View allAll Photos Tagged Genetics
When the Blue-Yellow family gathers for a family portrait outside the Pindale Aquarium, one notices an excellent (albeit oversimplified) example of mendelian genetics in their progeny make-up:
little blue (AA),
little yellow (aa)
and the twins (Aa and aA, they are not identical ~grin~) :)
"100 possibilities project" clothespins
83/100
scavenger16 patterns in nature........ this is probably a real stretch ~grin~
365colours.......green
happy cliche saturday
cliche: using overly simple examples to explain complex scientific principles. (but it sure was FUN)
................dithering again-- pulled it out of cliche saturday
Some screenshots from a tool that I am building to visualize flue genomic data.
Some screenshots from a tool that I am building to visualize flue genomic data.
For more information, read this blog post:
blog.blprnt.com/blog/blprnt/open-science-h1n1-processing-...
I would have bought this photograph even if it didn't have its wonderful deterioration, but I just love the way it is breaking down. I guess I'm not a very good salesperson for Photoshop. From the flea market, 2/19/06.
GSA-Genetics Society of America
Title: An ancient regulator of sex development
Author: Marisa Wexler
Photo Credit: Ho-Wen Chen
Date: March 12, 2019.
Citation: genestogenomes.org/an-ancient-regulator-of-sex-development/
A Wnt protein involved in the formation of the human ovary plays an important role in female zebrafish sex development.
Even though zebrafish are a well-studied research model, how these fish develop into males or females remains rather obscure—in part because the sex of lab strains is not determined by sex chromosomes. Research published in GENETICS reveals that one of the genes critical for proper development of mammalian ovaries seems to play a related role in zebrafish, providing insight into this major difference between fish and mammals.
In humans and other mammals, Wnt4 encodes a signaling molecule that antagonizes the male-promoting signal FGF9 and is crucial to the proper development of the ovaries. Though they lack an Fgf9 ortholog, zebrafish have two Wnt4-like genes—wnt4a and wnt4b—so Kossack et al. investigated whether these genes might play a role in zebrafish sexual development.
The authors first performed a phylogenetic analysis to better understand how the Wnt4-like genes have changed over evolutionary time. Zebrafish have two such genes, while mammals only have one, so two evolutionary scenarios are possible: either fish gained an extra copy of the gene, or mammals lost a copy that was originally present. The authors found that the latter scenario is more likely; both reptiles and birds have two such genes, making it more likely that mammals lost one of their copies. Further analysis revealed that wnt4a is likely the ortholog of mammalian Wnt4, whereas wnt4b was lost in mammals after they split from birds.
The authors next used RT-PCR to examine the expression patterns of the two genes. They detected wnt4a, but not wnt4b, in the ovaries of female zebrafish. In contrast, only wnt4b was detected in the testis of male fish. They also found that wnt4a was dynamically expressed in gonads during development in a non-sex-specific manner.
Fish mutant for wnt4a develop predominantly—though not exclusively—as males, which supports the idea that wnt4a is involved in either differentiation into a female or the maintenance of a female phenotype throughout development. Analysis of mutant embryos during development suggests that wnt4a likely promotes female development since most mutant embryos developed as males rather than “reverting” to males from an initially female phenotype.
Interestingly, wnt4a mutants were unable to produce progeny when mated to each other or to wild-type. Closer inspection revealed that wnt4a-mutant fish of both sexes had malformed reproductive tracts. Even though viable eggs and sperm could be obtained from their gonads, mutant fish were unable to release their gametes, preventing them from reproducing.
These findings suggest that Wnt4-like genes have been involved in female development of a diverse array of animals—including our fishy ancestors that swam the oceans some 450 millions year ago.
CITATION:
Female Sex Development and Reproductive Duct Formation Depend on Wnt4a in Zebrafish
Michelle E. Kossack, Samantha K. High, Rachel E. Hopton, Yi-lin Yan, John H. Postlethwait, Bruce W. Draper
GENETICS January 2019 211: 219-233; doi.org/10.1534/genetics.118.301620
Bonefish researchers Chris Haak (Fisheries Conservation Foundation) and Liz Wallace (FWRI) sort through the catch from a light trap.
Photo by Cindy Pinder
this happens when you have white and purple next to each other, you get ‘mixed’ ones: whurple??? LOLOLOL
Asters are almost the latest flowers in the garden here, still some colour though, despite the weather.
The most popular asters are Monte Casino and the New England aster.
Asters range in colours of white, red, pink, purple, lavender and blue.
Like the sunflower, the Aster flower is actually a collection of very tiny tubular flowers, which are grouped together in a central disk and surrounded by a “ray” of flowers or petals.
Asters are one of the easiest garden perennials to cultivate.
Have a wonderful day, and thank you for your time and comments, M, (*_*)
For more: www.indigo2photography.com
IT IS STRICTLY FORBIDDEN (BY LAW!!!) TO USE ANY OF MY image or TEXT on websites, blogs or any other media without my explicit permission. © All rights reserved
blooms, Asters, petals, plenty, floral, flowers, purple, mixed, colourful, black-background, colour, design, square, Magda indigo
I've been calling these sunflowers, but they may be rosin weed. Sigh. These yellow flowers are a bit like little brown shore birds - they don't have ID tags. At least, they don't have ID tags that I can figure out.
Anyway, you can see the top flowers are double blossoms - they have more petals than the bloom below. This is because the genes that says to produce the stamen and carpel are either deleted or damaged, and so more petals are produced. That's all the science I have for today!
More than 200 Iowa high school seniors and their parents attended the 2nd annual UI Health Sciences Visit Day on Saturday, June 21, 2014. Students chose from 14 different interactive health sciences sessions to engage in hands-on STEM learning to consider careers in science, medicine, and health care and learn more about the University of Iowa! The rising seniors studied cancer tumors, the genetics of flies, emergency room simulations, blood typing, assessing and diagnosing patients just to name a few experiences.
www.medical-explorer.com/genetics.php
Genetics is the science of genes, heredity, and the variation of organisms.
This is a linocut portrait of Gregor Mendel (1822-1884), scientist and Augustinian friar who posthumously gained fame for establishing many of the rules of heredity, fundamental to modern genetics. By carefully crossbreeeding pea plants and tracing seven characterististics (plant height, pod shape and colour, seed shape and colour, and flower position and colour) he was able to deduce what are now referred to as the laws of Mendelian inheritance. He coined the terms "recessive" and "dominant" traits. Some of his findings are subtlely alluded to in the layout of the pea flowers, like a Punnett square depicting a cross between two pea plants heterozygous for purple and white blossoms. The first edition is a variable run of 8 prints, each 11" by 14" (28 cm by 35.6 cm), on ivory Japanese kozo paper with "chine-collé" white and mauve paper.
genetics... its really a crap shoot... ant facially resembles v but he is built like and has that odd analytical bent like his dad...
now katie's face resembles mine... but her soul and her build come from her mother... she's sure to be 6 ft...
and all that blond hair... well that's mine :-)
#AbFav_HANDS_FEET_LEGS_
GENETICS... MOTHER and SON.
A mum in the park, Amitah Bachan, the famous Indian actor in the studio and Willem in communion with his zither.
And finally, smoking a cigaret for the 1940-ties portrait.
Wishing you a beautiful day and as always, thanks for visiting, Magda (*_*)
For more: www.indigo2photography.com
IT IS STRICTLY FORBIDDEN (BY LAW!!!) TO USE ANY OF MY image or TEXT on websites, blogs or any other media without my explicit permission. © All rights reserved
GENETICS, portrait, two, family, mother, woman, boy, son, studio, strong, face, hands, black and white, Ilford film, vertical, monochrome, Hasselblad, Magda-indigo
#AbFav_HANDS_FEET_LEGS_
GENETICS... MOTHER and SON.
I got this lovely family with 3 children and their dog in the studio.
Whilst the two youngest looked very much like their dad, her eldest son was just the spitting image of his mum.
AAAhhh, the Art of genetics.
I placed their hands like that in this portrait because often I find they play such a vital role and say so much.
Taken on the Hasselblad, which has square negs, so it is slightly cropped.
Scanned from the print.
The 80mm Zeiss lens and the film Ilford Delta 100.
A mum in the park, Amitah Bachan, the famous Indian actor in the studio and Willem in communion with his zither.
And finally, smoking a cigaret for the 1940-ties portrait.
Wishing you a beautiful day and as always, thanks for visiting, Magda (*_*)
For more: www.indigo2photography.com
IT IS STRICTLY FORBIDDEN (BY LAW!!!) TO USE ANY OF MY image or TEXT on websites, blogs or any other media without my explicit permission. © All rights reserved
GENETICS, portrait, two, family, mother, woman, boy, son, studio, strong, face, hands, black and white, Ilford film, vertical, monochrome, Hasselblad, Magda-indigo
Scientists can now modify genes or replace faulty genes with healthy ones to treat, cure or prevent a disease or medical condition. Gene therapy can be performed both inside and outside the body. This graphic illustrates in simple terms how gene therapy works inside the body. Read the FDA Consumer Update to learn more.
This infographic is an original FDA creation and in the public domain. It may be shared, downloaded and redistributed without restriction. Credit to FDA is appreciated but not required.
FDA infographic by Michael J. Ermarth
Ouzo's typical itinerary in placing the ball for the next throw. I love that snow alows me to capture it on camera. Here you can see how he came from where I was standing, performed almost a full clockwise circle, placed the ball, finished the circle and placed himself on balance from the ball and I :)
I visit my father once a year, as we live in different countries. I’m his oldest son (of 6 kids), he is a retired professor of engineering and he tried to make an engineer of me to , but got with lucky an architect ,(as a mutual agreement we made years ago). I showed him my flickr gallery, he liked the landscapes, (very few) and he doesn't know how to react with some of the , (on his words) "the abstract stuff, that in his view is 99% of the stream.
I told him that I haven't used yet the remote control of my new camera. He got excited about testing it, specialy reading the manual of instructions. We did a first try and just clicked this photo.
Eric Vilain, Professor of Human Genetics, Pediatrics and Urology; Director, Center for Gender-Based Biology, David Geffen School of Medicine at UCLA, USA speaking during the Session: Ask About: The Genetics of Sex and Gender at the Annual Meeting 2017 of the World Economic Forum in Davos, January 18, 2017
Copyright by World Economic Forum / Sikarin Thanachaiary
Picture taken 1983 - digitally captured from paper print
__________________________________________
The orangutans (also spelled orang-utan, orangutang, or orang-utang) are the two exclusively Asian species of extant great apes. Native to Indonesia and Malaysia, orangutans are currently found in only the rainforests of Borneo and Sumatra. Classified in the genus Pongo, orangutans were considered to be one species. Since 1996, they have been divided into two species: the Bornean orangutan (P. pygmaeus) and the Sumatran orangutan (P. abelii). In addition, the Bornean species is divided into three subspecies.
Based on genome sequencing, the two extant orangutan species evidently diverged around 400,000 years ago. The orangutans are also the only surviving species of the subfamily Ponginae, which also included several other species, such as the three extinct species of the genus Gigantopithecus, including the largest known primate Gigantopithecus blacki. The ancestors of the Ponginae subfamily split from the main ape line in Africa 16 to 19 million years ago (mya) and spread into Asia.
Orangutans are the most arboreal of the great apes and spend most of their time in trees. Their hair is typically reddish-brown, instead of the brown or black hair typical of chimpanzees and gorillas. Males and females differ in size and appearance. Dominant adult males have distinctive cheek pads and produce long calls that attract females and intimidate rivals. Younger males do not have these characteristics and resemble adult females. Orangutans are the most solitary of the great apes, with social bonds occurring primarily between mothers and their dependent offspring, who stay together for the first two years. Fruit is the most important component of an orangutan's diet; however, the apes will also eat vegetation, bark, honey, insects and even bird eggs. They can live over 30 years in both the wild and captivity.
Orangutans are among the most intelligent primates; they use a variety of sophisticated tools and construct elaborate sleeping nests each night from branches and foliage. The apes have been extensively studied for their learning abilities. There may even be distinctive cultures within populations. Field studies of the apes were pioneered by primatologist Birutė Galdikas. Both orangutan species are considered to be endangered, with the Sumatran orangutan being critically endangered. Human activities have caused severe declines in the populations and ranges of both species. Threats to wild orangutan populations include poaching, habitat destruction, and the illegal pet trade. Several conservation and rehabilitation organisations are dedicated to the survival of orangutans in the wild.
ETYMOLOGY
The name "orangutan" (also written orang-utan, orang utan, orangutang, and ourang-outang) is derived from the Malay and Indonesian words orang meaning "person" and hutan meaning "forest", thus "person of the forest". Orang Hutan was originally not used to refer to apes, but to forest-dwelling humans.
The Malay words used to refer specifically to the ape are maias and mawas, but it is unclear if those words refer to just orangutans, or to all apes in general. The first attestation of the word to name the Asian ape is in Dutch physician Jacobus Bontius' 1631 Historiae naturalis et medicae Indiae orientalis – he reported that Malays had informed him the ape was able to talk, but preferred not to "lest he be compelled to labour". The word appeared in several German-language descriptions of Indonesian zoology in the 17th century. The likely origin of the word comes specifically from the Banjarese variety of Malay.
Cribb et al. (2014) suggest that Bontius' account referred not to apes (which were not known from Java) but rather to humans suffering some serious medical condition (most likely endemic cretinism) and that his use of the word was misunderstood by Nicolaes Tulp, who was the first to use the term in a publication.
The word was first attested in English in 1691 in the form orang-outang, and variants with -ng instead of -n as in the Malay original are found in many languages. This spelling (and pronunciation) has remained in use in English up to the present, but has come to be regarded as incorrect. The loss of "h" in Utan and the shift from n to -ng has been taken to suggest that the term entered English through Portuguese. In 1869, British naturalist Alfred Russel Wallace, co-creator of modern evolutionary theory, published his account of Malaysia's wildlife: The Malay Archipelago: The Land of the Orang-Utan and the Bird of Paradise.
The name of the genus, Pongo, comes from a 16th-century account by Andrew Battell, an English sailor held prisoner by the Portuguese in Angola, which describes two anthropoid "monsters" named Pongo and Engeco. He is now believed to have been describing gorillas, but in the 18th century, the terms orangutan and pongo were used for all great apes. Lacépède used the term Pongo for the genus following the German botanist Friedrich von Wurmb who sent a skeleton from the Indies to Europe.
TAXONOMY, PHYLOGENY AND GENETICS
The two orangutan species are the only extant members of the subfamily Ponginae. This subfamily also included the extinct genera Lufengpithecus, which lived in southern China and Thailand 2–8 mya, and Sivapithecus, which lived India and Pakistan from 12.5 mya until 8.5 mya. These apes likely lived in drier and cooler environments than orangutans do today. Khoratpithecus piriyai, which lived in Thailand 5–7 mya, is believed to have been the closest known relative of the orangutans. The largest known primate, Gigantopithecus, was also a member of Ponginae and lived in China, India and Vietnam from 5 mya to 100,000 years ago.
Within apes (superfamily Hominoidea), the gibbons diverged during the early Miocene (between 19.7 and 24.1 mya, according to molecular evidence) and the orangutans split from the African great ape lineage between 15.7 and 19.3 mya.
HISTORY OF ORANGUTAN TAXONOMY
The orangutan was first described scientifically in the Systema Naturae of Linnaeus as Simia satyrus. The populations on the two islands were classified as subspecies until 1996, when they were elevated to full species status, and the three distinct populations on Borneo were elevated to subspecies. The population currently listed as P. p. wurmbii may be closer to the Sumatran orangutan than the other Bornean orangutan subspecies. If confirmed, abelii would be a subspecies of P. wurmbii (Tiedeman, 1808).
Regardless, the type locality of P. pygmaeus has not been established beyond doubts, and may be from the population currently listed as P. wurmbii (in which case P. wurmbii would be a junior synonym of P. pygmaeus, while one of the names currently considered a junior synonym of P. pygmaeus would take precedence for the northwest Bornean taxon). To further confuse, the name P. morio, as well as some suggested junior synonyms, may be junior synonyms of the P. pygmaeus subspecies, thus leaving the east Bornean populations unnamed.
In addition, some fossils described under the name P. hooijeri have been found in Vietnam, and multiple fossil subspecies have been described from several parts of southeastern Asia. It is unclear if these belong to P. pygmaeus or
P. abelii or, in fact, represent distinct species.
GENOMICS
The Sumatran orangutan genome was sequenced in January 2011. Following humans and chimpanzees, the Sumatran orangutan has become the third species of hominid to have its genome sequenced. Subsequently, the Bornean species would have its genome sequenced. Genetic diversity was found to be lower in Bornean orangutans (P. pygmaeus) than in Sumatran ones (P. abelii), despite the fact that Borneo is home to six or seven times as many orangutans as Sumatra.
The comparison has shown these two species diverged around 400,000 years ago, more recently than was previously thought. Also, the orangutan genome was found to have evolved much more slowly than chimpanzee and human DNA. Previously, the species was estimated to have diverged 2.9 to 4.9 mya. The researchers hope these data may help conservationists save the endangered ape, and also prove useful in further understanding of human genetic diseases.
Bornean orangutans have 48 diploid chromosomes.
ANATOMY AND PHYSIOLOGY
An orangutan has a large, bulky body, a thick neck, very long, strong arms, short, bowed legs, and no tail. It is mostly covered with long, reddish-brown hair and grey-black skin. Sumatran orangutans have more sparse and lighter-coloured coats. The orangutan has a large head with a prominent mouth area. Though largely hairless, their faces can develop some hair in males, giving them a moustache.
Adult males have large cheek flaps to show their dominance to other males. The cheek flaps are made mostly of fatty tissue and are supported by the musculature of the face. Mature males' throat pouches allow them to make loud calls. The species display significant sexual dimorphism; females typically stand 115 cm tall and weigh around 37 kg, while flanged adult males stand 136 cm tall and weigh 75 kg. A male orangutan has an arm span of about 2 m.
Orangutan hands are similar to human hands; they have four long fingers and an opposable thumb. However, the joint and tendon arrangement in the orangutans' hands produces two adaptations that are significant for arboreal locomotion. The resting configuration of the fingers is curved, creating a suspensory hook grip. Additionally, without the use of the thumb, the fingers and hands can grip tightly around objects with a small diameter by resting the tops of the fingers against the inside of the palm, creating a double-locked grip.
Their feet have four long toes and an opposable big toe. Orangutans can grasp things with both their hands and their feet. Their fingers and toes are curved, allowing them to get a better grip on branches. Since their hip joints have the same flexibility as their shoulder and arm joints, orangutans have less restriction in the movements of their legs than humans have. Unlike gorillas and chimpanzees, orangutans are not true knuckle-walkers, and are instead fist-walkers.
ECOLOGY AND BEHAVIOUR
Orangutans live in primary and old secondary forests, particularly dipterocarp forests and peat swamp forests. Both species can be found in mountainous and lowland swampy areas. Sumatran orangutans live at elevations as high as 1500 m, while Bornean orangutans live no higher than 1000 m. Other habitats used by orangutans include grasslands, cultivated fields, gardens, young secondary forest, and shallow lakes. Orangutans are the most arboreal of the great apes, spending nearly all their time in the trees.Most of the day is spent feeding, resting, and travelling. They start the day feeding for 2–3 hours in the morning. They rest during midday then travel in the late afternoon. When evening arrives, they begin to prepare their nests for the night. Orangutans do not swim, although they have been recorded wading in water. The main predators of orangutans are tigers. Other predators include clouded leopards, wild dogs and crocodiles. The absence of tigers on Borneo may explain why Bornean orangutans can be found on the ground more often than their Sumatran relatives.
DIET
Orangutans are opportunistic foragers, and their diets vary markedly from month to month. Fruit makes up 65–90% of the orangutan diet, and those with sugary or fatty pulp are favoured. Ficus fruits are commonly eaten and are easy to harvest and digest. Lowland dipterocarp forests are preferred by orangutans because of their plentiful fruit. Bornean orangutans consume at least 317 different food items that include young leaves, shoots, bark, insects, honey and bird eggs.
A decade-long study of urine and faecal samples at the Gunung Palung Orangutan Conservation Project in West Kalimantan has shown that orangutans give birth during and after the high fruit season (though not every year), during which they consume various abundant fruits, totalling up to 11,000 calories per day. In the low-fruit season, they eat whatever fruit is available in addition to tree bark and leaves, with daily intake at only 2,000 calories. Together with a long lactation period, orangutans also have a long birth interval.
Orangutans are thought to be the sole fruit disperser for some plant species including the climber species Strychnos ignatii which contains the toxic alkaloid strychnine. It does not appear to have any effect on orangutans except for excessive saliva production.
Geophagy, the practice of eating soil or rock, has been observed in orangutans. There are three main reasons for this dietary behaviour: for the addition of mineral nutrients to their diet; for the ingestion of clay minerals that can absorb toxic substances; or to treat a disorder such as diarrhoea. Orangutans also use plants of the genus Commelina as an anti-inflammatory balm.
SOCIAL LIFE
Orangutans live a more solitary lifestyle than the other great apes. Most social bonds occur between adult females and their dependent and weaned offspring. Adult males and independent adolescents of both sexes tend to live alone. Orangutan societies are made up of resident and transient individuals of both sexes. Resident females live with their offspring in defined home ranges that overlap with those of other adult females, which may be their immediate relatives. One to several resident female home ranges are encompassed within the home range of a resident male, who is their main mating partner.
Transient males and females move widely. Orangutans usually travel alone, but they may travel in small groups in their subadult years. However, this behaviour ends at adulthood. The social structure of the orangutan can be best described as solitary but social. Interactions between adult females range from friendly to avoidance to antagonistic. Resident males may have overlapping ranges and interactions between them tend to be hostile.
During dispersal, females tend to settle in home ranges that overlap with their mothers. However, they do not seem to have any special social bonds with them. Males disperse much farther from their mothers and enter into a transient phase. This phase lasts until a male can challenge and displace a dominant, resident male from his home range. Adult males dominate sub-adult males.
Both resident and transient orangutans aggregate on large fruiting trees to feed. The fruits tend to be abundant, so competition is low and individuals may engage in social interactions. Orangutans will also form travelling groups with members moving between different food sources. These groups tend to be made of only a few individuals. They also tend to be consortships between an adult male and female.
COMMUNICATION
Orangutans communicate with various sounds. Males will make long calls, both to attract females and advertise themselves to other males. Both sexes will try to intimidate conspecifics with a series of low guttural noises known collectively as the "rolling call". When annoyed, an orangutan will suck in air through pursed lips, making a kissing sound that is hence known as the "kiss squeak". Infants make soft hoots when distressed. Orangutans are also known to blow raspberries.
NESTING
Orangutans build nests specialized for both day or night use. These are carefully constructed; young orangutans learn from observing their mother's nest-building behaviour. In fact, nest-building is a leading cause in young orangutans leaving their mother for the first time. From six months of age onwards, orangutans practice nest-building and gain proficiency by the time they are three years old.
Construction of a night nest is done by following a sequence of steps. Initially, a suitable tree is located, orangutans being selective about sites though many tree species are used. The nest is then built by pulling together branches under them and joining them at a point. After the foundation has been built, the orangutan bends smaller, leafy branches onto the foundation; this serves the purpose of and is termed the "mattress". After this, orangutans stand and braid the tips of branches into the mattress. Doing this increases the stability of the nest and forms the final act of nest-building. In addition, orangutans may add additional features, such as "pillows", "blankets", "roofs" and "bunk-beds" to their nests.
REPRODUCTION AND PARENTING
Males mature at around 15 years of age, by which time they have fully descended testicles and can reproduce. However, they exhibit arrested development by not developing the distinctive cheek pads, pronounced throat pouches, long fur, or long-calls until they are between 15 and 20 years old. The development of these characteristics depends largely on the absence of a resident male.
Males without them are known as unflanged males in contrast to the more developed flanged males. The transformation from unflanged to flanged can occur very quickly. Unflanged and flanged males have two different mating strategies. Flanged males attract oestrous females with their characteristic long calls. Those calls may also suppress development in younger males. Unflanged males wander widely in search of oestrous females and upon finding one, will force copulation on her. While both strategies are successful, females prefer to mate with flanged males and seek their company for protection against unflanged males. Resident males may form consortships with females that can last days, weeks or months after copulation.
Female orangutans experience their first ovulatory cycle around 5.8–11.1 years. These occur earlier in females with more body fat. Like other great apes, female orangutans enter a period of infertility during adolescence which may last for 1–4 years. Female orangutans also have a 22– to 30-day menstrual cycle. Gestation lasts for 9 months, with females giving birth to their first offspring between the ages of 14 and 15 years.
Female orangutans have eight-year intervals between births, the longest interbirth intervals among the great apes. Unlike many other primates, male orangutans do not seem to practice infanticide. This may be because they cannot ensure they will sire a female's next offspring because she does not immediately begin ovulating again after her infant dies.
Male orangutans play almost no role in raising the young. Females do most of the caring and socializing of the young. A female often has an older offspring with her to help in socializing the infant. Infant orangutans are completely dependent on their mothers for the first two years of their lives. The mother will carry the infant during travelling, as well as feed it and sleep with it in the same night nest. For the first four months, the infant is carried on its belly and never relieves physical contact. In the following months, the time an infant spends with its mother decreases.
When an orangutan reaches the age of two, its climbing skills improve and it will travel through the canopy holding hands with other orangutans, a behaviour known as "buddy travel". Orangutans are juveniles from about two to five years of age and will start to temporarily move away from their mothers. Juveniles are usually weaned at about four years of age. Adolescent orangutans will socialize with their peers while still having contact with their mothers. Typically, orangutans live over 30 years in both the wild and captivity.
INTELLIGENCE
Orangutans are among the most intelligent primates. Experiments suggest they can figure out some invisible displacement problems with a representational strategy. In addition, Zoo Atlanta has a touch-screen computer where their two Sumatran orangutans play games. Scientists hope the data they collect will help researchers learn about socialising patterns, such as whether the apes learn behaviours through trial and error or by mimicry, and point to new conservation strategies.
A 2008 study of two orangutans at the Leipzig Zoo showed orangutans can use "calculated reciprocity", which involves weighing the costs and benefits of gift exchanges and keeping track of these over time. Orangutans are the first nonhuman species documented to do so. Orangutans are very technically adept nest builders, making a new nest each evening in only in 5 to 6 minutes and choosing branches which they know can support their body weight.
TOOL USE AND CULTURE
Tool use in orangutans was observed by primatologist Birutė Galdikas in ex-captive populations. In addition, evidence of sophisticated tool manufacture and use in the wild was reported from a population of orangutans in Suaq Balimbing (Pongo abelii) in 1996. These orangutans developed a tool kit for use in foraging that consisted of both insect-extraction tools for use in the hollows of trees and seed-extraction tools for harvesting seeds from hard-husked fruit. The orangutans adjusted their tools according to the nature of the task at hand, and preference was given to oral tool use. This preference was also found in an experimental study of captive orangutans (P. pygmaeus).
Primatologist Carel P. van Schaik and biological anthropologist Cheryl D. Knott further investigated tool use in different wild orangutan populations. They compared geographic variations in tool use related to the processing of Neesia fruit. The orangutans of Suaq Balimbing (P. abelii) were found to be avid users of insect and seed-extraction tools when compared to other wild orangutans. The scientists suggested these differences are cultural. The orangutans at Suaq Balimbing live in dense groups and are socially tolerant; this creates good conditions for social transmission. Further evidence that highly social orangutans are more likely to exhibit cultural behaviours came from a study of leaf-carrying behaviours of ex-captive orangutans that were being rehabilitated on the island of Kaja in Borneo.
Wild orangutans (P. pygmaeus wurmbii) in Tuanan, Borneo, were reported to use tools in acoustic communication. They use leaves to amplify the kiss squeak sounds they produce. The apes may employ this method of amplification to deceive the listener into believing they are larger animals.
In 2003, researchers from six different orangutan field sites who used the same behavioural coding scheme compared the behaviours of the animals from the different sites. They found the different orangutan populations behaved differently. The evidence suggested the differences were cultural: first, the extent of the differences increased with distance, suggesting cultural diffusion was occurring, and second, the size of the orangutans' cultural repertoire increased according to the amount of social contact present within the group. Social contact facilitates cultural transmission.
POSSIBLE LINGUISTIC CAPABILITIES
A study of orangutan symbolic capability was conducted from 1973 to 1975 by zoologist Gary L. Shapiro with Aazk, a juvenile female orangutan at the Fresno City Zoo (now Chaffee Zoo) in Fresno, California. The study employed the techniques of psychologist David Premack, who used plastic tokens to teach linguistic skills to the chimpanzee, Sarah. Shapiro continued to examine the linguistic and learning abilities of ex-captive orangutans in Tanjung Puting National Park, in Indonesian Borneo, between 1978 and 1980.
During that time, Shapiro instructed ex-captive orangutans in the acquisition and use of signs following the techniques of psychologists R. Allen Gardner and Beatrix Gardner, who taught the chimpanzee, Washoe, in the late 1960s. In the only signing study ever conducted in a great ape's natural environment, Shapiro home-reared Princess, a juvenile female, which learned nearly 40 signs (according to the criteria of sign acquisition used by psychologist Francine Patterson with Koko, the gorilla) and trained Rinnie, a free-ranging adult female orangutan, which learned nearly 30 signs over a two-year period. For his dissertation study, Shapiro examined the factors influencing sign learning by four juvenile orangutans over a 15-month period.
ORANGUTANS AND HUMANS
Orangutans were known to the native people of Sumatra and Borneo for millennia. While some communities hunted them for food and decoration, others placed taboos on such practices. In central Borneo, some traditional folk beliefs consider it bad luck to look in the face of an orangutan. Some folk tales involve orangutans mating with and kidnapping humans. There are even stories of hunters being seduced by female orangutans.
Europeans became aware of the existence of the orangutan possibly as early as the 17th century. European explorers in Borneo hunted them extensively during the 19th century. The first accurate description of orangutans was given by Dutch anatomist Petrus Camper, who observed the animals and dissected some specimens.
Little was known about their behaviour until the field studies of Birutė Galdikas, who became a leading authority on the apes. When she arrived in Borneo, Galdikas settled into a primitive bark and thatch hut, at a site she dubbed Camp Leakey, near the edge of the Java Sea. Despite numerous hardships, she remained there for over 30 years and became an outspoken advocate for orangutans and the preservation of their rainforest habitat, which is rapidly being devastated by loggers, palm oil plantations, gold miners, and unnatural forest fires.
Galdikas's conservation efforts have extended well beyond advocacy, largely focusing on rehabilitation of the many orphaned orangutans turned over to her for care. Galdikas is considered to be one of Leakey's Angels, along with Jane Goodall and Dian Fossey. According to the World Wildlife Fund, half of the habitat of the Bornean orangutan has been lost since 1994.
A persistent folktale on Sumatra and Borneo and in popular culture, is that male orangutans display sexual attraction to human women, and may even forcibly copulate with them. The only serious, but anecdotal, report of such an incident taking place, is primatologist Birutė Galdikas' report that her cook was sexually assaulted by a male orangutan. This orangutan, though, was raised in captivity and may have suffered from a skewed species identity, and forced copulation is a standard mating strategy for low-ranking male orangutans.
A female orangutan was rescued from a village brothel in Kareng Pangi village, Central Kalimantan, in 2003. The orangutan was shaved and chained for sexual purposes. Since being freed, the orangutan, named Pony, has been living with the Borneo Orangutan Survival Foundation. She has been re-socialised to live with other orang-utans.
LEGAL STATUS
In December 2014, Argentina became the first country to recognize a non-human primate as having legal rights when it ruled that an orangutan named Sandra at the Buenos Aires Zoo must be moved to a sanctuary in Brazil in order to provide her "partial or controlled freedom". Although animal rights groups interpreted the ruling as applicable to all species in captivity, legal specialists considered the ruling only applicable to hominid apes due to their genetic similarities to humans.
CONSERVATION
CONSERVATION STATUS
The Sumatran and Bornean species are both critically endangered according to the IUCN Red List of mammals, and both are listed on Appendix I of CITES.
The Bornean orangutan population declined by 60% in the past 60 years and is projected to decline by 82% over 75 years. Its range has become patchy throughout Borneo, being largely extirpated from various parts of the island, including the southeast. The largest remaining population is found in the forest around the Sabangau River, but this environment is at risk.
Sumatran orangutan populations declined by 80% in 75 years. This species is now found only in the northern part of Sumatra, with most of the population inhabiting the Leuser Ecosystem. In late March 2012, some of the last Sumatran orangutans in northern Sumatra were reported to be threatened with approaching forest fires and might be wiped out entirely within a matter of weeks.
Estimates between 2000 and 2003 found 7,300 Sumatran orangutans and between 45,000 and 69,000 Bornean orangutans remain in the wild. A 2007 study by the Government of Indonesia noted a total wild population of 61,234 orangutans, 54,567 of which were found on the island of Borneo in 2004.
During the early 2000s, orangutan habitat has decreased rapidly due to logging and forest fires, as well as fragmentation by roads. A major factor in that period of time has been the conversion of vast areas of tropical forest to palm oil plantations in response to international demand. Palm oil is used for cooking, cosmetics, mechanics, and biodiesel. Hunting is also a major problem as is the illegal pet trade.
Orangutans may be killed for the bushmeat trade, crop protection, or for use for traditional medicine. Orangutan bones are secretly traded in souvenir shops in several cities in Kalimantan, Indonesia. Mother orangutans are killed so their infants can be sold as pets, and many of these infants die without the help of their mother. Since 2004, several pet orangutans were confiscated by local authorities and sent to rehabilitation centres.
CONSERVATION CENTRES AND ORGANISATIONS
A number of organisations are working for the rescue, rehabilitation and reintroduction of orangutans. The largest of these is the Borneo Orangutan Survival Foundation, founded by conservationist Willie Smits. It is audited by a multinational auditor company and operates a number of large projects, such as the Nyaru Menteng Rehabilitation Program founded by conservationist Lone Drøscher Nielsen.
Other major conservation centres in Indonesia include those at Tanjung Puting National Park and Sebangau National Park in Central Kalimantan, Kutai in East Kalimantan, Gunung Palung National Park in West Kalimantan, and Bukit Lawang in the Gunung Leuser National Park on the border of Aceh and North Sumatra. In Malaysia, conservation areas include Semenggoh Wildlife Centre in Sarawak and Matang Wildlife Centre also in Sarawak, and the Sepilok Orang Utan Sanctuary near Sandakan in Sabah. Major conservation centres that are headquartered outside of the orangutan's home countries; include Frankfurt Zoological Society, Orangutan Foundation International, which was founded by Birutė Galdikas, and the Australian Orangutan Project.
Conservation organisations such as Orangutan Land Trust work with the palm oil industry to improve sustainability and encourages the industry to establish conservation areas for orangutans. It works to bring different stakeholders together to achieve conservation of the species and its habitat.
WIKIPEDIA
The cat (Felis catus), commonly referred to as the domestic cat or house cat, is the only domesticated species in the family Felidae. Recent advances in archaeology and genetics have shown that the domestication of the cat occurred in the Near East around 7500 BC. It is commonly kept as a house pet and farm cat, but also ranges freely as a feral cat avoiding human contact. It is valued by humans for companionship and its ability to kill vermin. Because of its retractable claws it is adapted to killing small prey like mice and rats. It has a strong flexible body, quick reflexes, sharp teeth, and its night vision and sense of smell are well developed. It is a social species, but a solitary hunter and a crepuscular predator. Cat communication includes vocalizations like meowing, purring, trilling, hissing, growling, and grunting as well as cat body language. It can hear sounds too faint or too high in frequency for human ears, such as those made by small mammals. It also secretes and perceives pheromones.
Female domestic cats can have kittens from spring to late autumn in temperate zones and throughout the year in equatorial regions, with litter sizes often ranging from two to five kittens. Domestic cats are bred and shown at events as registered pedigreed cats, a hobby known as cat fancy. Animal population control of cats may be achieved by spaying and neutering, but their proliferation and the abandonment of pets has resulted in large numbers of feral cats worldwide, contributing to the extinction of bird, mammal and reptile species.
As of 2017, the domestic cat was the second most popular pet in the United States, with 95.6 million cats owned and around 42 million households owning at least one cat. In the United Kingdom, 26% of adults have a cat, with an estimated population of 10.9 million pet cats as of 2020. As of 2021, there were an estimated 220 million owned and 480 million stray cats in the world.
Etymology and naming
The origin of the English word cat, Old English catt, is thought to be the Late Latin word cattus, which was first used at the beginning of the 6th century. The Late Latin word may be derived from an unidentified African language. The Nubian word kaddîska 'wildcat' and Nobiin kadīs are possible sources or cognates. The Nubian word may be a loan from Arabic قَطّ qaṭṭ ~ قِطّ qiṭṭ.
The forms might also have derived from an ancient Germanic word that was imported into Latin and then into Greek, Syriac, and Arabic. The word may be derived from Germanic and Northern European languages, and ultimately be borrowed from Uralic, cf. Northern Sámi gáđfi, 'female stoat', and Hungarian hölgy, 'lady, female stoat'; from Proto-Uralic *käďwä, 'female (of a furred animal)'.
The English puss, extended as pussy and pussycat, is attested from the 16th century and may have been introduced from Dutch poes or from Low German puuskatte, related to Swedish kattepus, or Norwegian pus, pusekatt. Similar forms exist in Lithuanian puižė and Irish puisín or puiscín. The etymology of this word is unknown, but it may have arisen from a sound used to attract a cat.
A male cat is called a tom or tomcat (or a gib, if neutered). A female is called a queen or a molly, if spayed, especially in a cat-breeding context. A juvenile cat is referred to as a kitten. In Early Modern English, the word kitten was interchangeable with the now-obsolete word catling.
A group of cats can be referred to as a clowder or a glaring.
Taxonomy
The scientific name Felis catus was proposed by Carl Linnaeus in 1758 for a domestic cat. Felis catus domesticus was proposed by Johann Christian Polycarp Erxleben in 1777. Felis daemon proposed by Konstantin Satunin in 1904 was a black cat from the Transcaucasus, later identified as a domestic cat.
In 2003, the International Commission on Zoological Nomenclature ruled that the domestic cat is a distinct species, namely Felis catus. In 2007, it was considered a subspecies, F. silvestris catus, of the European wildcat (F. silvestris) following results of phylogenetic research. In 2017, the IUCN Cat Classification Taskforce followed the recommendation of the ICZN in regarding the domestic cat as a distinct species, Felis catus.
Evolution
Main article: Cat evolution
The domestic cat is a member of the Felidae, a family that had a common ancestor about 10 to 15 million years ago. The evolutionary radiation of the Felidae began in Asia during the Miocene around 8.38 to 14.45 million years ago. Analysis of mitochondrial DNA of all Felidae species indicates a radiation at 6.46 to 16.76 million years ago. The genus Felis genetically diverged from other Felidae around 6 to 7 million years ago. Results of phylogenetic research shows that the wild members of this genus evolved through sympatric or parapatric speciation, whereas the domestic cat evolved through artificial selection. The domestic cat and its closest wild ancestor are diploid and both possess 38 chromosomes and roughly 20,000 genes.
Domestication
See also: Domestication of the cat and Cats in ancient Egypt
It was long thought that the domestication of the cat began in ancient Egypt, where cats were venerated from around 3100 BC, However, the earliest known indication for the taming of an African wildcat was excavated close by a human Neolithic grave in Shillourokambos, southern Cyprus, dating to about 7500–7200 BC. Since there is no evidence of native mammalian fauna on Cyprus, the inhabitants of this Neolithic village most likely brought the cat and other wild mammals to the island from the Middle Eastern mainland. Scientists therefore assume that African wildcats were attracted to early human settlements in the Fertile Crescent by rodents, in particular the house mouse (Mus musculus), and were tamed by Neolithic farmers. This mutual relationship between early farmers and tamed cats lasted thousands of years. As agricultural practices spread, so did tame and domesticated cats. Wildcats of Egypt contributed to the maternal gene pool of the domestic cat at a later time.
The earliest known evidence for the occurrence of the domestic cat in Greece dates to around 1200 BC. Greek, Phoenician, Carthaginian and Etruscan traders introduced domestic cats to southern Europe. During the Roman Empire they were introduced to Corsica and Sardinia before the beginning of the 1st millennium. By the 5th century BC, they were familiar animals around settlements in Magna Graecia and Etruria. By the end of the Western Roman Empire in the 5th century, the Egyptian domestic cat lineage had arrived in a Baltic Sea port in northern Germany.
The leopard cat (Prionailurus bengalensis) was tamed independently in China around 5500 BC. This line of partially domesticated cats leaves no trace in the domestic cat populations of today.
During domestication, cats have undergone only minor changes in anatomy and behavior, and they are still capable of surviving in the wild. Several natural behaviors and characteristics of wildcats may have pre-adapted them for domestication as pets. These traits include their small size, social nature, obvious body language, love of play, and high intelligence. Captive Leopardus cats may also display affectionate behavior toward humans but were not domesticated. House cats often mate with feral cats. Hybridisation between domestic and other Felinae species is also possible, producing hybrids such as the Kellas cat in Scotland.
Development of cat breeds started in the mid 19th century. An analysis of the domestic cat genome revealed that the ancestral wildcat genome was significantly altered in the process of domestication, as specific mutations were selected to develop cat breeds. Most breeds are founded on random-bred domestic cats. Genetic diversity of these breeds varies between regions, and is lowest in purebred populations, which show more than 20 deleterious genetic disorders.
Characteristics
Main article: Cat anatomy
Size
The domestic cat has a smaller skull and shorter bones than the European wildcat. It averages about 46 cm (18 in) in head-to-body length and 23–25 cm (9.1–9.8 in) in height, with about 30 cm (12 in) long tails. Males are larger than females. Adult domestic cats typically weigh 4–5 kg (8.8–11.0 lb).
Skeleton
Cats have seven cervical vertebrae (as do most mammals); 13 thoracic vertebrae (humans have 12); seven lumbar vertebrae (humans have five); three sacral vertebrae (as do most mammals, but humans have five); and a variable number of caudal vertebrae in the tail (humans have only three to five vestigial caudal vertebrae, fused into an internal coccyx). The extra lumbar and thoracic vertebrae account for the cat's spinal mobility and flexibility. Attached to the spine are 13 ribs, the shoulder, and the pelvis. Unlike human arms, cat forelimbs are attached to the shoulder by free-floating clavicle bones which allow them to pass their body through any space into which they can fit their head.
Skull
The cat skull is unusual among mammals in having very large eye sockets and a powerful specialized jaw. Within the jaw, cats have teeth adapted for killing prey and tearing meat. When it overpowers its prey, a cat delivers a lethal neck bite with its two long canine teeth, inserting them between two of the prey's vertebrae and severing its spinal cord, causing irreversible paralysis and death. Compared to other felines, domestic cats have narrowly spaced canine teeth relative to the size of their jaw, which is an adaptation to their preferred prey of small rodents, which have small vertebrae.
The premolar and first molar together compose the carnassial pair on each side of the mouth, which efficiently shears meat into small pieces, like a pair of scissors. These are vital in feeding, since cats' small molars cannot chew food effectively, and cats are largely incapable of mastication.: Cats tend to have better teeth than most humans, with decay generally less likely because of a thicker protective layer of enamel, a less damaging saliva, less retention of food particles between teeth, and a diet mostly devoid of sugar. Nonetheless, they are subject to occasional tooth loss and infection.
Claws
Cats have protractible and retractable claws. In their normal, relaxed position, the claws are sheathed with the skin and fur around the paw's toe pads. This keeps the claws sharp by preventing wear from contact with the ground and allows for the silent stalking of prey. The claws on the forefeet are typically sharper than those on the hindfeet. Cats can voluntarily extend their claws on one or more paws. They may extend their claws in hunting or self-defense, climbing, kneading, or for extra traction on soft surfaces. Cats shed the outside layer of their claw sheaths when scratching rough surfaces.
Most cats have five claws on their front paws and four on their rear paws. The dewclaw is proximal to the other claws. More proximally is a protrusion which appears to be a sixth "finger". This special feature of the front paws on the inside of the wrists has no function in normal walking but is thought to be an antiskidding device used while jumping. Some cat breeds are prone to having extra digits ("polydactyly"). Polydactylous cats occur along North America's northeast coast and in Great Britain.
Ambulation
The cat is digitigrade. It walks on the toes, with the bones of the feet making up the lower part of the visible leg. Unlike most mammals, it uses a "pacing" gait and moves both legs on one side of the body before the legs on the other side. It registers directly by placing each hind paw close to the track of the corresponding fore paw, minimizing noise and visible tracks. This also provides sure footing for hind paws when navigating rough terrain. As it speeds up from walking to trotting, its gait changes to a "diagonal" gait: The diagonally opposite hind and fore legs move simultaneously.
Balance
Cats are generally fond of sitting in high places or perching. A higher place may serve as a concealed site from which to hunt; domestic cats strike prey by pouncing from a perch such as a tree branch. Another possible explanation is that height gives the cat a better observation point, allowing it to survey its territory. A cat falling from heights of up to 3 m (9.8 ft) can right itself and land on its paws.
During a fall from a high place, a cat reflexively twists its body and rights itself to land on its feet using its acute sense of balance and flexibility. This reflex is known as the cat righting reflex. A cat always rights itself in the same way during a fall, if it has enough time to do so, which is the case in falls of 90 cm (3.0 ft) or more. How cats are able to right themselves when falling has been investigated as the "falling cat problem".
Coats
Main article: Cat coat genetics
The cat family (Felidae) can pass down many colors and patterns to their offspring. The domestic cat genes MC1R and ASIP allow for the variety of color in coats. The feline ASIP gene consists of three coding exons. Three novel microsatellite markers linked to ASIP were isolated from a domestic cat BAC clone containing this gene and were used to perform linkage analysis in a pedigree of 89 domestic cats that segregated for melanism.[citation needed]
Senses
Main article: Cat senses
Vision
A cat's nictitating membrane shown as it blinks
Cats have excellent night vision and can see at only one-sixth the light level required for human vision. This is partly the result of cat eyes having a tapetum lucidum, which reflects any light that passes through the retina back into the eye, thereby increasing the eye's sensitivity to dim light. Large pupils are an adaptation to dim light. The domestic cat has slit pupils, which allow it to focus bright light without chromatic aberration. At low light, a cat's pupils expand to cover most of the exposed surface of its eyes. The domestic cat has rather poor color vision and only two types of cone cells, optimized for sensitivity to blue and yellowish green; its ability to distinguish between red and green is limited. A response to middle wavelengths from a system other than the rod cells might be due to a third type of cone. This appears to be an adaptation to low light levels rather than representing true trichromatic vision. Cats also have a nictitating membrane, allowing them to blink without hindering their vision.
Hearing
The domestic cat's hearing is most acute in the range of 500 Hz to 32 kHz. It can detect an extremely broad range of frequencies ranging from 55 Hz to 79 kHz, whereas humans can only detect frequencies between 20 Hz and 20 kHz. It can hear a range of 10.5 octaves, while humans and dogs can hear ranges of about 9 octaves. Its hearing sensitivity is enhanced by its large movable outer ears, the pinnae, which amplify sounds and help detect the location of a noise. It can detect ultrasound, which enables it to detect ultrasonic calls made by rodent prey. Recent research has shown that cats have socio-spatial cognitive abilities to create mental maps of owners' locations based on hearing owners' voices.
Smell
Cats have an acute sense of smell, due in part to their well-developed olfactory bulb and a large surface of olfactory mucosa, about 5.8 cm2 (0.90 in2) in area, which is about twice that of humans. Cats and many other animals have a Jacobson's organ in their mouths that is used in the behavioral process of flehmening. It allows them to sense certain aromas in a way that humans cannot. Cats are sensitive to pheromones such as 3-mercapto-3-methylbutan-1-ol, which they use to communicate through urine spraying and marking with scent glands. Many cats also respond strongly to plants that contain nepetalactone, especially catnip, as they can detect that substance at less than one part per billion. About 70–80% of cats are affected by nepetalactone. This response is also produced by other plants, such as silver vine (Actinidia polygama) and the herb valerian; it may be caused by the smell of these plants mimicking a pheromone and stimulating cats' social or sexual behaviors.
Taste
Cats have relatively few taste buds compared to humans (470 or so versus more than 9,000 on the human tongue). Domestic and wild cats share a taste receptor gene mutation that keeps their sweet taste buds from binding to sugary molecules, leaving them with no ability to taste sweetness. They, however, possess taste bud receptors specialized for acids, amino acids like protein, and bitter tastes. Their taste buds possess the receptors needed to detect umami. However, these receptors contain molecular changes that make the cat taste of umami different from that of humans. In humans, they detect the amino acids of glutamic acid and aspartic acid, but in cats they instead detect nucleotides, in this case inosine monophosphate and l-Histidine. These nucleotides are particularly enriched in tuna. This has been argued is why cats find tuna so palatable: as put by researchers into cat taste, "the specific combination of the high IMP and free l-Histidine contents of tuna" .. "produces a strong umami taste synergy that is highly preferred by cats". One of the researchers involved in this research has further claimed, "I think umami is as important for cats as sweet is for humans".[87]
Cats also have a distinct temperature preference for their food, preferring food with a temperature around 38 °C (100 °F) which is similar to that of a fresh kill; some cats reject cold food (which would signal to the cat that the "prey" item is long dead and therefore possibly toxic or decomposing).
Whiskers
To aid with navigation and sensation, cats have dozens of movable whiskers (vibrissae) over their body, especially their faces. These provide information on the width of gaps and on the location of objects in the dark, both by touching objects directly and by sensing air currents; they also trigger protective blink reflexes to protect the eyes from damage.: 47
Behavior
See also: Cat behavior
Outdoor cats are active both day and night, although they tend to be slightly more active at night.[88] Domestic cats spend the majority of their time in the vicinity of their homes but can range many hundreds of meters from this central point. They establish territories that vary considerably in size, in one study ranging 7–28 ha (17–69 acres). The timing of cats' activity is quite flexible and varied but being low-light predators, they are generally crepuscular, which means they tend to be more active near dawn and dusk. However, house cats' behavior is also influenced by human activity and they may adapt to their owners' sleeping patterns to some extent.
Cats conserve energy by sleeping more than most animals, especially as they grow older. The daily duration of sleep varies, usually between 12 and 16 hours, with 13 and 14 being the average. Some cats can sleep as much as 20 hours. The term "cat nap" for a short rest refers to the cat's tendency to fall asleep (lightly) for a brief period. While asleep, cats experience short periods of rapid eye movement sleep often accompanied by muscle twitches, which suggests they are dreaming.
Sociability
The social behavior of the domestic cat ranges from widely dispersed individuals to feral cat colonies that gather around a food source, based on groups of co-operating females. Within such groups, one cat is usually dominant over the others. Each cat in a colony holds a distinct territory, with sexually active males having the largest territories, which are about 10 times larger than those of female cats and may overlap with several females' territories. These territories are marked by urine spraying, by rubbing objects at head height with secretions from facial glands, and by defecation. Between these territories are neutral areas where cats watch and greet one another without territorial conflicts. Outside these neutral areas, territory holders usually chase away stranger cats, at first by staring, hissing, and growling and, if that does not work, by short but noisy and violent attacks. Despite this colonial organization, cats do not have a social survival strategy or a herd behavior, and always hunt alone.
Life in proximity to humans and other domestic animals has led to a symbiotic social adaptation in cats, and cats may express great affection toward humans or other animals. Ethologically, a cat's human keeper functions as if a mother surrogate. Adult cats live their lives in a kind of extended kittenhood, a form of behavioral neoteny. Their high-pitched sounds may mimic the cries of a hungry human infant, making them particularly difficult for humans to ignore. Some pet cats are poorly socialized. In particular, older cats show aggressiveness toward newly arrived kittens, which include biting and scratching; this type of behavior is known as feline asocial aggression.
Redirected aggression is a common form of aggression which can occur in multiple cat households. In redirected aggression there is usually something that agitates the cat: this could be a sight, sound, or another source of stimuli which causes a heightened level of anxiety or arousal. If the cat cannot attack the stimuli, it may direct anger elsewhere by attacking or directing aggression to the nearest cat, dog, human or other being.
Domestic cats' scent rubbing behavior toward humans or other cats is thought to be a feline means for social bonding.
Communication
Main article: Cat communication
Domestic cats use many vocalizations for communication, including purring, trilling, hissing, growling/snarling, grunting, and several different forms of meowing. Their body language, including position of ears and tail, relaxation of the whole body, and kneading of the paws, are all indicators of mood. The tail and ears are particularly important social signal mechanisms in cats. A raised tail indicates a friendly greeting, and flattened ears indicate hostility. Tail-raising also indicates the cat's position in the group's social hierarchy, with dominant individuals raising their tails less often than subordinate ones. Feral cats are generally silent.: 208 Nose-to-nose touching is also a common greeting and may be followed by social grooming, which is solicited by one of the cats raising and tilting its head.
Purring may have developed as an evolutionary advantage as a signaling mechanism of reassurance between mother cats and nursing kittens, who are thought to use it as a care-soliciting signal. Post-nursing cats also often purr as a sign of contentment: when being petted, becoming relaxed, or eating. Even though purring is popularly interpreted as indicative of pleasure, it has been recorded in a wide variety of circumstances, most of which involve physical contact between the cat and another, presumably trusted individual. Some cats have been observed to purr continuously when chronically ill or in apparent pain.
The exact mechanism by which cats purr has long been elusive, but it has been proposed that purring is generated via a series of sudden build-ups and releases of pressure as the glottis is opened and closed, which causes the vocal folds to separate forcefully. The laryngeal muscles in control of the glottis are thought to be driven by a neural oscillator which generates a cycle of contraction and release every 30–40 milliseconds (giving a frequency of 33 to 25 Hz).
Domestic cats observed in a rescue facility have total of 276 distinct facial expressions based on 26 different facial movements; each facial expression corresponds to different social functions that are likely influenced by domestication.
Grooming
Cats are known for spending considerable amounts of time licking their coats to keep them clean. The cat's tongue has backward-facing spines about 500 μm long, which are called papillae. These contain keratin which makes them rigid so the papillae act like a hairbrush. Some cats, particularly longhaired cats, occasionally regurgitate hairballs of fur that have collected in their stomachs from grooming. These clumps of fur are usually sausage-shaped and about 2–3 cm (0.79–1.18 in) long. Hairballs can be prevented with remedies that ease elimination of the hair through the gut, as well as regular grooming of the coat with a comb or stiff brush.
Fighting
Among domestic cats, males are more likely to fight than females. Among feral cats, the most common reason for cat fighting is competition between two males to mate with a female. In such cases, most fights are won by the heavier male. Another common reason for fighting in domestic cats is the difficulty of establishing territories within a small home. Female cats also fight over territory or to defend their kittens. Neutering will decrease or eliminate this behavior in many cases, suggesting that the behavior is linked to sex hormones.
When cats become aggressive, they try to make themselves appear larger and more threatening by raising their fur, arching their backs, turning sideways and hissing or spitting. Often, the ears are pointed down and back to avoid damage to the inner ear and potentially listen for any changes behind them while focused forward. Cats may also vocalize loudly and bare their teeth in an effort to further intimidate their opponents. Fights usually consist of grappling and delivering powerful slaps to the face and body with the forepaws as well as bites. Cats also throw themselves to the ground in a defensive posture to rake their opponent's belly with their powerful hind legs.
Serious damage is rare, as the fights are usually short in duration, with the loser running away with little more than a few scratches to the face and ears. Fights for mating rights are typically more severe and injuries may include deep puncture wounds and lacerations. Normally, serious injuries from fighting are limited to infections of scratches and bites, though these can occasionally kill cats if untreated. In addition, bites are probably the main route of transmission of feline immunodeficiency virus. Sexually active males are usually involved in many fights during their lives, and often have decidedly battered faces with obvious scars and cuts to their ears and nose. Cats are willing to threaten animals larger than them to defend their territory, such as dogs and foxes.
Hunting and feeding
See also: Cat food
The shape and structure of cats' cheeks is insufficient to allow them to take in liquids using suction. Therefore, when drinking they lap with the tongue to draw liquid upward into their mouths. Lapping at a rate of four times a second, the cat touches the smooth tip of its tongue to the surface of the water, and quickly retracts it like a corkscrew, drawing water upward.
Feral cats and free-fed house cats consume several small meals in a day. The frequency and size of meals varies between individuals. They select food based on its temperature, smell and texture; they dislike chilled foods and respond most strongly to moist foods rich in amino acids, which are similar to meat. Cats reject novel flavors (a response termed neophobia) and learn quickly to avoid foods that have tasted unpleasant in the past. It is also a common misconception that cats like milk/cream, as they tend to avoid sweet food and milk. Most adult cats are lactose intolerant; the sugar in milk is not easily digested and may cause soft stools or diarrhea. Some also develop odd eating habits and like to eat or chew on things like wool, plastic, cables, paper, string, aluminum foil, or even coal. This condition, pica, can threaten their health, depending on the amount and toxicity of the items eaten.
Cats hunt small prey, primarily birds and rodents, and are often used as a form of pest control. Other common small creatures such as lizards and snakes may also become prey. Cats use two hunting strategies, either stalking prey actively, or waiting in ambush until an animal comes close enough to be captured. The strategy used depends on the prey species in the area, with cats waiting in ambush outside burrows, but tending to actively stalk birds.: 153 Domestic cats are a major predator of wildlife in the United States, killing an estimated 1.3 to 4.0 billion birds and 6.3 to 22.3 billion mammals annually.
Certain species appear more susceptible than others; in one English village, for example, 30% of house sparrow mortality was linked to the domestic cat. In the recovery of ringed robins (Erithacus rubecula) and dunnocks (Prunella modularis) in Britain, 31% of deaths were a result of cat predation. In parts of North America, the presence of larger carnivores such as coyotes which prey on cats and other small predators reduces the effect of predation by cats and other small predators such as opossums and raccoons on bird numbers and variety.
Perhaps the best-known element of cats' hunting behavior, which is commonly misunderstood and often appalls cat owners because it looks like torture, is that cats often appear to "play" with prey by releasing and recapturing it. This cat and mouse behavior is due to an instinctive imperative to ensure that the prey is weak enough to be killed without endangering the cat.
Another poorly understood element of cat hunting behavior is the presentation of prey to human guardians. One explanation is that cats adopt humans into their social group and share excess kill with others in the group according to the dominance hierarchy, in which humans are reacted to as if they are at or near the top. Another explanation is that they attempt to teach their guardians to hunt or to help their human as if feeding "an elderly cat, or an inept kitten". This hypothesis is inconsistent with the fact that male cats also bring home prey, despite males having negligible involvement in raising kittens.:
Play
Main article: Cat play and toys
Domestic cats, especially young kittens, are known for their love of play. This behavior mimics hunting and is important in helping kittens learn to stalk, capture, and kill prey. Cats also engage in play fighting, with each other and with humans. This behavior may be a way for cats to practice the skills needed for real combat, and might also reduce any fear they associate with launching attacks on other animals.
Cats also tend to play with toys more when they are hungry. Owing to the close similarity between play and hunting, cats prefer to play with objects that resemble prey, such as small furry toys that move rapidly, but rapidly lose interest. They become habituated to a toy they have played with before. String is often used as a toy, but if it is eaten, it can become caught at the base of the cat's tongue and then move into the intestines, a medical emergency which can cause serious illness, even death. Owing to the risks posed by cats eating string, it is sometimes replaced with a laser pointer's dot, which cats may chase.
Reproduction
See also: Kitten
The cat secretes and perceives pheromones. Female cats, called queens, are polyestrous with several estrus cycles during a year, lasting usually 21 days. They are usually ready to mate between early February and August in northern temperate zones and throughout the year in equatorial regions.
Several males, called tomcats, are attracted to a female in heat. They fight over her, and the victor wins the right to mate. At first, the female rejects the male, but eventually, the female allows the male to mate. The female utters a loud yowl as the male pulls out of her because a male cat's penis has a band of about 120–150 backward-pointing penile spines, which are about 1 mm (0.039 in) long; upon withdrawal of the penis, the spines may provide the female with increased sexual stimulation, which acts to induce ovulation.
After mating, the female cleans her vulva thoroughly. If a male attempts to mate with her at this point, the female attacks him. After about 20 to 30 minutes, once the female is finished grooming, the cycle will repeat. Because ovulation is not always triggered by a single mating, females may not be impregnated by the first male with which they mate. Furthermore, cats are superfecund; that is, a female may mate with more than one male when she is in heat, with the result that different kittens in a litter may have different fathers.
The morula forms 124 hours after conception. At 148 hours, early blastocysts form. At 10–12 days, implantation occurs. The gestation of queens lasts between 64 and 67 days, with an average of 65 days.
Data on the reproductive capacity of more than 2,300 free-ranging queens were collected during a study between May 1998 and October 2000. They had one to six kittens per litter, with an average of three kittens. They produced a mean of 1.4 litters per year, but a maximum of three litters in a year. Of 169 kittens, 127 died before they were six months old due to a trauma caused in most cases by dog attacks and road accidents. The first litter is usually smaller than subsequent litters. Kittens are weaned between six and seven weeks of age. Queens normally reach sexual maturity at 5–10 months, and males at 5–7 months. This varies depending on breed. Kittens reach puberty at the age of 9–10 months.
Cats are ready to go to new homes at about 12 weeks of age, when they are ready to leave their mother. They can be surgically sterilized (spayed or castrated) as early as seven weeks to limit unwanted reproduction. This surgery also prevents undesirable sex-related behavior, such as aggression, territory marking (spraying urine) in males and yowling (calling) in females. Traditionally, this surgery was performed at around six to nine months of age, but it is increasingly being performed before puberty, at about three to six months. In the United States, about 80% of household cats are neutered.
Lifespan and health
Main articles: Cat health and Aging in cats
The average lifespan of pet cats has risen in recent decades. In the early 1980s, it was about seven years,: 33 rising to 9.4 years in 1995: 33 and an average of about 13 years as of 2014 and 2023. Some cats have been reported as surviving into their 30s, with the oldest known cat dying at a verified age of 38.
Neutering increases life expectancy: one study found castrated male cats live twice as long as intact males, while spayed female cats live 62% longer than intact females.: 35 Having a cat neutered confers health benefits, because castrated males cannot develop testicular cancer, spayed females cannot develop uterine or ovarian cancer, and both have a reduced risk of mammary cancer.
Disease
Main article: List of feline diseases
About 250 heritable genetic disorders have been identified in cats, many similar to human inborn errors of metabolism. The high level of similarity among the metabolism of mammals allows many of these feline diseases to be diagnosed using genetic tests that were originally developed for use in humans, as well as the use of cats as animal models in the study of the human diseases. Diseases affecting domestic cats include acute infections, parasitic infestations, injuries, and chronic diseases such as kidney disease, thyroid disease, and arthritis. Vaccinations are available for many infectious diseases, as are treatments to eliminate parasites such as worms, ticks, and fleas.
Ecology
Habitats
The domestic cat is a cosmopolitan species and occurs across much of the world. It is adaptable and now present on all continents except Antarctica, and on 118 of the 131 main groups of islands, even on the isolated Kerguelen Islands. Due to its ability to thrive in almost any terrestrial habitat, it is among the world's most invasive species. It lives on small islands with no human inhabitants. Feral cats can live in forests, grasslands, tundra, coastal areas, agricultural land, scrublands, urban areas, and wetlands.
The unwantedness that leads to the domestic cat being treated as an invasive species is twofold. On one hand, as it is little altered from the wildcat, it can readily interbreed with the wildcat. This hybridization poses a danger to the genetic distinctiveness of some wildcat populations, particularly in Scotland and Hungary, possibly also the Iberian Peninsula, and where protected natural areas are close to human-dominated landscapes, such as Kruger National Park in South Africa. However, its introduction to places where no native felines are present also contributes to the decline of native species.
Ferality
Main article: Feral cat
Feral cats are domestic cats that were born in or have reverted to a wild state. They are unfamiliar with and wary of humans and roam freely in urban and rural areas. The numbers of feral cats is not known, but estimates of the United States feral population range from 25 to 60 million. Feral cats may live alone, but most are found in large colonies, which occupy a specific territory and are usually associated with a source of food. Famous feral cat colonies are found in Rome around the Colosseum and Forum Romanum, with cats at some of these sites being fed and given medical attention by volunteers.
Public attitudes toward feral cats vary widely, from seeing them as free-ranging pets to regarding them as vermin.
Some feral cats can be successfully socialized and 're-tamed' for adoption; young cats, especially kittens and cats that have had prior experience and contact with humans are the most receptive to these efforts.
Impact on wildlife
Main article: Cat predation on wildlife
On islands, birds can contribute as much as 60% of a cat's diet. In nearly all cases, the cat cannot be identified as the sole cause for reducing the numbers of island birds, and in some instances, eradication of cats has caused a "mesopredator release" effect; where the suppression of top carnivores creates an abundance of smaller predators that cause a severe decline in their shared prey. Domestic cats are a contributing factor to the decline of many species, a factor that has ultimately led, in some cases, to extinction. The South Island piopio, Chatham rail, and the New Zealand merganser are a few from a long list, with the most extreme case being the flightless Lyall's wren, which was driven to extinction only a few years after its discovery. One feral cat in New Zealand killed 102 New Zealand lesser short-tailed bats in seven days. In the US, feral and free-ranging domestic cats kill an estimated 6.3 – 22.3 billion mammals annually.
In Australia, the impact of cats on mammal populations is even greater than the impact of habitat loss. More than one million reptiles are killed by feral cats each day, representing 258 species. Cats have contributed to the extinction of the Navassa curly-tailed lizard and Chioninia coctei.
Interaction with humans
Main article: Human interaction with cats
Cats are common pets throughout the world, and their worldwide population as of 2007 exceeded 500 million. As of 2017, the domestic cat was the second most popular pet in the United States, with 95.6 million cats owned and around 42 million households owning at least one cat. In the United Kingdom, 26% of adults have a cat, with an estimated population of 10.9 million pet cats as of 2020. As of 2021, there were an estimated 220 million owned and 480 million stray cats in the world.
Cats have been used for millennia to control rodents, notably around grain stores and aboard ships, and both uses extend to the present day.
As well as being kept as pets, cats are also used in the international fur trade and leather industries for making coats, hats, blankets, stuffed toys, shoes, gloves, and musical instruments. About 24 cats are needed to make a cat-fur coat. This use has been outlawed in the United States since 2000 and in the European Union (as well as the United Kingdom) since 2007.
Cat pelts have been used for superstitious purposes as part of the practice of witchcraft, and are still made into blankets in Switzerland as traditional medicine thought to cure rheumatism.
A few attempts to build a cat census have been made over the years, both through associations or national and international organizations (such as that of the Canadian Federation of Humane Societies) and over the Internet, but such a task does not seem simple to achieve. General estimates for the global population of domestic cats range widely from anywhere between 200 million to 600 million. Walter Chandoha made his career photographing cats after his 1949 images of Loco, an especially charming stray taken in, were published around the world. He is reported to have photographed 90,000 cats during his career and maintained an archive of 225,000 images that he drew from for publications during his lifetime.
Shows
Main article: Cat show
A cat show is a judged event in which the owners of cats compete to win titles in various cat-registering organizations by entering their cats to be judged after a breed standard. It is often required that a cat must be healthy and vaccinated in order to participate in a cat show. Both pedigreed and non-purebred companion ("moggy") cats are admissible, although the rules differ depending on the organization. Competing cats are compared to the applicable breed standard, and assessed for temperament.
Infection
Main article: Feline zoonosis
Cats can be infected or infested with viruses, bacteria, fungus, protozoans, arthropods or worms that can transmit diseases to humans. In some cases, the cat exhibits no symptoms of the disease. The same disease can then become evident in a human. The likelihood that a person will become diseased depends on the age and immune status of the person. Humans who have cats living in their home or in close association are more likely to become infected. Others might also acquire infections from cat feces and parasites exiting the cat's body. Some of the infections of most concern include salmonella, cat-scratch disease and toxoplasmosis.
History and mythology
Main articles: Cultural depictions of cats and Cats in ancient Egypt
In ancient Egypt, cats were worshipped, and the goddess Bastet often depicted in cat form, sometimes taking on the war-like aspect of a lioness. The Greek historian Herodotus reported that killing a cat was forbidden, and when a household cat died, the entire family mourned and shaved their eyebrows. Families took their dead cats to the sacred city of Bubastis, where they were embalmed and buried in sacred repositories. Herodotus expressed astonishment at the domestic cats in Egypt, because he had only ever seen wildcats.
Ancient Greeks and Romans kept weasels as pets, which were seen as the ideal rodent-killers. The earliest unmistakable evidence of the Greeks having domestic cats comes from two coins from Magna Graecia dating to the mid-fifth century BC showing Iokastos and Phalanthos, the legendary founders of Rhegion and Taras respectively, playing with their pet cats. The usual ancient Greek word for 'cat' was ailouros, meaning 'thing with the waving tail'. Cats are rarely mentioned in ancient Greek literature. Aristotle remarked in his History of Animals that "female cats are naturally lecherous." The Greeks later syncretized their own goddess Artemis with the Egyptian goddess Bastet, adopting Bastet's associations with cats and ascribing them to Artemis. In Ovid's Metamorphoses, when the deities flee to Egypt and take animal forms, the goddess Diana turns into a cat.
Cats eventually displaced weasels as the pest control of choice because they were more pleasant to have around the house and were more enthusiastic hunters of mice. During the Middle Ages, many of Artemis's associations with cats were grafted onto the Virgin Mary. Cats are often shown in icons of Annunciation and of the Holy Family and, according to Italian folklore, on the same night that Mary gave birth to Jesus, a cat in Bethlehem gave birth to a kitten. Domestic cats were spread throughout much of the rest of the world during the Age of Discovery, as ships' cats were carried on sailing ships to control shipboard rodents and as good-luck charms.
Several ancient religions believed cats are exalted souls, companions or guides for humans, that are all-knowing but mute so they cannot influence decisions made by humans. In Japan, the maneki neko cat is a symbol of good fortune. In Norse mythology, Freyja, the goddess of love, beauty, and fertility, is depicted as riding a chariot drawn by cats. In Jewish legend, the first cat was living in the house of the first man Adam as a pet that got rid of mice. The cat was once partnering with the first dog before the latter broke an oath they had made which resulted in enmity between the descendants of these two animals. It is also written that neither cats nor foxes are represented in the water, while every other animal has an incarnation species in the water. Although no species are sacred in Islam, cats are revered by Muslims. Some Western writers have stated Muhammad had a favorite cat, Muezza. He is reported to have loved cats so much, "he would do without his cloak rather than disturb one that was sleeping on it". The story has no origin in early Muslim writers, and seems to confuse a story of a later Sufi saint, Ahmed ar-Rifa'i, centuries after Muhammad. One of the companions of Muhammad was known as Abu Hurayrah ("father of the kitten"), in reference to his documented affection to cats.
Superstitions and rituals
Many cultures have negative superstitions about cats. An example would be the belief that encountering a black cat ("crossing one's path") leads to bad luck, or that cats are witches' familiars used to augment a witch's powers and skills. The killing of cats in Medieval Ypres, Belgium, is commemorated in the innocuous present-day Kattenstoet (cat parade). In mid-16th century France, cats would be burnt alive as a form of entertainment, particularly during midsummer festivals. According to Norman Davies, the assembled people "shrieked with laughter as the animals, howling with pain, were singed, roasted, and finally carbonized". The remaining ashes were sometimes taken back home by the people for good luck.
According to a myth in many cultures, cats have multiple lives. In many countries, they are believed to have nine lives, but in Italy, Germany, Greece, Brazil and some Spanish-speaking regions, they are said to have seven lives, while in Arabic traditions, the number of lives is six. An early mention of the myth can be found in John Heywood's The Proverbs of John Heywood (1546)
Husband, (quoth she), ye studie, be merrie now,
And even as ye thinke now, so come to yow.
Nay not so, (quoth he), for my thought to tell right,
I thinke how you lay groning, wife, all last night.
Husband, a groning horse and a groning wife
Never faile their master, (quoth she), for my life.
No wife, a woman hath nine lives like a cat.
The myth is attributed to the natural suppleness and swiftness cats exhibit to escape life-threatening situations. Also lending credence to this myth is the fact that falling cats often land on their feet, using an instinctive righting reflex to twist their bodies around. Nonetheless, cats can still be injured or killed by a high fall.