado do Paraná.
Aspectos Silviculturais
Possui a tendência a crescer reto e sem bifurcações quando plantado em reflorestamento misto, pois é espécie monopodial. A desrrama se faz muito bem e a cicatrização é boa. Sendo assim, dificilmente encopa quando nova, a não ser que seja plantado em parques e jardins.
Ao ser utilizada em arborização urbana, o ipê amarelo requer podas de condução com freqüência mediana.
Espécie heliófila apresenta a pleno sol ramificação cimosa, registrando-se assim dicotomia para gema apical. Deve ser preconizada, para seu melhor aproveitamento madeireiro, podas de formação usuais (INQUE et al., 1983).
Produção de Mudas
A propagação deve realizada através de enxertia.
Os frutos devem ser coletados antes da dispersão, para evitar a perda de sementes. Após a coleta as sementes são postas em ambiente ventilado e a extração é feita manualmente. As sementes do ipê amarelo são ortodoxas, mantendo a viabilidade natural por até 3 meses em sala e por até 9 meses em vidro fechado, em câmara fria.
A condução das mudas deve ser feita a pleno sol. A muda atinge cerca de 30 cm em 9 meses, apresentando tolerância ao sol 3 semanas após a germinação.
Sementes
Os ipês, espécies do gênero Tabebuia, produzem uma grande quantidade de sementes leves, aladas com pequenas reservas, e que perdem a viabilidade em poucos dias após a sua coleta. A sua conservação vem sendo estudada em termos de determinação da condição ideal de armazenamento, e tem demonstrado a importância de se conhecer o comportamento da espécie quando armazenada com diferentes teores de umidade inicial, e a umidade de equilíbrio crítica para a espécie (KANO; MÁRQUEZ & KAGEYAMA, 1978).
As levíssimas sementes aladas da espécie não necessitam de quebra de dormência. Podem apenas ser expostas ao sol por cerca de 6 horas e semeadas diretamente nos saquinhos. A quebra natural leva cerca de 3 meses e a quebra na câmara leva 9 meses. A germinação ocorre após 30 dias e de 80%.
As sementes são ortodoxas e há aproximadamente 87000 sementes em cada quilo.
Preço da Madeira no Mercado
O preço médio do metro cúbico de pranchas de ipê no Estado do Pará cotado em Julho e Agosto de 2005 foi de R$1.200,00 o preço mínimo, R$ 1509,35 o médio e R$ 2.000,00 o preço máximo (CEPEA,2005).
The Danaid Eggfly or Mimic (Hypolimnas misippus) is a widespread species of nymphalid butterfly. It is well known for polymorphism and mimicry. Males are blackish with distinctive white spots that are fringed in blue. Females are in multiple forms that include male like forms while others appear like the toxic Danaus chrysippus and Danaus plexippus butterfles. They are found across Africa, Asia and Australia.
From Wikipedia, the free encyclopedia
Ipê Amarelo, Tabebuia [chrysotricha or ochracea].
Ipê-amarelo em Brasília, Brasil.
This tree is in Brasília, Capital of Brazil.
Text, in english, from Wikipedia, the free encyclopedia
"Trumpet tree" redirects here. This term is occasionally used for the Shield-leaved Pumpwood (Cecropia peltata).
Tabebuia
Flowering Araguaney or ipê-amarelo (Tabebuia chrysantha) in central Brazil
Scientific classification
Kingdom: Plantae
(unranked): Angiosperms
(unranked): Eudicots
(unranked): Asterids
Order: Lamiales
Family: Bignoniaceae
Tribe: Tecomeae
Genus: Tabebuia
Gomez
Species
Nearly 100.
Tabebuia is a neotropical genus of about 100 species in the tribe Tecomeae of the family Bignoniaceae. The species range from northern Mexico and the Antilles south to northern Argentina and central Venezuela, including the Caribbean islands of Hispaniola (Dominican Republic and Haiti) and Cuba. Well-known common names include Ipê, Poui, trumpet trees and pau d'arco.
They are large shrubs and trees growing to 5 to 50 m (16 to 160 ft.) tall depending on the species; many species are dry-season deciduous but some are evergreen. The leaves are opposite pairs, complex or palmately compound with 3–7 leaflets.
Tabebuia is a notable flowering tree. The flowers are 3 to 11 cm (1 to 4 in.) wide and are produced in dense clusters. They present a cupular calyx campanulate to tubular, truncate, bilabiate or 5-lobed. Corolla colors vary between species ranging from white, light pink, yellow, lavender, magenta, or red. The outside texture of the flower tube is either glabrous or pubescentThe fruit is a dehiscent pod, 10 to 50 cm (4 to 20 in.) long, containing numerous—in some species winged—seeds. These pods often remain on the tree through dry season until the beginning of the rainy.
Species in this genus are important as timber trees. The wood is used for furniture, decking, and other outdoor uses. It is increasingly popular as a decking material due to its insect resistance and durability. By 2007, FSC-certified ipê wood had become readily available on the market, although certificates are occasionally forged.
Tabebuia is widely used as ornamental tree in the tropics in landscaping gardens, public squares, and boulevards due to its impressive and colorful flowering. Many flowers appear on still leafless stems at the end of the dry season, making the floral display more conspicuous. They are useful as honey plants for bees, and are popular with certain hummingbirds. Naturalist Madhaviah Krishnan on the other hand once famously took offense at ipé grown in India, where it is not native.
Lapacho teaThe bark of several species has medical properties. The bark is dried, shredded, and then boiled making a bitter or sour-tasting brownish-colored tea. Tea from the inner bark of Pink Ipê (T. impetiginosa) is known as Lapacho or Taheebo. Its main active principles are lapachol, quercetin, and other flavonoids. It is also available in pill form. The herbal remedy is typically used during flu and cold season and for easing smoker's cough. It apparently works as expectorant, by promoting the lungs to cough up and free deeply embedded mucus and contaminants. However, lapachol is rather toxic and therefore a more topical use e.g. as antibiotic or pesticide may be advisable. Other species with significant folk medical use are T. alba and Yellow Lapacho (T. serratifolia)
Tabebuia heteropoda, T. incana, and other species are occasionally used as an additive to the entheogenic drink Ayahuasca.
Mycosphaerella tabebuiae, a plant pathogenic sac fungus, was first discovered on an ipê tree.
Tabebuia alba
Tabebuia anafensis
Tabebuia arimaoensis
Tabebuia aurea – Caribbean Trumpet Tree
Tabebuia bilbergii
Tabebuia bibracteolata
Tabebuia cassinoides
Tabebuia chrysantha – Araguaney, Yellow Ipê, tajibo (Bolivia), ipê-amarelo (Brazil), cañaguate (N Colombia)
Tabebuia chrysotricha – Golden Trumpet Tree
Tabebuia donnell-smithii Rose – Gold Tree, "Prima Vera", Cortez blanco (El Salvador), San Juan (Honduras), palo blanco (Guatemala),duranga (Mexico)
A native of Mexico and Central Americas, considered one of the most colorful of all Central American trees. The leaves are deciduous. Masses of golden-yellow flowers cover the crown after the leaves are shed.
Tabebuia dubia
Tabebuia ecuadorensis
Tabebuia elongata
Tabebuia furfuracea
Tabebuia geminiflora Rizz. & Mattos
Tabebuia guayacan (Seem.) Hemsl.
Tabebuia haemantha
Tabebuia heptaphylla (Vell.) Toledo – tajy
Tabebuia heterophylla – roble prieto
Tabebuia heteropoda
Tabebuia hypoleuca
Tabebuia impetiginosa – Pink Ipê, Pink Lapacho, ipê-cavatã, ipê-comum, ipê-reto, ipê-rosa, ipê-roxo-damata, pau d'arco-roxo, peúva, piúva (Brazil), lapacho negro (Spanish); not "brazilwood"
Tabebuia incana
Tabebuia jackiana
Tabebuia lapacho – lapacho amarillo
Tabebuia orinocensis A.H. Gentry[verification needed]
Tabebuia ochracea
Tabebuia oligolepis
Tabebuia pallida – Cuban Pink Trumpet Tree
Tabebuia platyantha
Tabebuia polymorpha
Tabebuia rosea (Bertol.) DC.[verification needed] (= T. pentaphylla (L.) Hemsley) – Pink Poui, Pink Tecoma, apama, apamate, matilisguate
A popular street tree in tropical cities because of its multi-annular masses of light pink to purple flowers and modest size. The roots are not especially destructive for roads and sidewalks. It is the national tree of El Salvador and the state tree of Cojedes, Venezuela
Tabebuia roseo-alba – White Ipê, ipê-branco (Brazil), lapacho blanco
Tabebuia serratifolia – Yellow Lapacho, Yellow Poui, ipê-roxo (Brazil)
Tabebuia shaferi
Tabebuia striata
Tabebuia subtilis Sprague & Sandwith
Tabebuia umbellata
Tabebuia vellosoi Toledo
Ipê-do-cerrado
Texto, em português, da Wikipédia, a enciclopédia livre.
Ipê-do-cerrado
Classificação científica
Reino: Plantae
Divisão: Magnoliophyta
Classe: Magnoliopsida
Subclasse: Asteridae
Ordem: Lamiales
Família: Bignoniaceae
Género: Tabebuia
Espécie: T. ochracea
Nome binomial
Tabebuia ochracea
(Cham.) Standl. 1832
Sinónimos
Bignonia tomentosa Pav. ex DC.
Handroanthus ochraceus (Cham.) Mattos
Tabebuia chrysantha (Jacq.) G. Nicholson
Tabebuia hypodictyon A. DC.) Standl.
Tabebuia neochrysantha A.H. Gentry
Tabebuia ochracea subsp. heteropoda (A. DC.) A.H. Gentry
Tabebuia ochracea subsp. neochrysantha (A.H. Gentry) A.H. Gentry
Tecoma campinae Kraenzl.
ecoma grandiceps Kraenzl.
Tecoma hassleri Sprague
Tecoma hemmendorffiana Kraenzl.
Tecoma heteropoda A. DC.
Tecoma hypodictyon A. DC.
Tecoma ochracea Cham.
Ipê-do-cerrado é um dos nomes populares da Tabebuia ochracea (Cham.) Standl. 1832, nativa do cerrado brasileiro, no estados de Amazonas, Pará, Maranhão, Piauí, Ceará, Pernambuco, Bahia, Espírito Santo, Goiás, Mato Grosso, Mato Grosso do Sul, Minas Gerais, Rio de Janeiro, São Paulo e Paraná.
Está na lista de espécies ameaçadas do estado de São Paulo, onde é encontrda também no domínio da Mata Atlântica[1].
Ocorre também na Argentina, Paraguai, Bolívia, Equador, Peru, Venezuela, Guiana, El Salvador, Guatemala e Panamá[2].
Há uma espécie homônima descrita por A.H. Gentry em 1992.
Outros nomes populares: ipê-amarelo, ipê-cascudo, ipê-do-campo, ipê-pardo, pau-d'arco-do-campo, piúva, tarumã.
Características
Altura de 6 a 14 m. Tronco tortuso com até 50 cm de diâmetro. Folhas pilosas em ambas as faces, mais na inferior, que é mais clara.
Planta decídua, heliófita, xerófita, nativa do cerrado em solos bem drenados.
Floresce de julho a setembro. Os frutos amadurecem de setembro a outubro.
FloresProduz grande quantidade de sementes leves, aladas com pequenas reservas, e que perdem a viabilidade em menos de 90 dias após coleta. A sua conservação vem sendo estudada em termos de determinação da condição ideal de armazenamento, e tem demonstrado a importância de se conhecer o comportamento da espécie quando armazenada com diferentes teores de umidade inicial, e a umidade de equilíbrio crítica para a espécie (KANO; MÁRQUEZ & KAGEYAMA, 1978). As levíssimas sementes aladas da espécie não necessitam de quebra de dormência. Podem apenas ser expostas ao sol por cerca de 6 horas e semeadas diretamente nos saquinhos. A germinação ocorre após 30 dias e de 80%. As sementes são ortodoxas e há aproximadamente 72 000 sementes em cada quilo.
O desenvolvimento da planta é rápido.
Como outros ipês, a madeira é usada em tacos, assoalhos, e em dormentes e postes. Presta-se também para peças torneadas e instrumento musicais.
Tabebuia alba (Ipê-Amarelo)
Texto, em português, produzido pela Acadêmica Giovana Beatriz Theodoro Marto
Supervisão e orientação do Prof. Luiz Ernesto George Barrichelo e do Eng. Paulo Henrique Müller
Atualizado em 10/07/2006
O ipê amarelo é a árvore brasileira mais conhecida, a mais cultivada e, sem dúvida nenhuma, a mais bela. É na verdade um complexo de nove ou dez espécies com características mais ou menos semelhantes, com flores brancas, amarelas ou roxas. Não há região do país onde não exista pelo menos uma espécie dele, porém a existência do ipê em habitat natural nos dias atuais é rara entre a maioria das espécies (LORENZI,2000).
A espécie Tabebuia alba, nativa do Brasil, é uma das espécies do gênero Tabebuia que possui “Ipê Amarelo” como nome popular. O nome alba provém de albus (branco em latim) e é devido ao tomento branco dos ramos e folhas novas.
As árvores desta espécie proporcionam um belo espetáculo com sua bela floração na arborização de ruas em algumas cidades brasileiras. São lindas árvores que embelezam e promovem um colorido no final do inverno. Existe uma crença popular de que quando o ipê-amarelo floresce não vão ocorrer mais geadas. Infelizmente, a espécie é considerada vulnerável quanto à ameaça de extinção.
A Tabebuia alba, natural do semi-árido alagoano está adaptada a todas as regiões fisiográficas, levando o governo, por meio do Decreto nº 6239, a transformar a espécie como a árvore símbolo do estado, estando, pois sob a sua tutela, não mais podendo ser suprimida de seus habitats naturais.
Taxonomia
Família: Bignoniaceae
Espécie: Tabebuia Alba (Chamiso) Sandwith
Sinonímia botânica: Handroanthus albus (Chamiso) Mattos; Tecoma alba Chamisso
Outros nomes vulgares: ipê-amarelo, ipê, aipê, ipê-branco, ipê-mamono, ipê-mandioca, ipê-ouro, ipê-pardo, ipê-vacariano, ipê-tabaco, ipê-do-cerrado, ipê-dourado, ipê-da-serra, ipezeiro, pau-d’arco-amarelo, taipoca.
Aspectos Ecológicos
O ipê-amarelo é uma espécie heliófita (Planta adaptada ao crescimento em ambiente aberto ou exposto à luz direta) e decídua (que perde as folhas em determinada época do ano). Pertence ao grupo das espécies secundárias iniciais (DURIGAN & NOGUEIRA, 1990).
Abrange a Floresta Pluvial da Mata Atlântica e da Floresta Latifoliada Semidecídua, ocorrendo principalmente no interior da Floresta Primária Densa. É característica de sub-bosques dos pinhais, onde há regeneração regular.
Informações Botânicas
Morfologia
As árvores de Tabebuia alba possuem cerca de 30 metros de altura. O tronco é reto ou levemente tortuoso, com fuste de 5 a 8 m de altura. A casca externa é grisáceo-grossa, possuindo fissuras longitudinais esparas e profundas. A coloração desta é cinza-rosa intenso, com camadas fibrosas, muito resistentes e finas, porém bem distintas.
Com ramos grossos, tortuosos e compridos, o ipê-amarelo possui copa alongada e alargada na base. As raízes de sustentação e absorção são vigorosas e profundas.
As folhas, deciduais, são opostas, digitadas e compostas. A face superior destas folhas é verde-escura, e, a face inferior, acinzentada, sendo ambas as faces tomentosas. Os pecíolos das folhas medem de 2,5 a 10 cm de comprimento. Os folíolos, geralmente, apresentam-se em número de 5 a 7, possuindo de 7 a 18 cm de comprimento por 2 a 6 cm de largura. Quando jovem estes folíolos são densamente pilosos em ambas as faces. O ápice destes é pontiagudo, com base arredondada e margem serreada.
As flores, grandes e lanceoladas, são de coloração amarelo-ouro. Possuem em média 8X15 cm.
Quanto aos frutos, estes possuem forma de cápsula bivalvar e são secos e deiscentes. Do tipo síliqua, lembram uma vagem. Medem de 15 a 30 cm de comprimento por 1,5 a 2,5 cm de largura. As valvas são finamente tomentosas com pêlos ramificados. Possuem grande quantidade de sementes.
As sementes são membranáceas brilhantes e esbranquiçadas, de coloração marrom. Possuem de 2 a 3 cm de comprimento por 7 a 9 mm de largura e são aladas.
Reprodução
A espécie é caducifólia e a queda das folhas coincide com o período de floração. A floração inicia-se no final de agosto, podendo ocorrer alguma variação devido a fenômenos climáticos. Como a espécie floresce no final do inverno é influenciada pela intensidade do mesmo. Quanto mais frio e seco for o inverno, maior será a intensidade da florada do ipê amarelo.
As flores por sua exuberância, atraem abelhas e pássaros, principalmente beija-flores que são importantes agentes polinizadores. Segundo CARVALHO (2003), a espécie possui como vetor de polinização a abelha mamangava (Bombus morio).
As sementes são dispersas pelo vento.
A planta é hermafrodita, e frutifica nos meses de setembro, outubro, novembro, dezembro, janeiro e fevereiro, dependendo da sua localização. Em cultivo, a espécie inicia o processo reprodutivo após o terceiro ano.
Ocorrência Natural
Ocorre naturalmente na Floresta Estaciobal Semidecicual, Floresta de Araucária e no Cerrado.
Segundo o IBGE, a Tabebuia alba (Cham.) Sandw. é uma árvore do Cerrado, Cerradão e Mata Seca. Apresentando-se nos campos secos (savana gramíneo-lenhosa), próximo às escarpas.
Clima
Segundo a classificação de Köppen, o ipê-amarelo abrange locais de clima tropical (Aw), subtropical úmido (Cfa), sutropical de altitude (Cwa e Cwb) e temperado.
A T.alba pode tolerar até 81 geadas em um ano. Ocorre em locais onde a temperatura média anual varia de 14,4ºC como mínimo e 22,4ºC como máximo.
Solo
A espécie prefere solos úmidos, com drenagem lenta e geralmente não muito ondulados (LONGHI, 1995).
Aparece em terras de boa à média fertilidade, em solos profundos ou rasos, nas matas e raramente cerradões (NOGUEIRA, 1977).
Pragas e Doenças
De acordo com CARVALHO (2003), possui como praga a espécie de coleópteros Cydianerus bohemani da família Curculionoideae e um outro coleóptero da família Chrysomellidae. Apesar da constatação de elevados índices populacionais do primeiro, os danos ocasionados até o momento são leves. Nas praças e ruas de Curitiba - PR, 31% das árvores foram atacadas pela Cochonilha Ceroplastes grandis.
ZIDKO (2002), ao estudar no município de Piracicaba a associação de coleópteros em espécies arbóreas, verificou a presença de insetos adultos da espécie Sitophilus linearis da família de coleópteros, Curculionidae, em estruturas reprodutivas. Os insetos adultos da espécie emergiram das vagens do ipê, danificando as sementes desta espécie nativa.
ANDRADE (1928) assinalou diversas espécies de Cerambycidae atacando essências florestais vivas, como ingazeiro, cinamomo, cangerana, cedro, caixeta, jacarandá, araribá, jatobá, entre outras como o ipê amarelo.
A Madeira
A Tabebuia alba produz madeira de grande durabilidade e resistência ao apodrecimento (LONGHI,1995).
MANIERI (1970) caracteriza o cerne desta espécie como de cor pardo-havana-claro, pardo-havan-escuro, ou pardo-acastanhado, com reflexos esverdeados. A superfície da madeira é irregularmente lustrosa, lisa ao tato, possuindo textura media e grã-direita.
Com densidade entre 0,90 e 1,15 grama por centímetro cúbico, a madeira é muito dura (LORENZI, 1992), apresentando grande dificuldade ao serrar.
A madeira possui cheiro e gosto distintos. Segundo LORENZI (1992), o cheiro característico é devido à presença da substância lapachol, ou ipeína.
Usos da Madeira
Sendo pesada, com cerne escuro, adquire grande valor comercial na marcenaria e carpintaria. Também é utilizada para fabricação de dormentes, moirões, pontes, postes, eixos de roda, varais de carroça, moendas de cana, etc.
Produtos Não-Madeireiros
A entrecasca do ipê-amarelo possui propriedades terapêuticas como adstringente, usada no tratamento de garganta e estomatites. É também usada como diurético.
O ipê-amarelo possui flores melíferas e que maduras podem ser utilizadas na alimentação humana.
Outros Usos
É comumente utilizada em paisagismo de parques e jardins pela beleza e porte. Além disso, é muito utilizada na arborização urbana.
Segundo MOREIRA & SOUZA (1987), o ipê-amarelo costuma povoar as beiras dos rios sendo, portanto, indicado para recomposição de matas ciliares. MARTINS (1986), também cita a espécie para recomposição de matas ciliares da Floresta Estacional Semidecidual, abrangendo alguns municípios das regiões Norte, Noroeste e parte do Oeste do Estado do Paraná.
Aspectos Silviculturais
Possui a tendência a crescer reto e sem bifurcações quando plantado em reflorestamento misto, pois é espécie monopodial. A desrrama se faz muito bem e a cicatrização é boa. Sendo assim, dificilmente encopa quando nova, a não ser que seja plantado em parques e jardins.
Ao ser utilizada em arborização urbana, o ipê amarelo requer podas de condução com freqüência mediana.
Espécie heliófila apresenta a pleno sol ramificação cimosa, registrando-se assim dicotomia para gema apical. Deve ser preconizada, para seu melhor aproveitamento madeireiro, podas de formação usuais (INQUE et al., 1983).
Produção de Mudas
A propagação deve realizada através de enxertia.
Os frutos devem ser coletados antes da dispersão, para evitar a perda de sementes. Após a coleta as sementes são postas em ambiente ventilado e a extração é feita manualmente. As sementes do ipê amarelo são ortodoxas, mantendo a viabilidade natural por até 3 meses em sala e por até 9 meses em vidro fechado, em câmara fria.
A condução das mudas deve ser feita a pleno sol. A muda atinge cerca de 30 cm em 9 meses, apresentando tolerância ao sol 3 semanas após a germinação.
Sementes
Os ipês, espécies do gênero Tabebuia, produzem uma grande quantidade de sementes leves, aladas com pequenas reservas, e que perdem a viabilidade em poucos dias após a sua coleta. A sua conservação vem sendo estudada em termos de determinação da condição ideal de armazenamento, e tem demonstrado a importância de se conhecer o comportamento da espécie quando armazenada com diferentes teores de umidade inicial, e a umidade de equilíbrio crítica para a espécie (KANO; MÁRQUEZ & KAGEYAMA, 1978).
As levíssimas sementes aladas da espécie não necessitam de quebra de dormência. Podem apenas ser expostas ao sol por cerca de 6 horas e semeadas diretamente nos saquinhos. A quebra natural leva cerca de 3 meses e a quebra na câmara leva 9 meses. A germinação ocorre após 30 dias e de 80%.
As sementes são ortodoxas e há aproximadamente 87000 sementes em cada quilo.
Preço da Madeira no Mercado
O preço médio do metro cúbico de pranchas de ipê no Estado do Pará cotado em Julho e Agosto de 2005 foi de R$1.200,00 o preço mínimo, R$ 1509,35 o médio e R$ 2.000,00 o preço máximo (CEPEA,2005).
Los Jameos del Agua, situados en el norte de la isla de Lanzarote, es uno de los siete Centros de Arte, Cultura y Turismo del Cabildo de Lanzarote.
Se trata de una intervención espacial creada a partir de una serie de jameos naturales por el artista lanzaroteño César Manrique y pretende mostrar al visitante un espacio para la contemplación de la naturaleza apenas intervenida por el hombre.
Los Jameos del Agua, al igual que la Cueva de los Verdes, se localizan en el interior del túnel volcánico producido por la erupción del Volcán de la Corona. Los Jameos del Agua se encuentran situados en la sección de este túnel más cercano a la costa.
Está formado por al menos tres jameos o aberturas en el terreno. El “Jameo Chico” por donde se realiza el acceso al interior, el “Jameo Grande” y un tercero, denominado “Jameo de la Cazuela”.
Los "Jameos del Agua" es el primer Centro de Arte, Cultura y Turismo creado por César Manrique, y es el reflejo de uno de sus pilares creativos: la armonía entre la naturaleza y la creación artística.
A principios de los años sesenta, y debido al abandono del entorno, se hace necesario acometer labores de limpieza y de acondicionamiento. Aunque las obras se dilataron en el tiempo, podemos señalar el año 1966 como fecha de la apertura de las primeras fases al público.
La especial morfología del tubo volcánico provocó que se realizaran numerosos cambios del proyecto inicial, explorando nuevas alternativas creativas y procurando que las soluciones adoptadas fuesen las más adecuadas.
Será en 1977, después de más de una década de trabajos, cuando quede conformada la estructura general de los Jameos del Agua. En estas mismas fechas se procede a la inauguración oficial del centro, incluyendo el Auditorio, y se considera conclusa la obra.
Aun así, y con posterioridad, se realizan nuevas instalaciones para funciones concretas como el espacio museístico de la “Casa de los Volcanes”. Este centro se dedica, desde 1987, a una labor científica y didáctica sobre la vulcanología.
Los “Jameos del Agua” son muy importantes desde el punto de vista ecológico, ya que existe una especie de cangrejo única y endémica, los cangrejitos ciegos (Munidopsis polymorpha) un cangrejo de apenas un centímetro de longitud, albino y ciego.
Estos cangrejos son muy sensibles a los cambios de la laguna (procedente de agua de mar) por lo que el ruido y la luz les afecta. Son muy sensibles al óxido, que puede llegar a matarlos, por lo que está prohibido tirar monedas al agua.
Más información:
www.turismolanzarote.com/centros-turisticos/jameos-del-ag...
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Narcissus is a genus of predominantly spring flowering perennial plants of the amaryllis family, Amaryllidaceae. Various common names including daffodil, narcissus, and jonquil are used to describe all or some members of the genus. Narcissus has conspicuous flowers with six petal-like tepals surmounted by a cup- or trumpet-shaped corona. The flowers are generally white and yellow (also orange or pink in garden varieties), with either uniform or contrasting coloured tepals and corona.
Narcissus were well known in ancient civilisation, both medicinally and botanically, but formally described by Linnaeus in his Species Plantarum (1753). The genus is generally considered to have about ten sections with approximately 50 species. The number of species has varied, depending on how they are classified, due to similarity between species and hybridisation. The genus arose some time in the Late Oligocene to Early Miocene epochs, in the Iberian peninsula and adjacent areas of southwest Europe. The exact origin of the name Narcissus is unknown, but it is often linked to a Greek word (ancient Greek ναρκῶ narkō, "to make numb") and the myth of the youth of that name who fell in love with his own reflection. The English word "daffodil" appears to be derived from "asphodel", with which it was commonly compared.
The species are native to meadows and woods in southern Europe and North Africa with a centre of diversity in the Western Mediterranean, particularly the Iberian peninsula. Both wild and cultivated plants have naturalised widely, and were introduced into the Far East prior to the tenth century. Narcissi tend to be long-lived bulbs, which propagate by division, but are also insect-pollinated. Known pests, diseases and disorders include viruses, fungi, the larvae of flies, mites and nematodes. Some Narcissus species have become extinct, while others are threatened by increasing urbanisation and tourism.
Historical accounts suggest narcissi have been cultivated from the earliest times, but became increasingly popular in Europe after the 16th century and by the late 19th century were an important commercial crop centred primarily in the Netherlands. Today narcissi are popular as cut flowers and as ornamental plants in private and public gardens. The long history of breeding has resulted in thousands of different cultivars. For horticultural purposes, narcissi are classified into divisions, covering a wide range of shapes and colours. Like other members of their family, narcissi produce a number of different alkaloids, which provide some protection for the plant, but may be poisonous if accidentally ingested. This property has been exploited for medicinal use in traditional healing and has resulted in the production of galantamine for the treatment of Alzheimer's dementia. Long celebrated in art and literature, narcissi are associated with a number of themes in different cultures, ranging from death to good fortune, and as symbols of spring.
The daffodil is the national flower of Wales and the symbol of cancer charities in many countries. The appearance of the wild flowers in spring is associated with festivals in many places.
Narcissus is a genus of perennial herbaceous bulbiferous geophytes, which die back after flowering to an underground storage bulb. They regrow in the following year from brown-skinned ovoid bulbs with pronounced necks, and reach heights of 5–80 centimetres (2.0–31.5 in) depending on the species. Dwarf species such as N. asturiensis have a maximum height of 5–8 centimetres (2.0–3.1 in), while Narcissus tazetta may grow as tall as 80 centimetres (31 in).
The plants are scapose, having a single central leafless hollow flower stem (scape). Several green or blue-green, narrow, strap-shaped leaves arise from the bulb. The plant stem usually bears a solitary flower, but occasionally a cluster of flowers (umbel). The flowers, which are usually conspicuous and white or yellow, sometimes both or rarely green, consist of a perianth of three parts. Closest to the stem (proximal) is a floral tube above the ovary, then an outer ring composed of six tepals (undifferentiated sepals and petals), and a central disc to conical shaped corona. The flowers may hang down (pendant), or be erect. There are six pollen bearing stamens surrounding a central style. The ovary is inferior (below the floral parts) consisting of three chambers (trilocular). The fruit consists of a dry capsule that splits (dehisces) releasing numerous black seeds.
The bulb lies dormant after the leaves and flower stem die back and has contractile roots that pull it down further into the soil. The flower stem and leaves form in the bulb, to emerge the following season. Most species are dormant from summer to late winter, flowering in the spring, though a few species are autumn flowering.
The pale brown-skinned ovoid tunicate bulbs have a membranous tunic and a corky stem (base or basal) plate from which arise the adventitious root hairs in a ring around the edge, which grow up to 40 mm in length. Above the stem plate is the storage organ consisting of bulb scales, surrounding the previous flower stalk and the terminal bud. The scales are of two types, true storage organs and the bases of the foliage leaves. These have a thicker tip and a scar from where the leaf lamina became detached. The innermost leaf scale is semicircular only partly enveloping the flower stalk (semisheathed).(see Hanks Fig 1.3). The bulb may contain a number of branched bulb units, each with two to three true scales and two to three leaf bases. Each bulb unit has a life of about four years.
Once the leaves die back in summer, the roots also wither. After some years, the roots shorten pulling the bulbs deeper into the ground (contractile roots). The bulbs develop from the inside, pushing the older layers outwards which become brown and dry, forming an outer shell, the tunic or skin. Up to 60 layers have been counted in some wild species. While the plant appears dormant above the ground the flower stalk which will start to grow in the following spring, develops within the bulb surrounded by two to three deciduous leaves and their sheaths. The flower stem lies in the axil of the second true leaf.
The single leafless Plant stem stem or scape, appearing from early to late spring depending on the species, bears from 1 to 20 blooms. Stem shape depends on the species, some are highly compressed with a visible seam, while others are rounded. The stems are upright and located at the centre of the leaves. In a few species such as N. hedraeanthus the stem is oblique. The stem is hollow in the upper portion but towards the bulb is more solid and filled with a spongy material.
Narcissus plants have one to several basal leaf leaves which are linear, ligulate or strap-shaped (long and narrow), sometimes channelled adaxially to semiterete, and may (pedicellate) or may not (sessile) have a petiole stalk. The leaves are flat and broad to cylindrical at the base and arise from the bulb. The emerging plant generally has two leaves, but the mature plant usually three, rarely four, and they are covered with a cutin containing cuticle, giving them a waxy appearance. Leaf colour is light green to blue-green. In the mature plant, the leaves extend higher than the flower stem, but in some species, the leaves are low-hanging. The leaf base is encased in a colorless sheath. After flowering, the leaves turn yellow and die back once the seed pod (fruit) is ripe.
Jonquils usually have dark green, round, rush-like leaves.
The inflorescence is scapose, the single stem or scape bearing either a solitary flower or forming an umbel with up to 20 blooms. Species bearing a solitary flower include section Bulbocodium and most of section Pseudonarcissus. Umbellate species have a fleshy racemose inflorescence (unbranched, with short floral stalks) with 2 to 15 or 20 flowers, such as N. papyraceus (see illustration, left) and N. tazetta (see Table I). The flower arrangement on the inflorescence may be either with (pedicellate) or without (sessile) floral stalks.
Prior to opening, the flower buds are enveloped and protected in a thin dry papery or membranous (scarious) spathe. The spathe consists of a singular bract that is ribbed, and which remains wrapped around the base of the open flower. As the bud grows, the spathe splits longitudinally. Bracteoles are small or absent.
The flowers of Narcissus are hermaphroditic (bisexual), have three parts (tripartite), and are sometimes fragrant (see Fragrances). The flower symmetry is actinomorphic (radial) to slightly zygomorphic (bilateral) due to declinate-ascending stamens (curving downwards, then bent up at the tip). Narcissus flowers are characterised by their, usually conspicuous, corona (trumpet).
The three major floral parts (in all species except N. cavanillesii in which the corona is virtually absent - Table I: Section Tapeinanthus) are;
(i) the proximal floral tube (hypanthium),
(ii) the surrounding free tepals, and
(iii) the more distal corona (paraperigon, paraperigonium).
All three parts may be considered to be components of the perianth (perigon, perigonium). The perianth arises above the apex of the inferior ovary, its base forming the hypanthial floral tube.
The floral tube is formed by fusion of the basal segments of the tepals (proximally connate). Its shape is from an inverted cone (obconic) to funnel-shaped (funneliform) or cylindrical, and is surmounted by the more distal corona. Floral tubes can range from long and narrow sections Apodanthi and Jonquilla to rudimentary (N. cavanillesii).
Surrounding the floral tube and corona and reflexed (bent back) from the rest of the perianth are the six spreading tepals or floral leaves, in two whorls which may be distally ascending, reflexed (folded back), or lanceolate. Like many monocotyledons, the perianth is homochlamydeous, which is undifferentiated into separate calyx (sepals) and corolla (petals), but rather has six tepals. The three outer tepal segments may be considered sepals, and the three inner segments petals. The transition point between the floral tube and the corona is marked by the insertion of the free tepals on the fused perianth.
The corona, or paracorolla, is variously described as bell-shaped (funneliform, trumpet), bowl-shaped (cupular, crateriform, cup-shaped) or disc-shaped with margins that are often frilled, and is free from the stamens. Rarely is the corona a simple callose (hardened, thickened) ring. The corona is formed during floral development as a tubular outgrowth from stamens which fuse into a tubular structure, the anthers becoming reduced. At its base, the fragrances which attract pollinators are formed. All species produce nectar at the top of the ovary. Coronal morphology varies from the tiny pigmented disk of N. serotinus (see Table I) or the rudimentary structure in N. cavanillesii to the elongated trumpets of section Pseudonarcissus (trumpet daffodils, Table I).
While the perianth may point forwards, in some species such as N. cyclamineus it is folded back (reflexed, see illustration, left), while in some other species such as N. bulbocodium (Table I), it is reduced to a few barely visible pointed segments with a prominent corona.
The colour of the perianth is white, yellow or bicoloured, with the exception of the night flowering N. viridiflorus which is green. In addition the corona of N. poeticus has a red crenulate margin (see Table I). Flower diameter varies from 12 mm (N. bulbocodium) to over 125 mm (N. nobilis=N. pseudonarcissus subsp. nobilis).
Flower orientation varies from pendent or deflexed (hanging down) as in N. triandrus (see illustration, left), through declinate-ascendant as in N. alpestris = N. pseudonarcissus subsp. moschatus, horizontal (patent, spreading) such as N. gaditanus or N. poeticus, erect as in N. cavanillesii, N. serotinus and N. rupicola (Table I), or intermediate between these positions (erecto-patent).
The flowers of Narcissus demonstrate exceptional floral diversity and sexual polymorphism, primarily by corona size and floral tube length, associated with pollinator groups (see for instance Figs. 1 and 2 in Graham and Barrett). Barrett and Harder (2005) describe three separate floral patterns;
"Daffodil" form
"Paperwhite" form
"Triandrus" form.
The predominant patterns are the 'daffodil' and 'paperwhite' forms, while the "triandrus" form is less common. Each corresponds to a different group of pollinators (See Pollination).
The "daffodil" form, which includes sections Pseudonarcissus and Bulbocodium, has a relatively short, broad or highly funnelform tube (funnel-like), which grades into an elongated corona, which is large and funnelform, forming a broad, cylindrical or trumpet-shaped perianth. Section Pseudonarcissus consists of relatively large flowers with a corolla length of around 50 mm, generally solitary but rarely in inflorescences of 2–4 flowers. They have wide greenish floral tubes with funnel-shaped bright yellow coronas. The six tepals sometimes differ in colour from the corona and may be cream coloured to pale yellow.
The "paperwhite" form, including sections Jonquilla, Apodanthi and Narcissus, has a relatively long, narrow tube and a short, shallow, flaring corona. The flower is horizontal and fragrant.
The "triandrus" form is seen in only two species, N. albimarginatus (a Moroccan endemic) and N. triandrus. It combines features of both the "daffodil" and "paperwhite" forms, with a well-developed, long, narrow tube and an extended bell-shaped corona of almost equal length. The flowers are pendent.
Androecium
There are six stamens in one to two rows (whorls), with the filaments separate from the corona, attached at the throat or base of the tube (epipetalous), often of two separate lengths, straight or declinate-ascending (curving downwards, then upwards). The anthers are basifixed (attached at their base).
Gynoecium
The ovary is inferior (below the floral parts) and trilocular (three chambered) and there is a pistil with a minutely three lobed stigma and filiform (thread like) style, which is often exserted (extending beyond the tube).
Fruit
The fruit consists of dehiscent loculicidal capsules (splitting between the locules) that are ellipsoid to subglobose (almost spherical) in shape and are papery to leathery in texture.
Seeds
The fruit contains numerous subglobose seeds which are round and swollen with a hard coat, sometimes with an attached elaiosome. The testa is black and the pericarp dry.
Most species have 12 ovules and 36 seeds, although some species such as N. bulbocodium have more, up to a maximum of 60. Seeds take five to six weeks to mature. The seeds of sections Jonquilla and Bulbocodium are wedge-shaped and matte black, while those of other sections are ovate and glossy black. A gust of wind or contact with a passing animal is sufficient to disperse the mature seeds.
Chromosomes
Chromosome numbers include 2n=14, 22, 26, with numerous aneuploid and polyploid derivatives. The basic chromosome number is 7, with the exception of N. tazetta, N. elegans and N. broussonetii in which it is 10 or 11; this subgenus (Hermione) was in fact characterised by this characteristic. Polyploid species include N. papyraceus (4x=22) and N. dubius (6x=50).
Phytochemistry
Alkaloids
As with all Amarylidaceae genera, Narcissus contains unique isoquinoline alkaloids. The first alkaloid to be identified was lycorine, from N. pseudonarcissus in 1877. These are considered a protective adaptation and are utilised in the classification of species. Nearly 100 alkaloids have been identified in the genus, about a third of all known Amaryllidaceae alkaloids, although not all species have been tested. Of the nine alkaloid ring types identified in the family, Narcissus species most commonly demonstrate the presence of alkaloids from within the Lycorine (lycorine, galanthine, pluviine) and Homolycorine (homolycorine, lycorenine) groups. Hemanthamine, tazettine, narciclasine, montanine and galantamine alkaloids are also represented. The alkaloid profile of any plant varies with time, location, and developmental stage. Narcissus also contain fructans and low molecular weight glucomannan in the leaves and plant stems.
Fragrances
Fragrances are predominantly monoterpene isoprenoids, with a small amount of benzenoids, although N. jonquilla has both equally represented. Another exception is N. cuatrecasasii which produces mainly fatty acid derivatives. The basic monoterpene precursor is geranyl pyrophosphate, and the commonest monoterpenes are limonene, myrcene, and trans-β-ocimene. Most benzenoids are non-methoxylated, while a few species contain methoxylated forms (ethers), e.g. N. bujei. Other ingredient include indole, isopentenoids and very small amounts of sesquiterpenes. Fragrance patterns can be correlated with pollinators, and fall into three main groups (see Pollination).
The taxonomy of Narcissus is complex, and still not fully resolved. Known to the ancients, the genus name appears in Graeco-Roman literature, although their interest was as much medicinal as botanical. It is unclear which species the ancients were familiar with. Although frequently mentioned in Mediaeval and Renaissance texts it was not formally described till the work of Linnaeus in 1753. By 1789 it had been grouped into a family (Narcissi) but shortly thereafter this was renamed Amaryllideae, from which comes the modern placement within Amaryllidaceae, although for a while it was considered part of Liliaceae.
Many of the species now considered to be Narcissus were in separate genera during the nineteenth century, and the situation was further confused by the inclusion of many cultivated varieties. By 1875 the current circumscription was relatively settled. By 2004 phylogenetic studies had allowed the place of Narcissus within its fairly large family to be established, nested within a series of subfamilies (Amaryllidoideae) and tribes (Narcisseae). It shares its position in the latter tribe with Sternbergia.
The infrageneric classification has been even more complex and many schemes of subgenera, sections, subsections and series have been proposed, although all had certain similarities. Most authorities now consider there to be 10 – 11 sections based on phylogenetic evidence. The problems have largely arisen from the diversity of the wild species, frequent natural hybridisation and extensive cultivation with escape and subsequent naturalisation. The number of species has varied anywhere from 16 to nearly 160, but is probably around 50 – 60.
The genus appeared some time in the Late Oligocene to Early Miocene eras, around 24 million years ago, in the Iberian peninsula. While the exact origin of the word Narcissus is unknown it is frequently linked to its fragrance which was thought to be narcotic, and to the legend of the youth of that name who fell in love with his reflection. In the English language the common name Daffodil appears to be derived from the Asphodel with which it was commonly compared.
Early
Narcissus was first described by Theophrastus (Θεόφραστος, c 371 - c 287 BC) in his Historia Plantarum (Greek: Περὶ φυτῶν ἱστορία) as νάρκισσος, referring to N. poeticus, but comparing it to Asphodelus (ασφοδελωδες). Theophrastus' description was frequently referred to at length by later authors writing in Latin such as Pliny the Elder (Gaius Plinius Secundus, 23 AD – 79 AD) from whom came the Latin form narcissus (see also Culture). Pliny's account is from his Natural History (Latin: Naturalis Historia). Like his contemporaries, his interests were as much therapeutic as botanical. Another much-cited Greek authority was Dioscorides (Διοσκουρίδης, 40 AD – 90 AD) in his De Materia Medica (Greek: Περὶ ὕλης ἰατρικῆς). Both authors were to remain influential until at least the Renaissance, given that their descriptions went beyond the merely botanical, to the therapeutic (see also Antiquity).
An early European reference is found in the work of Albert Magnus (c. 1200 – 1280), who noted in his De vegetabilibus et plantis the similarity to the leek. William Turner in his A New Herball (1551) cites all three extensively in his description of the plant and its properties.It was to remain to Linnaeus in 1753 to formally describe and name Narcissus as a genus in his Species Plantarum, at which time there were six known species (N. poeticus, N. pseudonarcissus, N. bulbocodium, N. serotinus, N. jonquilla and N. tazetta).[1] At that time, Linnaeus loosely grouped it together with 50 other genera into his Hexandria monogynia.
Modern
It was de Jussieu in 1789 who first formally created a 'family' (Narcissi), as the seventh 'Ordo' (Order) of the third class (Stamina epigyna) of Monocots in which Narcissus and 15 other genera were placed. The use of the term Ordo at that time was closer to what we now understand as Family, rather than Order. The family has undergone much reorganisation since then, but in 1805 it was renamed after a different genus in the family, Amaryllis, as 'Amaryllideae' by Jaume St.-Hilaire and has retained that association since. Jaume St.-Hilaire divided the family into two unnamed sections and recognised five species of Narcissus, omitting N. serotinus.
De Candolle brought together Linnaeus' genera and Jussieau's families into a systematic taxonomy for the first time, but included Narcissus (together with Amaryllis) in the Liliaceae in his Flore française (1805-1815) rather than Amaryllidaceae, a family he had not yet recognised. Shortly thereafter he separated the 'Amaryllidées' from 'Liliacées' (1813), though attributing the term to Brown's 'Amaryllideae' in the latter's Prodromus (1810) rather than St.-Hilaire's 'Amaryllidées'. He also provided the text to the first four volumes of Redouté illustrations in the latter's Les liliacées between 1805 and 1808 (see illustration here of N. candidissimus).
Historically both wide and narrow interpretations of the genus have been proposed. In the nineteenth century genus splitting was common, favouring the narrow view. Haworth (1831) using a narrow view treated many species as separate genera, as did Salisbury (1866). These authors listed various species in related genera such as Queltia (hybrids), Ajax (=Pseudonarcissus) and Hermione (=Tazettae), sixteen in all in Haworth's classification. In contrast, Herbert (1837) took a very wide view reducing Harworth's sixteen genera to six. Herbert, treating the Amaryllidacea as an 'order' as was common then, considered the narcissi to be a suborder, the Narcisseae, the six genera being Corbularia, Ajax, Ganymedes, Queltia, Narcissus and Hermione and his relatively narrow circumscription of Narcissus having only three species. Later Spach (1846) took an even wider view bringing most of Harworth's genera into the genus Narcissus, but as separate subgenera. By the time that Baker (1875) wrote his monograph all of the genera with one exception were included as Narcissus. The exception was the monotypic group Tapeinanthus which various subsequent authors have chosen to either exclude (e.g. Cullen 1986) or include (e.g. Webb 1978, 1980). Today it is nearly always included.
The eventual position of Narcissus within the Amaryllidaceae family only became settled in the twenty-first century with the advent of phylogenetic analysis and the Angiosperm Phylogeny Group system. The genus Narcissus belongs to the Narcisseae tribe, one of 13 within the Amaryllidoideae subfamily of the Amaryllidaceae. It is one of two sister clades corresponding to genera in the Narcisseae, being distinguished from Sternbergia by the presence of a paraperigonium, and is monophyletic
The infrageneric phylogeny of Narcissus still remains relatively unsettled. The taxonomy has proved very complex and difficult to resolve, particularly for the Pseudonarcissus group. This is due to a number of factors, including the diversity of the wild species, the ease with which natural hybridisation occurs, and extensive cultivation and breeding accompanied by escape and naturalisation.
De Candolle, in the first systematic taxonomy of Narcissus, arranged the species into named groups, and those names (Faux-Narcisse or Pseudonarcissus, Poétiques, Tazettes, Bulbocodiens, Jonquilles) have largely endured for the various subdivisions since and bear his name. The evolution of classification was confused by including many unknown or garden varieties, until Baker (1875) made the important distinction of excluding all specimens except the wild species from his system. He then grouped all of the earlier related genera as sections under one genus, Narcissus, the exception being the monotypic Tapeinanthus. Consequently, the number of accepted species has varied widely.
A common modern classification system has been that of Fernandes (1951, 1968, 1975) based on cytology, as modified by Blanchard (1990) and Mathew (2002), although in some countries such as Germany, the system of Meyer (1966) was preferred. Fernandes described two subgenera based on basal chromosome number, Hermione, n = 5 (11) and Narcissus, n = 7 (13). He further subdivided these into ten sections (Apodanthi, Aurelia, Bulbocodii, Ganymedes, Jonquillae, Narcissus, Pseudonarcissi, Serotini, Tapeinanthus, Tazettae), as did Blanchard later.
In contrast to Fernandes, Webb's treatment of the genus for the Flora Europaea (1978, 1980) prioritised morphology over genetics, and abandoned the subgenera ranks. He also restored De Candolle's original nomenclature, and made a number of changes to section Jonquilla, merging the existing subsections, reducing Apodanthi to a subsection of Jonquilla, and moving N. viridiflorus from Jonquilla to a new monotypic section of its own (Chloranthi). Finally, he divided Pseudonarcissus into two subsections. Blanchard (1990), whose Narcissus: a guide to wild daffodils has been very influential, adopted a simple approach, restoring Apodanthae, and based largely on ten sections alone.
The Royal Horticultural Society (RHS) currently lists ten sections, based on Fernandes (1968), three of which are monotypic (contain only one species), while two others only containing two species. Most species are placed in Pseudonarcissus While infrageneric groupings within Narcissus have been relatively constant, their status (genera, subgenera, sections, subsections, series, species) has not. Some authors treat some sections as being further subdivided into subsections, e.g. Tazettae (3 subsections). These subdivisions correspond roughly to the popular names for narcissi types, e.g. Trumpet Daffodils, Tazettas, Pheasant's Eyes, Hoop Petticoats, Jonquils.
While Webb had simply divided the genus into sections, Mathew found this unsatisfactory, implying every section had equal status. He adapted both Fernandes and Webb to devise a more hierarchical scheme he believed better reflected the interrelationships within the genus. Mathew's scheme consists of three subgenera (Narcissus, Hermione and Corbularia). The first two subgenera were then divided into five and two sections respectively. He then further subdivided two of the sections (subgenus Narcissus section Jonquillae, and subgenus Hermione section Hermione) into three subsections each. Finally, he divided section Hermione subsection Hermione further into two series, Hermione and Albiflorae. While lacking a phylogenetic basis, the system is still in use in horticulture. For instance the Pacific Bulb Society uses his numbering system for classifying species.
The phylogenetic analysis of Graham and Barrett (2004) supported the infrageneric division of Narcissus into two clades corresponding to the subgenera Hermione and Narcissus, but does not support monophyly of all sections, with only Apodanthi demonstrating clear monophyly, corresponding to Clade III of Graham and Barrett, although some other clades corresponded approximately to known sections. These authors examined 36 taxa of the 65 listed then, and a later extended analysis by Rønsted et al. (2008) with five additional taxa confirmed this pattern.
A very large (375 accessions) molecular analysis by Zonneveld (2008) utilising nuclear DNA content sought to reduce some of the paraphyly identified by Graham and Barrett. This led to a revision of the sectional structure, shifting some species between sections, eliminating one section and creating two new ones. In subgenus Hermione, Aurelia was merged with Tazettae. In subgenus Narcissus section Jonquillae subsection Juncifolii was elevated to sectional rank, thus resolving the paraphyly in this section observed by Graham and Barrett in Clade II due to this anomalous subsection, the remaining species being in subsection Jonquillae, which was monophyletic. The relatively large section Pseudonarcissi was divided by splitting off a new section, Nevadensis (species from southern Spain) leaving species from France, northern Spain and Portugal in the parent section. At the same time Fernández-Casas (2008) proposed a new monotypic section Angustini to accommodate Narcissus deficiens, placing it within subgenus Hermione.
While Graham and Barrett (2004) had determined that subgenus Hermione was monophyletic, using a much larger accession Santos-Gally et al. (2011) did not. However the former had excluded species of hybrid origins, while the latter included both N. dubius and N. tortifolius. If these two species are excluded (forming a clade with subgenus Narcissus) then Hermione can be considered monophyletic, although as a section of Hermione, Tazettae is not monophyletic. They also confirmed the monophyly of Apodanthi.
Some so-called nothosections have been proposed, predominantly by Fernández-Casas, to accommodate natural ('ancient') hybrids (nothospecies).
Subgenera and sections
Showing revisions by Zonnefeld (2008)
subgenus Hermione (Haw.) Spach.
(Aurelia (Gay) Baker (monotypic) - merged with Tazettae (2008)
Serotini Parlatore (2 species)
Tazettae de Candolle (16 species) syn. Hermione (Salisbury) Sprengel, in Fernandes' scheme. Incorporating Aurelia (2008)
subgenus Narcissus L.
Apodanthi A. Fernandes (6 species)
Bulbocodium de Candolle (11 species)
Ganymedes (Haworth) Schultes f. (monotypic)
Jonquillae de Candolle (8 species)
Juncifolii (A. Fern.) Zonn. sect. nov. (2008)
Narcissus L. (2 species)
Nevadensis Zonn. sect. nov. (2008)
Pseudonarcissus de Candolle (36 species) Trumpet daffodils
Tapeinanthus (Herbert) Traub (monotypic)
Species
Estimates of the number of species in Narcissus have varied widely, from anywhere between 16 and nearly 160, even in the modern era. Linnaeus originally included six species in 1753. By the time of the 14th edition of the Systema Naturae in 1784, there were fourteen. The 1819 Encyclopaedia Londinensis lists sixteen (see illustration here of three species) and by 1831 Adrian Haworth had described 150 species.
Much of the variation lies in the definition of species, and whether closely related taxa are considered separate species or subspecies. Thus, a very wide view of each species, such as Webb's results in few species, while a very narrow view such as that of Fernandes results in a larger number. Another factor is the status of hybrids, given natural hybridisation, with a distinction between 'ancient hybrids' and 'recent hybrids'. The term 'ancient hybrid' refers to hybrids found growing over a large area, and therefore now considered as separate species, while 'recent hybrid' refers to solitary plants found amongst their parents, with a more restricted range.
In the twentieth century Fernandes (1951) accepted 22 species, on which were based the 27 species listed by Webb in the 1980 Flora Europaea. By 1968, Fernandes had accepted 63 species, and by 1990 Blanchard listed 65 species, and Erhardt 66 in 1993. In 2006 the Royal Horticultural Society's (RHS) International Daffodil Register and Classified List listed 87 species, while Zonneveld's genetic study (2008) resulted in only 36. As of September 2014, the World Checklist of Selected Plant Families accepts 52 species, along with at least 60 hybrids, while the RHS has 81 accepted names in its October 2014 list.
Evolution
Within the Narcisseae, Narcissus (western Mediterranean) diverged from Sternbergia (Eurasia) some time in the Late Oligocene to Early Miocene eras, around 29.3–18.1 Ma, with a best estimate of 23.6 Ma. Later the genus divided into the two subgenera (Hermione and Narcissus) between 27.4 and 16.1 Ma (21.4 Ma). The divisions between the sections of Hermione then took place during the Miocene period 19.9–7.8 Ma.
Narcissus appears to have arisen in the area of the Iberian peninsula, southern France and northwestern Italy, and within this area most sections of the genus appeared, with only a few taxa being dispersed to North Africa at a time when the African and West European platforms were closer together. Subgenus Hermione in turn arose in the southwestern mediterranean and north west Africa. However, these are reconstructions, the Amaryllidaceae lacking a fossil record.
Names and etymology
The derivation of the Latin narcissus (Greek: νάρκισσος) is unknown. It may be a loanword from another language; for instance, it is said to be related to the Sanskrit word nark, meaning 'hell'. It is frequently linked to the Greek myth of Narcissus described by Ovid in his Metamorphoses, who became so obsessed with his own reflection that as he knelt and gazed into a pool of water, he fell into the water and drowned. In some variations, he died of starvation and thirst. In both versions, the narcissus plant sprang from where he died. Although Ovid appeared to describe the plant we now know as Narcissus there is no evidence for this popular derivation, and the person's name may have come from the flower's name. The Poet's Narcissus (N. poeticus), which grows in Greece, has a fragrance that has been described as intoxicating. This explanation is largely discredited due to lack of proof. Pliny wrote that the plant 'narce narcissum dictum, non a fabuloso puero' ('named narcissus from narce, not from the legendary youth'), i.e. that it was named for its narcotic properties (ναρκάω narkao, 'I grow numb' in Greek), not from the legend. Furthermore, there were accounts of narcissi growing, such as in the legend of Persephone, long before the story of Narcissus appeared (see Greek culture). It has also been suggested that daffodils bending over streams evoked the image of the youth admiring his own reflection in the water.
Linnaeus used the Latin name for the plant in formally describing the genus, although Matthias de l'Obel had previously used the name in describing various species of Narcissi in his Icones stirpium of 1591, and other publications, as had Clusius in Rariorum stirpium (1576).
The plural form of the common name narcissus has caused some confusion. British English sources such as the Shorter Oxford English Dictionary give two alternate forms, narcissi and narcissuses. In contrast, in American English the Merriam-Webster Dictionary provides for a third form, narcissus, used for both singular and plural. The Oxford dictionaries only list this third form under American English, although the Cambridge Dictionary allows of all three in the same order. However, Garner's Modern American Usage states that narcissi is the commonest form, narcissuses being excessively sibilant. For similar reasons, Fowler prefers narcissi in British English usage. Neither support narcissus as a plural form. Common names such as narcissus do not capitalise the first letter in contrast to the person of that name and the Latin genus name.
The name Narcissus (feminine Narcissa) was not uncommon in Roman times, such as Tiberius Claudius Narcissus, a Roman official in Claudius' time, an early New Testament Christian in Rome and later bishops and saints.
Daffodil
The word daffodil was unknown in the English language before the sixteenth century. The name is derived from an earlier affodell, a variant of asphodel. In classical Greek literature the narcissus is frequently referred to as the asphodel, such as the meadows of the Elysian fields in Homer (see Antiquity). Asphodel in turn appears to be a loanword coming from French via Mediaeval Latin affodilus from Classical Latin asphodilus and ultimately the Greek asphodelos (Greek: ἀσφόδελος). The reason for the introduction of the initial d is not known, although a probable source is an etymological merging from the Dutch article de, as in de affodil, or English the, as th'affodil or t'affodil, hence daffodil, and in French de and affodil to form fleur d'aphrodille and daphrodille.
From at least the 16th century, daffadown dilly and daffydowndilly have appeared as playful synonyms of the name. In common parlance and in historical documents, the term daffodil may refer specifically to populations or specimens of the wild daffodil, N. pseudonarcissus. H. N. Ellacombe suggests this may be from Saffon Lilly, citing Prior in support, though admittedly conjectural.
Lady Wilkinson (1858), who provides an extensive discussion of the etymology of the various names for this plant, suggests a very different origin, namely the Old English word affodyle (that which cometh early), citing a 14th-century (but likely originally much earlier) manuscript in support of this theory, and which appears to describe a plant resembling the daffodil. Ellacombe provides further support for this from a fifteenth century English translation of Palladius that also refers to it.
Jonquil
The name jonquil is said to be a corruption via French from the Latin juncifolius meaning 'rush-leaf' (Juncaceae) and its use is generally restricted to those species and cultivars which have rush like leaves, e.g. N. juncifolius.
Other
A profusion of names have attached themselves in the English language, either to the genus as a whole or to individual species or groups of species such as sections. These include narcissus, jonquil, Lent lily, Lenten lily, lide lily, yellow lily, wort or wyrt, Julians, glens, Lent cocks, corn flower, bell rose, asphodel, Solomon's lily, gracy day, haverdrils, giggary, cowslip, and crow foot.
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Femme Hmong Noir, marché de Sapa, nord du Vietnam
Avec la ligne de train et désormais l'autoroute la reliant à Hanoï, Sapa est devenue une destination touristique très fréquentée et a beaucoup perdu de son charme, même si les promenades dans les rizières alentour restent bien agréables. Le petit marché permanent sous la halle permet de croiser encore de pittoresques personnages telle cette femme dans son costume Hmong Noir
Avec les Dao Rouge, les Hmong Noir constituent l'une des ethnies les mieux représentées dans la région de Sapa.
Les Hmong encore appelés Méo, ou Miao , sont originaires des régions montagneuses du sud de la Chine (principalement la province du Guizhou), où ils sont encore présents ainsi qu'au nord du Viêt Nam et du Laos.
Le souci de préserver leur identité culturelle et leur indépendance les ont amenés à s'engager dans divers conflits. Au XXe siècle, en particulier, ils aidèrent les Français pendant la guerre d'Indochine puis les Américains pendant la la guerre du Vietnam. A l’avènement des régimes communistes dans ces pays un nombre important de Hmong se sont réfugiés dans des pays d'accueil, principalement les États-Unis, la France et l'Australie. Mais la majeure partie d’entre eux vit encore en Asie du Sud-Est
Les Hmongs sont animistes ou chrétiens. La langue hmong appartient à la famille des langues hmong-mien, encore appelée « miao-yao »
Les costumes traditionnels de cette ethnie sont très polymorphes mais ils ont en commun la richesse du décor brodé
Polymorph's head was hand cast by me so she is green right the way through! Yay! She didn't turn out too badly I think :) What colour should I make next? :P Her body is an obitsu :)
Juvenile Hepatic bird (brown/red) and Grey Morph at old Newcastle train station,
East Coast Nature Reserve (ECNR) 30-07-/-01-08-2016
Newcastle County Wicklow
Nikon D7000 Camera & Sigma 150-600 mm f/5.0-6.3 Sport Lens
Scientific classification
Domain:Eukaryota
Kingdom:Animalia
Phylum:Chordata
Class:Aves
Order:Cuculiformes
Family:Cuculidae
Genus:Cuculus
Species:C. canorus
Binomial name
Cuculus canorus | [UK] Cuckoo | [FR] Coucou gris | [DE] Kuckuck | [ES] Cuco Europeo | [IT] Cuculo eurasiatico | [NL] Koekoek | [IRL] Cuach
Status: Widespread summer visitor to Ireland from April to August.
Conservation Concern: Green-listed in Ireland. The European population is currently evaluated as secure.
Identification: Despite its obvious song, relatively infrequently seen. In flight, can be mistaken for a bird of prey such as Sparrowhawk, but has rapid wingbeats below the horizontal plane - ie. the wings are not raised above the body. Adult male Cuckoos are a uniform grey on the head, neck, back, wings and tail. The underparts are white with black barring. Adult females can appear in one of two forms. The so-called grey-morph resembles the adult male plumage, but has throat and breast barred black and white with yellowish wash. The rufous-morph has the grey replaced by rufous, with strong black barring on the wings, back and tail. Juvenile Cuckoos resemble the female rufous-morph, but are darker brown above.
Similar Species: Sparrowhawk
Call: The song is probably one of the most recognisable and well-known of all Irish bird species. The male gives a distinctive “wuck-oo”, which is occasionally doubled “wuck-uck-ooo”. The female has a distinctive bubbling “pupupupu”. The song period is late April to late June.
Diet: Mainly caterpillars and other insects.
Breeding: Widespread in Ireland, favouring open areas which hold their main Irish host species – Meadow Pipit. Has a remarkable breeding biology unlike any other Irish breeding species.
Wintering: Cuckoos winter in central and southern Africa.
To minimise the chance of being recognised and thus attacked by the birds they are trying to parasitize, female cuckoos have evolved different guises.
The common cuckoo (Cuculus canorus) lays its eggs in the nests of other birds. On hatching, the young cuckoo ejects the host's eggs and chicks from the nest, so the hosts end up raising a cuckoo chick rather than a brood of their own. To fight back, reed warblers (a common host across Europe) have a first line of defence: they attack, or ‘mob’, the female cuckoo, which reduces the chance that their nest is parasitized.
To deter the warbler from attacking, the colouring of the grey cuckoo mimics sparrow hawks, a common predator of reed warblers. However, other females are bright rufous (brownish-red). The presence of alternate colour morphs in the same species is rare in birds, but frequent among the females of parasitic cuckoo species. The new research shows that this is another cuckoo trick: cuckoos combat reed warbler mobbing by coming in different guises.
In the study, the researchers manipulated local frequencies of the more common grey colour cuckoo and the less common (in the United Kingdom) rufous colour cuckoo by placing models of the birds at neighbouring nests. They then recorded how the experience of watching their neighbours mob changed reed warbler responses to both cuckoos and a sparrow hawk at their own nest.
They found that reed warblers increased their mobbing, but only to the cuckoo morph that their neighbours had mobbed. Therefore, as one cuckoo morph increases in frequency, local host populations will become alerted specifically to that morph. This means the alternate morph will be more likely to slip past host defences and lay undetected. This is the first time that ‘social learning’ has been documented in the evolution of mimicry as well as the evolution of different observable characteristics - such as colour - in the same species (called polymorphism).
From the University of Cambridge “When mimicry becomes less effective, evolving to look completely different can be a successful trick. Our research shows that individuals assess disguises not only from personal experience, but also by observing others. However, because their learning is so specific, this social learning then selects for alternative cuckoo disguises and the arms race continues.”.
“It’s well known that cuckoos have evolved various egg types which mimic those of their hosts in order to combat rejection. This research shows that cuckoos have also evolved alternate female morphs to sneak through the hosts' defences. This explains why many species which use mimicry, such as the cuckoo, evolve different guises.”
A mineral is a naturally-occurring, solid, inorganic, crystalline substance having a fairly definite chemical composition and having fairly definite physical properties. At its simplest, a mineral is a naturally-occurring solid chemical. Currently, there are over 6100 named and described minerals - about 200 of them are common and about 20 of them are very common. Mineral classification is based on anion chemistry. Major categories of minerals are: elements, sulfides, oxides, halides, carbonates, sulfates, phosphates, and silicates.
Elements are fundamental substances of matter - matter that is composed of the same types of atoms. At present, 118 elements are known (four of them are still unnamed). Of these, 98 occur naturally on Earth (hydrogen to californium). Most of these occur in rocks & minerals, although some occur in very small, trace amounts. Only some elements occur in their native elemental state as minerals.
To find a native element in nature, it must be relatively non-reactive and there must be some concentration process. Metallic, semimetallic (metalloid), and nonmetallic elements are known in their native state.
The element carbon occurs principally in its native state as graphite (C) and diamond (C). Graphite is the common & far less valuable polymorph of carbon. A scarce polymorph of carbon is diamond. The physical properties of diamond and graphite couldn’t be more different, considering they have the same chemistry. Diamond has a nonmetallic, adamantine luster, typically occurs in cubic or octahedral (double-pyramid) crystals, or subspherical to irregularly-shaped masses, and is extremely hard (H≡10). Diamonds can be almost any color, but are typically clearish, grayish, or yellowish. Many diamonds are noticeably fluorescent under black light (ultraviolet light), but the color and intensity of fluorescence varies. Some diamonds are phosphorescent - under certain conditions, they glow for a short interval on their own.
Very rarely, diamond is a rock-forming mineral (see diamondite - www.flickr.com/photos/jsjgeology/14618393527).
The rough diamonds seen here are from a Cretaceous-aged lamproite body at Crater of Diamonds State Park in Arkansas. At the site, diamondiferous lamproitic rocks of the Prairie Creek Lamproite are exposed (actually, it’s just a field of rocky dirt). Diamonds are found regularly by tourists. The Prairie Creek Lamproite is an ultramafic pipe that intruded Proterozoic-aged (Grenvillian) basement rocks. Prairie Creek rocks are Cretaceous in age and date to about 106 million years old.
A strange myth has emerged over the years that says Arkansas diamonds are harder than diamonds from elsewhere. This is not true. Diamonds have a hardness of 10 on the Mohs Scale, whether they're from Arkansas or not.
--------------------------------------------
From park signage:
THE “CRATER” PIPE
The “Crater” diamond deposits are the result of a series of volcanic eruptions that carried a diamond-bearing magma called peridotite to the surface. This molten rock slowly cooled, forming a solid, vertical shaft or pipe.
Later, secondary eruptions occurred that shattered much of the peridotite. The fragmented mixture left by the eruptions formed a volcanic breccia that comprises the majority of the searching area. The intrusive peridotite, and a tuff and fine-grained breccia, also formed by the eruptions, cover the remainder of the mine surface.
STRATIGRAPHIC PROFILE
The volcanic eruptions forming the “Crater’s” pipe had to force their way upward through the layers of rock and sediment formed during Paleozoic and Mesozoic times.
Diamonds are located throughout the pipe - from the surface to an undetermined depth (possibly several thousand meters). The mine is simply the exposed surface area of the pipe.
TOPOGRAPHY
Surface features at the “Crater” today reflect the depositional, erosional, and weathering processes which have occurred since the pipe’s formation. Erosional forces, such as wind and water, have erased the original surface features of the pipe and surrounding rock formations.
Most of the exposed volcanic rocks tend to weather rapidly forming soils of various colors. It is in these soils that we find diamonds today.
In 1948, Glen L. Martin leased property to build a diamond processing plant. Martin’s plant operation lasted only eight months after recovering less than 250 carats of diamonds. The original location of the Martin Plant has been preserved. Its location lies just south of the Visitor Center.
DAZZLING DIAMONDS FROM THE CRATER
Over 85,000 Crater diamonds have been found. Each year an average of 600 are found.
1924 - The Uncle Sam, 40.23 carats
1956 - The Star of Arkansas, 15.33 carats
1956 - The Eisenhower, 6.11 carats
1960 - The Gary Moore, 6.43 carats
Over 24,000 diamonds have been found since Crater of Diamonds State Park was established in 1972.
1975 - The Amarillo Starlight, 163.7 carats
1978 - The Lamle Diamond, 8.61 carats
1981 - The Star of Shreveport, 8.82 carats
1986 - The Conell Diamond, 7.95 carats
1990 - The Strawn-Wagner, 3.03 carats
1997 - The Cooper Diamonds, 6.72 & 6.0 carats
1998 - The Dickinson/Stevens, 7.28 carats
An end to a mystery
Since 1906, when John Huddleston discovered the first diamonds in this field, people have wondered what was beneath this green soil surface. Just how deep and how large is this diamond deposit?
From 1990 to 1997, Arkansas State Parks and several international commercial mining companies conducted tests to determine the underground size and shape of the diamond deposit, and to evaluate the grade of diamonds from the mine.
Phase I testing determined that this is the world’s 8th largest diamond deposit, in size and shape. It extends over 660 feet into the earth. In phase II, 210 diamonds totaling 45.7 carats were recovered. This verified that a significant number of diamonds occur within this large deposit. However, the grade was less than commercially profitable at the time and interest in further testing diminished.
--------------------------------------------
Geologic provenance: Prairie Creek Lamproite (Prairie Creek Pipe), mid-Albian Stage, late Early Cretaceous, ~106 Ma
Locality: Crater of Diamonds State Park, just south of the town of Murfreesboro, Pike County, southwestern Arkansas, USA (34° 01’ 55” to 34° 02’ 05” North latitude, 93° 40’ 18” to 93° 40’ 25” West longitude)
Femme Dao Longue Tunique sur le marché de Quyet Tien, petite ville de la région de Hagiang, nord du Vietnam
Les Dao (ou Dzao) sont, comme les Hmongs, très nombreux et l'ethnie est très polymorphe : Dao Rouge, Dao à Tunique, Dao Noir, Dao à pantalon blanc… .J'en ai déjà posté plusieurs exemples.
Emigrée de Chine du sud depuis le XIIIème siècle, l’ethnie Dao habite au Nord du Vietnam dans les provinces de la moyenne et haute région. Elle occupe des terres à toute altitude et vit en bons voisins avec d’autres ethnies comme les Hmong, Tay, Thaï, … Ses ressources principales proviennent de l’agriculture, (riz et mais)
Souvent chez les femmes, les cheveux sont relevés en chignon sur la nuque ou rasés sur le pourtour à l’exception d’une touffe au sommet de la tête
Pratiquant le culte des ancêtres, les Dao adhèrent également au Taoïsme.
Les femmes Dao Tunique (ou Longues Tunique) portent donc une longue tunique noire rehaussée à la ceinture et à la base de parement rouge vif. Elles portent une coiffe colorée. On les rencontre surtout dans la région de Hagiang.
Cuculus canorus
[order] Cuculiformes | [family] Cuculidae | [latin] Cuculus canorus | [UK] Cuckoo | [FR] Coucou gris | [DE] Kuckuck | [ES] Cuco Europeo | [IT] Cuculo eurasiatico | [NL] Koekoek | [IRL] Cuach
Status: Widespread summer visitor to Ireland from April to August.
Conservation Concern: Green-listed in Ireland. The European population is currently evaluated as secure.
Identification: Despite its obvious song, relatively infrequently seen. In flight, can be mistaken for a bird of prey such as Sparrowhawk, but has rapid wingbeats below the horizontal plane - ie. the wings are not raised above the body. Adult male Cuckoos are a uniform grey on the head, neck, back, wings and tail. The underparts are white with black barring. Adult females can appear in one of two forms. The so-called grey-morph resembles the adult male plumage, but has throat and breast barred black and white with yellowish wash. The rufous-morph has the grey replaced by rufous, with strong black barring on the wings, back and tail. Juvenile Cuckoos resemble the female rufous-morph, but are darker brown above.
Similar Species: Sparrowhawk
Call: The song is probably one of the most recognisable and well-known of all Irish bird species. The male gives a distinctive “wuck-oo”, which is occasionally doubled “wuck-uck-ooo”. The female has a distinctive bubbling “pupupupu”. The song period is late April to late June.
Diet: Mainly caterpillars and other insects.
Breeding: Widespread in Ireland, favouring open areas which hold their main Irish host species – Meadow Pipit. Has a remarkable breeding biology unlike any other Irish breeding species.
Wintering: Cuckoos winter in central and southern Africa.
To minimise the chance of being recognised and thus attacked by the birds they are trying to parasitize, female cuckoos have evolved different guises.
The common cuckoo (Cuculus canorus) lays its eggs in the nests of other birds. On hatching, the young cuckoo ejects the host's eggs and chicks from the nest, so the hosts end up raising a cuckoo chick rather than a brood of their own. To fight back, reed warblers (a common host across Europe) have a first line of defence: they attack, or ‘mob’, the female cuckoo, which reduces the chance that their nest is parasitized.
To deter the warbler from attacking, the colouring of the grey cuckoo mimics sparrow hawks, a common predator of reed warblers. However, other females are bright rufous (brownish-red). The presence of alternate colour morphs in the same species is rare in birds, but frequent among the females of parasitic cuckoo species. The new research shows that this is another cuckoo trick: cuckoos combat reed warbler mobbing by coming in different guises.
In the study, the researchers manipulated local frequencies of the more common grey colour cuckoo and the less common (in the United Kingdom) rufous colour cuckoo by placing models of the birds at neighbouring nests. They then recorded how the experience of watching their neighbours mob changed reed warbler responses to both cuckoos and a sparrow hawk at their own nest.
They found that reed warblers increased their mobbing, but only to the cuckoo morph that their neighbours had mobbed. Therefore, as one cuckoo morph increases in frequency, local host populations will become alerted specifically to that morph. This means the alternate morph will be more likely to slip past host defences and lay undetected. This is the first time that ‘social learning’ has been documented in the evolution of mimicry as well as the evolution of different observable characteristics - such as colour - in the same species (called polymorphism).
From the University of Cambridge “When mimicry becomes less effective, evolving to look completely different can be a successful trick. Our research shows that individuals assess disguises not only from personal experience, but also by observing others. However, because their learning is so specific, this social learning then selects for alternative cuckoo disguises and the arms race continues.”.
“It’s well known that cuckoos have evolved various egg types which mimic those of their hosts in order to combat rejection. This research shows that cuckoos have also evolved alternate female morphs to sneak through the hosts' defences. This explains why many species which use mimicry, such as the cuckoo, evolve different guises.”
Y también en Twiter
No solo a lo largo de su vida, sino también, casi en un instante, Euglena polymorpha puede cambiar completamente de apariencia y de cuerpo de delfín. En un giro de segundos puede volverse esfera, igual que un minúsculo planeta intensamente verde, como si hubiese concentrado en su cuerpo el color teñido y turbio del agua que hoy la rodea y donde se descubre en ella por primera vez, sumándose así a las casi 40 nuevas citas de euglenófitas que desde este proyecto se han señalado para el Lago de Sanabria.
Euglena polymorpha es la alfarera de su cuerpo, moldea su masa verde, dejando un rastro pálido en su larga cola, y mientras se retuerce y se amasa avanzando, vigila atenta con su ojo el cambio de los fondos del Lago de Sanabria que como en ella todo es cambiante y verde.
Descrita en 1902 por Dangeard, es fácil confundir a esta Euglena con otras hermanas a las que se asemeja mucho, como Euglena proxima, Euglena variabilis o incluso Euglena caudata. Pero hay algo en ella que permite distinguirla con relativa facilidad y es que, aparte de su tamaño, ligeramente superior a ellas, y que puede llegar a alcanzar hasta las 90 micras de longitud y las 26 de anchura, la forma cónica de su cuerpo hace que sus contornos se vayan estrechando de forma muy progresiva hasta su extremo posterior que remata en un apéndice cónico y transparente.
Otra de las características de esta Euglena cambiante es que puede perder el flagelo con relativa facilidad y cuando esto sucede como hoy, sus movimientos reptantes y sus contorsiones suelen ser muy marcados y poco suaves.
La película que envuelve el flexible cuerpo de Euglena polymorpha es muy plástica, se puede deformar de forma intensa y puede aparecer marcada por gruesos surcos cuando se retuerce. Los numerosos cloroplastos que se encuentran en su interior y que le confieren ese aspecto tan granuloso pueden presentar un contorno relativamente regular en forma de disco o mostrar los bordes desflecados en anchos lóbulos.
En cualquier caso, como muchas otras euglenas Euglena polymorpha parece vigilante y observa todo con su intenso ojo de rubí. Euglena es alga, pero también es diminuto gusano verde. Flagelado, planta y animal a la vez con esencia de quimera. Su cuerpo, mágicamente se transforma en juego de pececillo verde a odre de plastilina.
Dentro del amplísimo grupo de las euglenas, de las que se han descrito cerca de 552, Euglena polymorpha es una de las 133 taxonómicamente aceptadas y no es de las más comunes.
Euglena polymorpha es de distribución cosmoplita, y suele habitar en diferentes masas de agua dulce, grandes y pequeñas, y hoy desde el fondo de este Lago, camuflada de verde entre estas aguas que fueron limpias como el aire.
La fotografía que mostramos, realizada a 400 aumentos empleando la técnica de contraste de interferencia, se ha tomado sobre una muestra de fondo, a cinco metros de profundidad, recogida en las inmediaciones de la Isla de Moras por Juan Carlos Feijó, Pablo, Lucas Patrick y Alberto Martínez el día 17 de junio de 2017 en el Lago de Sanabria (Zamora), desde el catamarán Helios Sanabria el primer catamarán construido en el Planeta propulsado por energía eólica y solar.
LIBRO: Lago de Sanabria 2015, presente y futuro de un ecosistema en desequilibrio
Presentación ponencia congreso internacional de Limnología de la AIL
Informes de contaminación en el Lago de Sanabria
informe de evolución de la contaminación en el Lago de Sanabria
Adult with substantial, white, dorsal line on tail continuing as a broken line towards the hump (1) over the pericardium.
North-east Scotland, February 2020. © C. Rickard.
Full SPECIES DESCRIPTION BELOW
PDF available at www.researchgate.net/publication/361247558_Coryphella_ver...
Sets of OTHER SPECIES at: www.flickr.com/photos/56388191@N08/collections/
Coryphella verrucosa (M. Sars, 1829).
Including forms Coryphella verrucosa verrucosa (M. Sars, 1829) and Coryphella verrucosa rufibranchialis (G. Johnston, 1832).
Current taxonomy: World Register of Marine Species www.marinespecies.org/aphia.php?p=taxdetails&id=139987
Synonyms: Eolidia verrucosa M. Sars, 1829; Eolis rufibranchialis G. Johnston, 1832; Coryphella rufibranchialis (G. Johnston, 1832); Coryphella rufibranchialis chocolata Balch, 1908: Coryphella pseudoverrucosa Martynov, Sanamyan & Korshunova, 2015; Flabellina verrucosa (M. Sars, 1829); Flabellina pseudoverrucosa (Martynov, Sanamyan & Korshunova, 2015).
GLOSSARY BELOW
Preface
The following description is primarily of Coryphella verrucosa rufibranchialis, the only form found in Britain except Shetland, where the Scandinavian form with radically different cerata, Coryphella verrucosa verrucosa, also occurs. The cerata of the two forms are described separately; other features are similar on both forms.
Description
The length, excluding appendages, may reach 25 mm, occasionally 35 mm. The body is translucent white, revealing white ovotestes when mature fig. 01 flic.kr/p/2nqKAan . The tail has an opaque white, substantial, dorsal line. On well grown adults the dorsal line may continue, often with breaks, as far the hump over the pericardium fig. 02 flic.kr/p/2nqLAnc , but often ( usually on Coryphella verrucosa verrucosa) the dorsum is obscured by cerata fig. 03 flic.kr/p/2nqLPHR . The dorsal line often does not extend the full length of the tail on young specimens fig. 04 flic.kr/p/2nqLPGt . On translucent, immature specimens, the rectum ending at the anus below the second group of cerata on the right, the pink jaw, the circum-oesophageal nerve ring bearing black internal eyes and the rhinophoral ganglia on nerve cords connected to the nerve ring may be discernible fig. 05 flic.kr/p/2nqKA2M .
The cerata on C. v, rufibranchialis are slender, almost linear, about as long as the width of the body, and held at varying angles, often across each other fig. 04 flic.kr/p/2nqLPGt . They are arranged in five to seven groups of diagonal rows on each side of the body fig. 01 flic.kr/p/2nqKAan & fig. 06 flic.kr/p/2nqEmdW , rising from a notal ridge which does not continue between ceratal groups fig. 12 flic.kr/p/2nqKGcQ . The digestive gland within the cerata can be various shades of red fig. 02 flic.kr/p/2nqLAnc or red-brown fig. 01 flic.kr/p/2nqKAan . A narrow, opaque white, subterminal pigment band encircles the translucent white tip containing cnidosacs fig. 01 flic.kr/p/2nqKAan & fig. 06 flic.kr/p/2nqEmdW , and there may be a translucent gap between the band and the digestive gland.
The cerata on C. v. verrucosa are stout and cylindrical with a slightly narrower base and an obtuse tip fig. 07 flic.kr/p/2nqN8kd . They are barely as long as half the width of the body and are held erect and packed densely together (Sars, 1829) fig. 03 flic.kr/p/2nqLPHR . They are attached along either side of the body, but it looks as if the whole back [apart from hump over pericardium] is occupied by them (Sars, 1829). They are so packed together that it is not possible to discern an arrangement of groups or lines (Sars, 1829). The digestive gland can be dark brown fig. 08 flic.kr/p/2nqEmco , rust-brown fig. 09 flic.kr/p/2nqLPCL or white fig. 03 flic.kr/p/2nqLPHR . The whole tip is covered by a substantial cap of opaque white, except for a small, translucent apex which, when viewed end on, can reveal the brown digestive gland fig. 09 flic.kr/p/2nqLPCL as the ‘brown spot’ recorded by Sars (1829).
The wrinkled translucent whitish rhinophores fig. 10 flic.kr/p/2nqKzXt are slightly yellowish fig. 04 flic.kr/p/2nqLPGt , sometimes pinkish, with an opaque white line along the distal third.
Translucent white oral tentacles arise from the upper surface of the head fig. 06 flic.kr/p/2nqEmdW . They have an opaque white line along the distal third fig. 04 flic.kr/p/2nqLPGt . The prominent outer parts of the mouth occupy most of the head ventrally fig. 11 flic.kr/p/2nqN8gL . The internal pink jaw can sometimes be discerned in the translucent head as a pink band in lateral view fig. 06 flic.kr/p/2nqEmdW and a forward pointing ‘V’ in dorsal view fig. 04 flic.kr/p/2nqLPGt .
The long, slender, translucent white foot is widest at the bilaminate anterior from whence it tapers gradually to a fine posterior point fig. 12 flic.kr/p/2nqKGcQ . It has propodial swellings which Thompson and Brown (1984) illustrate with a drawing and description of them as ‘propodial tentacles’ projecting ‘about half the width of the foot on either side’. This degree of extension has not been observed by I.F.S. and the greatest extension visible in images in this account is about 17% of body width in dorsal view fig. 04 flic.kr/p/2nqLPGt . The drawing in Alder and Hancock (1844-1854) shows propodial tentacles 30% of body width, the image in Sars (1829) shows barely any swelling and some photos show swellings that can hardly be described as tentacles fig. 11 flic.kr/p/2nqN8gL .
Key identification features
Of Eolis rufibranchialis Alder and Hancock (1844-1854) wrote “This beautiful species has been much misunderstood. - - we suspect that he [G. Johnston, 1832] has included more than one species under the - - name - - and the same course has been followed by other authors.” Over a century later, uncertainty remains, especially if other similar species or forms from Norway and America are included.
Coryphella verrucosa rufibranchialis
1) Cerata as long as the width of the body, and held at varying angles, often across each other fig. 04 flic.kr/p/2nqLPGt .
2) Cerata have a narrow, subterminal, opaque white ring fig. 01 flic.kr/p/2nqKAan & fig. 06 flic.kr/p/2nqEmdW .
3) Substantial white dorsal line on tail fig. 04 flic.kr/p/2nqLPGt , sometimes a broken, white dorsal line on dorsum fig. 02 flic.kr/p/2nqLAnc .
4) Propodial expansions sometimes form small propodial tentacles.
5) Recorded from Scandinavia, Britain, Bretagne and also Atlantic and Pacific coasts of North America, Bering Sea and Sea of Japan (Thompson and Brown 1984).
Coryphella verrucosa verrucosa
1) Cerata barely as long as half the width of the body, and held erect and packed densely together fig. 03 flic.kr/p/2nqLPHR & fig. 07 flic.kr/p/2nqN8kd .
2) Cerata have substantial cap of opaque white, except for a small, translucent apex fig. 09 flic.kr/p/2nqLPCL .
3) Continuous white dorsal line on tail. Dorsal body, apart from hump over pericardium, concealed by dense tubercles fig. 03 flic.kr/p/2nqLPHR & fig. 08 flic.kr/p/2nqEmco .
4) Small propodial expansions.
5) Confined to Scandinavia and Shetland.
Intermediate between C. v. rufibranchialis and C. v. verrucosa
Occasionally forms intermediate between C. v. verrucosa and C. v. rufibranchialis have been reported in Scandinavia where the two forms are sympatric. Some may have been individuals re-growing cerata after damage.
Similar species
Coryphella gracilis (Alder and Hancock, 1844)
fig. 13 flic.kr/p/2nqKzV9 & fig. 14 flic.kr/p/2nqLPwi .
1) Cerata length about same as body width. Narrowed at base so sometimes elliptical, and less slender and flexible than cerata on C. v. rufibranchialis. Arise from a pronounced notal ridge containing the digestive gland.
2) Digestive gland in cerata rather granular, colour varies red to, often, brownish orange or green. Narrow subterminal opaque white ring on cerata.
3) Opaque white dorsal line on tail is sometimes fragmentary or missing. No line on rest of body. Matures at small size, so white ovotestes are visible when 8 mm long, at which size ovotestes are usually missing in juvenile C. verrucosa.
4) Distinct tapering propodial tentacles, length nearly as great as body width.
5) Scattered records around Britain and Ireland. Few or none in southern half of east coast England and north-east Irish Sea.
Coryphella browni Picton, 1980. fig. 15 flic.kr/p/2nqKG8M & fig. 16 flic.kr/p/2nqEm1B .
1) Cerata length up to twice the body width.
2) Broad white band on cerata distally, but no pigment on apex.
3) Interrupted white medial line, often reduced to a few dots or none, on tail. No line on rest of body.
4) Distinct tapering propodial tentacles, length about 25% of body width.
5) Scattered records around Britain and Ireland. Few or none in southern half of east coast England and north-east Irish Sea.
Carronella pellucida (Alder & Hancock, 1843). fig. 17 flic.kr/p/2nqKG62 & fig. 18 flic.kr/p/2nqKG5a .
1) Cerata length greater than width of body.
2) White pigment covers entire distal end of cerata.
3) Opaque white dorsal line on tail. No line on rest of body.
4) Long tapering propodial tentacles as long as, or longer than, width of body.
5) Scandinavia and Scotland; the few English records on NBN are probably misidentifications.
Other similar species
A similar, uncertain species, which occurs in Scotland, Scandinavia and possibly Iceland, feeds on the scyphistoma polyp phase of Aurelia aurita (Linnaeus, 1758)
fig. 19 flic.kr/p/2nqKG4y .
In Scandinavia the following similar species, some of which are poorly known, also need consideration: Berghia norvegica Odhner, 1939; Coryphella borealis Odhner, 1922 Coryphella nobilis A. E. Verrill, 1880 and Ziminella salmonacea (Couthouy, 1838). Some are illustrated at www.seawater.no/fauna/mollusca/norvegica.html . Some of these and other species need consideration in North America.
Habits and ecology
C. verrucosa lives on rocky substrate at LWS and sublittorally (C. v. verrucosa to 450 metres in Scandinavia) where strong currents favour its principle prey; Tubularia indivisa. Many other hydroid species are eaten, especially by juveniles. Like other nudibranchs, it is a simultaneous hermaphrodite. The spawn of both forms is a thin line arranged as a smooth spiral on flat rock fig. 09 flic.kr/p/2nqLPCL , or looped around hydroids. C. v. rufibranchialis breeds from April to June in Britain (Thompson & Brown, 1984). Veliger larvae drift for about ten days before metamorphosing.
Distribution and status
C. v. rufibranchialis, or species or forms resembling it, is recorded, often as C. verrucosa, from Scandinavia, Britain, Bretagne and also Atlantic and Pacific coasts of North America fig. 20 flic.kr/p/2nqN86L , Bering Sea and Sea of Japan (Thompson and Brown 1984). It is fairly frequent sublittorally in Britain and Ireland, and sometimes at LWS. C. v. verrucosa is confined to Shetland fig. 21 flic.kr/p/2nqN84G and Scandinavia, where it is frequent and sympatric with C. v. rufibranchialis.
Appendix, taxonomic history
Coryphella verrucosa (as Eolidia verrucosa) was first described and illustrated fig. 07 flic.kr/p/2nqN8kd by Sars (1829) from specimens in Norway, where it is still frequently found fig. 03 flic.kr/p/2nqLPHR .
Eolis rufibranchialis was first described by Johnston (1832) from a specimen in Berwick, England. He described the cerata as long, thus differing from the short stout cerata of Eolidia verrucosa Sars, 1829.
Alder and Hancock (1844-1854) described and illustrated Eolis rufibranchialis Johnston, 1832 more fully fig. 10 flic.kr/p/2nqKzXt .
Winckworth (1932) brought the two species together as Coryphella verrucosa (M. Sars, 1829) in Norway and subspecies Coryphella verrucosa rufibranchialis (Johnston, 1832).
Odhner (1939) did as Winckworth, using the names Coryphella verrucosa verrucosa and Coryphella verrucosa rufibranchialis for the forms, both of which occur in Norway. Odhner wrote “ Since Løyning (1922) has found, in living material, transitions between C. verrucosa M. Sars and C. rufibranchialis Johnston, and since I have observed the same in Trondheimsfjord, it is evident that the two forms cannot be upheld as distinct species. - - The older name is that of Sars and has to be accepted instead of the generally used one, quite as Winckworth (1932) has done: the two extreme forms are thus given the names Coryphella verrucosa verrucosa and Coryphella verrucosa rufibranchialis, as to the short or elongate form of the papillae [cerata].”
Thompson and Brown (1976) used the form names of Odhner, but they gave an image of Coryphella verrucosa rufibranchialis the label Coryphella verrucosa var. verrucosa, unaware that the latter is absent from Britain, except Shetland
Picton (1980) described a new species, Coryphella browni, which previously would probably have been identified as Coryphella verrucosa rufibranchialis.
Thompson and Brown (1984) omitted use of the variety category and combined both forms, along with other forms from Atlantic and Pacific coasts of North America, Arctic Russia and the Sea of Japan, under the species name Coryphella verrucosa (M. Sars, 1829). Their description is of, only, Coryphella verrucosa rufibranchialis, and they stated their disagreement with Odhner’s accurate description of the cerata of Coryphella verrucosa verrucosa, presumably, because they had not seen live specimens of it with its very different cerata.
None of the images of C. verrucosa from north-east America fig. 20 flic.kr/p/2nqN86L on iNaturalist (accessed May 2022) are of the C. v. verrucosa form. Some match the morphology of European C. v. rufibranchialis.
Eriksson et al. (2006) wrote, “The conspecificity of two forms of Flabellina verrucosa (M. Sars, 1829), one form with short and one with long cerata, was tested by sequencing the mitochondrial COI and the nuclear 5.8S-ITS2 genes. We could not establish any genetic differences between the two forms and conclude that they belong to the same species.” But, examination of other parts of the DNA sequence might show differences which correlate with the morphological differences (B. Picton, 2022. pers. comm., 1 June).
The sighting of intermediate forms and of mating between the two forms fig. 08 flic.kr/p/2nqEmco adds weight to regarding them as a single species, but such sightings are much rarer than would be expected if interbreeding were free and unhindered. Mating of different species of nudibranch with each other is occasionally observed and, as with them, it is unknown if mating of C. v. verrucosa with C. v. rufibranchialis produces viable offspring capable of reproduction.
World Register of Marine Species (accessed 26 May 2022) accepts as valid the names Coryphella verrucosa (M. Sars, 1829) and, as a subspecies, Coryphella verrucosa rufibranchialis (G. Johnston, 1832).
As the two forms are sympatric in Scandinavia and Shetland, yet retain their distinctness, there remains some uncertainty about their exact relationship. It would be a source for future interest if they were recorded separately.
Acknowledgements
For use of images I gratefully thank Billy Arthur, George Brown, Jeff Goddard, Asbjørn Hansen flic.kr/s/87eY , Nathan Jeffery, Tim Nicholson, Poul Rasmussen, Chris Rickard and Erling Svensen. I thank Simon Taylor for specimens. For valuable advice and help with literature I thank Bernard Picton and Miquel Pontes.
References and links
Alder, J. & Hancock, A. 1845-1855. A monograph of the British nudibranchiate mollusca. London, Ray Society. Family 3, plate 14. www.biodiversitylibrary.org/item/131598#page/338/mode/1up
Balch, F. N. 1909. A spring collecting trip. Notes on New England nudibranchs II. The Nautilus. 23(3): 33-38., [Original description of Coryphella rufibranchialis var. chocolata Balch, 1909]
p 35-36 www.biodiversitylibrary.org/page/5314618
page(s): 35-36, 38
Eriksson, R., Nuygren, A. & Sundberg, P. 2006. Genetic evidence of phenotypic polymorphism in the aeolid nudibranch Flabellina verrucosa (M. Sars, 1829) (Opisthobranchia: Nudibranchia). Organisms, Diversity and Evolution 6:1, 71-76
www.sciencedirect.com/science/article/pii/S1439609205000772
iNaturalist, Observations of Coryphella verrucosa. (accessed 25 May 2022) www.inaturalist.org/observations?taxon_id=633193
Johnston, G. 1832. Illustrations in British zoology London Mag. nat. His. 5: 428-429.
www.biodiversitylibrary.org/item/19519#page/464/mode/1up
Korshunova, T., Martynov, A., Bakken, T., Evertsen, J., Fletcher, K., Mudianta, W.I., Saito, H., Lundin, K., Schrödl, M. and Picton B. 2017. Polyphyly of the traditional family Flabellinidae affects a major group of Nudibranchia: aeolidacean taxonomic reassessment with descriptions of several new families, genera, and species (Mollusca, Gastropoda). ZooKeys 717: 1–139. doi.org/10.3897/zookeys.717.21885
Løyning, P. 1922. Nudibranch faunaen i Drøbaksundet I. Fam. Aeolididae, Skrifter utgrit av det Norske Vidensk. Akad. i Oslo
Odhner, N.H. 1922. Norwegian opisthobranchiate mollusca in the collection of the Zoologial Museum of Kristiana. Nyt. Mag. Naturv. 60: 1-47 [Coryphella rufibranchialis p30] www.biodiversitylibrary.org/item/109307#page/38/mode/1up
Odhner, N.H. 1939. Opisthobranchiate Mollusca from the western and northern coasts of Norway. Det Kgl. Norske Videnskabers Selskabs Skrifter 1: 1-92.
Sars, M. 1829. Bidrag til söedyrenes naturhistorie. 1: 9-12 & Plate 2 figs. 1-4 Bergen.
gdz.sub.uni-goettingen.de/id/PPN612886522?tify=
Thompson, T.E. & Brown, G.H. 1976. British opisthobranch molluscs. London, Academic Press.
Thompson, T.E. & Brown, G.H. 1984. Biology of opisthobranch molluscs 2. London, Ray Society.
Winckworth, R. 1932. The British marine mollusca. J. Conch. Lond. 19: 211-252.
Picton, B. E. (1980). A new species of Coryphella (Gastropoda: Opisthobranchia) from the British Isles. Irish Naturalists' Journal. 20: 15-19.
Thompson, T.E. & Brown, G.H. 1984. Biology of opisthobranch molluscs 2. London, Ray Society.
Glossary
cerata = (sing. ceras) lobes on dorsum of aeolids and some other seaslugs.
circum-oesophageal nerve ring = ganglia-bearing nerve cord which encircles the oesophagus.
cnidocytes = explosive stinging cells of Cnidaria. en.wikipedia.org/wiki/Cnidocyte
cnidosac = storage capsule at tips of cerata of Aeolidiidae, but not Dotidae, for ingested unexploded cnidocytes.
digestive gland = large organ in gastropods which acts like the liver and pancreas in mammals to absorb food.
distal = away from centre of body or from point of attachment.
ganglia = (sing. ganglion) knots on a nerve cord containing sensory cell bodies that conduct impulses to innervate organs of the body.
lamellae = small plates on rhinophores.
LWS = low water spring tide, and level it falls to; two periods of a few days each month when tide falls lowest.
notal = (adj.) of the back.
notal ridge = ridge on dorsal surface.
notum = (of seaslugs) the dorsal surface.
ovotestis = (pl. ovotestes) hermaphrodite organ serving as both ovary and testis.
pericardium = sac containing heart, sometimes visible as a raised mound behind rhinophores in aeolid sea slugs.
propodial tentacles = tentacular, lateral extensions on anterior of the foot.
propodium = anterior portion of gastropod foot. (adj. propodial).
rhinophores = chemo-receptor tentacles on top of head of nudibranch.
veliger = shelled larva of marine gastropod or bivalve mollusc which moves by action of cilia on a velum (bilobed flap). Stage may be passed in plankton or within liquid-filled egg-capsule.
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Beija-flor Tesoura (Eupetomena macroura) - Swallow-tailed-Hummingbird
A text In English:
The Swallow-tailed Hummingbird, so called from its forked tail, is one of the largest hummingbirds in cities and gardens, but it also occurs in gallery forests, bushy pastures and edges of woods or coppices. It is green, except for the blue head and upper breast, turning to iridescent purple according to the direction of light; it has dark wings and a heavy black bill. The tail is dark blue with the external feathers longer than central ones. It is very aggressive and attacks other hummingbirds that dare to visit flowers in certain trees. Where the flowers are available for many months, the individual is fiercely territorial, but generally needs to search soon for other flowering plants. It flies to catch small insets on or under leaves in the gallery forests or woodlands. The female builds a small cup-shaped nest saddled on a branch, not far from the main trunk in the shade of leaves. Perched on favorite branches, the male can utter long but low chirps. Once in a while, it interrupts these singing sessions to feed, and flies back for more song or to clean the plumage. They occur from the Guianas and Amazon River to Paraguay and southeastern Peru. They can get along with partially deforested zones, but may disappear with intensive agriculture and with the development of treeless cities.
Um texto em Português:
Beija-flor Tesoura (Eupetomena macroura), fotografado em Brasília-DF, Brasil.
Eupetomena macroura (Gmelin, 1788): tesoura; swallow-tailed hummingbird c.
Destaca-se das espécies estudadas pelo maior porte e pela cauda comprida e bifurcada, o que lhe valeu o nome popular. Como é comum entre os beija-flores, é uma espécie agressiva que disputa com outras o seu território e fontes de alimento.
Nidificação: o ninho, em forma de tigela, é assentado numa forquilha de arbusto ou árvores, a cerca de 2 a 3 m do solo. O material utilizado na construção é composto por fibras vegetais incluindo painas, musgos e liquens, aderidos externamente com teias de aranhas.
Hábitat: capoeiras, cerrados, borda de matas e jardins.
Tamanho: 17,0 cm
A SEGUIR UM TEXTO ENCONTRADO E REPRODUZIDO DO ENDEREÇO nationalgeographic.abril.uol.com.br/ng/edicoes/83/reporta... DA NATIONAL GEOGRAFIC:
Prodígios da micro-engenharia, os beija-flores são os campeões dos pesos-leves entre as aves
Uma faísca safira, um frêmito de asas, e o minúsculo pássaro - ou seria um inseto? - some como miragem fugaz. Reaparece instantes depois, agora num ângulo melhor. É pássaro mesmo, um dervixe do tamanho do meu polegar com asas que batem 80 vertiginosas vezes por segundo, produzindo um zumbido quase inaudível. As penas da cauda, à guisa de leme, delicadamente direcionam o vôo em três direções. Ele fita a trombeta de uma vistosa flor alaranjada e do bico fino como agulha projeta uma língua delgada feito linha. Um raio de Sol ricocheteia de suas penas iridescentes. A cor refletida deslumbra como uma pedra preciosa contra uma janela ensolarada. Não admira que os beija-flores sejam tão queridos e que tanta gente já tenha tropeçado ao tentar descrevê-los. Nem mesmo circunspectos cientistas resistem a termos como "belo", "magnífico", "exótico".
Surpresa maior é o fato de o aparentemente frágil beija-flor ser uma das mais resistentes criaturas do reino animal. Cerca de 330 espécies prosperam em ambientes diversos, muitos deles brutais: do Alasca à Argentina, do deserto do Arizona à costa de Nova Scotia, da Amazônia à linha nevada acima dos 4,5 mil metros nos Andes (misteriosamente, essas aves só são encontradas no Novo Mundo).
"Eles vivem no limite do que é possível aos vertebrados, e com maestria", diz Karl Schuchmann, ornitólogo do Instituto Zoológico Alexander Koenig e do Fundo Brehm, na Alemanha. Schuchmann ouviu falar de um beija-flor que viveu 17 anos em cativeiro. "Imagine a resistência de um organismo de 5 ou 6 gramas para viver tanto tempo!", diz ele espantado. Em média, o minúsculo coração de um beija-flor bate cerca de 500 vezes por minuto (em repouso!). Assim, o desse pequeno cativo teria batido meio bilhão de vezes, quase o dobro do total de uma pessoa de 70 anos.
Mas esses passarinhos são duráveis apenas em vida. Quando morrem, seus ossos delicados e ocos quase nunca se fossilizam. Daí o assombro causado pela recente descoberta de um amontoado de fósseis de aves que talvez inclua um beija-flor ancestral de 30 milhões de anos. Como os beija-flores modernos, os espécimes fósseis tinham o bico longo e fino e os ossos superiores das asas mais curtos, terminando em uma saliência arredondada que talvez lhes permitisse fazer a rotação na articulação do ombro e parar no ar.
A outra surpresa foi o local do achado: no sul da Alemanha, longe do território dos beija-flores atuais. Para alguns cientistas, essa descoberta mostra que já existiram beija-flores fora das Américas, mas se extinguiram. Ou quem sabe os fósseis não fossem de beija-flor. Os céticos, entre eles Schuchmann, afirmam que muitas vezes, ao longo da evolução, outros grupos de aves adquiriram características semelhantes às do beija-flor. Os verdadeiros beija-flores, diz Schuchmann, evoluíram nas florestas do leste do Brasil, onde competiam com insetos pelo néctar das flores.
"O Brasil foi o laboratório do protótipo", diz o ornitólogo. "E o modelo funcionou." O beija-flor tornou-se a obra-prima da microengenharia da natureza. Aperfeiçoou sua habilidade de parar no ar há dezenas de milhões de anos para competir por parte das flores do Novo Mundo.
"Eles são uma ponte entre o mundo das aves e o dos insetos", diz Doug Altshuler, da Universidade da Califórnia em Riverside. Altshuler, que estuda o vôo dos beija-flores, examinou os movimentos das asas do pássaro. Observou que, nele, os impulsos elétricos propulsores dos músculos das asas lembram mais os dos insetos que os das aves. Talvez por isso o beija-flor produza tanta energia por batida de asas: mais, por unidade de massa, que qualquer outro vertebrado. Altshuler também analisou os trajetos neurais do beija-flor, que funcionam com a mesma vertiginosa velocidade encontrada nas aves mais ágeis, como seu primo mais próximo, o andorinhão. "São incríveis; uns pequenos Frankesteins", compara.
Certamente eles sabem intimidar: grama por grama, talvez sejam os maiores confrontadores da natureza. "O vocabulário do beija-flor deve ser 100% composto de palavrões", graceja Sheri Williamson, naturalista do Southeastern Arizona Bird Observatory. A agressão do beija-flor nasce de ferozes instintos territoriais moldados à necessidade de sugar néctar a cada poucos minutos. Os beija-flores competem desafiando e ameaçando uns aos outros. Postam-se face a face no ar, rodopiam, mergulham na direção da grama e voam de ré, em danças de dominância que terminam tão subitamente quanto começam.
O melhor lugar para vermos tais batalhas é nas montanhas, especialmente no Equador, em que ricos ecossistemas se apresentam em suas várias altitudes. Sheri supõe que o sentido norte-sul das cordilheiras americanas também crie rotas favoráveis à migração para onde haja constante suprimento de flores. O que contrasta, diz ela, com as barreiras naturais que se estendem de leste a oeste na África, como o Saara e o Mediterrâneo.
Algumas espécies de beija-flor, porém, adaptaram-se a atravessar vastidões planas, onde o alimento é escasso. Antes de sua intrépida migração da primavera para os Estados Unidos e o Canadá, os beija-flores-de-garganta-vermelha reúnem-se no México e empanturram-se de insetos e néctar. Armazenam gordura e duplicam de peso em uma semana. Em seguida, atravessam o golfo do México, voando 800 quilômetros sem escalas por 20 horas, até a costa distante.
A região próxima à linha do equador é um reino de beija-flores. Quem sai do aeroporto de Quito, no Equador, pode ser logo saudado por um cintilante beija-flor-violeta, com pintura de guerra de manchas púrpura iridescentes nos lados da face. A leste da cidade, nas cabeceiras da bacia Amazônica, o beija-flor-bico-de-espada esvoaça na mata portando o bico mais longo de todas as aves em proporção a seu tamanho: mais de metade do comprimento total do animal. Nas encostas do Cotopaxi, um vulcão ao sul de Quito, o beija-flor-do-chimborazo foi avistado acima dos 4,5 mil metros. Ali ele passa a noite entorpecido em cavernas, pois desacelera seu ritmo metabólico o suficiente para não morrer de fome antes de amanhecer. Mais tarde, aquecido pelo Sol, ele recomeça a se alimentar.
"Quem estuda beija-flores fica irremediavelmente enfeitiçado", diz Sheri Williamson. "São criaturinhas sedutoras. Tentei resistir, mas agora tenho sangue de beija-flor correndo nas veias."
Canon EOS Digital D50
Text, in english, from Wikipedia, the free encyclopedia
"Trumpet tree" redirects here. This term is occasionally used for the Shield-leaved Pumpwood (Cecropia peltata).
Tabebuia
Flowering Araguaney or ipê-amarelo (Tabebuia chrysantha) in central Brazil
Scientific classification
Kingdom: Plantae
(unranked): Angiosperms
(unranked): Eudicots
(unranked): Asterids
Order: Lamiales
Family: Bignoniaceae
Tribe: Tecomeae
Genus: Tabebuia
Gomez
Species
Nearly 100.
Tabebuia is a neotropical genus of about 100 species in the tribe Tecomeae of the family Bignoniaceae. The species range from northern Mexico and the Antilles south to northern Argentina and central Venezuela, including the Caribbean islands of Hispaniola (Dominican Republic and Haiti) and Cuba. Well-known common names include Ipê, Poui, trumpet trees and pau d'arco.
They are large shrubs and trees growing to 5 to 50 m (16 to 160 ft.) tall depending on the species; many species are dry-season deciduous but some are evergreen. The leaves are opposite pairs, complex or palmately compound with 3–7 leaflets.
Tabebuia is a notable flowering tree. The flowers are 3 to 11 cm (1 to 4 in.) wide and are produced in dense clusters. They present a cupular calyx campanulate to tubular, truncate, bilabiate or 5-lobed. Corolla colors vary between species ranging from white, light pink, yellow, lavender, magenta, or red. The outside texture of the flower tube is either glabrous or pubescentThe fruit is a dehiscent pod, 10 to 50 cm (4 to 20 in.) long, containing numerous—in some species winged—seeds. These pods often remain on the tree through dry season until the beginning of the rainy.
Species in this genus are important as timber trees. The wood is used for furniture, decking, and other outdoor uses. It is increasingly popular as a decking material due to its insect resistance and durability. By 2007, FSC-certified ipê wood had become readily available on the market, although certificates are occasionally forged.
Tabebuia is widely used as ornamental tree in the tropics in landscaping gardens, public squares, and boulevards due to its impressive and colorful flowering. Many flowers appear on still leafless stems at the end of the dry season, making the floral display more conspicuous. They are useful as honey plants for bees, and are popular with certain hummingbirds. Naturalist Madhaviah Krishnan on the other hand once famously took offense at ipé grown in India, where it is not native.
Lapacho teaThe bark of several species has medical properties. The bark is dried, shredded, and then boiled making a bitter or sour-tasting brownish-colored tea. Tea from the inner bark of Pink Ipê (T. impetiginosa) is known as Lapacho or Taheebo. Its main active principles are lapachol, quercetin, and other flavonoids. It is also available in pill form. The herbal remedy is typically used during flu and cold season and for easing smoker's cough. It apparently works as expectorant, by promoting the lungs to cough up and free deeply embedded mucus and contaminants. However, lapachol is rather toxic and therefore a more topical use e.g. as antibiotic or pesticide may be advisable. Other species with significant folk medical use are T. alba and Yellow Lapacho (T. serratifolia)
Tabebuia heteropoda, T. incana, and other species are occasionally used as an additive to the entheogenic drink Ayahuasca.
Mycosphaerella tabebuiae, a plant pathogenic sac fungus, was first discovered on an ipê tree.
Tabebuia alba
Tabebuia anafensis
Tabebuia arimaoensis
Tabebuia aurea – Caribbean Trumpet Tree
Tabebuia bilbergii
Tabebuia bibracteolata
Tabebuia cassinoides
Tabebuia chrysantha – Araguaney, Yellow Ipê, tajibo (Bolivia), ipê-amarelo (Brazil), cañaguate (N Colombia)
Tabebuia chrysotricha – Golden Trumpet Tree
Tabebuia donnell-smithii Rose – Gold Tree, "Prima Vera", Cortez blanco (El Salvador), San Juan (Honduras), palo blanco (Guatemala),duranga (Mexico)
A native of Mexico and Central Americas, considered one of the most colorful of all Central American trees. The leaves are deciduous. Masses of golden-yellow flowers cover the crown after the leaves are shed.
Tabebuia dubia
Tabebuia ecuadorensis
Tabebuia elongata
Tabebuia furfuracea
Tabebuia geminiflora Rizz. & Mattos
Tabebuia guayacan (Seem.) Hemsl.
Tabebuia haemantha
Tabebuia heptaphylla (Vell.) Toledo – tajy
Tabebuia heterophylla – roble prieto
Tabebuia heteropoda
Tabebuia hypoleuca
Tabebuia impetiginosa – Pink Ipê, Pink Lapacho, ipê-cavatã, ipê-comum, ipê-reto, ipê-rosa, ipê-roxo-damata, pau d'arco-roxo, peúva, piúva (Brazil), lapacho negro (Spanish); not "brazilwood"
Tabebuia incana
Tabebuia jackiana
Tabebuia lapacho – lapacho amarillo
Tabebuia orinocensis A.H. Gentry[verification needed]
Tabebuia ochracea
Tabebuia oligolepis
Tabebuia pallida – Cuban Pink Trumpet Tree
Tabebuia platyantha
Tabebuia polymorpha
Tabebuia rosea (Bertol.) DC.[verification needed] (= T. pentaphylla (L.) Hemsley) – Pink Poui, Pink Tecoma, apama, apamate, matilisguate
A popular street tree in tropical cities because of its multi-annular masses of light pink to purple flowers and modest size. The roots are not especially destructive for roads and sidewalks. It is the national tree of El Salvador and the state tree of Cojedes, Venezuela
Tabebuia roseo-alba – White Ipê, ipê-branco (Brazil), lapacho blanco
Tabebuia serratifolia – Yellow Lapacho, Yellow Poui, ipê-roxo (Brazil)
Tabebuia shaferi
Tabebuia striata
Tabebuia subtilis Sprague & Sandwith
Tabebuia umbellata
Tabebuia vellosoi Toledo
Ipê-do-cerrado
Texto, em português, da Wikipédia, a enciclopédia livre.
Ipê-do-cerrado
Classificação científica
Reino: Plantae
Divisão: Magnoliophyta
Classe: Magnoliopsida
Subclasse: Asteridae
Ordem: Lamiales
Família: Bignoniaceae
Género: Tabebuia
Espécie: T. ochracea
Nome binomial
Tabebuia ochracea
(Cham.) Standl. 1832
Sinónimos
Bignonia tomentosa Pav. ex DC.
Handroanthus ochraceus (Cham.) Mattos
Tabebuia chrysantha (Jacq.) G. Nicholson
Tabebuia hypodictyon A. DC.) Standl.
Tabebuia neochrysantha A.H. Gentry
Tabebuia ochracea subsp. heteropoda (A. DC.) A.H. Gentry
Tabebuia ochracea subsp. neochrysantha (A.H. Gentry) A.H. Gentry
Tecoma campinae Kraenzl.
ecoma grandiceps Kraenzl.
Tecoma hassleri Sprague
Tecoma hemmendorffiana Kraenzl.
Tecoma heteropoda A. DC.
Tecoma hypodictyon A. DC.
Tecoma ochracea Cham.
Ipê-do-cerrado é um dos nomes populares da Tabebuia ochracea (Cham.) Standl. 1832, nativa do cerrado brasileiro, no estados de Amazonas, Pará, Maranhão, Piauí, Ceará, Pernambuco, Bahia, Espírito Santo, Goiás, Mato Grosso, Mato Grosso do Sul, Minas Gerais, Rio de Janeiro, São Paulo e Paraná.
Está na lista de espécies ameaçadas do estado de São Paulo, onde é encontrda também no domínio da Mata Atlântica[1].
Ocorre também na Argentina, Paraguai, Bolívia, Equador, Peru, Venezuela, Guiana, El Salvador, Guatemala e Panamá[2].
Há uma espécie homônima descrita por A.H. Gentry em 1992.
Outros nomes populares: ipê-amarelo, ipê-cascudo, ipê-do-campo, ipê-pardo, pau-d'arco-do-campo, piúva, tarumã.
Características
Altura de 6 a 14 m. Tronco tortuso com até 50 cm de diâmetro. Folhas pilosas em ambas as faces, mais na inferior, que é mais clara.
Planta decídua, heliófita, xerófita, nativa do cerrado em solos bem drenados.
Floresce de julho a setembro. Os frutos amadurecem de setembro a outubro.
FloresProduz grande quantidade de sementes leves, aladas com pequenas reservas, e que perdem a viabilidade em menos de 90 dias após coleta. A sua conservação vem sendo estudada em termos de determinação da condição ideal de armazenamento, e tem demonstrado a importância de se conhecer o comportamento da espécie quando armazenada com diferentes teores de umidade inicial, e a umidade de equilíbrio crítica para a espécie (KANO; MÁRQUEZ & KAGEYAMA, 1978). As levíssimas sementes aladas da espécie não necessitam de quebra de dormência. Podem apenas ser expostas ao sol por cerca de 6 horas e semeadas diretamente nos saquinhos. A germinação ocorre após 30 dias e de 80%. As sementes são ortodoxas e há aproximadamente 72 000 sementes em cada quilo.
O desenvolvimento da planta é rápido.
Como outros ipês, a madeira é usada em tacos, assoalhos, e em dormentes e postes. Presta-se também para peças torneadas e instrumento musicais.
Tabebuia alba (Ipê-Amarelo)
Texto, em português, produzido pela Acadêmica Giovana Beatriz Theodoro Marto
Supervisão e orientação do Prof. Luiz Ernesto George Barrichelo e do Eng. Paulo Henrique Müller
Atualizado em 10/07/2006
O ipê amarelo é a árvore brasileira mais conhecida, a mais cultivada e, sem dúvida nenhuma, a mais bela. É na verdade um complexo de nove ou dez espécies com características mais ou menos semelhantes, com flores brancas, amarelas ou roxas. Não há região do país onde não exista pelo menos uma espécie dele, porém a existência do ipê em habitat natural nos dias atuais é rara entre a maioria das espécies (LORENZI,2000).
A espécie Tabebuia alba, nativa do Brasil, é uma das espécies do gênero Tabebuia que possui “Ipê Amarelo” como nome popular. O nome alba provém de albus (branco em latim) e é devido ao tomento branco dos ramos e folhas novas.
As árvores desta espécie proporcionam um belo espetáculo com sua bela floração na arborização de ruas em algumas cidades brasileiras. São lindas árvores que embelezam e promovem um colorido no final do inverno. Existe uma crença popular de que quando o ipê-amarelo floresce não vão ocorrer mais geadas. Infelizmente, a espécie é considerada vulnerável quanto à ameaça de extinção.
A Tabebuia alba, natural do semi-árido alagoano está adaptada a todas as regiões fisiográficas, levando o governo, por meio do Decreto nº 6239, a transformar a espécie como a árvore símbolo do estado, estando, pois sob a sua tutela, não mais podendo ser suprimida de seus habitats naturais.
Taxonomia
Família: Bignoniaceae
Espécie: Tabebuia Alba (Chamiso) Sandwith
Sinonímia botânica: Handroanthus albus (Chamiso) Mattos; Tecoma alba Chamisso
Outros nomes vulgares: ipê-amarelo, ipê, aipê, ipê-branco, ipê-mamono, ipê-mandioca, ipê-ouro, ipê-pardo, ipê-vacariano, ipê-tabaco, ipê-do-cerrado, ipê-dourado, ipê-da-serra, ipezeiro, pau-d’arco-amarelo, taipoca.
Aspectos Ecológicos
O ipê-amarelo é uma espécie heliófita (Planta adaptada ao crescimento em ambiente aberto ou exposto à luz direta) e decídua (que perde as folhas em determinada época do ano). Pertence ao grupo das espécies secundárias iniciais (DURIGAN & NOGUEIRA, 1990).
Abrange a Floresta Pluvial da Mata Atlântica e da Floresta Latifoliada Semidecídua, ocorrendo principalmente no interior da Floresta Primária Densa. É característica de sub-bosques dos pinhais, onde há regeneração regular.
Informações Botânicas
Morfologia
As árvores de Tabebuia alba possuem cerca de 30 metros de altura. O tronco é reto ou levemente tortuoso, com fuste de 5 a 8 m de altura. A casca externa é grisáceo-grossa, possuindo fissuras longitudinais esparas e profundas. A coloração desta é cinza-rosa intenso, com camadas fibrosas, muito resistentes e finas, porém bem distintas.
Com ramos grossos, tortuosos e compridos, o ipê-amarelo possui copa alongada e alargada na base. As raízes de sustentação e absorção são vigorosas e profundas.
As folhas, deciduais, são opostas, digitadas e compostas. A face superior destas folhas é verde-escura, e, a face inferior, acinzentada, sendo ambas as faces tomentosas. Os pecíolos das folhas medem de 2,5 a 10 cm de comprimento. Os folíolos, geralmente, apresentam-se em número de 5 a 7, possuindo de 7 a 18 cm de comprimento por 2 a 6 cm de largura. Quando jovem estes folíolos são densamente pilosos em ambas as faces. O ápice destes é pontiagudo, com base arredondada e margem serreada.
As flores, grandes e lanceoladas, são de coloração amarelo-ouro. Possuem em média 8X15 cm.
Quanto aos frutos, estes possuem forma de cápsula bivalvar e são secos e deiscentes. Do tipo síliqua, lembram uma vagem. Medem de 15 a 30 cm de comprimento por 1,5 a 2,5 cm de largura. As valvas são finamente tomentosas com pêlos ramificados. Possuem grande quantidade de sementes.
As sementes são membranáceas brilhantes e esbranquiçadas, de coloração marrom. Possuem de 2 a 3 cm de comprimento por 7 a 9 mm de largura e são aladas.
Reprodução
A espécie é caducifólia e a queda das folhas coincide com o período de floração. A floração inicia-se no final de agosto, podendo ocorrer alguma variação devido a fenômenos climáticos. Como a espécie floresce no final do inverno é influenciada pela intensidade do mesmo. Quanto mais frio e seco for o inverno, maior será a intensidade da florada do ipê amarelo.
As flores por sua exuberância, atraem abelhas e pássaros, principalmente beija-flores que são importantes agentes polinizadores. Segundo CARVALHO (2003), a espécie possui como vetor de polinização a abelha mamangava (Bombus morio).
As sementes são dispersas pelo vento.
A planta é hermafrodita, e frutifica nos meses de setembro, outubro, novembro, dezembro, janeiro e fevereiro, dependendo da sua localização. Em cultivo, a espécie inicia o processo reprodutivo após o terceiro ano.
Ocorrência Natural
Ocorre naturalmente na Floresta Estaciobal Semidecicual, Floresta de Araucária e no Cerrado.
Segundo o IBGE, a Tabebuia alba (Cham.) Sandw. é uma árvore do Cerrado, Cerradão e Mata Seca. Apresentando-se nos campos secos (savana gramíneo-lenhosa), próximo às escarpas.
Clima
Segundo a classificação de Köppen, o ipê-amarelo abrange locais de clima tropical (Aw), subtropical úmido (Cfa), sutropical de altitude (Cwa e Cwb) e temperado.
A T.alba pode tolerar até 81 geadas em um ano. Ocorre em locais onde a temperatura média anual varia de 14,4ºC como mínimo e 22,4ºC como máximo.
Solo
A espécie prefere solos úmidos, com drenagem lenta e geralmente não muito ondulados (LONGHI, 1995).
Aparece em terras de boa à média fertilidade, em solos profundos ou rasos, nas matas e raramente cerradões (NOGUEIRA, 1977).
Pragas e Doenças
De acordo com CARVALHO (2003), possui como praga a espécie de coleópteros Cydianerus bohemani da família Curculionoideae e um outro coleóptero da família Chrysomellidae. Apesar da constatação de elevados índices populacionais do primeiro, os danos ocasionados até o momento são leves. Nas praças e ruas de Curitiba - PR, 31% das árvores foram atacadas pela Cochonilha Ceroplastes grandis.
ZIDKO (2002), ao estudar no município de Piracicaba a associação de coleópteros em espécies arbóreas, verificou a presença de insetos adultos da espécie Sitophilus linearis da família de coleópteros, Curculionidae, em estruturas reprodutivas. Os insetos adultos da espécie emergiram das vagens do ipê, danificando as sementes desta espécie nativa.
ANDRADE (1928) assinalou diversas espécies de Cerambycidae atacando essências florestais vivas, como ingazeiro, cinamomo, cangerana, cedro, caixeta, jacarandá, araribá, jatobá, entre outras como o ipê amarelo.
A Madeira
A Tabebuia alba produz madeira de grande durabilidade e resistência ao apodrecimento (LONGHI,1995).
MANIERI (1970) caracteriza o cerne desta espécie como de cor pardo-havana-claro, pardo-havan-escuro, ou pardo-acastanhado, com reflexos esverdeados. A superfície da madeira é irregularmente lustrosa, lisa ao tato, possuindo textura media e grã-direita.
Com densidade entre 0,90 e 1,15 grama por centímetro cúbico, a madeira é muito dura (LORENZI, 1992), apresentando grande dificuldade ao serrar.
A madeira possui cheiro e gosto distintos. Segundo LORENZI (1992), o cheiro característico é devido à presença da substância lapachol, ou ipeína.
Usos da Madeira
Sendo pesada, com cerne escuro, adquire grande valor comercial na marcenaria e carpintaria. Também é utilizada para fabricação de dormentes, moirões, pontes, postes, eixos de roda, varais de carroça, moendas de cana, etc.
Produtos Não-Madeireiros
A entrecasca do ipê-amarelo possui propriedades terapêuticas como adstringente, usada no tratamento de garganta e estomatites. É também usada como diurético.
O ipê-amarelo possui flores melíferas e que maduras podem ser utilizadas na alimentação humana.
Outros Usos
É comumente utilizada em paisagismo de parques e jardins pela beleza e porte. Além disso, é muito utilizada na arborização urbana.
Segundo MOREIRA & SOUZA (1987), o ipê-amarelo costuma povoar as beiras dos rios sendo, portanto, indicado para recomposição de matas ciliares. MARTINS (1986), também cita a espécie para recomposição de matas ciliares da Floresta Estacional Semidecidual, abrangendo alguns municípios das regiões Norte, Noroeste e parte do Oeste do Estado do Paraná.
Aspectos Silviculturais
Possui a tendência a crescer reto e sem bifurcações quando plantado em reflorestamento misto, pois é espécie monopodial. A desrrama se faz muito bem e a cicatrização é boa. Sendo assim, dificilmente encopa quando nova, a não ser que seja plantado em parques e jardins.
Ao ser utilizada em arborização urbana, o ipê amarelo requer podas de condução com freqüência mediana.
Espécie heliófila apresenta a pleno sol ramificação cimosa, registrando-se assim dicotomia para gema apical. Deve ser preconizada, para seu melhor aproveitamento madeireiro, podas de formação usuais (INQUE et al., 1983).
Produção de Mudas
A propagação deve realizada através de enxertia.
Os frutos devem ser coletados antes da dispersão, para evitar a perda de sementes. Após a coleta as sementes são postas em ambiente ventilado e a extração é feita manualmente. As sementes do ipê amarelo são ortodoxas, mantendo a viabilidade natural por até 3 meses em sala e por até 9 meses em vidro fechado, em câmara fria.
A condução das mudas deve ser feita a pleno sol. A muda atinge cerca de 30 cm em 9 meses, apresentando tolerância ao sol 3 semanas após a germinação.
Sementes
Os ipês, espécies do gênero Tabebuia, produzem uma grande quantidade de sementes leves, aladas com pequenas reservas, e que perdem a viabilidade em poucos dias após a sua coleta. A sua conservação vem sendo estudada em termos de determinação da condição ideal de armazenamento, e tem demonstrado a importância de se conhecer o comportamento da espécie quando armazenada com diferentes teores de umidade inicial, e a umidade de equilíbrio crítica para a espécie (KANO; MÁRQUEZ & KAGEYAMA, 1978).
As levíssimas sementes aladas da espécie não necessitam de quebra de dormência. Podem apenas ser expostas ao sol por cerca de 6 horas e semeadas diretamente nos saquinhos. A quebra natural leva cerca de 3 meses e a quebra na câmara leva 9 meses. A germinação ocorre após 30 dias e de 80%.
As sementes são ortodoxas e há aproximadamente 87000 sementes em cada quilo.
Preço da Madeira no Mercado
O preço médio do metro cúbico de pranchas de ipê no Estado do Pará cotado em Julho e Agosto de 2005 foi de R$1.200,00 o preço mínimo, R$ 1509,35 o médio e R$ 2.000,00 o preço máximo (CEPEA,2005).
Papilio dardanus (the African Swallowtail, Mocker Swallowtail or Flying Handkerchief), is a species of butterfly in the family Papilionidae (the Swallowtails). The species is broadly distributed throughout sub-Saharan Africa. The British entomologist E. B. Poulton described it as "the most interesting butterfly in the world".
The species shows polymorphism in wing appearance, though this is limited to females, which are often given as an example of (Batesian) mimicry in insects. This female-limited mimicry was first described in 1869 by Roland Trimen. Males have a more or less uniform appearance throughout the species' range, but females come in at least 14 varieties or morphs. Some female morphs share a very similar pattern of colouration with various species of distasteful butterfly (e.g. from the family Danaidae), while others have been found that mimic male appearance (andromorphs). The persistence of these various morphs or different types of females may be explained by frequency dependent selection. Cook et al. suggest that Batesian mimics gain a fitness advantage by avoiding predators, but suffer harrassment from males (see sexual conflict), whereas andromorphs (male mimics) are vulnerable to predation but are not harrassed by male mating attempts.
Such female-limited Batesian mimicry is not unique to this species, even in the genus Papilio. For instance Papilio memnon shows a similar case of polymorphism in females. Similarly, male mimicry has been observed in another insect, a damselfly (Ischnura ramburi) which also appears to have evolved camouflage to avoid sexual coercion by males.
The viviparous lizard or common lizard, Zootoca vivipara (formerly Lacerta vivipara), is a Eurasian lizard. It lives farther north than any other reptile species, and most populations are viviparous (giving birth to live young), rather than laying eggs as most other lizards do. It is the only species in the monotypic genus Zootoca.
Zootoca vivipara can be seen in a variety of different colors. Female Zootoca vivipara undergo color polymorphism (biology) more commonly than males. A female lizard's display differs in ventral coloration, ranging from pale yellow to bright orange and a mixed coloration. There have been many hypothesis for the genetic cause of this polymorphic coloration. These hypothesis test for coloration due to thermoregulation, predator avoidance, and social cues, specifically sexual reproduction. Through an experiment conducted by Vercken et al., color polymorphism in viviparous lizard is caused by social cues, rather than the other hypotheses. More specifically, the ventral coloration that is seen in female lizards is associated with patterns of sexual reproduction and sex allocation.
The length of the body is less than 12 cm (5 in) (excluding the tail). The tail is up to twice as long as the body, although it is often partially or wholly lost. The limbs are short, and the head is rather round. Males have more slender bodies than females. The neck and the tail are thick. The collar and other scales seem jagged.
The colour and patterning of this species is variable. The main colour is typically medium brown, but it can be also grey, olive brown or black. Females may have dark stripes on their flanks and down the middle of their backs. Sometimes females also have light-coloured stripes, or dark and light spots along the sides of their backs. Most males and some females have dark spots in their undersides. Males have brightly coloured undersides – typically yellow or orange, but more rarely red. Females have paler, whitish underparts. The throat is white, sometimes blue.
(Wikipedia)
[Camponotus Mayr 1861: 1,083+†29 (IT: 19+†0) spp (41.2-0.0 mya)]
Camponotus is an extremely large and complex, globally distributed genus. At present, nearly 500 sspp belonging to 45 sgg have been described and it could well be the largest ant genus of all. The enormous species richness, high levels of intraspecific and geographic variation and polymorphism render the taxonomy of Camponotus one of the most complex and difficult. Revisionary studies are generally confined to species groups and/or small geographical regions. These ants live in a variety of habitats and microhabitats and the sheer size of the genus makes any characterisation of their biology challenging. Nests are built in the ground, in rotten branches or twigs, or rarely into living wood and most spp possess a highly generalistic diet.
REFERENCES
Zonotrichia albicollis has two genetically-determined but behaviorally distinct color-morphs: tan-striped and white-striped. These white-striped individuals, who are typically more aggressive and prone to polyamory, typically mate with more restrained, monogamous tan-stripes, producing both types of offspring and thus maintaining a balanced polymorphism in a regional population. Keene, NH (April 29, 2017)
[Camponotus Mayr 1861: 1,083+†29 (IT: 19+†0) spp (41.2-0.0 mya)]
Parabiotic of Crematogaster scutellaris, Lasius emarginatus, Lasius lasioides, Pheidole pallidula, Solenopsis fugax, Tapinoma magnum.
Camponotus is an extremely large and complex, globally distributed genus. At present, nearly 500 sspp belonging to 45 sgg have been described and it could well be the largest ant genus of all. The enormous species richness, high levels of intraspecific and geographic variation and polymorphism render the taxonomy of Camponotus one of the most complex and difficult. Revisionary studies are generally confined to species groups and/or small geographical regions. These ants live in a variety of habitats and microhabitats and the sheer size of the genus makes any characterisation of their biology challenging. Nests are built in the ground, in rotten branches or twigs, or rarely into living wood and most spp possess a highly generalistic diet.
REFERENCES
Narcissus is a genus of predominantly spring flowering perennial plants of the amaryllis family, Amaryllidaceae. Various common names including daffodil, narcissus, and jonquil are used to describe all or some members of the genus. Narcissus has conspicuous flowers with six petal-like tepals surmounted by a cup- or trumpet-shaped corona. The flowers are generally white and yellow (also orange or pink in garden varieties), with either uniform or contrasting coloured tepals and corona.
Narcissus were well known in ancient civilisation, both medicinally and botanically, but formally described by Linnaeus in his Species Plantarum (1753). The genus is generally considered to have about ten sections with approximately 50 species. The number of species has varied, depending on how they are classified, due to similarity between species and hybridisation. The genus arose some time in the Late Oligocene to Early Miocene epochs, in the Iberian peninsula and adjacent areas of southwest Europe. The exact origin of the name Narcissus is unknown, but it is often linked to a Greek word (ancient Greek ναρκῶ narkō, "to make numb") and the myth of the youth of that name who fell in love with his own reflection. The English word "daffodil" appears to be derived from "asphodel", with which it was commonly compared.
The species are native to meadows and woods in southern Europe and North Africa with a centre of diversity in the Western Mediterranean, particularly the Iberian peninsula. Both wild and cultivated plants have naturalised widely, and were introduced into the Far East prior to the tenth century. Narcissi tend to be long-lived bulbs, which propagate by division, but are also insect-pollinated. Known pests, diseases and disorders include viruses, fungi, the larvae of flies, mites and nematodes. Some Narcissus species have become extinct, while others are threatened by increasing urbanisation and tourism.
Historical accounts suggest narcissi have been cultivated from the earliest times, but became increasingly popular in Europe after the 16th century and by the late 19th century were an important commercial crop centred primarily in the Netherlands. Today narcissi are popular as cut flowers and as ornamental plants in private and public gardens. The long history of breeding has resulted in thousands of different cultivars. For horticultural purposes, narcissi are classified into divisions, covering a wide range of shapes and colours. Like other members of their family, narcissi produce a number of different alkaloids, which provide some protection for the plant, but may be poisonous if accidentally ingested. This property has been exploited for medicinal use in traditional healing and has resulted in the production of galantamine for the treatment of Alzheimer's dementia. Long celebrated in art and literature, narcissi are associated with a number of themes in different cultures, ranging from death to good fortune, and as symbols of spring.
The daffodil is the national flower of Wales and the symbol of cancer charities in many countries. The appearance of the wild flowers in spring is associated with festivals in many places.
Narcissus is a genus of perennial herbaceous bulbiferous geophytes, which die back after flowering to an underground storage bulb. They regrow in the following year from brown-skinned ovoid bulbs with pronounced necks, and reach heights of 5–80 centimetres (2.0–31.5 in) depending on the species. Dwarf species such as N. asturiensis have a maximum height of 5–8 centimetres (2.0–3.1 in), while Narcissus tazetta may grow as tall as 80 centimetres (31 in).
The plants are scapose, having a single central leafless hollow flower stem (scape). Several green or blue-green, narrow, strap-shaped leaves arise from the bulb. The plant stem usually bears a solitary flower, but occasionally a cluster of flowers (umbel). The flowers, which are usually conspicuous and white or yellow, sometimes both or rarely green, consist of a perianth of three parts. Closest to the stem (proximal) is a floral tube above the ovary, then an outer ring composed of six tepals (undifferentiated sepals and petals), and a central disc to conical shaped corona. The flowers may hang down (pendant), or be erect. There are six pollen bearing stamens surrounding a central style. The ovary is inferior (below the floral parts) consisting of three chambers (trilocular). The fruit consists of a dry capsule that splits (dehisces) releasing numerous black seeds.
The bulb lies dormant after the leaves and flower stem die back and has contractile roots that pull it down further into the soil. The flower stem and leaves form in the bulb, to emerge the following season. Most species are dormant from summer to late winter, flowering in the spring, though a few species are autumn flowering.
The pale brown-skinned ovoid tunicate bulbs have a membranous tunic and a corky stem (base or basal) plate from which arise the adventitious root hairs in a ring around the edge, which grow up to 40 mm in length. Above the stem plate is the storage organ consisting of bulb scales, surrounding the previous flower stalk and the terminal bud. The scales are of two types, true storage organs and the bases of the foliage leaves. These have a thicker tip and a scar from where the leaf lamina became detached. The innermost leaf scale is semicircular only partly enveloping the flower stalk (semisheathed).(see Hanks Fig 1.3). The bulb may contain a number of branched bulb units, each with two to three true scales and two to three leaf bases. Each bulb unit has a life of about four years.
Once the leaves die back in summer, the roots also wither. After some years, the roots shorten pulling the bulbs deeper into the ground (contractile roots). The bulbs develop from the inside, pushing the older layers outwards which become brown and dry, forming an outer shell, the tunic or skin. Up to 60 layers have been counted in some wild species. While the plant appears dormant above the ground the flower stalk which will start to grow in the following spring, develops within the bulb surrounded by two to three deciduous leaves and their sheaths. The flower stem lies in the axil of the second true leaf.
The single leafless Plant stem stem or scape, appearing from early to late spring depending on the species, bears from 1 to 20 blooms. Stem shape depends on the species, some are highly compressed with a visible seam, while others are rounded. The stems are upright and located at the centre of the leaves. In a few species such as N. hedraeanthus the stem is oblique. The stem is hollow in the upper portion but towards the bulb is more solid and filled with a spongy material.
Narcissus plants have one to several basal leaf leaves which are linear, ligulate or strap-shaped (long and narrow), sometimes channelled adaxially to semiterete, and may (pedicellate) or may not (sessile) have a petiole stalk. The leaves are flat and broad to cylindrical at the base and arise from the bulb. The emerging plant generally has two leaves, but the mature plant usually three, rarely four, and they are covered with a cutin containing cuticle, giving them a waxy appearance. Leaf colour is light green to blue-green. In the mature plant, the leaves extend higher than the flower stem, but in some species, the leaves are low-hanging. The leaf base is encased in a colorless sheath. After flowering, the leaves turn yellow and die back once the seed pod (fruit) is ripe.
Jonquils usually have dark green, round, rush-like leaves.
The inflorescence is scapose, the single stem or scape bearing either a solitary flower or forming an umbel with up to 20 blooms. Species bearing a solitary flower include section Bulbocodium and most of section Pseudonarcissus. Umbellate species have a fleshy racemose inflorescence (unbranched, with short floral stalks) with 2 to 15 or 20 flowers, such as N. papyraceus (see illustration, left) and N. tazetta (see Table I). The flower arrangement on the inflorescence may be either with (pedicellate) or without (sessile) floral stalks.
Prior to opening, the flower buds are enveloped and protected in a thin dry papery or membranous (scarious) spathe. The spathe consists of a singular bract that is ribbed, and which remains wrapped around the base of the open flower. As the bud grows, the spathe splits longitudinally. Bracteoles are small or absent.
The flowers of Narcissus are hermaphroditic (bisexual), have three parts (tripartite), and are sometimes fragrant (see Fragrances). The flower symmetry is actinomorphic (radial) to slightly zygomorphic (bilateral) due to declinate-ascending stamens (curving downwards, then bent up at the tip). Narcissus flowers are characterised by their, usually conspicuous, corona (trumpet).
The three major floral parts (in all species except N. cavanillesii in which the corona is virtually absent - Table I: Section Tapeinanthus) are;
(i) the proximal floral tube (hypanthium),
(ii) the surrounding free tepals, and
(iii) the more distal corona (paraperigon, paraperigonium).
All three parts may be considered to be components of the perianth (perigon, perigonium). The perianth arises above the apex of the inferior ovary, its base forming the hypanthial floral tube.
The floral tube is formed by fusion of the basal segments of the tepals (proximally connate). Its shape is from an inverted cone (obconic) to funnel-shaped (funneliform) or cylindrical, and is surmounted by the more distal corona. Floral tubes can range from long and narrow sections Apodanthi and Jonquilla to rudimentary (N. cavanillesii).
Surrounding the floral tube and corona and reflexed (bent back) from the rest of the perianth are the six spreading tepals or floral leaves, in two whorls which may be distally ascending, reflexed (folded back), or lanceolate. Like many monocotyledons, the perianth is homochlamydeous, which is undifferentiated into separate calyx (sepals) and corolla (petals), but rather has six tepals. The three outer tepal segments may be considered sepals, and the three inner segments petals. The transition point between the floral tube and the corona is marked by the insertion of the free tepals on the fused perianth.
The corona, or paracorolla, is variously described as bell-shaped (funneliform, trumpet), bowl-shaped (cupular, crateriform, cup-shaped) or disc-shaped with margins that are often frilled, and is free from the stamens. Rarely is the corona a simple callose (hardened, thickened) ring. The corona is formed during floral development as a tubular outgrowth from stamens which fuse into a tubular structure, the anthers becoming reduced. At its base, the fragrances which attract pollinators are formed. All species produce nectar at the top of the ovary. Coronal morphology varies from the tiny pigmented disk of N. serotinus (see Table I) or the rudimentary structure in N. cavanillesii to the elongated trumpets of section Pseudonarcissus (trumpet daffodils, Table I).
While the perianth may point forwards, in some species such as N. cyclamineus it is folded back (reflexed, see illustration, left), while in some other species such as N. bulbocodium (Table I), it is reduced to a few barely visible pointed segments with a prominent corona.
The colour of the perianth is white, yellow or bicoloured, with the exception of the night flowering N. viridiflorus which is green. In addition the corona of N. poeticus has a red crenulate margin (see Table I). Flower diameter varies from 12 mm (N. bulbocodium) to over 125 mm (N. nobilis=N. pseudonarcissus subsp. nobilis).
Flower orientation varies from pendent or deflexed (hanging down) as in N. triandrus (see illustration, left), through declinate-ascendant as in N. alpestris = N. pseudonarcissus subsp. moschatus, horizontal (patent, spreading) such as N. gaditanus or N. poeticus, erect as in N. cavanillesii, N. serotinus and N. rupicola (Table I), or intermediate between these positions (erecto-patent).
The flowers of Narcissus demonstrate exceptional floral diversity and sexual polymorphism, primarily by corona size and floral tube length, associated with pollinator groups (see for instance Figs. 1 and 2 in Graham and Barrett). Barrett and Harder (2005) describe three separate floral patterns;
"Daffodil" form
"Paperwhite" form
"Triandrus" form.
The predominant patterns are the 'daffodil' and 'paperwhite' forms, while the "triandrus" form is less common. Each corresponds to a different group of pollinators (See Pollination).
The "daffodil" form, which includes sections Pseudonarcissus and Bulbocodium, has a relatively short, broad or highly funnelform tube (funnel-like), which grades into an elongated corona, which is large and funnelform, forming a broad, cylindrical or trumpet-shaped perianth. Section Pseudonarcissus consists of relatively large flowers with a corolla length of around 50 mm, generally solitary but rarely in inflorescences of 2–4 flowers. They have wide greenish floral tubes with funnel-shaped bright yellow coronas. The six tepals sometimes differ in colour from the corona and may be cream coloured to pale yellow.
The "paperwhite" form, including sections Jonquilla, Apodanthi and Narcissus, has a relatively long, narrow tube and a short, shallow, flaring corona. The flower is horizontal and fragrant.
The "triandrus" form is seen in only two species, N. albimarginatus (a Moroccan endemic) and N. triandrus. It combines features of both the "daffodil" and "paperwhite" forms, with a well-developed, long, narrow tube and an extended bell-shaped corona of almost equal length. The flowers are pendent.
Androecium
There are six stamens in one to two rows (whorls), with the filaments separate from the corona, attached at the throat or base of the tube (epipetalous), often of two separate lengths, straight or declinate-ascending (curving downwards, then upwards). The anthers are basifixed (attached at their base).
Gynoecium
The ovary is inferior (below the floral parts) and trilocular (three chambered) and there is a pistil with a minutely three lobed stigma and filiform (thread like) style, which is often exserted (extending beyond the tube).
Fruit
The fruit consists of dehiscent loculicidal capsules (splitting between the locules) that are ellipsoid to subglobose (almost spherical) in shape and are papery to leathery in texture.
Seeds
The fruit contains numerous subglobose seeds which are round and swollen with a hard coat, sometimes with an attached elaiosome. The testa is black and the pericarp dry.
Most species have 12 ovules and 36 seeds, although some species such as N. bulbocodium have more, up to a maximum of 60. Seeds take five to six weeks to mature. The seeds of sections Jonquilla and Bulbocodium are wedge-shaped and matte black, while those of other sections are ovate and glossy black. A gust of wind or contact with a passing animal is sufficient to disperse the mature seeds.
Chromosomes
Chromosome numbers include 2n=14, 22, 26, with numerous aneuploid and polyploid derivatives. The basic chromosome number is 7, with the exception of N. tazetta, N. elegans and N. broussonetii in which it is 10 or 11; this subgenus (Hermione) was in fact characterised by this characteristic. Polyploid species include N. papyraceus (4x=22) and N. dubius (6x=50).
Phytochemistry
Alkaloids
As with all Amarylidaceae genera, Narcissus contains unique isoquinoline alkaloids. The first alkaloid to be identified was lycorine, from N. pseudonarcissus in 1877. These are considered a protective adaptation and are utilised in the classification of species. Nearly 100 alkaloids have been identified in the genus, about a third of all known Amaryllidaceae alkaloids, although not all species have been tested. Of the nine alkaloid ring types identified in the family, Narcissus species most commonly demonstrate the presence of alkaloids from within the Lycorine (lycorine, galanthine, pluviine) and Homolycorine (homolycorine, lycorenine) groups. Hemanthamine, tazettine, narciclasine, montanine and galantamine alkaloids are also represented. The alkaloid profile of any plant varies with time, location, and developmental stage. Narcissus also contain fructans and low molecular weight glucomannan in the leaves and plant stems.
Fragrances
Fragrances are predominantly monoterpene isoprenoids, with a small amount of benzenoids, although N. jonquilla has both equally represented. Another exception is N. cuatrecasasii which produces mainly fatty acid derivatives. The basic monoterpene precursor is geranyl pyrophosphate, and the commonest monoterpenes are limonene, myrcene, and trans-β-ocimene. Most benzenoids are non-methoxylated, while a few species contain methoxylated forms (ethers), e.g. N. bujei. Other ingredient include indole, isopentenoids and very small amounts of sesquiterpenes. Fragrance patterns can be correlated with pollinators, and fall into three main groups (see Pollination).
The taxonomy of Narcissus is complex, and still not fully resolved. Known to the ancients, the genus name appears in Graeco-Roman literature, although their interest was as much medicinal as botanical. It is unclear which species the ancients were familiar with. Although frequently mentioned in Mediaeval and Renaissance texts it was not formally described till the work of Linnaeus in 1753. By 1789 it had been grouped into a family (Narcissi) but shortly thereafter this was renamed Amaryllideae, from which comes the modern placement within Amaryllidaceae, although for a while it was considered part of Liliaceae.
Many of the species now considered to be Narcissus were in separate genera during the nineteenth century, and the situation was further confused by the inclusion of many cultivated varieties. By 1875 the current circumscription was relatively settled. By 2004 phylogenetic studies had allowed the place of Narcissus within its fairly large family to be established, nested within a series of subfamilies (Amaryllidoideae) and tribes (Narcisseae). It shares its position in the latter tribe with Sternbergia.
The infrageneric classification has been even more complex and many schemes of subgenera, sections, subsections and series have been proposed, although all had certain similarities. Most authorities now consider there to be 10 – 11 sections based on phylogenetic evidence. The problems have largely arisen from the diversity of the wild species, frequent natural hybridisation and extensive cultivation with escape and subsequent naturalisation. The number of species has varied anywhere from 16 to nearly 160, but is probably around 50 – 60.
The genus appeared some time in the Late Oligocene to Early Miocene eras, around 24 million years ago, in the Iberian peninsula. While the exact origin of the word Narcissus is unknown it is frequently linked to its fragrance which was thought to be narcotic, and to the legend of the youth of that name who fell in love with his reflection. In the English language the common name Daffodil appears to be derived from the Asphodel with which it was commonly compared.
Early
Narcissus was first described by Theophrastus (Θεόφραστος, c 371 - c 287 BC) in his Historia Plantarum (Greek: Περὶ φυτῶν ἱστορία) as νάρκισσος, referring to N. poeticus, but comparing it to Asphodelus (ασφοδελωδες). Theophrastus' description was frequently referred to at length by later authors writing in Latin such as Pliny the Elder (Gaius Plinius Secundus, 23 AD – 79 AD) from whom came the Latin form narcissus (see also Culture). Pliny's account is from his Natural History (Latin: Naturalis Historia). Like his contemporaries, his interests were as much therapeutic as botanical. Another much-cited Greek authority was Dioscorides (Διοσκουρίδης, 40 AD – 90 AD) in his De Materia Medica (Greek: Περὶ ὕλης ἰατρικῆς). Both authors were to remain influential until at least the Renaissance, given that their descriptions went beyond the merely botanical, to the therapeutic (see also Antiquity).
An early European reference is found in the work of Albert Magnus (c. 1200 – 1280), who noted in his De vegetabilibus et plantis the similarity to the leek. William Turner in his A New Herball (1551) cites all three extensively in his description of the plant and its properties.It was to remain to Linnaeus in 1753 to formally describe and name Narcissus as a genus in his Species Plantarum, at which time there were six known species (N. poeticus, N. pseudonarcissus, N. bulbocodium, N. serotinus, N. jonquilla and N. tazetta).[1] At that time, Linnaeus loosely grouped it together with 50 other genera into his Hexandria monogynia.
Modern
It was de Jussieu in 1789 who first formally created a 'family' (Narcissi), as the seventh 'Ordo' (Order) of the third class (Stamina epigyna) of Monocots in which Narcissus and 15 other genera were placed. The use of the term Ordo at that time was closer to what we now understand as Family, rather than Order. The family has undergone much reorganisation since then, but in 1805 it was renamed after a different genus in the family, Amaryllis, as 'Amaryllideae' by Jaume St.-Hilaire and has retained that association since. Jaume St.-Hilaire divided the family into two unnamed sections and recognised five species of Narcissus, omitting N. serotinus.
De Candolle brought together Linnaeus' genera and Jussieau's families into a systematic taxonomy for the first time, but included Narcissus (together with Amaryllis) in the Liliaceae in his Flore française (1805-1815) rather than Amaryllidaceae, a family he had not yet recognised. Shortly thereafter he separated the 'Amaryllidées' from 'Liliacées' (1813), though attributing the term to Brown's 'Amaryllideae' in the latter's Prodromus (1810) rather than St.-Hilaire's 'Amaryllidées'. He also provided the text to the first four volumes of Redouté illustrations in the latter's Les liliacées between 1805 and 1808 (see illustration here of N. candidissimus).
Historically both wide and narrow interpretations of the genus have been proposed. In the nineteenth century genus splitting was common, favouring the narrow view. Haworth (1831) using a narrow view treated many species as separate genera, as did Salisbury (1866). These authors listed various species in related genera such as Queltia (hybrids), Ajax (=Pseudonarcissus) and Hermione (=Tazettae), sixteen in all in Haworth's classification. In contrast, Herbert (1837) took a very wide view reducing Harworth's sixteen genera to six. Herbert, treating the Amaryllidacea as an 'order' as was common then, considered the narcissi to be a suborder, the Narcisseae, the six genera being Corbularia, Ajax, Ganymedes, Queltia, Narcissus and Hermione and his relatively narrow circumscription of Narcissus having only three species. Later Spach (1846) took an even wider view bringing most of Harworth's genera into the genus Narcissus, but as separate subgenera. By the time that Baker (1875) wrote his monograph all of the genera with one exception were included as Narcissus. The exception was the monotypic group Tapeinanthus which various subsequent authors have chosen to either exclude (e.g. Cullen 1986) or include (e.g. Webb 1978, 1980). Today it is nearly always included.
The eventual position of Narcissus within the Amaryllidaceae family only became settled in the twenty-first century with the advent of phylogenetic analysis and the Angiosperm Phylogeny Group system. The genus Narcissus belongs to the Narcisseae tribe, one of 13 within the Amaryllidoideae subfamily of the Amaryllidaceae. It is one of two sister clades corresponding to genera in the Narcisseae, being distinguished from Sternbergia by the presence of a paraperigonium, and is monophyletic
The infrageneric phylogeny of Narcissus still remains relatively unsettled. The taxonomy has proved very complex and difficult to resolve, particularly for the Pseudonarcissus group. This is due to a number of factors, including the diversity of the wild species, the ease with which natural hybridisation occurs, and extensive cultivation and breeding accompanied by escape and naturalisation.
De Candolle, in the first systematic taxonomy of Narcissus, arranged the species into named groups, and those names (Faux-Narcisse or Pseudonarcissus, Poétiques, Tazettes, Bulbocodiens, Jonquilles) have largely endured for the various subdivisions since and bear his name. The evolution of classification was confused by including many unknown or garden varieties, until Baker (1875) made the important distinction of excluding all specimens except the wild species from his system. He then grouped all of the earlier related genera as sections under one genus, Narcissus, the exception being the monotypic Tapeinanthus. Consequently, the number of accepted species has varied widely.
A common modern classification system has been that of Fernandes (1951, 1968, 1975) based on cytology, as modified by Blanchard (1990) and Mathew (2002), although in some countries such as Germany, the system of Meyer (1966) was preferred. Fernandes described two subgenera based on basal chromosome number, Hermione, n = 5 (11) and Narcissus, n = 7 (13). He further subdivided these into ten sections (Apodanthi, Aurelia, Bulbocodii, Ganymedes, Jonquillae, Narcissus, Pseudonarcissi, Serotini, Tapeinanthus, Tazettae), as did Blanchard later.
In contrast to Fernandes, Webb's treatment of the genus for the Flora Europaea (1978, 1980) prioritised morphology over genetics, and abandoned the subgenera ranks. He also restored De Candolle's original nomenclature, and made a number of changes to section Jonquilla, merging the existing subsections, reducing Apodanthi to a subsection of Jonquilla, and moving N. viridiflorus from Jonquilla to a new monotypic section of its own (Chloranthi). Finally, he divided Pseudonarcissus into two subsections. Blanchard (1990), whose Narcissus: a guide to wild daffodils has been very influential, adopted a simple approach, restoring Apodanthae, and based largely on ten sections alone.
The Royal Horticultural Society (RHS) currently lists ten sections, based on Fernandes (1968), three of which are monotypic (contain only one species), while two others only containing two species. Most species are placed in Pseudonarcissus While infrageneric groupings within Narcissus have been relatively constant, their status (genera, subgenera, sections, subsections, series, species) has not. Some authors treat some sections as being further subdivided into subsections, e.g. Tazettae (3 subsections). These subdivisions correspond roughly to the popular names for narcissi types, e.g. Trumpet Daffodils, Tazettas, Pheasant's Eyes, Hoop Petticoats, Jonquils.
While Webb had simply divided the genus into sections, Mathew found this unsatisfactory, implying every section had equal status. He adapted both Fernandes and Webb to devise a more hierarchical scheme he believed better reflected the interrelationships within the genus. Mathew's scheme consists of three subgenera (Narcissus, Hermione and Corbularia). The first two subgenera were then divided into five and two sections respectively. He then further subdivided two of the sections (subgenus Narcissus section Jonquillae, and subgenus Hermione section Hermione) into three subsections each. Finally, he divided section Hermione subsection Hermione further into two series, Hermione and Albiflorae. While lacking a phylogenetic basis, the system is still in use in horticulture. For instance the Pacific Bulb Society uses his numbering system for classifying species.
The phylogenetic analysis of Graham and Barrett (2004) supported the infrageneric division of Narcissus into two clades corresponding to the subgenera Hermione and Narcissus, but does not support monophyly of all sections, with only Apodanthi demonstrating clear monophyly, corresponding to Clade III of Graham and Barrett, although some other clades corresponded approximately to known sections. These authors examined 36 taxa of the 65 listed then, and a later extended analysis by Rønsted et al. (2008) with five additional taxa confirmed this pattern.
A very large (375 accessions) molecular analysis by Zonneveld (2008) utilising nuclear DNA content sought to reduce some of the paraphyly identified by Graham and Barrett. This led to a revision of the sectional structure, shifting some species between sections, eliminating one section and creating two new ones. In subgenus Hermione, Aurelia was merged with Tazettae. In subgenus Narcissus section Jonquillae subsection Juncifolii was elevated to sectional rank, thus resolving the paraphyly in this section observed by Graham and Barrett in Clade II due to this anomalous subsection, the remaining species being in subsection Jonquillae, which was monophyletic. The relatively large section Pseudonarcissi was divided by splitting off a new section, Nevadensis (species from southern Spain) leaving species from France, northern Spain and Portugal in the parent section. At the same time Fernández-Casas (2008) proposed a new monotypic section Angustini to accommodate Narcissus deficiens, placing it within subgenus Hermione.
While Graham and Barrett (2004) had determined that subgenus Hermione was monophyletic, using a much larger accession Santos-Gally et al. (2011) did not. However the former had excluded species of hybrid origins, while the latter included both N. dubius and N. tortifolius. If these two species are excluded (forming a clade with subgenus Narcissus) then Hermione can be considered monophyletic, although as a section of Hermione, Tazettae is not monophyletic. They also confirmed the monophyly of Apodanthi.
Some so-called nothosections have been proposed, predominantly by Fernández-Casas, to accommodate natural ('ancient') hybrids (nothospecies).
Subgenera and sections
Showing revisions by Zonnefeld (2008)
subgenus Hermione (Haw.) Spach.
(Aurelia (Gay) Baker (monotypic) - merged with Tazettae (2008)
Serotini Parlatore (2 species)
Tazettae de Candolle (16 species) syn. Hermione (Salisbury) Sprengel, in Fernandes' scheme. Incorporating Aurelia (2008)
subgenus Narcissus L.
Apodanthi A. Fernandes (6 species)
Bulbocodium de Candolle (11 species)
Ganymedes (Haworth) Schultes f. (monotypic)
Jonquillae de Candolle (8 species)
Juncifolii (A. Fern.) Zonn. sect. nov. (2008)
Narcissus L. (2 species)
Nevadensis Zonn. sect. nov. (2008)
Pseudonarcissus de Candolle (36 species) Trumpet daffodils
Tapeinanthus (Herbert) Traub (monotypic)
Species
Estimates of the number of species in Narcissus have varied widely, from anywhere between 16 and nearly 160, even in the modern era. Linnaeus originally included six species in 1753. By the time of the 14th edition of the Systema Naturae in 1784, there were fourteen. The 1819 Encyclopaedia Londinensis lists sixteen (see illustration here of three species) and by 1831 Adrian Haworth had described 150 species.
Much of the variation lies in the definition of species, and whether closely related taxa are considered separate species or subspecies. Thus, a very wide view of each species, such as Webb's results in few species, while a very narrow view such as that of Fernandes results in a larger number. Another factor is the status of hybrids, given natural hybridisation, with a distinction between 'ancient hybrids' and 'recent hybrids'. The term 'ancient hybrid' refers to hybrids found growing over a large area, and therefore now considered as separate species, while 'recent hybrid' refers to solitary plants found amongst their parents, with a more restricted range.
In the twentieth century Fernandes (1951) accepted 22 species, on which were based the 27 species listed by Webb in the 1980 Flora Europaea. By 1968, Fernandes had accepted 63 species, and by 1990 Blanchard listed 65 species, and Erhardt 66 in 1993. In 2006 the Royal Horticultural Society's (RHS) International Daffodil Register and Classified List listed 87 species, while Zonneveld's genetic study (2008) resulted in only 36. As of September 2014, the World Checklist of Selected Plant Families accepts 52 species, along with at least 60 hybrids, while the RHS has 81 accepted names in its October 2014 list.
Evolution
Within the Narcisseae, Narcissus (western Mediterranean) diverged from Sternbergia (Eurasia) some time in the Late Oligocene to Early Miocene eras, around 29.3–18.1 Ma, with a best estimate of 23.6 Ma. Later the genus divided into the two subgenera (Hermione and Narcissus) between 27.4 and 16.1 Ma (21.4 Ma). The divisions between the sections of Hermione then took place during the Miocene period 19.9–7.8 Ma.
Narcissus appears to have arisen in the area of the Iberian peninsula, southern France and northwestern Italy, and within this area most sections of the genus appeared, with only a few taxa being dispersed to North Africa at a time when the African and West European platforms were closer together. Subgenus Hermione in turn arose in the southwestern mediterranean and north west Africa. However, these are reconstructions, the Amaryllidaceae lacking a fossil record.
Names and etymology
The derivation of the Latin narcissus (Greek: νάρκισσος) is unknown. It may be a loanword from another language; for instance, it is said to be related to the Sanskrit word nark, meaning 'hell'. It is frequently linked to the Greek myth of Narcissus described by Ovid in his Metamorphoses, who became so obsessed with his own reflection that as he knelt and gazed into a pool of water, he fell into the water and drowned. In some variations, he died of starvation and thirst. In both versions, the narcissus plant sprang from where he died. Although Ovid appeared to describe the plant we now know as Narcissus there is no evidence for this popular derivation, and the person's name may have come from the flower's name. The Poet's Narcissus (N. poeticus), which grows in Greece, has a fragrance that has been described as intoxicating. This explanation is largely discredited due to lack of proof. Pliny wrote that the plant 'narce narcissum dictum, non a fabuloso puero' ('named narcissus from narce, not from the legendary youth'), i.e. that it was named for its narcotic properties (ναρκάω narkao, 'I grow numb' in Greek), not from the legend. Furthermore, there were accounts of narcissi growing, such as in the legend of Persephone, long before the story of Narcissus appeared (see Greek culture). It has also been suggested that daffodils bending over streams evoked the image of the youth admiring his own reflection in the water.
Linnaeus used the Latin name for the plant in formally describing the genus, although Matthias de l'Obel had previously used the name in describing various species of Narcissi in his Icones stirpium of 1591, and other publications, as had Clusius in Rariorum stirpium (1576).
The plural form of the common name narcissus has caused some confusion. British English sources such as the Shorter Oxford English Dictionary give two alternate forms, narcissi and narcissuses. In contrast, in American English the Merriam-Webster Dictionary provides for a third form, narcissus, used for both singular and plural. The Oxford dictionaries only list this third form under American English, although the Cambridge Dictionary allows of all three in the same order. However, Garner's Modern American Usage states that narcissi is the commonest form, narcissuses being excessively sibilant. For similar reasons, Fowler prefers narcissi in British English usage. Neither support narcissus as a plural form. Common names such as narcissus do not capitalise the first letter in contrast to the person of that name and the Latin genus name.
The name Narcissus (feminine Narcissa) was not uncommon in Roman times, such as Tiberius Claudius Narcissus, a Roman official in Claudius' time, an early New Testament Christian in Rome and later bishops and saints.
Daffodil
The word daffodil was unknown in the English language before the sixteenth century. The name is derived from an earlier affodell, a variant of asphodel. In classical Greek literature the narcissus is frequently referred to as the asphodel, such as the meadows of the Elysian fields in Homer (see Antiquity). Asphodel in turn appears to be a loanword coming from French via Mediaeval Latin affodilus from Classical Latin asphodilus and ultimately the Greek asphodelos (Greek: ἀσφόδελος). The reason for the introduction of the initial d is not known, although a probable source is an etymological merging from the Dutch article de, as in de affodil, or English the, as th'affodil or t'affodil, hence daffodil, and in French de and affodil to form fleur d'aphrodille and daphrodille.
From at least the 16th century, daffadown dilly and daffydowndilly have appeared as playful synonyms of the name. In common parlance and in historical documents, the term daffodil may refer specifically to populations or specimens of the wild daffodil, N. pseudonarcissus. H. N. Ellacombe suggests this may be from Saffon Lilly, citing Prior in support, though admittedly conjectural.
Lady Wilkinson (1858), who provides an extensive discussion of the etymology of the various names for this plant, suggests a very different origin, namely the Old English word affodyle (that which cometh early), citing a 14th-century (but likely originally much earlier) manuscript in support of this theory, and which appears to describe a plant resembling the daffodil. Ellacombe provides further support for this from a fifteenth century English translation of Palladius that also refers to it.
Jonquil
The name jonquil is said to be a corruption via French from the Latin juncifolius meaning 'rush-leaf' (Juncaceae) and its use is generally restricted to those species and cultivars which have rush like leaves, e.g. N. juncifolius.
Other
A profusion of names have attached themselves in the English language, either to the genus as a whole or to individual species or groups of species such as sections. These include narcissus, jonquil, Lent lily, Lenten lily, lide lily, yellow lily, wort or wyrt, Julians, glens, Lent cocks, corn flower, bell rose, asphodel, Solomon's lily, gracy day, haverdrils, giggary, cowslip, and crow foot.
A very very variable in color (polymorphic) species in the stinkbug family from South America. I have a couple of other examples, but there seems to be no limit to the color variations this species has. How lovely and mysterious. Not sure what the research is on this thing, but it must be an interesting story. This is what you find when you dig around in the National Collection at the Smithsonian. Sadly all the specimens are old as there is little collecting going on these days.
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All photographs are public domain, feel free to download and use as you wish.
Photography Information: Canon Mark II 5D, Zerene Stacker, Stackshot Sled, 65mm Canon MP-E 1-5X macro lens, Twin Macro Flash in Styrofoam Cooler, F5.0, ISO 100, Shutter Speed 200
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A very very variable in color (polymorphic) species in the stinkbug family from South America. I have a couple of other examples, but there seems to be no limit to the color variations this species has. How lovely and mysterious. Not sure what the research is on this thing, but it must be an interesting story. This is what you find when you dig around in the National Collection at the Smithsonian. Sadly all the specimens are old as there is little collecting going on these days.
~~~~~~~~~~{{{{{{0}}}}}}~~~~~~~~~~
All photographs are public domain, feel free to download and use as you wish.
Photography Information: Canon Mark II 5D, Zerene Stacker, Stackshot Sled, 65mm Canon MP-E 1-5X macro lens, Twin Macro Flash in Styrofoam Cooler, F5.0, ISO 100, Shutter Speed 200
Beauty is truth, truth beauty - that is all
Ye know on earth and all ye need to know
" Ode on a Grecian Urn"
John Keats
You can also follow us on Instagram account USGSBIML Want some Useful Links to the Techniques We Use? Well now here you go Citizen:
Art Photo Book: Bees: An Up-Close Look at Pollinators Around the World
www.qbookshop.com/products/216627/9780760347386/Bees.html...
Basic USGSBIML set up:
www.youtube.com/watch?v=S-_yvIsucOY
USGSBIML Photoshopping Technique: Note that we now have added using the burn tool at 50% opacity set to shadows to clean up the halos that bleed into the black background from "hot" color sections of the picture.
www.youtube.com/watch?v=Bdmx_8zqvN4
PDF of Basic USGSBIML Photography Set Up:
ftp://ftpext.usgs.gov/pub/er/md/laurel/Droege/How%20to%20Take%20MacroPhotographs%20of%20Insects%20BIML%20Lab2.pdf
Google Hangout Demonstration of Techniques:
plus.google.com/events/c5569losvskrv2nu606ltof8odo
or
www.youtube.com/watch?v=4c15neFttoU
Excellent Technical Form on Stacking:
Contact information:
Sam Droege
sdroege@usgs.gov
301 497 5840
Cuculus canorus
[order] Cuculiformes | [family] Cuculidae | [latin] Cuculus canorus | [UK] Cuckoo | [FR] Coucou gris | [DE] Kuckuck | [ES] Cuco Europeo | [IT] Cuculo eurasiatico | [NL] Koekoek | [IRL] Cuach
Status: Widespread summer visitor to Ireland from April to August.
Conservation Concern: Green-listed in Ireland. The European population is currently evaluated as secure.
Identification: Despite its obvious song, relatively infrequently seen. In flight, can be mistaken for a bird of prey such as Sparrowhawk, but has rapid wingbeats below the horizontal plane - ie. the wings are not raised above the body. Adult male Cuckoos are a uniform grey on the head, neck, back, wings and tail. The underparts are white with black barring. Adult females can appear in one of two forms. The so-called grey-morph resembles the adult male plumage, but has throat and breast barred black and white with yellowish wash. The rufous-morph has the grey replaced by rufous, with strong black barring on the wings, back and tail. Juvenile Cuckoos resemble the female rufous-morph, but are darker brown above.
Similar Species: Sparrowhawk
Call: The song is probably one of the most recognisable and well-known of all Irish bird species. The male gives a distinctive “wuck-oo”, which is occasionally doubled “wuck-uck-ooo”. The female has a distinctive bubbling “pupupupu”. The song period is late April to late June.
Diet: Mainly caterpillars and other insects.
Breeding: Widespread in Ireland, favouring open areas which hold their main Irish host species – Meadow Pipit. Has a remarkable breeding biology unlike any other Irish breeding species.
Wintering: Cuckoos winter in central and southern Africa.
To minimise the chance of being recognised and thus attacked by the birds they are trying to parasitize, female cuckoos have evolved different guises.
The common cuckoo (Cuculus canorus) lays its eggs in the nests of other birds. On hatching, the young cuckoo ejects the host's eggs and chicks from the nest, so the hosts end up raising a cuckoo chick rather than a brood of their own. To fight back, reed warblers (a common host across Europe) have a first line of defence: they attack, or ‘mob’, the female cuckoo, which reduces the chance that their nest is parasitized.
To deter the warbler from attacking, the colouring of the grey cuckoo mimics sparrow hawks, a common predator of reed warblers. However, other females are bright rufous (brownish-red). The presence of alternate colour morphs in the same species is rare in birds, but frequent among the females of parasitic cuckoo species. The new research shows that this is another cuckoo trick: cuckoos combat reed warbler mobbing by coming in different guises.
In the study, the researchers manipulated local frequencies of the more common grey colour cuckoo and the less common (in the United Kingdom) rufous colour cuckoo by placing models of the birds at neighbouring nests. They then recorded how the experience of watching their neighbours mob changed reed warbler responses to both cuckoos and a sparrow hawk at their own nest.
They found that reed warblers increased their mobbing, but only to the cuckoo morph that their neighbours had mobbed. Therefore, as one cuckoo morph increases in frequency, local host populations will become alerted specifically to that morph. This means the alternate morph will be more likely to slip past host defences and lay undetected. This is the first time that ‘social learning’ has been documented in the evolution of mimicry as well as the evolution of different observable characteristics - such as colour - in the same species (called polymorphism).
From the University of Cambridge “When mimicry becomes less effective, evolving to look completely different can be a successful trick. Our research shows that individuals assess disguises not only from personal experience, but also by observing others. However, because their learning is so specific, this social learning then selects for alternative cuckoo disguises and the arms race continues.”.
“It’s well known that cuckoos have evolved various egg types which mimic those of their hosts in order to combat rejection. This research shows that cuckoos have also evolved alternate female morphs to sneak through the hosts' defenses. This explains why many species which use mimicry, such as the cuckoo, evolve different guises.”
This is the first record of Fedia graciliflora Fish. and C.A. May in Crete. It is an example of fruits and seeds polymorphism which has three astonishingly different kinds of fruits that show adaptations to dispersal by wind and water, ants, and larger animals, respectively. It has dipsersed from Africa to S,\. Spain, S.France Italy and Malta.
The Lizard Lifeboat Station can refer to several Royal National Lifeboat Institution lifeboat stations located on the Lizard in Cornwall, United Kingdom. The first was established at the southernmost point of the peninsula in 1859. Since then successive stations have all been in operation at different locations on The Lizard. The current station is located at Kilcobben Cove 0.5 mi (0.80 km) east of the village of Lizard.
The lifeboat stations have all covered the westerly approaches to the English Channel; with up to 400 ships-a-day, it is one of the busiest shipping lanes in the world. The lifeboat service has saved many lives over the past 150 years.
The RNLI established its first lifeboat at the southern tip of The Lizard in 1859. The station, which cost £120 to build, was located atop the cliffs above Polpeor Cove about 0.6 mi (0.97 km) south of the village of Lizard. It was inaugurated after the 740-ton steamer, Czar, foundered on the Vrogue Rock, off Bass Point on 22 January 1859. The government transport ship was taking ammunition and uniforms to Malta. Fishing crews from Cadgwith and Church Cove saved some of the crew but the captain and his family drowned. Following the tragedy, a Mrs Agar of Lanhydrock donated money to buy the first Lizard Lifeboat (Anna Maria).
However the location of the first lifeboat station on the cliff above Polpeor Cove was not ideal as it made launches a long and precarious operation in rough sea and weather. On 2 January 1866 the lifeboat broke up after it was launched on exercise during a storm. It was pushed on to rocks causing the death of its Coxswain Peter Mitchell and crew members Richard Harris and Nicholas Stevens. As a tribute to the loss, the RNLI gave £130 to the local lifeboat fund. (Location: 49°57′32.53″N 5°12′22.77″W)
In 1885 a larger station was built above the high-water mark lower down in Polpeor Cove to house a larger lifeboat.[1] The existing smaller craft was moved to a new station at Church Cove just east of Lizard village. (Location: 49°57′33.51″N 5°12′24.53″W)
The final lifeboat station within Polpeor Cove was completed in 1914. The large concrete building had an integrated slipway which meant the lifeboat was able to launch directly into the sea. However this could prove hazardous in rough conditions because of the number of rocks in the cove. The exposed position of the station also meant it required a great deal of expense to maintain its general upkeep.
In order to relaunch the lifeboat, a recovery system was used to haul it back into the boat house. First ropes were places around a natural rock pillar in the sea in order to turn the stern of the boat towards land. A giant wheel – at the rear of the station – was then used to winch the boat back up the slipway.
The result of these difficulties meant the RNLI was forced to spend money repairing the station and the lifeboats from time to time. By 1958, with the need to employ larger and faster lifeboats due to the growth in maritime commerce, the RNLI decided to close Polpeor Cove because of its operating limitations. The RNLI eventually chose Kilcobben Cove as it new location for The Lizard Lifeboat station because it was sufficiently protected to allow safe launches in all conditions. Polpeor Cove closed in 1961. (Location: 49°57′33.56″N 5°12′25.57″W)
In 1867 the RNLI placed a second lifeboat at the fishing village of Cadgwith on the east side of the Lizard. This service ran until 1963, when it was finally closed. The station was then integrated with the service at The Lizard's Kilcobben Cove. (Location: 49°59′14.25″N 5°10′46.56″W)
Church Cove station, which was built at a cost of £300, opened in 1885. It housed the original lifeboat from the first station at Polpeor Cove when that was replaced with a new building and a larger vessel. Church Cove station, which was 0.4 mi (0.64 km) from the village of Landewednack was used in conjunction with Polpeor Cove for 14 years until it was closed and sold off in 1899. (Location: 49°58′16.49″N 5°11′18.62″W)
The RNLI decided that a new station on The Lizard would be built at Kilcobben Cove 1.25 mi (2.01 km) east of The Lizard lighthouse. Construction was a major civil engineering project because the station and its slipway were built on a cliff just above the waterline. The station, which cost £90,000, was opened on 7 July, 1961 by the Duke of Edinburgh, who also named the new Barnett-class lifeboat Duke of Cornwall ON952. Due to the steepness of the cliff, a funicular railway carries the lifeboat crew down to the boathouse.
The lifeboat station was originally called The Lizard-Cadgwith Lifeboat Station because it recognised the merging of the two former services based at Polpeor Cove and Cadgwith. This name was officially changed in 1987 to The Lizard Lifeboat Station. In 1988 the station and the slipway required adaptation with the arrival of a Tyne-class lifeboat called David Robinson ON1145.
In 2010 the original station was demolished because it could not accommodate the latest Tamar-class lifeboats. During the rebuilding the lifeboat was kept moored afloat off Cadgwith. On 5 May 2012, the new station was officially opened by Admiral the Lord Boyce, chairman of the RNLI. On the same day, the station's new Tamar-class boat, which had replaced David Robinson the year before, was named Rose in a ceremony by the-then Lord Lieutenant of Cornwall, Lady Mary Holborow.
The Lizard (Cornish: An Lysardh) is a peninsula in southern Cornwall, England, United Kingdom. The most southerly point of the British mainland is near Lizard Point at SW 701115; Lizard village, also known as The Lizard, is the most southerly on the British mainland, and is in the civil parish of Landewednack, the most southerly parish. The valleys of the River Helford and Loe Pool form the northern boundary, with the rest of the peninsula surrounded by sea. The area measures about 14 by 14 miles (23 km × 23 km). The Lizard is one of England's natural regions and has been designated as a National Character Area 157 by Natural England. The peninsula is known for its geology and for its rare plants and lies within the Cornwall Area of Outstanding Natural Beauty (AONB).
The Lizard's coast is particularly hazardous to shipping and the seaways round the peninsula were historically known as the "Graveyard of Ships" (see below). The Lizard Lighthouse was built at Lizard Point in 1752 and the RNLI operates The Lizard lifeboat station.
Etymology
The name "Lizard" is most probably a corruption of the Cornish name "Lys Ardh", meaning "high court"; it is purely coincidental that much of the peninsula is composed of serpentinite-bearing rock. The peninsula's original name may have been the Celtic Bridanoc, from Britannakon ("the "British one"), preserved in the name of the former village of Predannack, now site of Predannack Airfield.
History
There is evidence of early habitation with several burial mounds and stones. Part of the peninsula is known as the Meneage (land of the monks).
Helston, the nearest town to the Lizard peninsula, is said to have once headed the estuary of the River Cober, before it was cut off from the sea by Loe Bar in the 13th century. It is speculated that Helston was once a port, but no records exist. Geomorphologists believe the bar was most likely formed by rising sea levels, after the last ice age, blocking the river and creating a barrier beach. The beach is formed mostly of flint and the nearest source is found offshore under the drowned terraces of the former river that flowed between England and France, and now under the English Channel. The medieval port of Helston was at Gweek, possibly from around 1260 onwards, on the Helford river which exported tin and copper. Helston was believed to be in existence in the sixth century, around the River Cober (Dowr Kohar). The name comes from the Cornish "hen lis" or "old court" and "ton" added later to denote a Saxon manor; the Domesday Book refers to it as Henliston (which survives as the name of a road in the town). It was granted its charter by King John in 1201. It was here that tin ingots were weighed to determine the duty due to the Duke of Cornwall when a number of stannary towns were authorised by royal decree.
The royal manor of Winnianton, which was held by King William I at the time of the Domesday Book (1086), was also the head manor of the hundred of Kerrier and the largest estate in Cornwall. It was assessed as having fifteen hides before 1066. At the time of Domesday there was land for sixty ploughs, but in the lord's land there were two ploughs and in the lands held by villeins twenty-four ploughs. There were twenty-four villeins, forty-one freedmen, thirty-three smallholders and fourteen slaves. There was 6 acres (24,000 m2), eight square leagues of pasture and half a square league of woodland. The livestock was fourteen unbroken mares, three cattle and one hundred and twenty-eight sheep (in total 145 beasts); its value was £12 annually. 11 of the hides were held by the Count of Mortain and there is more arable and pasture and 13 more persons are recorded: Rinsey, Trelowarren, Mawgan-in-Meneage and seventeen other lands are also recorded under Winnianton.
Mullion has the 15th century church of St Mellanus, and the Old Inn from the 16th century. The harbour was completed in 1895 and financed by Lord Robartes of Lanhydrock as a recompense to the fishermen for several disastrous pilchard seasons.
The small church of St Peter in Coverack, built in 1885 for £500, has a serpentinite pulpit.
The Great Western Railway operated a road motor service to The Lizard from Helston railway station. Commencing on 17 August 1903, it was the first successful British railway-run bus service and was initially provided as a cheaper alternative to a proposed light railway.
The Solar eclipse of 11 August 1999 departed the UK mainland from the Lizard.
The transatlantic record run of the unaccompanied one hand sailor Thomas Coville within less than 5 days in his sailboat Sodebo Ultim from New York, USA, to Europe landed here on 15 July 2017.
Nautical
The Lizard has been the site of many maritime disasters. It forms a natural obstacle to entry and exit of Falmouth and its naturally deep estuary. At Lizard Point stands the Lizard Lighthouse. In fact, the light was erected by Sir John Killigrew by his own expense: It was built at the cost of "20 nobles a year" for 30 years, but it caused an uproar over the following years, as King James I considered charging vessels to pass. This caused so many problems that the lighthouse was demolished, but was successfully rebuilt in 1751 by order of Thomas Fonnereau and remains almost unchanged today. Further east lie The Manacles, near Porthoustock: 1+1⁄2 square miles (4 km2) of jagged rocks just beneath the waves.
In 1721 the Royal Anne Galley, an oared frigate, was wrecked at Lizard Point. Of a crew of 185 only three survived; lost was Lord Belhaven who was en route to take up the Governorship of Barbados.
A 44-gun frigate, HMS Anson, was wrecked at Loe Bar in 1807. Although it wrecked close to shore, many lost their lives in the storm. This inspired Henry Trengrouse to invent the rocket-fired line, later to become the Breeches buoy.
The transport ship Dispatch ran aground on the Manacles in 1809 on its return from the Peninsular War, losing 104 men from the 7th Hussars. The following day, with local villagers still attempting a rescue, the Cruizer-class brig-sloop HMS Primrose hit the northern end of these rocks. The only survivor of its 126 officers, men and boys was a drummer boy.
5 Sept 1856 the Cherubim and Ocean Home collided off Lizard Point
The SS Mohegan, a 6,889 GRT passenger liner, also hit the Manacles in 1898 with the loss of 106 lives.
The American passenger liner Paris was stranded on the Manacles in 1899, with no loss of life.
The biggest rescue in the RNLI's history was 17 March 1907 when the 12,000-tonne liner SS Suevic hit the Maenheere Reef near Lizard Point in Cornwall. In a strong gale and dense fog RNLI lifeboat volunteers rescued 456 passengers, including 70 babies. Crews from the Lizard, Cadgwith, Coverack and Porthleven rowed out repeatedly for 16 hours to rescue all of the people on board. Six silver RNLI medals were later awarded, two to Suevic crew members.
The Battle at the Lizard, a naval battle, took place off The Lizard on 21 October 1707.
Smuggling was a regular, and often necessary, way of life in these parts, despite the efforts of coastguards or "Preventive men". In 1801, the king's pardon was offered to any smuggler giving information on the Mullion musket men involved in a gunfight with the crew of HM Gun Vessel Hecate.
Avionic
In the First World War a Naval Air Station was established at Bonython, flying mainly blimps used for spotting U-boats. One was sunk and several probably damaged by bombs dropped by the blimps. The airfield site is now occupied by the wind farm.
RAF Predannack Down (see Predannack Airfield) was a Second World War airbase, from which Coastal Command squadrons flew anti-submarine sorties into the Bay of Biscay as well as convoy support in the western English Channel. The runways still exist and the site is used by a local Air Cadet Volunteergliding Squadron 626VGS and as an emergency/relief base for RNAS Culdrose (HMS Seahawk).
RNAS Culdrose is Europe's largest helicopter base, and currently hosts the Training and Operational Conversion Unit operating the EH101 "Merlin" helicopter. It is also the home base for Merlin Squadrons embarked upon Royal Navy warships, the Westland Sea King airborne early warning (AEW) variant helicopter, a Search And Rescue (Sea King, again) helicopter flight, and some BAe Hawk T.1 trainer jets used for training purposes by the Royal Navy. The base also operates some other types of fixed wing aircraft for calibration and other training purposes. As befits the base's name, a non-flying example of a Hawker Sea Hawk forms the main gate guardian static display. RNAS Culdrose is a major contributor to the economy of The Lizard area.
Political
The Lizard peninsula is in the St Ives parliamentary constituency (which comprises the whole of the former district of Penwith and the southern part of the former district of Kerrier). However, the parishes northeast of the Helford River are in Camborne and Redruth parliamentary constituency
To the north, The Lizard peninsula is bordered by the civil parishes of Breage, Porthleven, Sithney, Helston, Wendron, Gweek and – across the Helford River – by Constantine, Kerrier and Mawnan.
The parishes on the peninsula proper are (west to east):
Northern parishes:
Gunwalloe
Cury
Mawgan-in-Meneage
St Martin-in-Meneage
Manaccan
St Anthony-in-Meneage
Southern parishes:
Mullion
Grade-Ruan
St Keverne
Landewednack
The Lizard's political history includes the 1497 Cornish rebellion which began in St Keverne. The village blacksmith Michael Joseph (Michael An Gof in Cornish, meaning blacksmith) led the uprising, protesting against the punitive taxes levied by Henry VII to pay for the war against the Scots. The uprising was routed on its march to London and the two leaders, Michael Joseph and Thomas Flamank, were subsequently hanged, drawn and quartered.
Technological
Titanium was discovered here by the Reverend William Gregor in 1791.
In 1869, John Pender formed the Falmouth Gibraltar and Malta Telegraph company, intending to connect India to England with an undersea cable. Although intended to land at Falmouth, the final landing point was Porthcurno near Land's End.
In 1900 Guglielmo Marconi stayed the Housel Bay Hotel in his quest to locate a coastal radio station to receive signals from ships equipped with his apparatus. He leased a plot "in the wheat field adjoining the hotel" where the Lizard Wireless Telegraph Station still stands today. Recently restored by the National Trust, it looks as it did in January 1901, when Marconi received the distance record signals of 186 miles (299 km) from his transmitter station at Niton, Isle of Wight. The Lizard Wireless Station is the oldest Marconi station to survive in its original state, and is located to the west of the Lloyds Signal Station in what appears to be a wooden hut. On 12 December 1901 Poldhu Point was the site of the first trans Atlantic, wireless signal radio communication when Marconi sent a signal to St John's, Newfoundland. The technology is one of the key advances to the development of radio, television, satellites and the internet.
A radar station called RAF Dry Tree was built during World War II. The site was later chosen for the Telstar project in 1962; its rocky foundations, clear atmosphere and extreme southerly location being uniquely suitable. This became the Goonhilly satellite earth station, now owned by Goonhilly Earth Station Ltd. Some important developments in television satellite transmission were made at Goonhilly station. A wind farm exists near to the Goonhilly station site.
Geology
Known as the Lizard Complex, the peninsula's geology is the best preserved example of an exposed ophiolite in the United Kingdom.
An ophiolite is a suite of geological formations which represent a slice through a section of ocean crust (including the upper level of the mantle) thrust onto the continental crust.
The Lizard formations comprise three main units; the serpentinites, the "oceanic complex" and the metamorphic basement. The serpentinite contains significant samples of the serpentine polymorph lizardite, which were named after the Lizard complex in 1955.
Ecology
Several nature sites exist on the Lizard Peninsula; Predannack nature reserve, Mullion Island, Goonhilly Downs, and the Cornish Seal Sanctuary at Gweek. An area of the Lizard covering 16.62 square kilometres (6.42 sq mi) is designated a national nature reserve because of its coastal grasslands and heaths and inland heaths. The peninsula contains 3 main Sites of Special Scientific Interest (SSSI), both noted for their endangered insects and plants, as well as their geology. The first is East Lizard Heathlands SSSI, the second is Caerthillian to Kennack SSSI and the third is West Lizard SSSI, of which the important wetland, Hayle Kimbro Pool, forms a part of.
The area is also home to one of England's rarest breeding birds — the Cornish chough. This species of corvid is distinctive due to its red beak and legs and haunting "chee-aw" call. Choughs were extinct in Cornwall but returned naturally in 2001 and began breeding on Lizard in 2002 following a concerted effort by the National Trust, English Nature and the RSPB.
The Lizard contains some of the most specialised flora of any area in Britain, including many Red Data Book plant species. Of particular note is the Cornish heath, Erica vagans, that occurs in abundance here, but which is found nowhere else in Britain. There are more than 600 species of flowering plants on the Lizard, nearly a quarter of all UK species. The reason for this richness is partly because of the many different and unusual Lizard rocks on the Lizard Peninsula. But above all, it is a coming together of multiple factors: a very mild maritime climate, but one prone to gales and salt winds; waterlogged and boggy soils, but ones that often parch and dry out in the summer; soils of greatly contrasting fertility and pH; and lastly man's influence. Any single factor taken on its own would influence the flora; taken together, they combine, overlap and interact. Contrasting plant communities grow side-by-side in a mosaic that changes within a few metres but also changes markedly over time with the cycle of heath fires. It's not so much that conditions are ideal for growth, but that there is such a variety of different, difficult conditions. Each habitat, with its own combination of factors, attracts its own specialist plants. It is also one of the few places where the rare formicine ant, Formica exsecta, (the narrow-headed ant), can be found.
Portrayal in literature, film and music
Daphne du Maurier based many novels on this part of Cornwall, including Frenchman's Creek.
The Lizard was featured on the BBC television programme Seven Natural Wonders as one of the wonders of the South West, and on the BBC series Coast.
In James Clavell's novel Shōgun, ship's pilot Vasco Rodrigues challenges John Blackthorne to recite the latitude of the Lizard to verify that Blackthorne is the Pilot of the Dutch vessel Erasmus.
The Jennifer McQuiston 2015 novel The Spinster's Guide to Scandalous Behavior is set primarily in the fictional village Lizard Bay on the Lizard in the mid-nineteenth century.
In the television adaptation of "Horatio Hornblower", an order is given to "Weather the Lizard" in the episode Hornblower:Mutiny.
"Lizard Point" is also a track on the 1982 album Ambient 4: On Land released by Brian Eno.
The book series "Fenton House" by Ben Cheetham is set on the Lizard Peninsula.
Femme de l'ethnie Dao Longue Tunique sur le marché de Quyet Tien dans la région de Hagiang, nord du Vietnam
Les Dao (ou Dzao) sont, comme les Hmongs, très nombreux et l'ethnie est très polymorphe : Dao Rouge, Dao à Tunique, Dao Noir, Dao à pantalon blanc… .J'en ai déjà posté plusieurs exemples.
Emigrée de Chine du sud depuis le XIIIème siècle, l’ethnie Dao habite au Nord du Vietnam dans les provinces de la moyenne et haute région. Elle occupe des terres à toute altitude et vit en bons voisins avec d’autres ethnies comme les Hmong, Tay, Thaï, … Ses ressources principales proviennent de l’agriculture, (riz et mais)
Souvent chez les femmes, les cheveux sont relevés en chignon sur la nuque ou rasés sur le pourtour à l’exception d’une touffe au sommet de la tête
Pratiquant le culte des ancêtres, les Dao adhèrent également au Taoïsme.
Les femmes Dao Tunique (ou Longues Tunique) portent donc une longue tunique noire rehaussée à la ceinture et à la base de parement rouge vif. Elles portent une coiffe colorée. On les rencontre surtout dans la région de Hagiang.
Vesta is the largest and brightest asteroid in the asteroid belt and the second largest body overall (after the dwarf planet Ceres), with an average diameter of about 525 km (326 miles). That is pretty close to the size of the State of Colorado.
A couple of billion years ago two massive impacts ejected part of Vesta’s mass and some of that material landed here on Earth as HED (howardite, eucrite, and diogenite) meteorites. It’s astonishing to consider that when you look at photographs of Vesta, or eve through a powerful telescope, the actual craters from which the HED meteorites were blasted out can easily be seen, such as Rheasilvia which is over 300 miles wide.
Diogenites: “Vast underground chambers of magma churned and roiled, eventually cooling to the point at which orthopyroxene crystals froze out of the liquid rock. Insulated by kilometers of overlying rock and magma, they cooled slowly, growing into enormous sizes before settling downwards into vast piles of large crystals at the base of the underground caverns.” (Meteorite, p.102)
From the Meteorite Bulletin for NWA 7831:
History: Found buried in the ground near Chouichiyat in the Western Sahara on March 3, 2013, and excavated by a team of local people.
Physical characteristics: composed of translucent yellow-green crystals of orthopyroxene with pale orange weathering products along numerous fractures. Much of the material disintegrated into fragments upon excavation. The specimen is composed almost entirely of translucent,
yellow-green orthopyroxene with very sparse, tiny included grains of Ni-free metal, troilite, chromite, anorthite, silica polymorph and clinopyroxene. (see comments below for 2024 discovery related to the anorthite).
Stone meteorite - Diogenite, Achondrite: 5.5” by 4” by 2”, 1,301g, spent the past 17 years in the Michael Farmer "meteorite hunter" collection
Cuculus canorus
[order] Cuculiformes | [family] Cuculidae | [latin] Cuculus canorus | [UK] Cuckoo | [FR] Coucou gris | [DE] Kuckuck | [ES] Cuco Europeo | [IT] Cuculo eurasiatico | [NL] Koekoek | [IRL] Cuach
Status: Widespread summer visitor to Ireland from April to August.
Conservation Concern: Green-listed in Ireland. The European population is currently evaluated as secure.
Identification: Despite its obvious song, relatively infrequently seen. In flight, can be mistaken for a bird of prey such as Sparrowhawk, but has rapid wingbeats below the horizontal plane - ie. the wings are not raised above the body. Adult male Cuckoos are a uniform grey on the head, neck, back, wings and tail. The underparts are white with black barring. Adult females can appear in one of two forms. The so-called grey-morph resembles the adult male plumage, but has throat and breast barred black and white with yellowish wash. The rufous-morph has the grey replaced by rufous, with strong black barring on the wings, back and tail. Juvenile Cuckoos resemble the female rufous-morph, but are darker brown above.
Similar Species: Sparrowhawk
Call: The song is probably one of the most recognisable and well-known of all Irish bird species. The male gives a distinctive “wuck-oo”, which is occasionally doubled “wuck-uck-ooo”. The female has a distinctive bubbling “pupupupu”. The song period is late April to late June.
Diet: Mainly caterpillars and other insects.
Breeding: Widespread in Ireland, favouring open areas which hold their main Irish host species – Meadow Pipit. Has a remarkable breeding biology unlike any other Irish breeding species.
Wintering: Cuckoos winter in central and southern Africa.
To minimise the chance of being recognised and thus attacked by the birds they are trying to parasitize, female cuckoos have evolved different guises.
The common cuckoo (Cuculus canorus) lays its eggs in the nests of other birds. On hatching, the young cuckoo ejects the host's eggs and chicks from the nest, so the hosts end up raising a cuckoo chick rather than a brood of their own. To fight back, reed warblers (a common host across Europe) have a first line of defence: they attack, or ‘mob’, the female cuckoo, which reduces the chance that their nest is parasitized.
To deter the warbler from attacking, the colouring of the grey cuckoo mimics sparrow hawks, a common predator of reed warblers. However, other females are bright rufous (brownish-red). The presence of alternate colour morphs in the same species is rare in birds, but frequent among the females of parasitic cuckoo species. The new research shows that this is another cuckoo trick: cuckoos combat reed warbler mobbing by coming in different guises.
In the study, the researchers manipulated local frequencies of the more common grey colour cuckoo and the less common (in the United Kingdom) rufous colour cuckoo by placing models of the birds at neighbouring nests. They then recorded how the experience of watching their neighbours mob changed reed warbler responses to both cuckoos and a sparrow hawk at their own nest.
They found that reed warblers increased their mobbing, but only to the cuckoo morph that their neighbours had mobbed. Therefore, as one cuckoo morph increases in frequency, local host populations will become alerted specifically to that morph. This means the alternate morph will be more likely to slip past host defences and lay undetected. This is the first time that ‘social learning’ has been documented in the evolution of mimicry as well as the evolution of different observable characteristics - such as colour - in the same species (called polymorphism).
From the University of Cambridge “When mimicry becomes less effective, evolving to look completely different can be a successful trick. Our research shows that individuals assess disguises not only from personal experience, but also by observing others. However, because their learning is so specific, this social learning then selects for alternative cuckoo disguises and the arms race continues.”.
“It’s well known that cuckoos have evolved various egg types which mimic those of their hosts in order to combat rejection. This research shows that cuckoos have also evolved alternate female morphs to sneak through the hosts' defenses. This explains why many species which use mimicry, such as the cuckoo, evolve different guises.”
EMERSON,LAKE AND PALMER-From the Beginning(SUBTITULADA)
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Femme Hmong Noir dans une rue de Sapa, nord du Vietnam
Avec la ligne de train et désormais l'autoroute la reliant à Hanoï, Sapa est devenue une destination touristique très fréquentée et a beaucoup perdu de son charme, même si les promenades dans les rizières alentour restent bien agréables. On y croise cependant de pittoresques personnages telle cette femme souriante dans son costume Hmong Noir
Avec les Dao Rouge, les Hmong Noir constituent l'une des ethnies les mieux représentées dans la région de Sapa.
Les Hmong encore appelés Méo, ou Miao , sont originaires des régions montagneuses du sud de la Chine (principalement la province du Guizhou), où ils sont encore présents ainsi qu'au nord du Viêt Nam et du Laos.
Le souci de préserver leur identité culturelle et leur indépendance les ont amenés à s'engager dans divers conflits. Au XXe siècle, en particulier, ils aidèrent les Français pendant la guerre d'Indochine puis les Américains pendant la la guerre du Vietnam. A l’avènement des régimes communistes dans ces pays un nombre important de Hmong se sont réfugiés dans des pays d'accueil, principalement les États-Unis, la France et l'Australie. Mais la majeure partie d’entre eux vit encore en Asie du Sud-Est
Les Hmongs sont animistes ou chrétiens. La langue hmong appartient à la famille des langues hmong-mien, encore appelée « miao-yao »
Les costumes traditionnels de cette ethnie sont très polymorphes mais ils ont en commun la richesse du décor brodé bien que celui de cette femmes Hmong Noir soit très sobre.