NZ has the only polymorphic species

The lesser goldfinch or dark-backed goldfinch (Spinus psaltria) is a very small songbird of the Americas. Together with its relatives the American goldfinch and Lawrence's goldfinch, it forms the American goldfinches clade in the genus Spinus sensu stricto.

 

The American goldfinches can be distinguished by the males having a black (rarely green) forehead, whereas the latter is (like the rest of the face) red or yellow in the European goldfinch and its relatives. North American males are markedly polymorphic and 5 subspecies are often named; at least 2 of them seem to represent a less-progressed stage in evolution however.

 

This petite species is not only the smallest North American Spinus finch, it may be the smallest true finch in the world.Some sources list more subtropical Spinus species as slightly smaller on average, including the Andean siskin. This species ranges from 9 to 12 cm (3.5 to 4.7 in) in length and can weigh from 8 to 11.5 g (0.28 to 0.41 oz).Among standard measurements, the wing chord is 5.5 to 7 cm (2.2 to 2.8 in), the tail is 3.9 to 4.7 cm (1.5 to 1.9 in), the bill is 0.9 to 1.1 cm (0.35 to 0.43 in) and the tarsus is 1.1 to 1.2 cm (0.43 to 0.47 in).[4] There is a slight NW-SE cline in size, with the largest birds from Mexico and south being up to one-fifth larger than the smallest from the extreme NW of its range; this effect is more pronounced in females. There is also considerable variation in the amount of black on head and back in males, and thus three subspecies have been proposed. But this variation too seem to be simple and clinal changes in allele frequency, and thus the "subspecies" might be better considered morphs or geographical forms.

 

Males are easily recognized by their bright yellow underparts and big white patches in the tail (outer rectrices) and on the wings (the base of the primaries). They range from having solid black from the back to the upper head including the ear-coverts to having these regions medium green; each of the back, crown and ear regions varies in darkness rather independently though as a rule the ears are not darker than the rest. In most of the range dark psaltria birds (Arkansas goldfinch) predominate. The light birds are termed hesperophilus and are most common in the far western U.S. and northwestern Mexico.

 

Los Angeles. California.

The gyrfalcon (Falco rusticolus), the largest of the falcon species, is a bird of prey. The abbreviation gyr is also used. It breeds on Arctic coasts and tundra, and the islands of northern North America and the Eurosiberian region.

 

It is mainly a resident there also, but some gyrfalcons disperse more widely after the breeding season, or in winter. Individual vagrancy can take birds for long distances. Its plumage varies with location, with birds being coloured from all-white to dark brown.

 

These colour variations are called morphs. Like other falcons, it shows sexual dimorphism, with the female much larger than the male. For centuries, the gyrfalcon has been valued as a hunting bird. Typical prey includes the ptarmigan and waterfowl, which it may take in flight; it also takes fish and mammals.

 

The gyrfalcon is the largest falcon in the world, being about the same size as the largest buteos (buzzards) but probably slightly heavier. Males are 48 to 61 centimetres (19 to 24 inches) long, weigh 805 to 1,350 grams (1 pound 12+1⁄2 ounces to 2 pounds 15+1⁄2 ounces), with average weights reported as 1,130 or 1,170 g (2 lb 8 oz or 2 lb 9+1⁄2 oz) and have a wingspan from 110 to 130 cm (43 to 51 in).

 

Females are bulkier and larger, at 51 to 65 cm (20 to 25+1⁄2 in) long, 124 to 160 cm (49 to 63 in) wingspan, and of 1,180 to 2,100 g (2 lb 9+1⁄2 oz to 4 lb 10 oz) weight, with average weights of 1,585 or 1,752 g (3 lb 8 oz or 3 lb 13+3⁄4 oz).[8][9] An outsized female from eastern Siberia was found to have scaled 2,600 g (5 lb 12 oz).

 

Among standard measurements, the wing chord is 34.5 to 41 cm (13.6 to 16.1 in), the tail is 19.5 to 29 cm (7.7 to 11.4 in), the culmen is 2 to 2.8 cm (3⁄4 to 1+1⁄8 in) and the tarsus is 4.9 to 7.5 cm (1+7⁄8 to 3 in). The gyrfalcon is larger, broader-winged and longer-tailed than the peregrine falcon, which it is known to compete with (and occasionally hunt). It differs from the buzzard in general structure, having pointed wings.

 

The gyrfalcon is a very polymorphic species, so its plumage varies greatly. The archetypal morphs are called "white", "silver", "brown", and "black", though they can be coloured on a spectrum from all-white to very dark. The brown form of the gyrfalcon is distinguished from the peregrine by the cream streaking on the nape and crown and by the absence of a well-defined malar stripe and cap.

 

The black morph has a strongly black-spotted underside, rather than finely barred as in the peregrine. White form gyrfalcons are the only predominantly white falcons. Silver gyrfalcons resemble a light grey lanner falcon of larger size. The species shows no sex-based colour differences; juveniles are darker and browner than adults.

 

The black color seems to be sex-linked and to occur mostly in females; it proved difficult for breeders to get males darker than the dark side of slate grey. A color variety that arose in captive breeding is "black chick"

 

This image was taken at Dunrobin Castle in Scotland during a "Falconry Display"

The Great Mormon (Papilio memnon) is a large butterfly that belongs to the swallowtail family and is found in southern Asia. It is widely distributed and has thirteen subspecies. Furthermore, within these groupings, there are four male and many female forms (both tailed and tailless), the females being highly polymorphic and many of them being mimics of unpalatable butterflies. This species has been studied extensively for understanding the genetic basis for polymorphy and Batesian mimicry. As many as twenty-six female forms are reported.

 

I believe this individual is either Papilio memnon pryeri or Papilio memnon thunbergi.

 

Pu'er, Yunnan, China

things went out of control...

The red-tailed hawk (Buteo jamaicensis) is a bird of prey that breeds throughout most of North America, from the interior of Alaska and northern Canada to as far south as Panama and the West Indies. It is one of the most common members within the genus of Buteo in North America or worldwide. The red-tailed hawk is one of three species colloquially known in the United States as the "chickenhawk", though it rarely preys on standard-sized chickens. The bird is sometimes also referred to as the red-tail for short, when the meaning is clear in context. Red-tailed hawks can acclimate to all the biomes within their range, occurring on the edges of non-ideal habitats such as dense forests and sandy deserts. The red-tailed hawk occupies a wide range of habitats and altitudes, including deserts, grasslands (from small meadows to the treed fringes of more extensive prairies), coniferous and deciduous forests, agricultural fields, and urban areas. Its latitudinal limits fall around the tree line in the subarctic and it is absent from the high Arctic. Generally it favors varied habitats with open woodland, woodland edge and open terrain. It is legally protected in Canada, Mexico, and the United States by the Migratory Bird Treaty Act.

 

The 14 recognized subspecies vary in appearance and range, varying most often in color, and in the west of North America, red-tails are particularly often strongly polymorphic, with individuals ranging from almost white to nearly all black. The subspecies Harlan's hawk (B. j. harlani) is sometimes considered a separate species (B. harlani). The red-tailed hawk is one of the largest members of the genus Buteo, typically weighing from 690 to 1,600 g (1.5 to 3.5 lb) and measuring 45–65 cm (18–26 in) in length, with a wingspan from 110–141 cm (3 ft 7 in – 4 ft 8 in). This species displays sexual dimorphism in size, with females averaging about 25% heavier than males.

 

The diet of red-tailed hawks is highly variable and reflects their status as opportunistic generalists, but in North America, they are most often predators of small mammals such as rodents of an immense diversity of families and species. Prey that is terrestrial and at least partially diurnal is preferred, so types such as ground squirrels are preferred where they naturally occur. Over much of the range, smallish rodents such as voles alternated with larger rabbits and hares often collectively form the bulk of the diet. Large numbers of birds and reptiles can occur in the diet in several areas, and can even be the primary foods. Meanwhile, amphibians, fish and invertebrates can seem rare in the hawk's regular diet, but they are not infrequently taken by immature hawks. Red-tailed hawks may survive on islands absent of native mammals on diets variously including invertebrates such as crabs, as well as lizards or birds. Like many Buteo species, they hunt from a perch most often, but can vary their hunting techniques where prey and habitat demand it. Because they are so common and easily trained as capable hunters, in the United States they are the most commonly captured hawks for falconry. Falconers are permitted to take only passage hawks (which have left the nest, are on their own, but are less than a year old) so as to not affect the breeding population. Passage red-tailed hawks are also preferred by falconers because they have not yet developed the adult behaviors that would make them more difficult to train.

 

Taxonomy

The red-tailed hawk was formally described in 1788 by German naturalist Johann Friedrich Gmelin under the binomial name Falco jamaicensis. Gmelin based his description on the "cream-coloured buzzard" described in 1781 by John Latham in his A General Synopsis of Birds. The type locality is Jamaica. The red-tailed hawk is now placed in the genus Buteo that was erected by French naturalist Bernard Germain de Lacépède in 1799.

 

The red-tailed hawk is a member of the subfamily Buteoninae, which includes about 55 currently recognized species. Unlike many lineages of accipitrids, which seemed to have radiated out of Africa or south Asia, the Buteoninae clearly originated in the Americas based on fossil records and current species distributions (more than 75% of the extant hawks from this lineage are found in the Americas). As a subfamily, the Buteoninae seem to be rather old based on genetic materials, with monophyletic genera bearing several million years of individual evolution. Diverse in plumage appearance, habitat, prey, and nesting preferences, buteonine hawks are nonetheless typically medium- to large-sized hawks with ample wings (while some fossil forms are very large, larger than any eagle alive today). The red-tailed hawk is a member of the genus Buteo, a group of medium-sized raptors with robust bodies and broad wings. Members of this genus are known as "buzzards" in Eurasia, but "hawks" in North America. Under current classification, the genus includes about 29 species, the second-most diverse of all extant accipitrid genera behind only Accipiter. The buzzards of Eurasia and Africa are mostly part of the genus Buteo, although two other small genera within the subfamily Buteoninae occur in Africa.

 

At one time, the rufous-tailed hawk (B. ventralis), distributed in Patagonia and some other areas of southern South America, was considered part of the red-tailed hawk species. With a massive distributional gap consisting of most of South America, the rufous-tailed hawk is considered a separate species now, but the two hawks still compromise a "species pair" or superspecies, as they are clearly closely related. The rufous-tailed hawk, while comparatively little studied, is very similar to the red-tailed hawk, being about the same size and possessing the same wing structure, and having more or less parallel nesting and hunting habits. Physically, however, rufous-tailed hawk adults do not attain a bright brick-red tail as do red-tailed hawks, instead retaining a dark brownish-cinnamon tail with many blackish crossbars similar to juvenile red-tailed hawks. Another, more well-known, close relative to the red-tailed hawk is the common buzzard (B. buteo), which has been considered as its Eurasian "broad ecological counterpart" and may also be within a species complex with red-tailed hawks. The common buzzard, in turn, is also part of a species complex with other Old World buzzards, namely the mountain buzzard (B. oreophilus), the forest buzzard (B. trizonatus ), and the Madagascar buzzard (B. brachypterus). All six species, although varying notably in size and plumage characteristics, in the alleged species complex that contains the red-tailed hawk share with it the feature of the blackish patagium marking, which is missing in most other Buteo spp.

 

Subspecies

At least 14 recognized subspecies of B. jamaicensis are described, which vary in range and in coloration. Not all authors accept every subspecies, though, particularly some of the insular races of the tropics (which differ only slightly in some cases from the nearest mainland forms) and particularly Krider's hawk, by far the most controversial red-tailed hawk race, as few authors agree on its suitability as a full-fledged subspecies.

 

ImageSubspeciesDistribution

Jamaican red-tailed hawk (B. j. jamaicensis)occurs throughout the West Indies (including Jamaica, Hispaniola, Puerto Rico and the Lesser Antilles) except for the Bahamas and Cuba.

Alaska red-tailed hawk (B. j. alascensis)breeds (probably resident) from southeastern coastal Alaska to Haida Gwaii and Vancouver Island in British Columbia.

Eastern red-tailed hawk (B. j. borealis)breeds from southeast Canada and Maine south through Texas and east to northern Florida.

Western red-tailed hawk (B. j. calurus)greatest longitudinal breeding distribution of any race of red-tailed hawk.

Central American red-tailed hawk (B. j. costaricensis)from Nicaragua to Panama.

Southwestern red-tailed hawk (B. j. fuertesi)breeds from northern Chihuahua to South Texas.

Tres Marias red-tailed hawk (B. j. fumosus)endemic to Islas Marías, Mexico.

Mexican Highlands red-tailed hawk (B. j. hadropus)native to the Mexican Highlands.

Harlan's hawk (B. j. harlani)breeds from central Alaska to northwestern Canada, with the largest number of birds breeding in the Yukon or western Alaska, reaching their southern limit in north-central British Columbia.

Red-tailed hawk (kemsiesi) (B. j. kemsiesi)a dark subspecies resident from Chiapas, Mexico, to Nicaragua.

Krider's hawk (B. j. kriderii)breeds from southern Alberta, southern Saskatchewan, southern Manitoba, and extreme western Ontario south to south-central Montana, Wyoming, western Nebraska, and western Minnesota.

Socorro red-tailed hawk (B. j. socorroensis)endemic to Socorro Island, Mexico.

Cuban red-tailed hawk (B. j. solitudinis)native to the Bahamas and Cuba.

Florida red-tailed hawk (B. j. umbrinus)occurs year-round in peninsular Florida north to as far Tampa Bay and the Kissimmee Prairie south throughout the rest of peninsular Florida south to the Florida Keys.

 

Description

Red-tailed hawk plumage can be variable, depending on the subspecies and the region. These color variations are morphs, and are not related to molting. The western North American population, B. j. calurus, is the most variable subspecies and has three main color morphs: light, dark, and intermediate or rufous. The dark and intermediate morphs constitute 10–20% of the population in the Western United States, but seem to constitute only 1–2% of B. j. calurus in western Canada. A whitish underbelly with a dark brown band across the belly, formed by horizontal streaks in feather patterning, is present in most color variations. This feature is variable in eastern hawks and generally absent in some light subspecies (i.e. B. j. fuertesi). Most adult red-tails have a dark-brown nape and upper head, which gives them a somewhat hooded appearance, while the throat can variably present a lighter brown "necklace". Especially in younger birds, the underside may be otherwise covered with dark-brown spotting, and some adults may too manifest this stippling. The back is usually a slightly darker brown than elsewhere with paler scapular feathers, ranging from tawny to white, forming a variable imperfect "V" on the back. The tail of most adults, which gives this species its name, is rufous brick-red above with a variably sized, black subterminal band and generally appears light buff-orange from below. In comparison, the typical pale immatures (i.e. less than two years old) typically have a mildly paler headed and tend to show a darker back than adults with more apparent pale wing-feather edges above (for descriptions of dark morph juveniles from B. j. calurus, which is also generally apt for description of rare dark morphs of other races, see under that subspecies description). In immature red-tailed hawks of all morphs, the tail is a light brown above with numerous small dark brown bars of roughly equal width, but these tend to be much broader on dark morph birds. Even in young red-tails, the tail may be a somewhat rufous tinge of brown. The bill is relatively short and dark, in the hooked shape characteristic of raptors, and the head can sometimes appear small in size against the thick body frame. The cere, the legs, and the feet of the red-tailed hawk are all yellow, as is the color of bare parts in many accipitrids of different lineages. Immature birds can be readily identified at close range by their yellowish irises. As the bird attains full maturity over the course of 3–4 years, the iris slowly darkens into a reddish-brown, which is the adult eye-color in all races. Seen in flight, adults usually have dark brown along the lower edge of the wings, against a mostly pale wing, which bares light brownish barring. Individually, the underwing coverts can range from all dark to off-whitish (most often more heavily streaked with brown) which contrasts with a distinctive black patagium marking. The wing coloring of adults and immatures is similar but for typical pale morph immatures having somewhat heavier brownish markings.

 

Though the markings and color vary across the subspecies, the basic appearance of the red-tailed hawk is relatively consistent.

 

Overall, this species is blocky and broad in shape, often appearing (and being) heavier than other Buteos of similar length. They are the heaviest Buteos on average in eastern North America, albeit scarcely ahead of the larger winged rough-legged buzzard (Buteo lagopus), and second only in size in the west to the ferruginous hawk (Buteo regalis). Red-tailed hawks may be anywhere from the fifth to the ninth heaviest Buteo in the world depending on what figures are used. However, in the northwestern United States, ferruginous hawk females are 35% heavier than female red-tails from the same area.[2] On average, western red-tailed hawks are relatively longer winged and lankier proportioned but are slightly less stocky, compact and heavy than eastern red-tailed hawks in North America. Eastern hawks may also have mildly larger talons and bills than western ones. Based on comparisons of morphology and function amongst all accipitrids, these features imply that western red-tails may need to vary their hunting more frequently to on the wing as the habitat diversifies to more open situations and presumably would hunt more variable and faster prey, whereas the birds of the east, which was historically well-wooded, are more dedicated perch hunters and can take somewhat larger prey but are likely more dedicated mammal hunters. In terms of size variation, red-tailed hawks run almost contrary to Bergmann's rule (i.e. that northern animals should be larger in relation than those closer to the Equator within a species) as one of the northernmost subspecies, B. j. alascensis, is the second smallest race based on linear dimensions and that two of the most southerly occurring races in the United States, B. j. fuertesi and B. j. umbrinus, respectively, are the largest proportioned of all red-tailed hawks. Red-tailed hawks tend have a relatively short but broad tails and thick, chunky wings. Although often described as long-winged, the proportional size of the wings is quite small and red-tails have high wing loading for a buteonine hawk. For comparison, two other widespread Buteo hawks in North America were found to weigh: 30 g (1.1 oz) for every square centimeter of wing area in the rough-legged buzzard (B. lagopus) and 44 g (1.6 oz)/cm2 in the red-shouldered hawk (B. lineatus). In contrast, the red-tailed hawk weighed considerably more for their wing area: 199 g (7.0 oz) per square cm.

 

As is the case with many raptors, the red-tailed hawk displays sexual dimorphism in size, as females are on average 25% larger than males. As is typical in large raptors, frequently reported mean body mass for red-tailed hawks is somewhat higher than expansive research reveals. Part of this weight variation is seasonal fluctuations; hawks tend to be heavier in winter than during migration or especially during the trying summer breeding season, and also due to clinal variation. Furthermore, immature hawks are usually lighter in mass than their adult counterparts despite having somewhat longer wings and tails. Male red-tailed hawks may weigh from 690 to 1,300 g (1.52 to 2.87 lb) and females may weigh 801 to 1,723 g (1.766 to 3.799 lb) (the lowest figure from a migrating female immature from Goshute Mountains, Nevada, the highest from a wintering female in Wisconsin). Some sources claim the largest females can weigh up to 2,000 g (4.4 lb), but whether this is in reference to wild hawks (as opposed to those in captivity or used for falconry) is not clear. The largest known survey of body mass in red-tailed hawks is still credited to Craighead and Craighead (1956), who found 100 males to average 1,028 g (2.266 lb) and 108 females to average 1,244 g (2.743 lb). However, these figures were apparently taken from labels on museum specimens, from natural history collections in Wisconsin and Pennsylvania, without note to the region, age, or subspecies of the specimens. However, 16 sources ranging in sample size from the aforementioned 208 specimens to only four hawks in Puerto Rico (with 9 of the 16 studies of migrating red-tails), showed that males weigh a mean of 860.2 g (1.896 lb) and females weigh a mean of 1,036.2 g (2.284 lb), about 15% lighter than prior species-wide published weights. Within the continental United States, typical weights of males can range from 840.8 g (1.854 lb) (for migrating males in Chelan County, Washington) to 1,031 g (2.273 lb) (for male hawks found dead in Massachusetts), and females ranged from 1,057.9 g (2.332 lb) (migrants in the Goshutes) to 1,373 g (3.027 lb) (for females diagnosed as B. j. borealis in western Kansas). Size variation in body mass reveals that the red-tailed hawk typically varies only a modest amount and that size differences are geographically inconsistent.[9][40]

 

Male red-tailed hawks can measure 45 to 60 cm (18 to 24 in) in total length, females measuring 48 to 65 cm (19 to 26 in) long. Their wingspan typically can range from 105 to 141 cm (3 ft 5 in to 4 ft 8 in), although the largest females may possible span up to 147 cm (4 ft 10 in). In the standard scientific method of measuring wing size, the wing chord is 325.1–444.5 mm (12.80–17.50 in) long. The tail measures 188 to 258.7 mm (7.40 to 10.19 in) in length. The exposed culmen was reported to range from 21.7 to 30.2 mm (0.85 to 1.19 in) and the tarsus averaged 74.7–95.8 mm (2.94–3.77 in) across the races. The middle toe (excluding talon) can range from 38.3 to 53.8 mm (1.51 to 2.12 in), with the hallux-claw (the talon of the rear toe, which has evolved to be the largest in accipitrids) measuring from 24.1 to 33.6 mm (0.95 to 1.32 in) in length.

 

Identification

Although they overlap in range with most other American diurnal raptors, identifying most mature red-tailed hawks to species is relatively straightforward, particularly if viewing a typical adult at a reasonable distance. The red-tailed hawk is the only North American hawk with a rufous tail and a blackish patagium marking on the leading edge of its wing (which is obscured only on dark morph adults and Harlan's hawks by similarly dark-colored feathers). Other larger adult Buteo spp. in North America usually have obvious distinct markings that are absent in red-tails, whether the rufous-brown "beard" of Swainson's hawks (B. swainsonii) or the colorful rufous belly and shoulder markings and striking black-and-white mantle of red-shouldered hawks (also the small "windows" seen at the end of their primaries).[ In perched individuals, even as silhouettes, the shape of large Buteo spp. may be distinctive, such as the wingtips overhanging the tail in several other species, but not in red-tails. North American Buteo spp. range from the dainty, compact builds of much smaller ones, such as broad-winged hawk (B. platypterus) to the heavyset, neckless look of ferruginous hawks or the rough-legged buzzards, which have a compact, smaller appearance than a red-tail in perched birds due to its small bill, short neck, and much shorter tarsi, while the opposite effect occurs in flying rough-legs with their much bigger wing area. In flight, most other large North American Buteo spp. are distinctly longer and more slender-winged than red-tailed hawks, with the much paler ferruginous hawk having peculiarly slender wings in relation to its massive, chunky body. Swainson's hawks are distinctly darker on the wing and ferruginous hawks are much paler-winged than typical red-tailed hawks. Pale morph adult ferruginous hawk can show mildly tawny-pink (but never truly rufous) upper tail, and like red-tails tend to have dark markings on underwing-coverts and can have a dark belly band, but compared to red-tailed hawks have a distinctly broader head, their remiges are much whiter looking with very small, dark primary tips, they lack the red-tail's diagnostic patagial marks and usually also lack the dark subterminal tail-band, and ferruginous hawks have totally feathered tarsi. With its whitish head, the ferruginous hawk is most similar to Krider's red-tailed hawks, especially in immature plumage, but the larger hawk has broader head and narrower wing shape, and the ferruginous immatures are paler underneath and on their legs. Several species share a belly band with the typical red-tailed hawk, but they vary from subtle (as in the ferruginous hawk) to solid blackish, the latter in most light-morph rough-legged buzzards. More difficult to identify among adult red-tails are their darkest variations, as most species of Buteo in North America also have dark morphs. Western dark morph red-tails (i.e. B. j. calurus) adults, however, retain the typical distinctive brick-red tail, which other species lack, and may stand out even more against the otherwise all chocolate-brown to black bird. Standard pale juveniles when perched show a whitish patch in the outer half of the upper surface of the wing, which other juvenile Buteo spp. lack.[ The most difficult to identify stages and plumage types are dark morph juveniles, Harlan's hawk and some Krider's hawks (the latter mainly with typical ferruginous hawks as mentioned). Some darker juveniles are similar enough to other Buteo juveniles that they "cannot be identified to species with any confidence under various field conditions." However, field identification techniques have advanced in the last few decades and most experienced hawk-watchers can distinguish even the most vexingly plumaged immature hawks, especially as the wing shapes of each species becomes apparent after seeing many. Harlan's hawks are most similar to dark morph rough-legged buzzards and dark morph ferruginous hawks. Wing shape is the most reliable identification tool for distinguishing Harlan's hawks from these, but also the pale streaking on the breast of Harlan's, which tends to be conspicuous in most individuals, and is lacking in the other hawks. Also, dark morph ferruginous hawks do not have the dark subterminal band of a Harlan's hawk, but do bear a black undertail covert lacking in Harlan's.

 

Vocalization

The cry of the red-tailed hawk is a 2- to 3-second, hoarse, rasping scream, variously transcribed as kree-eee-ar, tsee-eeee-arrr or sheeeeee, that begins at a high pitch and slurs downward. This cry is often described as sounding similar to a steam whistle. The red-tailed hawk frequently vocalizes while hunting or soaring, but vocalizes loudest and most persistently in defiance or anger, in response to a predator or a rival hawk's intrusion into its territory. At close range, it makes a croaking guh-runk, possibly as a warning sound. Nestlings may give peeping notes with a "soft, sleepy quality" that give way to occasional screams as they develop, but those are more likely to be a soft whistle rather than the harsh screams of the adults. Their latter hunger call, given from 11 days (as recorded in Alaska) to after fledgling (in California), is different, a two-syllabled, wailing klee-uk food cry exerted by the young when parents leave the nest or enter their field of vision. A strange mechanical sound "not very unlike the rush of distant water" has been reported as uttered in the midst of a sky-dance. A modified call of chirp-chwirk is given during courtship, while a low key, duck-like nasal gank may be given by pairs when they are relaxed.

 

The fierce, screaming cry of the adult red-tailed hawk is frequently used as a generic raptor sound effect in television shows and other media, even if the bird featured is not a red-tailed hawk. It is especially used in depictions of the bald eagle, which contributes to the common misconception that it is a bald eagle cry; actual bald eagle vocalizations are far softer and more chirpy than those of a red-tailed hawk.

 

Distribution and habitat

The red-tailed hawk is one of the most widely distributed of all raptors in the Americas. It occupies the largest breeding range of any diurnal raptor north of the Mexican border, just ahead of the American kestrel (Falco sparverius). While the peregrine falcon (Falco peregrinus) has a greater latitudinal distribution as a nester in North America, its range as a breeding species is far more sporadic and sparse than that of red-tailed hawks. The red-tailed hawk breeds from nearly north-central Alaska, the Yukon, and a considerable portion of the Northwest Territories, there reaching as far as a breeder as Inuvik, Mackenzie River Delta and skirting the southern shores of Great Bear Lake and Great Slave Lake. Thereafter in northern Canada, breeding red-tails continue to northern Saskatchewan and across to north-central Ontario east to central Quebec and the Maritime Provinces of Canada, and south continuously to Florida. No substantial gaps occur throughout the entire contiguous United States where breeding red-tailed hawks do not occur. Along the Pacific, their range includes all of Baja California, including Islas Marías, and Socorro Island in the Revillagigedo Islands. On the mainland, breeding red-tails are found continuously to Oaxaca, then experience a brief gap at the Isthmus of Tehuantepec thereafter subsequently continuing from Chiapas through central Guatemala on to northern Nicaragua. To the south, the population in highlands from Costa Rica to central Panama is isolated from breeding birds in Nicaragua. Further east, breeding red-tailed hawks occur in the West Indies in north Bahamas (i.e. Grand Bahama, Abaco and Andros) and all larger islands (such as Cuba, Jamaica, Hispaniola, and Puerto Rico) and into the northern Lesser Antilles (Virgin Islands, Saint Barthélemy, Saba, Saint Kitts, and Nevis, being rare as a resident on Saint Eustatius and are probably extinct on Saint Martin). Their typical winter range stretches from southern Canada south throughout the remainder of the breeding range.

 

Red-tailed hawks have shown the ability to become habituated to almost any habitat present in North and Central America. Their preferred habitat is mixed forest and field, largely woodland edge with tall trees or alternately high bluffs that may be used as nesting and perching sites. They occupy a wide range of habitats and altitudes, including deserts, grasslands, nearly any coastal or wetland habitat, mountains, foothills, coniferous and deciduous woodlands, and tropical rainforests. Agricultural fields and pastures, which are more often than not varied with groves, ridges, or streamside trees in most parts of America, may make nearly ideal habitat for breeding or wintering red-tails. They also adapt well to suburban areas especially ones with tall trees or any kind of parkland. Some red-tails may survive or even flourish in urban areas, usually hunting and roosting in available urban parks, cemeteries, road verges, and so on, and nesting freely either in trees or virtually any tall man-made structures. One famous urban red-tailed hawk, known as "Pale Male", became the subject of a nonfiction book, Red-Tails in Love: A Wildlife Drama in Central Park, and is the first known red-tail in decades to successfully nest and raise young in the crowded New York City borough of Manhattan. As studied in Syracuse, New York, the highway system has been very beneficial to red-tails as it juxtaposed trees and open areas and blocks human encroachment with fences, with the red-tailed hawks easily becoming acclimated to car traffic. The only practice that has a negative effect on the highway-occupying red-tails is the planting of exotic Phragmites, which may occasionally obscure otherwise ideal highway habitat.

 

In the northern Great Plains, the widespread practices of wildfire suppression and planting of exotic trees by humans has allowed groves of aspen and various other trees to invade what was once vast, almost continuous prairie grasslands, causing grassland obligates such as ferruginous hawks to decline and allowing parkland-favoring red-tails to flourish. To the contrary, clear-cutting of mature woodlands in New England, resulting in only fragmented and isolated stands of trees or low second growth remaining, was recorded to also benefit red-tailed hawks, despite being to the determent of breeding red-shouldered hawks. The red-tailed hawk, as a whole, rivals the peregrine falcon and the great horned owl among raptorial birds in the use of diverse habitats in North America. Beyond the high Arctic (as they discontinue as a breeder at the tree line), few other areas exist where red-tailed hawks are absent or rare in North and Central America. Some areas of unbroken forest, especially lowland tropical forests, rarely host red-tailed hawks, although they can occupy forested tropical highlands surprisingly well. In deserts, they can only occur where some variety of arborescent growth or ample rocky bluffs or canyons occur.

 

Behavior

The red-tailed hawk is highly conspicuous to humans in much of its daily behavior. Most birds in resident populations, which are well more than half of all red-tailed hawks, usually split nonbreeding-season activity between territorial soaring flight and sitting on a perch. Often, perching is for hunting purposes, but many sit on a tree branch for hours, occasionally stretching on a single wing or leg to keep limber, with no signs of hunting intent. Wintering typical pale-morph hawks in Arkansas were found to perch in open areas near the top of tall, isolated trees, whereas dark morphs more frequently perched in dense groups of trees. For many, and perhaps most, red-tailed hawks being mobbed by various birds is a daily concern and can effectively disrupt many of their daily behaviors. Mostly larger passerines, of multiple families from tyrant flycatchers to icterids, mob red-tails, despite other raptors, such as Accipiter hawks and falcons, being a notably greater danger to them. The most aggressive and dangerous attacker as such is likely to be various crows or other corvids, i.e. American crows (Corvus brachyrhynchos), because a mobbing group (or "murder") of them can number up to as many as 75 crows, which may cause grievous physical harm to a solitary hawk, and if the hawks are nesting, separate the parent hawks and endanger the eggs or nestlings within their nest to predation by crows. Birds that mob red-tailed hawks can tell how distended the hawk's crop is (i.e. the upper chest and throat area being puffy versus flat-feathered and sleek), thus mob more often when the hawk is presumably about to hunt.

 

Flight

In flight, this hawk soars with wings often in a slight dihedral, flapping as little as possible to conserve energy. Soaring is by far the most efficient method of flight for red-tailed hawks, so is used more often than not. Active flight is slow and deliberate, with deep wing beats. Wing beats are somewhat less rapid in active flight than in most other Buteo hawks, even heavier species such as ferruginous hawks tend to flap more swiftly, due to the morphology of the wings. In wind, it occasionally hovers on beating wings and remains stationary above the ground, but this flight method is rarely employed by this species. When soaring or flapping its wings, it typically travels from 32 to 64 km/h (20 to 40 mph), but when diving may exceed 190 km/h (120 mph). Although North American red-tailed hawks will occasionally hunt from flight, a great majority of flight by red-tails in this area is for non-hunting purpose. During nest defense, red-tailed hawks may be capable of surprisingly swift, vigorous flight, while repeatedly diving at perceived threats.

 

Migration

Red-tailed hawks are considered partial migrants, as in about the northern third of their distribution, which is most of their range in Canada and Alaska, they almost entirely vacate their breeding grounds. In coastal areas of the north, however, such as in the Pacific Northwest to southern Alaska and in Nova Scotia on the Atlantic, red-tailed hawks do not usually migrate. More or less, any area where snow cover is nearly continuous during the winter shows an extended absence of most red-tailed hawks, so some areas as far south as Montana may show strong seasonal vacancies of red-tails. In southern Michigan, immature red-tailed hawks tended to remain in winter only when voles were abundant. During relatively long, harsh winters in Michigan, many more young ones were reported in northeastern Mexico. To the opposite extreme, hawks residing as far north as Fairbanks, Alaska, may persevere through the winter on their home territory, as was recorded with one male over three consecutive years. Birds of any age tend to be territorial during winter but may shift ranges whenever food requirements demand it. Wintering birds tend to perch on inconspicuous tree perches, seeking shelter especially if they have a full crop or are in the midst of poor or overly windy weather. Adult wintering red-tails tend to perch more prominently than immatures do, which select lower or more secluded perches. Immatures are often missed in winter bird counts, unless they are being displaced by dominant adults. Generally, though, immatures can seem to recognize that they are less likely to be attacked by adults during winter and can perch surprisingly close to them. Age is the most significant consideration of wintering hawks' hierarchy, but size does factor in, as larger immatures (presumably usually females) are less likely to displaced than smaller ones. Dark adult red-tailed hawks appear to be harder to locate when perched than other red-tails. In Oklahoma, for example, wintering adult Harlan's hawks were rarely engaged in fights or chased by other red-tails. These hawks tended to gather in regional pockets and frequently the same ones occurred year-to-year. In general, migratory behavior is complex and reliant on each individual hawk's decision-making (i.e. whether prey populations are sufficient to entice the hawk to endure prolonged snow cover). During fall migration, departure may occur as soon as late September, but peak movements occur in late October and all of November in the United States, with migration ceasing after mid-December. The northernmost migrants may pass over resident red-tailed hawks in the contiguous United States, while the latter are still in the midst of brooding fledglings. Not infrequently, several autumn hawk watches in Ontario, Quebec, and the northern United States record 4,500–8,900 red-tailed hawks migrating through each fall, with records of up to 15,000 in a season at Hawk Ridge hawk watch in Duluth, Minnesota. Unlike some other Buteo spp., such as Swainson's hawks and broad-winged hawks, red-tailed hawks do not usually migrate in groups, instead passing by one-by-one, and only migrate on days when winds are favorable. Most migrants do not move past southern Mexico in late autumn, but a few North American migrants may annually move as far south as breeding red-tailed hawks happen to occur, i.e. in Central America to as far south Panama. However, a few records were reported of wintering migrant red-tails turning up in Colombia, the first records of them anywhere in South America. Spring northward movements may commence as early as late February, with peak numbers usually occurring in late March and early April. Seasonal counts may include up to 19,000 red-tails in spring at Derby Hill hawk watch, in Oswego, New York, sometimes more than 5,000 are recorded in a day there. The most northerly migratory individuals may not reach breeding grounds until June, even adults.

 

Immature hawks migrate later than adults in spring on average, but not, generally speaking, in autumn. In the northern Great Lakes, immatures return in late May to early June, when adults are already well into their nesting season and must find unoccupied ranges. In Alaska, adults tend to migrate before immatures in early to mid-September, to the contrary of other areas, probably as heavy snowfall begins. Yearlings that were banded in southwestern Idaho stayed for about 2 months after fledging, and then traveled long distances with a strong directional bias, with 9 of 12 recovered southeast of the study area- six of these moved south to coastal lowlands in Mexico] and as far as Guatemala, 4,205 km (2,613 mi) from their initial banding. In California, 35 hawks were banded as nestlings; 26 were recovered at less than 50 miles away, with multidirectional juvenile dispersals. Nestlings banded in Southern California sometimes actually traveled north as far as 1,190 km (740 mi) to Oregon, ranging to the opposite extreme as far as a banded bird from the Sierra Nevadas that moved 1,700 km (1,100 mi) south to Sinaloa. Nestlings banded in Green County, Wisconsin, did not travel very far comparatively by October–November, but by December, recoveries were found in states including Illinois, Iowa, Texas, Louisiana, and Florida.

 

Diet

The red-tailed hawk is carnivorous, and a highly opportunistic feeder. Nearly any small animal they encounter may be viewed as potential food. Their most common prey are small mammals such as rodents and lagomorphs, but they also consume birds, reptiles, fish, amphibians, and invertebrates. Prey varies considerably with regional and seasonal availability, but usually centers on rodents, accounting for up to 85% of a hawk's diet. In total, nearly 500 prey species have been recorded in their diet, almost as many as great horned owls have been recorded as taking. When 27 North American studies are reviewed, mammals make up 65.3% of the diet by frequency, 20.9% by birds, 10.8% by reptiles, 2.8% by invertebrates, and 0.2% by amphibians and fish. The geometric mean body mass of prey taken by red-tailed hawks in North America is about 187 g (6.6 oz) based on a pair of compilation studies from across the continent, regionally varying at least from 43.4 to 361.4 g (1.53 to 12.75 oz). Staple prey (excluding invertebrates) has been claimed to weigh from 15 to 2,114 g (0.033 to 4.661 lb), ranging roughly from the size of a small mouse or lizard to the size of a black-tailed jackrabbit (Lepus californicus). The daily food requirements range from 7 to 11.2% of their own body weight, so that about three voles or the equivalent weight are required daily for a typical range adult.

 

The talons and feet of red-tailed hawks are relatively large for a Buteo hawk; in an average-sized adult red-tail, the "hallux-claw" or rear talon, the largest claw on all accipitrids, averages about 29.7 mm (1.17 in). In fact, the talons of red-tails in some areas averaged of similar size to those of ferruginous hawks which can be considerably heavier and notably larger than those of the only slightly lighter Swainson's hawk. This species may exert an average of about 91 kg/cm2 (1,290 lbf/in2) of pressure through its feet. Owing to its morphology, red-tailed hawks generally can attack larger prey than other Buteo hawks typically can, and are capable of selecting the largest prey of up to their own size available at the time of hunting, though in all likelihood numerically most prey probably weighs on average about 20% of the hawk's own weight (as is typical of many birds of prey). Red-tailed hawks usually hunt by watching for prey activity from a high perch, also known as still hunting. Upon being spotted, prey is dropped down upon by the hawk. Red-tails often select the highest available perches within a given environment, since the greater the height they are at, the less flapping is required and the faster the downward glide they can attain toward nearby prey. If prey is closer than average, the hawk may glide at a steep downward angle with few flaps, if farther than average, it may flap a few swift wingbeats alternating with glides. Perch hunting is the most successful hunting method generally speaking for red-tailed hawks and can account for up to 83% of their daily activities (i.e. in winter). Wintering pairs may join and aseasonally may join forces to group hunt agile prey that they may have trouble catching by themselves, such as tree squirrels. This may consist of stalking opposites sides of a tree, to surround the squirrel and almost inevitably drive the rodent to be captured by one after being flushed by the other hawk.

 

The most common flighted hunting method for red-tail is to cruise around 10 to 50 m (33 to 164 ft) over the ground with flap-and-glide type flight, interspersed occasionally with harrier-like quarters over the ground. This method is less successful than perch hunting, but seems relatively useful for capturing small birds and may show the best results while hunting in hilly country. Hunting red-tailed hawks readily use trees, bushes, or rocks for concealment before making a surprise attack, even showing a partial ability to dodge among trees in an Accipiter-like fashion. Among thick stands of spruce in Alaska, a dodging hunting flight was thought to be unusually important to red-tails living in extensive areas of conifers, with hawks even coming to the ground and walking hurriedly in prey pursuit especially if the prey was large, a similar behavior to goshawks. Additional surprisingly swift aerial hunting has reported in red-tails that habitually hunt bats in Texas. Here, the bat-hunting specialists stooped with half-closed wings, quite falcon-like, plowing through the huge stream of bats exiting their cave roosts, then zooming upwards with a bat in its talons. These hawks also flew parallel closely to the stream, then veer sharply into it and seize a bat. In the neotropics, red-tails have shown the ability to dodge amongst forest canopy whilst hunting. In Kansas, red-tailed hawks were recorded sailing to catch flying insects, a hunting method more typical of a Swainson's hawk. Alternately, they may drop to the ground to forage for insects like grasshoppers and beetles as well as other invertebrates and probably amphibians and fish (except by water in the latter cases). Hunting afoot seems to be particularly prevalent among immatures. Young red-tailed hawks in northeastern Florida were recorded often extracting earthworms from near the surface of the ground and some had a crop full of earthworms after rains. Ground hunting is also quite common on Socorro Island, where no native land mammals occur, and invertebrates are more significant to their overall diet. A red-tailed hawk was observed to incorporate an unconventional killing method, which was drowning a heron immediately after capture. One red-tailed hawk was seen to try to grab a young ground squirrel and, upon missing it, screamed loudly, which in turn caused another young squirrel to break into a run, wherein it was captured. Whether this was an intentional hunting technique needs investigation. Upon capture, smaller prey is taken to a feeding perch, which is almost always lower than a hunting perch. Among small prey, rodents are often swallowed whole, as are shrews and small snakes, while birds are plucked and beheaded. Even prey as small as chipmunks may take two or three bites to consume. Larger mammals of transportable size are at times beheaded and have part of their fur discarded, then leftovers are either stored in a tree or fall to the ground. Large prey, especially if too heavy to transport on the wing, is often dragged to a secluded spot and dismantled in various ways. If they can successfully carry what remains to a low perch, they tend to feed until full and then discard the rest.

 

Mammals

Rodents are certainly the type of prey taken most often by frequency, but their contribution to prey biomass at nests can be regionally low, and the type, variety, and importance of rodent prey can be highly variable. In total, well over 100 rodent species have turned up the diet of red-tailed hawks. Rodents of extremely varied sizes may be hunted by red-tails, with species ranging in size from the 8.2 g (0.29 oz) eastern harvest mouse (Reithrodontomys humulis) to full grown muskrats (Ondatra zibethicus). At times, the red-tailed hawk is thought of as a semi-specialized vole-catcher, but voles are a subsistence food that is more or less taken until larger prey such as rabbits and squirrels can be captured. In an area of Michigan, immature hawks took almost entirely voles but adults were diversified feeders. Indeed, the 44.1 g (1.56 oz) meadow vole (Microtus pennsylvanicus) was the highest frequency prey species in 27 dietary studies across North America, accounting for up to 54% of the food at nests by frequency. It is quite rare for any one species to make up more than half of the food in any dietary study for red-tailed hawks. In total about 9 Microtus species are known in the overall diet, with 5 other voles and lemmings known to be included in their prey spectrum. Another well-represented species was the 27.9 g (0.98 oz) prairie vole (Microtus ochrogaster), which were the primary food, making up 26.4% of a sample of 1322, in eastern Kansas. While crepuscular in primary feeding activity, voles are known to be active both day and night, and so are reliable food for hawks than most non-squirrel rodents, which generally are nocturnal in activity. Indeed, most other microtine rodents are largely inaccessible to red-tailed hawks due to their strongly nocturnal foraging patterns, even though 24 species outside of voles and lemmings are known to be hunted. Woodrats are taken as important supplemental prey in some regions, being considerably larger than most other crictetid rodents, and some numbers of North American deermouse (Peromyscus maniculatus) may turn up. The largest representation of the latter species was contributing 11.9% of the diet in the Great Basin of Utah, making them the second best-represented prey species there. Considering this limited association with nocturnal rodents, the high importance of pocket gophers in the diet of red-tailed hawks is puzzling to many biologists, as these tend to be highly nocturnal and elusive by day, rarely leaving the confines of their burrow. At least 8 species of pocket gopher are included in the prey spectrum (not to mention 5 species of pocket mice). The 110 g (3.9 oz) northern pocket gopher (Thomomys talpoides) is particularly often reported and, by frequency, even turns up as the third most often recorded prey species in 27 American dietary studies. Presumably, hunting of pocket gophers by red-tails, which has possibly never been witnessed, occurs in dim light at first dawn and last light of dusk when they luck upon a gopher out foraging.

 

By far, the most important prey among rodents is squirrels, as they are almost fully diurnal. All told, nearly 50 species from the squirrel family have turned up as food. In particular, where they are distributed, ground squirrels are doubly attractive as a primary food source due to their ground-dwelling habits, as red-tails prefer to attack prey that is terrestrial. There are also many disadvantages to ground squirrels as prey: they can escape quickly to the security of their burrows, they tend to be highly social and they are very effective and fast in response to alarm calls, and a good deal of species enter hibernation that in the coldest climates can range up to a 6 to 9-month period (although those in warmer climates with little to no snowy weather often have brief dormancy and no true hibernation). Nonetheless, red-tailed hawks are devoted predators of ground squirrels, especially catching incautious ones as they go out foraging (which are often younger animals). A multi-year study conducted on San Joaquin Experimental Range in California, seemingly still the largest food study to date done for red-tailed hawks with 4031 items examined, showed that throughout the seasons the 722 g (1.592 lb) California ground squirrel (Otospermophilus beecheyi) was the most significant prey, accounting for 60.8% of the breeding season diet and about 27.2% of the diet for hawks year-around. Because of the extremely high density of red-tailed hawks on this range, some pairs came to specialize in diverse alternate prey, which consisted variously of kangaroo rats, lizards, snakes or chipmunks. One pair apparently lessened competition by focusing on pocket gophers instead despite being near the center of ground squirrel activity. In Snake River NCA, the primary food of red-tailed hawks was the 203.5 g (7.18 oz) Townsend's ground squirrel (Urocitellus townsendii), which made up nearly 21% of the food in 382 prey items across several years despite sharp spikes and crashes of the ground squirrel population there. The same species was the main food of red-tailed hawks in southeastern Washington, making up 31.2% of 170 items. An even closer predatory relationship was reported in the Centennial valley of Montana and south-central Montana, where 45.4% of 194 prey items and 40.2% of 261 items, respectively, of the food of red-tails consisted of the 455.7 g (1.005 lb) Richardson's ground squirrel (Urocitellus richardsonii). Locally in Rochester, Alberta, Richardson's ground squirrel, estimated to average 444 g (15.7 oz), were secondary in number to unidentified small rodents but red-tails in the region killed an estimated 22–60% of the area's ground squirrel, a large dent in the squirrel's population. Further east, ground squirrels are not so reliably distributed, but one study in southern Wisconsin, in one of several quite different dietary studies in that state, the 172.7 g (6.09 oz) thirteen-lined ground squirrel (Ictidomys tridecemlineatus) was the main prey species, making up 29.7% of the diet (from a sample of 165).

 

In Kluane Lake, Yukon, 750 g (1.65 lb) Arctic ground squirrels (Spermophilus parryii) were the main overall food for Harlan's red-tailed hawks, making up 30.8% of a sample of 1074 prey items. When these ground squirrels enter their long hibernation, the breeding Harlan's hawks migrate south for the winter. Nearly as important in Kluane Lake was the 200 g (7.1 oz) American red squirrel (Tamiasciurus hudsonicus), which constituted 29.8% of the above sample. Red squirrels are highly agile dwellers on dense spruce stands, which has caused biologists to ponder how the red-tailed hawks are able to routinely catch them. It is possible that the hawks catch them on the ground such as when squirrels are digging their caches, but theoretically, the dark color of the Harlan's hawks may allow them to ambush the squirrels within the forests locally more effectively. While American red squirrels turn up not infrequently as supplementary prey elsewhere in North America, other tree squirrels seem to be comparatively infrequently caught, at least during the summer breeding season. It is known that pairs of red-tailed hawks will cooperatively hunt tree squirrels at times, probably mostly between late fall and early spring. Fox squirrels (Sciurus niger), the largest of North America's tree squirrels at 800 g (1.8 lb), are relatively common supplemental prey but the lighter, presumably more agile 533 g (1.175 lb) eastern gray squirrel (Sciurus carolinensis) appears to be seldom caught based on dietary studies. While adult marmot may be difficult for red-tailed hawks to catch, young marmots are readily taken in numbers after weaning, such as a high frequency of yellow-bellied marmot (Marmota flaviventris) in Boulder, Colorado. Another grouping of squirrels but at the opposite end of the size spectrum for squirrels, the chipmunks are also mostly supplemental prey but are considered more easily caught than tree squirrels, considering that they are more habitual terrestrial foragers In central Ohio, eastern chipmunks (Tamias striatus), the largest species of chipmunk at an average weight of 96 g (3.4 oz), were actually the leading prey by number, making up 12.3% of a sample of 179 items.

 

Outside of rodents, the most important prey for North American red-tailed hawks is rabbits and hares, of which at least 13 species are included in their prey spectrum. By biomass and reproductive success within populations, these are certain to be their most significant food source (at least in North America). Adult Sylvilagus rabbits known to be hunted by red-tails can range from the 700 g (1.5 lb) brush rabbit (Sylvilagus bachmani) to the Tres Marias rabbit (Sylvilagus graysoni) at 1,470 g (3.24 lb) while all leporids hunted may range the 421.3 g (14.86 oz) pygmy rabbit (Brachylagus idahoensis) to hares and jackrabbits potentially up twice the hawk's own weight. While primarily crepuscular in peak activity, rabbits and hares often foraging both during day and night and so face almost constant predatory pressure from a diverse range of predators. Male red-tailed hawks or pairs which are talented rabbit hunters are likely to have higher than average productivity due to the size and nutrition of the meal ensuring healthy, fast-growing offspring. Most widely reported are the cottontails, which the three most common North America varieties softly grading into mostly allopatric ranges, being largely segregated by habitat preferences where they overlap in distribution. Namely, in descending order of reportage were: the eastern cottontail (Sylvilagus floridanus), the second most widely reported prey species overall in North America and with maximum percentage known in a given study was 26.4% in Oklahoma (out of 958 prey items), the mountain cottontail (Sylvilagus nuttallii), maximum representation being 17.6% out of a sample of 478 in Kaibab Plateau, Arizona and the desert cottontail (Sylvilagus audubonii), maximum representation being 22.4% out of a sample of 326 in west-central Arizona. Black-tailed jackrabbits (Lepus californicus) are even more intensely focused upon as a food source by the hawks found in the west, particularly the Great Basin. With the weight around 2,114 g (4.661 lb), adults of this species is the largest prey routinely hunted by red-tailed hawks. When jackrabbit numbers crash, red-tailed hawk productivity tends to decline as well. In northern Utah, black-tailed jackrabbits made up 55.3% of a sample of 329. Elsewhere, they are usually somewhat secondary by number.

 

In the boreal forests of Canada and Alaska, red-tails are fairly dependent on the snowshoe hare (Lepus americanus), falling somewhere behind the great horned owl and ahead of the Anerican goshawk in their regional reliance on this food source. The hunting preferences of red-tails who rely on snowshoe hares are variable. In Rochester, Alberta, 52% of snowshoe hares caught were adults, such prey estimated to average 1,287 g (2.837 lb), and adults, in some years, were six times more often taken than juvenile hares, which averaged an estimated 560 g (1.23 lb). 1.9–7.1% of adults in the regional population of Rochester were taken by red-tails, while only 0.3–0.8 of juvenile hares were taken by them. Despite their reliance on it, only 4% (against 53.4% of the biomass) of the food by frequency here was made up of hares. On the other hand, in Kluane Lake, Yukon, juvenile hares were taken roughly 11 times more often than adults, despite the larger size of adults here, averaging 1,406.6 g (3.101 lb), and that the overall prey base was less diverse at this more northerly clime. In both Rochester and Kluane Lake, the number of snowshoe hares taken was considerably lower than the number of ground squirrels taken. The differences in average characteristics of snowshoe hares that were hunted may be partially due to habitat (extent of bog openings to dense forest) or topography. Another member of the Lagomorpha order has been found in the diet include juvenile white-tailed jackrabbit (Lepus townsendii) and the much smaller American pika (Ochotona princeps), at 150 g (5.3 oz).

 

A diversity of mammals may be consumed opportunistically outside of the main food groups of rodents and leporids, but usually occur in low numbers. At least five species each are taken of shrews and moles, ranging in size from their smallest mammalian prey, the cinereus (Sorex cinereus) and least shrews (Cryptotis parva), which both weigh about 4.4 g (0.16 oz), to Townsend's mole (Scapanus townsendii), which weighs about 126 g (4.4 oz). A respectable number of the 90 g (3.2 oz) eastern mole (Scalopus aquaticus) were recorded in studies from Oklahoma and Kansas. Four species of bat have been recorded in their foods. The red-tailed hawks local to the large cave colonies of 12.3 g (0.43 oz) Mexican free-tailed bats (Tadarida brasiliensis) in Texas can show surprising agility, some of the same hawks spending their early evening and early morning hours in flight patrolling the cave entrances in order to stoop suddenly on these flighted mammals. Larger miscellaneous mammalian prey are either usually taken as juveniles, like the nine-banded armadillo (Dasypus novemcinctus), or largely as carrion, like the Virginia opossum (Didelphis virginiana). Small carnivorans may be taken, usually consisting of much smaller mustelids, like the least weasels (Mustela nivalis), stoats (Mustela erminea), and long-tailed weasels (Neogale frenata). slightly larger carnivores, such as small Indian mongooses (Herpestes auropunctatus), ringtails (Bassariscus astutus), small American minks (Neovison vison) and even adult striped skunk (Mephitis mephitis), which can be much larger than a fully grown hawk, was reportedly taken by red-tailed hawks. Additionally, red-tailed hawks are considered as potential predators of white-nosed coati (Nasua narica) and kit fox (Vulpes macrotis) Remains of exceptionally large carnivoran species, such as domestic cats (Felis catus), red fox ( Vulpes vulpes) and common raccoon (Procyon lotor) are sometimes found amongst their foods, but most are likely taken as juveniles or consumed only as carrion. Many of these medium-sized carnivorans are probably visited as roadkill, especially during the sparser winter months, but carrion has turned up more widely than previously thought. Some nests have been found (to the occasional "shock" of researchers) with body parts from large domestic stock like sheep (Ovis aries), pigs (Sus domesticus), horses (Equus caballus ) and cattle (Bos taurus) (not to mention wild varieties like deer), which red-tails must visit when freshly dead out on pastures and take a couple of talonfuls of meat. In one instance, a red-tailed hawk was observed to kill a small but seemingly healthy lamb. These are born heavier than most red-tails at 1,500 g (3.3 lb) but in this case, the hawk was scared away before it could consume its kill by the rifle fire of the shepherd who witnessed the instance.

 

Birds

Like most (but not all) Buteo hawks, red-tailed hawks do not primarily hunt birds in most areas, but can take them fairly often whenever they opportune upon some that are vulnerable. Birds are, by far, the most diverse class in the red-tailed hawk's prey spectrum, with well over 200 species known in their foods In most circumstances where birds become the main food of red-tailed hawks, it is in response to ample local populations of galliforms. As these are meaty, mostly terrestrial birds which usually run rather than fly from danger (although all wild species in North America are capable of flight), galliforms are ideal avian prey for red-tails. Some 23 species of galliforms are known to be taken by red-tailed hawks, about a third of these being species introduced by humans. Native quails of all five North American species may expect occasional losses. All 12 species of grouse native to North America are also occasionally included in their prey spectrum. In the state of Wisconsin, two large studies, from Waupun and Green County, found the main prey species to be the ring-necked pheasant (Phasianus colchicus), making up 22.7% of a sample of 176 and 33.8% of a sample of 139, respectively. With a body mass averaging 1,135 g (2.502 lb), adult pheasants are among the largest meals that male red-tails are likely to deliver short of adult rabbits and hares and therefore these nests tend to be relatively productive. Despite being not native to North America, pheasants usually live in a wild state. Chickens (Gallus gallus domesticus) are also taken throughout North America, with all Wisconsin studies also found large numbers of them, making up as much as 14.4% of the diet. Many studies reflect that free-ranging chickens are vulnerable to red-tailed hawks although somewhat lesser numbers are taken by them overall in comparison to nocturnal predators (i.e. owls and foxes) and goshawks. In Rochester, Alberta, fairly large numbers of ruffed grouse (Bonasa umbellus) were taken but relatively more juveniles were taken of this species than the two other main contributors to biomass here, snowshoe hare and Townsend's ground squirrel, as they are fairly independent early on and more readily available. Here the adult grouse was estimated to average 550 g (1.21 lb) against the average juvenile which in mid-summer averaged 170 g (6.0 oz).

 

Beyond galliforms, three other quite different families of birds make the most significant contributions to the red-tailed hawk's avian diet. None of these three families are known as particularly skilled or swift fliers, but are generally small enough that they would generally easily be more nimble in flight. One of these are the woodpeckers, if only for one species, the 131.6 g (4.64 oz) northern flicker (Colaptes auratus), which was the best represented bird species in the diet in 27 North American studies and was even the fourth most often detected prey species of all. Woodpeckers are often a favorite in the diet of large raptors as their relatively slow, undulating flight makes these relatively easy targets. The flicker in particular is a highly numerous species that has similar habitat preferences to red-tailed hawks, preferring fragmented landscapes with trees and openings or parkland-type wooded mosaics, and often forage on the ground for ants, which may make them even more susceptible. Varied other woodpecker species may turn up in their foods, from the smallest to the largest extant in North America, but are much more infrequently detected in dietary studies. Another family relatively often selected prey family are corvids, which despite their relatively large size, formidable mobbing abilities and intelligence are also slower than average fliers for passerines. 14 species of corvid are known to fall prey to red-tailed hawks. In the Kaibab Plateau, the 128 g (4.5 oz) Steller's jay (Cyanocitta stelleri) were the fourth most identified prey species (10.3% of the diet). 453 g (0.999 lb) American crows are also regularly detected supplemental prey in several areas. Even the huge common raven (Corvus corax), at 1,050 g (2.31 lb) at least as large as red-tailed hawk itself, may fall prey to red-tails, albeit very infrequently and only in a well-staged ambush. One of the most surprising heavy contributors are the icterids, despite their slightly smaller size and tendency to travel in large, wary flocks, 12 species are known to be hunted. One species pair, the meadowlarks, are most often selected as they do not flock in the same ways as many other icterids and often come to the ground, throughout their life history, rarely leaving about shrub-height. The 100.7 g (3.55 oz) western meadowlark (Sturnella neglecta), in particular, was the third most often detected bird prey species in North America. Red-winged blackbirds (Agelaius phoeniceus) which are probably too small, at an average weight of 52.4 g (1.85 oz), and fast for a red-tailed hawk to ever chase on the wing (and do travel in huge flocks, especially in winter) are nonetheless also quite often found in their diet, representing up to 8% of the local diet for red-tails. It is possible that males, which are generally bold and often select lofty perches from which to display, are most regularly ambushed. One bird species that often flocks with red-winged blackbirds in winter is even better represented in the red-tail's diet, the non-native 78 g (2.8 oz) European starling (Sturnus vulgaris), being the second most numerous avian prey species and seventh overall in North America. Although perhaps most vulnerable when caught unaware while calling atonally on a perch, a few starlings (or various blackbirds) may be caught by red-tails which test the agile, twisting murmurations of birds by flying conspicuously towards the flock, to intentionally disturb them and possibly detect lagging, injured individual birds that can be caught unlike healthy birds. However, this behavior has been implied rather than verified.

 

Over 50 passerine species from various other families beyond corvids, icterids and starlings are included in the red-tailed hawks' prey spectrum but are caught so infrequently as to generally not warrant individual mention. Non-passerine prey taken infrequently may include but are not limited to pigeons and doves, cuckoos, nightjars, kingfishers and parrots. However, of some interest, is the extreme size range of birds that may be preyed upon. Red-tailed hawks in Caribbean islands seem to catch small birds more frequently due to the paucity of vertebrate prey diversity here.

For the Elysee Palace in Paris, with the request of President Georges Pompidou Yaacov Agam created in 1972 a whole environmental of the Salon with the walls covered with polymorphic murals of changing images a kinetic ceiling, moving transparent colored doors and a kinetic carpet on which he placed a sculpture. This was subsequently transferred to the Pompidou centre in Paris.

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Maize (/meɪz/ MAYZ; Zea mays subsp. mays, from Spanish: maíz after Taino: mahiz), also known as corn (North American and Australian English), is a cereal grain first domesticated by indigenous peoples in southern Mexico about 10,000 years ago. The leafy stalk of the plant produces pollen inflorescences and separate ovuliferous inflorescences called ears that yield kernels or seeds, which are fruits.

 

Maize has become a staple food in many parts of the world, with the total production of maize surpassing that of wheat or rice. In addition to being consumed directly by humans (often in the form of masa), maize is also used for corn ethanol, animal feed and other maize products, such as corn starch and corn syrup. The six major types of maize are dent corn, flint corn, pod corn, popcorn, flour corn, and sweet corn.[5] Sugar-rich varieties called sweet corn are usually grown for human consumption as kernels, while field corn varieties are used for animal feed, various corn-based human food uses (including grinding into cornmeal or masa, pressing into corn oil, and fermentation and distillation into alcoholic beverages like bourbon whiskey), and as chemical feedstocks. Maize is also used in making ethanol and other biofuels.

 

Maize is widely cultivated throughout the world, and a greater weight of maize is produced each year than any other grain. In 2014, total world production was 1.04 billion tonnes. Maize is the most widely grown grain crop throughout the Americas, with 361 million metric tons grown in the United States alone in 2014. Genetically modified maize made up 85% of the maize planted in the United States in 2009. Subsidies in the United States help to account for its high level of cultivation of maize and its position as the largest producer in the world.

 

HISTORY

PRE-COLUMBIAN DEVELOPMENT

Maize is a cultigen; human intervention is required for it to propagate. Whether or not the kernels fall off the cob on their own is a key piece of evidence used in archaeology to distinguish domesticated maize from its naturally-propagating teosinte ancestor. Genetic evidence can also be used to determine when various lineages split.

 

Most historians believe maize was domesticated in the Tehuacán Valley of Mexico. Recent research in the early 21st century has modified this view somewhat; scholars now indicate the adjacent Balsas River Valley of south-central Mexico as the center of domestication.

 

An influential 2002 study by Matsuoka et al. has demonstrated that, rather than the multiple independent domestications model, all maize arose from a single domestication in southern Mexico about 9,000 years ago. The study also demonstrated that the oldest surviving maize types are those of the Mexican highlands. Later, maize spread from this region over the Americas along two major paths. This is consistent with a model based on the archaeological record suggesting that maize diversified in the highlands of Mexico before spreading to the lowlands.

 

Archaeologist Dolores Piperno has said:

 

A large corpus of data indicates that [maize] was dispersed into lower Central America by 7600 BP [5600 BC] and had moved into the inter-Andean valleys of Colombia between 7000 and 6000 BP [5000–4000 BC].

— Dolores Piperno, The Origins of Plant Cultivation and Domestication in the New World Tropics: Patterns, Process, and New Developments

 

Since then, even earlier dates have been published.

 

According to a genetic study by Embrapa, corn cultivation was introduced in South America from Mexico, in two great waves: the first, more than 6000 years ago, spread through the Andes. Evidence of cultivation in Peru has been found dating to about 6700 years ago. The second wave, about 2000 years ago, through the lowlands of South America.

 

The earliest maize plants grew only small, 25-millimetre-long (1 in) corn cobs, and only one per plant. In Jackson Spielvogel's view, many centuries of artificial selection (rather than the current view that maize was exploited by interplanting with teosinte) by the indigenous people of the Americas resulted in the development of maize plants capable of growing several cobs per plant, which were usually several centimetres/inches long each. The Olmec and Maya cultivated maize in numerous varieties throughout Mesoamerica; they cooked, ground and processed it through nixtamalization. It was believed that beginning about 2500 BC, the crop spread through much of the Americas. Research of the 21st century has established even earlier dates. The region developed a trade network based on surplus and varieties of maize crops.

 

Mapuches of south-central Chile cultivated maize along with quinoa and potatoes in pre-Hispanic times; however, potato was the staple food of most Mapuches, "specially in the southern and coastal [Mapuche] territories where maize did not reach maturity". Before the expansion of the Inca Empire maize was traded and transported as far south as 40°19' S in Melinquina, Lácar Department. In that location maize remains were found inside pottery dated to 730 ± 80 BP and 920 ± 60 BP. Probably this maize was brought across the Andes from Chile. The presence of maize in Guaitecas Archipelago (43°55' S), the southernmost outpost of pre-Hispanic agriculture, is reported by early Spanish explorers. However the Spanish may have misidentified the plant.

 

COLUMBIAN EXCHANGE

After the arrival of Europeans in 1492, Spanish settlers consumed maize, and explorers and traders carried it back to Europe and introduced it to other countries. Spanish settlers far preferred wheat bread to maize, cassava, or potatoes. Maize flour could not be substituted for wheat for communion bread, since in Christian belief only wheat could undergo transubstantiation and be transformed into the body of Christ. Some Spaniards worried that by eating indigenous foods, which they did not consider nutritious, they would weaken and risk turning into Indians. "In the view of Europeans, it was the food they ate, even more than the environment in which they lived, that gave Amerindians and Spaniards both their distinctive physical characteristics and their characteristic personalities." Despite these worries, Spaniards did consume maize. Archeological evidence from Florida sites indicate they cultivated it as well.

 

Maize spread to the rest of the world because of its ability to grow in diverse climates. It was cultivated in Spain just a few decades after Columbus's voyages and then spread to Italy, West Africa and elsewhere. Widespread cultivation most likely began in southern Spain in 1525, after which it quickly spread to the rest of the Spanish Empire including its territories in Italy (and, from there, to other Italian states). Maize had many advantages over wheat and barley; it yielded two and a half times the food energy per unit cultivated area, could be harvested in successive years from the same plot of land, and grew in wildly varying altitudes and climates, from relatively dry regions with only 250 mm (10 in) of annual rainfall to damp regions with over 5,000 mm (200 in). By the 17th century it was a common peasant food in Southwestern Europe, including Portugal, Spain, southern France, and Italy. By the 18th century, it was the chief food of the southern French and Italian peasantry, especially in the form of polenta in Italy.

Names

 

The word maize derives from the Spanish form of the indigenous Taíno word for the plant, mahiz. It is known by other names around the world.

 

The word "corn" outside the US, Canada, Australia, and New Zealand refers to any cereal crop, its meaning understood to vary geographically to refer to the local staple. In the United States,[30] Canada, Australia, and New Zealand, corn primarily means maize; this usage started as a shortening of "Indian corn". "Indian corn" primarily means maize (the staple grain of indigenous Americans), but can refer more specifically to multicolored "flint corn" used for decoration.

 

In places outside the US, Canada, Australia, and New Zealand, corn often refers to maize in culinary contexts. The narrower meaning is usually indicated by some additional word, as in sweet corn, sweetcorn, corn on the cob, baby corn, the puffed confection known as popcorn and the breakfast cereal known as corn flakes.

 

In Southern Africa, maize is commonly called mielie (Afrikaans) or mealie (English), words derived from the Portuguese word for maize, milho.

 

Maize is preferred in formal, scientific, and international usage because it refers specifically to this one grain, unlike corn, which has a complex variety of meanings that vary by context and geographic region. Maize is used by agricultural bodies and research institutes such as the FAO and CSIRO. National agricultural and industry associations often include the word maize in their name even in English-speaking countries where the local, informal word is something other than maize; for example, the Maize Association of Australia, the Indian Maize Development Association, the Kenya Maize Consortium and Maize Breeders Network, the National Maize Association of Nigeria, the Zimbabwe Seed Maize Association.

 

STRUCTURE AND PHYSIOLOGY

The maize plant is often 3 m (10 ft) in height, though some natural strains can grow 13 m (43 ft). The stem is commonly composed of 20 internodes of 18 cm (7 in) length. The leaves arise from the nodes, alternately on opposite sides on the stalk. A leaf, which grows from each node, is generally 9 cm (3+1⁄2 in) in width and 120 cm (3 ft 11 in) in length.

 

Ears develop above a few of the leaves in the midsection of the plant, between the stem and leaf sheath, elongating by around 3 mm (1⁄8 in) per day, to a length of 18 cm (7 in) with 60 cm (24 in) being the maximum alleged in the subspecies. They are female inflorescences, tightly enveloped by several layers of ear leaves commonly called husks. Certain varieties of maize have been bred to produce many additional developed ears. These are the source of the "baby corn" used as a vegetable in Asian cuisine.

 

The apex of the stem ends in the tassel, an inflorescence of male flowers. When the tassel is mature and conditions are suitably warm and dry, anthers on the tassel dehisce and release pollen. Maize pollen is anemophilous (dispersed by wind), and because of its large settling velocity, most pollen falls within a few meters of the tassel.

 

Elongated stigmas, called silks, emerge from the whorl of husk leaves at the end of the ear. They are often pale yellow and 18 cm (7 in) in length, like tufts of hair in appearance. At the end of each is a carpel, which may develop into a "kernel" if fertilized by a pollen grain. The pericarp of the fruit is fused with the seed coat referred to as "caryopsis", typical of the grasses, and the entire kernel is often referred to as the "seed". The cob is close to a multiple fruit in structure, except that the individual fruits (the kernels) never fuse into a single mass. The grains are about the size of peas, and adhere in regular rows around a white, pithy substance, which forms the ear. The maximum size of kernels is reputedly 2.5 cm (1 in). An ear commonly holds 600 kernels. They are of various colors: blackish, bluish-gray, purple, green, red, white and yellow. When ground into flour, maize yields more flour with much less bran than wheat does. It lacks the protein gluten of wheat and, therefore, makes baked goods with poor rising capability. A genetic variant that accumulates more sugar and less starch in the ear is consumed as a vegetable and is called sweet corn. Young ears can be consumed raw, with the cob and silk, but as the plant matures (usually during the summer months), the cob becomes tougher and the silk dries to inedibility. By the end of the growing season, the kernels dry out and become difficult to chew without cooking them tender first in boiling water.

 

Planting density affects multiple aspects of maize. Modern farming techniques in developed countries usually rely on dense planting, which produces one ear per stalk. Stands of silage maize are yet denser,[citation needed] and achieve a lower percentage of ears and more plant matter.

 

Maize is a facultative short-day plant and flowers in a certain number of growing degree days > 10 °C (50 °F) in the environment to which it is adapted. The magnitude of the influence that long nights have on the number of days that must pass before maize flowers is genetically prescribed and regulated by the phytochrome system.

Photoperiodicity can be eccentric in tropical cultivars such that the long days characteristic of higher latitudes allow the plants to grow so tall that they do not have enough time to produce seed before being killed by frost. These attributes, however, may prove useful in using tropical maize for biofuels.

 

Immature maize shoots accumulate a powerful antibiotic substance, 2,4-dihydroxy-7-methoxy-1,4-benzoxazin-3-one (DIMBOA). DIMBOA is a member of a group of hydroxamic acids (also known as benzoxazinoids) that serve as a natural defense against a wide range of pests, including insects, pathogenic fungi and bacteria. DIMBOA is also found in related grasses, particularly wheat. A maize mutant (bx) lacking DIMBOA is highly susceptible to attack by aphids and fungi. DIMBOA is also responsible for the relative resistance of immature maize to the European corn borer (family Crambidae). As maize matures, DIMBOA levels and resistance to the corn borer decline.

 

Because of its shallow roots, maize is susceptible to droughts, intolerant of nutrient-deficient soils, and prone to be uprooted by severe winds.

 

While yellow maizes derive their color from lutein and zeaxanthin, in red-colored maizes, the kernel coloration is due to anthocyanins and phlobaphenes. These latter substances are synthesized in the flavonoids synthetic pathway from polymerization of flavan-4-ols by the expression of maize pericarp color1 (p1) gene which encodes an R2R3 myb-like transcriptional activator of the A1 gene encoding for the dihydroflavonol 4-reductase (reducing dihydroflavonols into flavan-4-ols) while another gene (Suppressor of Pericarp Pigmentation 1 or SPP1) acts as a suppressor. The p1 gene encodes an Myb-homologous transcriptional activator of genes required for biosynthesis of red phlobaphene pigments, while the P1-wr allele specifies colorless kernel pericarp and red cobs, and unstable factor for orange1 (Ufo1) modifies P1-wr expression to confer pigmentation in kernel pericarp, as well as vegetative tissues, which normally do not accumulate significant amounts of phlobaphene pigments. The maize P gene encodes a Myb homolog that recognizes the sequence CCT/AACC, in sharp contrast with the C/TAACGG bound by vertebrate Myb proteins.

 

The ear leaf is the leaf most closely associated with a particular developing ear. This leaf and above contribute 70%[57] to 75% to 90% of grain fill. Therefore fungicide application is most important in that region in most disease environments.

 

ABNORMAL FLOWERS

Maize flowers may sometimes exhibit mutations that lead to the formation of female flowers in the tassel. These mutations, ts4 and Ts6, prohibit the development of the stamen while simultaneously promoting pistil development. This may cause inflorescences containing both male and female flowers, or hermaphrodite flowers.

 

GENETICS

Maize is an annual grass in the family Gramineae, which includes such plants as wheat, rye, barley, rice, sorghum, and sugarcane. There are two major species of the genus Zea (out of six total): Zea mays (maize) and Zea diploperennis, which is a perennial type of teosinte. The annual teosinte variety called Zea mays mexicana is the closest botanical relative to maize. It still grows in the wild as an annual in Mexico and Guatemala.

 

Many forms of maize are used for food, sometimes classified as various subspecies related to the amount of starch each has:

 

Flour corn: Zea mays var. amylacea

Popcorn: Zea mays var. everta

Dent corn : Zea mays var. indentata

Flint corn: Zea mays var. indurata

Sweet corn: Zea mays var. saccharata and Zea mays var. rugosa

Waxy corn: Zea mays var. ceratina

Amylomaize: Zea mays

Pod corn: Zea mays var. tunicata Larrañaga ex A. St. Hil.

Striped maize: Zea mays var. japonica

 

This system has been replaced (though not entirely displaced) over the last 60 years by multivariable classifications based on ever more data. Agronomic data were supplemented by botanical traits for a robust initial classification, then genetic, cytological, protein and DNA evidence was added. Now, the categories are forms (little used), races, racial complexes, and recently branches.

 

Maize is a diploid with 20 chromosomes (n=10). The combined length of the chromosomes is 1500 cM. Some of the maize chromosomes have what are known as "chromosomal knobs": highly repetitive heterochromatic domains that stain darkly. Individual knobs are polymorphic among strains of both maize and teosinte.

 

Barbara McClintock used these knob markers to validate her transposon theory of "jumping genes", for which she won the 1983 Nobel Prize in Physiology or Medicine. Maize is still an important model organism for genetics and developmental biology today.

 

The centromeres have two types of structural components, both of which are found only in the centromeres: Large arrays of CentC, a short satellite DNA; and a few of a family of retrotransposons. The B chromosome, unlike the others, contains an additional repeat which extends into neighboring areas of the chromosome. Centromeres can accidentally shrink during division and still function, although it is thought this will fail if it shrinks below a few hundred kilobase. Kinetochores contain RNA originating from centromeres. Centromere regions can become inactive, and can continue in that state if the chromosome still has another active one.

 

The Maize Genetics Cooperation Stock Center, funded by the USDA Agricultural Research Service and located in the Department of Crop Sciences at the University of Illinois at Urbana-Champaign, is a stock center of maize mutants. The total collection has nearly 80,000 samples. The bulk of the collection consists of several hundred named genes, plus additional gene combinations and other heritable variants. There are about 1000 chromosomal aberrations (e.g., translocations and inversions) and stocks with abnormal chromosome numbers (e.g., tetraploids). Genetic data describing the maize mutant stocks as well as myriad other data about maize genetics can be accessed at MaizeGDB, the Maize Genetics and Genomics Database.

 

In 2005, the US National Science Foundation (NSF), Department of Agriculture (USDA) and the Department of Energy (DOE) formed a consortium to sequence the B73 maize genome. The resulting DNA sequence data was deposited immediately into GenBank, a public repository for genome-sequence data. Sequences and genome annotations have also been made available throughout the project's lifetime at the project's official site.

 

Primary sequencing of the maize genome was completed in 2008. On November 20, 2009, the consortium published results of its sequencing effort in Science. The genome, 85% of which is composed of transposons, was found to contain 32,540 genes (By comparison, the human genome contains about 2.9 billion bases and 26,000 genes). Much of the maize genome has been duplicated and reshuffled by helitrons—group of rolling circle transposons.

 

In Z. mays and various other angiosperms the MADS-box motif is involved in floral development. Early study in several angiosperm models including Z. mays was the beginning of research into the molecular evolution of floral structure in general, as well as their role in nonflowering plants.

 

EVOLUTION

As with many plants and animals, Z. mays has a positive correlation between effective population size and the magnitude of selection pressure. Z. m. having an EPS of ~650,000, it clusters with others of about the same EPS, and has 79% of its amino acid sites under selection.

 

Recombination is a significant source of diversity in Z. mays. (Note that this finding supersedes previous studies which showed no such correlation.)

 

This recombination/diversity effect is seen throughout plants but is also found to not occur – or not as strongly – in regions of high gene density. This is likely the reason that domesticated Z. mays has not seen as much of an increase in diversity within areas of higher density as in regions of lower density, although there is more evidence in other plants.

 

Some lines of maize have undergone ancient polyploidy events, starting 11m years ago. Over that time ~72% of polyploid duplicated genes have been retained, which is higher than other plants with older polyploidy events. Thus maize may be due to lose more duplicate genes as time goes along, similar to the course followed by the genomes of other plants. If so - if gene loss has merely not occurred yet - that could explain the lack of observed positive selection and lower negative selection which are observed in otherwise similar plants, i.e. also naturally outcrossing and with similar effective population sizes.

 

Ploidy does not appear to influence EPS or magnitude of selection effect in maize.

 

BREEDING

Maize reproduces sexually each year. This randomly selects half the genes from a given plant to propagate to the next generation, meaning that desirable traits found in the crop (like high yield or good nutrition) can be lost in subsequent generations unless certain techniques are used.

 

Maize breeding in prehistory resulted in large plants producing large ears. Modern breeding began with individuals who selected highly productive varieties in their fields and then sold seed to other farmers. James L. Reid was one of the earliest and most successful developing Reid's Yellow Dent in the 1860s. These early efforts were based on mass selection. Later breeding efforts included ear to row selection (C. G. Hopkins c. 1896), hybrids made from selected inbred lines (G. H. Shull, 1909), and the highly successful double cross hybrids using four inbred lines (D. F. Jones c. 1918, 1922). University supported breeding programs were especially important in developing and introducing modern hybrids. By the 1930s, companies such as Pioneer devoted to production of hybrid maize had begun to influence long-term development. Internationally important seed banks such as the International Maize and Wheat Improvement Center (CIMMYT) and the US bank at the Maize Genetics Cooperation Stock Center University of Illinois at Urbana-Champaign maintain germplasm important for future crop development.

 

Since the 1940s the best strains of maize have been first-generation hybrids made from inbred strains that have been optimized for specific traits, such as yield, nutrition, drought, pest and disease tolerance. Both conventional cross-breeding and genetic modification have succeeded in increasing output and reducing the need for cropland, pesticides, water and fertilizer. There is conflicting evidence to support the hypothesis that maize yield potential has increased over the past few decades. This suggests that changes in yield potential are associated with leaf angle, lodging resistance, tolerance of high plant density, disease/pest tolerance, and other agronomic traits rather than increase of yield potential per individual plant.

 

Tropical landraces remain an important and underutilized source of resistance alleles for for disease and for herbivores. Notable discoveries of rare alleles for this purpose were made by Dao et al 2014 and Sood et al 2014.

 

GLOBAL PROGRAM

CIMMYT operates a conventional breeding program to provide optimized strains. The program began in the 1980s. Hybrid seeds are distributed in Africa by the Drought Tolerant Maize for Africa project.

 

GENETIC MODIFICATION

Genetically modified (GM) maize was one of the 26 GM crops grown commercially in 2016. The vast majority of this is Bt maize. Grown since 1997 in the United States and Canada, 92% of the US maize crop was genetically modified in 2016 and 33% of the worldwide maize crop was GM in 2016. As of 2011, Herbicide-tolerant maize varieties were grown in Argentina, Australia, Brazil, Canada, China, Colombia, El Salvador, the European Union, Honduras, Japan, Korea, Malaysia, Mexico, New Zealand, Philippines, the Russian Federation, Singapore, South Africa, Taiwan, Thailand, and the United States. Insect-resistant maize was grown in Argentina, Australia, Brazil, Canada, Chile, China, Colombia, Egypt, the European Union, Honduras, Japan, Korea, Malaysia, Mexico, New Zealand, Philippines, South Africa, Switzerland, Taiwan, the United States, and Uruguay.

 

In September 2000, up to $50 million worth of food products were recalled due to the presence of Starlink genetically modified corn, which had been approved only for animal consumption and had not been approved for human consumption, and was subsequently withdrawn from the market.

 

ORIGIN

Maize is the domesticated variant of teosinte. The two plants have dissimilar appearance, maize having a single tall stalk with multiple leaves and teosinte being a short, bushy plant. The difference between the two is largely controlled by differences in just two genes, called grassy tillers-1 (gt1, A0A317YEZ1) and teosinte branched-1 (tb1, Q93WI2).

 

Several theories had been proposed about the specific origin of maize in Mesoamerica:

 

It is a direct domestication of a Mexican annual teosinte, Zea mays ssp. parviglumis, native to the Balsas River valley in south-eastern Mexico, with up to 12% of its genetic material obtained from Zea mays ssp. mexicana through introgression.

It has been derived from hybridization between a small domesticated maize (a slightly changed form of a wild maize) and a teosinte of section Luxuriantes, either Z. luxurians or Z. diploperennis.

It has undergone two or more domestications either of a wild maize or of a teosinte. (The term "teosinte" describes all species and subspecies in the genus Zea, excluding Zea mays ssp. mays.)

It has evolved from a hybridization of Z. diploperennis by Tripsacum dactyloides.

 

In the late 1930s, Paul Mangelsdorf suggested that domesticated maize was the result of a hybridization event between an unknown wild maize and a species of Tripsacum, a related genus. This theory about the origin of maize has been refuted by modern genetic testing, which refutes Mangelsdorf's model and the fourth listed above. 

 

The teosinte origin theory was proposed by the Russian botanist Nikolai Ivanovich Vavilov in 1931 and the later American Nobel Prize-winner George Beadle in 1932.: 10  It is supported experimentally and by recent studies of the plants' genomes. Teosinte and maize can cross-breed and produce fertile offspring. A number of questions remain concerning the species, among them:

 

how the immense diversity of the species of sect. Zea originated,

how the tiny archaeological specimens of 3500–2700 BC could have been selected from a teosinte, and

how domestication could have proceeded without leaving remains of teosinte or maize with teosintoid traits earlier than the earliest known until recently, dating from ca. 1100 BC.

 

The domestication of maize is of particular interest to researchers—archaeologists, geneticists, ethnobotanists, geographers, etc. The process is thought by some to have started 7,500 to 12,000 years ago. Research from the 1950s to 1970s originally focused on the hypothesis that maize domestication occurred in the highlands between the states of Oaxaca and Jalisco, because the oldest archaeological remains of maize known at the time were found there.

Connection with 'parviglumis' subspecies

Genetic studies, published in 2004 by John Doebley, identified Zea mays ssp. parviglumis, native to the Balsas River valley in Mexico's southwestern highlands, and also known as Balsas teosinte, as being the crop wild relative that is genetically most similar to modern maize. This was confirmed by further studies, which refined this hypothesis somewhat. Archaeobotanical studies, published in 2009, point to the middle part of the Balsas River valley as the likely location of early domestication; this river is not very long, so these locations are not very distant. Stone milling tools with maize residue have been found in an 8,700 year old layer of deposits in a cave not far from Iguala, Guerrero.

 

Doebley was part of the team that first published, in 2002, that maize had been domesticated only once, about 9,000 years ago, and then spread throughout the Americas.

 

A primitive corn was being grown in southern Mexico, Central America, and northern South America 7,000 years ago. Archaeological remains of early maize ears, found at Guila Naquitz Cave in the Oaxaca Valley, date back roughly 6,250 years; the oldest ears from caves near Tehuacan, Puebla, 5,450 B.P.

 

Maize pollen dated to 7,300 B.P. from San Andres, Tabasco, on the Caribbean coast has also been recovered.

 

As maize was introduced to new cultures, new uses were developed and new varieties selected to better serve in those preparations. Maize was the staple food, or a major staple – along with squash, Andean region potato, quinoa, beans, and amaranth – of most pre-Columbian North American, Mesoamerican, South American, and Caribbean cultures. The Mesoamerican civilization, in particular, was deeply interrelated with maize. Its traditions and rituals involved all aspects of maize cultivation – from the planting to the food preparation. Maize formed the Mesoamerican people's identity.

 

It is unknown what precipitated its domestication, because the edible portion of the wild variety is too small, and hard to obtain, to be eaten directly, as each kernel is enclosed in a very hard bivalve shell.

 

In 1939, George Beadle demonstrated that the kernels of teosinte are readily "popped" for human consumption, like modern popcorn.[91] Some have argued it would have taken too many generations of selective breeding to produce large, compressed ears for efficient cultivation. However, studies of the hybrids readily made by intercrossing teosinte and modern maize suggest this objection is not well founded.

 

SPREADING TO THE NORTH

Around 4,500 ago, maize began to spread to the north; it was first cultivated in what is now the United States at several sites in New Mexico and Arizona, about 4,100 ago.

 

During the first millennium AD, maize cultivation spread more widely in the areas north. In particular, the large-scale adoption of maize agriculture and consumption in eastern North America took place about A.D. 900. Native Americans cleared large forest and grassland areas for the new crop.

 

In 2005, research by the USDA Forest Service suggested that the rise in maize cultivation 500 to 1,000 years ago in what is now the southeastern United States corresponded with a decline of freshwater mussels, which are very sensitive to environmental changes.

 

CULTIVATION

PLANTING

Because it is cold-intolerant, in the temperate zones maize must be planted in the spring. Its root system is generally shallow, so the plant is dependent on soil moisture. As a plant that uses C4 carbon fixation, maize is a considerably more water-efficient crop than plants that use C3 carbon fixation such as alfalfa and soybeans. Maize is most sensitive to drought at the time of silk emergence, when the flowers are ready for pollination. In the United States, a good harvest was traditionally predicted if the maize was "knee-high by the Fourth of July", although modern hybrids generally exceed this growth rate. Maize used for silage is harvested while the plant is green and the fruit immature. Sweet corn is harvested in the "milk stage", after pollination but before starch has formed, between late summer and early to mid-autumn. Field maize is left in the field until very late in the autumn to thoroughly dry the grain, and may, in fact, sometimes not be harvested until winter or even early spring. The importance of sufficient soil moisture is shown in many parts of Africa, where periodic drought regularly causes maize crop failure and consequent famine. Although it is grown mainly in wet, hot climates, it has been said to thrive in cold, hot, dry or wet conditions, meaning that it is an extremely versatile crop.

 

Maize was planted by the Native Americans in hills, in a complex system known to some as the Three Sisters. Maize provided support for beans, and the beans provided nitrogen derived from nitrogen-fixing rhizobia bacteria which live on the roots of beans and other legumes; and squashes provided ground cover to stop weeds and inhibit evaporation by providing shade over the soil. This method was replaced by single species hill planting where each hill 60–120 cm (2 ft 0 in–3 ft 11 in) apart was planted with three or four seeds, a method still used by home gardeners. A later technique was "checked maize", where hills were placed

 

1 m (40 in) apart in each direction, allowing cultivators to run through the field in two directions. In more arid lands, this was altered and seeds were planted in the bottom of 10–12 cm (4–4+1⁄2 in) deep furrows to collect water. Modern technique plants maize in rows which allows for cultivation while the plant is young, although the hill technique is still used in the maize fields of some Native American reservations. When maize is planted in rows, it also allows for planting of other crops between these rows to make more efficient use of land space.

 

In most regions today, maize grown in residential gardens is still often planted manually with a hoe, whereas maize grown commercially is no longer planted manually but rather is planted with a planter. In North America, fields are often planted in a two-crop rotation with a nitrogen-fixing crop, often alfalfa in cooler climates and soybeans in regions with longer summers. Sometimes a third crop, winter wheat, is added to the rotation.

 

Many of the maize varieties grown in the United States and Canada are hybrids. Often the varieties have been genetically modified to tolerate glyphosate or to provide protection against natural pests. Glyphosate is an herbicide which kills all plants except those with genetic tolerance. This genetic tolerance is very rarely found in nature.

 

In the midwestern United States, low-till or no-till farming techniques are usually used. In low-till, fields are covered once, maybe twice, with a tillage implement either ahead of crop planting or after the previous harvest. The fields are planted and fertilized. Weeds are controlled through the use of herbicides, and no cultivation tillage is done during the growing season. This technique reduces moisture evaporation from the soil, and thus provides more moisture for the crop. The technologies mentioned in the previous paragraph enable low-till and no-till farming. Weeds compete with the crop for moisture and nutrients, making them undesirable.

 

HARVESTING

Before the 20th century, all maize harvesting was by manual labour, by grazing, or by some combination of those. Whether the ears were hand-picked and the stover was grazed, or the whole plant was cut, gathered, and shocked, people and livestock did all the work. Between the 1890s and the 1970s, the technology of maize harvesting expanded greatly. Today, all such technologies, from entirely manual harvesting to entirely mechanized, are still in use to some degree, as appropriate to each farm's needs, although the thoroughly mechanized versions predominate, as they offer the lowest unit costs when scaled to large farm operations. For small farms, their unit cost can be too high, as their higher fixed cost cannot be amortized over as many units.[citation needed]

 

Before World War II, most maize in North America was harvested by hand. This involved a large number of workers and associated social events (husking or shucking bees). From the 1890s onward, some machinery became available to partially mechanize the processes, such as one- and two-row mechanical pickers (picking the ear, leaving the stover) and corn binders, which are reaper-binders designed specifically for maize (for example, Video on YouTube). The latter produce sheaves that can be shocked. By hand or mechanical picker, the entire ear is harvested, which then requires a separate operation of a maize sheller to remove the kernels from the ear. Whole ears of maize were often stored in corn cribs, and these whole ears are a sufficient form for some livestock feeding use. Today corn cribs with whole ears, and corn binders, are less common because most modern farms harvest the grain from the field with a combine and store it in bins. The combine with a corn head (with points and snap rolls instead of a reel) does not cut the stalk; it simply pulls the stalk down. The stalk continues downward and is crumpled into a mangled pile on the ground, where it usually is left to become organic matter for the soil. The ear of maize is too large to pass between slots in a plate as the snap rolls pull the stalk away, leaving only the ear and husk to enter the machinery. The combine separates the husk and the cob, keeping only the kernels.

When maize is a silage crop, the entire plant is usually chopped at once with a forage harvester (chopper) and ensiled in silos or polymer wrappers. Ensiling of sheaves cut by a corn binder was formerly common in some regions but has become uncommon. For storing grain in bins, the moisture of the grain must be sufficiently low to avoid spoiling. If the moisture content of the harvested grain is too high, grain dryers are used to reduce the moisture content by blowing heated air through the grain. This can require large amounts of energy in the form of combustible gases (propane or natural gas) and electricity to power the blowers.

 

PRODUCTION

Maize is widely cultivated throughout the world, and a greater weight of maize is produced each year than any other grain. In 2018, total world production was 1.15 billion tonnes, led by the United States with 34.2% of the total (table). China produced 22.4% of the global total.

 

UNITED STATES

In 2016, maize production was forecast to be over 380 million metric tons (15 billion bushels), an increase of 11% over 2014 American production. Based on conditions as of August 2016, the expected yield would be the highest ever for the United States. The area of harvested maize was forecast to be 35 million hectares (87 million acres), an increase of 7% over 2015. Maize is especially popular in Midwestern states such as Indiana, Iowa, and Illinois; in the latter, it was named the state's official grain in 2017.

 

STORAGE

Drying is vital to prevent or at least reduce mycotoxin contamination. Aspergillus and Fusarium spp. are the most common mycotoxin sources, but there are others. Altogether maize contaminants are so common, and this crop is so economically important, that maize mycotoxins are among the most important in agriculture in general.

 

USES

HUMAN FOOD

Maize and cornmeal (ground dried maize) constitute a staple food in many regions of the world. Maize is used to produce cornstarch, a common ingredient in home cooking and many industrialized food products. Maize starch can be hydrolyzed and enzymatically treated to produce syrups, particularly high fructose corn syrup, a sweetener; and also fermented and distilled to produce grain alcohol. Grain alcohol from maize is traditionally the source of Bourbon whiskey. Corn flour is used to make cornbread and other baked products.

 

In prehistoric times Mesoamerican women used a metate to process maize into ground cornmeal, allowing the preparation of foods that were more calorie dense than popcorn. After ceramic vessels were invented the Olmec people began to cook maize together with beans, improving the nutritional value of the staple meal. Although maize naturally contains niacin, an important nutrient, it was not bioavailable without the process of nixtamalization. The Maya used nixtamal meal to make varieties of porridges and tamales. The process was later used in the cuisine of the American South to prepare corn for grits and hominy.

 

Maize is a staple of Mexican cuisine. Masa (cornmeal treated with limewater) is the main ingredient for tortillas, atole and many other dishes of Central American food. It is the main ingredient of corn tortilla, tamales, pozole, atole and all the dishes based on them, like tacos, quesadillas, chilaquiles, enchiladas, tostadas and many more. In Mexico the fungus of maize, known as huitlacoche, is considered a delicacy.

 

Coarse maize meal is made into a thick porridge in many cultures: from the polenta of Italy, the angu of Brazil, the mămăligă of Romania, to cornmeal mush in the US (or hominy grits in the South) or the food called mieliepap in South Africa and sadza, nshima, ugali and other names in other parts of Africa. Introduced into Africa by the Portuguese in the 16th century, maize has become Africa's most important staple food crop. These are commonly eaten in the Southeastern United States, foods handed down from Native Americans, who called the dish sagamite.

 

Maize can also be harvested and consumed in the unripe state, when the kernels are fully grown but still soft. Unripe maize must usually be cooked to become palatable; this may be done by simply boiling or roasting the whole ears and eating the kernels right off the cob. Sweet corn, a genetic variety that is high in sugars and low in starch, is usually consumed in the unripe state. Such corn on the cob is a common dish in the United States, Canada, United Kingdom, Cyprus, some parts of South America, and the Balkans, but virtually unheard of in some European countries. Corn on the cob was hawked on the streets of early 19th-century New York City by poor, barefoot "Hot Corn Girls", who were thus the precursors of hot dog carts, churro wagons, and fruit stands seen on the streets of big cities today.

 

Within the United States, the usage of maize for human consumption constitutes only around 1/40th of the amount grown in the country. In the United States and Canada, maize is mostly grown to feed livestock, as forage, silage (made by fermentation of chopped green cornstalks), or grain. Maize meal is also a significant ingredient of some commercial animal food products.

 

NUTRITIONAL VALUE

Raw, yellow, sweet maize kernels are composed of 76% water, 19% carbohydrates, 3% protein, and 1% fat (table). In a 100-gram serving, maize kernels provide 86 calories and are a good source (10–19% of the Daily Value) of the B vitamins, thiamin, niacin (but see Pellagra warning below), pantothenic acid (B5) and folate (right table for raw, uncooked kernels, USDA Nutrient Database). In moderate amounts, they also supply dietary fiber and the essential minerals, magnesium and phosphorus whereas other nutrients are in low amounts (table).

 

Maize has suboptimal amounts of the essential amino acids tryptophan and lysine, which accounts for its lower status as a protein source. However, the proteins of beans and legumes complement those of maize.

 

FEED AND FODDER FOR LIVESTOCK

Maize is a major source of both grain feed and fodder for livestock. It is fed to the livestock in various ways. When it is used as a grain crop, the dried kernels are used as feed. They are often kept on the cob for storage in a corn crib, or they may be shelled off for storage in a grain bin. The farm that consumes the feed may produce it, purchase it on the market, or some of both. When the grain is used for feed, the rest of the plant (the corn stover) can be used later as fodder, bedding (litter), or soil amendment. When the whole maize plant (grain plus stalks and leaves) is used for fodder, it is usually chopped all at once and ensilaged, as digestibility and palatability are higher in the ensilaged form than in the dried form. Maize silage is one of the most valuable forages for ruminants. Before the advent of widespread ensilaging, it was traditional to gather the corn into shocks after harvesting, where it dried further. With or without a subsequent move to the cover of a barn, it was then stored for weeks to several months until fed to the livestock. Today ensilaging can occur not only in siloes but also in silage wrappers. However, in the tropics, maize can be harvested year-round and fed as green forage to the animals.

 

CHEMICALS

Starch from maize can also be made into plastics, fabrics, adhesives, and many other chemical products.

 

The corn steep liquor, a plentiful watery byproduct of maize wet milling process, is widely used in the biochemical industry and research as a culture medium to grow many kinds of microorganisms.

 

Chrysanthemin is found in purple corn and is used as a food coloring.

 

BIO-FUEL

"Feed maize" is being used increasingly for heating; specialized corn stoves (similar to wood stoves) are available and use either feed maize or wood pellets to generate heat. Maize cobs are also used as a biomass fuel source. Maize is relatively cheap and home-heating furnaces have been developed which use maize kernels as a fuel. They feature a large hopper that feeds the uniformly sized maize kernels (or wood pellets or cherry pits) into the fire.[citation needed]

 

Maize is increasingly used as a feedstock for the production of ethanol fuel.[120] When considering where to construct an ethanol plant, one of the site selection criteria is to ensure there is locally available feedstock. Ethanol is mixed with gasoline to decrease the amount of pollutants emitted when used to fuel motor vehicles. High fuel prices in mid-2007 led to higher demand for ethanol, which in turn led to higher prices paid to farmers for maize. This led to the 2007 harvest being one of the most profitable maize crops in modern history for farmers. Because of the relationship between fuel and maize, prices paid for the crop now tend to track the price of oil.

 

The price of food is affected to a certain degree by the use of maize for biofuel production. The cost of transportation, production, and marketing are a large portion (80%) of the price of food in the United States. Higher energy costs affect these costs, especially transportation. The increase in food prices the consumer has been seeing is mainly due to the higher energy cost. The effect of biofuel production on other food crop prices is indirect. Use of maize for biofuel production increases the demand, and therefore price of maize. This, in turn, results in farm acreage being diverted from other food crops to maize production. This reduces the supply of the other food crops and increases their prices.

 

Maize is widely used in Germany as a feedstock for biogas plants. Here the maize is harvested, shredded then placed in silage clamps from which it is fed into the biogas plants. This process makes use of the whole plant rather than simply using the kernels as in the production of fuel ethanol.

 

A biomass gasification power plant in Strem near Güssing, Burgenland, Austria, began in 2005. Research is being done to make diesel out of the biogas by the Fischer Tropsch method.

 

Increasingly, ethanol is being used at low concentrations (10% or less) as an additive in gasoline (gasohol) for motor fuels to increase the octane rating, lower pollutants, and reduce petroleum use (what is nowadays also known as "biofuels" and has been generating an intense debate regarding the human beings' necessity of new sources of energy, on the one hand, and the need to maintain, in regions such as Latin America, the food habits and culture which has been the essence of civilizations such as the one originated in Mesoamerica; the entry, January 2008, of maize among the commercial agreements of NAFTA has increased this debate, considering the bad labor conditions of workers in the fields, and mainly the fact that NAFTA "opened the doors to the import of maize from the United States, where the farmers who grow it receive multimillion-dollar subsidies and other government supports. ... According to OXFAM UK, after NAFTA went into effect, the price of maize in Mexico fell 70% between 1994 and 2001. The number of farm jobs dropped as well: from 8.1 million in 1993 to 6.8 million in 2002. Many of those who found themselves without work were small-scale maize growers."). However, introduction in the northern latitudes of the US of tropical maize for biofuels, and not for human or animal consumption, may potentially alleviate this.

 

COMMODITY

Maize is bought and sold by investors and price speculators as a tradable commodity using corn futures contracts. These "futures" are traded on the Chicago Board of Trade (CBOT) under ticker symbol C. They are delivered every year in March, May, July, September, and December.

Ornamental and other uses

 

Some forms of the plant are occasionally grown for ornamental use in the garden. For this purpose, variegated and colored leaf forms as well as those with colorful ears are used.

 

Corncobs can be hollowed out and treated to make inexpensive smoking pipes, first manufactured in the United States in 1869.

 

An unusual use for maize is to create a "corn maze" (or "maize maze") as a tourist attraction. The idea of a maize maze was introduced by the American Maze Company who created a maze in Pennsylvania in 1993. Traditional mazes are most commonly grown using yew hedges, but these take several years to mature. The rapid growth of a field of maize allows a maze to be laid out using GPS at the start of a growing season and for the maize to grow tall enough to obstruct a visitor's line of sight by the start of the summer. In Canada and the US, these are popular in many farming communities.

 

Maize kernels can be used in place of sand in a sandboxlike enclosure for children's play.

 

Stigmas from female maize flowers, popularly called corn silk, are sold as herbal supplements.

 

Maize is used as a fish bait, called "dough balls". It is particularly popular in Europe for coarse fishing.

 

Additionally, feed corn is sometimes used by hunters to bait animals such as deer or wild hogs.

 

UNITED STATES USAGE BREAKDOWN

The breakdown of usage of the 12.1-billion-bushel (307-million-tonne) 2008 US maize crop was as follows, according to the World Agricultural Supply and Demand Estimates Report by the USDA.In the US since 2009/2010, maize feedstock use for ethanol production has somewhat exceeded direct use for livestock feed; maize use for fuel ethanol was 5,130 million bushels (130 million tonnes) in the 2013/2014 marketing year.A fraction of the maize feedstock dry matter used for ethanol production is usefully recovered as DDGS (dried distillers grains with solubles). In the 2010/2011 marketing year, about 29.1 million tonnes of DDGS were fed to US livestock and poultry. Because starch utilization in fermentation for ethanol production leaves other grain constituents more concentrated in the residue, the feed value per kg of DDGS, with regard to ruminant-metabolizable energy and protein, exceeds that of the grain. Feed value for monogastric animals, such as swine and poultry, is somewhat lower than for ruminants.

 

HAZARDS

PELLAGRA

When maize was first introduced into farming systems other than those used by traditional native-American peoples, it was generally welcomed with enthusiasm for its productivity. However, a widespread problem of malnutrition soon arose wherever maize was introduced as a staple food. This was a mystery, since these types of malnutrition were not normally seen among the indigenous Americans, for whom maize was the principal staple food.

 

It was eventually discovered that the indigenous Americans had learned to soak maize in alkali — water (the process now known as nixtamalization) — made with ashes and lime (calcium oxide) since at least 1200–1500 BC by Mesoamericans. They did this to liberate the corn hulls, but (unbeknownst to natives or colonists) it coincidentally liberates the B-vitamin niacin, the lack of which was the underlying cause of the condition known as pellagra.

 

Maize was introduced into the diet of non-indigenous Americans without the necessary cultural knowledge acquired over thousands of years in the Americas. In the late 19th century, pellagra reached epidemic proportions in parts of the southern US, as medical researchers debated two theories for its origin: the deficiency theory (which was eventually shown to be true) said that pellagra was due to a deficiency of some nutrient, and the germ theory said that pellagra was caused by a germ transmitted by stable flies. A third theory, promoted by the eugenicist Charles Davenport, held that people only contracted pellagra if they were susceptible to it due to certain "constitutional, inheritable" traits of the affected individual.

 

Once alkali processing and dietary variety were understood and applied, pellagra disappeared in the developed world. The development of high lysine maize and the promotion of a more balanced diet have also contributed to its demise. Pellagra still exists today in food-poor areas and refugee camps where people survive on donated maize.

 

ALLERGY

Maize contains lipid transfer protein, an indigestible protein that survives cooking. This protein has been linked to a rare and understudied allergy to maize in humans. The allergic reaction can cause skin rash, swelling or itching of mucous membranes, diarrhea, vomiting, asthma and, in severe cases, anaphylaxis. It is unclear how common this allergy is in the general population.

 

MYCOTOXINS

Fungicide application does not reduce fungal growth or mycotoxin dramatically, although it can be a part of a successful reduction strategy. Among the most common toxins are those produced by Aspergillus and Fusarium spp. The most common toxins are aflatoxins, fumonisins, zearalenone, and ochratoxin A. Bt maize discourages insect vectors and by so doing it dramatically reduces concentrations of fumonisins, significantly reduces aflatoxins, but only mildly reduces others.

 

ART

Maize has been an essential crop in the Andes since the pre-Columbian era. The Moche culture from Northern Peru made ceramics from earth, water, and fire. This pottery was a sacred substance, formed in significant shapes and used to represent important themes. Maize was represented anthropomorphically as well as naturally.

 

In the United States, maize ears along with tobacco leaves are carved into the capitals of columns in the United States Capitol building. Maize itself is sometimes used for temporary architectural detailing when the intent is to celebrate the fall season, local agricultural productivity and culture. Bundles of dried maize stalks are often displayed along with pumpkins, gourds and straw in autumnal displays outside homes and businesses. A well-known example of architectural use is the Corn Palace in Mitchell, South Dakota, which uses cobs and ears of colored maize to implement a mural design that is recycled annually. Another well-known example is the Field of Corn sculpture in Dublin, Ohio, where hundreds of concrete ears of corn stand in a grassy field.

 

A maize stalk with two ripe ears is depicted on the reverse of the Croatian 1 lipa coin, minted since 1993.

 

WIKIPEDIA

By Alfred 23 Harth, 50 x 35, 1967.

 

Alfred 23 Harth’s early formation can be read as a remarkable intertwining of play, discipline, and conceptual awakening that would later come to characterize his multidisciplinary oeuvre. The boyhood dream of becoming an architect already contained a telling dialectic: on the one hand, the imaginative freedom of building ephemeral huts in the garden, one after another; on the other hand, the precision of constructing variations within given parameters. These garden architectures were not merely child’s play but may be understood retrospectively as proto-installations, temporary structures that mediated between imagination and actuality, an early rehearsal of the experimental crossings between construction, performance, and image that marked Harth’s mature practice.

A decisive rupture, almost an initiation ritual into modern art, occurred in 1958 when his elder brother Dietrich took him to a Dada exhibition in Frankfurt am Main. The timing was crucial: postwar Germany was only just beginning to reopen itself to the radical avant-gardes suppressed under fascism. For the young Harth, Dada presented not only a set of provocative images but also the lived possibility that art could destabilize categories, break down hierarchies, and operate conceptually as much as materially. The work The Navel—a simple black dot on white paper, accompanied by a title that displaced perception into language—sharpened this awareness. What mattered was not the mark itself but the dynamic between sign and referent, artwork and its commentary. The epiphany here was not aesthetic pleasure in the traditional sense but recognition of art as a space of thought, irony, and intellectual tension. This was nothing less than the beginning of a lifelong trajectory in which Harth would consistently return to the interface of sound, image, and idea.

In the following years, Harth immersed himself with voracity in every available art medium. School courses gave him the discipline of drawing, painting, and craft; his own appetite for performance led him to stage small situations, often masked or disguised, anticipating the performative interventions of the happening movement. The acquisition of his first camera at twelve extended his field into visual experimentation, while his pencil drawings of jazz musicians revealed both his growing fascination with musical improvisation and his awareness of biography as a narrative lens for art. What is striking here is the simultaneity of practices—music, drawing, performance, photography—that refused to be subordinated to a single discipline. Even before formal professional training, Harth was cultivating a polymorphic artistic identity in the spirit of the avant-garde.

The turn at fifteen to oil painting coincided with a parallel transformation in music: the gift of a tenor saxophone by his parents, an instrument that would guide him into deep engagement with jazz and improvisation. This was not simply the adoption of a hobby but the entry point into an emerging identity as a musician-artist, one who would soon refuse to see music and art as separated categories. Music, drawing, film, and conceptual play converged into a holistic practice that aligned with the growing international awareness of intermedia arts in the 1960s.

Attending the Goethe Gymnasium in his final school years refined this eclecticism. As an art-focused program with an ambition to train future cultural producers, it provided him with a sweeping introduction to international avant-garde currents, from Informel painting and Fluxus to Concept Art and experimental film. What Harth absorbed was not only technique but also a certain intellectual ecology: Frankfurt at that time was a city where cultural exchange, experimental music, and critical thought interacted dynamically. Together with Hubertus Gassner, who would later become a prominent museum director, Harth initiated happenings and other art events. Harth and founded the centrum freier cunst. Such a venture signaled more than youthful ambition: it represented the determination to create autonomous platforms for hybrid work when established institutions remained largely indifferent. Here Harth’s music group Just Music performed alongside conceptual and photo-based works, embodying an ethos of cross-disciplinary experimentation that paralleled international movements but arose organically from the Frankfurt milieu.

By the time of his Abitur in 1968, Harth embodied a paradoxical combination: on the one hand, a youthful openness to every medium, on the other, a growing self-awareness of art as critical practice. His decision to study design at the Werkkunstschule Offenbach, later shifting to art pedagogy at Goethe University, should not be misunderstood as a retreat into conventional paths. Rather, it reflects his strategy of grounding avant-garde impulses in a broader discourse of form and teaching. His musical activities expanded concurrently, so that life at this junction became an intense negotiation of study, performance, and conceptual inquiry. Alfred Harth's focus on synästhetic creation was indeed a significant aspect of his artistic approach at that time. He was interested in exploring synaesthesia beyond traditional media like TV, film, or theater, aiming to realize multisensory or synästhetic works that integrated sound, visual elements, and space in novel ways. This approach reflected his broader interest in breaking conventional boundaries of artistic disciplines and engaging the audience in immersive, multi-layered experiences that could not be confined to a single medium or format.

Looking back, one sees that Harth’s early trajectory established key themes of his later career: the refusal of boundaries between disciplines; the privileging of concept and idea over medium-specificity; the creation of autonomous spaces for collaboration beyond institutional frameworks; and, above all, the conviction that art is not an object but a process—often ephemeral, contingent, and dialogic. The boy who once built huts in his parent’s garden was already rehearsing the logic of variation and improvisation that would structure his later works across music, performance, and visual art. To trace these beginnings is to see how Harth’s career was less a matter of progression from one discipline to another than an ongoing movement across media, always oriented toward the space where form touches thought.

  

Tenerife

Icod de los Vinos

Butterfly garden

 

www.mariposario.com/en/

 

Papilio memnon, the great Mormon, is a large butterfly native to southern Asia that belongs to the swallowtail family. It is widely distributed and has thirteen subspecies. The female is polymorphic and with mimetic forms.

en.wikipedia.org/wiki/Papilio_memnon

This weaver ant might be doing some housekeeping job out of its nest.

 

Weaver Ants are those ants with reddish long bodies and very long legs that construct their nests by getting leaves together neatly. Multiple leaves are held together with the white fibers. Queen ant lays eggs on surface of these leaves internally and the pupa grows up in the shady cool place. This is how the weaver ant's nest looks like. This is a smaller specimen with two or three leaves woven together, but there are larger ones where four or more leaves are also strung together.

 

The ants do not have silk, but their larva does. However, larva cannot move around. So, the worker ants carry larva around and the little one spins enough silk to keep the leaves together as a house.

 

Oecophylla smaragdina is widespread in the Old World tropics and are present the most sophisticated nest-building activities of all weaver ants.The weaver ant (O. smaragdina) is a dominant canopy ant in tropical India and Australasia with colonies of up to 500 000 ants housed in nests made of leaves fastened together by larval silk and scattered across tens of trees. Workers draw leaves together, often forming long chains, and glue them together with larval silk. The colonies are very large and highly polydomous. Queens are pre-dominantly though not exclusively once-mated and colonies are usually single-queened, but most Northern Territory (Australia) colonies are polygynous. The workers are highly polymorphic (seen also in a fossilized colony), show complex polyethism, and present a much-studied rich pheromonal repertoire for the colony's tasks. Colony odor is partly learned, showing a "nasty neighbor" effect in reactions to other colonies of this highly territorial ant, and partly intrinsic to each individual. The odor varies over time and differs between the nests of a colony. Not surprisingly, Oecophylla ants are hosts to a variety of inquilines, such as spiders, which mimic the colony odor to escape detection. In addition, a constellation of Homoptera benefit from ant protection, yet the activities of the ants in controlling pest species make these ants beneficial insects (they are also human food in some areas) (adapted from Crozier et al., 2010). Reference: taxo4254.wikispaces.com/Oecophylla+smaragdina

Drosera cistiflora is a widespread highly variable and polymorphic species of sundew from the western Cape region of South Africa. there is tremendous variation in flower size and color as well as stem and leaf morphology. For this region, there are current research efforts to identify and reclassify this 'species' into multiple classifications. The late afternoon light provided a spectacular setting for these images.

The viviparous lizard or common lizard, Zootoca vivipara (formerly Lacerta vivipara), is a Eurasian lizard. It lives farther north than any other species of non-marine reptile, and most populations are viviparous (giving birth to live young), rather than laying eggs as most other lizards do. It is the only species in the monotypic genus Zootoca.[3]

 

Zootoca vivipara can be seen in a variety of different colors. Female Zootoca vivipara undergo color polymorphism (biology) more commonly than males. A female lizard's display differs in ventral coloration, ranging from pale yellow to bright orange and a mixed coloration. There have been many hypothesis for the genetic cause of this polymorphic coloration. These hypothesis test for coloration due to thermoregulation, predator avoidance, and social cues, specifically sexual reproduction. Through an experiment conducted by Vercken et al., color polymorphism in viviparous lizard is caused by social cues, rather than the other hypotheses. More specifically, the ventral coloration that is seen in female lizards is associated with patterns of sexual reproduction and sex allocation.[4]

 

.https://en.wikipedia.org/wiki/Viviparous_lizard

 

Ted Trueblood WMA

Grandview, Idaho

 

One of the most widespread and commonly observed birds of prey in North America, the Red-tailed Hawk (hereafter Red-tail) occupies a broad range of habitats from central Alaska south to Venezuela and east to the Virgin Islands. Breeding behavior, summer food habits, and habitat use have been well documented in many of these regions, but the taxonomic status of some populations remains unclear. The species varies greatly across its range, with up to 16 subspecies recognized by various authorities. Races are usually distinguished by ventral coloration, tail markings, and/or size, but there is no clear geographic trend in any of these characters. Some populations are polymorphic in ventral coloration (i.e., polychromatic), ranging from nearly white to nearly black, and extensive inter-gradation among adjacent subspecies complicates taxonomic relationships.

 

birdsna.org/Species-Account/bna/species/rethaw/introducti...

 

ebird.org/camerica/profile/MTAwNTY0Ng/world

Cladonia phyllophora Hoffm., syn.: Cladonia alcicornis var. phyllophora (Hoffm.) Malbr., Cladonia cervicornis f. phyllophora (Hoffm.) Dalla Torre & Sarnth., Cladonia degenerans (Flörke) Spreng

Family: Cladoniaceae

EN: Felt cladonia, DE: Beblätterte Becherflechte

Slo.: no name found

 

Dat.: Sept. 18. 2008

Lat.: 46.32403 Long.: 13.58408

Code: Bot_0297/2008_DSC3510

 

Habitat: Steep mountain slope, northwest aspect; among large boulders of a recent, large sock slide; in half shade; on sandy, calcareous ground; moderately humid place; protected from direct rain by overhanging rock; average precipitations ~ 3.000 mm/year, average temperature 7-9 deg C, elevations 750 m (2.450 feet), alpine phytogeographical region.

 

Substratum: sandy soil/raw hummus, among large calcareous boulders.

 

Place: Bovec basin, Northwest slopes of Mt. Javoršček, 1557 m; toward the end of a dirt forest road, East Julian Alps, Posočje, Slovenia EC.

 

Comment (relates to Flickr album Cladonia phyllophora): Browsing literature to determine the name of this find I've found only one or two candidates with podetia, which sometimes proliferate in more than two stores from cup margins. Cladonia rappii as well as Cladonia cervikornis/verticilata look similarly from far, but proliferate strictly from the center of the cups. Cladonia ramulosa may look similar too, but rarely (if at all) proliferates in more than two stores and is usually fertile with numerous conspicuous brown apothecia. None of several specimens found in this observation had podetia with apothecia.

 

The best, although not ideal, fit I've found seems to be Cladonia phyllophora. All sources agree that this taxon is highly polymorphic (google the pictures of it!). The taxon is also very variously interpreted by the authors (Ref. 7.). The description in literature, which seems the closest to this find, is in Brodo, Sharnoff, Sharnoff (2001) (Ref. 2.) mentioning gradually broadening and seemingly soft near the apex podetia having a slightly puffed-up aspect and cup margins richly decorated by small and thick squamules (see Fig. 4.) and brown pycnidia /see Fig.7.). The description in Smith at al (2009) (Ref. 1.) fits reasonably well too, particularly the description of the habit stated as 'often extensive more or less interlocking tiers of proliferating podetia'. However, many sources mention that the surface of the podetia near the base should be areolate with contrasting blackened decorticated and maculated areas (Ref. 1., Ref. 8.) or blackish podetia base (Ref. 7.), which is not the case in this find. Also substratum is usually cited as acid. This find apparently grew on a mixture of sandy soil and raw hummus deposited in gaps among large rock boulders (a few meters across) of a relatively recent large mountain rock slide. It seems possible that it was at least to some extent acid, however, the bedrock and the boulders themselves are no doubt calcareous. I am not sure my determination is correct, but, I am also not aware of a better alternative.

 

Ref.:

(1) C.W.Smith, et all, The lichens of Great Britain and Ireland,The British Lichen

Society,(2009), p 333.

(2) I.M. Brodo, S.D. Sharnoff, S.Sharnoff, Lichens of North America, Yale Uni. Press (2001), p 265.

(3) V. Wirth, Die Flechten Baden-Württembergs, Teil.1., Ulmer (1995), p 332.

(4) www.researchgate.net/publication/228358096_The_lichen_gen... (accessed May. 31. 2021)

(5) v3.boldsystems.org/index.php/Taxbrowser_Taxonpage?taxid=6... (accessed June 8. 2021)

(6) www.sharnoffphotos.com/lichensB/cladonia_phyllophora.html (accessed June 12. 2021)

(7) www.lichensmaritimes.org/index.php?task=fiche&lichen=... (accessed June 12. 2021)

(8) italic.units.it/index.php?procedure=taxonpage&num=814 (accessed June 14. 2021)

  

Innenpark. Bonifatius-Felsen. Felsen am Bonifatiusweg,

Inner Park. Boniface Rock. Rock on the Boniface-Way

 

Asplenium trichomanes subsp. pachyrachis (Christ) Lovis et Reichst.

Willdenowia 10: 18. 1980.

 

Asplenium trichomanes sublus. pachyrachis Christ

Farnkr. Schweiz 1 (2): 92. 1900.

Asplenium csikii Kümmerle & András.

Magyar Bot. Lapok 17. 110. 1919 (nomen), 20. 3, fig. 1923

Asplenium pachyrhachis Landolt

Fl. Indicativa 268 2010.

 

Seestern-Braunschwarz-Streifenfarn, Dickstieliger Braunstieliger Streifenfarn, Dickstieliger Brauner Streifenfarn

Lobed Maidenhair Spleenwort

 

Anmerkung:

Die Asplenium trichomanes Gruppe, ist ein sehr polymorpher, taxonomisch kritischer Spezieskomplex! Die Evolutionsgeschichte und Beziehungen zwischen den Taxa in dieser Gruppe wurden intensiv untersucht. Allerdings sind morphologische Variation und die Verteilung dieser Taxa unzureichend bekannt, da sie in der lokalen Floren oder Checklisten häufig nicht vorkommen. Die Gründe für die Vernachlässigung der Taxa innerhalb der Asplenium trichomanes Gruppe sind der Mangel an diagnostischen morphologische Merkmalen, das häufige gemeinsame Auftreten an ihren Standorten, sowie die Hybridisierung unter den Taxa. Die Asplenium trichomanes Gruppe umfasst zytologisch und ökologisch unterschiedliche Taxa mit fast weltweiter Verbreitung, die offenbar noch aktiv in der Entwicklung sind (L. Ekrt & M. Štech, 2008).

 

Annotation:

The Asplenium trichomanes group is a very polymorphic, taxonomically critical species complex! The evolutionary history and relationships between the taxa in this group have been extensively studied. However, morphological variation and distribution of these taxa are poorly understood because they are often absent in local floras or checklists. The reasons for the neglect of the taxa within the Asplenium trichomanes group are the lack of diagnostic morphological features, the frequent common occurrence at their sites, as well as the hybridization among the taxa. The Asplenium trichomanes group includes cytologically and ecologically diverse taxa with almost worldwide distribution, which apparently are still active in development (L. Ekrt & M. Štech, 2008).

Under Dew is Life

 

Description: Picture taken in a pet shop. No product nor the fish were acquired or bought, and the picture I took was merely for educational purposes in benefit of the species depicted.

 

They are also known as lowland cichlid, pearlscale cichlid, Texas blue or green Texas cichlid.

 

This is a Herichthys carpintis of the variation "Short Body" commonly known as Texas Short Body and belongs in the superclass Osteichthyes, class Actinopterygii, subclass Neopterygii, superorder Acanthopterygii, order Perciformes (Cichliformes?), suborder Labroidei, family Cichlidae, subfamily Cichlasomatinae and tribe Heroini. Bear in mind that this taxonomy was very confusing and there may be mistakes, not to mention that I skipped plenty of subdivisions within Neopterygii to make this shorter. If you saw any mistakes, please warn me so I can change this text accordingly. All I ask is for you to provide a reliable source to your correction of my mistake. "Guesses" or speculations won't make me immediately change the text, but are welcome.

 

Apparently, the subject portrayed is a male but through a picture it's hard to tell with 100% precision. Some females look a lot like the males, with many being mistaken for each other. The differentiation is more reliable by annalyzing the behaviour or through the observation of the sexual and excretory organs through a process called "venting" after the subject reaches sexual maturity. In general, the males are larger than the females and display a characteristic black spot on the center of the dorsal fin, but this can't be used to differentiate the sex of these fishes with 100% reliability.

 

Herichthys carpintis can be found in harder waters with a pH of 7,5 to 9 and temperatures between 18ºC and 28ºC, which varies with the seasons. They can also inhabit transparent waters with visibility inferior to 1 meter.

 

Sexual maturity is reached after the fish reaches around 10cm. They will try to reproduce as soon as the sexual maturity is achieved and will search for a smooth surface to lay the eggs. More than 800 eggs can be laid. The offspring are transferred between sites in crevices multiple times by the female to fend off predators. Both the males and the females will guard the offsprings for around 8 weeks.

 

This species is somewhat aggressive and are known to provoke problems with other creatures even when they are bigger than them, which may result in the death of the Herichthys carpintis, although this behaviour is more often seen when they are under captivity.

 

They are omnivorous and feed on debris, vegetable matter (they are diggers and will uproot vegetation), other fishes, small insects, gastropods, and so on.

 

This species is polymorphic, meaning they can present various patterns depending on their zone of distribution, which includes the Eastern coast of Mexico, until the Rio Soto La Marina to the North, and to the South until the Panuco's River Bay, Laguna de Tamiahua. They have been introduced in many places.

 

They live around 8 years under captivity. This expectancy is usually halved in nature. The individual portrayed measured approximately 13 or 14cm in length and approximately 9cm in height.

 

Sources:

 

www.ciclideos.com/herichthys-carpintis-f195.html

 

www.aquarismopaulista.com/texas-blue-herichthys-carpintis/

 

en.wikipedia.org/wiki/Polymorphism_(biology)

 

www.seriouslyfish.com/species/herichthys-carpintis/

 

PROJECT NOAH (Português): www.projectnoah.org/spottings/1400149398

©All photographs on this site are copyright: DESPITE STRAIGHT LINES (Paul Williams) 2011 – 2019 & GETTY IMAGES ®

  

No license is given nor granted in respect of the use of any copyrighted material on this site other than with the express written agreement of DESPITE STRAIGHT LINES (Paul Williams) ©

  

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Photograph taken taken at an altitude of One metres at 12:43pm on Monday 2nd September 2019, off 1461 Benvenuto Avenue in Brentwood Bay at the Victoria Butterfly gardens on Vancouver Island, British Columbia, Canada.

  

The Great Mormon butterfly (Papilio memnon) can grow to one hundred and fifty millimentres and is polymorphic, with four male and up to twenty six different female forms. It is native to Southern Asia.

 

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Nikon D850. Focal length 120mm Shutter speed 1/2000s Aperture f/4.0 iso2000 RAW (14 bit uncompressed) Image size L (8256 x 5504 FX). Focus mode AF-C focus 51 point with 3D- tracking. AF-Area mode single point & 73 point switchable. Exposure mode - Aperture priority exposure. Nikon Back button focusing enabled. Matrix metering. ISO Sensitivity: Auto. White balance: Natural light auto. Colour space Adobe RGB. Nikon Distortion control on. Picture control: Auto. High ISO NR on. Vignette control: normal. Active D-lighting Auto.

  

Nikkor AF-S 24-120mm f/4G ED VR. Lee SW150 MKII filter holder. Lee SW150 77mm screw in adapter. Lee SW150 circular polariser glass filter.Lee SW150 Filters field pouch.Nikon EN-EL15a battery.Mcoplus professional MB-D850 multi function battery grip 6960. Matin quick release neckstrap. My Memory 128GB Class 10 SDXC 80MB/s card. Lowepro Flipside 400 AW camera bag. Nikon GP-1 GPS module. Hoodman HEYENRG round eyepiece oversized eyecup.Manfrotto 055XPROB Tripod 3 Sections (Payload: 5.6kgs). Manfrotto 327RC2 Light Duty Grip Ball Magnesium Tripod Head (Payload: 5.5kgs). Manfrotto quick release plate 200PL-14. Jessops Tripod bag.Nikon MC-DC2 remote shutter release cable.

  

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LATITUDE: N 48d 33m 52.28s

LONGITUDE: W 123d 26m 21.44s

ALTITUDE: 69.0m

  

RAW (TIFF) FILE: 130.00MB NEF: 91.9MB

PROCESSED (JPeg) FILE: 51.10MB

     

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PROCESSING POWER:

 

Nikon D850 Firmware versions C 1.10 (9/05/2019) LD Distortion Data 2.017 (20/3/18) LF 1.00

 

HP 110-352na Desktop PC with AMD Quad-Core A6-5200 APU 64Bit processor. Radeon HD8400 graphics. 8 GB DDR3 Memory with 1TB Data storage. 64-bit Windows 10. Verbatim USB 2.0 1TB desktop hard drive. WD My Passport Ultra 1tb USB3 Portable hard drive. Nikon ViewNX-1 64bit (Version 1.3.1 11/07/2019). Nikon Capture NX-D 64bit (Version 1.4.7 15/03/2018). Nikon Picture Control Utility 2 (Version 1.3.2 15/03/2018). Adobe photoshop Elements 8 Version 8.0 64bit.

   

During July yellow pond lilies fill Isa Lake near Grant Pass in Yellowstone National Park, Wyoming. Isa Lake is perched atop one of 2 crossing of the Continental Divide that the Craig Pass Highway makes between West Thumb and Fishing Bridge. The little lake is covered with ice most of the year but in the summer it is filled with yellow waterlilies (Nuphar polysepalum). Isa lake is unusual in that that it has 2 outlets. One flows out on one side and goes to the Snake River tributaries and eventually to the Pacific while the outlet on the other side sends water to the Firehole River drainage and with time to the Mississippi and on to the Gulf of Mexico/ Atlantic. Even more unusual is the the configuration of the drainage. Because the local Divide runs roughly east-west at Isa's location, the east side drains to the Pacific and the west side of the lake drains to the Atlantic. That is the reverse of what one would expect.

 

The yellow water lily is also known as yellow pond lily, Indian pond lily, cow lily and spatterdock. I found some confusion when I tired to look up the scientific name of the yellow pond lily found in Yellowstone. The different species and subspecies names seemed to be being applied to same plant depending on the reference. The website ZipcodeZoo.com had the explanation: "The taxonomy of the genus is problematic. E. O. Beal (1956) departed dramatically from previous North American treatments in recognizing a single polymorphic species, Nuphar lutea (name of European origin ), with several subspecies formerly treated as species. Subsequent research (C. E. DePoe and E. O. Beal 1969; E. O. Beal and R. M. Southall 1977) has supported Beal's treatment for some southeastern subspecies, but most other taxa have not been studied as extensively. Beal's treatment, for the most part, has not been adopted in the Northeast and elsewhere in North America or in Europe. Molecular studies of Nuphar currently in progress (D. J. Padgett, pers. comm. ) have clearly shown the North American taxa to be distinct from the Eurasian Nuphar lutea; Beal's nomenclature under that taxon cannot be upheld. Continuing to treat those taxa at subspecific rank would require new combinations under Nuphar sagittifolia (Walter) Pursh, the oldest name that has hitherto been applied only in a geographically restricted sense. Until the molecular studies are completed, creating new names is premature. We therefore return to the previous treatment of the taxa as species."

 

This information was confirmed on the USDA's GRIN. A nomenclature change recognized by the USDA confirmed the name as Nuphar polysepalum.

 

zipcodezoo.com/Plants/N/Nuphar_polysepalum/

 

www.ars-grin.gov/cgi-bin/npgs/html/taxon.pl?25419

The confusion also extends to the genus nomenclature. ZipcodeZoo.com reports "Prior to conservation in its current sense, the name Nymphaea was frequently used for Nuphar. Although often treated as neuter, Nuphar was originally assigned the feminine gender".

 

So with the nomenclature issue at least partially understood, I did learn some very interesting things about the history of the yellow pond lily as a Native American food and medicine source from the US Forest Service site. "The mature seeds of the Yellow Pond Lily can be placed in a frying pan on an open fire and they will swell and pop open resembling popcorn in appearance and taste.The roots may be boiled or roasted, peeled, and then eaten as is or placed in a soup or stew. The core of the root is rich in starch. The root was also dried and ground into a meal and then made into flour by Native Americans.The Native Americans apparently had some difficulties in gathering this particular plant for food and medicine. The fleshy rootstocks grow under 4 to 5 feet of water. Some Indians would raid caches of the roots in muskrat houses, but usually they would dive for the root. Baked rootstocks of the Yellow Pond Lily were sliced and used as a poultice for sores. A decoction made by boiling rootstocks was added to bath water to treat rheumatism." (www.fs.usda.gov/detail/ipnf/learning/?cid=fsm9_019139)

 

U.S. Engineer, Hiram Chittenden, came across the unusual lake in 1891 as he was looking for a route for a new road between West Thumb and Old Faithful. He wrote a poem about the pond but it was not named until a couple years later in 1893 when Union Pacific Railroad official named it Isa for Isabel Jelke of Cincinnati, OH. Little is know about the reasons it was named after her or what her relationships were with the Railroad, Yellowstone National Park, the Lake itself or Hiram Chittenden (if any).

 

Because of the beautiful pond lilies this spot has been a favorite stop of the Jones Family over the generations and years.

Tenerife

Icod de los Vinos

Butterfly garden

 

www.mariposario.com/en/

 

Papilio memnon, the great Mormon, is a large butterfly native to southern Asia that belongs to the swallowtail family. It is widely distributed and has thirteen subspecies. The female is polymorphic and with mimetic forms.

en.wikipedia.org/wiki/Papilio_memnon

Backyard

st. louis

missouri

 

Roberto Cicchinè

 

piùmenoinfinito

 

testo e cura

Simonetta Angelini

    

L’inaugurazione si terrà domenica 19 settembre 2010 alle 18:00

La mostra termina il 17 ottobre

La Galleria Marconi è aperta tutti giorni dalle 16.00 alle 20.00, esclusa la domenica

   

La Galleria Marconi di Cupra Marittima riprende la propria attività espositiva, domenica 19 settembre alle 18.00, infatti si inaugura piùmenoinfinito, personale di Roberto Cicchinè che apre la rassegna Troppo (la Galleria Marconi esagera). La mostra, a cura ci Simonetta Angelini, autrice anche del testo critico, è anche l’occasione per presentare e inaugurare gli spazi rinnovati della Galleria.

 

La Galleria Marconi, sabato 9 ottobre 2010, partecipa con la mostra di Roberto Cicchinè, alla sesta edizione della Giornata del Contemporaneo, promossa da AMACI (Associazione dei Musei d’Arte Contemporanea Italiani).

“La microstoria, ciò che resta sospeso nella memoria, diviene segno minimale. Si dà per sottrazioni, per tracce, per mancanze, per minuti oggetti significanti circonfusi, dai contorni evanescenti come i ricordi

Il percorso di senso è pendolare, tra appartenenza e distanza, tra sospensione e trascorrere, tra intimità e condivisione.

Il linguaggio fotografico diviene quasi iconico, sintetico, è “indeterminato come solo l’esattezza sa essere”, procede con nettezza quasi matematica.

 

Il lavoro artistico sa declinarsi come opera aperta, partecipata, segno di un processo di condivisione tra l’artista, il suo mezzo espressivo, il curatore, il pubblico. Diviene un ipertesto, una mappa di navigazione, approdo e partenza per l’attivazione del pensiero, con un moto da navigatori della rete.

L’arte contemporanea e la rete sembrano somigliarsi: diffuse, plurali, interattive, ibride, polimorfe e polisemantiche, connettive. Il lavoro dell’artista sarà presentato come un’ interfaccia.

Le nuove tecnologie e la visibilità moltiplicata divengono instrumentum di condivisione relazione, di slittamento sensoriale, di una riconversione del senso, di dislocamento e trasversalità, di accessibilità del lavoro artistico ovunque e in ogni momento, di dilatazione dei significati, di contatto.

La partecipazione al processo creativo, i feedback reciproci tra l’artista e pubblico diventano possibili attraverso una critica partecipata, in cui il curatore condivide una relazione e si fa ermeneuta, “colui che porta fuori”.

Il pensiero critico appartiene a tutti, è al plurale, non viene imposto; diviene un punto di vista possibile, da verificare, da discutere. Attraverso le parole e le cose, il pubblico ha la possibilità di attivare il pensiero e lasciare tracce “estetiche”.

Il testo critico si avvarrà dell’inserimento nel corpo dell’intervento stesso di link a immagini, video, musiche, brani di film e fonti eterogenee, nel suggerire rimandi e suggestioni, approcci quasi sinestetici all’esperienza artistica, nell’apertura di nuovi percorsi partecipati, diffusi, plurali, circolari.

Il processo estetico è multimediale, multiplo; un viaggio da icononauti, da spettattori, attivatore di relazioni e pensiero, ibrido, di prelievo, di produzione collettiva di senso. Contro ogni unilateralità”.

(Simonetta Angelini)

 

Una nuova rassegna per il sedicesimo anno di attività, in una Galleria Marconi rinnovata nella forma ma fortemente radicata nella sostanza del proprio percorso di ricerca, già questo sembrerebbe Troppo. Ma non ci può bastare. La speranza è lavorare ancora con la coerenza, la testardaggine e la voglia di crescere, restando sempre al passo con i tempi, cogliendo idee e mutamenti della nostra epoca, e anche questo sembrerebbe Troppo. Non c’è arroganza, né immodestia, ma la rassegna 2010/2011 si intitola proprio Troppo (la Galleria Marconi esagera), perché in un mondo dove la crisi è presente e tocca purtroppo la vita di milioni di persone, dove tutti noi siamo sottoposti a tagli e a restrizioni dei diritti fondamentali, abbiamo avvertito il desiderio di debordare, non solo fisicamente.

 

Marconi Gallery of Cupra Marittima starts its exhibitions again. In fact the sole exposition piùmenoinfinito by Roberto Cicchiné will open on Sunday 19th September at 6.00 p.m. It starts the series Too Much (Marconi Gallery exagerates). The exposition, cured by Simonetta Angelini who is the author of the critical text too, is also the occasion to present and to inaugurate the renewed spaces of the Gallery.

 

“ The microhistory, what remains hanging in the memory, becomes a minimal sign. It' given through subtractions, tracks, lacks,, through bathed significant minute objects with evanescent contours like memories.

The sense path is pendular, between belonging and distance, between suspension and flowing, between intimacy and sharing.

The photographic language becomes nearly iconic, synthetic, it's “unspecified as only accuracy can be”, it goes on with a nearly mathematical clarity.

 

The art work can decline as an open work, sign of a sharing process among artist, his means of expression, curator, visitors. It becomes hypertext, a navigation map, leaving and arriving in order to activate one's thought, with a web surfer-like movement. Contemporary art and web seem to be similar: spred, plural, interactive, hybrid, polymorphic and polysemantic, connective. The work of the artist will be presented like an interface.

The new technologies and the multiplied visbility become instrumentum of sharing relationship, of sensory shift, of a reconversation of the sense, of displacement and transversality, of art work approachability everywhere and in any moment, of meaning extension, of contact.

The participation in the creative process, the mutual feedbacks between artist and visitors are possible trough a shared criticism, when the curator shares a relation and becomes hermeneut, “the one who carries out things”.

The critical thought belongs to everyone, it's plural, it is not imposed; it becomes a possible opinion, to be verified, to be discussed. Visitors have the possibility of thinking and leaving “aesthetic” traces through words and things.

The critical text will insert links and images,videos, music, film and heterogeneous sources extracts in its intervention, in order to suggest references and suggestions, nearly synergistic approaches to the artistic experience, in order to create new shared, common, plural and circular courses.

The aesthetic process is multimedia, multiple; a visitors voyage, activating relations and thinking, hybrid, withdrawing, producing a common sense. Against any one-sidedness”.

(Simonetta Angelini)

 

A new series for the sixteenth year of activity, in the renewed spaces of Marconi Gallery, but strongly roteed in its research course. All this would seem already Too much. But it cannot be sufficient. Our hope is to work still coherently, with our being willful and wanting to grow, always behind the times, capturing ideas and changes of our time. All this would seem Too much as well. There is neither arrogance nor immodesty but the 2010-2011 series is just entitled Too Much (Marconi Gallery exagerates), because in a world where there is the crisis that is involving million of people, where everyone of us is subjected to cuts and restrictions of the fundamental rights, we feel the desire to overflow, and not only physically.

  

scheda tecnica/technical card

 

curatore/curator by Simonetta Angelini

testo critico/art critic by Simonetta Angelini

 

ufficio stampa/ press agent Dario Ciferri

traduzione di/translation by Patrizia Isidori

 

relazioni esterne e promozione delle attività/ external relationship and promotion of activities Stefania Palanca

fotografia/photography Marco Biancucci©

riprese video/video shooting Stefano Abbadini

allestimenti/preparation Marco Croci

progetto grafico/graphic project maicol e mirco

webmaster

 

Vitis (grapevine) is a genus of 81 accepted species of vining plants in the flowering plant family Vitaceae. The genus consists of species predominantly from the Northern Hemisphere. It is economically important as the source of grapes, both for direct consumption of the fruit and for fermentation to produce wine. The study and cultivation of grapevines is called viticulture.

 

Most cultivated Vitis varieties are wind-pollinated with hermaphroditic flowers containing both male and female reproductive structures, while wild species are dioecious. These flowers are grouped in bunches called inflorescences. In many species, such as Vitis vinifera, each successfully pollinated flower becomes a grape berry with the inflorescence turning into a cluster of grapes. While the flowers of the grapevines are usually very small, the berries are often large and brightly colored with sweet flavors that attract birds and other animals to disperse the seeds contained within the berries.

 

Grapevines usually only produce fruit on shoots that came from buds that were developed during the previous growing season. In viticulture, this is one of the principles behind pruning the previous year's growth (or "One year old wood") that includes shoots that have turned hard and woody during the winter (after harvest in commercial viticulture). These vines will be pruned either into a cane which will support 8 to 15 buds or to a smaller spur which holds 2 to 3 buds.

 

Description

Flower buds are formed late in the growing season and overwinter for blooming in spring of the next year. They produce leaf-opposed cymes. Vitis is distinguished from other genera of Vitaceae by having petals which remain joined at the tip and detach from the base to fall together as a calyptra or 'cap'. The flowers are mostly bisexual, pentamerous, with a hypogynous disk. The calyx is greatly reduced or nonexistent in most species and the petals are joined together at the tip into one unit but separated at the base. The fruit is a berry, ovoid in shape and juicy, with a two-celled ovary each containing two ovules, thus normally producing four seeds per flower (or fewer by way of aborted embryos).

 

Other parts of the vine include the tendrils which are leaf-opposed, branched in Vitis vinifera, and are used to support the climbing plant by twining onto surrounding structures such as branches or the trellising of a vine-training system.

 

In the wild, all species of Vitis are normally dioecious, but under domestication, variants with perfect flowers appear to have been selected.

 

The genus Vitis is divided into two subgenera, Euvitis Planch. have 38 chromosomes (n=19) with berries borne on clusters and Muscadinia Planch. 40 (n=20) with small clusters.

 

Wild grapes can resemble the single-seeded Menispermum canadense (moonseed), which is toxic.

 

Species

Most Vitis species are found mostly in the temperate regions of the Northern Hemisphere in North America and eastern Asia, exceptions being a few in the tropics and the wine grape Vitis vinifera which originated in southern Europe and southwestern Asia. Grape species occur in widely different geographical areas and show a great diversity of form.

 

Their growth makes leaf collection challenging and polymorphic leaves make identification of species difficult. Mature grapevines can grow up to 48 centimetres (19 inches) in diameter at breast height and reach the upper canopy of trees more than 35 metres (115 feet) in height.

 

Many species are sufficiently closely related to allow easy interbreeding and the resultant interspecific hybrids are invariably fertile and vigorous. Thus the concept of a species is less well defined and more likely represents the identification of different ecotypes of Vitis that have evolved in distinct geographical and environmental circumstances.

 

The exact number of species is not certain. Plants of the World Online states 81 species are accepted, but lists 84. More than 65 species in Asia are poorly defined. Approximately 25 species are known in North America and just one, V. vinifera has Eurasian origins; some of the more notable include:

 

Vitis aestivalis, the summer grape, native to the Eastern United States, especially the Southeastern United States

Vitis amurensis, native to the Asian continent, including parts of Siberia and China

Vitis arizonica, The Arizona grape is native to Arizona, Utah, Nevada, California, New Mexico, Texas, and Northern Mexico.

Vitis berlandieri, native to the southern North America, primarily Texas, New Mexico and Arkansas. Primarily known for good tolerance against soils with a high content of lime, which can cause chlorosis in many vines of American origin

Vitis californica, the California wild grape, or Northern California grape, or Pacific grape, is a wild grape species widespread across much of California as well as southwestern Oregon

Vitis coignetiae, the crimson glory vine, a species from East Asia grown as an ornamental plant for its crimson autumn foliage

Vitis labrusca L., the fox grapevine, sometimes used for winemaking and for jam. Native to the Eastern United States and Canada. The Concord grape was derived by a cross with this species

Vitis riparia, the riverbank grapevine, sometimes used for winemaking and for jam. Native to the entire Eastern United States and north to Quebec

Vitis rotundifolia (syn. Muscadinia rotundifolia), the muscadine, used for jams and wine. Native to the Southeastern United States from Delaware to the Gulf of Mexico

Vitis rupestris, the rock grapevine, used for breeding of Phylloxera resistant rootstock. Native to the Southern United States

Vitis vinifera, the European grapevine. Native to the Mediterranean and Central Asia.

Vitis vulpina, the frost grape, native to the Eastern United States, from Massachusetts to Florida, and west to Nebraska, Kansas, and Texas Treated by some as a synonym of V. riparia.

 

Plants of the World Online also includes:

Vitis acerifolia Raf.

Vitis amoena Z.H. Chen, Feng Chen & WW.Y. Xie

Vitis baihuashanensis M.S.Kang & D.Z.Lu

Vitis balansana Planch.

Vitis bashanica P.C.He

Vitis bellula (Rehder) W.T.Wang

Vitis betulifolia Diels & Gilg

Vitis biformis Rose

Vitis blancoi Munson

Vitis bloodworthiana Comeaux

Vitis bourgaeana Planch.

Vitis bryoniifolia Bunge

Vitis × champinii Planch.

Vitis chunganensis Hu

Vitis chungii F.P.Metcalf

Vitis cinerea (Engelm.) Millardet

Vitis davidi (Rom.Caill.) Foëx

Vitis × doaniana Munson ex Viala

Vitis erythrophylla W.T.Wang

Vitis fengqinensis C.L.Li

Vitis ficifolia Bunge

Vitis flavicosta Mickel & Beitel

Vitis flexuosa Thunb.

Vitis girdiana Munson

Vitis hancockii Hance

Vitis heyneana Schult.

Vitis hissarica Vassilcz.

Vitis hui W.C.Cheng

Vitis jaegeriana Comeaux

Vitis jinggangensis W.T.Wang

Vitis jinzhainensis X.S.Shen

Vitis kaihuaica Z.H.Chen, Feng Chen & W.Y Xie

Vitis kiusiana Momiy.

Vitis lanceolatifoliosa C.L.Li

Vitis longquanensis P.L.Chiu

Vitis luochengensis W.T.Wang

Vitis menghaiensis C.L.Li

Vitis mengziensis C.L.Li

Vitis metziana Miq.

Vitis monticola Buckley

Vitis mustangensis Buckley

Vitis nesbittiana Comeaux

Vitis × novae-angliae Fernald

Vitis novogranatensis Moldenke

Vitis nuristanica Vassilcz.

Vitis palmata Vahl

Vitis pedicellata M.A.Lawson

Vitis peninsularis M.E.Jones

Vitis piasezkii Maxim.

Vitis pilosonervia F.P.Metcalf

Vitis popenoei J.L.Fennell

Vitis pseudoreticulata W.T.Wang

Vitis quinlingensis P.C.He

Vitis retordii Rom.Caill. ex Planch.

Vitis romanetii Rom.Caill.

Vitis ruyuanensis C.L.Li

Vitis saccharifera Makino

Vitis shenxiensis C.L.Li

Vitis shizishanensis Z.Y.Ma, J.Wen, Q.Fu & X.Q.Liu

Vitis shuttleworthii House

Vitis silvestrii Pamp.

Vitis sinocinerea W.T.Wang

Vitis sinoternata W.T.Wang

Vitis tiliifolia Humb. & Bonpl. ex Schult.

Vitis tsoi Merr.

Vitis wenchowensis C.Ling

Vitis wenxianensis W.T.Wang

Vitis wilsoniae H.J.Veitch

Vitis wuhanensis C.L.Li

Vitis xunyangensis P.C.He

Vitis yunnanensis C.L.Li

Vitis zhejiang-adstricta P.L.Chiu

There are many cultivars of grapevines; most are cultivars of V. vinifera. One of them includes, Vitis 'Ornamental Grape'.

 

Hybrid grapes also exist, and these are primarily crosses between V. vinifera and one or more of V. labrusca, V. riparia or V. aestivalis. Hybrids tend to be less susceptible to frost and disease (notably phylloxera), but wine from some hybrids may have a little of the characteristic "foxy" taste of V. labrusca.

 

The Latin word Vitis is feminine,[19] and therefore adjectival species names take feminine forms, such as V. vinifera.

 

Ecology

Phylloxera is an American root aphid that devastated V. vinifera vineyards in Europe when accidentally introduced in the late 19th century. Attempts were made to breed in resistance from American species, but many winemakers and customers did not like the unusual flavour profile of the hybrid vines. However, V. vinifera grafts readily onto rootstocks of the American species and their hybrids with V. vinifera, and most commercial production of grapes now relies on such grafts.

 

Commercial distribution

According to the UN's Food and Agriculture Organization (FAO), 75,866 square kilometres of the world is dedicated to grapes. Approximately 71% of world grape production is used for wine, 27% as fresh fruit, and 2% as dried fruit. A portion of grape production goes to producing grape juice to be used as a sweetener for fruits canned "with no added sugar" and "100% natural". The area dedicated to vineyards is increasing by about 2% per year.

 

Domestic cultivation

Grapevines are widely cultivated by gardeners, and numerous suppliers cater specifically for this trade. The plants are valued for their decorative foliage, often colouring brightly in autumn; their ability to clothe walls, pergolas and arches, thus providing shade; and their fruits, which may be eaten as dessert or provide the basis for homemade wines. Popular varieties include:-

 

Buckland Sweetwater' (white dessert)

'Chardonnay' (white wine)

'Foster's Seedling' (white dessert)

'Grenache' (red wine)

'Muscat of Alexandria' (white dessert)

'Müller-Thurgau' (white wine)

'Phoenix' (white wine)

'Pinot noir' (red wine)

'Regent' (red wine)

'Schiava Grossa' (red dessert)

'Seyval blanc' (white wine)

'Tempranillo' (red wine)

 

The following varieties have gained the Royal Horticultural Society's Award of Garden Merit:-

'Boskoop Glory' (dessert/wine)

'Brant' (black dessert)

'Claret Cloak' or 'Frovit' (ornamental)

'New York Muscat' (black dessert)

'Purpurea' (ornamental)

 

Uses

The fruit of several Vitis species are grown commercially for consumption as fresh grapes and for fermentation into wine. Vitis vinifera is the most important such species.

 

The leaves of several species of grapevine are edible and are used in the production of dolmades and Vietnamese lot leaves.

 

Culture

The grapevine (typically Vitis vinifera) has been used as a symbol since ancient times. In Greek mythology, Dionysus (called Bacchus by the Romans) was god of the vintage and, therefore, a grapevine with bunches of the fruit are among his attributes. His attendants at the Bacchanalian festivals hence had the vine as an attribute, together with the thyrsus, the latter often entwined with vine branches. For the same reason, the Greek wine cup (cantharos) is commonly decorated with the vine and grapes, wine being drunk as a libation to the god.

 

The grapevine has a profound symbolic meaning in Jewish tradition and culture since antiquity. It is referenced 55 times in the Hebrew Bible (Old Testament), along with grapes and wine, which are also frequently mentioned (55 and 19, respectively). It is regarded as one of the Seven Species, and is employed several times in the Bible as a symbol of the Israelites as the chosen people. The grapevine has a prominent place in Jewish rituals: the wine was given a special blessing, "creator of the fruit of the vine", and the Kiddush blessing is recited over wine or grape juice on Shabbat and Jewish holidays. It is also employed in various parables and sayings in rabbinic literature. According to Josephus and the Mishnah, a golden vine was hung over the inner chamber of the Second Temple. The grapevine is featured on Hasmonean and Bar Kokhba revolt coinage, and as a decoration in mosaic floors of ancient synagogues.

 

In Christian iconography, the vine also frequently appears. It is mentioned several times in the New Testament. We have the parable of the kingdom of heaven likened to the father starting to engage laborers for his vineyard. The vine is used as symbol of Jesus Christ based on his own statement, "I am the true vine (John 15:1)." In that sense, a vine is placed as sole symbol on the tomb of Constantia, the sister of Constantine the Great, and elsewhere. In Byzantine art, the vine and grapes figure in early mosaics, and on the throne of Maximianus of Ravenna it is used as a decoration.

 

The vine and wheat ear have been frequently used as symbol of the blood and flesh of Christ, hence figuring as symbols (bread and wine) of the Eucharist and are found depicted on ostensories. Often the symbolic vine laden with grapes is found in ecclesiastical decorations with animals biting at the grapes. At times, the vine is used as symbol of temporal blessing.

 

In Mandaeism, uthras (angels or celestial beings) are often described as personified grapevines (gupna).

Magdalena the Astounding's spontaneous polymorphic aposematism was considered a divine warning by faithful followers in her village but physicians attributed it to lepidopteritis and proceeded with trepanation in an attempt to alleviate her symptoms. The debate continues.

 

Created for the 56th Contest on Man Ray: The Hole!

 

Thanks to Dr. Paul Grand for the Hole.

The Apalachicola Kingsnake is a polymorphic population of common kingsnakes that is restricted to the Apalachicola Lowlands of the Florida panhandle. They can exhibit a wide range of color patterns, but most have some combination of blotching and speckling. This is an adult male found in the Apalachicola National Forest.

As usual for owls, female screech-owls are usually larger than the males of their species, with owls of both sexes being compact in size and shape.

 

The Eastern Screech-owl, Megascops asio, is one of the smallest species of owls in North America. All of the birds in this genus are small and agile.

 

Screech-owls are generally coloured in various brownish hues with usually a whitish, patterned underside, which helps to camouflage them against the bark of trees. Some are polymorphic, occurring in a grayish- and a reddish-brown morph.

 

Went shooting again with www.raymondbarlow.com/

 

The red-tailed hawk (Buteo jamaicensis) is a bird of prey that breeds throughout most of North America, from the interior of Alaska and northern Canada to as far south as Panama and the West Indies. It is one of the most common members within the genus of Buteo in North America or worldwide. The red-tailed hawk is one of three species colloquially known in the United States as the "chickenhawk", though it rarely preys on standard-sized chickens. The bird is sometimes also referred to as the red-tail for short, when the meaning is clear in context. Red-tailed hawks can acclimate to all the biomes within their range, occurring on the edges of non-ideal habitats such as dense forests and sandy deserts. The red-tailed hawk occupies a wide range of habitats and altitudes, including deserts, grasslands (from small meadows to the treed fringes of more extensive prairies), coniferous and deciduous forests, agricultural fields, and urban areas. Its latitudinal limits fall around the tree line in the subarctic and it is absent from the high Arctic. Generally it favors varied habitats with open woodland, woodland edge and open terrain. It is legally protected in Canada, Mexico, and the United States by the Migratory Bird Treaty Act.

 

The 14 recognized subspecies vary in appearance and range, varying most often in color, and in the west of North America, red-tails are particularly often strongly polymorphic, with individuals ranging from almost white to nearly all black. The subspecies Harlan's hawk (B. j. harlani) is sometimes considered a separate species (B. harlani). The red-tailed hawk is one of the largest members of the genus Buteo, typically weighing from 690 to 1,600 g (1.5 to 3.5 lb) and measuring 45–65 cm (18–26 in) in length, with a wingspan from 110–141 cm (3 ft 7 in – 4 ft 8 in). This species displays sexual dimorphism in size, with females averaging about 25% heavier than males.

 

The diet of red-tailed hawks is highly variable and reflects their status as opportunistic generalists, but in North America, they are most often predators of small mammals such as rodents of an immense diversity of families and species. Prey that is terrestrial and at least partially diurnal is preferred, so types such as ground squirrels are preferred where they naturally occur. Over much of the range, smallish rodents such as voles alternated with larger rabbits and hares often collectively form the bulk of the diet. Large numbers of birds and reptiles can occur in the diet in several areas, and can even be the primary foods. Meanwhile, amphibians, fish and invertebrates can seem rare in the hawk's regular diet, but they are not infrequently taken by immature hawks. Red-tailed hawks may survive on islands absent of native mammals on diets variously including invertebrates such as crabs, as well as lizards or birds. Like many Buteo species, they hunt from a perch most often, but can vary their hunting techniques where prey and habitat demand it. Because they are so common and easily trained as capable hunters, in the United States they are the most commonly captured hawks for falconry. Falconers are permitted to take only passage hawks (which have left the nest, are on their own, but are less than a year old) so as to not affect the breeding population. Passage red-tailed hawks are also preferred by falconers because they have not yet developed the adult behaviors that would make them more difficult to train.

 

Taxonomy

The red-tailed hawk was formally described in 1788 by German naturalist Johann Friedrich Gmelin under the binomial name Falco jamaicensis. Gmelin based his description on the "cream-coloured buzzard" described in 1781 by John Latham in his A General Synopsis of Birds. The type locality is Jamaica. The red-tailed hawk is now placed in the genus Buteo that was erected by French naturalist Bernard Germain de Lacépède in 1799.

 

The red-tailed hawk is a member of the subfamily Buteoninae, which includes about 55 currently recognized species. Unlike many lineages of accipitrids, which seemed to have radiated out of Africa or south Asia, the Buteoninae clearly originated in the Americas based on fossil records and current species distributions (more than 75% of the extant hawks from this lineage are found in the Americas). As a subfamily, the Buteoninae seem to be rather old based on genetic materials, with monophyletic genera bearing several million years of individual evolution. Diverse in plumage appearance, habitat, prey, and nesting preferences, buteonine hawks are nonetheless typically medium- to large-sized hawks with ample wings (while some fossil forms are very large, larger than any eagle alive today). The red-tailed hawk is a member of the genus Buteo, a group of medium-sized raptors with robust bodies and broad wings. Members of this genus are known as "buzzards" in Eurasia, but "hawks" in North America. Under current classification, the genus includes about 29 species, the second-most diverse of all extant accipitrid genera behind only Accipiter. The buzzards of Eurasia and Africa are mostly part of the genus Buteo, although two other small genera within the subfamily Buteoninae occur in Africa.

 

At one time, the rufous-tailed hawk (B. ventralis), distributed in Patagonia and some other areas of southern South America, was considered part of the red-tailed hawk species. With a massive distributional gap consisting of most of South America, the rufous-tailed hawk is considered a separate species now, but the two hawks still compromise a "species pair" or superspecies, as they are clearly closely related. The rufous-tailed hawk, while comparatively little studied, is very similar to the red-tailed hawk, being about the same size and possessing the same wing structure, and having more or less parallel nesting and hunting habits. Physically, however, rufous-tailed hawk adults do not attain a bright brick-red tail as do red-tailed hawks, instead retaining a dark brownish-cinnamon tail with many blackish crossbars similar to juvenile red-tailed hawks. Another, more well-known, close relative to the red-tailed hawk is the common buzzard (B. buteo), which has been considered as its Eurasian "broad ecological counterpart" and may also be within a species complex with red-tailed hawks. The common buzzard, in turn, is also part of a species complex with other Old World buzzards, namely the mountain buzzard (B. oreophilus), the forest buzzard (B. trizonatus ), and the Madagascar buzzard (B. brachypterus). All six species, although varying notably in size and plumage characteristics, in the alleged species complex that contains the red-tailed hawk share with it the feature of the blackish patagium marking, which is missing in most other Buteo spp.

 

Subspecies

At least 14 recognized subspecies of B. jamaicensis are described, which vary in range and in coloration. Not all authors accept every subspecies, though, particularly some of the insular races of the tropics (which differ only slightly in some cases from the nearest mainland forms) and particularly Krider's hawk, by far the most controversial red-tailed hawk race, as few authors agree on its suitability as a full-fledged subspecies.

 

ImageSubspeciesDistribution

Jamaican red-tailed hawk (B. j. jamaicensis)occurs throughout the West Indies (including Jamaica, Hispaniola, Puerto Rico and the Lesser Antilles) except for the Bahamas and Cuba.

Alaska red-tailed hawk (B. j. alascensis)breeds (probably resident) from southeastern coastal Alaska to Haida Gwaii and Vancouver Island in British Columbia.

Eastern red-tailed hawk (B. j. borealis)breeds from southeast Canada and Maine south through Texas and east to northern Florida.

Western red-tailed hawk (B. j. calurus)greatest longitudinal breeding distribution of any race of red-tailed hawk.

Central American red-tailed hawk (B. j. costaricensis)from Nicaragua to Panama.

Southwestern red-tailed hawk (B. j. fuertesi)breeds from northern Chihuahua to South Texas.

Tres Marias red-tailed hawk (B. j. fumosus)endemic to Islas Marías, Mexico.

Mexican Highlands red-tailed hawk (B. j. hadropus)native to the Mexican Highlands.

Harlan's hawk (B. j. harlani)breeds from central Alaska to northwestern Canada, with the largest number of birds breeding in the Yukon or western Alaska, reaching their southern limit in north-central British Columbia.

Red-tailed hawk (kemsiesi) (B. j. kemsiesi)a dark subspecies resident from Chiapas, Mexico, to Nicaragua.

Krider's hawk (B. j. kriderii)breeds from southern Alberta, southern Saskatchewan, southern Manitoba, and extreme western Ontario south to south-central Montana, Wyoming, western Nebraska, and western Minnesota.

Socorro red-tailed hawk (B. j. socorroensis)endemic to Socorro Island, Mexico.

Cuban red-tailed hawk (B. j. solitudinis)native to the Bahamas and Cuba.

Florida red-tailed hawk (B. j. umbrinus)occurs year-round in peninsular Florida north to as far Tampa Bay and the Kissimmee Prairie south throughout the rest of peninsular Florida south to the Florida Keys.

 

Description

Red-tailed hawk plumage can be variable, depending on the subspecies and the region. These color variations are morphs, and are not related to molting. The western North American population, B. j. calurus, is the most variable subspecies and has three main color morphs: light, dark, and intermediate or rufous. The dark and intermediate morphs constitute 10–20% of the population in the Western United States, but seem to constitute only 1–2% of B. j. calurus in western Canada. A whitish underbelly with a dark brown band across the belly, formed by horizontal streaks in feather patterning, is present in most color variations. This feature is variable in eastern hawks and generally absent in some light subspecies (i.e. B. j. fuertesi). Most adult red-tails have a dark-brown nape and upper head, which gives them a somewhat hooded appearance, while the throat can variably present a lighter brown "necklace". Especially in younger birds, the underside may be otherwise covered with dark-brown spotting, and some adults may too manifest this stippling. The back is usually a slightly darker brown than elsewhere with paler scapular feathers, ranging from tawny to white, forming a variable imperfect "V" on the back. The tail of most adults, which gives this species its name, is rufous brick-red above with a variably sized, black subterminal band and generally appears light buff-orange from below. In comparison, the typical pale immatures (i.e. less than two years old) typically have a mildly paler headed and tend to show a darker back than adults with more apparent pale wing-feather edges above (for descriptions of dark morph juveniles from B. j. calurus, which is also generally apt for description of rare dark morphs of other races, see under that subspecies description). In immature red-tailed hawks of all morphs, the tail is a light brown above with numerous small dark brown bars of roughly equal width, but these tend to be much broader on dark morph birds. Even in young red-tails, the tail may be a somewhat rufous tinge of brown. The bill is relatively short and dark, in the hooked shape characteristic of raptors, and the head can sometimes appear small in size against the thick body frame. The cere, the legs, and the feet of the red-tailed hawk are all yellow, as is the color of bare parts in many accipitrids of different lineages. Immature birds can be readily identified at close range by their yellowish irises. As the bird attains full maturity over the course of 3–4 years, the iris slowly darkens into a reddish-brown, which is the adult eye-color in all races. Seen in flight, adults usually have dark brown along the lower edge of the wings, against a mostly pale wing, which bares light brownish barring. Individually, the underwing coverts can range from all dark to off-whitish (most often more heavily streaked with brown) which contrasts with a distinctive black patagium marking. The wing coloring of adults and immatures is similar but for typical pale morph immatures having somewhat heavier brownish markings.

 

Though the markings and color vary across the subspecies, the basic appearance of the red-tailed hawk is relatively consistent.

 

Overall, this species is blocky and broad in shape, often appearing (and being) heavier than other Buteos of similar length. They are the heaviest Buteos on average in eastern North America, albeit scarcely ahead of the larger winged rough-legged buzzard (Buteo lagopus), and second only in size in the west to the ferruginous hawk (Buteo regalis). Red-tailed hawks may be anywhere from the fifth to the ninth heaviest Buteo in the world depending on what figures are used. However, in the northwestern United States, ferruginous hawk females are 35% heavier than female red-tails from the same area.[2] On average, western red-tailed hawks are relatively longer winged and lankier proportioned but are slightly less stocky, compact and heavy than eastern red-tailed hawks in North America. Eastern hawks may also have mildly larger talons and bills than western ones. Based on comparisons of morphology and function amongst all accipitrids, these features imply that western red-tails may need to vary their hunting more frequently to on the wing as the habitat diversifies to more open situations and presumably would hunt more variable and faster prey, whereas the birds of the east, which was historically well-wooded, are more dedicated perch hunters and can take somewhat larger prey but are likely more dedicated mammal hunters. In terms of size variation, red-tailed hawks run almost contrary to Bergmann's rule (i.e. that northern animals should be larger in relation than those closer to the Equator within a species) as one of the northernmost subspecies, B. j. alascensis, is the second smallest race based on linear dimensions and that two of the most southerly occurring races in the United States, B. j. fuertesi and B. j. umbrinus, respectively, are the largest proportioned of all red-tailed hawks. Red-tailed hawks tend have a relatively short but broad tails and thick, chunky wings. Although often described as long-winged, the proportional size of the wings is quite small and red-tails have high wing loading for a buteonine hawk. For comparison, two other widespread Buteo hawks in North America were found to weigh: 30 g (1.1 oz) for every square centimeter of wing area in the rough-legged buzzard (B. lagopus) and 44 g (1.6 oz)/cm2 in the red-shouldered hawk (B. lineatus). In contrast, the red-tailed hawk weighed considerably more for their wing area: 199 g (7.0 oz) per square cm.

 

As is the case with many raptors, the red-tailed hawk displays sexual dimorphism in size, as females are on average 25% larger than males. As is typical in large raptors, frequently reported mean body mass for red-tailed hawks is somewhat higher than expansive research reveals. Part of this weight variation is seasonal fluctuations; hawks tend to be heavier in winter than during migration or especially during the trying summer breeding season, and also due to clinal variation. Furthermore, immature hawks are usually lighter in mass than their adult counterparts despite having somewhat longer wings and tails. Male red-tailed hawks may weigh from 690 to 1,300 g (1.52 to 2.87 lb) and females may weigh 801 to 1,723 g (1.766 to 3.799 lb) (the lowest figure from a migrating female immature from Goshute Mountains, Nevada, the highest from a wintering female in Wisconsin). Some sources claim the largest females can weigh up to 2,000 g (4.4 lb), but whether this is in reference to wild hawks (as opposed to those in captivity or used for falconry) is not clear. The largest known survey of body mass in red-tailed hawks is still credited to Craighead and Craighead (1956), who found 100 males to average 1,028 g (2.266 lb) and 108 females to average 1,244 g (2.743 lb). However, these figures were apparently taken from labels on museum specimens, from natural history collections in Wisconsin and Pennsylvania, without note to the region, age, or subspecies of the specimens. However, 16 sources ranging in sample size from the aforementioned 208 specimens to only four hawks in Puerto Rico (with 9 of the 16 studies of migrating red-tails), showed that males weigh a mean of 860.2 g (1.896 lb) and females weigh a mean of 1,036.2 g (2.284 lb), about 15% lighter than prior species-wide published weights. Within the continental United States, typical weights of males can range from 840.8 g (1.854 lb) (for migrating males in Chelan County, Washington) to 1,031 g (2.273 lb) (for male hawks found dead in Massachusetts), and females ranged from 1,057.9 g (2.332 lb) (migrants in the Goshutes) to 1,373 g (3.027 lb) (for females diagnosed as B. j. borealis in western Kansas). Size variation in body mass reveals that the red-tailed hawk typically varies only a modest amount and that size differences are geographically inconsistent.[9][40]

 

Male red-tailed hawks can measure 45 to 60 cm (18 to 24 in) in total length, females measuring 48 to 65 cm (19 to 26 in) long. Their wingspan typically can range from 105 to 141 cm (3 ft 5 in to 4 ft 8 in), although the largest females may possible span up to 147 cm (4 ft 10 in). In the standard scientific method of measuring wing size, the wing chord is 325.1–444.5 mm (12.80–17.50 in) long. The tail measures 188 to 258.7 mm (7.40 to 10.19 in) in length. The exposed culmen was reported to range from 21.7 to 30.2 mm (0.85 to 1.19 in) and the tarsus averaged 74.7–95.8 mm (2.94–3.77 in) across the races. The middle toe (excluding talon) can range from 38.3 to 53.8 mm (1.51 to 2.12 in), with the hallux-claw (the talon of the rear toe, which has evolved to be the largest in accipitrids) measuring from 24.1 to 33.6 mm (0.95 to 1.32 in) in length.

 

Identification

Although they overlap in range with most other American diurnal raptors, identifying most mature red-tailed hawks to species is relatively straightforward, particularly if viewing a typical adult at a reasonable distance. The red-tailed hawk is the only North American hawk with a rufous tail and a blackish patagium marking on the leading edge of its wing (which is obscured only on dark morph adults and Harlan's hawks by similarly dark-colored feathers). Other larger adult Buteo spp. in North America usually have obvious distinct markings that are absent in red-tails, whether the rufous-brown "beard" of Swainson's hawks (B. swainsonii) or the colorful rufous belly and shoulder markings and striking black-and-white mantle of red-shouldered hawks (also the small "windows" seen at the end of their primaries).[ In perched individuals, even as silhouettes, the shape of large Buteo spp. may be distinctive, such as the wingtips overhanging the tail in several other species, but not in red-tails. North American Buteo spp. range from the dainty, compact builds of much smaller ones, such as broad-winged hawk (B. platypterus) to the heavyset, neckless look of ferruginous hawks or the rough-legged buzzards, which have a compact, smaller appearance than a red-tail in perched birds due to its small bill, short neck, and much shorter tarsi, while the opposite effect occurs in flying rough-legs with their much bigger wing area. In flight, most other large North American Buteo spp. are distinctly longer and more slender-winged than red-tailed hawks, with the much paler ferruginous hawk having peculiarly slender wings in relation to its massive, chunky body. Swainson's hawks are distinctly darker on the wing and ferruginous hawks are much paler-winged than typical red-tailed hawks. Pale morph adult ferruginous hawk can show mildly tawny-pink (but never truly rufous) upper tail, and like red-tails tend to have dark markings on underwing-coverts and can have a dark belly band, but compared to red-tailed hawks have a distinctly broader head, their remiges are much whiter looking with very small, dark primary tips, they lack the red-tail's diagnostic patagial marks and usually also lack the dark subterminal tail-band, and ferruginous hawks have totally feathered tarsi. With its whitish head, the ferruginous hawk is most similar to Krider's red-tailed hawks, especially in immature plumage, but the larger hawk has broader head and narrower wing shape, and the ferruginous immatures are paler underneath and on their legs. Several species share a belly band with the typical red-tailed hawk, but they vary from subtle (as in the ferruginous hawk) to solid blackish, the latter in most light-morph rough-legged buzzards. More difficult to identify among adult red-tails are their darkest variations, as most species of Buteo in North America also have dark morphs. Western dark morph red-tails (i.e. B. j. calurus) adults, however, retain the typical distinctive brick-red tail, which other species lack, and may stand out even more against the otherwise all chocolate-brown to black bird. Standard pale juveniles when perched show a whitish patch in the outer half of the upper surface of the wing, which other juvenile Buteo spp. lack.[ The most difficult to identify stages and plumage types are dark morph juveniles, Harlan's hawk and some Krider's hawks (the latter mainly with typical ferruginous hawks as mentioned). Some darker juveniles are similar enough to other Buteo juveniles that they "cannot be identified to species with any confidence under various field conditions." However, field identification techniques have advanced in the last few decades and most experienced hawk-watchers can distinguish even the most vexingly plumaged immature hawks, especially as the wing shapes of each species becomes apparent after seeing many. Harlan's hawks are most similar to dark morph rough-legged buzzards and dark morph ferruginous hawks. Wing shape is the most reliable identification tool for distinguishing Harlan's hawks from these, but also the pale streaking on the breast of Harlan's, which tends to be conspicuous in most individuals, and is lacking in the other hawks. Also, dark morph ferruginous hawks do not have the dark subterminal band of a Harlan's hawk, but do bear a black undertail covert lacking in Harlan's.

 

Vocalization

The cry of the red-tailed hawk is a 2- to 3-second, hoarse, rasping scream, variously transcribed as kree-eee-ar, tsee-eeee-arrr or sheeeeee, that begins at a high pitch and slurs downward. This cry is often described as sounding similar to a steam whistle. The red-tailed hawk frequently vocalizes while hunting or soaring, but vocalizes loudest and most persistently in defiance or anger, in response to a predator or a rival hawk's intrusion into its territory. At close range, it makes a croaking guh-runk, possibly as a warning sound. Nestlings may give peeping notes with a "soft, sleepy quality" that give way to occasional screams as they develop, but those are more likely to be a soft whistle rather than the harsh screams of the adults. Their latter hunger call, given from 11 days (as recorded in Alaska) to after fledgling (in California), is different, a two-syllabled, wailing klee-uk food cry exerted by the young when parents leave the nest or enter their field of vision. A strange mechanical sound "not very unlike the rush of distant water" has been reported as uttered in the midst of a sky-dance. A modified call of chirp-chwirk is given during courtship, while a low key, duck-like nasal gank may be given by pairs when they are relaxed.

 

The fierce, screaming cry of the adult red-tailed hawk is frequently used as a generic raptor sound effect in television shows and other media, even if the bird featured is not a red-tailed hawk. It is especially used in depictions of the bald eagle, which contributes to the common misconception that it is a bald eagle cry; actual bald eagle vocalizations are far softer and more chirpy than those of a red-tailed hawk.

 

Distribution and habitat

The red-tailed hawk is one of the most widely distributed of all raptors in the Americas. It occupies the largest breeding range of any diurnal raptor north of the Mexican border, just ahead of the American kestrel (Falco sparverius). While the peregrine falcon (Falco peregrinus) has a greater latitudinal distribution as a nester in North America, its range as a breeding species is far more sporadic and sparse than that of red-tailed hawks. The red-tailed hawk breeds from nearly north-central Alaska, the Yukon, and a considerable portion of the Northwest Territories, there reaching as far as a breeder as Inuvik, Mackenzie River Delta and skirting the southern shores of Great Bear Lake and Great Slave Lake. Thereafter in northern Canada, breeding red-tails continue to northern Saskatchewan and across to north-central Ontario east to central Quebec and the Maritime Provinces of Canada, and south continuously to Florida. No substantial gaps occur throughout the entire contiguous United States where breeding red-tailed hawks do not occur. Along the Pacific, their range includes all of Baja California, including Islas Marías, and Socorro Island in the Revillagigedo Islands. On the mainland, breeding red-tails are found continuously to Oaxaca, then experience a brief gap at the Isthmus of Tehuantepec thereafter subsequently continuing from Chiapas through central Guatemala on to northern Nicaragua. To the south, the population in highlands from Costa Rica to central Panama is isolated from breeding birds in Nicaragua. Further east, breeding red-tailed hawks occur in the West Indies in north Bahamas (i.e. Grand Bahama, Abaco and Andros) and all larger islands (such as Cuba, Jamaica, Hispaniola, and Puerto Rico) and into the northern Lesser Antilles (Virgin Islands, Saint Barthélemy, Saba, Saint Kitts, and Nevis, being rare as a resident on Saint Eustatius and are probably extinct on Saint Martin). Their typical winter range stretches from southern Canada south throughout the remainder of the breeding range.

 

Red-tailed hawks have shown the ability to become habituated to almost any habitat present in North and Central America. Their preferred habitat is mixed forest and field, largely woodland edge with tall trees or alternately high bluffs that may be used as nesting and perching sites. They occupy a wide range of habitats and altitudes, including deserts, grasslands, nearly any coastal or wetland habitat, mountains, foothills, coniferous and deciduous woodlands, and tropical rainforests. Agricultural fields and pastures, which are more often than not varied with groves, ridges, or streamside trees in most parts of America, may make nearly ideal habitat for breeding or wintering red-tails. They also adapt well to suburban areas especially ones with tall trees or any kind of parkland. Some red-tails may survive or even flourish in urban areas, usually hunting and roosting in available urban parks, cemeteries, road verges, and so on, and nesting freely either in trees or virtually any tall man-made structures. One famous urban red-tailed hawk, known as "Pale Male", became the subject of a nonfiction book, Red-Tails in Love: A Wildlife Drama in Central Park, and is the first known red-tail in decades to successfully nest and raise young in the crowded New York City borough of Manhattan. As studied in Syracuse, New York, the highway system has been very beneficial to red-tails as it juxtaposed trees and open areas and blocks human encroachment with fences, with the red-tailed hawks easily becoming acclimated to car traffic. The only practice that has a negative effect on the highway-occupying red-tails is the planting of exotic Phragmites, which may occasionally obscure otherwise ideal highway habitat.

 

In the northern Great Plains, the widespread practices of wildfire suppression and planting of exotic trees by humans has allowed groves of aspen and various other trees to invade what was once vast, almost continuous prairie grasslands, causing grassland obligates such as ferruginous hawks to decline and allowing parkland-favoring red-tails to flourish. To the contrary, clear-cutting of mature woodlands in New England, resulting in only fragmented and isolated stands of trees or low second growth remaining, was recorded to also benefit red-tailed hawks, despite being to the determent of breeding red-shouldered hawks. The red-tailed hawk, as a whole, rivals the peregrine falcon and the great horned owl among raptorial birds in the use of diverse habitats in North America. Beyond the high Arctic (as they discontinue as a breeder at the tree line), few other areas exist where red-tailed hawks are absent or rare in North and Central America. Some areas of unbroken forest, especially lowland tropical forests, rarely host red-tailed hawks, although they can occupy forested tropical highlands surprisingly well. In deserts, they can only occur where some variety of arborescent growth or ample rocky bluffs or canyons occur.

 

Behavior

The red-tailed hawk is highly conspicuous to humans in much of its daily behavior. Most birds in resident populations, which are well more than half of all red-tailed hawks, usually split nonbreeding-season activity between territorial soaring flight and sitting on a perch. Often, perching is for hunting purposes, but many sit on a tree branch for hours, occasionally stretching on a single wing or leg to keep limber, with no signs of hunting intent. Wintering typical pale-morph hawks in Arkansas were found to perch in open areas near the top of tall, isolated trees, whereas dark morphs more frequently perched in dense groups of trees. For many, and perhaps most, red-tailed hawks being mobbed by various birds is a daily concern and can effectively disrupt many of their daily behaviors. Mostly larger passerines, of multiple families from tyrant flycatchers to icterids, mob red-tails, despite other raptors, such as Accipiter hawks and falcons, being a notably greater danger to them. The most aggressive and dangerous attacker as such is likely to be various crows or other corvids, i.e. American crows (Corvus brachyrhynchos), because a mobbing group (or "murder") of them can number up to as many as 75 crows, which may cause grievous physical harm to a solitary hawk, and if the hawks are nesting, separate the parent hawks and endanger the eggs or nestlings within their nest to predation by crows. Birds that mob red-tailed hawks can tell how distended the hawk's crop is (i.e. the upper chest and throat area being puffy versus flat-feathered and sleek), thus mob more often when the hawk is presumably about to hunt.

 

Flight

In flight, this hawk soars with wings often in a slight dihedral, flapping as little as possible to conserve energy. Soaring is by far the most efficient method of flight for red-tailed hawks, so is used more often than not. Active flight is slow and deliberate, with deep wing beats. Wing beats are somewhat less rapid in active flight than in most other Buteo hawks, even heavier species such as ferruginous hawks tend to flap more swiftly, due to the morphology of the wings. In wind, it occasionally hovers on beating wings and remains stationary above the ground, but this flight method is rarely employed by this species. When soaring or flapping its wings, it typically travels from 32 to 64 km/h (20 to 40 mph), but when diving may exceed 190 km/h (120 mph). Although North American red-tailed hawks will occasionally hunt from flight, a great majority of flight by red-tails in this area is for non-hunting purpose. During nest defense, red-tailed hawks may be capable of surprisingly swift, vigorous flight, while repeatedly diving at perceived threats.

 

Migration

Red-tailed hawks are considered partial migrants, as in about the northern third of their distribution, which is most of their range in Canada and Alaska, they almost entirely vacate their breeding grounds. In coastal areas of the north, however, such as in the Pacific Northwest to southern Alaska and in Nova Scotia on the Atlantic, red-tailed hawks do not usually migrate. More or less, any area where snow cover is nearly continuous during the winter shows an extended absence of most red-tailed hawks, so some areas as far south as Montana may show strong seasonal vacancies of red-tails. In southern Michigan, immature red-tailed hawks tended to remain in winter only when voles were abundant. During relatively long, harsh winters in Michigan, many more young ones were reported in northeastern Mexico. To the opposite extreme, hawks residing as far north as Fairbanks, Alaska, may persevere through the winter on their home territory, as was recorded with one male over three consecutive years. Birds of any age tend to be territorial during winter but may shift ranges whenever food requirements demand it. Wintering birds tend to perch on inconspicuous tree perches, seeking shelter especially if they have a full crop or are in the midst of poor or overly windy weather. Adult wintering red-tails tend to perch more prominently than immatures do, which select lower or more secluded perches. Immatures are often missed in winter bird counts, unless they are being displaced by dominant adults. Generally, though, immatures can seem to recognize that they are less likely to be attacked by adults during winter and can perch surprisingly close to them. Age is the most significant consideration of wintering hawks' hierarchy, but size does factor in, as larger immatures (presumably usually females) are less likely to displaced than smaller ones. Dark adult red-tailed hawks appear to be harder to locate when perched than other red-tails. In Oklahoma, for example, wintering adult Harlan's hawks were rarely engaged in fights or chased by other red-tails. These hawks tended to gather in regional pockets and frequently the same ones occurred year-to-year. In general, migratory behavior is complex and reliant on each individual hawk's decision-making (i.e. whether prey populations are sufficient to entice the hawk to endure prolonged snow cover). During fall migration, departure may occur as soon as late September, but peak movements occur in late October and all of November in the United States, with migration ceasing after mid-December. The northernmost migrants may pass over resident red-tailed hawks in the contiguous United States, while the latter are still in the midst of brooding fledglings. Not infrequently, several autumn hawk watches in Ontario, Quebec, and the northern United States record 4,500–8,900 red-tailed hawks migrating through each fall, with records of up to 15,000 in a season at Hawk Ridge hawk watch in Duluth, Minnesota. Unlike some other Buteo spp., such as Swainson's hawks and broad-winged hawks, red-tailed hawks do not usually migrate in groups, instead passing by one-by-one, and only migrate on days when winds are favorable. Most migrants do not move past southern Mexico in late autumn, but a few North American migrants may annually move as far south as breeding red-tailed hawks happen to occur, i.e. in Central America to as far south Panama. However, a few records were reported of wintering migrant red-tails turning up in Colombia, the first records of them anywhere in South America. Spring northward movements may commence as early as late February, with peak numbers usually occurring in late March and early April. Seasonal counts may include up to 19,000 red-tails in spring at Derby Hill hawk watch, in Oswego, New York, sometimes more than 5,000 are recorded in a day there. The most northerly migratory individuals may not reach breeding grounds until June, even adults.

 

Immature hawks migrate later than adults in spring on average, but not, generally speaking, in autumn. In the northern Great Lakes, immatures return in late May to early June, when adults are already well into their nesting season and must find unoccupied ranges. In Alaska, adults tend to migrate before immatures in early to mid-September, to the contrary of other areas, probably as heavy snowfall begins. Yearlings that were banded in southwestern Idaho stayed for about 2 months after fledging, and then traveled long distances with a strong directional bias, with 9 of 12 recovered southeast of the study area- six of these moved south to coastal lowlands in Mexico] and as far as Guatemala, 4,205 km (2,613 mi) from their initial banding. In California, 35 hawks were banded as nestlings; 26 were recovered at less than 50 miles away, with multidirectional juvenile dispersals. Nestlings banded in Southern California sometimes actually traveled north as far as 1,190 km (740 mi) to Oregon, ranging to the opposite extreme as far as a banded bird from the Sierra Nevadas that moved 1,700 km (1,100 mi) south to Sinaloa. Nestlings banded in Green County, Wisconsin, did not travel very far comparatively by October–November, but by December, recoveries were found in states including Illinois, Iowa, Texas, Louisiana, and Florida.

 

Diet

The red-tailed hawk is carnivorous, and a highly opportunistic feeder. Nearly any small animal they encounter may be viewed as potential food. Their most common prey are small mammals such as rodents and lagomorphs, but they also consume birds, reptiles, fish, amphibians, and invertebrates. Prey varies considerably with regional and seasonal availability, but usually centers on rodents, accounting for up to 85% of a hawk's diet. In total, nearly 500 prey species have been recorded in their diet, almost as many as great horned owls have been recorded as taking. When 27 North American studies are reviewed, mammals make up 65.3% of the diet by frequency, 20.9% by birds, 10.8% by reptiles, 2.8% by invertebrates, and 0.2% by amphibians and fish. The geometric mean body mass of prey taken by red-tailed hawks in North America is about 187 g (6.6 oz) based on a pair of compilation studies from across the continent, regionally varying at least from 43.4 to 361.4 g (1.53 to 12.75 oz). Staple prey (excluding invertebrates) has been claimed to weigh from 15 to 2,114 g (0.033 to 4.661 lb), ranging roughly from the size of a small mouse or lizard to the size of a black-tailed jackrabbit (Lepus californicus). The daily food requirements range from 7 to 11.2% of their own body weight, so that about three voles or the equivalent weight are required daily for a typical range adult.

 

The talons and feet of red-tailed hawks are relatively large for a Buteo hawk; in an average-sized adult red-tail, the "hallux-claw" or rear talon, the largest claw on all accipitrids, averages about 29.7 mm (1.17 in). In fact, the talons of red-tails in some areas averaged of similar size to those of ferruginous hawks which can be considerably heavier and notably larger than those of the only slightly lighter Swainson's hawk. This species may exert an average of about 91 kg/cm2 (1,290 lbf/in2) of pressure through its feet. Owing to its morphology, red-tailed hawks generally can attack larger prey than other Buteo hawks typically can, and are capable of selecting the largest prey of up to their own size available at the time of hunting, though in all likelihood numerically most prey probably weighs on average about 20% of the hawk's own weight (as is typical of many birds of prey). Red-tailed hawks usually hunt by watching for prey activity from a high perch, also known as still hunting. Upon being spotted, prey is dropped down upon by the hawk. Red-tails often select the highest available perches within a given environment, since the greater the height they are at, the less flapping is required and the faster the downward glide they can attain toward nearby prey. If prey is closer than average, the hawk may glide at a steep downward angle with few flaps, if farther than average, it may flap a few swift wingbeats alternating with glides. Perch hunting is the most successful hunting method generally speaking for red-tailed hawks and can account for up to 83% of their daily activities (i.e. in winter). Wintering pairs may join and aseasonally may join forces to group hunt agile prey that they may have trouble catching by themselves, such as tree squirrels. This may consist of stalking opposites sides of a tree, to surround the squirrel and almost inevitably drive the rodent to be captured by one after being flushed by the other hawk.

 

The most common flighted hunting method for red-tail is to cruise around 10 to 50 m (33 to 164 ft) over the ground with flap-and-glide type flight, interspersed occasionally with harrier-like quarters over the ground. This method is less successful than perch hunting, but seems relatively useful for capturing small birds and may show the best results while hunting in hilly country. Hunting red-tailed hawks readily use trees, bushes, or rocks for concealment before making a surprise attack, even showing a partial ability to dodge among trees in an Accipiter-like fashion. Among thick stands of spruce in Alaska, a dodging hunting flight was thought to be unusually important to red-tails living in extensive areas of conifers, with hawks even coming to the ground and walking hurriedly in prey pursuit especially if the prey was large, a similar behavior to goshawks. Additional surprisingly swift aerial hunting has reported in red-tails that habitually hunt bats in Texas. Here, the bat-hunting specialists stooped with half-closed wings, quite falcon-like, plowing through the huge stream of bats exiting their cave roosts, then zooming upwards with a bat in its talons. These hawks also flew parallel closely to the stream, then veer sharply into it and seize a bat. In the neotropics, red-tails have shown the ability to dodge amongst forest canopy whilst hunting. In Kansas, red-tailed hawks were recorded sailing to catch flying insects, a hunting method more typical of a Swainson's hawk. Alternately, they may drop to the ground to forage for insects like grasshoppers and beetles as well as other invertebrates and probably amphibians and fish (except by water in the latter cases). Hunting afoot seems to be particularly prevalent among immatures. Young red-tailed hawks in northeastern Florida were recorded often extracting earthworms from near the surface of the ground and some had a crop full of earthworms after rains. Ground hunting is also quite common on Socorro Island, where no native land mammals occur, and invertebrates are more significant to their overall diet. A red-tailed hawk was observed to incorporate an unconventional killing method, which was drowning a heron immediately after capture. One red-tailed hawk was seen to try to grab a young ground squirrel and, upon missing it, screamed loudly, which in turn caused another young squirrel to break into a run, wherein it was captured. Whether this was an intentional hunting technique needs investigation. Upon capture, smaller prey is taken to a feeding perch, which is almost always lower than a hunting perch. Among small prey, rodents are often swallowed whole, as are shrews and small snakes, while birds are plucked and beheaded. Even prey as small as chipmunks may take two or three bites to consume. Larger mammals of transportable size are at times beheaded and have part of their fur discarded, then leftovers are either stored in a tree or fall to the ground. Large prey, especially if too heavy to transport on the wing, is often dragged to a secluded spot and dismantled in various ways. If they can successfully carry what remains to a low perch, they tend to feed until full and then discard the rest.

 

Mammals

Rodents are certainly the type of prey taken most often by frequency, but their contribution to prey biomass at nests can be regionally low, and the type, variety, and importance of rodent prey can be highly variable. In total, well over 100 rodent species have turned up the diet of red-tailed hawks. Rodents of extremely varied sizes may be hunted by red-tails, with species ranging in size from the 8.2 g (0.29 oz) eastern harvest mouse (Reithrodontomys humulis) to full grown muskrats (Ondatra zibethicus). At times, the red-tailed hawk is thought of as a semi-specialized vole-catcher, but voles are a subsistence food that is more or less taken until larger prey such as rabbits and squirrels can be captured. In an area of Michigan, immature hawks took almost entirely voles but adults were diversified feeders. Indeed, the 44.1 g (1.56 oz) meadow vole (Microtus pennsylvanicus) was the highest frequency prey species in 27 dietary studies across North America, accounting for up to 54% of the food at nests by frequency. It is quite rare for any one species to make up more than half of the food in any dietary study for red-tailed hawks. In total about 9 Microtus species are known in the overall diet, with 5 other voles and lemmings known to be included in their prey spectrum. Another well-represented species was the 27.9 g (0.98 oz) prairie vole (Microtus ochrogaster), which were the primary food, making up 26.4% of a sample of 1322, in eastern Kansas. While crepuscular in primary feeding activity, voles are known to be active both day and night, and so are reliable food for hawks than most non-squirrel rodents, which generally are nocturnal in activity. Indeed, most other microtine rodents are largely inaccessible to red-tailed hawks due to their strongly nocturnal foraging patterns, even though 24 species outside of voles and lemmings are known to be hunted. Woodrats are taken as important supplemental prey in some regions, being considerably larger than most other crictetid rodents, and some numbers of North American deermouse (Peromyscus maniculatus) may turn up. The largest representation of the latter species was contributing 11.9% of the diet in the Great Basin of Utah, making them the second best-represented prey species there. Considering this limited association with nocturnal rodents, the high importance of pocket gophers in the diet of red-tailed hawks is puzzling to many biologists, as these tend to be highly nocturnal and elusive by day, rarely leaving the confines of their burrow. At least 8 species of pocket gopher are included in the prey spectrum (not to mention 5 species of pocket mice). The 110 g (3.9 oz) northern pocket gopher (Thomomys talpoides) is particularly often reported and, by frequency, even turns up as the third most often recorded prey species in 27 American dietary studies. Presumably, hunting of pocket gophers by red-tails, which has possibly never been witnessed, occurs in dim light at first dawn and last light of dusk when they luck upon a gopher out foraging.

 

By far, the most important prey among rodents is squirrels, as they are almost fully diurnal. All told, nearly 50 species from the squirrel family have turned up as food. In particular, where they are distributed, ground squirrels are doubly attractive as a primary food source due to their ground-dwelling habits, as red-tails prefer to attack prey that is terrestrial. There are also many disadvantages to ground squirrels as prey: they can escape quickly to the security of their burrows, they tend to be highly social and they are very effective and fast in response to alarm calls, and a good deal of species enter hibernation that in the coldest climates can range up to a 6 to 9-month period (although those in warmer climates with little to no snowy weather often have brief dormancy and no true hibernation). Nonetheless, red-tailed hawks are devoted predators of ground squirrels, especially catching incautious ones as they go out foraging (which are often younger animals). A multi-year study conducted on San Joaquin Experimental Range in California, seemingly still the largest food study to date done for red-tailed hawks with 4031 items examined, showed that throughout the seasons the 722 g (1.592 lb) California ground squirrel (Otospermophilus beecheyi) was the most significant prey, accounting for 60.8% of the breeding season diet and about 27.2% of the diet for hawks year-around. Because of the extremely high density of red-tailed hawks on this range, some pairs came to specialize in diverse alternate prey, which consisted variously of kangaroo rats, lizards, snakes or chipmunks. One pair apparently lessened competition by focusing on pocket gophers instead despite being near the center of ground squirrel activity. In Snake River NCA, the primary food of red-tailed hawks was the 203.5 g (7.18 oz) Townsend's ground squirrel (Urocitellus townsendii), which made up nearly 21% of the food in 382 prey items across several years despite sharp spikes and crashes of the ground squirrel population there. The same species was the main food of red-tailed hawks in southeastern Washington, making up 31.2% of 170 items. An even closer predatory relationship was reported in the Centennial valley of Montana and south-central Montana, where 45.4% of 194 prey items and 40.2% of 261 items, respectively, of the food of red-tails consisted of the 455.7 g (1.005 lb) Richardson's ground squirrel (Urocitellus richardsonii). Locally in Rochester, Alberta, Richardson's ground squirrel, estimated to average 444 g (15.7 oz), were secondary in number to unidentified small rodents but red-tails in the region killed an estimated 22–60% of the area's ground squirrel, a large dent in the squirrel's population. Further east, ground squirrels are not so reliably distributed, but one study in southern Wisconsin, in one of several quite different dietary studies in that state, the 172.7 g (6.09 oz) thirteen-lined ground squirrel (Ictidomys tridecemlineatus) was the main prey species, making up 29.7% of the diet (from a sample of 165).

 

In Kluane Lake, Yukon, 750 g (1.65 lb) Arctic ground squirrels (Spermophilus parryii) were the main overall food for Harlan's red-tailed hawks, making up 30.8% of a sample of 1074 prey items. When these ground squirrels enter their long hibernation, the breeding Harlan's hawks migrate south for the winter. Nearly as important in Kluane Lake was the 200 g (7.1 oz) American red squirrel (Tamiasciurus hudsonicus), which constituted 29.8% of the above sample. Red squirrels are highly agile dwellers on dense spruce stands, which has caused biologists to ponder how the red-tailed hawks are able to routinely catch them. It is possible that the hawks catch them on the ground such as when squirrels are digging their caches, but theoretically, the dark color of the Harlan's hawks may allow them to ambush the squirrels within the forests locally more effectively. While American red squirrels turn up not infrequently as supplementary prey elsewhere in North America, other tree squirrels seem to be comparatively infrequently caught, at least during the summer breeding season. It is known that pairs of red-tailed hawks will cooperatively hunt tree squirrels at times, probably mostly between late fall and early spring. Fox squirrels (Sciurus niger), the largest of North America's tree squirrels at 800 g (1.8 lb), are relatively common supplemental prey but the lighter, presumably more agile 533 g (1.175 lb) eastern gray squirrel (Sciurus carolinensis) appears to be seldom caught based on dietary studies. While adult marmot may be difficult for red-tailed hawks to catch, young marmots are readily taken in numbers after weaning, such as a high frequency of yellow-bellied marmot (Marmota flaviventris) in Boulder, Colorado. Another grouping of squirrels but at the opposite end of the size spectrum for squirrels, the chipmunks are also mostly supplemental prey but are considered more easily caught than tree squirrels, considering that they are more habitual terrestrial foragers In central Ohio, eastern chipmunks (Tamias striatus), the largest species of chipmunk at an average weight of 96 g (3.4 oz), were actually the leading prey by number, making up 12.3% of a sample of 179 items.

 

Outside of rodents, the most important prey for North American red-tailed hawks is rabbits and hares, of which at least 13 species are included in their prey spectrum. By biomass and reproductive success within populations, these are certain to be their most significant food source (at least in North America). Adult Sylvilagus rabbits known to be hunted by red-tails can range from the 700 g (1.5 lb) brush rabbit (Sylvilagus bachmani) to the Tres Marias rabbit (Sylvilagus graysoni) at 1,470 g (3.24 lb) while all leporids hunted may range the 421.3 g (14.86 oz) pygmy rabbit (Brachylagus idahoensis) to hares and jackrabbits potentially up twice the hawk's own weight. While primarily crepuscular in peak activity, rabbits and hares often foraging both during day and night and so face almost constant predatory pressure from a diverse range of predators. Male red-tailed hawks or pairs which are talented rabbit hunters are likely to have higher than average productivity due to the size and nutrition of the meal ensuring healthy, fast-growing offspring. Most widely reported are the cottontails, which the three most common North America varieties softly grading into mostly allopatric ranges, being largely segregated by habitat preferences where they overlap in distribution. Namely, in descending order of reportage were: the eastern cottontail (Sylvilagus floridanus), the second most widely reported prey species overall in North America and with maximum percentage known in a given study was 26.4% in Oklahoma (out of 958 prey items), the mountain cottontail (Sylvilagus nuttallii), maximum representation being 17.6% out of a sample of 478 in Kaibab Plateau, Arizona and the desert cottontail (Sylvilagus audubonii), maximum representation being 22.4% out of a sample of 326 in west-central Arizona. Black-tailed jackrabbits (Lepus californicus) are even more intensely focused upon as a food source by the hawks found in the west, particularly the Great Basin. With the weight around 2,114 g (4.661 lb), adults of this species is the largest prey routinely hunted by red-tailed hawks. When jackrabbit numbers crash, red-tailed hawk productivity tends to decline as well. In northern Utah, black-tailed jackrabbits made up 55.3% of a sample of 329. Elsewhere, they are usually somewhat secondary by number.

 

In the boreal forests of Canada and Alaska, red-tails are fairly dependent on the snowshoe hare (Lepus americanus), falling somewhere behind the great horned owl and ahead of the Anerican goshawk in their regional reliance on this food source. The hunting preferences of red-tails who rely on snowshoe hares are variable. In Rochester, Alberta, 52% of snowshoe hares caught were adults, such prey estimated to average 1,287 g (2.837 lb), and adults, in some years, were six times more often taken than juvenile hares, which averaged an estimated 560 g (1.23 lb). 1.9–7.1% of adults in the regional population of Rochester were taken by red-tails, while only 0.3–0.8 of juvenile hares were taken by them. Despite their reliance on it, only 4% (against 53.4% of the biomass) of the food by frequency here was made up of hares. On the other hand, in Kluane Lake, Yukon, juvenile hares were taken roughly 11 times more often than adults, despite the larger size of adults here, averaging 1,406.6 g (3.101 lb), and that the overall prey base was less diverse at this more northerly clime. In both Rochester and Kluane Lake, the number of snowshoe hares taken was considerably lower than the number of ground squirrels taken. The differences in average characteristics of snowshoe hares that were hunted may be partially due to habitat (extent of bog openings to dense forest) or topography. Another member of the Lagomorpha order has been found in the diet include juvenile white-tailed jackrabbit (Lepus townsendii) and the much smaller American pika (Ochotona princeps), at 150 g (5.3 oz).

 

A diversity of mammals may be consumed opportunistically outside of the main food groups of rodents and leporids, but usually occur in low numbers. At least five species each are taken of shrews and moles, ranging in size from their smallest mammalian prey, the cinereus (Sorex cinereus) and least shrews (Cryptotis parva), which both weigh about 4.4 g (0.16 oz), to Townsend's mole (Scapanus townsendii), which weighs about 126 g (4.4 oz). A respectable number of the 90 g (3.2 oz) eastern mole (Scalopus aquaticus) were recorded in studies from Oklahoma and Kansas. Four species of bat have been recorded in their foods. The red-tailed hawks local to the large cave colonies of 12.3 g (0.43 oz) Mexican free-tailed bats (Tadarida brasiliensis) in Texas can show surprising agility, some of the same hawks spending their early evening and early morning hours in flight patrolling the cave entrances in order to stoop suddenly on these flighted mammals. Larger miscellaneous mammalian prey are either usually taken as juveniles, like the nine-banded armadillo (Dasypus novemcinctus), or largely as carrion, like the Virginia opossum (Didelphis virginiana). Small carnivorans may be taken, usually consisting of much smaller mustelids, like the least weasels (Mustela nivalis), stoats (Mustela erminea), and long-tailed weasels (Neogale frenata). slightly larger carnivores, such as small Indian mongooses (Herpestes auropunctatus), ringtails (Bassariscus astutus), small American minks (Neovison vison) and even adult striped skunk (Mephitis mephitis), which can be much larger than a fully grown hawk, was reportedly taken by red-tailed hawks. Additionally, red-tailed hawks are considered as potential predators of white-nosed coati (Nasua narica) and kit fox (Vulpes macrotis) Remains of exceptionally large carnivoran species, such as domestic cats (Felis catus), red fox ( Vulpes vulpes) and common raccoon (Procyon lotor) are sometimes found amongst their foods, but most are likely taken as juveniles or consumed only as carrion. Many of these medium-sized carnivorans are probably visited as roadkill, especially during the sparser winter months, but carrion has turned up more widely than previously thought. Some nests have been found (to the occasional "shock" of researchers) with body parts from large domestic stock like sheep (Ovis aries), pigs (Sus domesticus), horses (Equus caballus ) and cattle (Bos taurus) (not to mention wild varieties like deer), which red-tails must visit when freshly dead out on pastures and take a couple of talonfuls of meat. In one instance, a red-tailed hawk was observed to kill a small but seemingly healthy lamb. These are born heavier than most red-tails at 1,500 g (3.3 lb) but in this case, the hawk was scared away before it could consume its kill by the rifle fire of the shepherd who witnessed the instance.

 

Birds

Like most (but not all) Buteo hawks, red-tailed hawks do not primarily hunt birds in most areas, but can take them fairly often whenever they opportune upon some that are vulnerable. Birds are, by far, the most diverse class in the red-tailed hawk's prey spectrum, with well over 200 species known in their foods In most circumstances where birds become the main food of red-tailed hawks, it is in response to ample local populations of galliforms. As these are meaty, mostly terrestrial birds which usually run rather than fly from danger (although all wild species in North America are capable of flight), galliforms are ideal avian prey for red-tails. Some 23 species of galliforms are known to be taken by red-tailed hawks, about a third of these being species introduced by humans. Native quails of all five North American species may expect occasional losses. All 12 species of grouse native to North America are also occasionally included in their prey spectrum. In the state of Wisconsin, two large studies, from Waupun and Green County, found the main prey species to be the ring-necked pheasant (Phasianus colchicus), making up 22.7% of a sample of 176 and 33.8% of a sample of 139, respectively. With a body mass averaging 1,135 g (2.502 lb), adult pheasants are among the largest meals that male red-tails are likely to deliver short of adult rabbits and hares and therefore these nests tend to be relatively productive. Despite being not native to North America, pheasants usually live in a wild state. Chickens (Gallus gallus domesticus) are also taken throughout North America, with all Wisconsin studies also found large numbers of them, making up as much as 14.4% of the diet. Many studies reflect that free-ranging chickens are vulnerable to red-tailed hawks although somewhat lesser numbers are taken by them overall in comparison to nocturnal predators (i.e. owls and foxes) and goshawks. In Rochester, Alberta, fairly large numbers of ruffed grouse (Bonasa umbellus) were taken but relatively more juveniles were taken of this species than the two other main contributors to biomass here, snowshoe hare and Townsend's ground squirrel, as they are fairly independent early on and more readily available. Here the adult grouse was estimated to average 550 g (1.21 lb) against the average juvenile which in mid-summer averaged 170 g (6.0 oz).

 

Beyond galliforms, three other quite different families of birds make the most significant contributions to the red-tailed hawk's avian diet. None of these three families are known as particularly skilled or swift fliers, but are generally small enough that they would generally easily be more nimble in flight. One of these are the woodpeckers, if only for one species, the 131.6 g (4.64 oz) northern flicker (Colaptes auratus), which was the best represented bird species in the diet in 27 North American studies and was even the fourth most often detected prey species of all. Woodpeckers are often a favorite in the diet of large raptors as their relatively slow, undulating flight makes these relatively easy targets. The flicker in particular is a highly numerous species that has similar habitat preferences to red-tailed hawks, preferring fragmented landscapes with trees and openings or parkland-type wooded mosaics, and often forage on the ground for ants, which may make them even more susceptible. Varied other woodpecker species may turn up in their foods, from the smallest to the largest extant in North America, but are much more infrequently detected in dietary studies. Another family relatively often selected prey family are corvids, which despite their relatively large size, formidable mobbing abilities and intelligence are also slower than average fliers for passerines. 14 species of corvid are known to fall prey to red-tailed hawks. In the Kaibab Plateau, the 128 g (4.5 oz) Steller's jay (Cyanocitta stelleri) were the fourth most identified prey species (10.3% of the diet). 453 g (0.999 lb) American crows are also regularly detected supplemental prey in several areas. Even the huge common raven (Corvus corax), at 1,050 g (2.31 lb) at least as large as red-tailed hawk itself, may fall prey to red-tails, albeit very infrequently and only in a well-staged ambush. One of the most surprising heavy contributors are the icterids, despite their slightly smaller size and tendency to travel in large, wary flocks, 12 species are known to be hunted. One species pair, the meadowlarks, are most often selected as they do not flock in the same ways as many other icterids and often come to the ground, throughout their life history, rarely leaving about shrub-height. The 100.7 g (3.55 oz) western meadowlark (Sturnella neglecta), in particular, was the third most often detected bird prey species in North America. Red-winged blackbirds (Agelaius phoeniceus) which are probably too small, at an average weight of 52.4 g (1.85 oz), and fast for a red-tailed hawk to ever chase on the wing (and do travel in huge flocks, especially in winter) are nonetheless also quite often found in their diet, representing up to 8% of the local diet for red-tails. It is possible that males, which are generally bold and often select lofty perches from which to display, are most regularly ambushed. One bird species that often flocks with red-winged blackbirds in winter is even better represented in the red-tail's diet, the non-native 78 g (2.8 oz) European starling (Sturnus vulgaris), being the second most numerous avian prey species and seventh overall in North America. Although perhaps most vulnerable when caught unaware while calling atonally on a perch, a few starlings (or various blackbirds) may be caught by red-tails which test the agile, twisting murmurations of birds by flying conspicuously towards the flock, to intentionally disturb them and possibly detect lagging, injured individual birds that can be caught unlike healthy birds. However, this behavior has been implied rather than verified.

 

Over 50 passerine species from various other families beyond corvids, icterids and starlings are included in the red-tailed hawks' prey spectrum but are caught so infrequently as to generally not warrant individual mention. Non-passerine prey taken infrequently may include but are not limited to pigeons and doves, cuckoos, nightjars, kingfishers and parrots. However, of some interest, is the extreme size range of birds that may be preyed upon. Red-tailed hawks in Caribbean islands seem to catch small birds more frequently due to the paucity of vertebrate prey diversity here.

spanwidth min.: 54 cm

spanwidth max.: 60 cm

size min.: 32 cm

size max.: 36 cm

Breeding

incubation min.: 11 days

incubation max.: 12 days

fledging min.: 17 days

fledging max.: 17 days

broods 15

eggs min.: 1

eggs max.: 25

 

Cuach

 

Status: Widespread summer visitor to Ireland from April to August.

 

Conservation Concern: Green-listed in Ireland. The European population is currently evaluated as secure.

 

Identification: Despite its obvious song, relatively infrequently seen. In flight, can be mistaken for a bird of prey such as Sparrowhawk, but has rapid wingbeats below the horizontal plane - ie. the wings are not raised above the body. Adult male Cuckoos are a uniform grey on the head, neck, back, wings and tail. The underparts are white with black barring. Adult females can appear in one of two forms. The so-called grey-morph resembles the adult male plumage, but has throat and breast barred black and white with yellowish wash. The rufous-morph has the grey replaced by rufous, with strong black barring on the wings, back and tail. Juvenile Cuckoos resemble the female rufous-morph, but are darker brown above.

 

Similar Species: Sparrowhawk

 

Call: The song is probably one of the most recognisable and well-known of all Irish bird species. The male gives a distinctive “wuck-oo”, which is occasionally doubled “wuck-uck-ooo”. The female has a distinctive bubbling “pupupupu”. The song period is late April to late June.

 

Diet: Mainly caterpillars and other insects.

 

Breeding: Widespread in Ireland, favouring open areas which hold their main Irish host species – Meadow Pipit. Has a remarkable breeding biology unlike any other Irish breeding species.

 

Wintering: Cuckoos winter in central and southern Africa.

 

Where to See: Occurs throughout Ireland though nowhere especially common. Good areas to see Cuckoo are the Burren and Connemara, which hold the highest density of breeding pairs.

  

Physical characteristics

 

Forests and woodlands, both coniferous and deciduous, second growth, open wooded areas, wooded steppe, scrub, heathland, also meadows, reedbeds. Lowlands and moorlands and hill country to 2 km.

 

Habitat

 

Forests and woodlands, both coniferous and deciduous, second growth, open wooded areas, wooded steppe, scrub, heathland, also meadows, reedbeds. Lowlands and moorlands and hill country to 2 km. Food and Feeding

 

Other details

 

Cuculus canorus is a widespread summer visitor to Europe, which accounts for less than half of its global breeding range. Its European breeding population is very large (>4,200,000 pairs), and was stable between 1970-1990. Although there were declines in many western populations-most notably France-during 1990-2000, most populations in the east, including key ones in Russia and Romania, were stable, and the species underwent only a slight decline overall

 

Feeding

 

Diet based on insects, mainly caterpillars, also dragonflies, mayflies, damselflies, crickets, and cicadas. Sometimes, spiders, snails, rarely fruit. Preys on eggs and nestling of small birds.

 

Conservation

 

This species has a large range, with an estimated global Extent of Occurrence of 10,000,000 km². It has a large global population, including an estimated 8,400,000-17,000,000 individuals in Europe (BirdLife International in prep.). Global population trends have not been quantified, but populations appear to be stable so the species is not believed to approach the thresholds for the population decline criterion of the IUCN Red List (i.e. declining more than 30% in ten years or three generations). For these reasons, the species is evaluated as Least Concern. [conservation status from birdlife.org]

 

Breeding

 

May-Jun in NW Europe, Apr-May in Algeria, Apr-Jul in India and Myanmar. Brood-parasitic, hosts include many insectivorous songbird species, like: flycatchers, chats, warblers, pipits, wagtails and buntigs. Often mobbed by real or potential hosts near their nests. Eggs polymorphic in color and pattern, closely match those of host in color and pattern. Nestling period 17-18 days, evicts host's eggs and chicks.

 

Migration

 

Migratory in N of range, arriving in SW Britain mainly Apr - May, when occasionally recorded in small parties, and even in one flock of 50+ birds; also seasonal in hill country from Assam and Chin Hills to Shan States, where present Mar - Aug. Resident in tropical lowland areas of S Asia. Winter resident in sub-Saharan Africa and in Sri Lanka. W Palearctic populations migrate to Africa, where a Dutch-ringed juvenile found in Togo in Oct and a British-ringed juvenile found in Cameroon in Jan; migrants appear in N Senegal as early as late Jul through Oct; in W Africa nearly all records are in autumn ( Sept - Dec), birds apparently continuing on to C & S Africa. Race bangsi occurs on passage in W Africa, and winters S of equator from W Africa to L Tanganyika. Asian populations of nominate canorus and bakeri winter in India, SE Asia and Philippines, also in Africa, but the extent of migration of Asian birds to Africa is unknown; some subtelephonus migrate through Middle East and occur in winter from Uganda and E Zaire to Zimbabwe, Mozambique and Natal. Mainly a passage migrant in Middle East, though some breed in region. Migrants also appear on islands in W Indian Ocean ( Seychelles, Aldabra). Nominate canorus accidental in Iceland, Faeroes, Azores, Madeira, Canary Is and Cape Verde Is, rarely also Alaska and eastern N America; one record of canorus in Indonesia, off W Java in winter. Autumn migration starts in August and continues until October. The main passage through Egypt is in September and the first half of October, with a peak in the third week of September (Goodman & Meininger 1989). Southward movement through Africa lasts from September to December and is linked to the occurrence of rainfall and the growth of cover.

 

Scops-owls are Strigidae (typical owls) belong to the genus Otus. Approximately 45 living species are known, but new ones are frequently recognized and unknown ones are still being discovered every few years or so, especially in Indonesia. For most of the 20th century, this genus included the American screech-owls which are now again separated in Megascops based on a range of behavioral, biogeographical, morphological and DNA sequence data.

 

Scops-owls in the modern sense are restricted to the Old World, except for a single North American species - the Flammulated Owl - that is only provisionally placed here and is likely to be moved out of Otus eventually. See below for details.

 

As usual for owls, female scops-owls are usually larger than the males of their species, with owls of both sexes being compact in size and shape. All of the birds in this genus are small and agile. Scops-owls are colored in various brownish hues, sometimes with a lighter underside and/or face, which helps to camouflage them against the bark of trees. Some are polymorphic, occurring in a greyish- and a reddish-brown morph.

The Sphingidae are amongst the largest, most easily recognised, and best known Lepidopterans. They are known variably as "hawk", "sphinx", "hummingbird" or "hornworm" moths.

Adults are medium- to very-large-sized, having a stout body with the abdomen typically tapering posteriorly. The forewing is narrow and hindwing relatively short, giving the moths a very streamlined profile. The proboscis usually is well developed, sometimes much longer than the body, and used to imbibe nectar while hovering in flight, hummingbird-style, and in some habitats sphingids have significant roles in pollination. Most are nocturnal, extremely strong fliers, amongst the fastest insects, and several are well-known long-distance migrants. Some genera are diurnal, and a few resemble bees.

The often polymorphic larvae are equally recognisable (in most cases) owing to a caudal horn or button. Pupation usually occurs without a cocoon, in soil or ground litter.

There are around 1450 described species in about 200 genera, distributed worldwide, but best represented in the tropics.

 

(text adapted from Encyclopedia of Insects (Second Edition), Jerry A. Powell, 2009)

 

Collage images from Pu'er, Yunnan, China

 

You can browse all of the Sphingid moth images in my photostream HERE...

 

see comments for additional image...

Tenagogerris euphrosyne

Family: Gerridae

Order: Hemiptera

 

The striders are hemipterans or "true-bugs" - insects that feed by piercing and sucking food through the proboscis.

 

In the first photo, this strider has located a small insect on the surface of the pond. It runs its antennae over it to determine suitability for eating. In the second photo, the front legs have clamped the prey, the proboscis has been lowered and the insect is being prepared for lunch!

 

The striders walk on water, thanks to hydrophobic hairs on the legs. The hairs extend across the whole body too, making the insect able to repel splashes of water. They are so successful in their niche, that the strider family, the Gerridae, are widespread with some 1700 species described, 10% of which are marine.

 

The Gerridae are polymorphic in that they can have wings in one generation, when there might be a need to relocate to a new water body. The next generation may not have wings however, if the current environment is stable.

  

DSC05051 - DSC05055 composite

Produjendo produciendo trabajando desde su multiforme organ-ization.

 

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Produsing producing working from its polymorphic organ-ization.

 

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fBwO cBwO eBwo

 

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View On Black

  

কালিম । Common Mormon (Female; Form: romulus) - Papilio polytes

 

This female form of the Common Mormon mimics the Common Rose very closely but lacks red markings on body. This is the commonest form wherever the Common Rose flies. The female of the Common Mormon is polymorphic. In South Asia, it has three forms or morphs. These are as follows: cyrus, stichius, romulus.

 

The Common Mormon (Papilio polytes) is a common species of swallowtail butterfly (family: papilionidae) widely distributed across Asia. Seen round the year throughout India from plains up to 2000m. This butterfly is known for the mimicry displayed by the numerous forms of its females.

  

Host Plant: Ixora coccinea; Rangan (রঙ্গন, Rugmini in Hindi, commonly known as the Jungle Geranium, Flame of the Woods, and Jungle Flame from Rubiaceae family) is an exotic bright red flower, bloom as a flower bunch comprises of lot of small red flowers at the top of branch. Each red flower has four petals and holds four yellow stamen(no filament) between the petals. Flower blooms more or less throughout the year, but best during the rainy season.

 

Tajpur Sea Beach, Bay of Bengal

Monsoon Images of Bengal, India

 

Sigma 150-600mm 5-6.3 DG HSM OS Contemporary plus Nikon D7000

The Variable Oystercatcher (Haematopus unicolour, torea-pango in Maori), a species of wading bird, is endemic to New Zealand. They are also known as 'red bills'. "Variable" refers to the frontal plumage, which ranges from pied through mottled to all black. They are polymorphic meaning they have different genetic variants. Blacker birds are more common in the south; all Stewart Island variable oystercatchers are black.

 

DISCRIPTION

Variables have pink legs, an orange eye ring and red beaks. Males are around 678 grams and females slightly larger at around 724 grams. Variables can be identified as they are slightly larger than the (black and white) South Island Pied Oystercatchers (SIPOs) which are around 550 grams. Occasionally totally black, but if they are pied (black and white) they can be easily confused with SIPOs. The variable species has less definition between the black and the white area, as well as a mottled band on the leading edges of the underwing. Variables also have a smaller white rump patch which is only a band across the base of the tail rather than a wide wedge shape reaching up to the middle of the back as in the SIPO. When mottled they are sometimes called 'smudgies'.

 

BREEDING

Variables are often seen in pairs on the coast all around New Zealand. Once mated pairs rarely divorce. They breed in North Island, South Island, Stewart Island, and Chatham Islands. They do not breed inland or beside rivers, although the SIPO does. They nest on the shore between rocks or on sand dunes by making a scrape out of the sand or shingle, sometimes lined with some seaweed. During breeding, the pair will defend their territory, sometimes aggressively. They usually lay 2 to 3 eggs, but they can lay up to 5. The eggs are typically stone coloured with small brown patches all over. Eggs hatch in 25 to 32 days. Chicks are well camouflaged by their colour and can fly in about 6 weeks. After breeding they may be seen within or on the edges of flocks of SIPO which also have vivid orange beaks. After breeding they may even form small flocks of their own. The bird lives up to about 27 years.

 

FOOD

They are noisy and talkative birds, and when in flight they make a high pitched 'kleep kleep' sound. The birds feed on molluscs, crabs and marine worms. After heavy rain, they sometime go inland in search of earthworms. They can open a shellfish by either hammering a hole in it or getting the bill between the two shells (of a bivalve) and twisting them apart.

Source: Wikipedia

 

Monographia Aranearum = Monographie der Spinnen /.

Nurnberg :Lechner,[182..

biodiversitylibrary.org/page/52001664

Maize (/meɪz/ MAYZ; Zea mays subsp. mays, from Spanish: maíz after Taino: mahiz), also known as corn (North American and Australian English), is a cereal grain first domesticated by indigenous peoples in southern Mexico about 10,000 years ago. The leafy stalk of the plant produces pollen inflorescences and separate ovuliferous inflorescences called ears that yield kernels or seeds, which are fruits.

 

Maize has become a staple food in many parts of the world, with the total production of maize surpassing that of wheat or rice. In addition to being consumed directly by humans (often in the form of masa), maize is also used for corn ethanol, animal feed and other maize products, such as corn starch and corn syrup. The six major types of maize are dent corn, flint corn, pod corn, popcorn, flour corn, and sweet corn. Sugar-rich varieties called sweet corn are usually grown for human consumption as kernels, while field corn varieties are used for animal feed, various corn-based human food uses (including grinding into cornmeal or masa, pressing into corn oil, and fermentation and distillation into alcoholic beverages like bourbon whiskey), and as chemical feedstocks. Maize is also used in making ethanol and other biofuels.

 

Maize is widely cultivated throughout the world, and a greater weight of maize is produced each year than any other grain. In 2014, total world production was 1.04 billion tonnes. Maize is the most widely grown grain crop throughout the Americas, with 361 million metric tons grown in the United States alone in 2014. Genetically modified maize made up 85% of the maize planted in the United States in 2009. Subsidies in the United States help to account for its high level of cultivation of maize and its position as the largest producer in the world.

 

HISTORY

PRE-COLUMBIAN DEVELOPMENT

Maize is a cultigen; human intervention is required for it to propagate. Whether or not the kernels fall off the cob on their own is a key piece of evidence used in archaeology to distinguish domesticated maize from its naturally-propagating teosinte ancestor. Genetic evidence can also be used to determine when various lineages split.

 

Most historians believe maize was domesticated in the Tehuacán Valley of Mexico. Recent research in the early 21st century has modified this view somewhat; scholars now indicate the adjacent Balsas River Valley of south-central Mexico as the center of domestication.

 

An influential 2002 study by Matsuoka et al. has demonstrated that, rather than the multiple independent domestications model, all maize arose from a single domestication in southern Mexico about 9,000 years ago. The study also demonstrated that the oldest surviving maize types are those of the Mexican highlands. Later, maize spread from this region over the Americas along two major paths. This is consistent with a model based on the archaeological record suggesting that maize diversified in the highlands of Mexico before spreading to the lowlands.

 

Archaeologist Dolores Piperno has said:

 

A large corpus of data indicates that [maize] was dispersed into lower Central America by 7600 BP [5600 BC] and had moved into the inter-Andean valleys of Colombia between 7000 and 6000 BP [5000–4000 BC].

— Dolores Piperno, The Origins of Plant Cultivation and Domestication in the New World Tropics: Patterns, Process, and New Developments

 

Since then, even earlier dates have been published.

 

According to a genetic study by Embrapa, corn cultivation was introduced in South America from Mexico, in two great waves: the first, more than 6000 years ago, spread through the Andes. Evidence of cultivation in Peru has been found dating to about 6700 years ago. The second wave, about 2000 years ago, through the lowlands of South America.

 

The earliest maize plants grew only small, 25-millimetre-long (1 in) corn cobs, and only one per plant. In Jackson Spielvogel's view, many centuries of artificial selection (rather than the current view that maize was exploited by interplanting with teosinte) by the indigenous people of the Americas resulted in the development of maize plants capable of growing several cobs per plant, which were usually several centimetres/inches long each. The Olmec and Maya cultivated maize in numerous varieties throughout Mesoamerica; they cooked, ground and processed it through nixtamalization. It was believed that beginning about 2500 BC, the crop spread through much of the Americas. Research of the 21st century has established even earlier dates. The region developed a trade network based on surplus and varieties of maize crops.

 

Mapuches of south-central Chile cultivated maize along with quinoa and potatoes in pre-Hispanic times; however, potato was the staple food of most Mapuches, "specially in the southern and coastal [Mapuche] territories where maize did not reach maturity". Before the expansion of the Inca Empire maize was traded and transported as far south as 40°19' S in Melinquina, Lácar Department. In that location maize remains were found inside pottery dated to 730 ± 80 BP and 920 ± 60 BP. Probably this maize was brought across the Andes from Chile. The presence of maize in Guaitecas Archipelago (43°55' S), the southernmost outpost of pre-Hispanic agriculture, is reported by early Spanish explorers. However the Spanish may have misidentified the plant.

 

COLUMBIAN EXCHANGE

After the arrival of Europeans in 1492, Spanish settlers consumed maize, and explorers and traders carried it back to Europe and introduced it to other countries. Spanish settlers far preferred wheat bread to maize, cassava, or potatoes. Maize flour could not be substituted for wheat for communion bread, since in Christian belief only wheat could undergo transubstantiation and be transformed into the body of Christ. Some Spaniards worried that by eating indigenous foods, which they did not consider nutritious, they would weaken and risk turning into Indians. "In the view of Europeans, it was the food they ate, even more than the environment in which they lived, that gave Amerindians and Spaniards both their distinctive physical characteristics and their characteristic personalities." Despite these worries, Spaniards did consume maize. Archeological evidence from Florida sites indicate they cultivated it as well.

 

Maize spread to the rest of the world because of its ability to grow in diverse climates. It was cultivated in Spain just a few decades after Columbus's voyages and then spread to Italy, West Africa and elsewhere. Widespread cultivation most likely began in southern Spain in 1525, after which it quickly spread to the rest of the Spanish Empire including its territories in Italy (and, from there, to other Italian states). Maize had many advantages over wheat and barley; it yielded two and a half times the food energy per unit cultivated area, could be harvested in successive years from the same plot of land, and grew in wildly varying altitudes and climates, from relatively dry regions with only 250 mm (10 in) of annual rainfall to damp regions with over 5,000 mm (200 in). By the 17th century it was a common peasant food in Southwestern Europe, including Portugal, Spain, southern France, and Italy. By the 18th century, it was the chief food of the southern French and Italian peasantry, especially in the form of polenta in Italy.

Names

 

The word maize derives from the Spanish form of the indigenous Taíno word for the plant, mahiz. It is known by other names around the world.

 

The word "corn" outside the US, Canada, Australia, and New Zealand refers to any cereal crop, its meaning understood to vary geographically to refer to the local staple. In the United States, Canada, Australia, and New Zealand, corn primarily means maize; this usage started as a shortening of "Indian corn". "Indian corn" primarily means maize (the staple grain of indigenous Americans), but can refer more specifically to multicolored "flint corn" used for decoration.

 

In places outside the US, Canada, Australia, and New Zealand, corn often refers to maize in culinary contexts. The narrower meaning is usually indicated by some additional word, as in sweet corn, sweetcorn, corn on the cob, baby corn, the puffed confection known as popcorn and the breakfast cereal known as corn flakes.

 

In Southern Africa, maize is commonly called mielie (Afrikaans) or mealie (English), words derived from the Portuguese word for maize, milho.

 

Maize is preferred in formal, scientific, and international usage because it refers specifically to this one grain, unlike corn, which has a complex variety of meanings that vary by context and geographic region. Maize is used by agricultural bodies and research institutes such as the FAO and CSIRO. National agricultural and industry associations often include the word maize in their name even in English-speaking countries where the local, informal word is something other than maize; for example, the Maize Association of Australia, the Indian Maize Development Association, the Kenya Maize Consortium and Maize Breeders Network, the National Maize Association of Nigeria, the Zimbabwe Seed Maize Association.

 

STRUCTURE AND PHYSIOLOGY

The maize plant is often 3 m (10 ft) in height, though some natural strains can grow 13 m (43 ft). The stem is commonly composed of 20 internodes of 18 cm (7 in) length. The leaves arise from the nodes, alternately on opposite sides on the stalk. A leaf, which grows from each node, is generally 9 cm (3+1⁄2 in) in width and 120 cm (3 ft 11 in) in length.

 

Ears develop above a few of the leaves in the midsection of the plant, between the stem and leaf sheath, elongating by around 3 mm (1⁄8 in) per day, to a length of 18 cm (7 in) with 60 cm (24 in) being the maximum alleged in the subspecies. They are female inflorescences, tightly enveloped by several layers of ear leaves commonly called husks. Certain varieties of maize have been bred to produce many additional developed ears. These are the source of the "baby corn" used as a vegetable in Asian cuisine.

 

The apex of the stem ends in the tassel, an inflorescence of male flowers. When the tassel is mature and conditions are suitably warm and dry, anthers on the tassel dehisce and release pollen. Maize pollen is anemophilous (dispersed by wind), and because of its large settling velocity, most pollen falls within a few meters of the tassel.

 

Elongated stigmas, called silks, emerge from the whorl of husk leaves at the end of the ear. They are often pale yellow and 18 cm (7 in) in length, like tufts of hair in appearance. At the end of each is a carpel, which may develop into a "kernel" if fertilized by a pollen grain. The pericarp of the fruit is fused with the seed coat referred to as "caryopsis", typical of the grasses, and the entire kernel is often referred to as the "seed". The cob is close to a multiple fruit in structure, except that the individual fruits (the kernels) never fuse into a single mass. The grains are about the size of peas, and adhere in regular rows around a white, pithy substance, which forms the ear. The maximum size of kernels is reputedly 2.5 cm (1 in). An ear commonly holds 600 kernels. They are of various colors: blackish, bluish-gray, purple, green, red, white and yellow. When ground into flour, maize yields more flour with much less bran than wheat does. It lacks the protein gluten of wheat and, therefore, makes baked goods with poor rising capability. A genetic variant that accumulates more sugar and less starch in the ear is consumed as a vegetable and is called sweet corn. Young ears can be consumed raw, with the cob and silk, but as the plant matures (usually during the summer months), the cob becomes tougher and the silk dries to inedibility. By the end of the growing season, the kernels dry out and become difficult to chew without cooking them tender first in boiling water.

 

Planting density affects multiple aspects of maize. Modern farming techniques in developed countries usually rely on dense planting, which produces one ear per stalk. Stands of silage maize are yet denser,[citation needed] and achieve a lower percentage of ears and more plant matter.

 

Maize is a facultative short-day plant and flowers in a certain number of growing degree days > 10 °C (50 °F) in the environment to which it is adapted. The magnitude of the influence that long nights have on the number of days that must pass before maize flowers is genetically prescribed and regulated by the phytochrome system.

Photoperiodicity can be eccentric in tropical cultivars such that the long days characteristic of higher latitudes allow the plants to grow so tall that they do not have enough time to produce seed before being killed by frost. These attributes, however, may prove useful in using tropical maize for biofuels.

 

Immature maize shoots accumulate a powerful antibiotic substance, 2,4-dihydroxy-7-methoxy-1,4-benzoxazin-3-one (DIMBOA). DIMBOA is a member of a group of hydroxamic acids (also known as benzoxazinoids) that serve as a natural defense against a wide range of pests, including insects, pathogenic fungi and bacteria. DIMBOA is also found in related grasses, particularly wheat. A maize mutant (bx) lacking DIMBOA is highly susceptible to attack by aphids and fungi. DIMBOA is also responsible for the relative resistance of immature maize to the European corn borer (family Crambidae). As maize matures, DIMBOA levels and resistance to the corn borer decline.

 

Because of its shallow roots, maize is susceptible to droughts, intolerant of nutrient-deficient soils, and prone to be uprooted by severe winds.

 

While yellow maizes derive their color from lutein and zeaxanthin, in red-colored maizes, the kernel coloration is due to anthocyanins and phlobaphenes. These latter substances are synthesized in the flavonoids synthetic pathway from polymerization of flavan-4-ols by the expression of maize pericarp color1 (p1) gene which encodes an R2R3 myb-like transcriptional activator of the A1 gene encoding for the dihydroflavonol 4-reductase (reducing dihydroflavonols into flavan-4-ols) while another gene (Suppressor of Pericarp Pigmentation 1 or SPP1) acts as a suppressor. The p1 gene encodes an Myb-homologous transcriptional activator of genes required for biosynthesis of red phlobaphene pigments, while the P1-wr allele specifies colorless kernel pericarp and red cobs, and unstable factor for orange1 (Ufo1) modifies P1-wr expression to confer pigmentation in kernel pericarp, as well as vegetative tissues, which normally do not accumulate significant amounts of phlobaphene pigments. The maize P gene encodes a Myb homolog that recognizes the sequence CCT/AACC, in sharp contrast with the C/TAACGG bound by vertebrate Myb proteins.

 

The ear leaf is the leaf most closely associated with a particular developing ear. This leaf and above contribute 70% to 75% to 90% of grain fill. Therefore fungicide application is most important in that region in most disease environments.

 

ABNORMAL FLOWERS

Maize flowers may sometimes exhibit mutations that lead to the formation of female flowers in the tassel. These mutations, ts4 and Ts6, prohibit the development of the stamen while simultaneously promoting pistil development. This may cause inflorescences containing both male and female flowers, or hermaphrodite flowers.

 

GENETICS

Maize is an annual grass in the family Gramineae, which includes such plants as wheat, rye, barley, rice, sorghum, and sugarcane. There are two major species of the genus Zea (out of six total): Zea mays (maize) and Zea diploperennis, which is a perennial type of teosinte. The annual teosinte variety called Zea mays mexicana is the closest botanical relative to maize. It still grows in the wild as an annual in Mexico and Guatemala.

 

Many forms of maize are used for food, sometimes classified as various subspecies related to the amount of starch each has:

 

Flour corn: Zea mays var. amylacea

Popcorn: Zea mays var. everta

Dent corn : Zea mays var. indentata

Flint corn: Zea mays var. indurata

Sweet corn: Zea mays var. saccharata and Zea mays var. rugosa

Waxy corn: Zea mays var. ceratina

Amylomaize: Zea mays

Pod corn: Zea mays var. tunicata Larrañaga ex A. St. Hil.

Striped maize: Zea mays var. japonica

 

This system has been replaced (though not entirely displaced) over the last 60 years by multivariable classifications based on ever more data. Agronomic data were supplemented by botanical traits for a robust initial classification, then genetic, cytological, protein and DNA evidence was added. Now, the categories are forms (little used), races, racial complexes, and recently branches.

 

Maize is a diploid with 20 chromosomes (n=10). The combined length of the chromosomes is 1500 cM. Some of the maize chromosomes have what are known as "chromosomal knobs": highly repetitive heterochromatic domains that stain darkly. Individual knobs are polymorphic among strains of both maize and teosinte.

 

Barbara McClintock used these knob markers to validate her transposon theory of "jumping genes", for which she won the 1983 Nobel Prize in Physiology or Medicine. Maize is still an important model organism for genetics and developmental biology today.

 

The centromeres have two types of structural components, both of which are found only in the centromeres: Large arrays of CentC, a short satellite DNA; and a few of a family of retrotransposons. The B chromosome, unlike the others, contains an additional repeat which extends into neighboring areas of the chromosome. Centromeres can accidentally shrink during division and still function, although it is thought this will fail if it shrinks below a few hundred kilobase. Kinetochores contain RNA originating from centromeres. Centromere regions can become inactive, and can continue in that state if the chromosome still has another active one.

 

The Maize Genetics Cooperation Stock Center, funded by the USDA Agricultural Research Service and located in the Department of Crop Sciences at the University of Illinois at Urbana-Champaign, is a stock center of maize mutants. The total collection has nearly 80,000 samples. The bulk of the collection consists of several hundred named genes, plus additional gene combinations and other heritable variants. There are about 1000 chromosomal aberrations (e.g., translocations and inversions) and stocks with abnormal chromosome numbers (e.g., tetraploids). Genetic data describing the maize mutant stocks as well as myriad other data about maize genetics can be accessed at MaizeGDB, the Maize Genetics and Genomics Database.

 

In 2005, the US National Science Foundation (NSF), Department of Agriculture (USDA) and the Department of Energy (DOE) formed a consortium to sequence the B73 maize genome. The resulting DNA sequence data was deposited immediately into GenBank, a public repository for genome-sequence data. Sequences and genome annotations have also been made available throughout the project's lifetime at the project's official site.

 

Primary sequencing of the maize genome was completed in 2008. On November 20, 2009, the consortium published results of its sequencing effort in Science. The genome, 85% of which is composed of transposons, was found to contain 32,540 genes (By comparison, the human genome contains about 2.9 billion bases and 26,000 genes). Much of the maize genome has been duplicated and reshuffled by helitrons—group of rolling circle transposons.

 

In Z. mays and various other angiosperms the MADS-box motif is involved in floral development. Early study in several angiosperm models including Z. mays was the beginning of research into the molecular evolution of floral structure in general, as well as their role in nonflowering plants.

 

EVOLUTION

As with many plants and animals, Z. mays has a positive correlation between effective population size and the magnitude of selection pressure. Z. m. having an EPS of ~650,000, it clusters with others of about the same EPS, and has 79% of its amino acid sites under selection.

 

Recombination is a significant source of diversity in Z. mays. (Note that this finding supersedes previous studies which showed no such correlation.)

 

This recombination/diversity effect is seen throughout plants but is also found to not occur – or not as strongly – in regions of high gene density. This is likely the reason that domesticated Z. mays has not seen as much of an increase in diversity within areas of higher density as in regions of lower density, although there is more evidence in other plants.

 

Some lines of maize have undergone ancient polyploidy events, starting 11m years ago. Over that time ~72% of polyploid duplicated genes have been retained, which is higher than other plants with older polyploidy events. Thus maize may be due to lose more duplicate genes as time goes along, similar to the course followed by the genomes of other plants. If so - if gene loss has merely not occurred yet - that could explain the lack of observed positive selection and lower negative selection which are observed in otherwise similar plants, i.e. also naturally outcrossing and with similar effective population sizes.

 

Ploidy does not appear to influence EPS or magnitude of selection effect in maize.

 

BREEDING

Maize reproduces sexually each year. This randomly selects half the genes from a given plant to propagate to the next generation, meaning that desirable traits found in the crop (like high yield or good nutrition) can be lost in subsequent generations unless certain techniques are used.

 

Maize breeding in prehistory resulted in large plants producing large ears. Modern breeding began with individuals who selected highly productive varieties in their fields and then sold seed to other farmers. James L. Reid was one of the earliest and most successful developing Reid's Yellow Dent in the 1860s. These early efforts were based on mass selection. Later breeding efforts included ear to row selection (C. G. Hopkins c. 1896), hybrids made from selected inbred lines (G. H. Shull, 1909), and the highly successful double cross hybrids using four inbred lines (D. F. Jones c. 1918, 1922). University supported breeding programs were especially important in developing and introducing modern hybrids. By the 1930s, companies such as Pioneer devoted to production of hybrid maize had begun to influence long-term development. Internationally important seed banks such as the International Maize and Wheat Improvement Center (CIMMYT) and the US bank at the Maize Genetics Cooperation Stock Center University of Illinois at Urbana-Champaign maintain germplasm important for future crop development.

 

Since the 1940s the best strains of maize have been first-generation hybrids made from inbred strains that have been optimized for specific traits, such as yield, nutrition, drought, pest and disease tolerance. Both conventional cross-breeding and genetic modification have succeeded in increasing output and reducing the need for cropland, pesticides, water and fertilizer. There is conflicting evidence to support the hypothesis that maize yield potential has increased over the past few decades. This suggests that changes in yield potential are associated with leaf angle, lodging resistance, tolerance of high plant density, disease/pest tolerance, and other agronomic traits rather than increase of yield potential per individual plant.

 

Tropical landraces remain an important and underutilized source of resistance alleles for for disease and for herbivores. Notable discoveries of rare alleles for this purpose were made by Dao et al 2014 and Sood et al 2014.

 

GLOBAL PROGRAM

CIMMYT operates a conventional breeding program to provide optimized strains. The program began in the 1980s. Hybrid seeds are distributed in Africa by the Drought Tolerant Maize for Africa project.

 

GENETIC MODIFICATION

Genetically modified (GM) maize was one of the 26 GM crops grown commercially in 2016. The vast majority of this is Bt maize. Grown since 1997 in the United States and Canada, 92% of the US maize crop was genetically modified in 2016 and 33% of the worldwide maize crop was GM in 2016. As of 2011, Herbicide-tolerant maize varieties were grown in Argentina, Australia, Brazil, Canada, China, Colombia, El Salvador, the European Union, Honduras, Japan, Korea, Malaysia, Mexico, New Zealand, Philippines, the Russian Federation, Singapore, South Africa, Taiwan, Thailand, and the United States. Insect-resistant maize was grown in Argentina, Australia, Brazil, Canada, Chile, China, Colombia, Egypt, the European Union, Honduras, Japan, Korea, Malaysia, Mexico, New Zealand, Philippines, South Africa, Switzerland, Taiwan, the United States, and Uruguay.

 

In September 2000, up to $50 million worth of food products were recalled due to the presence of Starlink genetically modified corn, which had been approved only for animal consumption and had not been approved for human consumption, and was subsequently withdrawn from the market.

 

ORIGIN

Maize is the domesticated variant of teosinte. The two plants have dissimilar appearance, maize having a single tall stalk with multiple leaves and teosinte being a short, bushy plant. The difference between the two is largely controlled by differences in just two genes, called grassy tillers-1 (gt1, A0A317YEZ1) and teosinte branched-1 (tb1, Q93WI2).

 

Several theories had been proposed about the specific origin of maize in Mesoamerica:

 

It is a direct domestication of a Mexican annual teosinte, Zea mays ssp. parviglumis, native to the Balsas River valley in south-eastern Mexico, with up to 12% of its genetic material obtained from Zea mays ssp. mexicana through introgression.

It has been derived from hybridization between a small domesticated maize (a slightly changed form of a wild maize) and a teosinte of section Luxuriantes, either Z. luxurians or Z. diploperennis.

It has undergone two or more domestications either of a wild maize or of a teosinte. (The term "teosinte" describes all species and subspecies in the genus Zea, excluding Zea mays ssp. mays.)

It has evolved from a hybridization of Z. diploperennis by Tripsacum dactyloides.

 

In the late 1930s, Paul Mangelsdorf suggested that domesticated maize was the result of a hybridization event between an unknown wild maize and a species of Tripsacum, a related genus. This theory about the origin of maize has been refuted by modern genetic testing, which refutes Mangelsdorf's model and the fourth listed above. 

 

The teosinte origin theory was proposed by the Russian botanist Nikolai Ivanovich Vavilov in 1931 and the later American Nobel Prize-winner George Beadle in 1932.: 10  It is supported experimentally and by recent studies of the plants' genomes. Teosinte and maize can cross-breed and produce fertile offspring. A number of questions remain concerning the species, among them:

 

how the immense diversity of the species of sect. Zea originated,

how the tiny archaeological specimens of 3500–2700 BC could have been selected from a teosinte, and

how domestication could have proceeded without leaving remains of teosinte or maize with teosintoid traits earlier than the earliest known until recently, dating from ca. 1100 BC.

 

The domestication of maize is of particular interest to researchers—archaeologists, geneticists, ethnobotanists, geographers, etc. The process is thought by some to have started 7,500 to 12,000 years ago. Research from the 1950s to 1970s originally focused on the hypothesis that maize domestication occurred in the highlands between the states of Oaxaca and Jalisco, because the oldest archaeological remains of maize known at the time were found there.

Connection with 'parviglumis' subspecies

Genetic studies, published in 2004 by John Doebley, identified Zea mays ssp. parviglumis, native to the Balsas River valley in Mexico's southwestern highlands, and also known as Balsas teosinte, as being the crop wild relative that is genetically most similar to modern maize. This was confirmed by further studies, which refined this hypothesis somewhat. Archaeobotanical studies, published in 2009, point to the middle part of the Balsas River valley as the likely location of early domestication; this river is not very long, so these locations are not very distant. Stone milling tools with maize residue have been found in an 8,700 year old layer of deposits in a cave not far from Iguala, Guerrero.

 

Doebley was part of the team that first published, in 2002, that maize had been domesticated only once, about 9,000 years ago, and then spread throughout the Americas.

 

A primitive corn was being grown in southern Mexico, Central America, and northern South America 7,000 years ago. Archaeological remains of early maize ears, found at Guila Naquitz Cave in the Oaxaca Valley, date back roughly 6,250 years; the oldest ears from caves near Tehuacan, Puebla, 5,450 B.P.

 

Maize pollen dated to 7,300 B.P. from San Andres, Tabasco, on the Caribbean coast has also been recovered.

 

As maize was introduced to new cultures, new uses were developed and new varieties selected to better serve in those preparations. Maize was the staple food, or a major staple – along with squash, Andean region potato, quinoa, beans, and amaranth – of most pre-Columbian North American, Mesoamerican, South American, and Caribbean cultures. The Mesoamerican civilization, in particular, was deeply interrelated with maize. Its traditions and rituals involved all aspects of maize cultivation – from the planting to the food preparation. Maize formed the Mesoamerican people's identity.

 

It is unknown what precipitated its domestication, because the edible portion of the wild variety is too small, and hard to obtain, to be eaten directly, as each kernel is enclosed in a very hard bivalve shell.

 

In 1939, George Beadle demonstrated that the kernels of teosinte are readily "popped" for human consumption, like modern popcorn. Some have argued it would have taken too many generations of selective breeding to produce large, compressed ears for efficient cultivation. However, studies of the hybrids readily made by intercrossing teosinte and modern maize suggest this objection is not well founded.

 

SPREADING TO THE NORTH

Around 4,500 ago, maize began to spread to the north; it was first cultivated in what is now the United States at several sites in New Mexico and Arizona, about 4,100 ago.

 

During the first millennium AD, maize cultivation spread more widely in the areas north. In particular, the large-scale adoption of maize agriculture and consumption in eastern North America took place about A.D. 900. Native Americans cleared large forest and grassland areas for the new crop.

 

In 2005, research by the USDA Forest Service suggested that the rise in maize cultivation 500 to 1,000 years ago in what is now the southeastern United States corresponded with a decline of freshwater mussels, which are very sensitive to environmental changes.

 

CULTIVATION

PLANTING

Because it is cold-intolerant, in the temperate zones maize must be planted in the spring. Its root system is generally shallow, so the plant is dependent on soil moisture. As a plant that uses C4 carbon fixation, maize is a considerably more water-efficient crop than plants that use C3 carbon fixation such as alfalfa and soybeans. Maize is most sensitive to drought at the time of silk emergence, when the flowers are ready for pollination. In the United States, a good harvest was traditionally predicted if the maize was "knee-high by the Fourth of July", although modern hybrids generally exceed this growth rate. Maize used for silage is harvested while the plant is green and the fruit immature. Sweet corn is harvested in the "milk stage", after pollination but before starch has formed, between late summer and early to mid-autumn. Field maize is left in the field until very late in the autumn to thoroughly dry the grain, and may, in fact, sometimes not be harvested until winter or even early spring. The importance of sufficient soil moisture is shown in many parts of Africa, where periodic drought regularly causes maize crop failure and consequent famine. Although it is grown mainly in wet, hot climates, it has been said to thrive in cold, hot, dry or wet conditions, meaning that it is an extremely versatile crop.

 

Maize was planted by the Native Americans in hills, in a complex system known to some as the Three Sisters. Maize provided support for beans, and the beans provided nitrogen derived from nitrogen-fixing rhizobia bacteria which live on the roots of beans and other legumes; and squashes provided ground cover to stop weeds and inhibit evaporation by providing shade over the soil. This method was replaced by single species hill planting where each hill 60–120 cm (2 ft 0 in–3 ft 11 in) apart was planted with three or four seeds, a method still used by home gardeners. A later technique was "checked maize", where hills were placed

 

1 m (40 in) apart in each direction, allowing cultivators to run through the field in two directions. In more arid lands, this was altered and seeds were planted in the bottom of 10–12 cm (4–4+1⁄2 in) deep furrows to collect water. Modern technique plants maize in rows which allows for cultivation while the plant is young, although the hill technique is still used in the maize fields of some Native American reservations. When maize is planted in rows, it also allows for planting of other crops between these rows to make more efficient use of land space.

 

In most regions today, maize grown in residential gardens is still often planted manually with a hoe, whereas maize grown commercially is no longer planted manually but rather is planted with a planter. In North America, fields are often planted in a two-crop rotation with a nitrogen-fixing crop, often alfalfa in cooler climates and soybeans in regions with longer summers. Sometimes a third crop, winter wheat, is added to the rotation.

 

Many of the maize varieties grown in the United States and Canada are hybrids. Often the varieties have been genetically modified to tolerate glyphosate or to provide protection against natural pests. Glyphosate is an herbicide which kills all plants except those with genetic tolerance. This genetic tolerance is very rarely found in nature.

 

In the midwestern United States, low-till or no-till farming techniques are usually used. In low-till, fields are covered once, maybe twice, with a tillage implement either ahead of crop planting or after the previous harvest. The fields are planted and fertilized. Weeds are controlled through the use of herbicides, and no cultivation tillage is done during the growing season. This technique reduces moisture evaporation from the soil, and thus provides more moisture for the crop. The technologies mentioned in the previous paragraph enable low-till and no-till farming. Weeds compete with the crop for moisture and nutrients, making them undesirable.

 

HARVESTING

Before the 20th century, all maize harvesting was by manual labour, by grazing, or by some combination of those. Whether the ears were hand-picked and the stover was grazed, or the whole plant was cut, gathered, and shocked, people and livestock did all the work. Between the 1890s and the 1970s, the technology of maize harvesting expanded greatly. Today, all such technologies, from entirely manual harvesting to entirely mechanized, are still in use to some degree, as appropriate to each farm's needs, although the thoroughly mechanized versions predominate, as they offer the lowest unit costs when scaled to large farm operations. For small farms, their unit cost can be too high, as their higher fixed cost cannot be amortized over as many units.[citation needed]

 

Before World War II, most maize in North America was harvested by hand. This involved a large number of workers and associated social events (husking or shucking bees). From the 1890s onward, some machinery became available to partially mechanize the processes, such as one- and two-row mechanical pickers (picking the ear, leaving the stover) and corn binders, which are reaper-binders designed specifically for maize (for example, Video on YouTube). The latter produce sheaves that can be shocked. By hand or mechanical picker, the entire ear is harvested, which then requires a separate operation of a maize sheller to remove the kernels from the ear. Whole ears of maize were often stored in corn cribs, and these whole ears are a sufficient form for some livestock feeding use. Today corn cribs with whole ears, and corn binders, are less common because most modern farms harvest the grain from the field with a combine and store it in bins. The combine with a corn head (with points and snap rolls instead of a reel) does not cut the stalk; it simply pulls the stalk down. The stalk continues downward and is crumpled into a mangled pile on the ground, where it usually is left to become organic matter for the soil. The ear of maize is too large to pass between slots in a plate as the snap rolls pull the stalk away, leaving only the ear and husk to enter the machinery. The combine separates the husk and the cob, keeping only the kernels.

When maize is a silage crop, the entire plant is usually chopped at once with a forage harvester (chopper) and ensiled in silos or polymer wrappers. Ensiling of sheaves cut by a corn binder was formerly common in some regions but has become uncommon. For storing grain in bins, the moisture of the grain must be sufficiently low to avoid spoiling. If the moisture content of the harvested grain is too high, grain dryers are used to reduce the moisture content by blowing heated air through the grain. This can require large amounts of energy in the form of combustible gases (propane or natural gas) and electricity to power the blowers.

 

PRODUCTION

Maize is widely cultivated throughout the world, and a greater weight of maize is produced each year than any other grain. In 2018, total world production was 1.15 billion tonnes, led by the United States with 34.2% of the total (table). China produced 22.4% of the global total.

 

UNITED STATES

In 2016, maize production was forecast to be over 380 million metric tons (15 billion bushels), an increase of 11% over 2014 American production. Based on conditions as of August 2016, the expected yield would be the highest ever for the United States. The area of harvested maize was forecast to be 35 million hectares (87 million acres), an increase of 7% over 2015. Maize is especially popular in Midwestern states such as Indiana, Iowa, and Illinois; in the latter, it was named the state's official grain in 2017.

 

STORAGE

Drying is vital to prevent or at least reduce mycotoxin contamination. Aspergillus and Fusarium spp. are the most common mycotoxin sources, but there are others. Altogether maize contaminants are so common, and this crop is so economically important, that maize mycotoxins are among the most important in agriculture in general.

 

USES

HUMAN FOOD

Maize and cornmeal (ground dried maize) constitute a staple food in many regions of the world. Maize is used to produce cornstarch, a common ingredient in home cooking and many industrialized food products. Maize starch can be hydrolyzed and enzymatically treated to produce syrups, particularly high fructose corn syrup, a sweetener; and also fermented and distilled to produce grain alcohol. Grain alcohol from maize is traditionally the source of Bourbon whiskey. Corn flour is used to make cornbread and other baked products.

 

In prehistoric times Mesoamerican women used a metate to process maize into ground cornmeal, allowing the preparation of foods that were more calorie dense than popcorn. After ceramic vessels were invented the Olmec people began to cook maize together with beans, improving the nutritional value of the staple meal. Although maize naturally contains niacin, an important nutrient, it was not bioavailable without the process of nixtamalization. The Maya used nixtamal meal to make varieties of porridges and tamales. The process was later used in the cuisine of the American South to prepare corn for grits and hominy.

 

Maize is a staple of Mexican cuisine. Masa (cornmeal treated with limewater) is the main ingredient for tortillas, atole and many other dishes of Central American food. It is the main ingredient of corn tortilla, tamales, pozole, atole and all the dishes based on them, like tacos, quesadillas, chilaquiles, enchiladas, tostadas and many more. In Mexico the fungus of maize, known as huitlacoche, is considered a delicacy.

 

Coarse maize meal is made into a thick porridge in many cultures: from the polenta of Italy, the angu of Brazil, the mămăligă of Romania, to cornmeal mush in the US (or hominy grits in the South) or the food called mieliepap in South Africa and sadza, nshima, ugali and other names in other parts of Africa. Introduced into Africa by the Portuguese in the 16th century, maize has become Africa's most important staple food crop. These are commonly eaten in the Southeastern United States, foods handed down from Native Americans, who called the dish sagamite.

 

Maize can also be harvested and consumed in the unripe state, when the kernels are fully grown but still soft. Unripe maize must usually be cooked to become palatable; this may be done by simply boiling or roasting the whole ears and eating the kernels right off the cob. Sweet corn, a genetic variety that is high in sugars and low in starch, is usually consumed in the unripe state. Such corn on the cob is a common dish in the United States, Canada, United Kingdom, Cyprus, some parts of South America, and the Balkans, but virtually unheard of in some European countries. Corn on the cob was hawked on the streets of early 19th-century New York City by poor, barefoot "Hot Corn Girls", who were thus the precursors of hot dog carts, churro wagons, and fruit stands seen on the streets of big cities today.

 

Within the United States, the usage of maize for human consumption constitutes only around 1/40th of the amount grown in the country. In the United States and Canada, maize is mostly grown to feed livestock, as forage, silage (made by fermentation of chopped green cornstalks), or grain. Maize meal is also a significant ingredient of some commercial animal food products.

 

NUTRITIONAL VALUE

Raw, yellow, sweet maize kernels are composed of 76% water, 19% carbohydrates, 3% protein, and 1% fat (table). In a 100-gram serving, maize kernels provide 86 calories and are a good source (10–19% of the Daily Value) of the B vitamins, thiamin, niacin (but see Pellagra warning below), pantothenic acid (B5) and folate (right table for raw, uncooked kernels, USDA Nutrient Database). In moderate amounts, they also supply dietary fiber and the essential minerals, magnesium and phosphorus whereas other nutrients are in low amounts (table).

 

Maize has suboptimal amounts of the essential amino acids tryptophan and lysine, which accounts for its lower status as a protein source. However, the proteins of beans and legumes complement those of maize.

 

FEED AND FODDER FOR LIVESTOCK

Maize is a major source of both grain feed and fodder for livestock. It is fed to the livestock in various ways. When it is used as a grain crop, the dried kernels are used as feed. They are often kept on the cob for storage in a corn crib, or they may be shelled off for storage in a grain bin. The farm that consumes the feed may produce it, purchase it on the market, or some of both. When the grain is used for feed, the rest of the plant (the corn stover) can be used later as fodder, bedding (litter), or soil amendment. When the whole maize plant (grain plus stalks and leaves) is used for fodder, it is usually chopped all at once and ensilaged, as digestibility and palatability are higher in the ensilaged form than in the dried form. Maize silage is one of the most valuable forages for ruminants. Before the advent of widespread ensilaging, it was traditional to gather the corn into shocks after harvesting, where it dried further. With or without a subsequent move to the cover of a barn, it was then stored for weeks to several months until fed to the livestock. Today ensilaging can occur not only in siloes but also in silage wrappers. However, in the tropics, maize can be harvested year-round and fed as green forage to the animals.

 

CHEMICALS

Starch from maize can also be made into plastics, fabrics, adhesives, and many other chemical products.

 

The corn steep liquor, a plentiful watery byproduct of maize wet milling process, is widely used in the biochemical industry and research as a culture medium to grow many kinds of microorganisms.

 

Chrysanthemin is found in purple corn and is used as a food coloring.

 

BIO-FUEL

"Feed maize" is being used increasingly for heating; specialized corn stoves (similar to wood stoves) are available and use either feed maize or wood pellets to generate heat. Maize cobs are also used as a biomass fuel source. Maize is relatively cheap and home-heating furnaces have been developed which use maize kernels as a fuel. They feature a large hopper that feeds the uniformly sized maize kernels (or wood pellets or cherry pits) into the fire.[citation needed]

 

Maize is increasingly used as a feedstock for the production of ethanol fuel. When considering where to construct an ethanol plant, one of the site selection criteria is to ensure there is locally available feedstock. Ethanol is mixed with gasoline to decrease the amount of pollutants emitted when used to fuel motor vehicles. High fuel prices in mid-2007 led to higher demand for ethanol, which in turn led to higher prices paid to farmers for maize. This led to the 2007 harvest being one of the most profitable maize crops in modern history for farmers. Because of the relationship between fuel and maize, prices paid for the crop now tend to track the price of oil.

 

The price of food is affected to a certain degree by the use of maize for biofuel production. The cost of transportation, production, and marketing are a large portion (80%) of the price of food in the United States. Higher energy costs affect these costs, especially transportation. The increase in food prices the consumer has been seeing is mainly due to the higher energy cost. The effect of biofuel production on other food crop prices is indirect. Use of maize for biofuel production increases the demand, and therefore price of maize. This, in turn, results in farm acreage being diverted from other food crops to maize production. This reduces the supply of the other food crops and increases their prices.

 

Maize is widely used in Germany as a feedstock for biogas plants. Here the maize is harvested, shredded then placed in silage clamps from which it is fed into the biogas plants. This process makes use of the whole plant rather than simply using the kernels as in the production of fuel ethanol.

 

A biomass gasification power plant in Strem near Güssing, Burgenland, Austria, began in 2005. Research is being done to make diesel out of the biogas by the Fischer Tropsch method.

 

Increasingly, ethanol is being used at low concentrations (10% or less) as an additive in gasoline (gasohol) for motor fuels to increase the octane rating, lower pollutants, and reduce petroleum use (what is nowadays also known as "biofuels" and has been generating an intense debate regarding the human beings' necessity of new sources of energy, on the one hand, and the need to maintain, in regions such as Latin America, the food habits and culture which has been the essence of civilizations such as the one originated in Mesoamerica; the entry, January 2008, of maize among the commercial agreements of NAFTA has increased this debate, considering the bad labor conditions of workers in the fields, and mainly the fact that NAFTA "opened the doors to the import of maize from the United States, where the farmers who grow it receive multimillion-dollar subsidies and other government supports. ... According to OXFAM UK, after NAFTA went into effect, the price of maize in Mexico fell 70% between 1994 and 2001. The number of farm jobs dropped as well: from 8.1 million in 1993 to 6.8 million in 2002. Many of those who found themselves without work were small-scale maize growers."). However, introduction in the northern latitudes of the US of tropical maize for biofuels, and not for human or animal consumption, may potentially alleviate this.

 

COMMODITY

Maize is bought and sold by investors and price speculators as a tradable commodity using corn futures contracts. These "futures" are traded on the Chicago Board of Trade (CBOT) under ticker symbol C. They are delivered every year in March, May, July, September, and December.

 

ORNAMENTAL AND OTHER USES

Some forms of the plant are occasionally grown for ornamental use in the garden. For this purpose, variegated and colored leaf forms as well as those with colorful ears are used.

 

Corncobs can be hollowed out and treated to make inexpensive smoking pipes, first manufactured in the United States in 1869.

 

An unusual use for maize is to create a "corn maze" (or "maize maze") as a tourist attraction. The idea of a maize maze was introduced by the American Maze Company who created a maze in Pennsylvania in 1993. Traditional mazes are most commonly grown using yew hedges, but these take several years to mature. The rapid growth of a field of maize allows a maze to be laid out using GPS at the start of a growing season and for the maize to grow tall enough to obstruct a visitor's line of sight by the start of the summer. In Canada and the US, these are popular in many farming communities.

 

Maize kernels can be used in place of sand in a sandboxlike enclosure for children's play.

 

Stigmas from female maize flowers, popularly called corn silk, are sold as herbal supplements.

 

Maize is used as a fish bait, called "dough balls". It is particularly popular in Europe for coarse fishing.

 

Additionally, feed corn is sometimes used by hunters to bait animals such as deer or wild hogs.

 

UNITED STATES USAGE BREAKDOWN

The breakdown of usage of the 12.1-billion-bushel (307-million-tonne) 2008 US maize crop was as follows, according to the World Agricultural Supply and Demand Estimates Report by the USDA.In the US since 2009/2010, maize feedstock use for ethanol production has somewhat exceeded direct use for livestock feed; maize use for fuel ethanol was 5,130 million bushels (130 million tonnes) in the 2013/2014 marketing year.A fraction of the maize feedstock dry matter used for ethanol production is usefully recovered as DDGS (dried distillers grains with solubles). In the 2010/2011 marketing year, about 29.1 million tonnes of DDGS were fed to US livestock and poultry. Because starch utilization in fermentation for ethanol production leaves other grain constituents more concentrated in the residue, the feed value per kg of DDGS, with regard to ruminant-metabolizable energy and protein, exceeds that of the grain. Feed value for monogastric animals, such as swine and poultry, is somewhat lower than for ruminants.

 

HAZARDS

PELLAGRA

When maize was first introduced into farming systems other than those used by traditional native-American peoples, it was generally welcomed with enthusiasm for its productivity. However, a widespread problem of malnutrition soon arose wherever maize was introduced as a staple food. This was a mystery, since these types of malnutrition were not normally seen among the indigenous Americans, for whom maize was the principal staple food.

 

It was eventually discovered that the indigenous Americans had learned to soak maize in alkali — water (the process now known as nixtamalization) — made with ashes and lime (calcium oxide) since at least 1200–1500 BC by Mesoamericans. They did this to liberate the corn hulls, but (unbeknownst to natives or colonists) it coincidentally liberates the B-vitamin niacin, the lack of which was the underlying cause of the condition known as pellagra.

 

Maize was introduced into the diet of non-indigenous Americans without the necessary cultural knowledge acquired over thousands of years in the Americas. In the late 19th century, pellagra reached epidemic proportions in parts of the southern US, as medical researchers debated two theories for its origin: the deficiency theory (which was eventually shown to be true) said that pellagra was due to a deficiency of some nutrient, and the germ theory said that pellagra was caused by a germ transmitted by stable flies. A third theory, promoted by the eugenicist Charles Davenport, held that people only contracted pellagra if they were susceptible to it due to certain "constitutional, inheritable" traits of the affected individual.

 

Once alkali processing and dietary variety were understood and applied, pellagra disappeared in the developed world. The development of high lysine maize and the promotion of a more balanced diet have also contributed to its demise. Pellagra still exists today in food-poor areas and refugee camps where people survive on donated maize.

 

ALLERGY

Maize contains lipid transfer protein, an indigestible protein that survives cooking. This protein has been linked to a rare and understudied allergy to maize in humans. The allergic reaction can cause skin rash, swelling or itching of mucous membranes, diarrhea, vomiting, asthma and, in severe cases, anaphylaxis. It is unclear how common this allergy is in the general population.

 

MYCOTOXINS

Fungicide application does not reduce fungal growth or mycotoxin dramatically, although it can be a part of a successful reduction strategy. Among the most common toxins are those produced by Aspergillus and Fusarium spp. The most common toxins are aflatoxins, fumonisins, zearalenone, and ochratoxin A. Bt maize discourages insect vectors and by so doing it dramatically reduces concentrations of fumonisins, significantly reduces aflatoxins, but only mildly reduces others.

 

ART

Maize has been an essential crop in the Andes since the pre-Columbian era. The Moche culture from Northern Peru made ceramics from earth, water, and fire. This pottery was a sacred substance, formed in significant shapes and used to represent important themes. Maize was represented anthropomorphically as well as naturally.

 

In the United States, maize ears along with tobacco leaves are carved into the capitals of columns in the United States Capitol building. Maize itself is sometimes used for temporary architectural detailing when the intent is to celebrate the fall season, local agricultural productivity and culture. Bundles of dried maize stalks are often displayed along with pumpkins, gourds and straw in autumnal displays outside homes and businesses. A well-known example of architectural use is the Corn Palace in Mitchell, South Dakota, which uses cobs and ears of colored maize to implement a mural design that is recycled annually. Another well-known example is the Field of Corn sculpture in Dublin, Ohio, where hundreds of concrete ears of corn stand in a grassy field.

 

A maize stalk with two ripe ears is depicted on the reverse of the Croatian 1 lipa coin, minted since 1993.

 

WIKIPEDIA

Maize (/meɪz/ MAYZ; Zea mays subsp. mays, from Spanish: maíz after Taino: mahiz), also known as corn (North American and Australian English), is a cereal grain first domesticated by indigenous peoples in southern Mexico about 10,000 years ago. The leafy stalk of the plant produces pollen inflorescences and separate ovuliferous inflorescences called ears that yield kernels or seeds, which are fruits.

 

Maize has become a staple food in many parts of the world, with the total production of maize surpassing that of wheat or rice. In addition to being consumed directly by humans (often in the form of masa), maize is also used for corn ethanol, animal feed and other maize products, such as corn starch and corn syrup. The six major types of maize are dent corn, flint corn, pod corn, popcorn, flour corn, and sweet corn. Sugar-rich varieties called sweet corn are usually grown for human consumption as kernels, while field corn varieties are used for animal feed, various corn-based human food uses (including grinding into cornmeal or masa, pressing into corn oil, and fermentation and distillation into alcoholic beverages like bourbon whiskey), and as chemical feedstocks. Maize is also used in making ethanol and other biofuels.

 

Maize is widely cultivated throughout the world, and a greater weight of maize is produced each year than any other grain. In 2014, total world production was 1.04 billion tonnes. Maize is the most widely grown grain crop throughout the Americas, with 361 million metric tons grown in the United States alone in 2014. Genetically modified maize made up 85% of the maize planted in the United States in 2009. Subsidies in the United States help to account for its high level of cultivation of maize and its position as the largest producer in the world.

 

HISTORY

PRE-COLUMBIAN DEVELOPMENT

Maize is a cultigen; human intervention is required for it to propagate. Whether or not the kernels fall off the cob on their own is a key piece of evidence used in archaeology to distinguish domesticated maize from its naturally-propagating teosinte ancestor. Genetic evidence can also be used to determine when various lineages split.

 

Most historians believe maize was domesticated in the Tehuacán Valley of Mexico. Recent research in the early 21st century has modified this view somewhat; scholars now indicate the adjacent Balsas River Valley of south-central Mexico as the center of domestication.

 

An influential 2002 study by Matsuoka et al. has demonstrated that, rather than the multiple independent domestications model, all maize arose from a single domestication in southern Mexico about 9,000 years ago. The study also demonstrated that the oldest surviving maize types are those of the Mexican highlands. Later, maize spread from this region over the Americas along two major paths. This is consistent with a model based on the archaeological record suggesting that maize diversified in the highlands of Mexico before spreading to the lowlands.

 

Archaeologist Dolores Piperno has said:

 

A large corpus of data indicates that [maize] was dispersed into lower Central America by 7600 BP [5600 BC] and had moved into the inter-Andean valleys of Colombia between 7000 and 6000 BP [5000–4000 BC].

— Dolores Piperno, The Origins of Plant Cultivation and Domestication in the New World Tropics: Patterns, Process, and New Developments

 

Since then, even earlier dates have been published.

 

According to a genetic study by Embrapa, corn cultivation was introduced in South America from Mexico, in two great waves: the first, more than 6000 years ago, spread through the Andes. Evidence of cultivation in Peru has been found dating to about 6700 years ago. The second wave, about 2000 years ago, through the lowlands of South America.

 

The earliest maize plants grew only small, 25-millimetre-long (1 in) corn cobs, and only one per plant. In Jackson Spielvogel's view, many centuries of artificial selection (rather than the current view that maize was exploited by interplanting with teosinte) by the indigenous people of the Americas resulted in the development of maize plants capable of growing several cobs per plant, which were usually several centimetres/inches long each. The Olmec and Maya cultivated maize in numerous varieties throughout Mesoamerica; they cooked, ground and processed it through nixtamalization. It was believed that beginning about 2500 BC, the crop spread through much of the Americas. Research of the 21st century has established even earlier dates. The region developed a trade network based on surplus and varieties of maize crops.

 

Mapuches of south-central Chile cultivated maize along with quinoa and potatoes in pre-Hispanic times; however, potato was the staple food of most Mapuches, "specially in the southern and coastal [Mapuche] territories where maize did not reach maturity". Before the expansion of the Inca Empire maize was traded and transported as far south as 40°19' S in Melinquina, Lácar Department. In that location maize remains were found inside pottery dated to 730 ± 80 BP and 920 ± 60 BP. Probably this maize was brought across the Andes from Chile. The presence of maize in Guaitecas Archipelago (43°55' S), the southernmost outpost of pre-Hispanic agriculture, is reported by early Spanish explorers. However the Spanish may have misidentified the plant.

 

COLUMBIAN EXCHANGE

After the arrival of Europeans in 1492, Spanish settlers consumed maize, and explorers and traders carried it back to Europe and introduced it to other countries. Spanish settlers far preferred wheat bread to maize, cassava, or potatoes. Maize flour could not be substituted for wheat for communion bread, since in Christian belief only wheat could undergo transubstantiation and be transformed into the body of Christ. Some Spaniards worried that by eating indigenous foods, which they did not consider nutritious, they would weaken and risk turning into Indians. "In the view of Europeans, it was the food they ate, even more than the environment in which they lived, that gave Amerindians and Spaniards both their distinctive physical characteristics and their characteristic personalities." Despite these worries, Spaniards did consume maize. Archeological evidence from Florida sites indicate they cultivated it as well.

 

Maize spread to the rest of the world because of its ability to grow in diverse climates. It was cultivated in Spain just a few decades after Columbus's voyages and then spread to Italy, West Africa and elsewhere. Widespread cultivation most likely began in southern Spain in 1525, after which it quickly spread to the rest of the Spanish Empire including its territories in Italy (and, from there, to other Italian states). Maize had many advantages over wheat and barley; it yielded two and a half times the food energy per unit cultivated area, could be harvested in successive years from the same plot of land, and grew in wildly varying altitudes and climates, from relatively dry regions with only 250 mm (10 in) of annual rainfall to damp regions with over 5,000 mm (200 in). By the 17th century it was a common peasant food in Southwestern Europe, including Portugal, Spain, southern France, and Italy. By the 18th century, it was the chief food of the southern French and Italian peasantry, especially in the form of polenta in Italy.

Names

 

The word maize derives from the Spanish form of the indigenous Taíno word for the plant, mahiz. It is known by other names around the world.

 

The word "corn" outside the US, Canada, Australia, and New Zealand refers to any cereal crop, its meaning understood to vary geographically to refer to the local staple. In the United States, Canada, Australia, and New Zealand, corn primarily means maize; this usage started as a shortening of "Indian corn". "Indian corn" primarily means maize (the staple grain of indigenous Americans), but can refer more specifically to multicolored "flint corn" used for decoration.

 

In places outside the US, Canada, Australia, and New Zealand, corn often refers to maize in culinary contexts. The narrower meaning is usually indicated by some additional word, as in sweet corn, sweetcorn, corn on the cob, baby corn, the puffed confection known as popcorn and the breakfast cereal known as corn flakes.

 

In Southern Africa, maize is commonly called mielie (Afrikaans) or mealie (English), words derived from the Portuguese word for maize, milho.

 

Maize is preferred in formal, scientific, and international usage because it refers specifically to this one grain, unlike corn, which has a complex variety of meanings that vary by context and geographic region. Maize is used by agricultural bodies and research institutes such as the FAO and CSIRO. National agricultural and industry associations often include the word maize in their name even in English-speaking countries where the local, informal word is something other than maize; for example, the Maize Association of Australia, the Indian Maize Development Association, the Kenya Maize Consortium and Maize Breeders Network, the National Maize Association of Nigeria, the Zimbabwe Seed Maize Association.

 

STRUCTURE AND PHYSIOLOGY

The maize plant is often 3 m (10 ft) in height, though some natural strains can grow 13 m (43 ft). The stem is commonly composed of 20 internodes of 18 cm (7 in) length. The leaves arise from the nodes, alternately on opposite sides on the stalk. A leaf, which grows from each node, is generally 9 cm (3+1⁄2 in) in width and 120 cm (3 ft 11 in) in length.

 

Ears develop above a few of the leaves in the midsection of the plant, between the stem and leaf sheath, elongating by around 3 mm (1⁄8 in) per day, to a length of 18 cm (7 in) with 60 cm (24 in) being the maximum alleged in the subspecies. They are female inflorescences, tightly enveloped by several layers of ear leaves commonly called husks. Certain varieties of maize have been bred to produce many additional developed ears. These are the source of the "baby corn" used as a vegetable in Asian cuisine.

 

The apex of the stem ends in the tassel, an inflorescence of male flowers. When the tassel is mature and conditions are suitably warm and dry, anthers on the tassel dehisce and release pollen. Maize pollen is anemophilous (dispersed by wind), and because of its large settling velocity, most pollen falls within a few meters of the tassel.

 

Elongated stigmas, called silks, emerge from the whorl of husk leaves at the end of the ear. They are often pale yellow and 18 cm (7 in) in length, like tufts of hair in appearance. At the end of each is a carpel, which may develop into a "kernel" if fertilized by a pollen grain. The pericarp of the fruit is fused with the seed coat referred to as "caryopsis", typical of the grasses, and the entire kernel is often referred to as the "seed". The cob is close to a multiple fruit in structure, except that the individual fruits (the kernels) never fuse into a single mass. The grains are about the size of peas, and adhere in regular rows around a white, pithy substance, which forms the ear. The maximum size of kernels is reputedly 2.5 cm (1 in). An ear commonly holds 600 kernels. They are of various colors: blackish, bluish-gray, purple, green, red, white and yellow. When ground into flour, maize yields more flour with much less bran than wheat does. It lacks the protein gluten of wheat and, therefore, makes baked goods with poor rising capability. A genetic variant that accumulates more sugar and less starch in the ear is consumed as a vegetable and is called sweet corn. Young ears can be consumed raw, with the cob and silk, but as the plant matures (usually during the summer months), the cob becomes tougher and the silk dries to inedibility. By the end of the growing season, the kernels dry out and become difficult to chew without cooking them tender first in boiling water.

 

Planting density affects multiple aspects of maize. Modern farming techniques in developed countries usually rely on dense planting, which produces one ear per stalk. Stands of silage maize are yet denser,[citation needed] and achieve a lower percentage of ears and more plant matter.

 

Maize is a facultative short-day plant and flowers in a certain number of growing degree days > 10 °C (50 °F) in the environment to which it is adapted. The magnitude of the influence that long nights have on the number of days that must pass before maize flowers is genetically prescribed and regulated by the phytochrome system.

Photoperiodicity can be eccentric in tropical cultivars such that the long days characteristic of higher latitudes allow the plants to grow so tall that they do not have enough time to produce seed before being killed by frost. These attributes, however, may prove useful in using tropical maize for biofuels.

 

Immature maize shoots accumulate a powerful antibiotic substance, 2,4-dihydroxy-7-methoxy-1,4-benzoxazin-3-one (DIMBOA). DIMBOA is a member of a group of hydroxamic acids (also known as benzoxazinoids) that serve as a natural defense against a wide range of pests, including insects, pathogenic fungi and bacteria. DIMBOA is also found in related grasses, particularly wheat. A maize mutant (bx) lacking DIMBOA is highly susceptible to attack by aphids and fungi. DIMBOA is also responsible for the relative resistance of immature maize to the European corn borer (family Crambidae). As maize matures, DIMBOA levels and resistance to the corn borer decline.

 

Because of its shallow roots, maize is susceptible to droughts, intolerant of nutrient-deficient soils, and prone to be uprooted by severe winds.

 

While yellow maizes derive their color from lutein and zeaxanthin, in red-colored maizes, the kernel coloration is due to anthocyanins and phlobaphenes. These latter substances are synthesized in the flavonoids synthetic pathway from polymerization of flavan-4-ols by the expression of maize pericarp color1 (p1) gene which encodes an R2R3 myb-like transcriptional activator of the A1 gene encoding for the dihydroflavonol 4-reductase (reducing dihydroflavonols into flavan-4-ols) while another gene (Suppressor of Pericarp Pigmentation 1 or SPP1) acts as a suppressor. The p1 gene encodes an Myb-homologous transcriptional activator of genes required for biosynthesis of red phlobaphene pigments, while the P1-wr allele specifies colorless kernel pericarp and red cobs, and unstable factor for orange1 (Ufo1) modifies P1-wr expression to confer pigmentation in kernel pericarp, as well as vegetative tissues, which normally do not accumulate significant amounts of phlobaphene pigments. The maize P gene encodes a Myb homolog that recognizes the sequence CCT/AACC, in sharp contrast with the C/TAACGG bound by vertebrate Myb proteins.

 

The ear leaf is the leaf most closely associated with a particular developing ear. This leaf and above contribute 70% to 75% to 90% of grain fill. Therefore fungicide application is most important in that region in most disease environments.

 

ABNORMAL FLOWERS

Maize flowers may sometimes exhibit mutations that lead to the formation of female flowers in the tassel. These mutations, ts4 and Ts6, prohibit the development of the stamen while simultaneously promoting pistil development. This may cause inflorescences containing both male and female flowers, or hermaphrodite flowers.

 

GENETICS

Maize is an annual grass in the family Gramineae, which includes such plants as wheat, rye, barley, rice, sorghum, and sugarcane. There are two major species of the genus Zea (out of six total): Zea mays (maize) and Zea diploperennis, which is a perennial type of teosinte. The annual teosinte variety called Zea mays mexicana is the closest botanical relative to maize. It still grows in the wild as an annual in Mexico and Guatemala.

 

Many forms of maize are used for food, sometimes classified as various subspecies related to the amount of starch each has:

 

Flour corn: Zea mays var. amylacea

Popcorn: Zea mays var. everta

Dent corn : Zea mays var. indentata

Flint corn: Zea mays var. indurata

Sweet corn: Zea mays var. saccharata and Zea mays var. rugosa

Waxy corn: Zea mays var. ceratina

Amylomaize: Zea mays

Pod corn: Zea mays var. tunicata Larrañaga ex A. St. Hil.

Striped maize: Zea mays var. japonica

 

This system has been replaced (though not entirely displaced) over the last 60 years by multivariable classifications based on ever more data. Agronomic data were supplemented by botanical traits for a robust initial classification, then genetic, cytological, protein and DNA evidence was added. Now, the categories are forms (little used), races, racial complexes, and recently branches.

 

Maize is a diploid with 20 chromosomes (n=10). The combined length of the chromosomes is 1500 cM. Some of the maize chromosomes have what are known as "chromosomal knobs": highly repetitive heterochromatic domains that stain darkly. Individual knobs are polymorphic among strains of both maize and teosinte.

 

Barbara McClintock used these knob markers to validate her transposon theory of "jumping genes", for which she won the 1983 Nobel Prize in Physiology or Medicine. Maize is still an important model organism for genetics and developmental biology today.

 

The centromeres have two types of structural components, both of which are found only in the centromeres: Large arrays of CentC, a short satellite DNA; and a few of a family of retrotransposons. The B chromosome, unlike the others, contains an additional repeat which extends into neighboring areas of the chromosome. Centromeres can accidentally shrink during division and still function, although it is thought this will fail if it shrinks below a few hundred kilobase. Kinetochores contain RNA originating from centromeres. Centromere regions can become inactive, and can continue in that state if the chromosome still has another active one.

 

The Maize Genetics Cooperation Stock Center, funded by the USDA Agricultural Research Service and located in the Department of Crop Sciences at the University of Illinois at Urbana-Champaign, is a stock center of maize mutants. The total collection has nearly 80,000 samples. The bulk of the collection consists of several hundred named genes, plus additional gene combinations and other heritable variants. There are about 1000 chromosomal aberrations (e.g., translocations and inversions) and stocks with abnormal chromosome numbers (e.g., tetraploids). Genetic data describing the maize mutant stocks as well as myriad other data about maize genetics can be accessed at MaizeGDB, the Maize Genetics and Genomics Database.

 

In 2005, the US National Science Foundation (NSF), Department of Agriculture (USDA) and the Department of Energy (DOE) formed a consortium to sequence the B73 maize genome. The resulting DNA sequence data was deposited immediately into GenBank, a public repository for genome-sequence data. Sequences and genome annotations have also been made available throughout the project's lifetime at the project's official site.

 

Primary sequencing of the maize genome was completed in 2008. On November 20, 2009, the consortium published results of its sequencing effort in Science. The genome, 85% of which is composed of transposons, was found to contain 32,540 genes (By comparison, the human genome contains about 2.9 billion bases and 26,000 genes). Much of the maize genome has been duplicated and reshuffled by helitrons—group of rolling circle transposons.

 

In Z. mays and various other angiosperms the MADS-box motif is involved in floral development. Early study in several angiosperm models including Z. mays was the beginning of research into the molecular evolution of floral structure in general, as well as their role in nonflowering plants.

 

EVOLUTION

As with many plants and animals, Z. mays has a positive correlation between effective population size and the magnitude of selection pressure. Z. m. having an EPS of ~650,000, it clusters with others of about the same EPS, and has 79% of its amino acid sites under selection.

 

Recombination is a significant source of diversity in Z. mays. (Note that this finding supersedes previous studies which showed no such correlation.)

 

This recombination/diversity effect is seen throughout plants but is also found to not occur – or not as strongly – in regions of high gene density. This is likely the reason that domesticated Z. mays has not seen as much of an increase in diversity within areas of higher density as in regions of lower density, although there is more evidence in other plants.

 

Some lines of maize have undergone ancient polyploidy events, starting 11m years ago. Over that time ~72% of polyploid duplicated genes have been retained, which is higher than other plants with older polyploidy events. Thus maize may be due to lose more duplicate genes as time goes along, similar to the course followed by the genomes of other plants. If so - if gene loss has merely not occurred yet - that could explain the lack of observed positive selection and lower negative selection which are observed in otherwise similar plants, i.e. also naturally outcrossing and with similar effective population sizes.

 

Ploidy does not appear to influence EPS or magnitude of selection effect in maize.

 

BREEDING

Maize reproduces sexually each year. This randomly selects half the genes from a given plant to propagate to the next generation, meaning that desirable traits found in the crop (like high yield or good nutrition) can be lost in subsequent generations unless certain techniques are used.

 

Maize breeding in prehistory resulted in large plants producing large ears. Modern breeding began with individuals who selected highly productive varieties in their fields and then sold seed to other farmers. James L. Reid was one of the earliest and most successful developing Reid's Yellow Dent in the 1860s. These early efforts were based on mass selection. Later breeding efforts included ear to row selection (C. G. Hopkins c. 1896), hybrids made from selected inbred lines (G. H. Shull, 1909), and the highly successful double cross hybrids using four inbred lines (D. F. Jones c. 1918, 1922). University supported breeding programs were especially important in developing and introducing modern hybrids. By the 1930s, companies such as Pioneer devoted to production of hybrid maize had begun to influence long-term development. Internationally important seed banks such as the International Maize and Wheat Improvement Center (CIMMYT) and the US bank at the Maize Genetics Cooperation Stock Center University of Illinois at Urbana-Champaign maintain germplasm important for future crop development.

 

Since the 1940s the best strains of maize have been first-generation hybrids made from inbred strains that have been optimized for specific traits, such as yield, nutrition, drought, pest and disease tolerance. Both conventional cross-breeding and genetic modification have succeeded in increasing output and reducing the need for cropland, pesticides, water and fertilizer. There is conflicting evidence to support the hypothesis that maize yield potential has increased over the past few decades. This suggests that changes in yield potential are associated with leaf angle, lodging resistance, tolerance of high plant density, disease/pest tolerance, and other agronomic traits rather than increase of yield potential per individual plant.

 

Tropical landraces remain an important and underutilized source of resistance alleles for for disease and for herbivores. Notable discoveries of rare alleles for this purpose were made by Dao et al 2014 and Sood et al 2014.

 

GLOBAL PROGRAM

CIMMYT operates a conventional breeding program to provide optimized strains. The program began in the 1980s. Hybrid seeds are distributed in Africa by the Drought Tolerant Maize for Africa project.

 

GENETIC MODIFICATION

Genetically modified (GM) maize was one of the 26 GM crops grown commercially in 2016. The vast majority of this is Bt maize. Grown since 1997 in the United States and Canada, 92% of the US maize crop was genetically modified in 2016 and 33% of the worldwide maize crop was GM in 2016. As of 2011, Herbicide-tolerant maize varieties were grown in Argentina, Australia, Brazil, Canada, China, Colombia, El Salvador, the European Union, Honduras, Japan, Korea, Malaysia, Mexico, New Zealand, Philippines, the Russian Federation, Singapore, South Africa, Taiwan, Thailand, and the United States. Insect-resistant maize was grown in Argentina, Australia, Brazil, Canada, Chile, China, Colombia, Egypt, the European Union, Honduras, Japan, Korea, Malaysia, Mexico, New Zealand, Philippines, South Africa, Switzerland, Taiwan, the United States, and Uruguay.

 

In September 2000, up to $50 million worth of food products were recalled due to the presence of Starlink genetically modified corn, which had been approved only for animal consumption and had not been approved for human consumption, and was subsequently withdrawn from the market.

 

ORIGIN

Maize is the domesticated variant of teosinte. The two plants have dissimilar appearance, maize having a single tall stalk with multiple leaves and teosinte being a short, bushy plant. The difference between the two is largely controlled by differences in just two genes, called grassy tillers-1 (gt1, A0A317YEZ1) and teosinte branched-1 (tb1, Q93WI2).

 

Several theories had been proposed about the specific origin of maize in Mesoamerica:

 

It is a direct domestication of a Mexican annual teosinte, Zea mays ssp. parviglumis, native to the Balsas River valley in south-eastern Mexico, with up to 12% of its genetic material obtained from Zea mays ssp. mexicana through introgression.

It has been derived from hybridization between a small domesticated maize (a slightly changed form of a wild maize) and a teosinte of section Luxuriantes, either Z. luxurians or Z. diploperennis.

It has undergone two or more domestications either of a wild maize or of a teosinte. (The term "teosinte" describes all species and subspecies in the genus Zea, excluding Zea mays ssp. mays.)

It has evolved from a hybridization of Z. diploperennis by Tripsacum dactyloides.

 

In the late 1930s, Paul Mangelsdorf suggested that domesticated maize was the result of a hybridization event between an unknown wild maize and a species of Tripsacum, a related genus. This theory about the origin of maize has been refuted by modern genetic testing, which refutes Mangelsdorf's model and the fourth listed above. 

 

The teosinte origin theory was proposed by the Russian botanist Nikolai Ivanovich Vavilov in 1931 and the later American Nobel Prize-winner George Beadle in 1932.: 10  It is supported experimentally and by recent studies of the plants' genomes. Teosinte and maize can cross-breed and produce fertile offspring. A number of questions remain concerning the species, among them:

 

how the immense diversity of the species of sect. Zea originated,

how the tiny archaeological specimens of 3500–2700 BC could have been selected from a teosinte, and

how domestication could have proceeded without leaving remains of teosinte or maize with teosintoid traits earlier than the earliest known until recently, dating from ca. 1100 BC.

 

The domestication of maize is of particular interest to researchers—archaeologists, geneticists, ethnobotanists, geographers, etc. The process is thought by some to have started 7,500 to 12,000 years ago. Research from the 1950s to 1970s originally focused on the hypothesis that maize domestication occurred in the highlands between the states of Oaxaca and Jalisco, because the oldest archaeological remains of maize known at the time were found there.

Connection with 'parviglumis' subspecies

Genetic studies, published in 2004 by John Doebley, identified Zea mays ssp. parviglumis, native to the Balsas River valley in Mexico's southwestern highlands, and also known as Balsas teosinte, as being the crop wild relative that is genetically most similar to modern maize. This was confirmed by further studies, which refined this hypothesis somewhat. Archaeobotanical studies, published in 2009, point to the middle part of the Balsas River valley as the likely location of early domestication; this river is not very long, so these locations are not very distant. Stone milling tools with maize residue have been found in an 8,700 year old layer of deposits in a cave not far from Iguala, Guerrero.

 

Doebley was part of the team that first published, in 2002, that maize had been domesticated only once, about 9,000 years ago, and then spread throughout the Americas.

 

A primitive corn was being grown in southern Mexico, Central America, and northern South America 7,000 years ago. Archaeological remains of early maize ears, found at Guila Naquitz Cave in the Oaxaca Valley, date back roughly 6,250 years; the oldest ears from caves near Tehuacan, Puebla, 5,450 B.P.

 

Maize pollen dated to 7,300 B.P. from San Andres, Tabasco, on the Caribbean coast has also been recovered.

 

As maize was introduced to new cultures, new uses were developed and new varieties selected to better serve in those preparations. Maize was the staple food, or a major staple – along with squash, Andean region potato, quinoa, beans, and amaranth – of most pre-Columbian North American, Mesoamerican, South American, and Caribbean cultures. The Mesoamerican civilization, in particular, was deeply interrelated with maize. Its traditions and rituals involved all aspects of maize cultivation – from the planting to the food preparation. Maize formed the Mesoamerican people's identity.

 

It is unknown what precipitated its domestication, because the edible portion of the wild variety is too small, and hard to obtain, to be eaten directly, as each kernel is enclosed in a very hard bivalve shell.

 

In 1939, George Beadle demonstrated that the kernels of teosinte are readily "popped" for human consumption, like modern popcorn. Some have argued it would have taken too many generations of selective breeding to produce large, compressed ears for efficient cultivation. However, studies of the hybrids readily made by intercrossing teosinte and modern maize suggest this objection is not well founded.

 

SPREADING TO THE NORTH

Around 4,500 ago, maize began to spread to the north; it was first cultivated in what is now the United States at several sites in New Mexico and Arizona, about 4,100 ago.

 

During the first millennium AD, maize cultivation spread more widely in the areas north. In particular, the large-scale adoption of maize agriculture and consumption in eastern North America took place about A.D. 900. Native Americans cleared large forest and grassland areas for the new crop.

 

In 2005, research by the USDA Forest Service suggested that the rise in maize cultivation 500 to 1,000 years ago in what is now the southeastern United States corresponded with a decline of freshwater mussels, which are very sensitive to environmental changes.

 

CULTIVATION

PLANTING

Because it is cold-intolerant, in the temperate zones maize must be planted in the spring. Its root system is generally shallow, so the plant is dependent on soil moisture. As a plant that uses C4 carbon fixation, maize is a considerably more water-efficient crop than plants that use C3 carbon fixation such as alfalfa and soybeans. Maize is most sensitive to drought at the time of silk emergence, when the flowers are ready for pollination. In the United States, a good harvest was traditionally predicted if the maize was "knee-high by the Fourth of July", although modern hybrids generally exceed this growth rate. Maize used for silage is harvested while the plant is green and the fruit immature. Sweet corn is harvested in the "milk stage", after pollination but before starch has formed, between late summer and early to mid-autumn. Field maize is left in the field until very late in the autumn to thoroughly dry the grain, and may, in fact, sometimes not be harvested until winter or even early spring. The importance of sufficient soil moisture is shown in many parts of Africa, where periodic drought regularly causes maize crop failure and consequent famine. Although it is grown mainly in wet, hot climates, it has been said to thrive in cold, hot, dry or wet conditions, meaning that it is an extremely versatile crop.

 

Maize was planted by the Native Americans in hills, in a complex system known to some as the Three Sisters. Maize provided support for beans, and the beans provided nitrogen derived from nitrogen-fixing rhizobia bacteria which live on the roots of beans and other legumes; and squashes provided ground cover to stop weeds and inhibit evaporation by providing shade over the soil. This method was replaced by single species hill planting where each hill 60–120 cm (2 ft 0 in–3 ft 11 in) apart was planted with three or four seeds, a method still used by home gardeners. A later technique was "checked maize", where hills were placed

 

1 m (40 in) apart in each direction, allowing cultivators to run through the field in two directions. In more arid lands, this was altered and seeds were planted in the bottom of 10–12 cm (4–4+1⁄2 in) deep furrows to collect water. Modern technique plants maize in rows which allows for cultivation while the plant is young, although the hill technique is still used in the maize fields of some Native American reservations. When maize is planted in rows, it also allows for planting of other crops between these rows to make more efficient use of land space.

 

In most regions today, maize grown in residential gardens is still often planted manually with a hoe, whereas maize grown commercially is no longer planted manually but rather is planted with a planter. In North America, fields are often planted in a two-crop rotation with a nitrogen-fixing crop, often alfalfa in cooler climates and soybeans in regions with longer summers. Sometimes a third crop, winter wheat, is added to the rotation.

 

Many of the maize varieties grown in the United States and Canada are hybrids. Often the varieties have been genetically modified to tolerate glyphosate or to provide protection against natural pests. Glyphosate is an herbicide which kills all plants except those with genetic tolerance. This genetic tolerance is very rarely found in nature.

 

In the midwestern United States, low-till or no-till farming techniques are usually used. In low-till, fields are covered once, maybe twice, with a tillage implement either ahead of crop planting or after the previous harvest. The fields are planted and fertilized. Weeds are controlled through the use of herbicides, and no cultivation tillage is done during the growing season. This technique reduces moisture evaporation from the soil, and thus provides more moisture for the crop. The technologies mentioned in the previous paragraph enable low-till and no-till farming. Weeds compete with the crop for moisture and nutrients, making them undesirable.

 

HARVESTING

Before the 20th century, all maize harvesting was by manual labour, by grazing, or by some combination of those. Whether the ears were hand-picked and the stover was grazed, or the whole plant was cut, gathered, and shocked, people and livestock did all the work. Between the 1890s and the 1970s, the technology of maize harvesting expanded greatly. Today, all such technologies, from entirely manual harvesting to entirely mechanized, are still in use to some degree, as appropriate to each farm's needs, although the thoroughly mechanized versions predominate, as they offer the lowest unit costs when scaled to large farm operations. For small farms, their unit cost can be too high, as their higher fixed cost cannot be amortized over as many units.[citation needed]

 

Before World War II, most maize in North America was harvested by hand. This involved a large number of workers and associated social events (husking or shucking bees). From the 1890s onward, some machinery became available to partially mechanize the processes, such as one- and two-row mechanical pickers (picking the ear, leaving the stover) and corn binders, which are reaper-binders designed specifically for maize (for example, Video on YouTube). The latter produce sheaves that can be shocked. By hand or mechanical picker, the entire ear is harvested, which then requires a separate operation of a maize sheller to remove the kernels from the ear. Whole ears of maize were often stored in corn cribs, and these whole ears are a sufficient form for some livestock feeding use. Today corn cribs with whole ears, and corn binders, are less common because most modern farms harvest the grain from the field with a combine and store it in bins. The combine with a corn head (with points and snap rolls instead of a reel) does not cut the stalk; it simply pulls the stalk down. The stalk continues downward and is crumpled into a mangled pile on the ground, where it usually is left to become organic matter for the soil. The ear of maize is too large to pass between slots in a plate as the snap rolls pull the stalk away, leaving only the ear and husk to enter the machinery. The combine separates the husk and the cob, keeping only the kernels.

When maize is a silage crop, the entire plant is usually chopped at once with a forage harvester (chopper) and ensiled in silos or polymer wrappers. Ensiling of sheaves cut by a corn binder was formerly common in some regions but has become uncommon. For storing grain in bins, the moisture of the grain must be sufficiently low to avoid spoiling. If the moisture content of the harvested grain is too high, grain dryers are used to reduce the moisture content by blowing heated air through the grain. This can require large amounts of energy in the form of combustible gases (propane or natural gas) and electricity to power the blowers.

 

PRODUCTION

Maize is widely cultivated throughout the world, and a greater weight of maize is produced each year than any other grain. In 2018, total world production was 1.15 billion tonnes, led by the United States with 34.2% of the total (table). China produced 22.4% of the global total.

 

UNITED STATES

In 2016, maize production was forecast to be over 380 million metric tons (15 billion bushels), an increase of 11% over 2014 American production. Based on conditions as of August 2016, the expected yield would be the highest ever for the United States. The area of harvested maize was forecast to be 35 million hectares (87 million acres), an increase of 7% over 2015. Maize is especially popular in Midwestern states such as Indiana, Iowa, and Illinois; in the latter, it was named the state's official grain in 2017.

 

STORAGE

Drying is vital to prevent or at least reduce mycotoxin contamination. Aspergillus and Fusarium spp. are the most common mycotoxin sources, but there are others. Altogether maize contaminants are so common, and this crop is so economically important, that maize mycotoxins are among the most important in agriculture in general.

 

USES

HUMAN FOOD

Maize and cornmeal (ground dried maize) constitute a staple food in many regions of the world. Maize is used to produce cornstarch, a common ingredient in home cooking and many industrialized food products. Maize starch can be hydrolyzed and enzymatically treated to produce syrups, particularly high fructose corn syrup, a sweetener; and also fermented and distilled to produce grain alcohol. Grain alcohol from maize is traditionally the source of Bourbon whiskey. Corn flour is used to make cornbread and other baked products.

 

In prehistoric times Mesoamerican women used a metate to process maize into ground cornmeal, allowing the preparation of foods that were more calorie dense than popcorn. After ceramic vessels were invented the Olmec people began to cook maize together with beans, improving the nutritional value of the staple meal. Although maize naturally contains niacin, an important nutrient, it was not bioavailable without the process of nixtamalization. The Maya used nixtamal meal to make varieties of porridges and tamales. The process was later used in the cuisine of the American South to prepare corn for grits and hominy.

 

Maize is a staple of Mexican cuisine. Masa (cornmeal treated with limewater) is the main ingredient for tortillas, atole and many other dishes of Central American food. It is the main ingredient of corn tortilla, tamales, pozole, atole and all the dishes based on them, like tacos, quesadillas, chilaquiles, enchiladas, tostadas and many more. In Mexico the fungus of maize, known as huitlacoche, is considered a delicacy.

 

Coarse maize meal is made into a thick porridge in many cultures: from the polenta of Italy, the angu of Brazil, the mămăligă of Romania, to cornmeal mush in the US (or hominy grits in the South) or the food called mieliepap in South Africa and sadza, nshima, ugali and other names in other parts of Africa. Introduced into Africa by the Portuguese in the 16th century, maize has become Africa's most important staple food crop. These are commonly eaten in the Southeastern United States, foods handed down from Native Americans, who called the dish sagamite.

 

Maize can also be harvested and consumed in the unripe state, when the kernels are fully grown but still soft. Unripe maize must usually be cooked to become palatable; this may be done by simply boiling or roasting the whole ears and eating the kernels right off the cob. Sweet corn, a genetic variety that is high in sugars and low in starch, is usually consumed in the unripe state. Such corn on the cob is a common dish in the United States, Canada, United Kingdom, Cyprus, some parts of South America, and the Balkans, but virtually unheard of in some European countries. Corn on the cob was hawked on the streets of early 19th-century New York City by poor, barefoot "Hot Corn Girls", who were thus the precursors of hot dog carts, churro wagons, and fruit stands seen on the streets of big cities today.

 

Within the United States, the usage of maize for human consumption constitutes only around 1/40th of the amount grown in the country. In the United States and Canada, maize is mostly grown to feed livestock, as forage, silage (made by fermentation of chopped green cornstalks), or grain. Maize meal is also a significant ingredient of some commercial animal food products.

 

NUTRITIONAL VALUE

Raw, yellow, sweet maize kernels are composed of 76% water, 19% carbohydrates, 3% protein, and 1% fat (table). In a 100-gram serving, maize kernels provide 86 calories and are a good source (10–19% of the Daily Value) of the B vitamins, thiamin, niacin (but see Pellagra warning below), pantothenic acid (B5) and folate (right table for raw, uncooked kernels, USDA Nutrient Database). In moderate amounts, they also supply dietary fiber and the essential minerals, magnesium and phosphorus whereas other nutrients are in low amounts (table).

 

Maize has suboptimal amounts of the essential amino acids tryptophan and lysine, which accounts for its lower status as a protein source. However, the proteins of beans and legumes complement those of maize.

 

FEED AND FODDER FOR LIVESTOCK

Maize is a major source of both grain feed and fodder for livestock. It is fed to the livestock in various ways. When it is used as a grain crop, the dried kernels are used as feed. They are often kept on the cob for storage in a corn crib, or they may be shelled off for storage in a grain bin. The farm that consumes the feed may produce it, purchase it on the market, or some of both. When the grain is used for feed, the rest of the plant (the corn stover) can be used later as fodder, bedding (litter), or soil amendment. When the whole maize plant (grain plus stalks and leaves) is used for fodder, it is usually chopped all at once and ensilaged, as digestibility and palatability are higher in the ensilaged form than in the dried form. Maize silage is one of the most valuable forages for ruminants. Before the advent of widespread ensilaging, it was traditional to gather the corn into shocks after harvesting, where it dried further. With or without a subsequent move to the cover of a barn, it was then stored for weeks to several months until fed to the livestock. Today ensilaging can occur not only in siloes but also in silage wrappers. However, in the tropics, maize can be harvested year-round and fed as green forage to the animals.

 

CHEMICALS

Starch from maize can also be made into plastics, fabrics, adhesives, and many other chemical products.

 

The corn steep liquor, a plentiful watery byproduct of maize wet milling process, is widely used in the biochemical industry and research as a culture medium to grow many kinds of microorganisms.

 

Chrysanthemin is found in purple corn and is used as a food coloring.

 

BIO-FUEL

"Feed maize" is being used increasingly for heating; specialized corn stoves (similar to wood stoves) are available and use either feed maize or wood pellets to generate heat. Maize cobs are also used as a biomass fuel source. Maize is relatively cheap and home-heating furnaces have been developed which use maize kernels as a fuel. They feature a large hopper that feeds the uniformly sized maize kernels (or wood pellets or cherry pits) into the fire.[citation needed]

 

Maize is increasingly used as a feedstock for the production of ethanol fuel. When considering where to construct an ethanol plant, one of the site selection criteria is to ensure there is locally available feedstock. Ethanol is mixed with gasoline to decrease the amount of pollutants emitted when used to fuel motor vehicles. High fuel prices in mid-2007 led to higher demand for ethanol, which in turn led to higher prices paid to farmers for maize. This led to the 2007 harvest being one of the most profitable maize crops in modern history for farmers. Because of the relationship between fuel and maize, prices paid for the crop now tend to track the price of oil.

 

The price of food is affected to a certain degree by the use of maize for biofuel production. The cost of transportation, production, and marketing are a large portion (80%) of the price of food in the United States. Higher energy costs affect these costs, especially transportation. The increase in food prices the consumer has been seeing is mainly due to the higher energy cost. The effect of biofuel production on other food crop prices is indirect. Use of maize for biofuel production increases the demand, and therefore price of maize. This, in turn, results in farm acreage being diverted from other food crops to maize production. This reduces the supply of the other food crops and increases their prices.

 

Maize is widely used in Germany as a feedstock for biogas plants. Here the maize is harvested, shredded then placed in silage clamps from which it is fed into the biogas plants. This process makes use of the whole plant rather than simply using the kernels as in the production of fuel ethanol.

 

A biomass gasification power plant in Strem near Güssing, Burgenland, Austria, began in 2005. Research is being done to make diesel out of the biogas by the Fischer Tropsch method.

 

Increasingly, ethanol is being used at low concentrations (10% or less) as an additive in gasoline (gasohol) for motor fuels to increase the octane rating, lower pollutants, and reduce petroleum use (what is nowadays also known as "biofuels" and has been generating an intense debate regarding the human beings' necessity of new sources of energy, on the one hand, and the need to maintain, in regions such as Latin America, the food habits and culture which has been the essence of civilizations such as the one originated in Mesoamerica; the entry, January 2008, of maize among the commercial agreements of NAFTA has increased this debate, considering the bad labor conditions of workers in the fields, and mainly the fact that NAFTA "opened the doors to the import of maize from the United States, where the farmers who grow it receive multimillion-dollar subsidies and other government supports. ... According to OXFAM UK, after NAFTA went into effect, the price of maize in Mexico fell 70% between 1994 and 2001. The number of farm jobs dropped as well: from 8.1 million in 1993 to 6.8 million in 2002. Many of those who found themselves without work were small-scale maize growers."). However, introduction in the northern latitudes of the US of tropical maize for biofuels, and not for human or animal consumption, may potentially alleviate this.

 

COMMODITY

Maize is bought and sold by investors and price speculators as a tradable commodity using corn futures contracts. These "futures" are traded on the Chicago Board of Trade (CBOT) under ticker symbol C. They are delivered every year in March, May, July, September, and December.

 

ORNAMENTAL AND OTHER USES

Some forms of the plant are occasionally grown for ornamental use in the garden. For this purpose, variegated and colored leaf forms as well as those with colorful ears are used.

 

Corncobs can be hollowed out and treated to make inexpensive smoking pipes, first manufactured in the United States in 1869.

 

An unusual use for maize is to create a "corn maze" (or "maize maze") as a tourist attraction. The idea of a maize maze was introduced by the American Maze Company who created a maze in Pennsylvania in 1993. Traditional mazes are most commonly grown using yew hedges, but these take several years to mature. The rapid growth of a field of maize allows a maze to be laid out using GPS at the start of a growing season and for the maize to grow tall enough to obstruct a visitor's line of sight by the start of the summer. In Canada and the US, these are popular in many farming communities.

 

Maize kernels can be used in place of sand in a sandboxlike enclosure for children's play.

 

Stigmas from female maize flowers, popularly called corn silk, are sold as herbal supplements.

 

Maize is used as a fish bait, called "dough balls". It is particularly popular in Europe for coarse fishing.

 

Additionally, feed corn is sometimes used by hunters to bait animals such as deer or wild hogs.

 

UNITED STATES USAGE BREAKDOWN

The breakdown of usage of the 12.1-billion-bushel (307-million-tonne) 2008 US maize crop was as follows, according to the World Agricultural Supply and Demand Estimates Report by the USDA.In the US since 2009/2010, maize feedstock use for ethanol production has somewhat exceeded direct use for livestock feed; maize use for fuel ethanol was 5,130 million bushels (130 million tonnes) in the 2013/2014 marketing year.A fraction of the maize feedstock dry matter used for ethanol production is usefully recovered as DDGS (dried distillers grains with solubles). In the 2010/2011 marketing year, about 29.1 million tonnes of DDGS were fed to US livestock and poultry. Because starch utilization in fermentation for ethanol production leaves other grain constituents more concentrated in the residue, the feed value per kg of DDGS, with regard to ruminant-metabolizable energy and protein, exceeds that of the grain. Feed value for monogastric animals, such as swine and poultry, is somewhat lower than for ruminants.

 

HAZARDS

PELLAGRA

When maize was first introduced into farming systems other than those used by traditional native-American peoples, it was generally welcomed with enthusiasm for its productivity. However, a widespread problem of malnutrition soon arose wherever maize was introduced as a staple food. This was a mystery, since these types of malnutrition were not normally seen among the indigenous Americans, for whom maize was the principal staple food.

 

It was eventually discovered that the indigenous Americans had learned to soak maize in alkali — water (the process now known as nixtamalization) — made with ashes and lime (calcium oxide) since at least 1200–1500 BC by Mesoamericans. They did this to liberate the corn hulls, but (unbeknownst to natives or colonists) it coincidentally liberates the B-vitamin niacin, the lack of which was the underlying cause of the condition known as pellagra.

 

Maize was introduced into the diet of non-indigenous Americans without the necessary cultural knowledge acquired over thousands of years in the Americas. In the late 19th century, pellagra reached epidemic proportions in parts of the southern US, as medical researchers debated two theories for its origin: the deficiency theory (which was eventually shown to be true) said that pellagra was due to a deficiency of some nutrient, and the germ theory said that pellagra was caused by a germ transmitted by stable flies. A third theory, promoted by the eugenicist Charles Davenport, held that people only contracted pellagra if they were susceptible to it due to certain "constitutional, inheritable" traits of the affected individual.

 

Once alkali processing and dietary variety were understood and applied, pellagra disappeared in the developed world. The development of high lysine maize and the promotion of a more balanced diet have also contributed to its demise. Pellagra still exists today in food-poor areas and refugee camps where people survive on donated maize.

 

ALLERGY

Maize contains lipid transfer protein, an indigestible protein that survives cooking. This protein has been linked to a rare and understudied allergy to maize in humans. The allergic reaction can cause skin rash, swelling or itching of mucous membranes, diarrhea, vomiting, asthma and, in severe cases, anaphylaxis. It is unclear how common this allergy is in the general population.

 

MYCOTOXINS

Fungicide application does not reduce fungal growth or mycotoxin dramatically, although it can be a part of a successful reduction strategy. Among the most common toxins are those produced by Aspergillus and Fusarium spp. The most common toxins are aflatoxins, fumonisins, zearalenone, and ochratoxin A. Bt maize discourages insect vectors and by so doing it dramatically reduces concentrations of fumonisins, significantly reduces aflatoxins, but only mildly reduces others.

 

ART

Maize has been an essential crop in the Andes since the pre-Columbian era. The Moche culture from Northern Peru made ceramics from earth, water, and fire. This pottery was a sacred substance, formed in significant shapes and used to represent important themes. Maize was represented anthropomorphically as well as naturally.

 

In the United States, maize ears along with tobacco leaves are carved into the capitals of columns in the United States Capitol building. Maize itself is sometimes used for temporary architectural detailing when the intent is to celebrate the fall season, local agricultural productivity and culture. Bundles of dried maize stalks are often displayed along with pumpkins, gourds and straw in autumnal displays outside homes and businesses. A well-known example of architectural use is the Corn Palace in Mitchell, South Dakota, which uses cobs and ears of colored maize to implement a mural design that is recycled annually. Another well-known example is the Field of Corn sculpture in Dublin, Ohio, where hundreds of concrete ears of corn stand in a grassy field.

 

A maize stalk with two ripe ears is depicted on the reverse of the Croatian 1 lipa coin, minted since 1993.

 

WIKIPEDIA

リコリス・ロンギチュバ ‘ピュア・ホワイト’

Lycoris longituba Y.C.Hsu et G.J.Fan, 1974 ‘Pure White’

(Imported at JAPAN, 1979 from China, By Mr. Kaneko, Japan.)

Very rare plants. 2n=16=6M+10T

 

Endemic to China (Jiangsu and Anhui). Very polymorphic species. Variable in flower color and shape. Plants with yellow flowers classified as a variety, "Lycoris longituba Y.C.Hsu et G.J.Fan, 1974". Very long tepatube and its fragrance are marked chracteristics of this species. Leaves lanceolate, to 63 cm long and to 4 cm wide, somewhat fleshy and pale green, appearing in early spring and wither up in May. Scape, 60-80 cm hight, appearing in July to August. Spathe, to 5 cm long. Pedicel 1.5-4.5 cm, tepaltube 4.5-6.0 cm. Tepals 7-9.5 cm long, . Stamens 6-7.5 cm, shorter than tepals. Style 7-9.5 cm, nearly equal to or slightly exceeding tepals. Before that it was far carrying out this plant in a scientific statement, it was indicated by U.S. an Advanced Horticulturalist Mr. Sam Coldwell from that it was a new species. The formal scientific statement of "Lycoris longituba" was announced by Y.C.Hsu & G.J.Fan in 1974. However, this plant was introduced into Japan and was already grown in the 1930s. It by misconception of scholar Dr. Inariyama (1948) of Japan "Lycoris x straminea Lindl., 1848" said. When it depended on research of scholar Dr. Kurita of Japan, it became clear that the individual of this plants is one of the variations of "Lycoris longituba Y.C.Hsu et G.J.Fan, 1974"

 

この植物は、学術記載をされるはるか以前から、アメリカの先進的園芸家・Mr. Sam Coldwell により新種ではないかと指摘をされていた。「Lycoris longituba」の正式な学術記載は Y.C.Hsu & G.J.Fan によって1974年になされた。しかし、既に1930年代にこの植物は日本に渡来して栽培されていた。日本の学者 Dr. Inariyama(1948)の誤認で「Lycoris × straminea Lindl., 1848」と言われていた。日本の学者 Dr. Kurita の研究に依ると、この画像の個体は「Lycoris longituba Y.C.Hsu et G.J.Fan, 1974」のバリエーションの一つだと判明した。

 

この個体はリコリス中でも最も大型の部類で、主に種子繁殖をし、非常に自家受粉しやすいが、その反面、分球はしずらく、1球が10球までになるには凡そ30年はかかった。ヒガンバナが30年で1000球に分けつする事と比較すると恐ろしいほどスローモーな分けつ力である。此の個体は、金子氏が40年前に中国から導入したヒガンバナ類の中から選抜した個体で、他に麦藁色の個体が明治期にもたらされていたが、在来の物より鑑賞価値は高い。春出葉型なので、短い期間しか葉が無い為、その間には良く日光に当て肥培をすることが肝心で有る。自然交雑種のナツズイセンの片親である事がDr. Kurita の研究で判明した。自然界でのリコリス・ロンギチュバは、白、ピンク、麦藁色など幅があり、黄花の物は別途変種扱いされている。此の個体自体は、蘂にやや色が乗るが、これほどに白い個体は少なく、殆どがやや濁っているのが殆どであり、選抜した意味は大きい。本種は大球性で他のリコリスよりもやや深く土中に球根が潜る。開花は、是から得られたナツズイセンよりも1ヶ月早く、7月中には開花する。

  

SONY NEX-7

Nikon Ai AF Micro Nikkor ED 200mm F4D (IF)

(Product Year : 1993)

pOLymORpHic hUMansCApE by Nandita Kumar

 

"The tiny LCD screens in this interactive piece portray scenes of urban life. The screens react to movement, switching from restful scenes of nature to images of uncontrolled urban sprawl. The artist is intrigued by how urban environments shape our consciousness. She highlights India’s mega-metropoles, where uncontrolled urbanization is exceeding sustainable limits, resulting in slums and impacting negatively on our wellbeing."

 

arsguide.kiasma.fi/en/artwork/nandita-kumar/

  

# 5 World Environment Day

117 Pictures in 2017

white-throated sparrow [photo taken thru window]

Maize (/meɪz/ MAYZ; Zea mays subsp. mays, from Spanish: maíz after Taino: mahiz), also known as corn (North American and Australian English), is a cereal grain first domesticated by indigenous peoples in southern Mexico about 10,000 years ago. The leafy stalk of the plant produces pollen inflorescences and separate ovuliferous inflorescences called ears that yield kernels or seeds, which are fruits.

 

Maize has become a staple food in many parts of the world, with the total production of maize surpassing that of wheat or rice. In addition to being consumed directly by humans (often in the form of masa), maize is also used for corn ethanol, animal feed and other maize products, such as corn starch and corn syrup. The six major types of maize are dent corn, flint corn, pod corn, popcorn, flour corn, and sweet corn. Sugar-rich varieties called sweet corn are usually grown for human consumption as kernels, while field corn varieties are used for animal feed, various corn-based human food uses (including grinding into cornmeal or masa, pressing into corn oil, and fermentation and distillation into alcoholic beverages like bourbon whiskey), and as chemical feedstocks. Maize is also used in making ethanol and other biofuels.

 

Maize is widely cultivated throughout the world, and a greater weight of maize is produced each year than any other grain. In 2014, total world production was 1.04 billion tonnes. Maize is the most widely grown grain crop throughout the Americas, with 361 million metric tons grown in the United States alone in 2014. Genetically modified maize made up 85% of the maize planted in the United States in 2009. Subsidies in the United States help to account for its high level of cultivation of maize and its position as the largest producer in the world.

 

HISTORY

PRE-COLUMBIAN DEVELOPMENT

Maize is a cultigen; human intervention is required for it to propagate. Whether or not the kernels fall off the cob on their own is a key piece of evidence used in archaeology to distinguish domesticated maize from its naturally-propagating teosinte ancestor. Genetic evidence can also be used to determine when various lineages split.

 

Most historians believe maize was domesticated in the Tehuacán Valley of Mexico. Recent research in the early 21st century has modified this view somewhat; scholars now indicate the adjacent Balsas River Valley of south-central Mexico as the center of domestication.

 

An influential 2002 study by Matsuoka et al. has demonstrated that, rather than the multiple independent domestications model, all maize arose from a single domestication in southern Mexico about 9,000 years ago. The study also demonstrated that the oldest surviving maize types are those of the Mexican highlands. Later, maize spread from this region over the Americas along two major paths. This is consistent with a model based on the archaeological record suggesting that maize diversified in the highlands of Mexico before spreading to the lowlands.

 

Archaeologist Dolores Piperno has said:

 

A large corpus of data indicates that [maize] was dispersed into lower Central America by 7600 BP [5600 BC] and had moved into the inter-Andean valleys of Colombia between 7000 and 6000 BP [5000–4000 BC].

— Dolores Piperno, The Origins of Plant Cultivation and Domestication in the New World Tropics: Patterns, Process, and New Developments

 

Since then, even earlier dates have been published.

 

According to a genetic study by Embrapa, corn cultivation was introduced in South America from Mexico, in two great waves: the first, more than 6000 years ago, spread through the Andes. Evidence of cultivation in Peru has been found dating to about 6700 years ago. The second wave, about 2000 years ago, through the lowlands of South America.

 

The earliest maize plants grew only small, 25-millimetre-long (1 in) corn cobs, and only one per plant. In Jackson Spielvogel's view, many centuries of artificial selection (rather than the current view that maize was exploited by interplanting with teosinte) by the indigenous people of the Americas resulted in the development of maize plants capable of growing several cobs per plant, which were usually several centimetres/inches long each. The Olmec and Maya cultivated maize in numerous varieties throughout Mesoamerica; they cooked, ground and processed it through nixtamalization. It was believed that beginning about 2500 BC, the crop spread through much of the Americas. Research of the 21st century has established even earlier dates. The region developed a trade network based on surplus and varieties of maize crops.

 

Mapuches of south-central Chile cultivated maize along with quinoa and potatoes in pre-Hispanic times; however, potato was the staple food of most Mapuches, "specially in the southern and coastal [Mapuche] territories where maize did not reach maturity". Before the expansion of the Inca Empire maize was traded and transported as far south as 40°19' S in Melinquina, Lácar Department. In that location maize remains were found inside pottery dated to 730 ± 80 BP and 920 ± 60 BP. Probably this maize was brought across the Andes from Chile. The presence of maize in Guaitecas Archipelago (43°55' S), the southernmost outpost of pre-Hispanic agriculture, is reported by early Spanish explorers. However the Spanish may have misidentified the plant.

 

COLUMBIAN EXCHANGE

After the arrival of Europeans in 1492, Spanish settlers consumed maize, and explorers and traders carried it back to Europe and introduced it to other countries. Spanish settlers far preferred wheat bread to maize, cassava, or potatoes. Maize flour could not be substituted for wheat for communion bread, since in Christian belief only wheat could undergo transubstantiation and be transformed into the body of Christ. Some Spaniards worried that by eating indigenous foods, which they did not consider nutritious, they would weaken and risk turning into Indians. "In the view of Europeans, it was the food they ate, even more than the environment in which they lived, that gave Amerindians and Spaniards both their distinctive physical characteristics and their characteristic personalities." Despite these worries, Spaniards did consume maize. Archeological evidence from Florida sites indicate they cultivated it as well.

 

Maize spread to the rest of the world because of its ability to grow in diverse climates. It was cultivated in Spain just a few decades after Columbus's voyages and then spread to Italy, West Africa and elsewhere. Widespread cultivation most likely began in southern Spain in 1525, after which it quickly spread to the rest of the Spanish Empire including its territories in Italy (and, from there, to other Italian states). Maize had many advantages over wheat and barley; it yielded two and a half times the food energy per unit cultivated area, could be harvested in successive years from the same plot of land, and grew in wildly varying altitudes and climates, from relatively dry regions with only 250 mm (10 in) of annual rainfall to damp regions with over 5,000 mm (200 in). By the 17th century it was a common peasant food in Southwestern Europe, including Portugal, Spain, southern France, and Italy. By the 18th century, it was the chief food of the southern French and Italian peasantry, especially in the form of polenta in Italy.

Names

 

The word maize derives from the Spanish form of the indigenous Taíno word for the plant, mahiz. It is known by other names around the world.

 

The word "corn" outside the US, Canada, Australia, and New Zealand refers to any cereal crop, its meaning understood to vary geographically to refer to the local staple. In the United States, Canada, Australia, and New Zealand, corn primarily means maize; this usage started as a shortening of "Indian corn". "Indian corn" primarily means maize (the staple grain of indigenous Americans), but can refer more specifically to multicolored "flint corn" used for decoration.

 

In places outside the US, Canada, Australia, and New Zealand, corn often refers to maize in culinary contexts. The narrower meaning is usually indicated by some additional word, as in sweet corn, sweetcorn, corn on the cob, baby corn, the puffed confection known as popcorn and the breakfast cereal known as corn flakes.

 

In Southern Africa, maize is commonly called mielie (Afrikaans) or mealie (English), words derived from the Portuguese word for maize, milho.

 

Maize is preferred in formal, scientific, and international usage because it refers specifically to this one grain, unlike corn, which has a complex variety of meanings that vary by context and geographic region. Maize is used by agricultural bodies and research institutes such as the FAO and CSIRO. National agricultural and industry associations often include the word maize in their name even in English-speaking countries where the local, informal word is something other than maize; for example, the Maize Association of Australia, the Indian Maize Development Association, the Kenya Maize Consortium and Maize Breeders Network, the National Maize Association of Nigeria, the Zimbabwe Seed Maize Association.

 

STRUCTURE AND PHYSIOLOGY

The maize plant is often 3 m (10 ft) in height, though some natural strains can grow 13 m (43 ft). The stem is commonly composed of 20 internodes of 18 cm (7 in) length. The leaves arise from the nodes, alternately on opposite sides on the stalk. A leaf, which grows from each node, is generally 9 cm (3+1⁄2 in) in width and 120 cm (3 ft 11 in) in length.

 

Ears develop above a few of the leaves in the midsection of the plant, between the stem and leaf sheath, elongating by around 3 mm (1⁄8 in) per day, to a length of 18 cm (7 in) with 60 cm (24 in) being the maximum alleged in the subspecies. They are female inflorescences, tightly enveloped by several layers of ear leaves commonly called husks. Certain varieties of maize have been bred to produce many additional developed ears. These are the source of the "baby corn" used as a vegetable in Asian cuisine.

 

The apex of the stem ends in the tassel, an inflorescence of male flowers. When the tassel is mature and conditions are suitably warm and dry, anthers on the tassel dehisce and release pollen. Maize pollen is anemophilous (dispersed by wind), and because of its large settling velocity, most pollen falls within a few meters of the tassel.

 

Elongated stigmas, called silks, emerge from the whorl of husk leaves at the end of the ear. They are often pale yellow and 18 cm (7 in) in length, like tufts of hair in appearance. At the end of each is a carpel, which may develop into a "kernel" if fertilized by a pollen grain. The pericarp of the fruit is fused with the seed coat referred to as "caryopsis", typical of the grasses, and the entire kernel is often referred to as the "seed". The cob is close to a multiple fruit in structure, except that the individual fruits (the kernels) never fuse into a single mass. The grains are about the size of peas, and adhere in regular rows around a white, pithy substance, which forms the ear. The maximum size of kernels is reputedly 2.5 cm (1 in). An ear commonly holds 600 kernels. They are of various colors: blackish, bluish-gray, purple, green, red, white and yellow. When ground into flour, maize yields more flour with much less bran than wheat does. It lacks the protein gluten of wheat and, therefore, makes baked goods with poor rising capability. A genetic variant that accumulates more sugar and less starch in the ear is consumed as a vegetable and is called sweet corn. Young ears can be consumed raw, with the cob and silk, but as the plant matures (usually during the summer months), the cob becomes tougher and the silk dries to inedibility. By the end of the growing season, the kernels dry out and become difficult to chew without cooking them tender first in boiling water.

 

Planting density affects multiple aspects of maize. Modern farming techniques in developed countries usually rely on dense planting, which produces one ear per stalk. Stands of silage maize are yet denser,[citation needed] and achieve a lower percentage of ears and more plant matter.

 

Maize is a facultative short-day plant and flowers in a certain number of growing degree days > 10 °C (50 °F) in the environment to which it is adapted. The magnitude of the influence that long nights have on the number of days that must pass before maize flowers is genetically prescribed and regulated by the phytochrome system.

Photoperiodicity can be eccentric in tropical cultivars such that the long days characteristic of higher latitudes allow the plants to grow so tall that they do not have enough time to produce seed before being killed by frost. These attributes, however, may prove useful in using tropical maize for biofuels.

 

Immature maize shoots accumulate a powerful antibiotic substance, 2,4-dihydroxy-7-methoxy-1,4-benzoxazin-3-one (DIMBOA). DIMBOA is a member of a group of hydroxamic acids (also known as benzoxazinoids) that serve as a natural defense against a wide range of pests, including insects, pathogenic fungi and bacteria. DIMBOA is also found in related grasses, particularly wheat. A maize mutant (bx) lacking DIMBOA is highly susceptible to attack by aphids and fungi. DIMBOA is also responsible for the relative resistance of immature maize to the European corn borer (family Crambidae). As maize matures, DIMBOA levels and resistance to the corn borer decline.

 

Because of its shallow roots, maize is susceptible to droughts, intolerant of nutrient-deficient soils, and prone to be uprooted by severe winds.

 

While yellow maizes derive their color from lutein and zeaxanthin, in red-colored maizes, the kernel coloration is due to anthocyanins and phlobaphenes. These latter substances are synthesized in the flavonoids synthetic pathway from polymerization of flavan-4-ols by the expression of maize pericarp color1 (p1) gene which encodes an R2R3 myb-like transcriptional activator of the A1 gene encoding for the dihydroflavonol 4-reductase (reducing dihydroflavonols into flavan-4-ols) while another gene (Suppressor of Pericarp Pigmentation 1 or SPP1) acts as a suppressor. The p1 gene encodes an Myb-homologous transcriptional activator of genes required for biosynthesis of red phlobaphene pigments, while the P1-wr allele specifies colorless kernel pericarp and red cobs, and unstable factor for orange1 (Ufo1) modifies P1-wr expression to confer pigmentation in kernel pericarp, as well as vegetative tissues, which normally do not accumulate significant amounts of phlobaphene pigments. The maize P gene encodes a Myb homolog that recognizes the sequence CCT/AACC, in sharp contrast with the C/TAACGG bound by vertebrate Myb proteins.

 

The ear leaf is the leaf most closely associated with a particular developing ear. This leaf and above contribute 70% to 75% to 90% of grain fill. Therefore fungicide application is most important in that region in most disease environments.

 

ABNORMAL FLOWERS

Maize flowers may sometimes exhibit mutations that lead to the formation of female flowers in the tassel. These mutations, ts4 and Ts6, prohibit the development of the stamen while simultaneously promoting pistil development. This may cause inflorescences containing both male and female flowers, or hermaphrodite flowers.

 

GENETICS

Maize is an annual grass in the family Gramineae, which includes such plants as wheat, rye, barley, rice, sorghum, and sugarcane. There are two major species of the genus Zea (out of six total): Zea mays (maize) and Zea diploperennis, which is a perennial type of teosinte. The annual teosinte variety called Zea mays mexicana is the closest botanical relative to maize. It still grows in the wild as an annual in Mexico and Guatemala.

 

Many forms of maize are used for food, sometimes classified as various subspecies related to the amount of starch each has:

 

Flour corn: Zea mays var. amylacea

Popcorn: Zea mays var. everta

Dent corn : Zea mays var. indentata

Flint corn: Zea mays var. indurata

Sweet corn: Zea mays var. saccharata and Zea mays var. rugosa

Waxy corn: Zea mays var. ceratina

Amylomaize: Zea mays

Pod corn: Zea mays var. tunicata Larrañaga ex A. St. Hil.

Striped maize: Zea mays var. japonica

 

This system has been replaced (though not entirely displaced) over the last 60 years by multivariable classifications based on ever more data. Agronomic data were supplemented by botanical traits for a robust initial classification, then genetic, cytological, protein and DNA evidence was added. Now, the categories are forms (little used), races, racial complexes, and recently branches.

 

Maize is a diploid with 20 chromosomes (n=10). The combined length of the chromosomes is 1500 cM. Some of the maize chromosomes have what are known as "chromosomal knobs": highly repetitive heterochromatic domains that stain darkly. Individual knobs are polymorphic among strains of both maize and teosinte.

 

Barbara McClintock used these knob markers to validate her transposon theory of "jumping genes", for which she won the 1983 Nobel Prize in Physiology or Medicine. Maize is still an important model organism for genetics and developmental biology today.

 

The centromeres have two types of structural components, both of which are found only in the centromeres: Large arrays of CentC, a short satellite DNA; and a few of a family of retrotransposons. The B chromosome, unlike the others, contains an additional repeat which extends into neighboring areas of the chromosome. Centromeres can accidentally shrink during division and still function, although it is thought this will fail if it shrinks below a few hundred kilobase. Kinetochores contain RNA originating from centromeres. Centromere regions can become inactive, and can continue in that state if the chromosome still has another active one.

 

The Maize Genetics Cooperation Stock Center, funded by the USDA Agricultural Research Service and located in the Department of Crop Sciences at the University of Illinois at Urbana-Champaign, is a stock center of maize mutants. The total collection has nearly 80,000 samples. The bulk of the collection consists of several hundred named genes, plus additional gene combinations and other heritable variants. There are about 1000 chromosomal aberrations (e.g., translocations and inversions) and stocks with abnormal chromosome numbers (e.g., tetraploids). Genetic data describing the maize mutant stocks as well as myriad other data about maize genetics can be accessed at MaizeGDB, the Maize Genetics and Genomics Database.

 

In 2005, the US National Science Foundation (NSF), Department of Agriculture (USDA) and the Department of Energy (DOE) formed a consortium to sequence the B73 maize genome. The resulting DNA sequence data was deposited immediately into GenBank, a public repository for genome-sequence data. Sequences and genome annotations have also been made available throughout the project's lifetime at the project's official site.

 

Primary sequencing of the maize genome was completed in 2008. On November 20, 2009, the consortium published results of its sequencing effort in Science. The genome, 85% of which is composed of transposons, was found to contain 32,540 genes (By comparison, the human genome contains about 2.9 billion bases and 26,000 genes). Much of the maize genome has been duplicated and reshuffled by helitrons—group of rolling circle transposons.

 

In Z. mays and various other angiosperms the MADS-box motif is involved in floral development. Early study in several angiosperm models including Z. mays was the beginning of research into the molecular evolution of floral structure in general, as well as their role in nonflowering plants.

 

EVOLUTION

As with many plants and animals, Z. mays has a positive correlation between effective population size and the magnitude of selection pressure. Z. m. having an EPS of ~650,000, it clusters with others of about the same EPS, and has 79% of its amino acid sites under selection.

 

Recombination is a significant source of diversity in Z. mays. (Note that this finding supersedes previous studies which showed no such correlation.)

 

This recombination/diversity effect is seen throughout plants but is also found to not occur – or not as strongly – in regions of high gene density. This is likely the reason that domesticated Z. mays has not seen as much of an increase in diversity within areas of higher density as in regions of lower density, although there is more evidence in other plants.

 

Some lines of maize have undergone ancient polyploidy events, starting 11m years ago. Over that time ~72% of polyploid duplicated genes have been retained, which is higher than other plants with older polyploidy events. Thus maize may be due to lose more duplicate genes as time goes along, similar to the course followed by the genomes of other plants. If so - if gene loss has merely not occurred yet - that could explain the lack of observed positive selection and lower negative selection which are observed in otherwise similar plants, i.e. also naturally outcrossing and with similar effective population sizes.

 

Ploidy does not appear to influence EPS or magnitude of selection effect in maize.

 

BREEDING

Maize reproduces sexually each year. This randomly selects half the genes from a given plant to propagate to the next generation, meaning that desirable traits found in the crop (like high yield or good nutrition) can be lost in subsequent generations unless certain techniques are used.

 

Maize breeding in prehistory resulted in large plants producing large ears. Modern breeding began with individuals who selected highly productive varieties in their fields and then sold seed to other farmers. James L. Reid was one of the earliest and most successful developing Reid's Yellow Dent in the 1860s. These early efforts were based on mass selection. Later breeding efforts included ear to row selection (C. G. Hopkins c. 1896), hybrids made from selected inbred lines (G. H. Shull, 1909), and the highly successful double cross hybrids using four inbred lines (D. F. Jones c. 1918, 1922). University supported breeding programs were especially important in developing and introducing modern hybrids. By the 1930s, companies such as Pioneer devoted to production of hybrid maize had begun to influence long-term development. Internationally important seed banks such as the International Maize and Wheat Improvement Center (CIMMYT) and the US bank at the Maize Genetics Cooperation Stock Center University of Illinois at Urbana-Champaign maintain germplasm important for future crop development.

 

Since the 1940s the best strains of maize have been first-generation hybrids made from inbred strains that have been optimized for specific traits, such as yield, nutrition, drought, pest and disease tolerance. Both conventional cross-breeding and genetic modification have succeeded in increasing output and reducing the need for cropland, pesticides, water and fertilizer. There is conflicting evidence to support the hypothesis that maize yield potential has increased over the past few decades. This suggests that changes in yield potential are associated with leaf angle, lodging resistance, tolerance of high plant density, disease/pest tolerance, and other agronomic traits rather than increase of yield potential per individual plant.

 

Tropical landraces remain an important and underutilized source of resistance alleles for for disease and for herbivores. Notable discoveries of rare alleles for this purpose were made by Dao et al 2014 and Sood et al 2014.

 

GLOBAL PROGRAM

CIMMYT operates a conventional breeding program to provide optimized strains. The program began in the 1980s. Hybrid seeds are distributed in Africa by the Drought Tolerant Maize for Africa project.

 

GENETIC MODIFICATION

Genetically modified (GM) maize was one of the 26 GM crops grown commercially in 2016. The vast majority of this is Bt maize. Grown since 1997 in the United States and Canada, 92% of the US maize crop was genetically modified in 2016 and 33% of the worldwide maize crop was GM in 2016. As of 2011, Herbicide-tolerant maize varieties were grown in Argentina, Australia, Brazil, Canada, China, Colombia, El Salvador, the European Union, Honduras, Japan, Korea, Malaysia, Mexico, New Zealand, Philippines, the Russian Federation, Singapore, South Africa, Taiwan, Thailand, and the United States. Insect-resistant maize was grown in Argentina, Australia, Brazil, Canada, Chile, China, Colombia, Egypt, the European Union, Honduras, Japan, Korea, Malaysia, Mexico, New Zealand, Philippines, South Africa, Switzerland, Taiwan, the United States, and Uruguay.

 

In September 2000, up to $50 million worth of food products were recalled due to the presence of Starlink genetically modified corn, which had been approved only for animal consumption and had not been approved for human consumption, and was subsequently withdrawn from the market.

 

ORIGIN

Maize is the domesticated variant of teosinte. The two plants have dissimilar appearance, maize having a single tall stalk with multiple leaves and teosinte being a short, bushy plant. The difference between the two is largely controlled by differences in just two genes, called grassy tillers-1 (gt1, A0A317YEZ1) and teosinte branched-1 (tb1, Q93WI2).

 

Several theories had been proposed about the specific origin of maize in Mesoamerica:

 

It is a direct domestication of a Mexican annual teosinte, Zea mays ssp. parviglumis, native to the Balsas River valley in south-eastern Mexico, with up to 12% of its genetic material obtained from Zea mays ssp. mexicana through introgression.

It has been derived from hybridization between a small domesticated maize (a slightly changed form of a wild maize) and a teosinte of section Luxuriantes, either Z. luxurians or Z. diploperennis.

It has undergone two or more domestications either of a wild maize or of a teosinte. (The term "teosinte" describes all species and subspecies in the genus Zea, excluding Zea mays ssp. mays.)

It has evolved from a hybridization of Z. diploperennis by Tripsacum dactyloides.

 

In the late 1930s, Paul Mangelsdorf suggested that domesticated maize was the result of a hybridization event between an unknown wild maize and a species of Tripsacum, a related genus. This theory about the origin of maize has been refuted by modern genetic testing, which refutes Mangelsdorf's model and the fourth listed above. 

 

The teosinte origin theory was proposed by the Russian botanist Nikolai Ivanovich Vavilov in 1931 and the later American Nobel Prize-winner George Beadle in 1932.: 10  It is supported experimentally and by recent studies of the plants' genomes. Teosinte and maize can cross-breed and produce fertile offspring. A number of questions remain concerning the species, among them:

 

how the immense diversity of the species of sect. Zea originated,

how the tiny archaeological specimens of 3500–2700 BC could have been selected from a teosinte, and

how domestication could have proceeded without leaving remains of teosinte or maize with teosintoid traits earlier than the earliest known until recently, dating from ca. 1100 BC.

 

The domestication of maize is of particular interest to researchers—archaeologists, geneticists, ethnobotanists, geographers, etc. The process is thought by some to have started 7,500 to 12,000 years ago. Research from the 1950s to 1970s originally focused on the hypothesis that maize domestication occurred in the highlands between the states of Oaxaca and Jalisco, because the oldest archaeological remains of maize known at the time were found there.

Connection with 'parviglumis' subspecies

Genetic studies, published in 2004 by John Doebley, identified Zea mays ssp. parviglumis, native to the Balsas River valley in Mexico's southwestern highlands, and also known as Balsas teosinte, as being the crop wild relative that is genetically most similar to modern maize. This was confirmed by further studies, which refined this hypothesis somewhat. Archaeobotanical studies, published in 2009, point to the middle part of the Balsas River valley as the likely location of early domestication; this river is not very long, so these locations are not very distant. Stone milling tools with maize residue have been found in an 8,700 year old layer of deposits in a cave not far from Iguala, Guerrero.

 

Doebley was part of the team that first published, in 2002, that maize had been domesticated only once, about 9,000 years ago, and then spread throughout the Americas.

 

A primitive corn was being grown in southern Mexico, Central America, and northern South America 7,000 years ago. Archaeological remains of early maize ears, found at Guila Naquitz Cave in the Oaxaca Valley, date back roughly 6,250 years; the oldest ears from caves near Tehuacan, Puebla, 5,450 B.P.

 

Maize pollen dated to 7,300 B.P. from San Andres, Tabasco, on the Caribbean coast has also been recovered.

 

As maize was introduced to new cultures, new uses were developed and new varieties selected to better serve in those preparations. Maize was the staple food, or a major staple – along with squash, Andean region potato, quinoa, beans, and amaranth – of most pre-Columbian North American, Mesoamerican, South American, and Caribbean cultures. The Mesoamerican civilization, in particular, was deeply interrelated with maize. Its traditions and rituals involved all aspects of maize cultivation – from the planting to the food preparation. Maize formed the Mesoamerican people's identity.

 

It is unknown what precipitated its domestication, because the edible portion of the wild variety is too small, and hard to obtain, to be eaten directly, as each kernel is enclosed in a very hard bivalve shell.

 

In 1939, George Beadle demonstrated that the kernels of teosinte are readily "popped" for human consumption, like modern popcorn. Some have argued it would have taken too many generations of selective breeding to produce large, compressed ears for efficient cultivation. However, studies of the hybrids readily made by intercrossing teosinte and modern maize suggest this objection is not well founded.

 

SPREADING TO THE NORTH

Around 4,500 ago, maize began to spread to the north; it was first cultivated in what is now the United States at several sites in New Mexico and Arizona, about 4,100 ago.

 

During the first millennium AD, maize cultivation spread more widely in the areas north. In particular, the large-scale adoption of maize agriculture and consumption in eastern North America took place about A.D. 900. Native Americans cleared large forest and grassland areas for the new crop.

 

In 2005, research by the USDA Forest Service suggested that the rise in maize cultivation 500 to 1,000 years ago in what is now the southeastern United States corresponded with a decline of freshwater mussels, which are very sensitive to environmental changes.

 

CULTIVATION

PLANTING

Because it is cold-intolerant, in the temperate zones maize must be planted in the spring. Its root system is generally shallow, so the plant is dependent on soil moisture. As a plant that uses C4 carbon fixation, maize is a considerably more water-efficient crop than plants that use C3 carbon fixation such as alfalfa and soybeans. Maize is most sensitive to drought at the time of silk emergence, when the flowers are ready for pollination. In the United States, a good harvest was traditionally predicted if the maize was "knee-high by the Fourth of July", although modern hybrids generally exceed this growth rate. Maize used for silage is harvested while the plant is green and the fruit immature. Sweet corn is harvested in the "milk stage", after pollination but before starch has formed, between late summer and early to mid-autumn. Field maize is left in the field until very late in the autumn to thoroughly dry the grain, and may, in fact, sometimes not be harvested until winter or even early spring. The importance of sufficient soil moisture is shown in many parts of Africa, where periodic drought regularly causes maize crop failure and consequent famine. Although it is grown mainly in wet, hot climates, it has been said to thrive in cold, hot, dry or wet conditions, meaning that it is an extremely versatile crop.

 

Maize was planted by the Native Americans in hills, in a complex system known to some as the Three Sisters. Maize provided support for beans, and the beans provided nitrogen derived from nitrogen-fixing rhizobia bacteria which live on the roots of beans and other legumes; and squashes provided ground cover to stop weeds and inhibit evaporation by providing shade over the soil. This method was replaced by single species hill planting where each hill 60–120 cm (2 ft 0 in–3 ft 11 in) apart was planted with three or four seeds, a method still used by home gardeners. A later technique was "checked maize", where hills were placed

 

1 m (40 in) apart in each direction, allowing cultivators to run through the field in two directions. In more arid lands, this was altered and seeds were planted in the bottom of 10–12 cm (4–4+1⁄2 in) deep furrows to collect water. Modern technique plants maize in rows which allows for cultivation while the plant is young, although the hill technique is still used in the maize fields of some Native American reservations. When maize is planted in rows, it also allows for planting of other crops between these rows to make more efficient use of land space.

 

In most regions today, maize grown in residential gardens is still often planted manually with a hoe, whereas maize grown commercially is no longer planted manually but rather is planted with a planter. In North America, fields are often planted in a two-crop rotation with a nitrogen-fixing crop, often alfalfa in cooler climates and soybeans in regions with longer summers. Sometimes a third crop, winter wheat, is added to the rotation.

 

Many of the maize varieties grown in the United States and Canada are hybrids. Often the varieties have been genetically modified to tolerate glyphosate or to provide protection against natural pests. Glyphosate is an herbicide which kills all plants except those with genetic tolerance. This genetic tolerance is very rarely found in nature.

 

In the midwestern United States, low-till or no-till farming techniques are usually used. In low-till, fields are covered once, maybe twice, with a tillage implement either ahead of crop planting or after the previous harvest. The fields are planted and fertilized. Weeds are controlled through the use of herbicides, and no cultivation tillage is done during the growing season. This technique reduces moisture evaporation from the soil, and thus provides more moisture for the crop. The technologies mentioned in the previous paragraph enable low-till and no-till farming. Weeds compete with the crop for moisture and nutrients, making them undesirable.

 

HARVESTING

Before the 20th century, all maize harvesting was by manual labour, by grazing, or by some combination of those. Whether the ears were hand-picked and the stover was grazed, or the whole plant was cut, gathered, and shocked, people and livestock did all the work. Between the 1890s and the 1970s, the technology of maize harvesting expanded greatly. Today, all such technologies, from entirely manual harvesting to entirely mechanized, are still in use to some degree, as appropriate to each farm's needs, although the thoroughly mechanized versions predominate, as they offer the lowest unit costs when scaled to large farm operations. For small farms, their unit cost can be too high, as their higher fixed cost cannot be amortized over as many units.[citation needed]

 

Before World War II, most maize in North America was harvested by hand. This involved a large number of workers and associated social events (husking or shucking bees). From the 1890s onward, some machinery became available to partially mechanize the processes, such as one- and two-row mechanical pickers (picking the ear, leaving the stover) and corn binders, which are reaper-binders designed specifically for maize (for example, Video on YouTube). The latter produce sheaves that can be shocked. By hand or mechanical picker, the entire ear is harvested, which then requires a separate operation of a maize sheller to remove the kernels from the ear. Whole ears of maize were often stored in corn cribs, and these whole ears are a sufficient form for some livestock feeding use. Today corn cribs with whole ears, and corn binders, are less common because most modern farms harvest the grain from the field with a combine and store it in bins. The combine with a corn head (with points and snap rolls instead of a reel) does not cut the stalk; it simply pulls the stalk down. The stalk continues downward and is crumpled into a mangled pile on the ground, where it usually is left to become organic matter for the soil. The ear of maize is too large to pass between slots in a plate as the snap rolls pull the stalk away, leaving only the ear and husk to enter the machinery. The combine separates the husk and the cob, keeping only the kernels.

When maize is a silage crop, the entire plant is usually chopped at once with a forage harvester (chopper) and ensiled in silos or polymer wrappers. Ensiling of sheaves cut by a corn binder was formerly common in some regions but has become uncommon. For storing grain in bins, the moisture of the grain must be sufficiently low to avoid spoiling. If the moisture content of the harvested grain is too high, grain dryers are used to reduce the moisture content by blowing heated air through the grain. This can require large amounts of energy in the form of combustible gases (propane or natural gas) and electricity to power the blowers.

 

PRODUCTION

Maize is widely cultivated throughout the world, and a greater weight of maize is produced each year than any other grain. In 2018, total world production was 1.15 billion tonnes, led by the United States with 34.2% of the total (table). China produced 22.4% of the global total.

 

UNITED STATES

In 2016, maize production was forecast to be over 380 million metric tons (15 billion bushels), an increase of 11% over 2014 American production. Based on conditions as of August 2016, the expected yield would be the highest ever for the United States. The area of harvested maize was forecast to be 35 million hectares (87 million acres), an increase of 7% over 2015. Maize is especially popular in Midwestern states such as Indiana, Iowa, and Illinois; in the latter, it was named the state's official grain in 2017.

 

STORAGE

Drying is vital to prevent or at least reduce mycotoxin contamination. Aspergillus and Fusarium spp. are the most common mycotoxin sources, but there are others. Altogether maize contaminants are so common, and this crop is so economically important, that maize mycotoxins are among the most important in agriculture in general.

 

USES

HUMAN FOOD

Maize and cornmeal (ground dried maize) constitute a staple food in many regions of the world. Maize is used to produce cornstarch, a common ingredient in home cooking and many industrialized food products. Maize starch can be hydrolyzed and enzymatically treated to produce syrups, particularly high fructose corn syrup, a sweetener; and also fermented and distilled to produce grain alcohol. Grain alcohol from maize is traditionally the source of Bourbon whiskey. Corn flour is used to make cornbread and other baked products.

 

In prehistoric times Mesoamerican women used a metate to process maize into ground cornmeal, allowing the preparation of foods that were more calorie dense than popcorn. After ceramic vessels were invented the Olmec people began to cook maize together with beans, improving the nutritional value of the staple meal. Although maize naturally contains niacin, an important nutrient, it was not bioavailable without the process of nixtamalization. The Maya used nixtamal meal to make varieties of porridges and tamales. The process was later used in the cuisine of the American South to prepare corn for grits and hominy.

 

Maize is a staple of Mexican cuisine. Masa (cornmeal treated with limewater) is the main ingredient for tortillas, atole and many other dishes of Central American food. It is the main ingredient of corn tortilla, tamales, pozole, atole and all the dishes based on them, like tacos, quesadillas, chilaquiles, enchiladas, tostadas and many more. In Mexico the fungus of maize, known as huitlacoche, is considered a delicacy.

 

Coarse maize meal is made into a thick porridge in many cultures: from the polenta of Italy, the angu of Brazil, the mămăligă of Romania, to cornmeal mush in the US (or hominy grits in the South) or the food called mieliepap in South Africa and sadza, nshima, ugali and other names in other parts of Africa. Introduced into Africa by the Portuguese in the 16th century, maize has become Africa's most important staple food crop. These are commonly eaten in the Southeastern United States, foods handed down from Native Americans, who called the dish sagamite.

 

Maize can also be harvested and consumed in the unripe state, when the kernels are fully grown but still soft. Unripe maize must usually be cooked to become palatable; this may be done by simply boiling or roasting the whole ears and eating the kernels right off the cob. Sweet corn, a genetic variety that is high in sugars and low in starch, is usually consumed in the unripe state. Such corn on the cob is a common dish in the United States, Canada, United Kingdom, Cyprus, some parts of South America, and the Balkans, but virtually unheard of in some European countries. Corn on the cob was hawked on the streets of early 19th-century New York City by poor, barefoot "Hot Corn Girls", who were thus the precursors of hot dog carts, churro wagons, and fruit stands seen on the streets of big cities today.

 

Within the United States, the usage of maize for human consumption constitutes only around 1/40th of the amount grown in the country. In the United States and Canada, maize is mostly grown to feed livestock, as forage, silage (made by fermentation of chopped green cornstalks), or grain. Maize meal is also a significant ingredient of some commercial animal food products.

 

NUTRITIONAL VALUE

Raw, yellow, sweet maize kernels are composed of 76% water, 19% carbohydrates, 3% protein, and 1% fat (table). In a 100-gram serving, maize kernels provide 86 calories and are a good source (10–19% of the Daily Value) of the B vitamins, thiamin, niacin (but see Pellagra warning below), pantothenic acid (B5) and folate (right table for raw, uncooked kernels, USDA Nutrient Database). In moderate amounts, they also supply dietary fiber and the essential minerals, magnesium and phosphorus whereas other nutrients are in low amounts (table).

 

Maize has suboptimal amounts of the essential amino acids tryptophan and lysine, which accounts for its lower status as a protein source. However, the proteins of beans and legumes complement those of maize.

 

FEED AND FODDER FOR LIVESTOCK

Maize is a major source of both grain feed and fodder for livestock. It is fed to the livestock in various ways. When it is used as a grain crop, the dried kernels are used as feed. They are often kept on the cob for storage in a corn crib, or they may be shelled off for storage in a grain bin. The farm that consumes the feed may produce it, purchase it on the market, or some of both. When the grain is used for feed, the rest of the plant (the corn stover) can be used later as fodder, bedding (litter), or soil amendment. When the whole maize plant (grain plus stalks and leaves) is used for fodder, it is usually chopped all at once and ensilaged, as digestibility and palatability are higher in the ensilaged form than in the dried form. Maize silage is one of the most valuable forages for ruminants. Before the advent of widespread ensilaging, it was traditional to gather the corn into shocks after harvesting, where it dried further. With or without a subsequent move to the cover of a barn, it was then stored for weeks to several months until fed to the livestock. Today ensilaging can occur not only in siloes but also in silage wrappers. However, in the tropics, maize can be harvested year-round and fed as green forage to the animals.

 

CHEMICALS

Starch from maize can also be made into plastics, fabrics, adhesives, and many other chemical products.

 

The corn steep liquor, a plentiful watery byproduct of maize wet milling process, is widely used in the biochemical industry and research as a culture medium to grow many kinds of microorganisms.

 

Chrysanthemin is found in purple corn and is used as a food coloring.

 

BIO-FUEL

"Feed maize" is being used increasingly for heating; specialized corn stoves (similar to wood stoves) are available and use either feed maize or wood pellets to generate heat. Maize cobs are also used as a biomass fuel source. Maize is relatively cheap and home-heating furnaces have been developed which use maize kernels as a fuel. They feature a large hopper that feeds the uniformly sized maize kernels (or wood pellets or cherry pits) into the fire.[citation needed]

 

Maize is increasingly used as a feedstock for the production of ethanol fuel. When considering where to construct an ethanol plant, one of the site selection criteria is to ensure there is locally available feedstock. Ethanol is mixed with gasoline to decrease the amount of pollutants emitted when used to fuel motor vehicles. High fuel prices in mid-2007 led to higher demand for ethanol, which in turn led to higher prices paid to farmers for maize. This led to the 2007 harvest being one of the most profitable maize crops in modern history for farmers. Because of the relationship between fuel and maize, prices paid for the crop now tend to track the price of oil.

 

The price of food is affected to a certain degree by the use of maize for biofuel production. The cost of transportation, production, and marketing are a large portion (80%) of the price of food in the United States. Higher energy costs affect these costs, especially transportation. The increase in food prices the consumer has been seeing is mainly due to the higher energy cost. The effect of biofuel production on other food crop prices is indirect. Use of maize for biofuel production increases the demand, and therefore price of maize. This, in turn, results in farm acreage being diverted from other food crops to maize production. This reduces the supply of the other food crops and increases their prices.

 

Maize is widely used in Germany as a feedstock for biogas plants. Here the maize is harvested, shredded then placed in silage clamps from which it is fed into the biogas plants. This process makes use of the whole plant rather than simply using the kernels as in the production of fuel ethanol.

 

A biomass gasification power plant in Strem near Güssing, Burgenland, Austria, began in 2005. Research is being done to make diesel out of the biogas by the Fischer Tropsch method.

 

Increasingly, ethanol is being used at low concentrations (10% or less) as an additive in gasoline (gasohol) for motor fuels to increase the octane rating, lower pollutants, and reduce petroleum use (what is nowadays also known as "biofuels" and has been generating an intense debate regarding the human beings' necessity of new sources of energy, on the one hand, and the need to maintain, in regions such as Latin America, the food habits and culture which has been the essence of civilizations such as the one originated in Mesoamerica; the entry, January 2008, of maize among the commercial agreements of NAFTA has increased this debate, considering the bad labor conditions of workers in the fields, and mainly the fact that NAFTA "opened the doors to the import of maize from the United States, where the farmers who grow it receive multimillion-dollar subsidies and other government supports. ... According to OXFAM UK, after NAFTA went into effect, the price of maize in Mexico fell 70% between 1994 and 2001. The number of farm jobs dropped as well: from 8.1 million in 1993 to 6.8 million in 2002. Many of those who found themselves without work were small-scale maize growers."). However, introduction in the northern latitudes of the US of tropical maize for biofuels, and not for human or animal consumption, may potentially alleviate this.

 

COMMODITY

Maize is bought and sold by investors and price speculators as a tradable commodity using corn futures contracts. These "futures" are traded on the Chicago Board of Trade (CBOT) under ticker symbol C. They are delivered every year in March, May, July, September, and December.

 

ORNAMENTAL AND OTHER USES

Some forms of the plant are occasionally grown for ornamental use in the garden. For this purpose, variegated and colored leaf forms as well as those with colorful ears are used.

 

Corncobs can be hollowed out and treated to make inexpensive smoking pipes, first manufactured in the United States in 1869.

 

An unusual use for maize is to create a "corn maze" (or "maize maze") as a tourist attraction. The idea of a maize maze was introduced by the American Maze Company who created a maze in Pennsylvania in 1993. Traditional mazes are most commonly grown using yew hedges, but these take several years to mature. The rapid growth of a field of maize allows a maze to be laid out using GPS at the start of a growing season and for the maize to grow tall enough to obstruct a visitor's line of sight by the start of the summer. In Canada and the US, these are popular in many farming communities.

 

Maize kernels can be used in place of sand in a sandboxlike enclosure for children's play.

 

Stigmas from female maize flowers, popularly called corn silk, are sold as herbal supplements.

 

Maize is used as a fish bait, called "dough balls". It is particularly popular in Europe for coarse fishing.

 

Additionally, feed corn is sometimes used by hunters to bait animals such as deer or wild hogs.

 

UNITED STATES USAGE BREAKDOWN

The breakdown of usage of the 12.1-billion-bushel (307-million-tonne) 2008 US maize crop was as follows, according to the World Agricultural Supply and Demand Estimates Report by the USDA.In the US since 2009/2010, maize feedstock use for ethanol production has somewhat exceeded direct use for livestock feed; maize use for fuel ethanol was 5,130 million bushels (130 million tonnes) in the 2013/2014 marketing year.A fraction of the maize feedstock dry matter used for ethanol production is usefully recovered as DDGS (dried distillers grains with solubles). In the 2010/2011 marketing year, about 29.1 million tonnes of DDGS were fed to US livestock and poultry. Because starch utilization in fermentation for ethanol production leaves other grain constituents more concentrated in the residue, the feed value per kg of DDGS, with regard to ruminant-metabolizable energy and protein, exceeds that of the grain. Feed value for monogastric animals, such as swine and poultry, is somewhat lower than for ruminants.

 

HAZARDS

PELLAGRA

When maize was first introduced into farming systems other than those used by traditional native-American peoples, it was generally welcomed with enthusiasm for its productivity. However, a widespread problem of malnutrition soon arose wherever maize was introduced as a staple food. This was a mystery, since these types of malnutrition were not normally seen among the indigenous Americans, for whom maize was the principal staple food.

 

It was eventually discovered that the indigenous Americans had learned to soak maize in alkali — water (the process now known as nixtamalization) — made with ashes and lime (calcium oxide) since at least 1200–1500 BC by Mesoamericans. They did this to liberate the corn hulls, but (unbeknownst to natives or colonists) it coincidentally liberates the B-vitamin niacin, the lack of which was the underlying cause of the condition known as pellagra.

 

Maize was introduced into the diet of non-indigenous Americans without the necessary cultural knowledge acquired over thousands of years in the Americas. In the late 19th century, pellagra reached epidemic proportions in parts of the southern US, as medical researchers debated two theories for its origin: the deficiency theory (which was eventually shown to be true) said that pellagra was due to a deficiency of some nutrient, and the germ theory said that pellagra was caused by a germ transmitted by stable flies. A third theory, promoted by the eugenicist Charles Davenport, held that people only contracted pellagra if they were susceptible to it due to certain "constitutional, inheritable" traits of the affected individual.

 

Once alkali processing and dietary variety were understood and applied, pellagra disappeared in the developed world. The development of high lysine maize and the promotion of a more balanced diet have also contributed to its demise. Pellagra still exists today in food-poor areas and refugee camps where people survive on donated maize.

 

ALLERGY

Maize contains lipid transfer protein, an indigestible protein that survives cooking. This protein has been linked to a rare and understudied allergy to maize in humans. The allergic reaction can cause skin rash, swelling or itching of mucous membranes, diarrhea, vomiting, asthma and, in severe cases, anaphylaxis. It is unclear how common this allergy is in the general population.

 

MYCOTOXINS

Fungicide application does not reduce fungal growth or mycotoxin dramatically, although it can be a part of a successful reduction strategy. Among the most common toxins are those produced by Aspergillus and Fusarium spp. The most common toxins are aflatoxins, fumonisins, zearalenone, and ochratoxin A. Bt maize discourages insect vectors and by so doing it dramatically reduces concentrations of fumonisins, significantly reduces aflatoxins, but only mildly reduces others.

 

ART

Maize has been an essential crop in the Andes since the pre-Columbian era. The Moche culture from Northern Peru made ceramics from earth, water, and fire. This pottery was a sacred substance, formed in significant shapes and used to represent important themes. Maize was represented anthropomorphically as well as naturally.

 

In the United States, maize ears along with tobacco leaves are carved into the capitals of columns in the United States Capitol building. Maize itself is sometimes used for temporary architectural detailing when the intent is to celebrate the fall season, local agricultural productivity and culture. Bundles of dried maize stalks are often displayed along with pumpkins, gourds and straw in autumnal displays outside homes and businesses. A well-known example of architectural use is the Corn Palace in Mitchell, South Dakota, which uses cobs and ears of colored maize to implement a mural design that is recycled annually. Another well-known example is the Field of Corn sculpture in Dublin, Ohio, where hundreds of concrete ears of corn stand in a grassy field.

 

A maize stalk with two ripe ears is depicted on the reverse of the Croatian 1 lipa coin, minted since 1993.

 

WIKIPEDIA

These snails are all the same species, Cepaea nemoralis the Brown-lipped Snail. They are shown here in three polymorphic variations.

Mais uma vez, muito obrigado ao Marcelo Cazani (Marcazani) pela identificação deste pássaro.

Once more, thank you very much Marcelo Cazani (Marcazani) for the identification of this bird.

 

A text, in english, from Wikipedia, the free encyclopedia:

See at en.wikipedia.org/wiki/American_Kestrel

Jump to: navigation, search

American Kestrel

Conservation status

 

Least Concern (IUCN 3.1)

Scientific classification

Kingdom: Animalia

Phylum: Chordata

Class: Aves

Order: Falconiformes

Family: Falconidae

Genus: Falco

Species: F. sparverius

Binomial name

Falco sparverius

Linnaeus, 1758

Synonyms

 

Cerchneis sparverius

Tinnunculus sparverius

 

The American Kestrel (Falco sparverius) is a small falcon. This bird was (and sometimes still is) colloquially known in North America as the "Sparrow Hawk". This name is misleading because it implies a connection with the Eurasian Sparrowhawk Accipiter nisus, which is unrelated; the latter is an accipiter rather than a falcon. Though both are diurnal raptors, they are only distantly related.

American Kestrels are widely distributed across the Americas. Their breeding range extends from central and western Alaska across northern Canada to Nova Scotia, and south throughout North America, into central Mexico, the Baja, and the Caribbean. They are local breeders in Central America and are widely distributed throughout South America.

Most of the birds breeding in Canada and the northern United States migrate south in the winter, although some males stay as year-round residents. It is a very rare vagrant to western Europe.

The American Kestrel is the smallest falcon in North America—about the size of an American Robin. Like all raptors, the American Kestrel is sexually dimorphic, although there is some overlap within the species. The female ranges in length from 23 to 28 centimeters (9-11 inches) with a wingspan of 53–61 centimeters (21–24 inches) and weighs an average of 120 grams (4.2 ounces). The length of the male varies between 20–25 centimeters (8–10 inches) with a wingspan ranging from 51–56 centimeters (20–22 inches) and weighing an average of 111 grams (3.9 ounces). These subtle differences are often difficult to discern in the field.

The coloration of the feathers, however, greatly varies between the sexes. Males have blue-grey secondary feathers on their wings, while the undersides are white with black barring. The back is rufous in coloration, with barring on the lower half. The belly and flanks are white with black spotting. The tail is also rufous, except for the outer rectrix set, which is white with a black subterminal band.

The back of the female American Kestrel is rufous with dark brown barring. The wings exhibit similar coloration and patterning to the back. The undersides of the females are white with rufous streaking. The tail of the female is noticeably different from the male, being rufous in color with numerous narrow dark brown or black bars. Juveniles exhibit coloration patterns similar to the adults.

In both sexes, the head is white with a bluish-grey top. There are also two narrow, vertical black facial markings on each side of the head; one below the eyes and one on the rear portion of the auriculars. Two black spots (ocelli) can be found on each side of the white or orangish nape. The wings are moderately long, fairly narrow, and taper to a point. While perched, the wingtips are noticeably shorter than the tail tip.

The American Kestrel has three basic vocalizations - the "klee" or "killy", the "whine", and the "chitter." The "klee" or "killy" is usually delivered as a rapid series - "killy, killy, killy, killy" when the kestrel is upset or excited. It is used at the apex of the dive display, during fights with other kestrels, and after unsuccessful hunting attempts.

The "whine" may last as long as one to two minutes and may be single or treble. The more intense the situation, the more likely the "whine" will move toward the treble extreme. "Whines" are given during courtship feeding and copulation. The treble whine is heard from breeding females and fledged hungry young.

The "chitter" is the most frequent vocalization in male - female interactions. Its volume and duration depends upon the stress or excitement of the situation. It is associated with friendly approaches and bodily contact between the sexes during breeding season. Occasionally a "chitter" follows a "whine."

Calling occurs throughout the day. Nestlings at two weeks can produce all three vocalizations. Female kestrels tend to have slightly lower pitched and harsher voices than males.

This bird is apparently not a true kestrel. mtDNA cytochrome b sequence analysis (Groombridge et al. 2002) indicates a Late Miocene split[1] between the ancestors of the American Kestrel, and those of the Common Kestrel and its closest relatives. The color pattern with its large areas of brown is reminiscent of kestrels, but the coloration of the head - notably the black ear patch, which is not found in any of the true kestrels - and the male's extensively gray wings are suggestive of a closer relationship with the hobbies, an informal grouping of falcons of usually average size.

Species such as the Merlin and the Aplomado Falcon are proposed as possible close relatives. Indeed, the Merlin is a highly polymorphic bird and although its grey tail and back are distinctive, certain morphs are the only birds that might conceivably be confused with American Kestrels. Conclusive evidence is lacking, and what can be said at present judging from the fairly noninclusive DNA sequence studies[2] is that the general relationships of the present species seem to lie with a number of rather basal "hobby" lineages, such as the Merlin and Aplomado Falcon mentioned already, or the Red-footed and Amur Falcons - or even the Peregrine Falcon lineage with its large species.

The American Kestrel is not very closely related to any of these groups, although it might be closer to the Aplomado Falcon (and its presumed close relatives, the Bat and Orange-breasted Falcons) than to any other living species (Wink et al. 1998) - an association that is also better supported by biogeography than a close relationship with the exclusively Old World true kestrels. It is nonetheless highly distinct in morphology from any of these and, interestingly, has a syrinx similar to the Peregrine and the hierofalcons[3].

In conclusion, until better evidence is available, it is best considered part of a radiation of falcon lineages that diversified around the North Atlantic at the end of the Miocene. Though several fossils of small falcons arte known from North America at roughly the correct time, the earliest testimony of the American Kestrel lineage is Pleistocene remains of the living species (Brodkorb 1964).

American Kestrels are found in a variety of habitats including parks, suburbs, open fields, forest edges and openings, alpine zones, grasslands, marshes, open areas on mountainsides, prairies, plains, deserts with giant cacti, and freeway and highway corridors.

In addition to requiring open space for hunting, American Kestrels seem to need perches for hunting from, cavities for nesting (either natural or man-made), and a sufficient food supply.

The American Kestrel is the only North American falcon to habitually hover with rapid wing beats, keeping its head motionless while scanning the ground for prey. The kestrel commonly perches along fences and powerlines. It glides with flat wings and wingtips curved upward. It occasionally soars in circles with its tail spread and its wings flat.

This falcon species is not long-lived. The oldest banded wild bird was 11 years and seven months old while a captive lived 17 years. A mortality rate average of 57 percent was found. First year mortality rates have declined since 1945 with a decrease in shooting. Major causes of death include collision with traffic, illegal shooting, and predation by other raptors, including the Red-tailed Hawk, Northern Goshawk, Cooper's Hawk, Peregrine Falcon, Barn Owl, and Great Horned Owl.

In summer, kestrels feed largely on grasshoppers, dragonflies, lizards, mice, and voles. They will also eat other small birds. Wintering birds feed primarily on rodents and birds. The birds characteristically hunt along roadsides from telephone wires, fence posts, trees or other convenient perches when not flying in search of food. When they are flying and looking for food they frequently hover with rapid wingbeats.

Because it feeds on both insects and vertebrates, the American Kestrel maintains fairly high population densities. It has a small breeding home range, from 1.75 square miles (4.5 km²) to 2 square miles (5.2 km²). Territory size has been estimated at 269 acres (1.1 km²) to 321 acres (1.3 km²) with much larger wintering home ranges.

Several hunting techniques are used by the American Kestrel. It will hover over one spot—when prey is sighted the kestrel will partly fold its wings and drop lower once or several times before striking. When the prey disappears the falcon will glide in a semicircle before turning back into the wind to hover again. It will also soar in circles, or figure eights, using the same stooping tactics as when hovering.

The kestrel commonly hunts from elevated perch sites, waiting for prey to move on the ground. The kestrel bobs its head and pumps its tail just before attacking.

Other prey capture techniques include direct pursuit, landing and flushing prey from the ground (especially for grasshoppers)and then taking them in flight, capturing flying insects from an elevated perch, and nest robbing including the burrows of Bank Swallows and the nests of Cliff Swallows. It is also an occasional bat catcher, taking bats from their tree roosts, or striking bats in flight from above or as the bats leave or enter caves. The kestrel will kill and cache food items.

The American Kestrel occasionally robs others of the same species. It has also been known to rob a shrike of its prey. Kestrels sometimes harass other hawks, and even Golden Eagles, in flight.

American Kestrels form pairs in which the bond is strong, tending toward permanence. Returning migrants commonly re-establish territories held the previous year. In one study[citation needed], a pair nested in the same tree for six consecutive years. Nesting occurs from late spring to late summer in North America, with incubation underway by the end of May[4]; in tropical South America the birds breed roughly from June onwards through to September or so[5].

Courtship begins shortly after the male establishes a territory. In early courtship, he may give the "dive display", a series of climbs and dives 33 to 66 feet (10 to 20 m) with 3-5 "klee" notes given near the peak of each climb. He may present the female with food during courtship feeding. He may entice her to the nest site by calling. He may "flutter-glide" toward her with quick and shallow wing beats while carrying food and she may also beg for food by flutter-gliding. The female initiates copulation by bowing with her tail in line with her body or slightly raised.

A cavity nester, American Kestrels will use holes in trees, rock cavities and crevices in cliffs, artificial nest boxes, or small spaces in buildings. The number of suitable breeding cavities limits this species' breeding density. The American Kestrel has adapted well to nest boxes. In one program, nest boxes were fixed to the backs of signs along a freeway thus allowing kestrels to breed in areas formerly devoid of nest sites. Pairs nesting in boxes on poles have much higher nesting success than pairs using boxes on trees. No nest is built inside. In nest boxes sawdust and wood shavings may be a suitable substrate for the eggs. Males and females defend the nest against intruders, with the male maintaining a small core territory and the female defending the nest cavity directly rather the surroundings[5].

Both sexes take turns incubating their eggs, a very rare situation among North American birds of prey where the female usually incubates exclusively. Correspondingly, both sexes develop bare oval patches on each side of their breasts where the warm bare skin can contact the eggs for warming. Eggs hatch 29 to 31 days after being laid.[6] There are from three to seven eggs laid, but four to six are average.[6] The eggs are typically short elliptical in shape, and are white or pinkish-white with an even covering of fine spots and flecks of brown shades, occasionally concentrating as a ring or a cap. They will renest if the first nest fails and have been reported to raise 2 broods per year in some of the southern states.

The young grow very quickly, becoming noisy between day 11 and 14 and assuming adult weight in about 2.5 weeks. The young fledge in 30 to 31 days.[7] [6]Early fledgling behavior varies. Broods typically stay together for a week or two. Some broods remain close to the nest area for a week or two while others travel throughout the parents' home range. Generally, young do not disperse more than 0.6 miles (one km) away from the nest area until two to four weeks old. Young disperse as hunting skills develop. Occasionally groups of older juveniles from various broods join together into flocks.

The American Kestrel can be double-brooded, particularly in the southern United States, in areas of abundant small mammals. Replacement clutches can be laid.

 

Um texto em português:

Falcão-americano ou quiriquiri (Falco sparverius) é um pequeno falcão (23-27 cm de comprimento e 85-140 g de peso). Tem uma ampla área de distribuição: desde o Alasca e Norte do Canadá até à ponta Sul da América do Sul (Terra do Fogo). Tem asas azul acizentadas. O dorso é avermelhado pontuado de preto. A cauda também é avermelhada possuindo uma larga lista preta. Na face possui 2 listas verticais que começam junto aos olhos e seguem para baixo. É um predador de pequeno tamanho, alimentando-se de insetos e microvertrebrados como roedores e pequenos pássaros; ocasionalmente caça morcegos.

Falcão é o nome genérico dado a várias aves da família Falconidae, mais estritamente aos animais classificados dentro do género Falco. O que diferencia os falcões das demais aves de rapina é o fato de terem evoluído no sentido de uma especialização no voo em velocidade (em oposição ao voo planado das águias e abutres e ao voo acrobático dos gaviões), facilitado pelas asas ponteagudas e finas, favorecendo a caça em espaços abertos – daí o fato dos falcões não serem aves de ambientes florestais, preferindo montanhas e penhascos, pradarias, estepes e desertos. Os falcões podem ser identificados, aliás, pelo fato de não planarem em correntes termais, como outras aves de rapina. O falcão-peregrino, especializado na caça de aves médias e grandes em voo, pode atingir 300 km/h em voo picado e é o animal mais rápido da terra. Diferentemente das águias e gaviões, que matam suas presas com os pés, os falcões utilizam as garras apenas para apreenderem a presa, matando-a depois com o bico por desconjuntamento das vértebras, para o que possuem um rebordo em forma de dente na mandíbula superior.

 

Na Idade Média, os falcões eram apreciados como animais de caça acessíveis apenas à elite.

The variable oystercatcher (Haematopus unicolor) is a species of wader in the family Haematopodidae. It is endemic to New Zealand. The Maori name is torea-pango] They are also known as 'red bills'.

 

"Variable" refers to the frontal plumage, which ranges from pied through mottled to all black. They are polymorphic meaning they have different genetic variants. Blacker birds are more common in the south.

 

All Stewart Island variable oystercatchers are black. They have pink legs, an orange eye ring and red beaks. They are often seen in pairs on the coast all around New Zealand. During breeding, the pair will defend their territory, sometimes aggressively. Once mated pairs rarely divorce.

 

After breeding they may be seen within flocks, or on the edges of flocks, of black and white South Island pied oystercatcher (SIPO) which also have vivid orange beaks. After breeding they may even form small flocks of their own.

 

Males are around 678 grams and females slightly larger at around 724 grams. Variables can be identified as they are slightly larger than the SIPO - SIPO are around 550 grams.

 

Occasionally totally black but if they are pied (black and white) they can be easily confused with SIPO. The variable species has less definition between the black and the white area, as well as a mottled band on the leading edges of the underwing. Variables also have a smaller white rump patch which is only a band across the base of the tail rather than a wide wedge shape reaching up to the middle of the back as in the SIPO.

 

When mottled they are sometimes called 'smudgies'. They feed on molluscs, crabs and marine worms. After heavy rain, they sometime go inland in search of earthworms. They can open a shellfish by either hammering a hole in it or getting the bill between the two shells (of a bivalve) and twisting them apart.

 

They breed in North Island, South Island, Stewart Island, and Chatham Islands. They do not breed inland or beside rivers although the SIPO does.

 

They nest on the shore between rocks or on sand dunes by making a scrape out of the sand or shingle, sometimes lined with some seaweed. When in flight they make a high pitched 'kleep kleep' sound.

 

They usually lay 2-3 eggs but they can lay up to 5. The eggs are typically stone coloured with small brown patches all over. Eggs hatch in 25–32 days. Chicks are well camouflaged by their colour and can fly in about 6 weeks. The bird lives up to about 27 years.

 

This image was taken on the jetty at Motuara Island in the Queen Charlotte Sound near Picton on the south Island of New Zealand.

  

リコリス・ロンギチュバ ‘ピュア・ホワイト’

Lycoris longituba Y.C.Hsu et G.J.Fan, 1974 ‘Pure White’

(Imported at JAPAN, 1979 from China, By Mr. Kaneko, Japan.)

Very rare plants. 2n=16=6M+10T

 

Endemic to China (Jiangsu and Anhui). Very polymorphic species. Variable in flower color and shape. Plants with yellow flowers classified as a variety, "Lycoris longituba Y.C.Hsu et G.J.Fan, 1974". Very long tepatube and its fragrance are marked chracteristics of this species. Leaves lanceolate, to 63 cm long and to 4 cm wide, somewhat fleshy and pale green, appearing in early spring and wither up in May. Scape, 60-80 cm hight, appearing in July to August. Spathe, to 5 cm long. Pedicel 1.5-4.5 cm, tepaltube 4.5-6.0 cm. Tepals 7-9.5 cm long, . Stamens 6-7.5 cm, shorter than tepals. Style 7-9.5 cm, nearly equal to or slightly exceeding tepals. Before that it was far carrying out this plant in a scientific statement, it was indicated by U.S. an Advanced Horticulturalist Mr. Sam Coldwell from that it was a new species. The formal scientific statement of "Lycoris longituba" was announced by Y.C.Hsu & G.J.Fan in 1974. However, this plant was introduced into Japan and was already grown in the 1930s. It by misconception of scholar Dr. Inariyama (1948) of Japan "Lycoris x straminea Lindl., 1848" said. When it depended on research of scholar Dr. Kurita of Japan, it became clear that the individual of this plants is one of the variations of "Lycoris longituba Y.C.Hsu et G.J.Fan, 1974"

 

この植物は、学術記載をされるはるか以前から、アメリカの先進的園芸家・Mr. Sam Coldwell により新種ではないかと指摘をされていた。「Lycoris longituba」の正式な学術記載は Y.C.Hsu & G.J.Fan によって1974年になされた。しかし、既に1930年代にこの植物は日本に渡来して栽培されていた。日本の学者 Dr. Inariyama(1948)の誤認で「Lycoris × straminea Lindl., 1848」と言われていた。日本の学者 Dr. Kurita の研究に依ると、この画像の個体は「Lycoris longituba Y.C.Hsu et G.J.Fan, 1974」のバリエーションの一つだと判明した。

 

この個体はリコリス中でも最も大型の部類で、主に種子繁殖をし、非常に自家受粉しやすいが、その反面、分球はしずらく、1球が10球までになるには凡そ30年はかかった。ヒガンバナが30年で1000球に分けつする事と比較すると恐ろしいほどスローモーな分けつ力である。此の個体は、金子氏が40年前に中国から導入したヒガンバナ類の中から選抜した個体で、他に麦藁色の個体が明治期にもたらされていたが、在来の物より鑑賞価値は高い。春出葉型なので、短い期間しか葉が無い為、その間には良く日光に当て肥培をすることが肝心で有る。自然交雑種のナツズイセンの片親である事がDr. Kurita の研究で判明した。自然界でのリコリス・ロンギチュバは、白、ピンク、麦藁色など幅があり、黄花の物は別途変種扱いされている。此の個体自体は、蘂にやや色が乗るが、これほどに白い個体は少なく、殆どがやや濁っているのが殆どであり、選抜した意味は大きい。本種は大球性で他のリコリスよりもやや深く土中に球根が潜る。開花は、是から得られたナツズイセンよりも1ヶ月早く、7月中には開花する。

  

SONY NEX-7

Nikon Ai AF Micro Nikkor ED 200mm F4D (IF)

(Product Year : 1993)

apparently butterflies with polymorphic female forms are called "mormons", as a joke on having more than one wife.

Pictured are the atomic-resolution structures of three amyloid polymorphs against a (falsely coloured) background image of the fibrils taken with a transmission electron microscope. Determining the fibril structures, and defining the major structural elements and interactions contributing to their hierarchical self-assembly, provides insight into the formation of polymorphic amyloid in a range of protein deposition disorders including Alzheimer’s and Parkinson’s diseases. Image courtesy of Anthony W. P. Fitzpatrick, Christopher A. Waudby, Daniel K. Clare, Michele Vendruscolo and Christopher M. Dobson.

 

Courtesy of Dr. Anthony Fitzpatrick

 

Image Details

Instrument used: Tecnai

 

Mais uma vez, muito obrigado ao Marcelo Cazani (Marcazani) pela identificação deste pássaro.

Once more, thank you very much Marcelo Cazani (Marcazani) for the identification of this bird.

 

A text, in english, from Wikipedia, the free encyclopedia:

See at en.wikipedia.org/wiki/American_Kestrel

Jump to: navigation, search

American Kestrel

Conservation status

 

Least Concern (IUCN 3.1)

Scientific classification

Kingdom: Animalia

Phylum: Chordata

Class: Aves

Order: Falconiformes

Family: Falconidae

Genus: Falco

Species: F. sparverius

Binomial name

Falco sparverius

Linnaeus, 1758

Synonyms

 

Cerchneis sparverius

Tinnunculus sparverius

 

The American Kestrel (Falco sparverius) is a small falcon. This bird was (and sometimes still is) colloquially known in North America as the "Sparrow Hawk". This name is misleading because it implies a connection with the Eurasian Sparrowhawk Accipiter nisus, which is unrelated; the latter is an accipiter rather than a falcon. Though both are diurnal raptors, they are only distantly related.

American Kestrels are widely distributed across the Americas. Their breeding range extends from central and western Alaska across northern Canada to Nova Scotia, and south throughout North America, into central Mexico, the Baja, and the Caribbean. They are local breeders in Central America and are widely distributed throughout South America.

Most of the birds breeding in Canada and the northern United States migrate south in the winter, although some males stay as year-round residents. It is a very rare vagrant to western Europe.

The American Kestrel is the smallest falcon in North America—about the size of an American Robin. Like all raptors, the American Kestrel is sexually dimorphic, although there is some overlap within the species. The female ranges in length from 23 to 28 centimeters (9-11 inches) with a wingspan of 53–61 centimeters (21–24 inches) and weighs an average of 120 grams (4.2 ounces). The length of the male varies between 20–25 centimeters (8–10 inches) with a wingspan ranging from 51–56 centimeters (20–22 inches) and weighing an average of 111 grams (3.9 ounces). These subtle differences are often difficult to discern in the field.

The coloration of the feathers, however, greatly varies between the sexes. Males have blue-grey secondary feathers on their wings, while the undersides are white with black barring. The back is rufous in coloration, with barring on the lower half. The belly and flanks are white with black spotting. The tail is also rufous, except for the outer rectrix set, which is white with a black subterminal band.

The back of the female American Kestrel is rufous with dark brown barring. The wings exhibit similar coloration and patterning to the back. The undersides of the females are white with rufous streaking. The tail of the female is noticeably different from the male, being rufous in color with numerous narrow dark brown or black bars. Juveniles exhibit coloration patterns similar to the adults.

In both sexes, the head is white with a bluish-grey top. There are also two narrow, vertical black facial markings on each side of the head; one below the eyes and one on the rear portion of the auriculars. Two black spots (ocelli) can be found on each side of the white or orangish nape. The wings are moderately long, fairly narrow, and taper to a point. While perched, the wingtips are noticeably shorter than the tail tip.

The American Kestrel has three basic vocalizations - the "klee" or "killy", the "whine", and the "chitter." The "klee" or "killy" is usually delivered as a rapid series - "killy, killy, killy, killy" when the kestrel is upset or excited. It is used at the apex of the dive display, during fights with other kestrels, and after unsuccessful hunting attempts.

The "whine" may last as long as one to two minutes and may be single or treble. The more intense the situation, the more likely the "whine" will move toward the treble extreme. "Whines" are given during courtship feeding and copulation. The treble whine is heard from breeding females and fledged hungry young.

The "chitter" is the most frequent vocalization in male - female interactions. Its volume and duration depends upon the stress or excitement of the situation. It is associated with friendly approaches and bodily contact between the sexes during breeding season. Occasionally a "chitter" follows a "whine."

Calling occurs throughout the day. Nestlings at two weeks can produce all three vocalizations. Female kestrels tend to have slightly lower pitched and harsher voices than males.

This bird is apparently not a true kestrel. mtDNA cytochrome b sequence analysis (Groombridge et al. 2002) indicates a Late Miocene split[1] between the ancestors of the American Kestrel, and those of the Common Kestrel and its closest relatives. The color pattern with its large areas of brown is reminiscent of kestrels, but the coloration of the head - notably the black ear patch, which is not found in any of the true kestrels - and the male's extensively gray wings are suggestive of a closer relationship with the hobbies, an informal grouping of falcons of usually average size.

Species such as the Merlin and the Aplomado Falcon are proposed as possible close relatives. Indeed, the Merlin is a highly polymorphic bird and although its grey tail and back are distinctive, certain morphs are the only birds that might conceivably be confused with American Kestrels. Conclusive evidence is lacking, and what can be said at present judging from the fairly noninclusive DNA sequence studies[2] is that the general relationships of the present species seem to lie with a number of rather basal "hobby" lineages, such as the Merlin and Aplomado Falcon mentioned already, or the Red-footed and Amur Falcons - or even the Peregrine Falcon lineage with its large species.

The American Kestrel is not very closely related to any of these groups, although it might be closer to the Aplomado Falcon (and its presumed close relatives, the Bat and Orange-breasted Falcons) than to any other living species (Wink et al. 1998) - an association that is also better supported by biogeography than a close relationship with the exclusively Old World true kestrels. It is nonetheless highly distinct in morphology from any of these and, interestingly, has a syrinx similar to the Peregrine and the hierofalcons[3].

In conclusion, until better evidence is available, it is best considered part of a radiation of falcon lineages that diversified around the North Atlantic at the end of the Miocene. Though several fossils of small falcons arte known from North America at roughly the correct time, the earliest testimony of the American Kestrel lineage is Pleistocene remains of the living species (Brodkorb 1964).

American Kestrels are found in a variety of habitats including parks, suburbs, open fields, forest edges and openings, alpine zones, grasslands, marshes, open areas on mountainsides, prairies, plains, deserts with giant cacti, and freeway and highway corridors.

In addition to requiring open space for hunting, American Kestrels seem to need perches for hunting from, cavities for nesting (either natural or man-made), and a sufficient food supply.

The American Kestrel is the only North American falcon to habitually hover with rapid wing beats, keeping its head motionless while scanning the ground for prey. The kestrel commonly perches along fences and powerlines. It glides with flat wings and wingtips curved upward. It occasionally soars in circles with its tail spread and its wings flat.

This falcon species is not long-lived. The oldest banded wild bird was 11 years and seven months old while a captive lived 17 years. A mortality rate average of 57 percent was found. First year mortality rates have declined since 1945 with a decrease in shooting. Major causes of death include collision with traffic, illegal shooting, and predation by other raptors, including the Red-tailed Hawk, Northern Goshawk, Cooper's Hawk, Peregrine Falcon, Barn Owl, and Great Horned Owl.

In summer, kestrels feed largely on grasshoppers, dragonflies, lizards, mice, and voles. They will also eat other small birds. Wintering birds feed primarily on rodents and birds. The birds characteristically hunt along roadsides from telephone wires, fence posts, trees or other convenient perches when not flying in search of food. When they are flying and looking for food they frequently hover with rapid wingbeats.

Because it feeds on both insects and vertebrates, the American Kestrel maintains fairly high population densities. It has a small breeding home range, from 1.75 square miles (4.5 km²) to 2 square miles (5.2 km²). Territory size has been estimated at 269 acres (1.1 km²) to 321 acres (1.3 km²) with much larger wintering home ranges.

Several hunting techniques are used by the American Kestrel. It will hover over one spot—when prey is sighted the kestrel will partly fold its wings and drop lower once or several times before striking. When the prey disappears the falcon will glide in a semicircle before turning back into the wind to hover again. It will also soar in circles, or figure eights, using the same stooping tactics as when hovering.

The kestrel commonly hunts from elevated perch sites, waiting for prey to move on the ground. The kestrel bobs its head and pumps its tail just before attacking.

Other prey capture techniques include direct pursuit, landing and flushing prey from the ground (especially for grasshoppers)and then taking them in flight, capturing flying insects from an elevated perch, and nest robbing including the burrows of Bank Swallows and the nests of Cliff Swallows. It is also an occasional bat catcher, taking bats from their tree roosts, or striking bats in flight from above or as the bats leave or enter caves. The kestrel will kill and cache food items.

The American Kestrel occasionally robs others of the same species. It has also been known to rob a shrike of its prey. Kestrels sometimes harass other hawks, and even Golden Eagles, in flight.

American Kestrels form pairs in which the bond is strong, tending toward permanence. Returning migrants commonly re-establish territories held the previous year. In one study[citation needed], a pair nested in the same tree for six consecutive years. Nesting occurs from late spring to late summer in North America, with incubation underway by the end of May[4]; in tropical South America the birds breed roughly from June onwards through to September or so[5].

Courtship begins shortly after the male establishes a territory. In early courtship, he may give the "dive display", a series of climbs and dives 33 to 66 feet (10 to 20 m) with 3-5 "klee" notes given near the peak of each climb. He may present the female with food during courtship feeding. He may entice her to the nest site by calling. He may "flutter-glide" toward her with quick and shallow wing beats while carrying food and she may also beg for food by flutter-gliding. The female initiates copulation by bowing with her tail in line with her body or slightly raised.

A cavity nester, American Kestrels will use holes in trees, rock cavities and crevices in cliffs, artificial nest boxes, or small spaces in buildings. The number of suitable breeding cavities limits this species' breeding density. The American Kestrel has adapted well to nest boxes. In one program, nest boxes were fixed to the backs of signs along a freeway thus allowing kestrels to breed in areas formerly devoid of nest sites. Pairs nesting in boxes on poles have much higher nesting success than pairs using boxes on trees. No nest is built inside. In nest boxes sawdust and wood shavings may be a suitable substrate for the eggs. Males and females defend the nest against intruders, with the male maintaining a small core territory and the female defending the nest cavity directly rather the surroundings[5].

Both sexes take turns incubating their eggs, a very rare situation among North American birds of prey where the female usually incubates exclusively. Correspondingly, both sexes develop bare oval patches on each side of their breasts where the warm bare skin can contact the eggs for warming. Eggs hatch 29 to 31 days after being laid.[6] There are from three to seven eggs laid, but four to six are average.[6] The eggs are typically short elliptical in shape, and are white or pinkish-white with an even covering of fine spots and flecks of brown shades, occasionally concentrating as a ring or a cap. They will renest if the first nest fails and have been reported to raise 2 broods per year in some of the southern states.

The young grow very quickly, becoming noisy between day 11 and 14 and assuming adult weight in about 2.5 weeks. The young fledge in 30 to 31 days.[7] [6]Early fledgling behavior varies. Broods typically stay together for a week or two. Some broods remain close to the nest area for a week or two while others travel throughout the parents' home range. Generally, young do not disperse more than 0.6 miles (one km) away from the nest area until two to four weeks old. Young disperse as hunting skills develop. Occasionally groups of older juveniles from various broods join together into flocks.

The American Kestrel can be double-brooded, particularly in the southern United States, in areas of abundant small mammals. Replacement clutches can be laid.

 

Um texto em português:

Falcão-americano ou quiriquiri (Falco sparverius) é um pequeno falcão (23-27 cm de comprimento e 85-140 g de peso). Tem uma ampla área de distribuição: desde o Alasca e Norte do Canadá até à ponta Sul da América do Sul (Terra do Fogo). Tem asas azul acizentadas. O dorso é avermelhado pontuado de preto. A cauda também é avermelhada possuindo uma larga lista preta. Na face possui 2 listas verticais que começam junto aos olhos e seguem para baixo. É um predador de pequeno tamanho, alimentando-se de insetos e microvertrebrados como roedores e pequenos pássaros; ocasionalmente caça morcegos.

Falcão é o nome genérico dado a várias aves da família Falconidae, mais estritamente aos animais classificados dentro do género Falco. O que diferencia os falcões das demais aves de rapina é o fato de terem evoluído no sentido de uma especialização no voo em velocidade (em oposição ao voo planado das águias e abutres e ao voo acrobático dos gaviões), facilitado pelas asas ponteagudas e finas, favorecendo a caça em espaços abertos – daí o fato dos falcões não serem aves de ambientes florestais, preferindo montanhas e penhascos, pradarias, estepes e desertos. Os falcões podem ser identificados, aliás, pelo fato de não planarem em correntes termais, como outras aves de rapina. O falcão-peregrino, especializado na caça de aves médias e grandes em voo, pode atingir 300 km/h em voo picado e é o animal mais rápido da terra. Diferentemente das águias e gaviões, que matam suas presas com os pés, os falcões utilizam as garras apenas para apreenderem a presa, matando-a depois com o bico por desconjuntamento das vértebras, para o que possuem um rebordo em forma de dente na mandíbula superior.

 

Na Idade Média, os falcões eram apreciados como animais de caça acessíveis apenas à elite.