View allAll Photos Tagged testicles

Go to the Book with image in the Internet Archive

Title: United States Naval Medical Bulletin Vol. 3, Nos. 1-4, 1909

Creator: U.S. Navy. Bureau of Medicine and Surgery

Publisher:

Sponsor:

Contributor:

Date: 1909

Language: eng

  

<p class="MsoNormal" style="margin-bottom:.0001pt;line-height:normal;">Table of Contents</p>

 

<p class="MsoNormal" style="margin-bottom:.0001pt;line-height:normal;"> </p>

 

<p class="MsoNormal" style="margin-bottom:.0001pt;line-height:normal;">Number 1</p>

 

<p class="MsoNormal" style="margin-bottom:.0001pt;line-height:normal;"> </p>

 

<p class="MsoNormal" style="margin-bottom:.0001pt;line-height:normal;">Preface vii</p>

 

<p class="MsoNormal" style="margin-bottom:.0001pt;line-height:normal;">Special articles 1</p>

 

<p class="MsoNormal" style="margin-bottom:.0001pt;line-height:normal;">The artificial illumination of naval vessels (a study in naval

hygiene), by J. D. Gatewood 1</p>

 

<p class="MsoNormal" style="margin-bottom:.0001pt;line-height:normal;">A simple operation for hemorrhoids, by H. F. Hull 22</p>

 

<p class="MsoNormal" style="margin-bottom:.0001pt;line-height:normal;">Suggested devices 25</p>

 

<p class="MsoNormal" style="margin-bottom:.0001pt;line-height:normal;">A metal suspensory, by W. B. Grove 25</p>

 

<p class="MsoNormal" style="margin-bottom:.0001pt;line-height:normal;">A short and accurate method of calculating the age in years and months,

by E. M. Brown 25</p>

 

<p class="MsoNormal" style="margin-bottom:.0001pt;line-height:normal;">Card for index system to be used in preparing smooth quarterly form

"X" at recruiting stations, etc., by C. R. Keen 27</p>

 

<p class="MsoNormal" style="margin-bottom:.0001pt;line-height:normal;">Clinical notes 29</p>

 

<p class="MsoNormal" style="margin-bottom:.0001pt;line-height:normal;">A case of angina Ludovici, by W. S. Pugh 29</p>

 

<p class="MsoNormal" style="margin-bottom:.0001pt;line-height:normal;">A case of Vincent's angina, by G. F. Clark 31</p>

 

<p class="MsoNormal" style="margin-bottom:.0001pt;line-height:normal;">Rupture of the iris; two cases, by R. K. Riggs 32</p>

 

<p class="MsoNormal" style="margin-bottom:.0001pt;line-height:normal;">Wood alcohol poisoning; 13 cases, 3 deaths, by R. A. Baehmann 33</p>

 

<p class="MsoNormal" style="margin-bottom:.0001pt;line-height:normal;">A case of virulent chancroids, by D. C. Gather 36</p>

 

<p class="MsoNormal" style="margin-bottom:.0001pt;line-height:normal;">A case of septicemia successfully treated with Steam's streptolytic

serum by M. F. Gates . 39</p>

 

<p class="MsoNormal" style="margin-bottom:.0001pt;line-height:normal;">An unusual case of undescended testicle, by E. M. Brown 39</p>

 

<p class="MsoNormal" style="margin-bottom:.0001pt;line-height:normal;">Current comment 41</p>

 

<p class="MsoNormal" style="margin-bottom:.0001pt;line-height:normal;">United States Pharmacopeial Convention 41</p>

 

<p class="MsoNormal" style="margin-bottom:.0001pt;line-height:normal;">Concerning extracts or abstracts for publication 4l</p>

 

<p class="MsoNormal" style="margin-bottom:.0001pt;line-height:normal;">Measuring the height of recruits 43</p>

 

<p class="MsoNormal" style="margin-bottom:.0001pt;line-height:normal;">Suggestions for the study of heat exhaustion 44</p>

 

<p class="MsoNormal" style="margin-bottom:.0001pt;line-height:normal;">Perfected routine of dosage, etc., in the treatment of tuberculosis by

the administration of mercury, by B. L. Wright 46</p>

 

<p class="MsoNormal" style="margin-bottom:.0001pt;line-height:normal;">Has the chemical examination of water practical value to the military medical

officer? by P. '.T'. Waldner 47</p>

 

<p class="MsoNormal" style="margin-bottom:.0001pt;line-height:normal;">An aid in throat and laryngeal examinations, by E. M. Brown 50</p>

 

<p class="MsoNormal" style="margin-bottom:.0001pt;line-height:normal;">Progress in medical sciences 51</p>

 

<p class="MsoNormal" style="margin-bottom:.0001pt;line-height:normal;">Laboratory —An anatomical peculiarity noted in specimens of hook worm

from Culebra 51</p>

 

<p class="MsoNormal" style="margin-bottom:.0001pt;line-height:normal;">Preliminary note on the lesions of anchylostomiasis in the intestines of

dogs, by O. J. Mink 51</p>

 

<p class="MsoNormal" style="margin-bottom:.0001pt;line-height:normal;">Preliminary note on nematode found in the liver of a wild rat, by O.

J.Mink 52</p>

 

<p class="MsoNormal" style="margin-bottom:.0001pt;line-height:normal;">Chemistry and pharmacy — Note on the disintegration of tablets;

influence of benzoic acid and benzoates on digestion and health: address on the

clinical examination of urine, with especial reference to estimation of urea;

determination of pepsin by the edestin test, E. W. Brown and P. J. Waldner 52</p>

 

<p class="MsoNormal" style="margin-bottom:.0001pt;line-height:normal;">Surgery —Review of advances; the operative treatment of recent

fractures of the femoral shaft; the treatment of fractures by mobilization and massage;

has surgical treatment lessened mortality from appendicitis; when to operate

for appendicitis; diffuse septic peritonitis, due to appendicitis; local

anesthesia of a limb by venous transfusion after expulsion of blood; on

narcosis under an artificially restricted circulation; the correlation of

glands with internal secretion; improved technique for the detection of

tubercle bacilli in the urine; relief of the wounded during battle, H. C. Curl

and H. W. Smith 54</p>

 

<p class="MsoNormal" style="margin-bottom:.0001pt;line-height:normal;">Pathology and bacteriology —On the so-called fatty degeneration of the adrenals;

three cases of squamous celled carcinoma of the gall bladder; the practical

value of the demonstration of spirochaeta pallida in the early diagnosis of

syphilis; C. 8. Butler and O. J. Mink 65</p>

 

<p class="MsoNormal" style="margin-bottom:.0001pt;line-height:normal;">Medical zoology — Plague in ground squirrels (a review); the prevalence

and distribution of the animal parasites of man in the Philippine Islands, with

a consideration of their possible influence on the public'</p>

 

<p class="MsoNormal" style="margin-bottom:.0001pt;line-height:normal;">health; preliminary note on a protozoan in yaws; the intestinal protozoa

of man, R. C. Holcomb • 67</p>

 

<p class="MsoNormal" style="margin-bottom:.0001pt;line-height:normal;">Tropical medicine — Ankylostomiasis in the Tropics; bilharziasis among women

and girls in Egypt; a report of several cases with unusual symptoms caused by

contact with some unknown variety of jellyfish; the diagnosis of latent

malaria; haemolysins and antihaemolytic substances in the blood of malarial

patients, E. R. Stitt 73</p>

 

<p class="MsoNormal" style="margin-bottom:.0001pt;line-height:normal;">General medicine —The direct inspection of the gastric mucous membrane;

toxemia from the standpoint of perverted metabolism; a rapid method of

test-meal removal, lavage, and inflation; the therapeutics of diseases which

involve the internal secretions (mercury in the treatment of tuberculosis — its

mode of action —a warning); Flexner's serum in the treatment of epidemic

cerebrospinal meningitis; vascular crises; the curative influence of extracts

of leucocytes upon infections in animals, R. M. Kennedy 77</p>

 

<p class="MsoNormal" style="margin-bottom:.0001pt;line-height:normal;">Hygiene and sanitation —Koch's standpoint with reference to the

question of the relation between human and bovine tuberculosis; the prevention of

tuberculosis; tropical lands and white races; sanitary report of the operations

of the naval expeditionary corps (German) in southwest Africa and in east

Africa; growth and naval military service; a study in measurements of cadets at

the naval school; on growth in height of youths serving their time in the army;

the value of fencing as a sport from hygienic and ethical point* of view; on-

the significance of the ophthalmo-reaction for the army; hematuria caused by a

parasite akin to bilharzia; the complex nature of typhoid etiology and the role

played by animals and man in the spread of the typhoid group of diseases; amoebae

carriers, H. G. Beyer 90</p>

 

<p class="MsoNormal" style="margin-bottom:.0001pt;line-height:normal;">Reports and letters 195</p>

 

<p class="MsoNormal" style="margin-bottom:.0001pt;line-height:normal;">Annual meeting of the American Pharmaceutical Association, Alrik Hammar,

delegate 105</p>

 

<p class="MsoNormal" style="margin-bottom:.0001pt;line-height:normal;">Report of an epidemic of typhoid on the U.S.S. Maine, by M. S.

Elliott.<span>  </span>106</p>

 

<p class="MsoNormal" style="margin-bottom:.0001pt;line-height:normal;">Report of an epidemic of grippe on the U. S. S. Charleston, by M. F.

Gates. 109</p>

 

<p class="MsoNormal" style="margin-bottom:.0001pt;line-height:normal;"> </p>

 

<p class="MsoNormal" style="margin-bottom:.0001pt;line-height:normal;">Number 2</p>

 

<p class="MsoNormal" style="margin-bottom:.0001pt;line-height:normal;"> </p>

 

<p class="MsoNormal" style="margin-bottom:.0001pt;line-height:normal;">Preface vii</p>

 

<p class="MsoNormal" style="margin-bottom:.0001pt;line-height:normal;">Special articles 111</p>

 

<p class="MsoNormal" style="margin-bottom:.0001pt;line-height:normal;">The treatment of tuberculosis and the results observed during the year 1908

(at the United States Naval Hospital, Las Animas, Colo.), by B. L. Wright 111</p>

 

<p class="MsoNormal" style="margin-bottom:.0001pt;line-height:normal;">Laboratory studies and observations during the year 1908 (at the United

States Naval Hospital, Las Animas, Colo.), by A. B. Clifford 114</p>

 

<p class="MsoNormal" style="margin-bottom:.0001pt;line-height:normal;">Tonsillar hypertrophy; a menace to the service, by B. F. Jenness 120</p>

 

<p class="MsoNormal" style="margin-bottom:.0001pt;line-height:normal;">The ice bag in the treatment of typhoid fever, by G. Tucker Smith 122</p>

 

<p class="MsoNormal" style="margin-bottom:.0001pt;line-height:normal;">Treatment of typhoid fever by colon irrigations, by the late C. G.

Alderman 124</p>

 

<p class="MsoNormal" style="margin-bottom:.0001pt;line-height:normal;">Suggested devices 129</p>

 

<p class="MsoNormal" style="margin-bottom:.0001pt;line-height:normal;">Description of a pit incinerator furnace, by R. C. Holcomb 129</p>

 

<p class="MsoNormal" style="margin-bottom:.0001pt;line-height:normal;">Clinical notes 131</p>

 

<p class="MsoNormal" style="margin-bottom:.0001pt;line-height:normal;">Report of a case of malignant endocarditis, following chancroid, by I.

Franklin Cohn 131</p>

 

<p class="MsoNormal" style="margin-bottom:.0001pt;line-height:normal;">A case of multiple infected wounds from bear bite, by C. C. Grieve 132</p>

 

<p class="MsoNormal" style="margin-bottom:.0001pt;line-height:normal;">A case presenting successive liver abscesses, by H. C. Curl and H. W. Smith

134</p>

 

<p class="MsoNormal" style="margin-bottom:.0001pt;line-height:normal;">Cerebro-spinal fever, by J. G. Field 135</p>

 

<p class="MsoNormal" style="margin-bottom:.0001pt;line-height:normal;">Current comment 141</p>

 

<p class="MsoNormal" style="margin-bottom:.0001pt;line-height:normal;">Gangosa in Haiti 141</p>

 

<p class="MsoNormal" style="margin-bottom:.0001pt;line-height:normal;">Hookworm disease in recruits from the Southern States 141</p>

 

<p class="MsoNormal" style="margin-bottom:.0001pt;line-height:normal;">Care of ears and eyes in the Japanese navy<span>  </span><span> </span>142</p>

 

<p class="MsoNormal" style="margin-bottom:.0001pt;line-height:normal;">The question of ear protection in the British navy 142</p>

 

<p class="MsoNormal" style="margin-bottom:.0001pt;line-height:normal;">Report relative to a series of experiments conducted on board the U. S.

S. Ohio during target practice, with "Plasticine" for the protection

of the ear drums during heavy gun fire, by W. M. Garton 142</p>

 

<p class="MsoNormal" style="margin-bottom:.0001pt;line-height:normal;">Hygienic rules, with particular reference to venereal prophylaxis, in

the Austro-Hungarian navy 144</p>

 

<p class="MsoNormal" style="margin-bottom:.0001pt;line-height:normal;">Experiments with gonococcic vaccine, by W. M. Garton 145</p>

 

<p class="MsoNormal" style="margin-bottom:.0001pt;line-height:normal;">Thyroidal enlargement among applicants for enlistment in the Northwest,

by W. A. Angwin 147</p>

 

<p class="MsoNormal" style="margin-bottom:.0001pt;line-height:normal;">Progress in medical sciences 148</p>

 

<p class="MsoNormal" style="margin-bottom:.0001pt;line-height:normal;">Laboratory — Sterilization of catgut, by H. W. Smith 148</p>

 

<p class="MsoNormal" style="margin-bottom:.0001pt;line-height:normal;">Chemistry and pharmacy — Fluidglycerates, pharmaceutical and physiological

aspect; the importance and significance of the chemical examination of the

gastric contents after a test meal, with a new method for estimating the

ferment activity of the gastric contents; demonstrations of enzymes and

antienzymes; studies on the chemistry of anaphylaxis; the clinical value of

viscosity determination; the viscosity of the blood; the detection and

quantitative determination of B-oxybutyric acid in the urine; a new method for

the quantitative estimation of albumin in the urine; concerning the diagnostic

value of Cammidge crystals in pancreatic diseases, E. W. Brown and P. J.

Waldner 150</p>

 

<p class="MsoNormal" style="margin-bottom:.0001pt;line-height:normal;">Surgery — Review of advances; cerebral decompression; operative treatment

of acute gonorrheal epididymitis; appendectomy in diffuse septic peritonitis;

concerning technique of skin grafting; treatment of hypertrophy of the prostate

by injections of alien blood; the value of the Cammidge reaction in the diagnosis

of pancreatic disease; the Cammidge reaction in experimental pancreatitis; the

syphilis case sheet; the thymus in Basedow's disease; the effect of mammalian

pituitary on tetany after parathyreoidectomy, and upon the pupil; hemorrhage in

jaundice controlled by blood transfusion; on the haematogenic origin of

purulent nephritis through the staphylococcus; the snapping hip; three cases of

liver abscess treated by aspiration and injection of quinine, H. C. Curl and H.

\V. Smith: 156</p>

 

<p class="MsoNormal" style="margin-bottom:.0001pt;line-height:normal;">Pathology and bacteriology — <span> </span>Widal’s

reaction with sterilized cultures; a new medium for typhoid work; report on a

further series of blood cultures from seventy-four cases of typhoid and

paratyphoid fever; the histology of liver tissue regeneration; typhoid bacilli

and gall bladder; the occurrence and distribution of the spirochaeta pallida in

congenital syphilis; experiments on the differentiation of cholera and

cholera-like vitrios by complement fixation;<span> 

</span>C. S. Butler and O. J. Mink 166</p>

 

<p class="MsoNormal" style="margin-bottom:.0001pt;line-height:normal;">Medical zoology —What is "schistosoma mansoni;" pulmonary

bilharziasis; filariasis and elephantiasis in southern Luzon; the diagnosis of African

tick fever from the examination of the blood; the parasite of</p>

 

<p class="MsoNormal" style="margin-bottom:.0001pt;line-height:normal;">Kula Azar and allied organisms; a new human nematode-strongylus gibsoni;

report of the Permanent Commission for the Suppression of Uncinariasis; on the

supposed occurrence of the filaria immitis in man, R. C. Holcomb 174</p>

 

<p class="MsoNormal" style="margin-bottom:.0001pt;line-height:normal;">Tropical medicine —An inquiry concerning the etiology of beriberi; have

trypanosomes an ultramicroscopical stage in their life history?; atoxyl as a

curative agent in malaria, E. R. Stitt 179</p>

 

<p class="MsoNormal" style="margin-bottom:.0001pt;line-height:normal;">General medicine —The treatment of acute inflammatory conditions by

Bier's hypertemia; treatment of tetanus with subarachnoid injections of

magnesium sulphate; the serum diagnosis of syphilis; tubercle bacilli in the

sputum; a summary of the most recently published work on the doctrine of

opsonins; experimental investigation on "simple continued fever," H.

M. Kennedy 182</p>

 

<p class="MsoNormal" style="margin-bottom:.0001pt;line-height:normal;">Hygiene and sanitation —On the application of heat for the purification

of water with troops in the field; catarrhal icterus of eberthian origin; the epidemic

of typhoid fever on H. M. S. Regina Elena; the treatment of sweat-foot in the

army; a contribution to our knowledge of the spread of cerebro-spinal

meningitis; on book disinfection on the large scale; the etiology of impetigo

contagiosa; tuberculosis in the British army and its prevention; symptoms that

may be attributed to soldering with the oxyhydrogen</p>

 

<p class="MsoNormal" style="margin-bottom:.0001pt;line-height:normal;">flame; tactics and the health of the army, H. G. Beyer 189</p>

 

<p class="MsoNormal" style="margin-bottom:.0001pt;line-height:normal;">Reports and letters 203</p>

 

<p class="MsoNormal" style="margin-bottom:.0001pt;line-height:normal;">Seventeenth annual meeting of the Association of Military Surgeons,

Manley H. Simons, delegate 203</p>

 

<p class="MsoNormal" style="margin-bottom:.0001pt;line-height:normal;">Report and recommendations of a board of officers, convened at the navy-yard,

Mare Island, Cal., on the precautionary methods <span> </span>to be taken to prevent the invasion of bubonic

plague at that station 205</p>

 

<p class="MsoNormal" style="margin-bottom:.0001pt;line-height:normal;"> </p>

 

<p class="MsoNormal" style="margin-bottom:.0001pt;line-height:normal;">Number 3</p>

 

<p class="MsoNormal" style="margin-bottom:.0001pt;line-height:normal;"> </p>

 

<p class="MsoNormal" style="margin-bottom:.0001pt;line-height:normal;">Preface VII</p>

 

<p class="MsoNormal" style="margin-bottom:.0001pt;line-height:normal;">Special articles 211</p>

 

<p class="MsoNormal" style="margin-bottom:.0001pt;line-height:normal;">Notes on the treatment of elephantiasis by the internal administration

of tinctuia ferri cbloridi, by P. S. Rossiter 211</p>

 

<p class="MsoNormal" style="margin-bottom:.0001pt;line-height:normal;">A few notes on syphilis, by W. J. Zalesky 215</p>

 

<p class="MsoNormal" style="margin-bottom:.0001pt;line-height:normal;">A note on the pathology of epidemic asthma, by O. J. Mink 222</p>

 

<p class="MsoNormal" style="margin-bottom:.0001pt;line-height:normal;">Report on sixteen cases of heat prostration, with remarks on etiology,

by A. G. Grunwell 223</p>

 

<p class="MsoNormal" style="margin-bottom:.0001pt;line-height:normal;">Reviews 231</p>

 

<p class="MsoNormal" style="margin-bottom:.0001pt;line-height:normal;">Liver abscess from the point of view of etiology and prophylaxis; pathology

and differential diagnosis; and treatment (3 papers), by G. B. Crow,, J. A. B.

Sinclair, and J. F. Cottle 231</p>

 

<p class="MsoNormal" style="margin-bottom:.0001pt;line-height:normal;">Suggested devices 245</p>

 

<p class="MsoNormal" style="margin-bottom:.0001pt;line-height:normal;">Appliances improvised on sick bay bunks, by C. M. De Valin 245</p>

 

<p class="MsoNormal" style="margin-bottom:.0001pt;line-height:normal;">Clinical notes 247</p>

 

<p class="MsoNormal" style="margin-bottom:.0001pt;line-height:normal;">A case of fracture of patella, with operation at sea, by N. J.

Blackwood.. 247</p>

 

<p class="MsoNormal" style="margin-bottom:.0001pt;line-height:normal;">A case of n current nasal hemorrhage, by Raymond Spear 250</p>

 

<p class="MsoNormal" style="margin-bottom:.0001pt;line-height:normal;">A case of traumatic pneumonia, by C. F. Sterne 252</p>

 

<p class="MsoNormal" style="margin-bottom:.0001pt;line-height:normal;">A case of liver abscess, by M. A. Stuart 254</p>

 

<p class="MsoNormal" style="margin-bottom:.0001pt;line-height:normal;">Current comment 255</p>

 

<p class="MsoNormal" style="margin-bottom:.0001pt;line-height:normal;">Hospital corps efficiency report 255</p>

 

<p class="MsoNormal" style="margin-bottom:.0001pt;line-height:normal;">Physical defects found on reexamination of recruits 255</p>

 

<p class="MsoNormal" style="margin-bottom:.0001pt;line-height:normal;">Some observations on the berthing of enlisted men of the navy, with suggestions

for improvement, by L. W. Curtis 256</p>

 

<p class="MsoNormal" style="margin-bottom:.0001pt;line-height:normal;">The value of a chemical examination of water, by E. R. Noyes 257</p>

 

<p class="MsoNormal" style="margin-bottom:.0001pt;line-height:normal;">Progress in medical sciences 267</p>

 

<p class="MsoNormal" style="margin-bottom:.0001pt;line-height:normal;">Laboratory —A method for the preparation of flat worms for study, by O.

J. Mink and A. H. Ebeling .. 267</p>

 

<p class="MsoNormal" style="margin-bottom:.0001pt;line-height:normal;">The formalin method for the clinical estimation of ammonia in the

urine, by E. W. Brown 269</p>

 

<p class="MsoNormal" style="margin-bottom:.0001pt;line-height:normal;">Bang's method for estimation of sugar in the urine; the Edestin method for

the estimation of pepsin in stomach contents 273</p>

 

<p class="MsoNormal" style="margin-bottom:.0001pt;line-height:normal;">Chemistry and pharmacy — Concerning the fractional precipitation of

albumin in the spinal fluid of normal cases luetics, functional and organic nervous

diseases and their bearing upon the differential diagnosis of dementia

paralytica, tabes dorsalis, tertiary and late syphilis; quantitative determination

of several sugars in the presence of each other in diabetic urines; the butyric

reaction for syphilis in man and in the monkey; excretion of amino acids in

pregnancy and after parturition; the relation between the protein content of

the blood serum and that of serous fluids; the further separation of antitoxin

from its associated proteins in horse serum, E. W. Brown and P. J. Waldner...276-279</p>

 

<p class="MsoNormal" style="margin-bottom:.0001pt;line-height:normal;">Surgery —The Hodgen splint; surgical anemia and resuscitation; mechanism

underlying artificial respiration; a new theory of surgical shock; carbon

dioxide snow in the treatment of augioma; bursitis subacromialis, or

periarthritis of the shoulder joint; report on the local anesthetics recommended

as substitutes for cocaine; further researches on the etiology of endemic

goiter; auto- and iso-transplantation, in dogs, of the parathyroid glandules;

partial, progressive, and complete occlusion of the aorta and other large

arteries in the dog by means of the metal band; C. F. Stokes, R. Spear, and H.

W. Smith 279-289</p>

 

<p class="MsoNormal" style="margin-bottom:.0001pt;line-height:normal;">Pathology and bacteriology —A simple method for the diagnosis of

syphilis; differential methods for detecting the typhoid bacilli in infected

water and milk; a peculiar intralobular cirrhosis of the liver produced by the protozoal

parasite of kala azar; the pathological anatomy of atoxyl poisoning; an

observation on the fate of B. Bulgaricus in the digestive tract of a monkey; a

contribution to the pathology of the spleen; a note, on the histology of a caue

of myelomatosis with Bence-Jones protein in</p>

 

<p class="MsoNormal" style="margin-bottom:.0001pt;line-height:normal;">the urine; a new method for the recognition of indol in media; the rapid

diagnosis of rabies (a new stain for negri bodies); C. S. Butler and O. J. Mink

289-297</p>

 

<p class="MsoNormal" style="margin-bottom:.0001pt;line-height:normal;">Medical zoology —Anew intestinal trematodeof man; some applications of the

precipitin reaction in the diagnosis of hydatid disease; bilharzia, hematobia,

and circumcision; trichocephaliasis; R. C. Holcomb ...... 297-306</p>

 

<p class="MsoNormal" style="margin-bottom:.0001pt;line-height:normal;">Tropical medicine — Rice and beriberi; on the etiology of ulcerative

granuloma of the pudenda; amaebic dysentery with abscess of the liver in a patient

who had never been out of England; E. R. Stitt 306-308</p>

 

<p class="MsoNormal" style="margin-bottom:.0001pt;line-height:normal;">General medicine —The dietetic treatment of diabetes; artificial

hyperemia in the treatment of pulmonary tuberculosis; remarks on the treatment of

gastric ulcer by immediate feeding; present status of the tuberculin tests; T.

W. Richards S0S-315</p>

 

<p class="MsoNormal" style="margin-bottom:.0001pt;line-height:normal;">Hygiene and sanitation — On 'a new and practical method of securing bodily

cleanliness for our men on board ship; on the heat-conducting power of linoleum

as compared to that of floors made of wood or of</p>

 

<p class="MsoNormal" style="margin-bottom:.0001pt;line-height:normal;">betone; on the discrimination of unrecognized diseases and on a disease

of overcrowding in ships, <span> </span>especially at

Malta; H. G. Beyer 315-320</p>

 

<p class="MsoNormal" style="margin-bottom:.0001pt;line-height:normal;">Reports and letters 321</p>

 

<p class="MsoNormal" style="margin-bottom:.0001pt;line-height:normal;">Guam; reports on health and sanitation for the years 1907 and 1908, by F.

E. McCullough and G. L. Angeny. 321</p>

 

<p class="MsoNormal" style="margin-bottom:.0001pt;line-height:normal;"> </p>

 

<p class="MsoNormal" style="margin-bottom:.0001pt;line-height:normal;">Number 4</p>

 

<p class="MsoNormal" style="margin-bottom:.0001pt;line-height:normal;"> </p>

 

<p class="MsoNormal" style="margin-bottom:.0001pt;line-height:normal;">Preface vii</p>

 

<p class="MsoNormal" style="margin-bottom:.0001pt;line-height:normal;">Special articles 335</p>

 

<p class="MsoNormal" style="margin-bottom:.0001pt;line-height:normal;">The hospital camp at Norfolk, Va., by P. A. Lovering 335</p>

 

<p class="MsoNormal" style="margin-bottom:.0001pt;line-height:normal;">The teaching of tropical medicine outside of the Tropics, by E. R.

Stitt 308</p>

 

<p class="MsoNormal" style="margin-bottom:.0001pt;line-height:normal;">Ethyl chloride as a general anaesthetic, by L. W. Johnson 344</p>

 

<p class="MsoNormal" style="margin-bottom:.0001pt;line-height:normal;">Chronic nephritis in recruits, by B. F. Jenness 347</p>

 

<p class="MsoNormal" style="margin-bottom:.0001pt;line-height:normal;">Supplementary report on the investigation of Samoan conjunctivitis, by P.

S. Rossiter 349</p>

 

<p class="MsoNormal" style="margin-bottom:.0001pt;line-height:normal;">Points on embalming practicable on board ship, by C. Schaffer 351</p>

 

<p class="MsoNormal" style="margin-bottom:.0001pt;line-height:normal;">Reviews 355</p>

 

<p class="MsoNormal" style="margin-bottom:.0001pt;line-height:normal;">Surgical shock; a review of recent literature, by H. W. Smith 355</p>

 

<p class="MsoNormal" style="margin-bottom:.0001pt;line-height:normal;">Suggested devices 365</p>

 

<p class="MsoNormal" style="margin-bottom:.0001pt;line-height:normal;">Installation of an X-ray apparatus on the U. S. S. Maryland, by A.

Farenholt 365</p>

 

<p class="MsoNormal" style="margin-bottom:.0001pt;line-height:normal;">Method of fumigation of vessels at Hamburg 368</p>

 

<p class="MsoNormal" style="margin-bottom:.0001pt;line-height:normal;">An oxygen apparatus 370</p>

 

<p class="MsoNormal" style="margin-bottom:.0001pt;line-height:normal;">An easily constructed bunk tray, by C. M. Oman 371</p>

 

<p class="MsoNormal" style="margin-bottom:.0001pt;line-height:normal;">Clinical notes 373</p>

 

<p class="MsoNormal" style="margin-bottom:.0001pt;line-height:normal;">Operations upon the kidney. United States naval hospital, New York, by G.

T. Smith 373</p>

 

<p class="MsoNormal" style="margin-bottom:.0001pt;line-height:normal;">A report on two cases of dentigerous cysts, by D. N. Carpenter 374</p>

 

<p class="MsoNormal" style="margin-bottom:.0001pt;line-height:normal;">A case of mammary development in the male, by E. M. Brown 376</p>

 

<p class="MsoNormal" style="margin-bottom:.0001pt;line-height:normal;">Operative treatment of epididymitis, by W. S. Pugh, Jr 376</p>

 

<p class="MsoNormal" style="margin-bottom:.0001pt;line-height:normal;">Two cases from report of U. S. S. Hancock—1908: (1) Retinal hemorrhage,

(2) myocarditis with rupture, by P. Leach 377</p>

 

<p class="MsoNormal" style="margin-bottom:.0001pt;line-height:normal;">A case of fracture of the skull; operation and recovery, by F. W. F.

Wieber. 378</p>

 

<p class="MsoNormal" style="margin-bottom:.0001pt;line-height:normal;">Carron oil in the treatment of otitis media suppurativa (acuta), by R.

E. Riggs 379</p>

 

<p class="MsoNormal" style="margin-bottom:.0001pt;line-height:normal;">Fracture of skull and gunshot wound of lung, with recovery, by W. S.

Pugh, Jr ..... 381</p>

 

<p class="MsoNormal" style="margin-bottom:.0001pt;line-height:normal;">Two unusual appendix cases, by R. R. Richardson 382</p>

 

<p class="MsoNormal" style="margin-bottom:.0001pt;line-height:normal;">Proctoclysis in typhoid fever, by C. F. Stokes 384</p>

 

<p class="MsoNormal" style="margin-bottom:.0001pt;line-height:normal;">Current comment 385</p>

 

<p class="MsoNormal" style="margin-bottom:.0001pt;line-height:normal;">Subscription price of the Bulletin 385</p>

 

<p class="MsoNormal" style="margin-bottom:.0001pt;line-height:normal;">Note on New York Post-Graduate Medical School 385</p>

 

<p class="MsoNormal" style="margin-bottom:.0001pt;line-height:normal;">Note on contributions to the Bulletin 385</p>

 

<p class="MsoNormal" style="margin-bottom:.0001pt;line-height:normal;">Note on annual meeting of American Medical Association on revision of pharmacopeia

386</p>

 

<p class="MsoNormal" style="margin-bottom:.0001pt;line-height:normal;">Note on inquiry concerning clothing in the Tropics 386</p>

 

<p class="MsoNormal" style="margin-bottom:.0001pt;line-height:normal;">Note on publicity concerning venereal disease in California 387</p>

 

<p class="MsoNormal" style="margin-bottom:.0001pt;line-height:normal;">Review of Gatewood's Naval Hygiene 387</p>

 

<p class="MsoNormal" style="margin-bottom:.0001pt;line-height:normal;">Note on the work at Tay Tay 388</p>

 

<p class="MsoNormal" style="margin-bottom:.0001pt;line-height:normal;">Medical examination of army recruits, by A. E. Peck 389</p>

 

<p class="MsoNormal" style="margin-bottom:.0001pt;line-height:normal;">Notes on the treatment of syphilis, by W. S. Hoen 391</p>

 

<p class="MsoNormal" style="margin-bottom:.0001pt;line-height:normal;">Views on the treatment of typhoid fever, by H. A. May 393</p>

 

<p class="MsoNormal" style="margin-bottom:.0001pt;line-height:normal;">Progress in medical sciences 397</p>

 

<p class="MsoNormal" style="margin-bottom:.0001pt;line-height:normal;">Laboratory —Benedict's method for the estimation of glucose in the

urine; estimation of uric acid in the urine, Folin-Schaffer; clinical method

for the estimation of uric acid, modification of the Folin-Schaffer process; test

for blood in the urine; two methods for the estimation of albumin in the urine,

by O. J. Mink and E. W. Brown 397</p>

 

<p class="MsoNormal" style="margin-bottom:.0001pt;line-height:normal;">Chemistry and pharmacy —The excretion in urine of sugars other than

glucose; experiments and experiences, pharmacological and clinical, with

digitalis, squill, and strophanthus; a reagent for the detection of reducing

sugars; on the antagonism of alcohol to carbolic acid ; the antitoxic activity

of iodine in tuberculosis; new experiments on the physiological action of

sulphuric ether; contribution to the physiology of the glands —further

contributions on the function of the spleen as an organ of iron metabolism;

modifications in the chemical composition of the blood serum in victims of

carbon dioxide poisoning, by P. J. Waldnerand C. Schaffer 402</p>

 

<p class="MsoNormal" style="margin-bottom:.0001pt;line-height:normal;">Pathology and bacteriology —Studies on typhoid fever; chloroform

poisoning — liver necrosis and repair; the importance of blood cultures in the

study of infections of otitic origin; the cultivation of the spirocheeta

pallidum; the cultivation of the bacillus leprae; the chemistry of the liver in

chloroform necrosis; the present status of the whooping-cough question; the

conveyance of whooping cough from man to animals by direct experiment; serology

of syphilis, by C. S. Butler and O. J. Mink 407</p>

 

<p class="MsoNormal" style="margin-bottom:.0001pt;line-height:normal;">Medical zoology — Schistosomiasis at Bahia; contribution to the study

of schistosomiasis in Bahia, Brazil; notes on malaria and kala-azar; endemic

amoebic dysentery in New York, with a review of its <span> </span>istribution in North America; filaria

(microfilaria) philippinensis; the distribution of filaria in the Philippine

Islands; acariens and cancers—acariens and leprosy; necator americanus in

Ceylon; anaemia due to trichocephalus dispar; study of the protozoa of J. H.

Wright in sixteen cases of Aleppo boil, by R. C. Holcomb 411</p>

 

<p class="MsoNormal" style="margin-bottom:.0001pt;line-height:normal;">Tropical medicine — Infantile kala-azar; on the identity of beri-beri

and epidemic dropsy; Malta fever in South Africa; leprosy in the Philippine

Islands and its treatment; the various types of plague and their clinical

manifestations, by C. S. Butler 417</p>

 

<p class="MsoNormal" style="margin-bottom:.0001pt;line-height:normal;">Hygiene and sanitation —The means by which infectious diseases are

transmitted; a critical study of the value of the measurements of chest expansion

and lung capacity; notes on the sanitation of yellow fever and malaria; the

house fly as a disease carrier, by H. G. Beyer 419</p>

 

<p class="MsoNormal" style="margin-bottom:.0001pt;line-height:normal;">General medicine —A study of the aural and laryngeal complications of

typhoid fever, especially as observed in hospital practice; the problem of

cancer considered from the standpoint of immunity; nine cases of typhoid fever

treated with an antiendotoxic serum, by T. W. Richards 425</p>

 

<p class="MsoNormal" style="margin-bottom:.0001pt;line-height:normal;">Surgery —Some practical points in the application of the bismuth paste

in chronic suppurative diseases; the sequence of the pathological changes in appendiceal

peritonitis; direct blood transfusion by means of paraffin-coated glass tubes;

the use of animal membrane in producing mobility in ankylosed joints, by C. F.

Stokes and R. Spear 431</p>

 

<p class="MsoNormal" style="margin-bottom:.0001pt;line-height:normal;">Reports and letters 489</p>

 

<p class="MsoNormal" style="margin-bottom:.0001pt;line-height:normal;">American Medical Association, by M. F. Gates 439</p>

 

<p class="MsoNormal" style="margin-bottom:.0001pt;line-height:normal;">Report on the Second International Conference for Revision of Nomenclature

of Diseases and Causes of Death, by F. L. Pleadwell 445</p>

 

<p class="MsoNormal" style="margin-bottom:.0001pt;line-height:normal;">Report upon medical relief measures at Messina, Sicily, by M. Donelson.

. 449</p>

 

<p class="MsoNormal" style="margin-bottom:.0001pt;line-height:normal;">Reports of medical relief measures at Adana, Turkey, by J. T. Miller

and L. W. McGuire 452</p>

  

If you have questions concerning reproductions, please contact the Contributing Library.

 

Note: The colors, contrast and appearance of these illustrations are unlikely to be true to life. They are derived from scanned images that have been enhanced for machine interpretation and have been altered from their originals.

 

Read/Download from the Internet Archive

 

See all images from this book

See all MHL images published in the same year

#A #AffordableHealthcareAct. #Obamacare #Masturbation #Twitter #TemperTantrum #BarackObama #statue #ovaloffice #disinfectant #bleach #chlorox #lysol #chlorinebleach #101 #wipes #DrTrump #Medusa #ClashoftheTitans #Snake #Rattlesnake #Slimeball #ForkedTongue #scales #hairdye #eastereggdye #PAAS #loreal #missclairol #DonaldTrump #narcissisticpersonalitydisorder #JaredKushner #DonaldTrumpJr #COVID19 #Coronavirus #Pandemic #DrAnthonySFauci #MikePence #testicle #penis #dick #balls #AsktheMortician #disease #pneumonia #infection #flu #JimmyKimmel #JimmyFallon #StephenColbert #SethMyer #minilogue #quarantine #sociadistancing #6feet #MitchMcConnell #handsanitizer #novelcoronavirus #trumplies #wipes #stayathome #workfromhome #homeschooling #deathtoll #StevenMnuchin #MikePompeo #testspositive #testsnegative #farright #freaks #ventilator #medicalmask #hospitalbed #temporarymorgue #WuhanChina, #novelcoronavirus #sociallydistant #selfquarantine #SeanHannity #FakeNews #infectiousdisease #FoxNews #epidemic #publichealthemergency #CDC #AndrewCuomo #CenterForDiseaseControl #CNN #flatteningthecurve #worldhealthorganization #recession #depression #contagious #WHO #narcissisticpersonalitydisorder #positive #negative #coronavirus #pandemic #COVID19 #infection #Dr.AnthonyFauci #trumpcausescancer #windmill #windpower #windturbine #antivaxxer #bigfoot #UFO #lochnessmonster #flyingsaucer #mythology #atlantis #bermudatriangle #flatearth #wikileaks #angels #fallenangels #45 #BiggestBaby #dirtydiaper #TheMuellerInvestigation #WilliamBarr #AttorneyGeneral #Report #TheEmperorsNewClothes #TheExorcist #Pazuzu #God #JesusChrist #LindaBlair #possessed #KingofDemons #SarahSanders #hypocrite #Exorcism #Pampers #TheWall #NeverGiveIn #Bullshit #dirtydiaper #diaperrash #PooPooBaDoop #Clemson #Wendys #burgerking #football #KellyanneConway #SarahHuckabee #Trolls #Kremlin #KremlinTroll #RussianTroll #RobertMueller #FISA #SteveBannon #MikePence #Twitter #Tweet #wiretap #Twit #wiretapped #Twat #dontaldtrump #WashingtonDC #MamaAyeshas #wallofpresidents #CIA #GOP #KKK #ISIS #FBI #BLM #LGBT #Russia #VladimirPutin #Russianinterference #AlternativeFacts #MicrowaveOven #Camera #sexdrugsandrockandroll #HillaryClinton #BernieSanders #BarackObama #PresidentoftheUnited #plannedparenthood #bigot #jihad #OsamabinLaden #DumpTrump #NotMyPresident #Dontee #DonteesInferno #thewalkingdead #republican #pedophile #WomensMarch #badhombre #conservative #rape #RiencePriebus #DonaldMcGahn #FrankGaffney #JeffSessions #GeneralJamesMattis #GeneralJohnKelly #StevenMnuchin #AndyPuzder #WilburRoss #CathyMcMorrisRodgers #MitchMcConnell #KTMcFarland #MikePompeo #NikkiHaley #LtGenMichaelFlynn #BenCarson #BetsyDeVos #TomPrice #ScottPruitt #SeemaVerma #PaulRyan #TrumpTower #MarriageEquality #KuKluxKlan #NewYorkCity #Hanksy #MelaniaTrump #BarronTrump #IvankaTrump #TiffanyTrump #EricTrump #DonaldTrumpJr #JaredKushner #conflictofinterest #emolument #RiggedElection #TemperTantrum #Tweet #Twitter #Twit #ManChild #DiaperBlowout #Trump #poop #turd #bigbaby #manindiapers #Inauguration #ScottBaio #TedNugent #TheRockettes #RadioCityMusicHall #MormonTabernacleChoir #Medusa #breitbart #lies #NationalEnquirer #douchebag #POS #Pussy #PussyGrabber

#terrorist #Taliban #jihad #MexicanWall #racism #jihad #nobannowall #confederateflag #Nazi #Islam #Freedom #AmericanNaziParty #TheRollingStones #Democrat #CivilRights #Idiot #abortion #tinfoilhatsociety #tyrant #foxnews #MerylStreep #Liberal #SaturdayNightLive #AlecBaldwin #MelissaMcCarthy #AdolfHitler #BenitoMussolini #Dictator #Megalomaniac #KingComplex #Demagogue #Narcissist #Delusional #Nuts #Oligarch #Populist #tyrant #Narcissistic #Autocracy #Oligarchy #DelusionsofGrandeur #GodComplex #MangoMussolini #DerPumpkinfuhrer #Apocalypse #NuclearButton #OvalOffice #civilliberties #goldenshowers #tinyhands #discrimination #TrumpGate #freedomandjusticeforall #TheBible #JesusChrist #The12Apostles #FredPhelps #GodHatesFags #WestboroBaptistChurch #RedNeck #ScienceFiction, #rapistsandmurderers #antiGay #homophobe #dinosaurs #religiousright #AmericanFamilyAssociation #hategroup #BruceJenner #CaitlynJenner #BarbieandKen #Mattel #PopeFrancis #QueenElizabeth #KeepYourPeckerUp #PatRobertson #BatteredWomanSyndrome #FranklinGraham #Cracker #JudyGarland #TheWizardofOz #BarbraStreisand #BettyWhite #MarilynMonroe #ValleyoftheDolls #PeytonPlace #DowntonAbbey #MaggieSmith #JudyDench #EvaGreen #MissPeregrine #DarylDixon #jabbathehutt #EmperorPalpatine #StarWars #StarTrek #RickGrimes #TeaParty #GlennBeck #RushLimbaugh #fakeNews #politicallyincorrect #BillMaher #AngelaMerkel #TheresaMay #RosieODonnell #MegynKelly #TheManchurianCandidate #BadCombOver #commemorativecoin #collectorsitem #ebay #buffalonewyork #artvoice #carlpaladino #byecarl #OutrageFatigue #hotair #weaponsofmassdestruction #motherofallbombs #farts #farting #robertmueller #bombingsyria #kellyanneconway #brettkavanaugh #sexualassault #harrassment #metoo #supremecourt #kanyewest #kimkardashian #idiot #incoherent

Nice little feature on museum gift shops in the November 26, 2006 issue of the West Sunday Magazine in the Los Angeles Times. The LA Library Store sells my line.

for Environment and Food Jakob Ellemann-Jensen interviewed as Denmark hosts World Food Summit at the start of September. This shot was used for the cover of Atingets magasin: www.altinget.dk/magasin/vis.aspx

 

A night photo of a portion of the rear of Wall Street's Charging Bull including its hind quarters and testicles. The iconic bronze sculpture by Arturo Di Modica is located in Bowling Green, on Broadway at Morris Street in lower Manhattan, New York.

  

ronablack.com

The Southern Right Whale. Hermanus, South Africa. Oct/2015

 

The southern right whale (Eubalaena australis) is a baleen whale, one of three species classified as right whales belonging to the genus Eubalaena. Like other right whales, the southern right whale is readily distinguished from others by the callosities on its head, a broad back without a dorsal fin, and a long arching mouth that begins above the eye. Its skin is very dark grey or black, occasionally with some white patches on the belly. The right whale's callosities appear white due to large colonies of cyamids (whale lice). It is almost indistinguishable from the closely related North Atlantic and the North Pacific right whales, displaying only minor skull differences. It may have fewer callosities on its head and more on its lower lips than the two northern species. Approximately 10,000 southern right whales are spread throughout the southern part of the Southern Hemisphere.

 

The size of an adult female is 15 m (49 ft)[12] and can weigh up to 47 tonnes (46 long tons; 52 short tons), with the larger records of 17.5–18 m (57–59 ft) in length and 80 tonnes (79 long tons; 88 short tons) in weight, making them slightly smaller than other right whales in Northern Hemisphere. The testicles of right whales are likely to be the largest of any animal, each weighing around 500 kg (1,100 lb). This suggests that sperm competition is important in the mating process. Right whales cannot cross the warm equatorial waters to connect with the other (sub)species and (inter)breed: their thick layers of insulating blubber make it impossible for them to dissipate their internal body heat in tropical waters. However, based on historical records and unconfirmed sightings in modern periods, E. australis actually do occur in equatorial waters.

 

The proportion and numbers of molten-coloured individuals are notable in this species compared with the other species in the Northern Hemisphere. Some whales remain white even after growing up.

 

A baleia-franca-austral (Eubalaena australis) é uma das três espécies de baleia-franca, pertencente ao género Eubalaena. Estima-se que haja cerca de 7500 exemplares desta baleia espalhadas pelo sul do Hemisfério Sul, numa faixa compreendida entre os 30º e os 55º de latitude. Pode atingir os 18 metros de comprimento e as 80 toneladas de peso.

 

A caça indiscriminada deste tipo de baleia, devido à quantidade de óleo possuída por exemplar, deixou-a quase em perigo de extinção. Desde o século XIX, a população destes animais foi reduzida em 90%. Actualmente estima-se que exista uma população que oscila entre os 7500 e 8000 indivíduos. Durante o inverno, as baleias escolhem as águas mais quentes do hemisfério sul para se reproduzirem, tais como os seguintes as costas da Península Valdés (na Patagónia), Austrália, África do Sul e Brasil.

I took this photo of Jambo during the daily show. He is a cute white lion! :)

In the Musei Capitolini, in the Hall of Eagles, there is a small-scale Diana of Ephesus, a copy of the cult statue of the shrine of Artemis at Ephesus, a marble sculpture with bronze inlays, enriched by the presence of flowers, bees and fertility symbols. It combines elements of Cybele, the Anatolian mother goddess, and Diana, the Roman goddess of the hunt and moon. The goddess' skin on this statue is black, possibly to emulate the colour of the aged wood of the original.

 

The sculpture depicts a female figure in a frontal position, arms extended, palms turned upwards (frequently seen in 'orans' - praying figures). Noteworthy for its detailed execution combined with stylistic simplicity, the piece exhibits typical elements of Hellenistic craftsmanship. The sculpture presents a divided personification with its lower half featuring carved animal and bird figures, likely symbolising Diana's role as the protector of nature. The upper portion portrays a female bust adorned with opulent jewelry and garments, intended to evoke Cybele's wealth.

 

The sculpture prominently features many pendulous objects arranged down the figure's chest - frequently thought of as breasts or dates but probably sacrificial bull testicles - a typical trait of Ephesian Diana, symbols of fertility and abundant life. The subject's face displays finely crafted detail, emitting a sense of tranquility.

 

Dated to the 2nd century CE, the sculpture embodies the period's Roman reinterpretation of Greek culture. It demonstrates an instance of religious syncretism, merging Anatolian, Greek, and Roman religious elements signifying Rome's geographic expansion and cultural assimilation during the era.

 

Remaining in unusually excellent condition, all sculptural details and craftsmanship are well-preserved and visible for inspection from all angles. This piece invites contemplation on the fluid evolution of religious iconography under diverse cultural influences.

 

Roman, 2nd century CE, marble and bronze.

 

Musei Capitolini, Rome (inv. MC1182)

A view of Loch Tarsan taken from the dam..

.

I'm not as sprightly as I once was so I managed to make quite a good job of mangling my testicles climbing over the gate which blocked the path over the dam :( .

Still, it was almost worth it :D

 

See where this picture was taken.. [?]

The wild boar (Sus scrofa), also known as the wild swine, common wild pig, Eurasian wild pig, or simply wild pig, is a suid native to much of Eurasia and North Africa, and has been introduced to the Americas and Oceania. The species is now one of the widest-ranging mammals in the world, as well as the most widespread suiform. It has been assessed as least concern on the IUCN Red List due to its wide range, high numbers, and adaptability to a diversity of habitats. It has become an invasive species in part of its introduced range. Wild boars probably originated in Southeast Asia during the Early Pleistocene and outcompeted other suid species as they spread throughout the Old World.

 

As of 2005, up to 16 subspecies are recognized, which are divided into four regional groupings based on skull height and lacrimal bone length. The species lives in matriarchal societies consisting of interrelated females and their young (both male and female). Fully grown males are usually solitary outside the breeding season. The wolf is the wild boar's main predator in most of its natural range except in the Far East and the Lesser Sunda Islands, where it is replaced by the tiger and Komodo dragon respectively. The wild boar has a long history of association with humans, having been the ancestor of most domestic pig breeds and a big-game animal for millennia. Boars have also re-hybridized in recent decades with feral pigs; these boar–pig hybrids have become a serious pest wild animal in the Americas and Australia.

 

Terminology

As true wild boars became extinct in Great Britain before the development of Modern English, the same terms are often used for both true wild boar and pigs, especially large or semi-wild ones. The English boar stems from the Old English bār, which is thought to be derived from the West Germanic bair, of unknown origin. Boar is sometimes used specifically to refer to males, and may also be used to refer to male domesticated pigs, especially breeding males that have not been castrated.

 

Sow, the traditional name for a female, again comes from Old English and Germanic; it stems from Proto-Indo-European, and is related to the Latin: sus and Greek hus, and more closely to the New High German Sau. The young may be called piglets or boarlets.

 

The animals' specific name scrofa is Latin for 'sow'.

 

Taxonomy and evolution

MtDNA studies indicate that the wild boar originated from islands in Southeast Asia such as Indonesia and the Philippines, and subsequently spread onto mainland Eurasia and North Africa. The earliest fossil finds of the species come from both Europe and Asia, and date back to the Early Pleistocene. By the late Villafranchian, S. scrofa largely displaced the related S. strozzii, a large, possibly swamp-adapted suid ancestral to the modern S. verrucosus throughout the Eurasian mainland, restricting it to insular Asia. Its closest wild relative is the bearded pig of Malacca and surrounding islands.

 

Domestication

With the exception of domestic pigs in Timor and Papua New Guinea (which appear to be of Sulawesi warty pig stock), the wild boar is the ancestor of most pig breeds. Archaeological evidence suggests that pigs were domesticated from wild boar as early as 13,000–12,700 BCE in the Near East in the Tigris Basin, being managed in the wild in a way similar to the way they are managed by some modern New Guineans. Remains of pigs have been dated to earlier than 11,400 BCE in Cyprus. Those animals must have been introduced from the mainland, which suggests domestication in the adjacent mainland by then. There was also a separate domestication in China, which took place about 8,000 years ago.

 

DNA evidence from sub-fossil remains of teeth and jawbones of Neolithic pigs shows that the first domestic pigs in Europe had been brought from the Near East. This stimulated the domestication of local European wild boars, resulting in a third domestication event with the Near Eastern genes dying out in European pig stock. Modern domesticated pigs have involved complex exchanges, with European domesticated lines being exported in turn to the ancient Near East. Historical records indicate that Asian pigs were introduced into Europe during the 18th and early 19th centuries. Domestic pigs tend to have much more developed hindquarters than their wild boar ancestors, to the point where 70% of their body weight is concentrated in the posterior, which is the opposite of wild boar, where most of the muscles are concentrated on the head and shoulders.

 

Synonymous species

The Heude's pig (Sus bucculentus), also known as the Indochinese warty pig or Vietnam warty pig, was an alleged pig species found in Laos and Vietnam. It was virtually unknown and was feared extinct, until the discovery of a skull from a recently killed individual in the Annamite Range, Laos, in 1995. Subsequent studies indicated that Sus bucculentus was not a valid taxon. As of 2022 the Mammal Diversity Database included it in Sus scrofa.

 

Description

The wild boar is a bulky, massively built suid with short and relatively thin legs. The trunk is short and robust, while the hindquarters are comparatively underdeveloped. The region behind the shoulder blades rises into a hump and the neck is short and thick to the point of being nearly immobile. The animal's head is very large, taking up to one-third of the body's entire length. The structure of the head is well suited for digging. The head acts as a plough, while the powerful neck muscles allow the animal to upturn considerable amounts of soil: it is capable of digging 8–10 cm (3.1–3.9 in) into frozen ground and can upturn rocks weighing 40–50 kg (88–110 lb). The eyes are small and deep-set and the ears long and broad. The species has well developed canine teeth, which protrude from the mouths of adult males. The medial hooves are larger and more elongated than the lateral ones and are capable of quick movements. The animal can run at a maximum speed of 40 km/h (25 mph) and jump at a height of 140–150 cm (55–59 in).

 

Sexual dimorphism is very pronounced in the species, with males being typically 5–10% larger and 20–30% heavier than females. Males also sport a mane running down the back, which is particularly apparent during autumn and winter. The canine teeth are also much more prominent in males and grow throughout life. The upper canines are relatively short and grow sideways early in life, though they gradually curve upwards. The lower canines are much sharper and longer, with the exposed parts measuring 10–12 cm (3.9–4.7 in) in length. In the breeding period, males develop a coating of subcutaneous tissue, which may be 2–3 cm (0.79–1.18 in) thick, extending from the shoulder blades to the rump, thus protecting vital organs during fights. Males sport a roughly egg-sized sack near the opening of the penis, which collects urine and emits a sharp odour. The function of this sack is not fully understood.

 

Adult size and weight is largely determined by environmental factors; boars living in arid areas with little productivity tend to attain smaller sizes than their counterparts inhabiting areas with abundant food and water. In most of Europe, males average 75–100 kg (165–220 lb) in weight, 75–80 cm (30–31 in) in shoulder height and 150 cm (59 in) in body length, whereas females average 60–80 kg (130–180 lb) in weight, 70 cm (28 in) in shoulder height and 140 cm (55 in) in body length. In Europe's Mediterranean regions, males may reach average weights as low as 50 kg (110 lb) and females 45 kg (99 lb), with shoulder heights of 63–65 cm (25–26 in). In the more productive areas of Eastern Europe, males average 110–130 kg (240–290 lb) in weight, 95 cm (37 in) in shoulder height and 160 cm (63 in) in body length, while females weigh 95 kg (209 lb), reach 85–90 cm (33–35 in) in shoulder height, and reach 145 cm (57 in) in body length. In Western and Central Europe, the largest males weigh 200 kg (440 lb) and females 120 kg (260 lb). In Northeastern Asia, large males can reach brown bear-like sizes, weighing 270 kg (600 lb) and measuring 110–118 cm (43–46 in) in shoulder height. Some adult males in Ussuriland and Manchuria have been recorded to weigh 300–350 kg (660–770 lb) and measure 125 cm (49 in) in shoulder height. Adults of this size are generally immune from wolf predation. Such giants are rare in modern times, due to past overhunting preventing animals from attaining their full growth.

 

The winter coat consists of long, coarse bristles underlaid with short brown downy fur. The length of these bristles varies along the body, with the shortest being around the face and limbs and the longest running along the back. These back bristles form the aforementioned mane prominent in males and stand erect when the animal is agitated. Colour is highly variable; specimens around Lake Balkhash are very lightly coloured, and can even be white, while some boars from Belarus and Ussuriland can be black. Some subspecies sport a light-coloured patch running backward from the corners of the mouth. Coat colour also varies with age, with piglets having light brown or rusty-brown fur with pale bands extending from the flanks and back.

 

The wild boar produces a number of different sounds which are divided into three categories:

 

Contact calls: Grunting noises which differ in intensity according to the situation. Adult males are usually silent, while females frequently grunt and piglets whine. When feeding, boars express their contentment through purring. Studies have shown that piglets imitate the sounds of their mother, thus different litters may have unique vocalisations.

Alarm calls: Warning cries emitted in response to threats. When frightened, boars make loud huffing ukh! ukh! sounds or emit screeches transcribed as gu-gu-gu.

Combat calls: High-pitched, piercing cries.

Its sense of smell is very well developed to the point that the animal is used for drug detection in Germany. Its hearing is also acute, though its eyesight is comparatively weak, lacking color vision and being unable to recognise a standing human 10–15 metres (33–49 ft) away.

 

Pigs are one of four known mammalian taxa which possess mutations in the nicotinic acetylcholine receptor that protect against snake venom. Mongooses, honey badgers, hedgehogs, and pigs all have modifications to the receptor pocket which prevents the snake venom α-neurotoxin from binding. These represent four separate, independent mutations.

 

Social behaviour and life cycle

Boars are typically social animals, living in female-dominated sounders consisting of barren sows and mothers with young led by an old matriarch. Male boars leave their sounder at the age of 8–15 months, while females either remain with their mothers or establish new territories nearby. Subadult males may live in loosely knit groups, while adult and elderly males tend to be solitary outside the breeding season.

 

The breeding period in most areas lasts from November to January, though most mating only lasts a month and a half. Prior to mating, the males develop their subcutaneous armour in preparation for confronting rivals. The testicles double in size and the glands secrete a foamy yellowish liquid. Once ready to reproduce, males travel long distances in search of a sounder of sows, eating little on the way. Once a sounder has been located, the male drives off all young animals and persistently chases the sows. At this point, the male fiercely fights potential rivals. A single male can mate with 5–10 sows. By the end of the rut, males are often badly mauled and have lost 20% of their body weight, with bite-induced injuries to the penis being common. The gestation period varies according to the age of the expecting mother. For first-time breeders, it lasts 114–130 days, while it lasts 133–140 days in older sows. Farrowing occurs between March and May, with litter sizes depending on the age and nutrition of the mother. The average litter consists of 4–6 piglets, with the maximum being 10–12. The piglets are whelped in a nest constructed from twigs, grasses and leaves. Should the mother die prematurely, the piglets are adopted by the other sows in the sounder.

 

Newborn piglets weigh around 600–1,000 grams, lacking underfur and bearing a single milk incisor and canine on each half of the jaw. There is intense competition between the piglets over the most milk-rich nipples, as the best-fed young grow faster and have stronger constitutions. The piglets do not leave the lair for their first week of life. Should the mother be absent, the piglets lie closely pressed to each other. By two weeks of age, the piglets begin accompanying their mother on her journeys. Should danger be detected, the piglets take cover or stand immobile, relying on their camouflage to keep them hidden. The neonatal coat fades after three months, with adult colouration being attained at eight months. Although the lactation period lasts 2.5–3.5 months, the piglets begin displaying adult feeding behaviours at the age of 2–3 weeks. The permanent dentition is fully formed by 1–2 years. With the exception of the canines in males, the teeth stop growing during the middle of the fourth year. The canines in old males continue to grow throughout their lives, curving strongly as they age. Sows attain sexual maturity at the age of one year, with males attaining it a year later. However, estrus usually first occurs after two years in sows, while males begin participating in the rut after 4–5 years, as they are not permitted to mate by the older males. The maximum lifespan in the wild is 10–14 years, though few specimens survive past 4–5 years. Boars in captivity have lived for 20 years.

 

Behaviour and ecology

The wild boar inhabits a diverse array of habitats from boreal taigas to deserts. In mountainous regions, it can even occupy alpine zones, occurring up to 1,900 m (6,200 ft) in the Carpathians, 2,600 m (8,500 ft) in the Caucasus and up to 3,600–4,000 m (11,800–13,100 ft) in the mountains in Central Asia and Kazakhstan. In order to survive in a given area, wild boars require a habitat fulfilling three conditions: heavily brushed areas providing shelter from predators, water for drinking and bathing purposes and an absence of regular snowfall.

 

The main habitats favored by boars in Europe are deciduous and mixed forests, with the most favorable areas consisting of forest composed of oak and beech enclosing marshes and meadows. In the Białowieża Forest, the animal's primary habitat consists of well-developed broad-leaved and mixed forests, along with marshy mixed forests, with coniferous forests and undergrowths being of secondary importance. Forests made up entirely of oak groves and beeches are used only during the fruit-bearing season. This is in contrast to the Caucasian and Transcaucasian mountain areas, where boars will occupy such fruit-bearing forests year-round. In the mountainous areas of the Russian Far East, the species inhabits nutpine groves, hilly mixed forests where Mongolian oak and Korean pine are present, swampy mixed taiga and coastal oak forests. In Transbaikalia, boars are restricted to river valleys with nut pine and shrubs. Boars are regularly encountered in pistachio groves in winter in some areas of Tajikistan and Turkmenistan, while in spring they migrate to open deserts; boar have also colonized deserts in several areas they have been introduced to.

 

On the islands of Komodo and Rinca, the boar mostly inhabits savanna or open monsoon forests, avoiding heavily forested areas unless pursued by humans. Wild boar are known to be competent swimmers, capable of covering long distances. In 2013, one boar was reported to have completed the 11-kilometre (7 mi) swim from France to Alderney in the Channel Islands. Due to concerns about disease, it was shot and incinerated.

 

Wild boar rest in shelters, which contain insulating material like spruce branches and dry hay. These resting places are occupied by whole families (though males lie separately) and are often located in the vicinity of streams, in swamp forests and in tall grass or shrub thickets. Boars never defecate in their shelters and will cover themselves with soil and pine needles when irritated by insects.

 

Diet

The wild boar is a highly versatile omnivore, whose diversity in choice of food is comparable to that of humans. Their foods can be divided into four categories:

 

Rhizomes, roots, tubers and bulbs, all of which are dug up throughout the year in the animal's whole range.

Nuts, berries and seeds, which are consumed when ripened and are dug up from the snow when necessary.

Leaves, bark, twigs and shoots, along with garbage.

Earthworms, insects, mollusks, fish, rodents, insectivores, bird eggs, lizards, snakes, frogs and carrion. Most of these prey items are taken in warm periods.

A 50 kg (110 lb) boar needs around 4,000–4,500 calories of food per day, though this required amount increases during winter and pregnancy, with the majority of its diet consisting of food items dug from the ground, like underground plant material and burrowing animals. Acorns and beechnuts are invariably its most important food items in temperate zones, as they are rich in the carbohydrates necessary for the buildup of fat reserves needed to survive lean periods. In Western Europe, underground plant material favoured by boars includes bracken, willow herb, bulbs, meadow herb roots and bulbs and the bulbs of cultivated crops. Such food is favoured in early spring and summer, but may also be eaten in autumn and winter during beechnut and acorn crop failures. Should regular wild foods become scarce, boars will eat tree bark and fungi, as well as visit cultivated potato and artichoke fields. Boar soil disturbance and foraging have been shown to facilitate invasive plants. Boars of the vittatus subspecies in Ujung Kulon National Park in Java differ from most other populations by their primarily frugivorous diet, which consists of 50 different fruit species, especially figs, thus making them important seed dispersers. The wild boar can consume numerous genera of poisonous plants without ill effect, including Aconitum, Anemone, Calla, Caltha, Ferula and Pteridium.

 

Boars may occasionally prey on small vertebrates like newborn deer fawns, leporids and galliform chicks. Boars inhabiting the Volga Delta and near some lakes and rivers of Kazakhstan have been recorded to feed extensively on fish like carp and Caspian roach. Boars in the former area will also feed on cormorant and heron chicks, bivalved molluscs, trapped muskrats and mice. There is at least one record of a boar killing and eating a bonnet macaque in southern India's Bandipur National Park, though this may have been a case of intraguild predation, brought on by interspecific competition for human handouts. There is also at least one recorded case of a group of wild boar attacking, killing, and eating an adult, healthy female axis deer (Axis axis) as a pack.

 

Predators

Piglets are vulnerable to attack from medium-sized felids like Eurasian lynx (Lynx lynx), jungle cats (Felis chaus), and snow leopards (Panthera uncia), as well as other carnivorans like brown bears (Ursus arctos) and yellow-throated martens (Martes flavigula).

 

The wolf (Canis lupus) is the main predator of wild boar throughout most of its range. A single wolf can kill around 50 to 80 boars of differing ages in one year. In Italy and Belarus' Belovezhskaya Pushcha National Park, boars are the wolf's primary prey, despite an abundance of alternative, less powerful ungulates. Wolves are particularly threatening during the winter, when deep snow impedes the boars' movements. In the Baltic regions, heavy snowfall can allow wolves to eliminate boars from an area almost completely. Wolves primarily target piglets and subadults and only rarely attack adult sows. Adult males are usually avoided entirely. Dholes (Cuon alpinus) may also prey on boars, to the point of keeping their numbers down in northwestern Bhutan, despite there being many more cattle in the area.

 

Leopards (Panthera pardus) are predators of wild boar in the Caucasus (particularly Transcaucasia), the Russian Far East, India, China and Iran. In most areas, boars constitute only a small part of the leopard's diet. However, in Iran's Sarigol National Park, boars are the second most frequently targeted prey species after mouflon (Ovis gmelini), though adult individuals are generally avoided, as they are above the leopard's preferred weight range of 10–40 kg (22–88 lb). This dependence on wild boar is largely due in part to the local leopard subspecies' large size.

 

Boars of all ages were once the primary prey of the tiger (Panthera tigris) in Transcaucasia, Kazakhstan, Middle Asia and the Far East up until the late 19th century. In modern times, tiger numbers are too low to have a limiting effect on boar populations. A single tiger can systematically destroy an entire sounder by preying on its members one by one, before moving on to another sounder. Tigers have been noted to chase boars for longer distances than with other prey. In two rare cases, boars were reported to gore a small tiger and a tigress to death in self-defense. A "large male tiger" died of wounds inflicted by an old wild boar it had killed in "a battle royal" between the two animals.

 

In the Amur region, wild boars are one of the two most important prey species for Siberian tigers, alongside the Manchurian wapiti (Cervus canadensis xanthopygus), with the two species collectively comprising roughly 80% of the felid's prey. In Sikhote Alin, a tiger can kill 30–34 boars a year. Studies of tigers in India indicate that boars are usually secondary in preference to various cervids and bovids, though when boars are targeted, healthy adults are caught more frequently than young and sick specimens.

 

On the islands of Komodo, Rinca and Flores, the boar's main predator is the Komodo dragon (Varanus komodoensis).

 

Distribution and habitat

Reconstructed range

The species originally occurred in North Africa and much of Eurasia; from the British Isles to Korea and the Sunda Islands. The northern limit of its range extended from southern Scandinavia to southern Siberia and Japan. Within this range, it was only absent in extremely dry deserts and alpine zones. It was once found in North Africa along the Nile valley up to Khartoum and north of the Sahara. The species occurs on a few Ionian and Aegean Islands, sometimes swimming between islands. The reconstructed northern boundary of the animal's Asian range ran from Lake Ladoga (at 60°N) through the area of Novgorod and Moscow into the southern Urals, where it reached 52°N. From there, the boundary passed Ishim and farther east the Irtysh at 56°N. In the eastern Baraba steppe (near Novosibirsk) the boundary turned steep south, encircled the Altai Mountains and went again eastward including the Tannu-Ola Mountains and Lake Baikal. From here, the boundary went slightly north of the Amur River eastward to its lower reaches at the Sea of Okhotsk. On Sakhalin, there are only fossil reports of wild boar. The southern boundaries in Europe and Asia were almost invariably identical to the seashores of these continents. It is absent in the dry regions of Mongolia from 44 to 46°N southward, in China westward of Sichuan and in India north of the Himalayas. It is absent in the higher elevations of the Pamir and the Tian Shan, though they do occur in the Tarim basin and on the lower slopes of the Tien Shan.

 

Present range

In recent centuries, the range of wild boar has changed dramatically, largely due to hunting by humans and more recently because of captive wild boar escaping into the wild. Prior to the 20th century, boar populations had declined in numerous areas, with British populations probably becoming extinct during the 13th century. In the warm period after the ice age, wild boar lived in the southern parts of Sweden and Norway and north of Lake Ladoga in Karelia. It was previously thought that the species did not live in Finland during prehistory because no prehistoric wild boar bones had been found within the borders of the country. It was not until 2013, when a wild boar bone was found in Askola, that the species was found to have lived in Finland more than 8,000 years ago. It is believed, however, that man prevented its establishment by hunting. In Denmark, the last boar was shot at the beginning of the 19th century, and by 1900 they were absent in Tunisia and Sudan and large areas of Germany, Austria and Italy. In Russia, they were extirpated in wide areas by the 1930s. The last boar in Egypt reportedly died on 20 December 1912 in the Giza Zoo, with wild populations having disappeared by 1894–1902. Prince Kamal el Dine Hussein attempted to repopulate Wadi El Natrun with boars of Hungarian stock, but they were quickly exterminated by poachers.

 

A revival of boar populations began in the middle of the 20th century. By 1950, wild boar had once again reached their original northern boundary in many parts of their Asiatic range. By 1960, they reached Leningrad and Moscow and by 1975, they were to be found in Archangelsk and Astrakhan. In the 1970s they again occurred in Denmark and Sweden, where captive animals escaped and now survive in the wild. In England, wild boar populations re-established themselves in the 1990s, after escaping from specialist farms that had imported European stock.

 

Status in Great Britain

By the 11th century, wild boars were apparently already becoming rare in Britain. A 1087 forestry law enacted by William the Conqueror punished through blinding the unlawful killing of a boar. Charles I attempted to reintroduce the species into the New Forest, but this population was exterminated in the 17th century during the English Civil War. Between their medieval extinction and the 1980s, when wild boar farming began, only a handful of captive wild boar, imported from the continent, were present in Britain. Occasional escapes of wild boar from wildlife parks have occurred as early as the 1970s, but since the early 1990s significant populations have re-established themselves after escapes from farms, the number of which has increased as the demand for meat from the species has grown. A 1998 MAFF (now DEFRA) study on wild boar living wild in Britain confirmed the presence of two populations of wild boar living in Britain; one in Kent/East Sussex and another in Dorset.

 

Another DEFRA report, in February 2008, confirmed the existence of these two sites as 'established breeding areas' and identified a third in Gloucestershire/Herefordshire; in the Forest of Dean/Ross on Wye area. A 'new breeding population' was also identified in Devon. There is another significant population in Dumfries and Galloway. Populations estimates were as follows:

 

The largest population, in Kent/East Sussex, was then estimated at 200 animals in the core distribution area.

The smallest, in west Dorset, was estimated to be fewer than 50 animals.

Since winter 2005–2006 significant escapes/releases have also resulted in animals colonizing areas around the fringes of Dartmoor, in Devon. These are considered as an additional single 'new breeding population' and currently estimated to be up to 100 animals.

Population estimates for the Forest of Dean are disputed as, at the time that the DEFRA population estimate was 100, a photo of a boar sounder in the forest near Staunton with over 33 animals visible was published and at about the same time over 30 boar were seen in a field near the original escape location of Weston under Penyard many kilometres or miles away. In early 2010 the Forestry Commission embarked on a cull, with the aim of reducing the boar population from an estimated 150 animals to 100. By August it was stated that efforts were being made to reduce the population from 200 to 90, but that only 25 had been killed. The failure to meet cull targets was confirmed in February 2011.

 

Wild boars have crossed the River Wye into Monmouthshire, Wales. Iolo Williams, the BBC Wales wildlife expert, attempted to film Welsh boar in late 2012. Many other sightings, across the UK, have also been reported. The effects of wild boar on the U.K.'s woodlands were discussed with Ralph Harmer of the Forestry Commission on the BBC Radio's Farming Today radio programme in 2011. The programme prompted activist writer George Monbiot to propose a thorough population study, followed by the introduction of permit-controlled culling.

 

In Scotland wild boar can be killed legally without a license and are culled by land managers as wild populations appear occasionally.

 

Introduction to North America

Wild boars are an invasive species in the Americas, having been introduced by European explorers and settlers in the 16th century to serve as a source of food. Wild boars now cause problems including out-competing native species for food, destroying the nests of ground-nesting species, killing fawns and young domestic livestock, destroying agricultural crops, eating tree seeds and seedlings, destroying native vegetation and wetlands through wallowing, damaging water quality, coming into violent conflict with humans and pets and carrying pig and human diseases including brucellosis, trichinosis and pseudorabies. In some jurisdictions, it is illegal to import, breed, release, possess, sell, distribute, trade, transport, hunt, or trap Eurasian boars. Hunting and trapping is done systematically, to increase the chance of eradication and to remove the incentive to illegally release boars, which have mostly been spread deliberately by sport hunters.

 

History

While domestic pigs, both captive and feral (popularly termed "razorbacks"), have been in North America since the earliest days of European colonization, pure wild boars were not introduced into the New World until the 19th century. The suids were released into the wild by wealthy landowners as big game animals. The initial introductions took place in fenced enclosures, though several escapes occurred, with the escapees sometimes intermixing with already established feral pig populations.

 

The first of these introductions occurred in New Hampshire in 1890. Thirteen wild boars from Germany were purchased by Austin Corbin from Carl Hagenbeck and released into a 9,500-hectare (23,000-acre) game preserve in Sullivan County. Several of these boars escaped, though they were quickly hunted down by locals. Two further introductions were made from the original stocking, with several escapes taking place due to breaches in the game preserve's fencing. These escapees have ranged widely, with some specimens having been observed crossing into Vermont.

 

In 1902, 15–20 wild boar from Germany were released into a 3,200-hectare (7,900-acre) estate in Hamilton County, New York. Several specimens escaped six years later, dispersing into the William C. Whitney Wilderness Area, with their descendants surviving for at least 20 years.

 

The most extensive boar introduction in the US took place in western North Carolina in 1912, when 13 boars of undetermined European origin were released into two fenced enclosures in a game preserve in Hooper Bald, Graham County. Most of the specimens remained in the preserve for the next decade, until a large-scale hunt caused the remaining animals to break through their confines and escape. Some of the boars migrated to Tennessee, where they intermixed with both free-ranging and feral pigs in the area. In 1924, a dozen Hooper Bald wild pigs were shipped to California and released in a property between Carmel Valley and the Los Padres National Forest. These hybrid boar were later used as breeding stock on various private and public lands throughout the state, as well as in other states like Florida, Georgia, South Carolina, West Virginia and Mississippi.

 

Several wild boars from Leon Springs and the San Antonio, Saint Louis and San Diego Zoos were released in the Powder Horn Ranch in Calhoun County, Texas, in 1939. These specimens escaped and established themselves in surrounding ranchlands and coastal areas, with some crossing the Espiritu Santo Bay and colonizing Matagorda Island. Descendants of the Powder Horn Ranch boars were later released onto San José Island and the coast of Chalmette, Louisiana.

 

Wild boar of unknown origin were stocked in a ranch in the Edwards Plateau in the 1940s, only to escape during a storm and hybridize with local feral pig populations, later spreading into neighboring counties.

 

Starting in the mid-1980s, several boars purchased from the San Diego Zoo and Tierpark Berlin were released into the United States. A decade later, more specimens from farms in Canada and Białowieża Forest were let loose. In recent years, wild pig populations have been reported in 44 states within the US, most of which are likely wild boar–feral hog hybrids. Pure wild boar populations may still be present, but are extremely localized.

 

Introduction and lack of control in South America

In South America, the European boar is believed to have been introduced for the first time in Argentina and Uruguay around the 20th century for breeding purposes. In Brazil, the creation of wild boar and hybrids started on a large scale in the mid-1990s. With the invasion of wild boar that crossed the border and entered Rio Grande do Sul around 1989, and the escape and intentional release by several Brazilian breeders in the late 1990s – in response to a IBAMA decision against the import and breeding of wild boar in 1998 – numerous feral species formed a growing population, which progressively advances in Brazilian territory. The species has no natural predators in Brazil, as it is an exotic species, in addition to breeding with the domestic pig, generating the so-called "javaporco" (neologism created to define this hybrid), factors that contribute to the exaggerated increase in the population. With its population in continuous and uncontrolled growth, without predators, the wild boar causes environmental damage, contributing to the aggradation of river and stream springs, attacking native species feeding on eggs and puppies, causing damage to fauna, flora and to agriculture and livestock, since it also attacks farm animals and can carry various diseases, including zoonosis.

 

Pest control in Brazil

As a form of control for the wild boar population (which is considered a pest and harmful species), hunting and killing are allowed for Collectors, Shooters and Hunters (CACs) duly registered by the environmental control agency, IBAMA, which, on the other hand, seeks to encourage the preservation of similar species of native peccaries, such as the "queixada" and the "caititu".

 

Effect on other habitats

Wild boars negatively impact other habitats through the destruction of the environment, or homes of wildlife. When wild boars invade new areas, they adapt to the new area by trampling and rooting, as well as displacing many saplings/nutrients. This causes a decrease in growing of many plants and trees. Water is also affected negatively by wild boars. When wild boars are active in streams, or small pools of water, it causes increased turbidity (excessive silt and particle suspension). In some cases, the fecal coliform concentration increases to dangerous levels because of wild boars. Aquatic wildlife is affected, more prominently fish, and amphibians. Wild boars have caused a great decrease in over 300 animal or plant species, 250 being endangered or threatened.

 

The boars cause many habitats to become less diverse because of their feeding behaviors and predation. Wild boars will dig up eggs of species and eat them, as well as killing other wildlife for food. When these boars compete with other species for resources, they usually come out successful. A study published in the Journal of Experimental Marine Biology and Ecology was conducted on the results of Feral Swine control. Only two years after the control started, the amount of turtle nests jumped from 57 to 143, and the turtle nest predation percent dropped from 74 to 15. They kill and eat deers, lizards, birds, snakes, and more. These boars are called "opportunist omnivores", which means they eat almost anything. This means they can survive almost anywhere. A big surplus of food and the ability to adapt to any new place causes lots of breeding. All of these factors make it difficult to get rid of wild boars. Wild boars also tend to carry diseases and numerous pathogens. This also adds to the decrease in diversity among species.

 

Diseases and parasites

Wild boars are known to host at least 20 different parasitic worm species, with maximum infections occurring in summer. Young animals are vulnerable to helminths like Metastrongylus, which are consumed by boars through earthworms and cause death by parasitising the lungs. Wild boar also carry parasites known to infect humans, including Gastrodiscoides, Trichinella spiralis, Taenia solium, Balantidium coli and Toxoplasma gondii. Wild boar in southern regions are frequently infested with ticks (Dermacentor, Rhipicephalus, and Hyalomma) and hog lice. The species also suffers from blood-sucking flies, which it escapes by bathing frequently or hiding in dense shrubs.

 

Swine plague spreads very quickly in wild boar, with epizootics being recorded in Germany, Poland, Hungary, Belarus, the Caucasus, the Far East, Kazakhstan and other regions. Foot-and-mouth disease can also take on epidemic proportions in boar populations. The species occasionally, but rarely contracts Pasteurellosis, hemorrhagic sepsis, tularemia, and anthrax. Wild boar may on occasion contract swine erysipelas through rodents or hog lice and ticks.

 

Relationships with humans

The wild boar features prominently in the cultures of Indo-European people, many of which saw the animal as embodying warrior virtues. Cultures throughout Europe and Asia Minor saw the killing of a boar as proof of one's valor and strength. Neolithic hunter gatherers depicted reliefs of ferocious wild boars on their temple pillars at Göbekli Tepe some 11,600 years ago. Virtually all heroes in Greek mythology fight or kill a boar at one point. The demigod Herakles' third labour involves the capture of the Erymanthian Boar, Theseus slays the wild sow Phaea, and a disguised Odysseus is recognised by his handmaiden Eurycleia by the scars inflicted on him by a boar during a hunt in his youth. To the mythical Hyperboreans, the boar represented spiritual authority. Several Greek myths use the boar as a symbol of darkness, death and winter. One example is the story of the youthful Adonis, who is killed by a boar and is permitted by Zeus to depart from Hades only during the spring and summer period. This theme also occurs in Irish and Egyptian mythology, where the animal is explicitly linked to the month of October, therefore autumn. This association likely arose from aspects of the boar's actual nature. Its dark colour was linked to the night, while its solitary habits, proclivity to consume crops and nocturnal nature were associated with evil. The foundation myth of Ephesus has the city being built over the site where Prince Androklos of Athens killed a boar Boars were frequently depicted on Greek funerary monuments alongside lions, representing gallant losers who have finally met their match, as opposed to victorious hunters as lions are. The theme of the doomed, yet valorous boar warrior also occurred in Hittite culture, where it was traditional to sacrifice a boar alongside a dog and a prisoner of war after a military defeat

 

The boar as a warrior also appears in Germanic cultures, with its image having been frequently engraved on shields and swords. They also feature on Germanic boar helmets, such as the Benty Grange helmet, where it was believed to offer protection to the wearer and has been theorised to have been used in spiritual transformations into swine, similar to berserkers. The boar features heavily in religious practice in Germanic paganism where it is closely associated with Freyr and has also been suggested to have been a totemic animal to the Swedes, especially to the Yngling royal dynasty who claimed descent from the god.

 

According to Tacitus, the Baltic Aesti featured boars on their helmets and may have also worn boar masks. The boar and pig were held in particularly high esteem by the Celts, who considered them to be their most important sacred animal. Some Celtic deities linked to boars include Moccus and Veteris. It has been suggested that some early myths surrounding the Welsh hero Culhwch involved the character being the son of a boar god. Nevertheless, the importance of the boar as a culinary item among Celtic tribes may have been exaggerated in popular culture by the Asterix series, as wild boar bones are rare among Celtic archaeological sites and the few that do occur show no signs of butchery, having probably been used in sacrificial rituals.

 

The boar also appears in Vedic mythology and Hindu mythology. A story present in the Brahmanas has the god Indra slaying an avaricious boar, who has stolen the treasure of the asuras, then giving its carcass to the god Vishnu, who offered it as a sacrifice to the gods. In the story's retelling in the Charaka Samhita, the boar is described as a form of Prajapati and is credited with having raised the Earth from the primeval waters. In the Ramayana and the Puranas, the same boar is portrayed as Varaha, an avatar of Vishnu.

 

In Japanese culture, the boar is widely seen as a fearsome and reckless animal, to the point that several words and expressions in Japanese referring to recklessness include references to boars. The boar is the last animal of the Oriental zodiac, with people born during the year of the Pig being said to embody the boar-like traits of determination and impetuosity. Among Japanese hunters, the boar's courage and defiance is a source of admiration and it is not uncommon for hunters and mountain people to name their sons after the animal inoshishi (猪). Boars are also seen as symbols of fertility and prosperity; in some regions, it is thought that boars are drawn to fields owned by families including pregnant women, and hunters with pregnant wives are thought to have greater chances of success when boar hunting. The animal's link to prosperity was illustrated by its inclusion on the ¥10 note during the Meiji period and it was once believed that a man could become wealthy by keeping a clump of boar hair in his wallet.

 

In the folklore of the Mongol Altai Uriankhai tribe, the wild boar was associated with the watery underworld, as it was thought that the spirits of the dead entered the animal's head, to be ultimately transported to the water. Prior to the conversion to Islam, the Kyrgyz people believed that they were descended from boars and thus did not eat pork. In Buryat mythology, the forefathers of the Buryats descended from heaven and were nourished by a boar. In China, the boar is the emblem of the Miao people.

 

The boar (sanglier) is frequently displayed in English, Scottish and Welsh heraldry. As with the lion, the boar is often shown as armed and langued. As with the bear, Scottish and Welsh heraldry displays the boar's head with the neck cropped, unlike the English version, which retains the neck. The white boar served as the badge of King Richard III of England, who distributed it among his northern retainers during his tenure as Duke of Gloucester.

 

As a game animal and food source

Humans have been hunting boar for millennia, the earliest artistic depictions of such activities dating back to the Upper Paleolithic. The animal was seen as a source of food among the Ancient Greeks, as well as a sporting challenge and source of epic narratives. The Romans inherited this tradition, with one of its first practitioners being Scipio Aemilianus. Boar hunting became particularly popular among the young nobility during the 3rd century BC as preparation for manhood and battle. A typical Roman boar hunting tactic involved surrounding a given area with large nets, then flushing the boar with dogs and immobilizing it with smaller nets. The animal would then be dispatched with a venabulum, a short spear with a crossguard at the base of the blade. More than their Greek predecessors, the Romans extensively took inspiration from boar hunting in their art and sculpture. With the ascension of Constantine the Great, boar hunting took on Christian allegorical themes, with the animal being portrayed as a "black beast" analogous to the dragon of Saint George.

 

Boar hunting continued after the fall of the Western Roman Empire, though the Germanic tribes considered the red deer to be a more noble and worthy quarry. The post-Roman nobility hunted boar as their predecessors did, but primarily as training for battle rather than sport. It was not uncommon for medieval hunters to deliberately hunt boars during the breeding season when the animals were more aggressive. During the Renaissance, when deforestation and the introduction of firearms reduced boar numbers, boar hunting became the sole prerogative of the nobility, one of many charges brought up against the rich during the German Peasants' War and the French Revolution.

 

During the mid-20th century, 7,000–8,000 boars were caught in the Caucasus, 6,000–7,000 in Kazakhstan and about 5,000 in Central Asia during the Soviet period, primarily through the use of dogs and beats. In Nepal, farmers and poachers eliminate boars by baiting balls of wheat flour containing explosives with kerosene oil, with the animals' chewing motions triggering the devices.

 

Wild boar can thrive in captivity, though piglets grow slowly and poorly without their mothers. Products derived from wild boar include meat, hide and bristles. Apicius devotes a whole chapter to the cooking of boar meat, providing 10 recipes involving roasting, boiling and what sauces to use. The Romans usually served boar meat with garum. Boar's head was the centrepiece of most medieval Christmas celebrations among the nobility. Although growing in popularity as a captive-bred source of food, the wild boar takes longer to mature than most domestic pigs and it is usually smaller and produces less meat. Nevertheless, wild boar meat is leaner and healthier than pork, being of higher nutritional value and having a much higher concentration of essential amino acids. Most meat-dressing organizations agree that a boar carcass should yield 50 kg (110 lb) of meat on average. Large specimens can yield 15–20 kg (33–44 lb) of fat, with some giants yielding 30 kg (66 lb) or more. A boar hide can measure 3 m2 (4,700 sq in) and can yield 350–1,000 grams (12–35 oz) of bristle and 400 grams (14 oz) of underwool.

 

Boars can be damaging to agriculture in situations where their natural habitat is sparse. Populations living on the outskirts of towns or farms can dig up potatoes and damage melons, watermelons and maize. However, they generally only encroach upon farms when natural food is scarce. In the Belovezh forest for example, 34–47% of the local boar population will enter fields in years of moderate availability of natural foods. While the role of boars in damaging crops is often exaggerated, cases are known of boar depredations causing famines, as was the case in Hachinohe, Japan in 1749, where 3,000 people died of what became known as the "wild boar famine". Still, within Japanese culture, the boar's status as vermin is expressed through its title as "king of pests" and the popular saying (addressed to young men in rural areas) "When you get married, choose a place with no wild boar."

 

In Central Europe, farmers typically repel boars through distraction or fright, while in Kazakhstan it is usual to employ guard dogs in plantations. However, research shows that when compared with other mitigation tactics, hunting is the only strategy to significantly reduce crop damage by boars. Although large boar populations can play an important role in limiting forest growth, they are also useful in keeping pest populations such as June bugs under control. The growth of urban areas and the corresponding decline in natural boar habitats has led to some sounders entering human habitations in search of food. As in natural conditions, sounders in peri-urban areas are matriarchal, though males tend to be much less represented and adults of both sexes can be up to 35% heavier than their forest-dwelling counterparts. As of 2010, at least 44 cities in 15 countries have experienced problems of some kind relating to the presence of habituated wild boar.

 

A 2023 study found that allowing wild pigs to forage on edible garbage in large regional landfills results in those animals getting physically large/heavier, having larger litters of piglets, and causing more wild pig-vehicle collisions in the vicinity of the landfill. The effects of letting these pigs scavenge in these landfills can present unique challenges to population management, control, public safety, and disease transmission. Wild pigs foraging on edible food waste in landfills has also been identified as a vector that facilitates the spread of African swine fever virus.

 

Attacks on humans

Actual attacks on humans are rare, but can be serious, resulting in penetrating injuries to the lower part of the body. They generally occur during the boars' rutting season from November to January, in agricultural areas bordering forests or on paths leading through forests. The animal typically attacks by charging and pointing its tusks towards the intended victim, with most injuries occurring on the thigh region. Once the initial attack is over, the boar steps back, takes position and attacks again if the victim is still moving, only ending once the victim is completely incapacitated.

 

Boar attacks on humans have been documented throughout history. The Romans and Ancient Greeks wrote of these attacks (Odysseus was wounded by a boar and Adonis was killed by one). A 2012 study compiling recorded attacks from 1825 to 2012 found accounts of 665 human victims of both wild boars and feral pigs, with the majority (19%) of attacks in the animal's native range occurring in India. Most of the attacks occurred in rural areas during the winter months in non-hunting contexts and were committed by solitary males.

 

Management

Managing wild boar is a pressing task in both native and invasive contexts as they can be disrupting to other systems when not addressed. Wild boar find their success through adaptation of daily patterns to circumvent threats. They avoid human contact through nocturnal lifestyles, despite the fact that they are not evolutionarily predisposed, and alter their diets substantially based on what is available. These "adaptive generalists", can survive in a variety of landscapes, making the prediction of their movement patterns and any potential close contact areas crucial to limiting damage. All of these qualities make them equally difficult to manage or limit.

 

Within Central Europe, the native habitat of the wild boar, there has been a push to re-evaluate interactions between wild boar and humans, with the priority of fostering positive engagement. Negative media and public perception of wild boar as "crop raiders" have made those living alongside them less willing to accept the economic damages of their behaviors, as wild boar are seen as pests. This media tone impacts management policy, with every 10 negative articles increasing wild boar policy activity by 6.7%. Contrary to this portrayal, wild boar, when managed well within their natural environments, can be a crucial part of forest ecosystems.

 

Defining the limits of proper management is difficult, but the exclusion of wild boar from rare environments is generally agreed upon, as when not properly managed, they can damage agricultural ventures and harm vulnerable plant life. These damages are estimated at $800 million yearly in environmental and financial costs for the United States alone. The breadth of this damage is due to prior inattention and lack of management tactics for extended lengths of time. Managing wild boar is a complex task, as it involves coordinating a combination of crop harvest techniques, fencing, toxic bait, corrals, and hunting. The most common tactic employed by private land owners in the United States is recreational hunting, however, this is generally not as effective on its own. Management strategies are most successful when they take into account reproduction, dispersion, and the differences between ideal resources for males and females.

 

According to a study, wild boars are causing soil disturbance that, among other problems, globally results in annual carbon dioxide emissions equivalent to that of ~1.1 million passenger vehicles (4.9 Mt, 0.01% of all GHG emissions as of 2022), implying that as of 2021 hunted boar meat – unlike other meat products – has beneficial effects on the environment even though the effect would diminish if boars are introduced for meat production and consistently retaining small populations of boars may be preferable.

© Todos los derechos reservados / © All rights reserved.

PaseAndo por Barakaldo (Vizcaya). Escultura dedicada a los trabajadores y a la industria.

 

Pulsa L para verlo en grande y caja de luz (fondo negro).

I've shared Dudette's photos and story many times. I first photo'd this deer in 2009 after a ranger told me about an antlered doe. In 2013, a photo revealed that Dudette is actually a buck without testicles. Behaves as a doe.

There are three city states in the world, which are not part of the nations by which they are surrounded. They are the Vatican, the District of Columbia and the City of London: Religion, Warfare and Money. This is why the Queen, when she attends a banquet at the Guildhall, or a service at St Paul's Cathedral, must pause to receive the Lord Mayor's consent before passing this point, and thereafter proceed behind him. The City is not part of British sovereign territory.

So who does it belong to? Not wishing to encourage the wrong sort of commenter, I'll leave it to you to look into this fascinating topic if you wish. The Middle and Inner Temple are off to the left and those are the Royal Courts of Justice across the road. The structure in the middle of the road marks the former position of Temple Bar ...the "Bar" to which members of the legal profession are "called". The nightmare creature, with its scaly body and leathern wings, which surmounts it, bears the emblem of the City, which is actually the flag of the Knights Templar. It doesn't exactly strike one as a benevolent, let alone a compassionate deity, does it? I didn't notice here, but ...and although I couldn't say why, this struck me as an interesting detail... its counterparts that mark the City boundary down on the Embankment each has a scrotum and testicles. I am not a zoologist, but I don't think reptiles have external gonads, do they?

It is like a giant stubble egg. Or a very big testicle. No, maybe not that. If one of mine felt like this then it would be straight to the doctors. Kind of wish I hadn't said that now. Just imagining if one of mine was that size is giving me very peculiar thoughts about whether or not my underwear would still fit. I don't think it would.

 

I'm not sure if that's really the way anyone should start a conversation. Not unless the conversation started with "Is that a giant testicle in your pocket or are you just plea....oh is it....um...bye". As all conversations about such things should go. Unless it's with a doctor. The last thing you want is them taking one look and bolting. No, that may be the second worse thing. The worst would be them coming back with some large, sharp and pointy.

 

This really is turning into a load of old bollocks, isn't it? Well, just the one bollock but still we should probably change the subject. Anyone been getting out? No, I don't suppose you have. Had any holi...no, you wont have those either. Have you donte something to you hair? Ah, neglect and hats. Yes, that's what I do too.

 

I think we'll leave it there for now.

#A #whatwouldjesusdo #TheBroncoBuster #FredericRemington #statue #ovaloffice #disinfectant #bleach #chlorox #lysol #chlorinebleach #101 #wipes #DrTrump #Medusa #ClashoftheTitans #Snake #Rattlesnake #Slimeball #ForkedTongue #scales #hairdye #eastereggdye #PAAS #loreal #missclairol #DonaldTrump #narcissisticpersonalitydisorder #JaredKushner #DonaldTrumpJr #COVID19 #Coronavirus #Pandemic #DrAnthonySFauci #MikePence #testicle #penis #dick #balls #AsktheMortician #disease #pneumonia #infection #flu #JimmyKimmel #JimmyFallon #StephenColbert #SethMyer #minilogue #quarantine #sociadistancing #6feet #MitchMcConnell #handsanitizer #novelcoronavirus #trumplies #wipes #stayathome #workfromhome #homeschooling #deathtoll #StevenMnuchin #MikePompeo #testspositive #testsnegative #farright #freaks #ventilator #medicalmask #hospitalbed #temporarymorgue #WuhanChina, #novelcoronavirus #sociallydistant #selfquarantine #SeanHannity #FakeNews #infectiousdisease #FoxNews #epidemic #publichealthemergency #CDC #AndrewCuomo #CenterForDiseaseControl #CNN #flatteningthecurve #worldhealthorganization #recession #depression #contagious #WHO #narcissisticpersonalitydisorder #positive #negative #coronavirus #pandemic #COVID19 #infection #Dr.AnthonyFauci #trumpcausescancer #windmill #windpower #windturbine #antivaxxer #bigfoot #UFO #lochnessmonster #flyingsaucer #mythology #atlantis #bermudatriangle #flatearth #wikileaks #angels #fallenangels #45 #BiggestBaby #dirtydiaper #TheMuellerInvestigation #WilliamBarr #AttorneyGeneral #Report #TheEmperorsNewClothes #TheExorcist #Pazuzu #God #JesusChrist #LindaBlair #possessed #KingofDemons #SarahSanders #hypocrite #Exorcism #Pampers #TheWall #NeverGiveIn #Bullshit #dirtydiaper #diaperrash #PooPooBaDoop #Clemson #Wendys #burgerking #football #KellyanneConway #SarahHuckabee #Trolls #Kremlin #KremlinTroll #RussianTroll #RobertMueller #FISA #SteveBannon #MikePence #Twitter #Tweet #wiretap #Twit #wiretapped #Twat #dontaldtrump #WashingtonDC #MamaAyeshas #wallofpresidents #CIA #GOP #KKK #ISIS #FBI #BLM #LGBT #Russia #VladimirPutin #Russianinterference #AlternativeFacts #MicrowaveOven #Camera #sexdrugsandrockandroll #HillaryClinton #BernieSanders #BarackObama #PresidentoftheUnited #plannedparenthood #bigot #jihad #OsamabinLaden #DumpTrump #NotMyPresident #Dontee #DonteesInferno #thewalkingdead #republican #pedophile #WomensMarch #badhombre #conservative #rape #RiencePriebus #DonaldMcGahn #FrankGaffney #JeffSessions #GeneralJamesMattis #GeneralJohnKelly #StevenMnuchin #AndyPuzder #WilburRoss #CathyMcMorrisRodgers #MitchMcConnell #KTMcFarland #MikePompeo #NikkiHaley #LtGenMichaelFlynn #BenCarson #BetsyDeVos #TomPrice #ScottPruitt #SeemaVerma #PaulRyan #TrumpTower #MarriageEquality #KuKluxKlan #NewYorkCity #Hanksy #MelaniaTrump #BarronTrump #IvankaTrump #TiffanyTrump #EricTrump #DonaldTrumpJr #JaredKushner #conflictofinterest #emolument #RiggedElection #TemperTantrum #Tweet #Twitter #Twit #ManChild #DiaperBlowout #Trump #poop #turd #bigbaby #manindiapers #Inauguration #ScottBaio #TedNugent #TheRockettes #RadioCityMusicHall #MormonTabernacleChoir #Medusa #breitbart #lies #NationalEnquirer #douchebag #POS #Pussy #PussyGrabber

#terrorist #Taliban #jihad #MexicanWall #racism #jihad #nobannowall #confederateflag #Nazi #Islam #Freedom #AmericanNaziParty #TheRollingStones #Democrat #CivilRights #Idiot #abortion #tinfoilhatsociety #tyrant #foxnews #MerylStreep #Liberal #SaturdayNightLive #AlecBaldwin #MelissaMcCarthy #AdolfHitler #BenitoMussolini #Dictator #Megalomaniac #KingComplex #Demagogue #Narcissist #Delusional #Nuts #Oligarch #Populist #tyrant #Narcissistic #Autocracy #Oligarchy #DelusionsofGrandeur #GodComplex #MangoMussolini #DerPumpkinfuhrer #Apocalypse #NuclearButton #OvalOffice #civilliberties #goldenshowers #tinyhands #discrimination #TrumpGate #freedomandjusticeforall #TheBible #JesusChrist #The12Apostles #FredPhelps #GodHatesFags #WestboroBaptistChurch #RedNeck #ScienceFiction, #rapistsandmurderers #antiGay #homophobe #dinosaurs #religiousright #AmericanFamilyAssociation #hategroup #BruceJenner #CaitlynJenner #BarbieandKen #Mattel #PopeFrancis #QueenElizabeth #KeepYourPeckerUp #PatRobertson #BatteredWomanSyndrome #FranklinGraham #Cracker #JudyGarland #TheWizardofOz #BarbraStreisand #BettyWhite #MarilynMonroe #ValleyoftheDolls #PeytonPlace #DowntonAbbey #MaggieSmith #JudyDench #EvaGreen #MissPeregrine #DarylDixon #jabbathehutt #EmperorPalpatine #StarWars #StarTrek #RickGrimes #TeaParty #GlennBeck #RushLimbaugh #fakeNews #politicallyincorrect #BillMaher #AngelaMerkel #TheresaMay #RosieODonnell #MegynKelly #TheManchurianCandidate #BadCombOver #commemorativecoin #collectorsitem #ebay #buffalonewyork #artvoice #carlpaladino #byecarl #OutrageFatigue #hotair #weaponsofmassdestruction #motherofallbombs #farts #farting #robertmueller #bombingsyria #kellyanneconway #brettkavanaugh #sexualassault #harrassment #metoo #supremecourt #kanyewest #kimkardashian #idiot #incoherent

UPDATE: I made it onto Gawker

I figure since London got the Dildo, Miami seems to have gotten the Testicle. If you don't know what I am talking about use Google

 

Tags

 

The Southern Right Whale. Hermanus, South Africa. Oct/2015

 

The southern right whale (Eubalaena australis) is a baleen whale, one of three species classified as right whales belonging to the genus Eubalaena. Like other right whales, the southern right whale is readily distinguished from others by the callosities on its head, a broad back without a dorsal fin, and a long arching mouth that begins above the eye. Its skin is very dark grey or black, occasionally with some white patches on the belly. The right whale's callosities appear white due to large colonies of cyamids (whale lice). It is almost indistinguishable from the closely related North Atlantic and the North Pacific right whales, displaying only minor skull differences. It may have fewer callosities on its head and more on its lower lips than the two northern species. Approximately 10,000 southern right whales are spread throughout the southern part of the Southern Hemisphere.

 

The size of an adult female is 15 m (49 ft)[12] and can weigh up to 47 tonnes (46 long tons; 52 short tons), with the larger records of 17.5–18 m (57–59 ft) in length and 80 tonnes (79 long tons; 88 short tons) in weight, making them slightly smaller than other right whales in Northern Hemisphere. The testicles of right whales are likely to be the largest of any animal, each weighing around 500 kg (1,100 lb). This suggests that sperm competition is important in the mating process. Right whales cannot cross the warm equatorial waters to connect with the other (sub)species and (inter)breed: their thick layers of insulating blubber make it impossible for them to dissipate their internal body heat in tropical waters. However, based on historical records and unconfirmed sightings in modern periods, E. australis actually do occur in equatorial waters.

 

The proportion and numbers of molten-coloured individuals are notable in this species compared with the other species in the Northern Hemisphere. Some whales remain white even after growing up.

 

A baleia-franca-austral (Eubalaena australis) é uma das três espécies de baleia-franca, pertencente ao género Eubalaena. Estima-se que haja cerca de 7500 exemplares desta baleia espalhadas pelo sul do Hemisfério Sul, numa faixa compreendida entre os 30º e os 55º de latitude. Pode atingir os 18 metros de comprimento e as 80 toneladas de peso.

 

A caça indiscriminada deste tipo de baleia, devido à quantidade de óleo possuída por exemplar, deixou-a quase em perigo de extinção. Desde o século XIX, a população destes animais foi reduzida em 90%. Actualmente estima-se que exista uma população que oscila entre os 7500 e 8000 indivíduos. Durante o inverno, as baleias escolhem as águas mais quentes do hemisfério sul para se reproduzirem, tais como os seguintes as costas da Península Valdés (na Patagónia), Austrália, África do Sul e Brasil.

Canon A1 35mm on Ilford HP5 400

 

#A #TheCapitolBuilding #WashingtonDC #cpac #Dildo #CecilBDeMille #TheTenCommandments #TrumpIncitesViolence #DomesticTerrorism #Terrorist #Coup #Sedition #HeadTerrorist #Riot #CapitolPolice #Curfew #TakeOver #ElectoralVote #HornedTerrorist #JakeAngeli #QAnon #Facebook #Instagram #Twitter #Shopify #blocked #banned #RudyGiuliani #Gold #Coins #HairDye #Farts #Fart #AirFreshener #HorribleSmell #Flatulence #TheVapors #Borat #AffordableHealthcareAct. #Obamacare #Masturbation #Twitter #TemperTantrum #BarackObama #statue #ovaloffice #disinfectant #bleach #chlorox #lysol #chlorinebleach #101 #wipes #DrTrump #Medusa #ClashoftheTitans #Snake #Rattlesnake #Slimeball #ForkedTongue #scales #hairdye #eastereggdye #PAAS #loreal #missclairol #DonaldTrump #narcissisticpersonalitydisorder #JaredKushner #DonaldTrumpJr #COVID19 #Coronavirus #Pandemic #DrAnthonySFauci #MikePence #testicle #penis #dick #balls #AsktheMortician #disease #pneumonia #infection #flu #JimmyKimmel #JimmyFallon #StephenColbert #SethMyer #minilogue #quarantine #sociadistancing #6feet #MitchMcConnell #handsanitizer #novelcoronavirus #trumplies #wipes #stayathome #workfromhome #homeschooling #deathtoll #StevenMnuchin #MikePompeo #testspositive #testsnegative #farright #freaks #ventilator #medicalmask #hospitalbed #temporarymorgue #WuhanChina, #novelcoronavirus #sociallydistant #selfquarantine #SeanHannity #FakeNews #infectiousdisease #FoxNews #epidemic #publichealthemergency #CDC #AndrewCuomo #CenterForDiseaseControl #CNN #flatteningthecurve #worldhealthorganization #recession #depression #contagious #WHO #narcissisticpersonalitydisorder #positive #negative #coronavirus #pandemic #COVID19 #infection #Dr.AnthonyFauci #trumpcausescancer #windmill #windpower #windturbine #antivaxxer #bigfoot #UFO #lochnessmonster #flyingsaucer #mythology #atlantis #bermudatriangle #flatearth #wikileaks #angels #fallenangels #45 #BiggestBaby #dirtydiaper #TheMuellerInvestigation #WilliamBarr #AttorneyGeneral #Report #TheEmperorsNewClothes #TheExorcist #Pazuzu #God #JesusChrist #LindaBlair #possessed #KingofDemons #SarahSanders #hypocrite #Exorcism #Pampers #TheWall #NeverGiveIn #Bullshit #dirtydiaper #diaperrash #PooPooBaDoop #Clemson #Wendys #burgerking #football #KellyanneConway #SarahHuckabee #Trolls #Kremlin #KremlinTroll #RussianTroll #RobertMueller #FISA #SteveBannon #MikePence #Twitter #Tweet #wiretap #Twit #wiretapped #Twat #dontaldtrump #WashingtonDC #MamaAyeshas #wallofpresidents #CIA #GOP #KKK #ISIS #FBI #BLM #LGBT #Russia #VladimirPutin #Russianinterference #AlternativeFacts #MicrowaveOven #Camera #sexdrugsandrockandroll #HillaryClinton #BernieSanders #BarackObama #PresidentoftheUnited #plannedparenthood #bigot #jihad #OsamabinLaden #DumpTrump #NotMyPresident #Dontee #DonteesInferno #thewalkingdead #republican #pedophile #WomensMarch #badhombre #conservative #rape #RiencePriebus #DonaldMcGahn #FrankGaffney #JeffSessions #GeneralJamesMattis #GeneralJohnKelly #StevenMnuchin #AndyPuzder #WilburRoss #CathyMcMorrisRodgers #MitchMcConnell #KTMcFarland #MikePompeo #NikkiHaley #LtGenMichaelFlynn #BenCarson #BetsyDeVos #TomPrice #ScottPruitt #SeemaVerma #PaulRyan #TrumpTower #MarriageEquality #KuKluxKlan #NewYorkCity #Hanksy #MelaniaTrump #BarronTrump #IvankaTrump #TiffanyTrump #EricTrump #DonaldTrumpJr #JaredKushner #conflictofinterest #emolument #RiggedElection #TemperTantrum #Tweet #Twitter #Twit #ManChild #DiaperBlowout #Trump #poop #turd #bigbaby #manindiapers #Inauguration #ScottBaio #TedNugent #TheRockettes #RadioCityMusicHall #MormonTabernacleChoir #Medusa #breitbart #lies #NationalEnquirer #douchebag #POS #Pussy #PussyGrabber

#terrorist #Taliban #jihad #MexicanWall #racism #jihad #nobannowall #confederateflag #Nazi #Islam #Freedom #AmericanNaziParty #TheRollingStones #Democrat #CivilRights #Idiot #abortion #tinfoilhatsociety #tyrant #foxnews #MerylStreep #Liberal #SaturdayNightLive #AlecBaldwin #MelissaMcCarthy #AdolfHitler #BenitoMussolini #Dictator #Megalomaniac #KingComplex #Demagogue #Narcissist #Delusional #Nuts #Oligarch #Populist #tyrant #Narcissistic #Autocracy #Oligarchy #DelusionsofGrandeur #GodComplex #MangoMussolini #DerPumpkinfuhrer #Apocalypse #NuclearButton #OvalOffice #civilliberties #goldenshowers #tinyhands #discrimination #TrumpGate #freedomandjusticeforall #TheBible #JesusChrist #The12Apostles #FredPhelps #GodHatesFags #WestboroBaptistChurch #RedNeck #ScienceFiction, #rapistsandmurderers #antiGay #homophobe #dinosaurs #religiousright #AmericanFamilyAssociation #hategroup #BruceJenner #CaitlynJenner #BarbieandKen #Mattel #PopeFrancis #QueenElizabeth #KeepYourPeckerUp #PatRobertson #BatteredWomanSyndrome #FranklinGraham #Cracker #JudyGarland #TheWizardofOz #BarbraStreisand #BettyWhite #MarilynMonroe #ValleyoftheDolls #PeytonPlace #DowntonAbbey #MaggieSmith #JudyDench #EvaGreen #MissPeregrine #DarylDixon #jabbathehutt #EmperorPalpatine #StarWars #StarTrek #RickGrimes #TeaParty #GlennBeck #RushLimbaugh #fakeNews #politicallyincorrect #BillMaher #AngelaMerkel #TheresaMay #RosieODonnell #MegynKelly #TheManchurianCandidate #BadCombOver #commemorativecoin #collectorsitem #ebay #buffalonewyork #artvoice #carlpaladino #byecarl #OutrageFatigue #hotair #weaponsofmassdestruction #motherofallbombs #farts #farting #robertmueller #bombingsyria #kellyanneconway #brettkavanaugh #sexualassault #harrassment #metoo #supremecourt #kanyewest #kimkardashian #idiot #incoherent #dumptrump2020 #rosegardenmassacre #ivankatrump #jaredKushner #Donaldtrumpjr #erictrump #Stephenmiller #mitchMcConnell #Williambarr #KellyAnneConway #KayleighMcEnany #LyingBitches #sarahhuckabeesanders #sexdrugsandrockandroll #Death #Dying #GrimReaper #AlternativeFacts #liarliarpantsonfire #masturbation #jerkoff #Disinfectant #Purell #Re-election2020 #God #Lysol #Coronavirus #SteveBannon #syphiliticskinlesions #Pandemic #DrAnthonySFauci #MikePence #Death #dying #AsktheMortician #pneumonia #infection #flu, #influenza, #quarantine #socialdistancing #6feet #HopeHicks #WilliamBarr #handsanitizer #trumpslies #wipes #narcissisticpersonalitydisorder #deathtoll #stayathome #workfromhome #homeschooling #ventilator #medicalmask #novelcoronavirus #sociallydistant #SeanHannity #FakeNews #TuckerCarlson #infectiousdisease #FoxNews #CNN #epidemic #CDC #AndrewCuomo #CenterForDiseaseControl #worldhealthorganization #recession #depression #contagious #ProudBoys #AffordableHealthcareAct. #Obamacare #Masturbation #Twitter #TemperTantrum #BarackObama #statue #ovaloffice #disinfectant #bleach #chlorox #lysol #chlorinebleach #101 #wipes #DrTrump #Medusa #ClashoftheTitans #Snake #Rattlesnake #Slimeball #ForkedTongue #scales #hairdye #eastereggdye #PAAS #loreal #missclairol #DonaldTrump #narcissisticpersonalitydisorder #JaredKushner #DonaldTrumpJr #COVID19 #Coronavirus #Pandemic #DrAnthonySFauci #MikePence #testicle #penis #dick #balls #AsktheMortician #disease #pneumonia #infection #flu #JimmyKimmel #JimmyFallon #StephenColbert #SethMyer #minilogue #quarantine #sociadistancing #6feet #MitchMcConnell #handsanitizer #novelcoronavirus #trumplies #wipes #stayathome #workfromhome #homeschooling #deathtoll #StevenMnuchin #MikePompeo #testspositive #testsnegative #farright #freaks #ventilator #medicalmask #hospitalbed #temporarymorgue #WuhanChina, #novelcoronavirus #sociallydistant #selfquarantine #SeanHannity #FakeNews #infectiousdisease #FoxNews #epidemic #publichealthemergency #CDC #AndrewCuomo #CenterForDiseaseControl #CNN #flatteningthecurve #worldhealthorganization #recession #depression #contagious #WHO #narcissisticpersonalitydisorder #positive #negative #coronavirus #pandemic #COVID19 #infection #Dr.AnthonyFauci #trumpcausescancer #windmill #windpower #windturbine #antivaxxer #bigfoot #UFO #lochnessmonster #flyingsaucer #mythology #atlantis #bermudatriangle #flatearth #wikileaks #angels #fallenangels #45 #BiggestBaby #dirtydiaper #TheMuellerInvestigation #WilliamBarr #AttorneyGeneral #Report #TheEmperorsNewClothes #TheExorcist #Pazuzu #God #JesusChrist #LindaBlair #possessed #KingofDemons #SarahSanders #hypocrite #Exorcism #Pampers #TheWall #NeverGiveIn #Bullshit #dirtydiaper #diaperrash #PooPooBaDoop #Clemson #Wendys #burgerking #football #KellyanneConway #SarahHuckabee #Trolls #Kremlin #KremlinTroll #RussianTroll #RobertMueller #FISA #SteveBannon #MikePence #Twitter #Tweet #wiretap #Twit #wiretapped #Twat #dontaldtrump #WashingtonDC #MamaAyeshas #wallofpresidents #CIA #GOP #KKK #ISIS #FBI #BLM #LGBT #Russia #VladimirPutin #Russianinterference #AlternativeFacts #MicrowaveOven #Camera #sexdrugsandrockandroll #HillaryClinton #BernieSanders #BarackObama #PresidentoftheUnited #plannedparenthood #bigot #jihad #OsamabinLaden #DumpTrump #NotMyPresident #Dontee #DonteesInferno #thewalkingdead #republican #pedophile #WomensMarch #badhombre #conservative #rape #RiencePriebus #DonaldMcGahn #FrankGaffney #JeffSessions #GeneralJamesMattis #GeneralJohnKelly #StevenMnuchin #AndyPuzder #WilburRoss #CathyMcMorrisRodgers #MitchMcConnell #KTMcFarland #MikePompeo #NikkiHaley #LtGenMichaelFlynn #BenCarson #BetsyDeVos #TomPrice #ScottPruitt #SeemaVerma #PaulRyan #TrumpTower #MarriageEquality #KuKluxKlan #NewYorkCity #Hanksy #MelaniaTrump #BarronTrump #IvankaTrump #TiffanyTrump #EricTrump #DonaldTrumpJr #JaredKushner #conflictofinterest #emolument #RiggedElection #TemperTantrum #Tweet #Twitter #Twit #ManChild #DiaperBlowout #Trump #poop #turd #bigbaby #manindiapers #Inauguration #ScottBaio #TedNugent #TheRockettes #RadioCityMusicHall #MormonTabernacleChoir #Medusa #breitbart #lies #NationalEnquirer #douchebag #POS #Pussy #PussyGrabber

A #terrorist #Taliban #jihad #MexicanWall #racism #jihad #nobannowall #confederateflag #Nazi #Islam #Freedom #AmericanNaziParty #TheRollingStones #Democrat #CivilRights #Idiot #abortion #tinfoilhatsociety #tyrant #foxnews #MerylStreep #Liberal #SaturdayNightLive #AlecBaldwin #MelissaMcCarthy #AdolfHitler #BenitoMussolini #Dictator #Megalomaniac #KingComplex #Demagogue #Narcissist #Delusional #Nuts #Oligarch #Populist #tyrant #Narcissistic #Autocracy #Oligarchy #DelusionsofGrandeur #GodComplex #MangoMussolini #DerPumpkinfuhrer #Apocalypse #NuclearButton #OvalOffice #civilliberties #goldenshowers #tinyhands #discrimination #TrumpGate #freedomandjusticeforall #TheBible #JesusChrist #The12Apostles #FredPhelps #GodHatesFags #WestboroBaptistChurch #RedNeck #ScienceFiction, #rapistsandmurderers #antiGay #homophobe #dinosaurs #religiousright #AmericanFamilyAssociation #hategroup #BruceJenner #CaitlynJenner #BarbieandKen #Mattel #PopeFrancis #QueenElizabeth #KeepYourPeckerUp #PatRobertson #BatteredWomanSyndrome #FranklinGraham #Cracker #JudyGarland #TheWizardofOz #BarbraStreisand #BettyWhite #MarilynMonroe #ValleyoftheDolls #PeytonPlace #DowntonAbbey #MaggieSmith #JudyDench #EvaGreen #MissPeregrine #DarylDixon #jabbathehutt #EmperorPalpatine #StarWars #StarTrek #RickGrimes #TeaParty #GlennBeck #RushLimbaugh #fakeNews #politicallyincorrect #BillMaher #AngelaMerkel #TheresaMay #RosieODonnell #MegynKelly #TheManchurianCandidate #BadCombOver #commemorativecoin #collectorsitem #ebay #buffalonewyork #artvoice #carlpaladino #byecarl #OutrageFatigue #hotair #weaponsofmassdestruction #motherofallbombs #farts #farting #robertmueller #bombingsyria #kellyanneconway #brettkavanaugh #sexualassault #harrassment #metoo #supremecourt #kanyewest #kimkardashian #idiot #incoherent #dumptrump2020 #rosegardenmassacre #ivankatrump #jaredKushner #Donaldtrumpjr #erictrump #Stephenmiller #mitchMcConnell #Williambarr #KellyAnneConway #KayleighMcEnany #LyingBitches #sarahhuckabeesanders #sexdrugsandrockandroll #Death #Dying #GrimReaper #AlternativeFacts #liarliarpantsonfire #masturbation #jerkoff #Disinfectant #Purell #Re-election2020 #God #Lysol #Coronavirus #SteveBannon #syphiliticskinlesions #Pandemic #DrAnthonySFauci #MikePence #Death #dying #AsktheMortician #pneumonia #infection #flu, #influenza, #quarantine #socialdistancing #6feet #HopeHicks #WilliamBarr #handsanitizer #trumpslies #wipes #narcissisticpersonalitydisorder #deathtoll #stayathome #workfromhome #homeschooling #ventilator #medicalmask #novelcoronavirus #sociallydistant #SeanHannity #FakeNews #TuckerCarlson #infectiousdisease #FoxNews #CNN #epidemic #CDC #AndrewCuomo #CenterForDiseaseControl #worldhealthorganization #recession #depression #contagious #ProudBoys

You may view more of my Orchids, by clicking here !

 

Orchidaceae is a diverse and widespread family of flowering plants with blooms that are often colourful and often fragrant, commonly known as the orchid family. Along with the Asteraceae, they are one of the two largest families of flowering plants, with between 21,950 and 26,049 currently accepted species, found in 880 genera. Selecting which of the two families is larger is still under debate, as concrete numbers on such enormous families are constantly in flux. Regardless, the number of orchid species equals more than twice the number of bird species, and about four times the number of mammal species. The family also encompasses about 6â11% of all seed plants.The largest genera are Bulbophyllum (2,000 species), Epidendrum (1,500 species), Dendrobium (1,400 species) and Pleurothallis (1,000 species). The family also includes Vanilla (the genus of the vanilla plant), Orchis (type genus), and many commonly cultivated plants such as Phalaenopsis and Cattleya. Moreover, since the introduction of tropical species in the 19th century, horticulturists have produced more than 100,000 hybrids and cultivars. The name comes from the Ancient Greek á½ÏÏÎ¹Ï (órkhis), literally meaning "testicle", because of the shape of the root. Carl Linnaeus classified the family as Orchidaceae. Orchid was introduced in 1845 by John Lindley in School Botany, due to an incorrect attempt to extract the Latin stem (orchis) from Orchidaceae. The Greek myth of Orchis explains the origin of the plants. Orchis, the son of a nymph and a satyr, came upon a festival of Dionysus (Bacchus) in the forest. He drank too much, and attempted to rape a priestess of Dionysus. For his insult, he was torn apart by the Bacchanalians. His father prayed for him to be restored, but the gods instead changed him into a flower.

These flowers were previously called Orchis, Satyrion (Satyrion feminina), or "ballockwort".

 

From Wikipedia, the free encyclopedia

Seen around the Pushkar fair.

#a #A #AffordableHealthcareAct. #Obamacare #Masturbation #Twitter #TemperTantrum #BarackObama #statue #ovaloffice #disinfectant #bleach #chlorox #lysol #chlorinebleach #101 #wipes #DrTrump #Medusa #ClashoftheTitans #Snake #Rattlesnake #Slimeball #ForkedTongue #scales #hairdye #eastereggdye #PAAS #loreal #missclairol #DonaldTrump #narcissisticpersonalitydisorder #JaredKushner #DonaldTrumpJr #COVID19 #Coronavirus #Pandemic #DrAnthonySFauci #MikePence #testicle #penis #dick #balls #AsktheMortician #disease #pneumonia #infection #flu #JimmyKimmel #JimmyFallon #StephenColbert #SethMyer #minilogue #quarantine #sociadistancing #6feet #MitchMcConnell #handsanitizer #novelcoronavirus #trumplies #wipes #stayathome #workfromhome #homeschooling #deathtoll #StevenMnuchin #MikePompeo #testspositive #testsnegative #farright #freaks #ventilator #medicalmask #hospitalbed #temporarymorgue #WuhanChina, #novelcoronavirus #sociallydistant #selfquarantine #SeanHannity #FakeNews #infectiousdisease #FoxNews #epidemic #publichealthemergency #CDC #AndrewCuomo #CenterForDiseaseControl #CNN #flatteningthecurve #worldhealthorganization #recession #depression #contagious #WHO #narcissisticpersonalitydisorder #positive #negative #coronavirus #pandemic #COVID19 #infection #Dr.AnthonyFauci #trumpcausescancer #windmill #windpower #windturbine #antivaxxer #bigfoot #UFO #lochnessmonster #flyingsaucer #mythology #atlantis #bermudatriangle #flatearth #wikileaks #angels #fallenangels #45 #BiggestBaby #dirtydiaper #TheMuellerInvestigation #WilliamBarr #AttorneyGeneral #Report #TheEmperorsNewClothes #TheExorcist #Pazuzu #God #JesusChrist #LindaBlair #possessed #KingofDemons #SarahSanders #hypocrite #Exorcism #Pampers #TheWall #NeverGiveIn #Bullshit #dirtydiaper #diaperrash #PooPooBaDoop #Clemson #Wendys #burgerking #football #KellyanneConway #SarahHuckabee #Trolls #Kremlin #KremlinTroll #RussianTroll #RobertMueller #FISA #SteveBannon #MikePence #Twitter #Tweet #wiretap #Twit #wiretapped #Twat #dontaldtrump #WashingtonDC #MamaAyeshas #wallofpresidents #CIA #GOP #KKK #ISIS #FBI #BLM #LGBT #Russia #VladimirPutin #Russianinterference #AlternativeFacts #MicrowaveOven #Camera #sexdrugsandrockandroll #HillaryClinton #BernieSanders #BarackObama #PresidentoftheUnited #plannedparenthood #bigot #jihad #OsamabinLaden #DumpTrump #NotMyPresident #Dontee #DonteesInferno #thewalkingdead #republican #pedophile #WomensMarch #badhombre #conservative #rape #RiencePriebus #DonaldMcGahn #FrankGaffney #JeffSessions #GeneralJamesMattis #GeneralJohnKelly #StevenMnuchin #AndyPuzder #WilburRoss #CathyMcMorrisRodgers #MitchMcConnell #KTMcFarland #MikePompeo #NikkiHaley #LtGenMichaelFlynn #BenCarson #BetsyDeVos #TomPrice #ScottPruitt #SeemaVerma #PaulRyan #TrumpTower #MarriageEquality #KuKluxKlan #NewYorkCity #Hanksy #MelaniaTrump #BarronTrump #IvankaTrump #TiffanyTrump #EricTrump #DonaldTrumpJr #JaredKushner #conflictofinterest #emolument #RiggedElection #TemperTantrum #Tweet #Twitter #Twit #ManChild #DiaperBlowout #Trump #poop #turd #bigbaby #manindiapers #Inauguration #ScottBaio #TedNugent #TheRockettes #RadioCityMusicHall #MormonTabernacleChoir #Medusa #breitbart #lies #NationalEnquirer #douchebag #POS #Pussy #PussyGrabber

#terrorist #Taliban #jihad #MexicanWall #racism #jihad #nobannowall #confederateflag #Nazi #Islam #Freedom #AmericanNaziParty #TheRollingStones #Democrat #CivilRights #Idiot #abortion #tinfoilhatsociety #tyrant #foxnews #MerylStreep #Liberal #SaturdayNightLive #AlecBaldwin #MelissaMcCarthy #AdolfHitler #BenitoMussolini #Dictator #Megalomaniac #KingComplex #Demagogue #Narcissist #Delusional #Nuts #Oligarch #Populist #tyrant #Narcissistic #Autocracy #Oligarchy #DelusionsofGrandeur #GodComplex #MangoMussolini #DerPumpkinfuhrer #Apocalypse #NuclearButton #OvalOffice #civilliberties #goldenshowers #tinyhands #discrimination #TrumpGate #freedomandjusticeforall #TheBible #JesusChrist #The12Apostles #FredPhelps #GodHatesFags #WestboroBaptistChurch #RedNeck #ScienceFiction, #rapistsandmurderers #antiGay #homophobe #dinosaurs #religiousright #AmericanFamilyAssociation #hategroup #BruceJenner #CaitlynJenner #BarbieandKen #Mattel #PopeFrancis #QueenElizabeth #KeepYourPeckerUp #PatRobertson #BatteredWomanSyndrome #FranklinGraham #Cracker #JudyGarland #TheWizardofOz #BarbraStreisand #BettyWhite #MarilynMonroe #ValleyoftheDolls #PeytonPlace #DowntonAbbey #MaggieSmith #JudyDench #EvaGreen #MissPeregrine #DarylDixon #jabbathehutt #EmperorPalpatine #StarWars #StarTrek #RickGrimes #TeaParty #GlennBeck #RushLimbaugh #fakeNews #politicallyincorrect #BillMaher #AngelaMerkel #TheresaMay #RosieODonnell #MegynKelly #TheManchurianCandidate #BadCombOver #commemorativecoin #collectorsitem #ebay #buffalonewyork #artvoice #carlpaladino #byecarl #OutrageFatigue #hotair #weaponsofmassdestruction #motherofallbombs #farts #farting #robertmueller #bombingsyria #kellyanneconway #brettkavanaugh #sexualassault #harrassment #metoo #supremecourt #kanyewest #kimkardashian #idiot #incoherent #dumptrump2020 #rosegardenmassacre #ivankatrump #jaredKushner #Donaldtrumpjr #erictrump #Stephenmiller #mitchMcConnell #Williambarr #KellyAnneConway #KayleighMcEnany #LyingBitches #sarahhuckabeesanders # sexdrugsandrockandroll #Death #Dying #GrimReaper #AlternativeFacts #liarliarpantsonfire #masturbation #jerkoff #Disinfectant #Purell #Re-election2020 #God #Lysol #Coronavirus #SteveBannon #syphiliticskinlesions #Pandemic #DrAnthonySFauci #MikePence #Death #dying #AsktheMortician #pneumonia #infection #flu, #influenza, #quarantine #socialdistancing #6feet #HopeHicks #WilliamBarr #handsanitizer #trumpslies #wipes #narcissisticpersonalitydisorder #deathtoll #stayathome #workfromhome #homeschooling #ventilator #medicalmask #novelcoronavirus #sociallydistant #SeanHannity #FakeNews #TuckerCarlson #infectiousdisease #FoxNews #CNN #epidemic #CDC #AndrewCuomo #CenterForDiseaseControl #worldhealthorganization #recession #depression #contagious

Guess who gets excited about Brussels sprouts? Personally, I was pretty keyed up too. Being a city boy, I had no idea they came like this. I thought you dug 'em up like turnips or something..

Voacanga globosa Testicle tree

_1270226.JPG BF

 

Testosterone

The chemical structure of testosterone.

A ball-and-stick model of testosterone.

Names

IUPAC name

17β-Hydroxyandrost-4-en-3-one

Systematic IUPAC name

(8R,9S,10R,13S,14S,17S)-17-Hydroxy-10,13-dimethyl-1,2,6,7,8,9,11,12,14,15,16,17-dodecahydrocyclopenta[a]phenanthren-3-one

Other names

Androst-4-en-17β-ol-3-one

Identifiers

CAS Number

58-22-0 ☑

3D model (JSmol)

Interactive image

ChEBI

CHEBI:17347 ☑

ChEMBL

ChEMBL386630 ☑

ChemSpider

5791 ☑

DrugBank

DB00624 ☑

ECHA InfoCard100.000.336

KEGG

D00075 ☑

PubChem CID

6013

UNII

3XMK78S47O ☑

InChI[show]

SMILES[show]

Properties

Chemical formula

C19H28O2

Molar mass288.431 g·mol−1

Melting point155 °C

Pharmacology

ATC code

G03BA03 (WHO)

License data

EU EMA: by INN

Routes of

administration

Transdermal (gel, cream, solution, patch), by mouth (as testosterone undecanoate), in the cheek, intranasal (gel), intramuscular injection (as esters), subcutaneous pellets

Pharmacokinetics:

Bioavailability

Oral: very low (due to extensive first pass metabolism)

Protein binding

97.0–99.5% (to SHBG and albumin)[1]

Metabolism

Liver (mainly reduction and conjugation)

Biological half-life

2–4 hours[citation needed]

Excretion

Urine (90%), feces (6%)

Except where otherwise noted, data are given for materials in their standard state (at 25 °C [77 °F], 100 kPa).

☑ verify (what is ☑☒ ?)

Infobox references

Testosterone is the primary male sex hormone and an anabolic steroid. In male humans, testosterone plays a key role in the development of male reproductive tissues such as testes and prostate, as well as promoting secondary sexual characteristics such as increased muscle and bone mass, and the growth of body hair.[2] In addition, testosterone is involved in health and well-being,[3] and the prevention of osteoporosis.[4] Insufficient levels of testosterone in men may lead to abnormalities including frailty and bone loss.

 

Testosterone is a steroid from the androstane class containing a keto and hydroxyl groups at the three and seventeen positions respectively. It is biosynthesized in several steps from cholesterol and is converted in the liver to inactive metabolites.[5] It exerts its action through binding to and activation of the androgen receptor.[5] In humans and most other vertebrates, testosterone is secreted primarily by the testicles of males and, to a lesser extent, the ovaries of females. On average, in adult males, levels of testosterone are about 7 to 8 times as great as in adult females.[6] As the metabolism of testosterone in males is more pronounced, the daily production is about 20 times greater in men.[7][8] Females are also more sensitive to the hormone.[9]

 

In addition to its role as a natural hormone, testosterone is used as a medication, for instance in the treatment of low testosterone levels in men and breast cancer in women.[10] Since testosterone levels decrease as men age, testosterone is sometimes used in older men to counteract this deficiency. It is also used illicitly to enhance physique and performance, for instance in athletes.

  

Contents

1Biological effects

1.1Before birth

1.2Early infancy

1.3Before puberty

1.4Pubertal

1.5Adult

1.6Aggression and criminality

1.7Brain

2Medical use

3Biological activity

3.1Steroid hormone activity

3.2Neurosteroid activity

4Biochemistry

4.1Biosynthesis

4.2Distribution

4.3Metabolism

4.4Levels

5Measurement

6History

7Other animals

8See also

9References

10Further reading

Biological effects[edit]

In general, androgens such as testosterone promote protein synthesis and thus growth of tissues with androgen receptors.[11] Testosterone can be described as having virilising and anabolic effects (though these categorical descriptions are somewhat arbitrary, as there is a great deal of mutual overlap between them).[12]

 

Anabolic effects include growth of muscle mass and strength, increased bone density and strength, and stimulation of linear growth and bone maturation.

Androgenic effects include maturation of the sex organs, particularly the penis and the formation of the scrotum in the fetus, and after birth (usually at puberty) a deepening of the voice, growth of facial hair (such as the beard) and axillary (underarm) hair. Many of these fall into the category of male secondary sex characteristics.

Testosterone effects can also be classified by the age of usual occurrence. For postnatal effects in both males and females, these are mostly dependent on the levels and duration of circulating free testosterone.

 

Before birth[edit]

Effects before birth are divided into two categories, classified in relation to the stages of development.

 

The first period occurs between 4 and 6 weeks of the gestation. Examples include genital virilisation such as midline fusion, phallic urethra, scrotal thinning and rugation, and phallic enlargement; although the role of testosterone is far smaller than that of dihydrotestosterone. There is also development of the prostate gland and seminal vesicles.

 

During the second trimester, androgen level is associated with sex formation.[13] This period affects the femininization or masculinization of the fetus and can be a better predictor of feminine or masculine behaviours such as sex typed behaviour than an adult's own levels. A mother's testosterone level during pregnancy is correlated with her daughter's sex-typical behavior as an adult, and the correlation is even stronger than with the daughter's own adult testosterone level.[14]

 

Early infancy[edit]

Early infancy androgen effects are the least understood. In the first weeks of life for male infants, testosterone levels rise. The levels remain in a pubertal range for a few months, but usually reach the barely detectable levels of childhood by 4–7 months of age.[15][16] The function of this rise in humans is unknown. It has been theorized that brain masculinization is occurring since no significant changes have been identified in other parts of the body.[17] The male brain is masculinized by the aromatization of testosterone into estrogen, which crosses the blood–brain barrier and enters the male brain, whereas female fetuses have α-fetoprotein, which binds the estrogen so that female brains are not affected.[18]

 

Before puberty[edit]

Before puberty effects of rising androgen levels occur in both boys and girls. These include adult-type body odor, increased oiliness of skin and hair, acne, pubarche (appearance of pubic hair), axillary hair (armpit hair), growth spurt, accelerated bone maturation, and facial hair.[19]

 

Pubertal[edit]

Pubertal effects begin to occur when androgen has been higher than normal adult female levels for months or years. In males, these are usual late pubertal effects, and occur in women after prolonged periods of heightened levels of free testosterone in the blood. The effects include:[19][20]

 

Growth of spermatogenic tissue in testicles, male fertility, penis or clitoris enlargement, increased libido and frequency of erection or clitoral engorgement occurs. Growth of jaw, brow, chin, and nose and remodeling of facial bone contours, in conjunction with human growth hormone occurs.[21] Completion of bone maturation and termination of growth. This occurs indirectly via estradiol metabolites and hence more gradually in men than women. Increased muscle strength and mass, shoulders become broader and rib cage expands, deepening of voice, growth of the Adam's apple. Enlargement of sebaceous glands. This might cause acne, subcutaneous fat in face decreases. Pubic hair extends to thighs and up toward umbilicus, development of facial hair (sideburns, beard, moustache), loss of scalp hair (androgenetic alopecia), increase in chest hair, periareolar hair, perianal hair, leg hair, armpit hair.

 

Adult[edit]

Testosterone is necessary for normal sperm development. It activates genes in Sertoli cells, which promote differentiation of spermatogonia. It regulates acute HPA (hypothalamic–pituitary–adrenal axis) response under dominance challenge.[22] Androgen including testosterone enhances muscle growth. Testosterone also regulates the population of thromboxane A2 receptors on megakaryocytes and platelets and hence platelet aggregation in humans.[23][24]

 

Adult testosterone effects are more clearly demonstrable in males than in females, but are likely important to both sexes. Some of these effects may decline as testosterone levels might decrease in the later decades of adult life.[25]

 

Health risks[edit]

Testosterone does not appear to increase the risk of developing prostate cancer. In people who have undergone testosterone deprivation therapy, testosterone increases beyond the castrate level have been shown to increase the rate of spread of an existing prostate cancer.[26][27][28]

 

Conflicting results have been obtained concerning the importance of testosterone in maintaining cardiovascular health.[29][30] Nevertheless, maintaining normal testosterone levels in elderly men has been shown to improve many parameters that are thought to reduce cardiovascular disease risk, such as increased lean body mass, decreased visceral fat mass, decreased total cholesterol, and glycemic control.[31]

 

High androgen levels are associated with menstrual cycle irregularities in both clinical populations and healthy women.[32]

 

Sexual arousal[edit]

See also: Hormones and sexual arousal

When testosterone and endorphins in ejaculated semen meet the cervical wall after sexual intercourse, females receive a spike in testosterone, endorphin, and oxytocin levels, and males after orgasm during copulation experience an increase in endorphins and a marked increase in oxytocin levels. This adds to the hospitable physiological environment in the female internal reproductive tract for conceiving, and later for nurturing the conceptus in the pre-embryonic stages, and stimulates feelings of love, desire, and paternal care in the male (this is the only time male oxytocin levels rival a female's).[citation needed]

 

Testosterone levels follow a nyctohemeral rhythm that peaks early each day, regardless of sexual activity.[33]

 

There are positive correlations between positive orgasm experience in women and testosterone levels where relaxation was a key perception of the experience. There is no correlation between testosterone and men's perceptions of their orgasm experience, and also no correlation between higher testosterone levels and greater sexual assertiveness in either sex.[34]

 

Sexual arousal and masturbation in women produce small increases in testosterone concentrations.[35] The plasma levels of various steroids significantly increase after masturbation in men and the testosterone levels correlate to those levels.[36]

 

Mammalian studies[edit]

Studies conducted in rats have indicated that their degree of sexual arousal is sensitive to reductions in testosterone. When testosterone-deprived rats were given medium levels of testosterone, their sexual behaviors (copulation, partner preference, etc.) resumed, but not when given low amounts of the same hormone. Therefore, these mammals may provide a model for studying clinical populations among humans suffering from sexual arousal deficits such as hypoactive sexual desire disorder.[37]

 

In every mammalian species examined demonstrated a marked increase in a male's testosterone level upon encountering a novel female. The reflexive testosterone increases in male mice is related to the male's initial level of sexual arousal.[38]

 

In non-human primates, it may be that testosterone in puberty stimulates sexual arousal, which allows the primate to increasingly seek out sexual experiences with females and thus creates a sexual preference for females.[39] Some research has also indicated that if testosterone is eliminated in an adult male human or other adult male primate's system, its sexual motivation decreases, but there is no corresponding decrease in ability to engage in sexual activity (mounting, ejaculating, etc.).[39]

 

In accordance with sperm competition theory, testosterone levels are shown to increase as a response to previously neutral stimuli when conditioned to become sexual in male rats.[40] This reaction engages penile reflexes (such as erection and ejaculation) that aid in sperm competition when more than one male is present in mating encounters, allowing for more production of successful sperm and a higher chance of reproduction.

 

Males[edit]

In men, higher levels of testosterone are associated with periods of sexual activity.[41][42]

 

Men who watch a sexually explicit movie have an average increase of 35% in testosterone, peaking at 60–90 minutes after the end of the film, but no increase is seen in men who watch sexually neutral films.[43] Men who watch sexually explicit films also report increased motivation, competitiveness, and decreased exhaustion.[44] A link has also been found between relaxation following sexual arousal and testosterone levels.[45]

 

Men's levels of testosterone, a hormone known to affect men's mating behaviour, changes depending on whether they are exposed to an ovulating or nonovulating woman's body odour. Men who are exposed to scents of ovulating women maintained a stable testosterone level that was higher than the testosterone level of men exposed to nonovulation cues. Testosterone levels and sexual arousal in men are heavily aware of hormone cycles in females.[46] This may be linked to the ovulatory shift hypothesis,[47] where males are adapted to respond to the ovulation cycles of females by sensing when they are most fertile and whereby females look for preferred male mates when they are the most fertile; both actions may be driven by hormones.

 

Females[edit]

Androgens may modulate the physiology of vaginal tissue and contribute to female genital sexual arousal.[48] Women's level of testosterone is higher when measured pre-intercourse vs pre-cuddling, as well as post-intercourse vs post-cuddling.[49] There is a time lag effect when testosterone is administered, on genital arousal in women. In addition, a continuous increase in vaginal sexual arousal may result in higher genital sensations and sexual appetitive behaviors.[50]

 

When females have a higher baseline level of testosterone, they have higher increases in sexual arousal levels but smaller increases in testosterone, indicating a ceiling effect on testosterone levels in females. Sexual thoughts also change the level of testosterone but not level of cortisol in the female body, and hormonal contraceptives may affect the variation in testosterone response to sexual thoughts.[51]

 

Testosterone may prove to be an effective treatment in female sexual arousal disorders,[52] and is available as a dermal patch. There is no FDA approved androgen preparation for the treatment of androgen insufficiency; however, it has been used off-label to treat low libido and sexual dysfunction in older women. Testosterone may be a treatment for postmenopausal women as long as they are effectively estrogenized.[52]

 

Romantic relationships[edit]

Falling in love decreases men's testosterone levels while increasing women's testosterone levels. There has been speculation that these changes in testosterone result in the temporary reduction of differences in behavior between the sexes.[53] However, it is suggested that after the "honeymoon phase" ends—about four years into a relationship—this change in testosterone levels is no longer apparent.[53] Men who produce less testosterone are more likely to be in a relationship[54] or married,[55] and men who produce more testosterone are more likely to divorce;[55] however, causality cannot be determined in this correlation. Marriage or commitment could cause a decrease in testosterone levels.[56] Single men who have not had relationship experience have lower testosterone levels than single men with experience. It is suggested that these single men with prior experience are in a more competitive state than their non-experienced counterparts.[57] Married men who engage in bond-maintenance activities such as spending the day with their spouse/and or child have no different testosterone levels compared to times when they do not engage in such activities. Collectively, these results suggest that the presence of competitive activities rather than bond-maintenance activities are more relevant to changes in testosterone levels.[58]

 

Men who produce more testosterone are more likely to engage in extramarital sex.[55] Testosterone levels do not rely on physical presence of a partner; testosterone levels of men engaging in same-city and long-distance relationships are similar.[54] Physical presence may be required for women who are in relationships for the testosterone–partner interaction, where same-city partnered women have lower testosterone levels than long-distance partnered women.[59]

 

Fatherhood[edit]

Fatherhood decreases testosterone levels in men, suggesting that the emotions and behavior tied to decreased testosterone promote paternal care. In humans and other species that utilize allomaternal care, paternal investment in offspring is beneficial to said offspring's survival because it allows the parental dyad to raise multiple children simultaneously. This increases the reproductive fitness of the parents, because their offspring are more likely to survive and reproduce. Paternal care increases offspring survival due to increased access to higher quality food and reduced physical and immunological threats.[60] This is particularly beneficial for humans since offspring are dependent on parents for extended periods of time and mothers have relatively short inter-birth intervals.[61] While extent of paternal care varies between cultures, higher investment in direct child care has been seen to be correlated with lower average testosterone levels as well as temporary fluctuations.[62] For instance, fluctuation in testosterone levels when a child is in distress has been found to be indicative of fathering styles. If a father's testosterone levels decrease in response to hearing their baby cry, it is an indication of empathizing with the baby. This is associated with increased nurturing behavior and better outcomes for the infant.[63]

 

Motivation[edit]

Testosterone levels play a major role in risk-taking during financial decisions.[64][65]

 

Aggression and criminality [edit]

See also: Aggression § Testosterone, and Biosocial criminology

Most studies support a link between adult criminality and testosterone, although the relationship is modest if examined separately for each sex. Nearly all studies of juvenile delinquency and testosterone are not significant. Most studies have also found testosterone to be associated with behaviors or personality traits linked with criminality such as antisocial behavior and alcoholism. Many studies have also been done on the relationship between more general aggressive behavior/feelings and testosterone. About half the studies have found a relationship and about half no relationship.[66]

 

Testosterone is only one of many factors that influence aggression and the effects of previous experience and environmental stimuli have been found to correlate more strongly. A few studies indicate that the testosterone derivative estradiol (one form of estrogen) might play an important role in male aggression.[66][67][68][69] Studies have also found that testosterone facilitates aggression by modulating vasopressin receptors in the hypothalamus.[70]

 

The sexual hormone can encourage fair behavior. For the study, subjects took part in a behavioral experiment where the distribution of a real amount of money was decided. The rules allowed both fair and unfair offers. The negotiating partner could subsequently accept or decline the offer. The fairer the offer, the less probable a refusal by the negotiating partner. If no agreement was reached, neither party earned anything. Test subjects with an artificially enhanced testosterone level generally made better, fairer offers than those who received placebos, thus reducing the risk of a rejection of their offer to a minimum. Two later studies have empirically confirmed these results.[71][72][73] However men with high testosterone were significantly 27% less generous in an ultimatum game.[74] The Annual NY Academy of Sciences has also found anabolic steroid use which increase testosterone to be higher in teenagers, and this was associated with increased violence.[75] Studies have also found administered testosterone to increase verbal aggression and anger in some participants.[76]

 

Testosterone is significantly correlated with aggression and competitive behaviour and is directly facilitated by the latter. There are two theories on the role of testosterone in aggression and competition.[77] The first one is the challenge hypothesis which states that testosterone would increase during puberty thus facilitating reproductive and competitive behaviour which would include aggression.[77] Thus it is the challenge of competition among males of the species that facilitates aggression and violence.[77] Studies conducted have found direct correlation between testosterone and dominance especially among the most violent criminals in prison who had the highest testosterone levels.[77] The same research also found fathers (those outside competitive environments) had the lowest testosterone levels compared to other males.[77]

 

The second theory is similar and is known as "evolutionary neuroandrogenic (ENA) theory of male aggression".[78][79] Testosterone and other androgens have evolved to masculinize a brain in order to be competitive even to the point of risking harm to the person and others. By doing so, individuals with masculinized brains as a result of pre-natal and adult life testosterone and androgens enhance their resource acquiring abilities in order to survive, attract and copulate with mates as much as possible.[78] The masculinization of the brain is not just mediated by testosterone levels at the adult stage, but also testosterone exposure in the womb as a fetus. Higher pre-natal testosterone indicated by a low digit ratio as well as adult testosterone levels increased risk of fouls or aggression among male players in a soccer game.[80] Studies have also found higher pre-natal testosterone or lower digit ratio to be correlated with higher aggression in males.[81][82][83][84][85]

 

The rise in testosterone levels during competition predicted aggression in males but not in females.[86] Subjects who interacted with hand guns and an experimental game showed rise in testosterone and aggression.[87] Natural selection might have evolved males to be more sensitive to competitive and status challenge situations and that the interacting roles of testosterone are the essential ingredient for aggressive behaviour in these situations.[88] Testosterone produces aggression by activating subcortical areas in the brain, which may also be inhibited or suppressed by social norms or familial situations while still manifesting in diverse intensities and ways through thoughts, anger, verbal aggression, competition, dominance and physical violence.[89] Testosterone mediates attraction to cruel and violent cues in men by promoting extended viewing of violent stimuli.[90] Testosterone specific structural brain characteristic can predict aggressive behaviour in individuals.[91]

 

Estradiol is known to correlate with aggression in male mice.[92] Moreover, the conversion of testosterone to estradiol regulates male aggression in sparrows during breeding season.[93] Rats who were given anabolic steroids that increase testosterone were also more physically aggressive to provocation as a result of "threat sensitivity".[94]

 

Brain[edit]

The brain is also affected by this sexual differentiation;[13] the enzyme aromatase converts testosterone into estradiol that is responsible for masculinization of the brain in male mice. In humans, masculinization of the fetal brain appears, by observation of gender preference in patients with congenital diseases of androgen formation or androgen receptor function, to be associated with functional androgen receptors.[95]

 

There are some differences between a male and female brain (possibly the result of different testosterone levels), one of them being size: the male human brain is, on average, larger.[96] Men were found to have a total myelinated fiber length of 176 000 km at the age of 20, whereas in women the total length was 149 000 km (approx. 15% less).[97]

 

No immediate short term effects on mood or behavior were found from the administration of supraphysiologic doses of testosterone for 10 weeks on 43 healthy men.[98] A correlation between testosterone and risk tolerance in career choice exists among women.[64][99]

 

Attention, memory, and spatial ability are key cognitive functions affected by testosterone in humans. Preliminary evidence suggests that low testosterone levels may be a risk factor for cognitive decline and possibly for dementia of the Alzheimer's type,[100][101][102][103] a key argument in life extension medicine for the use of testosterone in anti-aging therapies. Much of the literature, however, suggests a curvilinear or even quadratic relationship between spatial performance and circulating testosterone,[104] where both hypo- and hypersecretion (deficient- and excessive-secretion) of circulating androgens have negative effects on cognition.

 

Medical use[edit]

Main article: Testosterone (medication)

Testosterone is used as a medication for the treatment of males with too little or no natural testosterone production, certain forms of breast cancer,[10] and gender dysphoria in transgender men. This is known as hormone replacement therapy (HRT) or testosterone replacement therapy (TRT), which maintains serum testosterone levels in the normal range. Decline of testosterone production with age has led to interest in androgen replacement therapy.[105] It is unclear if the use of testosterone for low levels due to aging is beneficial or harmful.[106]

 

Testosterone is included in the World Health Organization's list of essential medicines, which are the most important medications needed in a basic health system.[107] It is available as a generic medication.[10] The price depends on the form of testosterone used.[108] It can be administered as a cream or transdermal patch that is applied to the skin, by injection into a muscle, as a tablet that is placed in the cheek, or by ingestion.[10]

 

Common side effects from testosterone medication include acne, swelling, and breast enlargement in males.[10] Serious side effects may include liver toxicity, heart disease, and behavioral changes.[10] Women and children who are exposed may develop virilization.[10] It is recommended that individuals with prostate cancer not use the medication.[10] It can cause harm if used during pregnancy or breastfeeding.[10]

 

Biological activity[edit]

Steroid hormone activity[edit]

The effects of testosterone in humans and other vertebrates occur by way of multiple mechanisms: by activation of the androgen receptor (directly or as DHT), and by conversion to estradiol and activation of certain estrogen receptors.[109][110] Androgens such as testosterone have also been found to bind to and activate membrane androgen receptors.[111][112][113]

 

Free testosterone (T) is transported into the cytoplasm of target tissue cells, where it can bind to the androgen receptor, or can be reduced to 5α-dihydrotestosterone (DHT) by the cytoplasmic enzyme 5α-reductase. DHT binds to the same androgen receptor even more strongly than testosterone, so that its androgenic potency is about 5 times that of T.[114] The T-receptor or DHT-receptor complex undergoes a structural change that allows it to move into the cell nucleus and bind directly to specific nucleotide sequences of the chromosomal DNA. The areas of binding are called hormone response elements (HREs), and influence transcriptional activity of certain genes, producing the androgen effects.

 

Androgen receptors occur in many different vertebrate body system tissues, and both males and females respond similarly to similar levels. Greatly differing amounts of testosterone prenatally, at puberty, and throughout life account for a share of biological differences between males and females.

 

The bones and the brain are two important tissues in humans where the primary effect of testosterone is by way of aromatization to estradiol. In the bones, estradiol accelerates ossification of cartilage into bone, leading to closure of the epiphyses and conclusion of growth. In the central nervous system, testosterone is aromatized to estradiol. Estradiol rather than testosterone serves as the most important feedback signal to the hypothalamus (especially affecting LH secretion).[115] In many mammals, prenatal or perinatal "masculinization" of the sexually dimorphic areas of the brain by estradiol derived from testosterone programs later male sexual behavior.[116]

 

Neurosteroid activity[edit]

Testosterone, via its active metabolite 3α-androstanediol, is a potent positive allosteric modulator of the GABAA receptor.[117]

 

Testosterone has been found to act as an antagonist of the TrkA and p75NTR, receptors for the neurotrophin nerve growth factor (NGF), with high affinity (around 5 nM).[118][119][120] In contrast to testosterone, DHEA and DHEA sulfate have been found to act as high-affinity agonists of these receptors.[118][119][120]

 

Testosterone is an antagonist of the sigma σ1 receptor (Ki = 1,014 or 201 nM).[121] However, the concentrations of testosterone required for binding the receptor are far above even total circulating concentrations of testosterone in adult males (which range between 10 and 35 nM).[122]

 

Biochemistry[edit]

 

Human steroidogenesis, showing testosterone near bottom.[123]

Biosynthesis[edit]

Like other steroid hormones, testosterone is derived from cholesterol (see figure).[124] The first step in the biosynthesis involves the oxidative cleavage of the side-chain of cholesterol by cholesterol side-chain cleavage enzyme (P450scc, CYP11A1), a mitochondrial cytochrome P450 oxidase with the loss of six carbon atoms to give pregnenolone. In the next step, two additional carbon atoms are removed by the CYP17A1 (17α-hydroxylase/17,20-lyase) enzyme in the endoplasmic reticulum to yield a variety of C19 steroids.[125] In addition, the 3β-hydroxyl group is oxidized by 3β-hydroxysteroid dehydrogenase to produce androstenedione. In the final and rate limiting step, the C17 keto group androstenedione is reduced by 17β-hydroxysteroid dehydrogenase to yield testosterone.

 

The largest amounts of testosterone (>95%) are produced by the testes in men,[2] while the adrenal glands account for most of the remainder. Testosterone is also synthesized in far smaller total quantities in women by the adrenal glands, thecal cells of the ovaries, and, during pregnancy, by the placenta.[126] In the testes, testosterone is produced by the Leydig cells.[127] The male generative glands also contain Sertoli cells, which require testosterone for spermatogenesis. Like most hormones, testosterone is supplied to target tissues in the blood where much of it is transported bound to a specific plasma protein, sex hormone-binding globulin (SHBG).

 

Regulation[edit]

 

Hypothalamic–pituitary–testicular axis

In males, testosterone is synthesized primarily in Leydig cells. The number of Leydig cells in turn is regulated by luteinizing hormone (LH) and follicle-stimulating hormone (FSH). In addition, the amount of testosterone produced by existing Leydig cells is under the control of LH, which regulates the expression of 17β-hydroxysteroid dehydrogenase.[128]

 

The amount of testosterone synthesized is regulated by the hypothalamic–pituitary–testicular axis (see figure to the right).[129] When testosterone levels are low, gonadotropin-releasing hormone (GnRH) is released by the hypothalamus, which in turn stimulates the pituitary gland to release FSH and LH. These latter two hormones stimulate the testis to synthesize testosterone. Finally, increasing levels of testosterone through a negative feedback loop act on the hypothalamus and pituitary to inhibit the release of GnRH and FSH/LH, respectively.

 

Factors affecting testosterone levels may include:

 

Age: Testosterone levels gradually reduce as men age.[130][131] This effect is sometimes referred to as andropause or late-onset hypogonadism.[132]

Exercise: Resistance training increases testosterone levels,[133] however, in older men, that increase can be avoided by protein ingestion.[134] Endurance training in men may lead to lower testosterone levels.[135]

Nutrients: Vitamin A deficiency may lead to sub-optimal plasma testosterone levels.[136] The secosteroid vitamin D in levels of 400–1000 IU/d (10–25 µg/d) raises testosterone levels.[137] Zinc deficiency lowers testosterone levels[138] but over-supplementation has no effect on serum testosterone.[139]

Weight loss: Reduction in weight may result in an increase in testosterone levels. Fat cells synthesize the enzyme aromatase, which converts testosterone, the male sex hormone, into estradiol, the female sex hormone.[140] However no clear association between body mass index and testosterone levels has been found.[141]

Miscellaneous: Sleep: (REM sleep) increases nocturnal testosterone levels.[142] Behavior: Dominance challenges can, in some cases, stimulate increased testosterone release in men.[143] Drugs: Natural or man-made antiandrogens including spearmint tea reduce testosterone levels.[144][145][146] Licorice can decrease the production of testosterone and this effect is greater in females.[147]

Distribution[edit]

The plasma protein binding of testosterone is 98.0 to 98.5%, with 1.5 to 2.0% free or unbound.[148] It is bound 65% to sex hormone-binding globulin (SHBG) and 33% bound weakly to albumin.[149]

 

Plasma protein binding of testosterone and dihydrotestosterone show

Metabolism[edit]

vte Testosterone metabolism in humans

Testosterone structures

The image above contains clickable linksTestosterone metabolism in humans. Conjugation (sulfation and glucuronidation) occurs both with testosterone and with all of the other steroids that have one or more available hydroxyl (-OH) groups in this diagram.

Both testosterone and 5α-DHT are metabolized mainly in the liver.[1][151] Approximately 50% of testosterone is metabolized via conjugation into testosterone glucuronide and to a lesser extent testosterone sulfate by glucuronosyltransferases and sulfotransferases, respectively.[1] An additional 40% of testosterone is metabolized in equal proportions into the 17-ketosteroids androsterone and etiocholanolone via the combined actions of 5α- and 5β-reductases, 3α-hydroxysteroid dehydrogenase, and 17β-HSD, in that order.[1][151][152] Androsterone and etiocholanolone are then glucuronidated and to a lesser extent sulfated similarly to testosterone.[1][151] The conjugates of testosterone and its hepatic metabolites are released from the liver into circulation and excreted in the urine and bile.[1][151][152] Only a small fraction (2%) of testosterone is excreted unchanged in the urine.[151]

 

In the hepatic 17-ketosteroid pathway of testosterone metabolism, testosterone is converted in the liver by 5α-reductase and 5β-reductase into 5α-DHT and the inactive 5β-DHT, respectively.[1][151] Then, 5α-DHT and 5β-DHT are converted by 3α-HSD into 3α-androstanediol and 3α-etiocholanediol, respectively.[1][151] Subsequently, 3α-androstanediol and 3α-etiocholanediol are converted by 17β-HSD into androsterone and etiocholanolone, which is followed by their conjugation and excretion.[1][151] 3β-Androstanediol and 3β-etiocholanediol can also be formed in this pathway when 5α-DHT and 5β-DHT are acted upon by 3β-HSD instead of 3α-HSD, respectively, and they can then be transformed into epiandrosterone and epietiocholanolone, respectively.[153][154] A small portion of approximately 3% of testosterone is reversibly converted in the liver into androstenedione by 17β-HSD.[152]

 

In addition to conjugation and the 17-ketosteroid pathway, testosterone can also be hydroxylated and oxidized in the liver by cytochrome P450 enzymes, including CYP3A4, CYP3A5, CYP2C9, CYP2C19, and CYP2D6.[155] 6β-Hydroxylation and to a lesser extent 16β-hydroxylation are the major transformations.[155] The 6β-hydroxylation of testosterone is catalyzed mainly by CYP3A4 and to a lesser extent CYP3A5 and is responsible for 75 to 80% of cytochrome P450-mediated testosterone metabolism.[155] In addition to 6β- and 16β-hydroxytestosterone, 1β-, 2α/β-, 11β-, and 15β-hydroxytestosterone are also formed as minor metabolites.[155][156] Certain cytochrome P450 enzymes such as CYP2C9 and CYP2C19 can also oxidize testosterone at the C17 position to form androstenedione.[155]

 

Two of the immediate metabolites of testosterone, 5α-DHT and estradiol, are biologically important and can be formed both in the liver and in extrahepatic tissues.[151] Approximately 5 to 7% of testosterone is converted by 5α-reductase into 5α-DHT, with circulating levels of 5α-DHT about 10% of those of testosterone, and approximately 0.3% of testosterone is converted into estradiol by aromatase.[2][151][157][158] 5α-Reductase is highly expressed in the male reproductive organs (including the prostate gland, seminal vesicles, and epididymides),[159] skin, hair follicles, and brain[160] and aromatase is highly expressed in adipose tissue, bone, and the brain.[161][162] As much as 90% of testosterone is converted into 5α-DHT in so-called androgenic tissues with high 5α-reductase expression,[152] and due to the several-fold greater potency of 5α-DHT as an AR agonist relative to testosterone,[163] it has been estimated that the effects of testosterone are potentiated 2- to 3-fold in such tissues.[164]

 

Levels[edit]

Total levels of testosterone in the body are 264 to 916 ng/dL in men age 19 to 39 years,[165] while mean testosterone levels in adult men have been reported as 630 ng/dL.[166] Levels of testosterone in men decline with age.[165] In women, mean levels of total testosterone have been reported to be 32.6 ng/dL.[167][168] In women with hyperandrogenism, mean levels of total testosterone have been reported to be 62.1 ng/dL.[167][168]

 

Testosterone levels in males and females show

Total testosterone levels in males throughout life show

 

Reference ranges for blood tests, showing adult male testosterone levels in light blue at center-left.

Measurement[edit]

Testosterone’s bioavailable concentration is commonly determined using the Vermeulen calculation or more precisely using the modified Vermeulen method,[174][175] which considers the dimeric form of sex-hormone-binding-globulin.[176]

 

Both methods use chemical equilibrium to derive the concentration of bioavailable testosterone: in circulation testosterone has two major binding partners, albumin (weakly bound) and sex-hormone-binding-globulin (strongly bound). These methods are described in detail in the accompanying figure.

  

Dimeric sex-hormone-binding-globulin with its testosterone ligands

  

Two methods for determining concentration of bioavailable testosterone.

History[edit]

A testicular action was linked to circulating blood fractions – now understood to be a family of androgenic hormones – in the early work on castration and testicular transplantation in fowl by Arnold Adolph Berthold (1803–1861).[177] Research on the action of testosterone received a brief boost in 1889, when the Harvard professor Charles-Édouard Brown-Séquard (1817–1894), then in Paris, self-injected subcutaneously a "rejuvenating elixir" consisting of an extract of dog and guinea pig testicle. He reported in The Lancet that his vigor and feeling of well-being were markedly restored but the effects were transient,[178] and Brown-Séquard's hopes for the compound were dashed. Suffering the ridicule of his colleagues, he abandoned his work on the mechanisms and effects of androgens in human beings.

 

In 1927, the University of Chicago's Professor of Physiologic Chemistry, Fred C. Koch, established easy access to a large source of bovine testicles — the Chicago stockyards — and recruited students willing to endure the tedious work of extracting their isolates. In that year, Koch and his student, Lemuel McGee, derived 20 mg of a substance from a supply of 40 pounds of bovine testicles that, when administered to castrated roosters, pigs and rats, remasculinized them.[179] The group of Ernst Laqueur at the University of Amsterdam purified testosterone from bovine testicles in a similar manner in 1934, but isolation of the hormone from animal tissues in amounts permitting serious study in humans was not feasible until three European pharmaceutical giants—Schering (Berlin, Germany), Organon (Oss, Netherlands) and Ciba (Basel, Switzerland)—began full-scale steroid research and development programs in the 1930s.

  

Nobel Prize winner, Leopold Ruzicka of Ciba, a pharmaceutical industry giant that synthesized testosterone.

The Organon group in the Netherlands were the first to isolate the hormone, identified in a May 1935 paper "On Crystalline Male Hormone from Testicles (Testosterone)".[180] They named the hormone testosterone, from the stems of testicle and sterol, and the suffix of ketone. The structure was worked out by Schering's Adolf Butenandt, at the Chemisches Institut of Technical University in Gdańsk.[181][182]

 

The chemical synthesis of testosterone from cholesterol was achieved in August that year by Butenandt and Hanisch.[183] Only a week later, the Ciba group in Zurich, Leopold Ruzicka (1887–1976) and A. Wettstein, published their synthesis of testosterone.[184] These independent partial syntheses of testosterone from a cholesterol base earned both Butenandt and Ruzicka the joint 1939 Nobel Prize in Chemistry.[182][185] Testosterone was identified as 17β-hydroxyandrost-4-en-3-one (C19H28O2), a solid polycyclic alcohol with a hydroxyl group at the 17th carbon atom. This also made it obvious that additional modifications on the synthesized testosterone could be made, i.e., esterification and alkylation.

 

The partial synthesis in the 1930s of abundant, potent testosterone esters permitted the characterization of the hormone's effects, so that Kochakian and Murlin (1936) were able to show that testosterone raised nitrogen retention (a mechanism central to anabolism) in the dog, after which Allan Kenyon's group[186] was able to demonstrate both anabolic and androgenic effects of testosterone propionate in eunuchoidal men, boys, and women. The period of the early 1930s to the 1950s has been called "The Golden Age of Steroid Chemistry",[187] and work during this period progressed quickly. Research in this golden age proved that this newly synthesized compound—testosterone—or rather family of compounds (for many derivatives were developed from 1940 to 1960), was a potent multiplier of muscle, strength, and well-being.[188]

 

Other animals[edit]

Testosterone is observed in most vertebrates. Testosterone and the classical nuclear androgen receptor first appeared in gnathostomes (jawed vertebrates).[189] Agnathans (jawless vertebrates) such as lampreys do not produce testosterone but instead use androstenedione as a male sex hormone.[190] Fish make a slightly different form called 11-ketotestosterone.[191] Its counterpart in insects is ecdysone.[192] The presence of these ubiquitous steroids in a wide range of animals suggest that sex hormones have an ancient evolutionary history.[193]

 

en.wikipedia.org/wiki/Testosterone

Testy Fest 2013

Normal testicle. Arrows point to the outer edge of the testicle.

No, not Finglands 1741 (N741 VBA) (fine vehicle without a doubt), but that rather distracting advert promoting awareness about testicular cancer, just off to the right.

 

Go on lads, have a quick check: you know it makes sense...

 

Manchester Oxford Road, 08/02/2008

London's City Hall is an unusual spherical shape. One local aptly described it as a "glass testicle."

Keep your testicles at their optimal operating temperature. Choose Nude!

Pt came in with pain and mass in right testicle. Scheduled for removal the next day, but he was very disappointed in the fact that he would not be allowed to see it, as it had to go to pathology, so he asked me to take a picture for him. Images and testicle taken with patient's permission.

1 2 ••• 6 7 9 11 12 ••• 79 80