Unit ON-01

Toa:Earth

Safeguard build

Pakari – active, stage 8 (polymorphic)

Tectonic Claws

 

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Decided to try to see if the lights in the kitchen could substitute for daylight in a pinch - results are not great but I think I finally have a cheap idea of how I can muzzle my flash for future pictures.

夢幻蝶影 蝶戀花

 

Dream of BUTTERFLY WITH

 

The Great Mormon (Papilio memnon) is a large butterfly that belongs to the swallowtail family and is found in southern Asia. It is widely distributed and has thirteen subspecies. The female is polymorphic and with mimetic forms.

 

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Thanks to every friend ! Wish you lucky !

Because of personal reasons,

I unable to respond to all friends one by one, apologize !

 

4/10 re

Agave difformis is a plant species of the genus of agave (Agave). An English common name is "Deformed Century Plant".

 

Agave difformis forms open rosettes with plant height of 70 to 100 cm and 100 to 150 cm width, making plenty of offshoots. The arrayed variable, polymorphic, linealischen, stiff, striped, green to yellow-green leaves are 50-80 cm long and 40-60 cm wide. The white to brown, wavy, narrow, horny leaf margins are toothed flexible. The strong brown to gray Enddorn is 1.5 to 3 cm long.

 

The eared, thin, waxy inflorescence is 3 to 5 m high. The flowers are bright green, yellow to pink. They are 30 to 40 mm long and appear at the top of the inflorescence. The flower tube is 2.5 to 3.5 mm long.

 

The variable-shaped, dark brown, dreikammerigen capsule fruits are 22-26 mm long and 12-15 mm wide. Black, irregularly shaped seeds , up to 5 mm long and 3 mm wide.

 

Agave difformis growing endemic in Mexico in the states of San Luis Potosi and Hidalgo in dry regions in limestone soils, on rocky slopes and in grasslands in 1550-1900 m altitude. It is associated with cacti and other succulents.

 

The first description by Alwin Berger was published in 1915.

 

Agave difformis is a member of the group Marginatae. The robust nature is characterized by the polymorphic, long sword-shaped leaves. It forms dark green median strip on both leaf surfaces. Agave difformis is in the Huntington Botanical Gardens in San Marino cultivated in California.

 

translate.google.co.uk/translate?hl=en&sl=de&u=ht...

La formica rossa (Formica rufa Linnaeus, 1758) è una formica diffusa nell'emisfero boreale. Dotata di livrea rosso ruggine, addome, zampe ed antenne di colorazione bruna, questa specie è priva di pungiglione, ma è capace di lanciare potenti getti di acido formico prodotto da un apparato addominale anche a 30 cm di distanza. Lunga fino a 8-10 millimetri, è dotata di grandi mandibole. La formica si nutre comunemente di invertebrati trovati intorno al nido e in particolare di afidi trovati sugli alberi, sebbene sia anche un vorace saprofago. Diffusa nei boschi europei di conifere, soprattutto di abeti e larici, costruisce nidi giganteschi, alti sino a 2 metri, che vengono trapiantati anche in altri boschi per la lotta contro gli insetti dannosi; si rinvengono in genere nelle aree di bosco meno fitto, dove possono essere raggiunti dai raggi del sole.

 

Formica rufa, also known as the red wood ant, southern wood ant, or horse ant, is a boreal member of the Formica rufa group of ants, and is the type species for that group. It is native to Europe and Anatolia but is also found in North America, in both coniferous and broad-leaf broken woodland and parkland. Workers are bicolored red and brownish-black, with a dorsal dark patch on the head and promensonotum, and are polymorphic, measuring 4.5–9 mm in length. They have large mandibles and like many other ant species they are able to spray formic acid from their abdomens as a defence. Formic acid was first extracted in 1671 by the English naturalist John Ray by distilling a large number of crushed ants of this species.

The ant's primary diet is aphid honeydew, but they also prey on invertebrates such as insects and arachnids; they are voracious scavengers. Foraging trails may extend 100 m. Larger workers have been observed to forage further away from the nest. F. rufa is commonly used in forestry and is often introduced into an area as a form of pest management.

 

Le cocciniglie o, impropriamente, coccidi (Coccoidea Handlirsch, 1903), sono una superfamiglia di insetti fitofagi compresi nell'ordine dei Rhynchota (sottordine Homoptera, sezione Sternorrhyncha). Il nome cocciniglia deriva dallo spagnolo cochinilla ("porcellino di terra"). Sono insetti esclusivamente fitomizi e costituiscono uno tra i più importanti raggruppamenti di insetti dannosi. La caratteristica generale che contraddistingue questi insetti è il marcato dimorfismo sessuale e la regressione morfologica, anatomica e funzionale delle femmine (neotenia). Il maschio ha un corpo di piccolissime dimensioni, esile e allungato, con le regioni corporee ben conformate. La femmina presenta particolari adattamenti morfologici, anatomici e funzionali che la rendono inconfondibile, per quanto un profano potrebbe non riconoscere in essa un insetto. La regressione che caratterizza le femmine è tale che fino ad alcuni secoli fa si riteneva si trattasse di galle vegetali. Apparentemente si potrebbe pensare ad una forma di vita poco evoluta, in realtà si tratta di un adattamento spinto al regime dietetico fitofago che rientra nei casi di neotenia. La caratteristica morfologica più evidente delle femmine adulte è la protezione formata dalle secrezioni esterne. Da un punto di vista chimico le secrezioni sono di varia natura e sono riconducibili alla cera, alla lacca, alla seta. Ogni gruppo sistematico produce uno specifico secreto che si dispone e si accumula in modo caratteristico al punto che, spesso, la forma e l'aspetto della secrezione sono fondamentali per determinare le cocciniglie a livello di famiglia, sottofamiglia o specie. Le secrezioni esterne sono anche il prodotto del metabolismo glucidico di questi insetti. Le cocciniglie si riproducono per via sessuale, ma non sono rari i casi di partenogenesi; in questo raggruppamento è presente anche l'unico caso di ermafroditismo riscontrato fra gli insetti (Icerya purchasi). Data la differente etologia nei due sessi, le femmine emettono feromoni sessuali per attirare i maschi. L'anatomia e la fisiologia dell'apparato digerente delle cocciniglie è l'espressione del forte adattamento alla fitofagia che presentano questi insetti e, più in generale, tutti gli Sternorrinchi (Afidi, Aleurodidi, Psille e Cocciniglie). La dieta di questi insetti ha un elevato rapporto carbonio/azoto, manca di alcuni componenti essenziali e la sostanziale sedentarietà che caratterizza la loro etologia non permette l'integrazione del fabbisogno nutritivo con l'assunzione di altri alimenti. Per soddisfare i fabbisogni proteici gli Sternorrinchi devono assumere grandi quantità di linfa (o di succhi vegetali in generale) e smaltire l'eccesso d'acqua e zuccheri con la produzione della melata. Questa funzione è svolta dalla camera filtrante, un vero e proprio organo dializzatore che effettua il by-pass dei glucidi. Le cocciniglie rappresentano uno dei raggruppamenti sistematici più dannosi all'agricoltura. Nonostante siano insetti piuttosto piccoli e dotati di un potenziale riproduttivo sostanzialmente inferiore a quello di altri fitofagi come gli Afidi, i Ditteri Tefritidi o gli Acari, diverse specie di cocciniglie possono facilmente essere responsabili di infestazioni di gravità tale da causare perdite considerevoli di prodotto.

Photographed at Los Altos, California - Sitting, no cover

 

=> Please click on the image to see the largest size. <=

 

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From Wikipedia: The lesser goldfinch (Spinus psaltria) is a very small songbird of the Americas. Together with its relatives the American goldfinch and Lawrence's goldfinch, it forms the American goldfinches clade in the genus Spinus sensu stricto.

 

The American goldfinches can be distinguished by the males having a black (rarely green) forehead, whereas the latter is (like the rest of the face) red or yellow in the European goldfinch and its relatives. North American males are markedly polymorphic and five subspecies are often named; at least two of them seem to represent a less-progressed stage in evolution, however.

 

This petite species is not only the smallest North American Spinus finch, it may be the smallest true finch in the world. Some sources list more subtropical Spinus species as slightly smaller on average, including the Andean siskin.[4] This species ranges from 9 to 12 cm (3.5 to 4.7 in) in length and can weigh from 8 to 11.5 g (0.28 to 0.41 oz). Among standard measurements, the wing chord is 5.5 to 7 cm (2.2 to 2.8 in), the tail is 3.9 to 4.7 cm (1.5 to 1.9 in), the bill is 0.9 to 1.1 cm (0.35 to 0.43 in) and the tarsus is 1.1 to 1.2 cm (0.43 to 0.47 in). There is a slight NW-SE cline in size, with the largest birds from Mexico and further south being up to one-fifth larger than the smallest from the extreme northwest of its range; this effect is more pronounced in females. There is also considerable variation in the amount of black on head and back in males, and thus three subspecies have been proposed. But this variation, too, seems to be simple and clinal changes in allele frequency, and thus the "subspecies" might be better considered morphs or geographic forms.

 

Males are easily recognized by their bright yellow underparts and big white patches in the tail (outer rectrices) and on the wings (the base of the primaries). They range from having solid black from the back to the upper head including the ear-coverts to having these regions medium green; each of the back, crown and ear regions varies in darkness rather independently though, as a rule, the ears are not darker than the rest. In most of the range, dark psaltria birds (Arkansas goldfinch) predominate. The light birds are termed hesperophilus (green-backed goldfinch) and are most common in the far western U.S. and northwestern Mexico.

 

Females' and immatures' upperparts are more or less grayish olive-green; their underparts are yellowish, buffier in immatures. They have only a narrow strip of white on the wings (with other white markings in some forms) and little or no white on the tail. They are best distinguished from other members of the genus by the combination of small size, upperparts without white or yellow, and dark gray bill. In all plumages, this bird can easily be taken for a New World warbler if the typical finch bill is not seen well.

 

Like other goldfinches, it has an undulating flight in which it frequently gives a call: in this case, a harsh chig chig chig. Another distinctive call is a very high-pitched, drawn-out whistle, often rising from one level pitch to another (teeeyeee) or falling (teeeyooo). The song is a prolonged warble or twitter, more phrased than that of the American goldfinch, often incorporating imitations of other species.

 

Distribution and ecology:

Lesser Goldfinch Landing, Santa Fe.jpg

This American goldfinch ranges from the southwestern United States (near the coast, as far north as extreme southwestern Washington) to Venezuela and Peru. It migrates from the colder parts of its U.S. range.

 

The lesser goldfinch often occurs in flocks or at least loose associations. It utilizes almost any habitat with trees or shrubs except for dense forest, and is common and conspicuous in many areas, often coming near houses. It is common at feeders in the Southwest United States and will come almost anywhere with thistle sock feeders. Flocks of at least six birds will often be seen at feeders. It feeds mostly on tree buds and weed seeds; geophagy has been observed in this species.

 

The nesting season is in summer in the temperate parts of its range; in the tropics it apparently breeds all-year round, perhaps less often in September/October. It lays three or four bluish white eggs in a cup nest made of fine plant materials such as lichens, rootlets, and strips of bark, placed in a bush or at low or middle levels in a tree.

 

6D6A4859-2CAFlkr

Being a Mediterranean myself I use to spend a lot of free time on a seashore.Recently I've met this fairy creature who guard the sea life using his polymorphic abilities.This phantom is against greedy fishermen !

Sophora chrysophylla, known as Māmane in Hawaiian, is a species of flowering plant in the pea and bean family, Fabaceae, that is endemic to Hawaii. It is highly polymorphic, growing as a shrub or tree, and able to reach a height of 15 m (49 ft) in tree form. Yellow flowers are produced in winter and spring.

Sophora is a genus of about 45 species of small trees and shrubs in the pea family Fabaceae. The species are native to southeast Europe, southern Asia, Australasia, various Pacific islands, western South America, the western United States, Florida and Puerto Rico. The generic name is derived from sophera, an Arabic name for a pea-flowered tree.

The New Zealand Sophora species are known as Kowhai.

 

リコリス・ロンギチュバ ‘ピュア・ホワイト’

Lycoris longituba Y.C.Hsu et G.J.Fan, 1974 ‘Pure White’

(Imported at JAPAN, 1979 from China, By Mr. Kaneko, Japan.)

Very rare plants. 2n=16=6M+10T

 

Endemic to China (Jiangsu and Anhui). Very polymorphic species. Variable in flower color and shape. Plants with yellow flowers classified as a variety, "Lycoris longituba Y.C.Hsu et G.J.Fan, 1974". Very long tepatube and its fragrance are marked chracteristics of this species. Leaves lanceolate, to 63 cm long and to 4 cm wide, somewhat fleshy and pale green, appearing in early spring and wither up in May. Scape, 60-80 cm hight, appearing in July to August. Spathe, to 5 cm long. Pedicel 1.5-4.5 cm, tepaltube 4.5-6.0 cm. Tepals 7-9.5 cm long, . Stamens 6-7.5 cm, shorter than tepals. Style 7-9.5 cm, nearly equal to or slightly exceeding tepals. Before that it was far carrying out this plant in a scientific statement, it was indicated by U.S. an Advanced Horticulturalist Mr. Sam Coldwell from that it was a new species. The formal scientific statement of "Lycoris longituba" was announced by Y.C.Hsu & G.J.Fan in 1974. However, this plant was introduced into Japan and was already grown in the 1930s. It by misconception of scholar Dr. Inariyama (1948) of Japan "Lycoris x straminea Lindl., 1848" said. When it depended on research of scholar Dr. Kurita of Japan, it became clear that the individual of this plants is one of the variations of "Lycoris longituba Y.C.Hsu et G.J.Fan, 1974"

 

この植物は、学術記載をされるはるか以前から、アメリカの先進的園芸家・Mr. Sam Coldwell により新種ではないかと指摘をされていた。「Lycoris longituba」の正式な学術記載は Y.C.Hsu & G.J.Fan によって1974年になされた。しかし、既に1930年代にこの植物は日本に渡来して栽培されていた。日本の学者 Dr. Inariyama(1948)の誤認で「Lycoris × straminea Lindl., 1848」と言われていた。日本の学者 Dr. Kurita の研究に依ると、この画像の個体は「Lycoris longituba Y.C.Hsu et G.J.Fan, 1974」のバリエーションの一つだと判明した。

 

この個体はリコリス中でも最も大型の部類で、主に種子繁殖をし、非常に自家受粉しやすいが、その反面、分球はしずらく、1球が10球までになるには凡そ30年はかかった。ヒガンバナが30年で1000球に分けつする事と比較すると恐ろしいほどスローモーな分けつ力である。此の個体は、金子氏が40年前に中国から導入したヒガンバナ類の中から選抜した個体で、他に麦藁色の個体が明治期にもたらされていたが、在来の物より鑑賞価値は高い。春出葉型なので、短い期間しか葉が無い為、その間には良く日光に当て肥培をすることが肝心で有る。自然交雑種のナツズイセンの片親である事がDr. Kurita の研究で判明した。自然界でのリコリス・ロンギチュバは、白、ピンク、麦藁色など幅があり、黄花の物は別途変種扱いされている。此の個体自体は、蘂にやや色が乗るが、これほどに白い個体は少なく、殆どがやや濁っているのが殆どであり、選抜した意味は大きい。本種は大球性で他のリコリスよりもやや深く土中に球根が潜る。開花は、是から得られたナツズイセンよりも1ヶ月早く、7月中には開花する。

  

SONY NEX-7

Canon New FD Macro 100mm F4 (Product Year : 1979)

Unit GA-01

Toa:Water

Safeguard build

Kaukau – active, stage 8 (polymorphic

Ocean Hooks

 

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Decided to try to see if the lights in the kitchen could substitute for daylight in a pinch - results are not great but I think I finally have a cheap idea of how I can muzzle my flash for future pictures.

In computer science, polymorphism is a programming language feature that allows values of different data types to be handled using a uniform interface. The concept of parametric polymorphism applies to both data types and functions. A function that can evaluate to or be applied to values of different types is known as a polymorphic function. A data type that can appear to be of a generalized type (e.g., a list with elements of arbitrary type) is designated polymorphic data type like the generalized type from which such specializations are made.

 

Source :Wikipedia

  

Innenpark. Bonifatius-Felsen. Felsen am Bonifatiusweg,

Inner Park. Boniface Rock. Rock on the Boniface-Way

 

Asplenium trichomanes subsp. pachyrachis (Christ) Lovis et Reichst.

Willdenowia 10: 18. 1980.

 

Asplenium trichomanes sublus. pachyrachis Christ

Farnkr. Schweiz 1 (2): 92. 1900.

Asplenium csikii Kümmerle & András.

Magyar Bot. Lapok 17. 110. 1919 (nomen), 20. 3, fig. 1923

Asplenium pachyrhachis Landolt

Fl. Indicativa 268 2010.

 

Seestern-Braunschwarz-Streifenfarn, Dickstieliger Braunstieliger Streifenfarn, Dickstieliger Brauner Streifenfarn

Lobed Maidenhair Spleenwort

 

Anmerkung:

Die Asplenium trichomanes Gruppe, ist ein sehr polymorpher, taxonomisch kritischer Spezieskomplex! Die Evolutionsgeschichte und Beziehungen zwischen den Taxa in dieser Gruppe wurden intensiv untersucht. Allerdings sind morphologische Variation und die Verteilung dieser Taxa unzureichend bekannt, da sie in der lokalen Floren oder Checklisten häufig nicht vorkommen. Die Gründe für die Vernachlässigung der Taxa innerhalb der Asplenium trichomanes Gruppe sind der Mangel an diagnostischen morphologische Merkmalen, das häufige gemeinsame Auftreten an ihren Standorten, sowie die Hybridisierung unter den Taxa. Die Asplenium trichomanes Gruppe umfasst zytologisch und ökologisch unterschiedliche Taxa mit fast weltweiter Verbreitung, die offenbar noch aktiv in der Entwicklung sind (L. Ekrt & M. Štech, 2008).

 

Annotation:

The Asplenium trichomanes group is a very polymorphic, taxonomically critical species complex! The evolutionary history and relationships between the taxa in this group have been extensively studied. However, morphological variation and distribution of these taxa are poorly understood because they are often absent in local floras or checklists. The reasons for the neglect of the taxa within the Asplenium trichomanes group are the lack of diagnostic morphological features, the frequent common occurrence at their sites, as well as the hybridization among the taxa. The Asplenium trichomanes group includes cytologically and ecologically diverse taxa with almost worldwide distribution, which apparently are still active in development (L. Ekrt & M. Štech, 2008).

কালিম । Common Mormon (Female; Form: romulus) - Papilio polytes

 

This female form of the Common Mormon mimics the Common Rose very closely but lacks red markings on body. This is the commonest form wherever the Common Rose flies. The female of the Common Mormon is polymorphic. In South Asia, it has three forms or morphs. These are as follows: cyrus, stichius, romulus.

 

The Common Mormon (Papilio polytes) is a common species of swallowtail butterfly (family: papilionidae) widely distributed across Asia. Seen round the year throughout India from plains up to 2000m. This butterfly is known for the mimicry displayed by the numerous forms of its females.

 

Host Plant: Putush, পুটুশ ( Lantana camara "Feston Rose")

Evergreen shrubs and perennial herbs with green leaves and spiny branches that bear globular flowers, intermittently year round. Flowerheads with yellow flowers in the center and fuchsia to purple flowers at the circumference. Poisonous plants though leaves are in traditional medicinal use.

 

Lantana camara is an invasive species and has covered large areas in India, Australia and much of Africa. It is believed to be indigenous to the Lower Rio Grande Valley of Texas in the United States.

 

Family: Papilionidae

 

Damodor River Bank, Durgapur

Monsoon Images of Bengal, India

 

Sigma 150-600mm 5-6.3 DG HSM OS Contemporary plus Nikon D7000

"The tiny LCD screens in this interactive piece portray scenes of urban life. The screens react to movement, switching from restful scenes of nature to images of uncontrolled urban sprawl. The artist is intrigued by how urban environments shape our consciousness. She highlights India’s mega-metropoles, where uncontrolled urbanization is exceeding sustainable limits, resulting in slums and impacting negatively on our wellbeing."

 

arsguide.kiasma.fi/en/artwork/nandita-kumar/

 

Common Mormon (Papilio polytes) - Male

 

A common species of swallowtail butterfly (family: papilionidae) widely distributed across Asia. Seen round the year throughout India from plains up to 2000m. This butterfly is known for the mimicry displayed by the numerous polymorphic forms of its females. These are as follows: cyrus, stichius, romulus.

 

This picture is taken in the indoor live butterfly garden of the Sensational Butterflies Exhibition, Natural History Museum, London

  

Maize (/meɪz/ MAYZ; Zea mays subsp. mays, from Spanish: maíz after Taino: mahiz), also known as corn (North American and Australian English), is a cereal grain first domesticated by indigenous peoples in southern Mexico about 10,000 years ago. The leafy stalk of the plant produces pollen inflorescences and separate ovuliferous inflorescences called ears that yield kernels or seeds, which are fruits.

 

Maize has become a staple food in many parts of the world, with the total production of maize surpassing that of wheat or rice. In addition to being consumed directly by humans (often in the form of masa), maize is also used for corn ethanol, animal feed and other maize products, such as corn starch and corn syrup. The six major types of maize are dent corn, flint corn, pod corn, popcorn, flour corn, and sweet corn. Sugar-rich varieties called sweet corn are usually grown for human consumption as kernels, while field corn varieties are used for animal feed, various corn-based human food uses (including grinding into cornmeal or masa, pressing into corn oil, and fermentation and distillation into alcoholic beverages like bourbon whiskey), and as chemical feedstocks. Maize is also used in making ethanol and other biofuels.

 

Maize is widely cultivated throughout the world, and a greater weight of maize is produced each year than any other grain. In 2014, total world production was 1.04 billion tonnes. Maize is the most widely grown grain crop throughout the Americas, with 361 million metric tons grown in the United States alone in 2014. Genetically modified maize made up 85% of the maize planted in the United States in 2009. Subsidies in the United States help to account for its high level of cultivation of maize and its position as the largest producer in the world.

 

HISTORY

PRE-COLUMBIAN DEVELOPMENT

Maize is a cultigen; human intervention is required for it to propagate. Whether or not the kernels fall off the cob on their own is a key piece of evidence used in archaeology to distinguish domesticated maize from its naturally-propagating teosinte ancestor. Genetic evidence can also be used to determine when various lineages split.

 

Most historians believe maize was domesticated in the Tehuacán Valley of Mexico. Recent research in the early 21st century has modified this view somewhat; scholars now indicate the adjacent Balsas River Valley of south-central Mexico as the center of domestication.

 

An influential 2002 study by Matsuoka et al. has demonstrated that, rather than the multiple independent domestications model, all maize arose from a single domestication in southern Mexico about 9,000 years ago. The study also demonstrated that the oldest surviving maize types are those of the Mexican highlands. Later, maize spread from this region over the Americas along two major paths. This is consistent with a model based on the archaeological record suggesting that maize diversified in the highlands of Mexico before spreading to the lowlands.

 

Archaeologist Dolores Piperno has said:

 

A large corpus of data indicates that [maize] was dispersed into lower Central America by 7600 BP [5600 BC] and had moved into the inter-Andean valleys of Colombia between 7000 and 6000 BP [5000–4000 BC].

— Dolores Piperno, The Origins of Plant Cultivation and Domestication in the New World Tropics: Patterns, Process, and New Developments

 

Since then, even earlier dates have been published.

 

According to a genetic study by Embrapa, corn cultivation was introduced in South America from Mexico, in two great waves: the first, more than 6000 years ago, spread through the Andes. Evidence of cultivation in Peru has been found dating to about 6700 years ago. The second wave, about 2000 years ago, through the lowlands of South America.

 

The earliest maize plants grew only small, 25-millimetre-long (1 in) corn cobs, and only one per plant. In Jackson Spielvogel's view, many centuries of artificial selection (rather than the current view that maize was exploited by interplanting with teosinte) by the indigenous people of the Americas resulted in the development of maize plants capable of growing several cobs per plant, which were usually several centimetres/inches long each. The Olmec and Maya cultivated maize in numerous varieties throughout Mesoamerica; they cooked, ground and processed it through nixtamalization. It was believed that beginning about 2500 BC, the crop spread through much of the Americas. Research of the 21st century has established even earlier dates. The region developed a trade network based on surplus and varieties of maize crops.

 

Mapuches of south-central Chile cultivated maize along with quinoa and potatoes in pre-Hispanic times; however, potato was the staple food of most Mapuches, "specially in the southern and coastal [Mapuche] territories where maize did not reach maturity". Before the expansion of the Inca Empire maize was traded and transported as far south as 40°19' S in Melinquina, Lácar Department. In that location maize remains were found inside pottery dated to 730 ± 80 BP and 920 ± 60 BP. Probably this maize was brought across the Andes from Chile. The presence of maize in Guaitecas Archipelago (43°55' S), the southernmost outpost of pre-Hispanic agriculture, is reported by early Spanish explorers. However the Spanish may have misidentified the plant.

 

COLUMBIAN EXCHANGE

After the arrival of Europeans in 1492, Spanish settlers consumed maize, and explorers and traders carried it back to Europe and introduced it to other countries. Spanish settlers far preferred wheat bread to maize, cassava, or potatoes. Maize flour could not be substituted for wheat for communion bread, since in Christian belief only wheat could undergo transubstantiation and be transformed into the body of Christ. Some Spaniards worried that by eating indigenous foods, which they did not consider nutritious, they would weaken and risk turning into Indians. "In the view of Europeans, it was the food they ate, even more than the environment in which they lived, that gave Amerindians and Spaniards both their distinctive physical characteristics and their characteristic personalities." Despite these worries, Spaniards did consume maize. Archeological evidence from Florida sites indicate they cultivated it as well.

 

Maize spread to the rest of the world because of its ability to grow in diverse climates. It was cultivated in Spain just a few decades after Columbus's voyages and then spread to Italy, West Africa and elsewhere. Widespread cultivation most likely began in southern Spain in 1525, after which it quickly spread to the rest of the Spanish Empire including its territories in Italy (and, from there, to other Italian states). Maize had many advantages over wheat and barley; it yielded two and a half times the food energy per unit cultivated area, could be harvested in successive years from the same plot of land, and grew in wildly varying altitudes and climates, from relatively dry regions with only 250 mm (10 in) of annual rainfall to damp regions with over 5,000 mm (200 in). By the 17th century it was a common peasant food in Southwestern Europe, including Portugal, Spain, southern France, and Italy. By the 18th century, it was the chief food of the southern French and Italian peasantry, especially in the form of polenta in Italy.

Names

 

The word maize derives from the Spanish form of the indigenous Taíno word for the plant, mahiz. It is known by other names around the world.

 

The word "corn" outside the US, Canada, Australia, and New Zealand refers to any cereal crop, its meaning understood to vary geographically to refer to the local staple. In the United States, Canada, Australia, and New Zealand, corn primarily means maize; this usage started as a shortening of "Indian corn". "Indian corn" primarily means maize (the staple grain of indigenous Americans), but can refer more specifically to multicolored "flint corn" used for decoration.

 

In places outside the US, Canada, Australia, and New Zealand, corn often refers to maize in culinary contexts. The narrower meaning is usually indicated by some additional word, as in sweet corn, sweetcorn, corn on the cob, baby corn, the puffed confection known as popcorn and the breakfast cereal known as corn flakes.

 

In Southern Africa, maize is commonly called mielie (Afrikaans) or mealie (English), words derived from the Portuguese word for maize, milho.

 

Maize is preferred in formal, scientific, and international usage because it refers specifically to this one grain, unlike corn, which has a complex variety of meanings that vary by context and geographic region. Maize is used by agricultural bodies and research institutes such as the FAO and CSIRO. National agricultural and industry associations often include the word maize in their name even in English-speaking countries where the local, informal word is something other than maize; for example, the Maize Association of Australia, the Indian Maize Development Association, the Kenya Maize Consortium and Maize Breeders Network, the National Maize Association of Nigeria, the Zimbabwe Seed Maize Association.

 

STRUCTURE AND PHYSIOLOGY

The maize plant is often 3 m (10 ft) in height, though some natural strains can grow 13 m (43 ft). The stem is commonly composed of 20 internodes of 18 cm (7 in) length. The leaves arise from the nodes, alternately on opposite sides on the stalk. A leaf, which grows from each node, is generally 9 cm (3+1⁄2 in) in width and 120 cm (3 ft 11 in) in length.

 

Ears develop above a few of the leaves in the midsection of the plant, between the stem and leaf sheath, elongating by around 3 mm (1⁄8 in) per day, to a length of 18 cm (7 in) with 60 cm (24 in) being the maximum alleged in the subspecies. They are female inflorescences, tightly enveloped by several layers of ear leaves commonly called husks. Certain varieties of maize have been bred to produce many additional developed ears. These are the source of the "baby corn" used as a vegetable in Asian cuisine.

 

The apex of the stem ends in the tassel, an inflorescence of male flowers. When the tassel is mature and conditions are suitably warm and dry, anthers on the tassel dehisce and release pollen. Maize pollen is anemophilous (dispersed by wind), and because of its large settling velocity, most pollen falls within a few meters of the tassel.

 

Elongated stigmas, called silks, emerge from the whorl of husk leaves at the end of the ear. They are often pale yellow and 18 cm (7 in) in length, like tufts of hair in appearance. At the end of each is a carpel, which may develop into a "kernel" if fertilized by a pollen grain. The pericarp of the fruit is fused with the seed coat referred to as "caryopsis", typical of the grasses, and the entire kernel is often referred to as the "seed". The cob is close to a multiple fruit in structure, except that the individual fruits (the kernels) never fuse into a single mass. The grains are about the size of peas, and adhere in regular rows around a white, pithy substance, which forms the ear. The maximum size of kernels is reputedly 2.5 cm (1 in). An ear commonly holds 600 kernels. They are of various colors: blackish, bluish-gray, purple, green, red, white and yellow. When ground into flour, maize yields more flour with much less bran than wheat does. It lacks the protein gluten of wheat and, therefore, makes baked goods with poor rising capability. A genetic variant that accumulates more sugar and less starch in the ear is consumed as a vegetable and is called sweet corn. Young ears can be consumed raw, with the cob and silk, but as the plant matures (usually during the summer months), the cob becomes tougher and the silk dries to inedibility. By the end of the growing season, the kernels dry out and become difficult to chew without cooking them tender first in boiling water.

 

Planting density affects multiple aspects of maize. Modern farming techniques in developed countries usually rely on dense planting, which produces one ear per stalk. Stands of silage maize are yet denser,[citation needed] and achieve a lower percentage of ears and more plant matter.

 

Maize is a facultative short-day plant and flowers in a certain number of growing degree days > 10 °C (50 °F) in the environment to which it is adapted. The magnitude of the influence that long nights have on the number of days that must pass before maize flowers is genetically prescribed and regulated by the phytochrome system.

Photoperiodicity can be eccentric in tropical cultivars such that the long days characteristic of higher latitudes allow the plants to grow so tall that they do not have enough time to produce seed before being killed by frost. These attributes, however, may prove useful in using tropical maize for biofuels.

 

Immature maize shoots accumulate a powerful antibiotic substance, 2,4-dihydroxy-7-methoxy-1,4-benzoxazin-3-one (DIMBOA). DIMBOA is a member of a group of hydroxamic acids (also known as benzoxazinoids) that serve as a natural defense against a wide range of pests, including insects, pathogenic fungi and bacteria. DIMBOA is also found in related grasses, particularly wheat. A maize mutant (bx) lacking DIMBOA is highly susceptible to attack by aphids and fungi. DIMBOA is also responsible for the relative resistance of immature maize to the European corn borer (family Crambidae). As maize matures, DIMBOA levels and resistance to the corn borer decline.

 

Because of its shallow roots, maize is susceptible to droughts, intolerant of nutrient-deficient soils, and prone to be uprooted by severe winds.

 

While yellow maizes derive their color from lutein and zeaxanthin, in red-colored maizes, the kernel coloration is due to anthocyanins and phlobaphenes. These latter substances are synthesized in the flavonoids synthetic pathway from polymerization of flavan-4-ols by the expression of maize pericarp color1 (p1) gene which encodes an R2R3 myb-like transcriptional activator of the A1 gene encoding for the dihydroflavonol 4-reductase (reducing dihydroflavonols into flavan-4-ols) while another gene (Suppressor of Pericarp Pigmentation 1 or SPP1) acts as a suppressor. The p1 gene encodes an Myb-homologous transcriptional activator of genes required for biosynthesis of red phlobaphene pigments, while the P1-wr allele specifies colorless kernel pericarp and red cobs, and unstable factor for orange1 (Ufo1) modifies P1-wr expression to confer pigmentation in kernel pericarp, as well as vegetative tissues, which normally do not accumulate significant amounts of phlobaphene pigments. The maize P gene encodes a Myb homolog that recognizes the sequence CCT/AACC, in sharp contrast with the C/TAACGG bound by vertebrate Myb proteins.

 

The ear leaf is the leaf most closely associated with a particular developing ear. This leaf and above contribute 70% to 75% to 90% of grain fill. Therefore fungicide application is most important in that region in most disease environments.

 

ABNORMAL FLOWERS

Maize flowers may sometimes exhibit mutations that lead to the formation of female flowers in the tassel. These mutations, ts4 and Ts6, prohibit the development of the stamen while simultaneously promoting pistil development. This may cause inflorescences containing both male and female flowers, or hermaphrodite flowers.

 

GENETICS

Maize is an annual grass in the family Gramineae, which includes such plants as wheat, rye, barley, rice, sorghum, and sugarcane. There are two major species of the genus Zea (out of six total): Zea mays (maize) and Zea diploperennis, which is a perennial type of teosinte. The annual teosinte variety called Zea mays mexicana is the closest botanical relative to maize. It still grows in the wild as an annual in Mexico and Guatemala.

 

Many forms of maize are used for food, sometimes classified as various subspecies related to the amount of starch each has:

 

Flour corn: Zea mays var. amylacea

Popcorn: Zea mays var. everta

Dent corn : Zea mays var. indentata

Flint corn: Zea mays var. indurata

Sweet corn: Zea mays var. saccharata and Zea mays var. rugosa

Waxy corn: Zea mays var. ceratina

Amylomaize: Zea mays

Pod corn: Zea mays var. tunicata Larrañaga ex A. St. Hil.

Striped maize: Zea mays var. japonica

 

This system has been replaced (though not entirely displaced) over the last 60 years by multivariable classifications based on ever more data. Agronomic data were supplemented by botanical traits for a robust initial classification, then genetic, cytological, protein and DNA evidence was added. Now, the categories are forms (little used), races, racial complexes, and recently branches.

 

Maize is a diploid with 20 chromosomes (n=10). The combined length of the chromosomes is 1500 cM. Some of the maize chromosomes have what are known as "chromosomal knobs": highly repetitive heterochromatic domains that stain darkly. Individual knobs are polymorphic among strains of both maize and teosinte.

 

Barbara McClintock used these knob markers to validate her transposon theory of "jumping genes", for which she won the 1983 Nobel Prize in Physiology or Medicine. Maize is still an important model organism for genetics and developmental biology today.

 

The centromeres have two types of structural components, both of which are found only in the centromeres: Large arrays of CentC, a short satellite DNA; and a few of a family of retrotransposons. The B chromosome, unlike the others, contains an additional repeat which extends into neighboring areas of the chromosome. Centromeres can accidentally shrink during division and still function, although it is thought this will fail if it shrinks below a few hundred kilobase. Kinetochores contain RNA originating from centromeres. Centromere regions can become inactive, and can continue in that state if the chromosome still has another active one.

 

The Maize Genetics Cooperation Stock Center, funded by the USDA Agricultural Research Service and located in the Department of Crop Sciences at the University of Illinois at Urbana-Champaign, is a stock center of maize mutants. The total collection has nearly 80,000 samples. The bulk of the collection consists of several hundred named genes, plus additional gene combinations and other heritable variants. There are about 1000 chromosomal aberrations (e.g., translocations and inversions) and stocks with abnormal chromosome numbers (e.g., tetraploids). Genetic data describing the maize mutant stocks as well as myriad other data about maize genetics can be accessed at MaizeGDB, the Maize Genetics and Genomics Database.

 

In 2005, the US National Science Foundation (NSF), Department of Agriculture (USDA) and the Department of Energy (DOE) formed a consortium to sequence the B73 maize genome. The resulting DNA sequence data was deposited immediately into GenBank, a public repository for genome-sequence data. Sequences and genome annotations have also been made available throughout the project's lifetime at the project's official site.

 

Primary sequencing of the maize genome was completed in 2008. On November 20, 2009, the consortium published results of its sequencing effort in Science. The genome, 85% of which is composed of transposons, was found to contain 32,540 genes (By comparison, the human genome contains about 2.9 billion bases and 26,000 genes). Much of the maize genome has been duplicated and reshuffled by helitrons—group of rolling circle transposons.

 

In Z. mays and various other angiosperms the MADS-box motif is involved in floral development. Early study in several angiosperm models including Z. mays was the beginning of research into the molecular evolution of floral structure in general, as well as their role in nonflowering plants.

 

EVOLUTION

As with many plants and animals, Z. mays has a positive correlation between effective population size and the magnitude of selection pressure. Z. m. having an EPS of ~650,000, it clusters with others of about the same EPS, and has 79% of its amino acid sites under selection.

 

Recombination is a significant source of diversity in Z. mays. (Note that this finding supersedes previous studies which showed no such correlation.)

 

This recombination/diversity effect is seen throughout plants but is also found to not occur – or not as strongly – in regions of high gene density. This is likely the reason that domesticated Z. mays has not seen as much of an increase in diversity within areas of higher density as in regions of lower density, although there is more evidence in other plants.

 

Some lines of maize have undergone ancient polyploidy events, starting 11m years ago. Over that time ~72% of polyploid duplicated genes have been retained, which is higher than other plants with older polyploidy events. Thus maize may be due to lose more duplicate genes as time goes along, similar to the course followed by the genomes of other plants. If so - if gene loss has merely not occurred yet - that could explain the lack of observed positive selection and lower negative selection which are observed in otherwise similar plants, i.e. also naturally outcrossing and with similar effective population sizes.

 

Ploidy does not appear to influence EPS or magnitude of selection effect in maize.

 

BREEDING

Maize reproduces sexually each year. This randomly selects half the genes from a given plant to propagate to the next generation, meaning that desirable traits found in the crop (like high yield or good nutrition) can be lost in subsequent generations unless certain techniques are used.

 

Maize breeding in prehistory resulted in large plants producing large ears. Modern breeding began with individuals who selected highly productive varieties in their fields and then sold seed to other farmers. James L. Reid was one of the earliest and most successful developing Reid's Yellow Dent in the 1860s. These early efforts were based on mass selection. Later breeding efforts included ear to row selection (C. G. Hopkins c. 1896), hybrids made from selected inbred lines (G. H. Shull, 1909), and the highly successful double cross hybrids using four inbred lines (D. F. Jones c. 1918, 1922). University supported breeding programs were especially important in developing and introducing modern hybrids. By the 1930s, companies such as Pioneer devoted to production of hybrid maize had begun to influence long-term development. Internationally important seed banks such as the International Maize and Wheat Improvement Center (CIMMYT) and the US bank at the Maize Genetics Cooperation Stock Center University of Illinois at Urbana-Champaign maintain germplasm important for future crop development.

 

Since the 1940s the best strains of maize have been first-generation hybrids made from inbred strains that have been optimized for specific traits, such as yield, nutrition, drought, pest and disease tolerance. Both conventional cross-breeding and genetic modification have succeeded in increasing output and reducing the need for cropland, pesticides, water and fertilizer. There is conflicting evidence to support the hypothesis that maize yield potential has increased over the past few decades. This suggests that changes in yield potential are associated with leaf angle, lodging resistance, tolerance of high plant density, disease/pest tolerance, and other agronomic traits rather than increase of yield potential per individual plant.

 

Tropical landraces remain an important and underutilized source of resistance alleles for for disease and for herbivores. Notable discoveries of rare alleles for this purpose were made by Dao et al 2014 and Sood et al 2014.

 

GLOBAL PROGRAM

CIMMYT operates a conventional breeding program to provide optimized strains. The program began in the 1980s. Hybrid seeds are distributed in Africa by the Drought Tolerant Maize for Africa project.

 

GENETIC MODIFICATION

Genetically modified (GM) maize was one of the 26 GM crops grown commercially in 2016. The vast majority of this is Bt maize. Grown since 1997 in the United States and Canada, 92% of the US maize crop was genetically modified in 2016 and 33% of the worldwide maize crop was GM in 2016. As of 2011, Herbicide-tolerant maize varieties were grown in Argentina, Australia, Brazil, Canada, China, Colombia, El Salvador, the European Union, Honduras, Japan, Korea, Malaysia, Mexico, New Zealand, Philippines, the Russian Federation, Singapore, South Africa, Taiwan, Thailand, and the United States. Insect-resistant maize was grown in Argentina, Australia, Brazil, Canada, Chile, China, Colombia, Egypt, the European Union, Honduras, Japan, Korea, Malaysia, Mexico, New Zealand, Philippines, South Africa, Switzerland, Taiwan, the United States, and Uruguay.

 

In September 2000, up to $50 million worth of food products were recalled due to the presence of Starlink genetically modified corn, which had been approved only for animal consumption and had not been approved for human consumption, and was subsequently withdrawn from the market.

 

ORIGIN

Maize is the domesticated variant of teosinte. The two plants have dissimilar appearance, maize having a single tall stalk with multiple leaves and teosinte being a short, bushy plant. The difference between the two is largely controlled by differences in just two genes, called grassy tillers-1 (gt1, A0A317YEZ1) and teosinte branched-1 (tb1, Q93WI2).

 

Several theories had been proposed about the specific origin of maize in Mesoamerica:

 

It is a direct domestication of a Mexican annual teosinte, Zea mays ssp. parviglumis, native to the Balsas River valley in south-eastern Mexico, with up to 12% of its genetic material obtained from Zea mays ssp. mexicana through introgression.

It has been derived from hybridization between a small domesticated maize (a slightly changed form of a wild maize) and a teosinte of section Luxuriantes, either Z. luxurians or Z. diploperennis.

It has undergone two or more domestications either of a wild maize or of a teosinte. (The term "teosinte" describes all species and subspecies in the genus Zea, excluding Zea mays ssp. mays.)

It has evolved from a hybridization of Z. diploperennis by Tripsacum dactyloides.

 

In the late 1930s, Paul Mangelsdorf suggested that domesticated maize was the result of a hybridization event between an unknown wild maize and a species of Tripsacum, a related genus. This theory about the origin of maize has been refuted by modern genetic testing, which refutes Mangelsdorf's model and the fourth listed above. 

 

The teosinte origin theory was proposed by the Russian botanist Nikolai Ivanovich Vavilov in 1931 and the later American Nobel Prize-winner George Beadle in 1932.: 10  It is supported experimentally and by recent studies of the plants' genomes. Teosinte and maize can cross-breed and produce fertile offspring. A number of questions remain concerning the species, among them:

 

how the immense diversity of the species of sect. Zea originated,

how the tiny archaeological specimens of 3500–2700 BC could have been selected from a teosinte, and

how domestication could have proceeded without leaving remains of teosinte or maize with teosintoid traits earlier than the earliest known until recently, dating from ca. 1100 BC.

 

The domestication of maize is of particular interest to researchers—archaeologists, geneticists, ethnobotanists, geographers, etc. The process is thought by some to have started 7,500 to 12,000 years ago. Research from the 1950s to 1970s originally focused on the hypothesis that maize domestication occurred in the highlands between the states of Oaxaca and Jalisco, because the oldest archaeological remains of maize known at the time were found there.

Connection with 'parviglumis' subspecies

Genetic studies, published in 2004 by John Doebley, identified Zea mays ssp. parviglumis, native to the Balsas River valley in Mexico's southwestern highlands, and also known as Balsas teosinte, as being the crop wild relative that is genetically most similar to modern maize. This was confirmed by further studies, which refined this hypothesis somewhat. Archaeobotanical studies, published in 2009, point to the middle part of the Balsas River valley as the likely location of early domestication; this river is not very long, so these locations are not very distant. Stone milling tools with maize residue have been found in an 8,700 year old layer of deposits in a cave not far from Iguala, Guerrero.

 

Doebley was part of the team that first published, in 2002, that maize had been domesticated only once, about 9,000 years ago, and then spread throughout the Americas.

 

A primitive corn was being grown in southern Mexico, Central America, and northern South America 7,000 years ago. Archaeological remains of early maize ears, found at Guila Naquitz Cave in the Oaxaca Valley, date back roughly 6,250 years; the oldest ears from caves near Tehuacan, Puebla, 5,450 B.P.

 

Maize pollen dated to 7,300 B.P. from San Andres, Tabasco, on the Caribbean coast has also been recovered.

 

As maize was introduced to new cultures, new uses were developed and new varieties selected to better serve in those preparations. Maize was the staple food, or a major staple – along with squash, Andean region potato, quinoa, beans, and amaranth – of most pre-Columbian North American, Mesoamerican, South American, and Caribbean cultures. The Mesoamerican civilization, in particular, was deeply interrelated with maize. Its traditions and rituals involved all aspects of maize cultivation – from the planting to the food preparation. Maize formed the Mesoamerican people's identity.

 

It is unknown what precipitated its domestication, because the edible portion of the wild variety is too small, and hard to obtain, to be eaten directly, as each kernel is enclosed in a very hard bivalve shell.

 

In 1939, George Beadle demonstrated that the kernels of teosinte are readily "popped" for human consumption, like modern popcorn. Some have argued it would have taken too many generations of selective breeding to produce large, compressed ears for efficient cultivation. However, studies of the hybrids readily made by intercrossing teosinte and modern maize suggest this objection is not well founded.

 

SPREADING TO THE NORTH

Around 4,500 ago, maize began to spread to the north; it was first cultivated in what is now the United States at several sites in New Mexico and Arizona, about 4,100 ago.

 

During the first millennium AD, maize cultivation spread more widely in the areas north. In particular, the large-scale adoption of maize agriculture and consumption in eastern North America took place about A.D. 900. Native Americans cleared large forest and grassland areas for the new crop.

 

In 2005, research by the USDA Forest Service suggested that the rise in maize cultivation 500 to 1,000 years ago in what is now the southeastern United States corresponded with a decline of freshwater mussels, which are very sensitive to environmental changes.

 

CULTIVATION

PLANTING

Because it is cold-intolerant, in the temperate zones maize must be planted in the spring. Its root system is generally shallow, so the plant is dependent on soil moisture. As a plant that uses C4 carbon fixation, maize is a considerably more water-efficient crop than plants that use C3 carbon fixation such as alfalfa and soybeans. Maize is most sensitive to drought at the time of silk emergence, when the flowers are ready for pollination. In the United States, a good harvest was traditionally predicted if the maize was "knee-high by the Fourth of July", although modern hybrids generally exceed this growth rate. Maize used for silage is harvested while the plant is green and the fruit immature. Sweet corn is harvested in the "milk stage", after pollination but before starch has formed, between late summer and early to mid-autumn. Field maize is left in the field until very late in the autumn to thoroughly dry the grain, and may, in fact, sometimes not be harvested until winter or even early spring. The importance of sufficient soil moisture is shown in many parts of Africa, where periodic drought regularly causes maize crop failure and consequent famine. Although it is grown mainly in wet, hot climates, it has been said to thrive in cold, hot, dry or wet conditions, meaning that it is an extremely versatile crop.

 

Maize was planted by the Native Americans in hills, in a complex system known to some as the Three Sisters. Maize provided support for beans, and the beans provided nitrogen derived from nitrogen-fixing rhizobia bacteria which live on the roots of beans and other legumes; and squashes provided ground cover to stop weeds and inhibit evaporation by providing shade over the soil. This method was replaced by single species hill planting where each hill 60–120 cm (2 ft 0 in–3 ft 11 in) apart was planted with three or four seeds, a method still used by home gardeners. A later technique was "checked maize", where hills were placed

 

1 m (40 in) apart in each direction, allowing cultivators to run through the field in two directions. In more arid lands, this was altered and seeds were planted in the bottom of 10–12 cm (4–4+1⁄2 in) deep furrows to collect water. Modern technique plants maize in rows which allows for cultivation while the plant is young, although the hill technique is still used in the maize fields of some Native American reservations. When maize is planted in rows, it also allows for planting of other crops between these rows to make more efficient use of land space.

 

In most regions today, maize grown in residential gardens is still often planted manually with a hoe, whereas maize grown commercially is no longer planted manually but rather is planted with a planter. In North America, fields are often planted in a two-crop rotation with a nitrogen-fixing crop, often alfalfa in cooler climates and soybeans in regions with longer summers. Sometimes a third crop, winter wheat, is added to the rotation.

 

Many of the maize varieties grown in the United States and Canada are hybrids. Often the varieties have been genetically modified to tolerate glyphosate or to provide protection against natural pests. Glyphosate is an herbicide which kills all plants except those with genetic tolerance. This genetic tolerance is very rarely found in nature.

 

In the midwestern United States, low-till or no-till farming techniques are usually used. In low-till, fields are covered once, maybe twice, with a tillage implement either ahead of crop planting or after the previous harvest. The fields are planted and fertilized. Weeds are controlled through the use of herbicides, and no cultivation tillage is done during the growing season. This technique reduces moisture evaporation from the soil, and thus provides more moisture for the crop. The technologies mentioned in the previous paragraph enable low-till and no-till farming. Weeds compete with the crop for moisture and nutrients, making them undesirable.

 

HARVESTING

Before the 20th century, all maize harvesting was by manual labour, by grazing, or by some combination of those. Whether the ears were hand-picked and the stover was grazed, or the whole plant was cut, gathered, and shocked, people and livestock did all the work. Between the 1890s and the 1970s, the technology of maize harvesting expanded greatly. Today, all such technologies, from entirely manual harvesting to entirely mechanized, are still in use to some degree, as appropriate to each farm's needs, although the thoroughly mechanized versions predominate, as they offer the lowest unit costs when scaled to large farm operations. For small farms, their unit cost can be too high, as their higher fixed cost cannot be amortized over as many units.[citation needed]

 

Before World War II, most maize in North America was harvested by hand. This involved a large number of workers and associated social events (husking or shucking bees). From the 1890s onward, some machinery became available to partially mechanize the processes, such as one- and two-row mechanical pickers (picking the ear, leaving the stover) and corn binders, which are reaper-binders designed specifically for maize (for example, Video on YouTube). The latter produce sheaves that can be shocked. By hand or mechanical picker, the entire ear is harvested, which then requires a separate operation of a maize sheller to remove the kernels from the ear. Whole ears of maize were often stored in corn cribs, and these whole ears are a sufficient form for some livestock feeding use. Today corn cribs with whole ears, and corn binders, are less common because most modern farms harvest the grain from the field with a combine and store it in bins. The combine with a corn head (with points and snap rolls instead of a reel) does not cut the stalk; it simply pulls the stalk down. The stalk continues downward and is crumpled into a mangled pile on the ground, where it usually is left to become organic matter for the soil. The ear of maize is too large to pass between slots in a plate as the snap rolls pull the stalk away, leaving only the ear and husk to enter the machinery. The combine separates the husk and the cob, keeping only the kernels.

When maize is a silage crop, the entire plant is usually chopped at once with a forage harvester (chopper) and ensiled in silos or polymer wrappers. Ensiling of sheaves cut by a corn binder was formerly common in some regions but has become uncommon. For storing grain in bins, the moisture of the grain must be sufficiently low to avoid spoiling. If the moisture content of the harvested grain is too high, grain dryers are used to reduce the moisture content by blowing heated air through the grain. This can require large amounts of energy in the form of combustible gases (propane or natural gas) and electricity to power the blowers.

 

PRODUCTION

Maize is widely cultivated throughout the world, and a greater weight of maize is produced each year than any other grain. In 2018, total world production was 1.15 billion tonnes, led by the United States with 34.2% of the total (table). China produced 22.4% of the global total.

 

UNITED STATES

In 2016, maize production was forecast to be over 380 million metric tons (15 billion bushels), an increase of 11% over 2014 American production. Based on conditions as of August 2016, the expected yield would be the highest ever for the United States. The area of harvested maize was forecast to be 35 million hectares (87 million acres), an increase of 7% over 2015. Maize is especially popular in Midwestern states such as Indiana, Iowa, and Illinois; in the latter, it was named the state's official grain in 2017.

 

STORAGE

Drying is vital to prevent or at least reduce mycotoxin contamination. Aspergillus and Fusarium spp. are the most common mycotoxin sources, but there are others. Altogether maize contaminants are so common, and this crop is so economically important, that maize mycotoxins are among the most important in agriculture in general.

 

USES

HUMAN FOOD

Maize and cornmeal (ground dried maize) constitute a staple food in many regions of the world. Maize is used to produce cornstarch, a common ingredient in home cooking and many industrialized food products. Maize starch can be hydrolyzed and enzymatically treated to produce syrups, particularly high fructose corn syrup, a sweetener; and also fermented and distilled to produce grain alcohol. Grain alcohol from maize is traditionally the source of Bourbon whiskey. Corn flour is used to make cornbread and other baked products.

 

In prehistoric times Mesoamerican women used a metate to process maize into ground cornmeal, allowing the preparation of foods that were more calorie dense than popcorn. After ceramic vessels were invented the Olmec people began to cook maize together with beans, improving the nutritional value of the staple meal. Although maize naturally contains niacin, an important nutrient, it was not bioavailable without the process of nixtamalization. The Maya used nixtamal meal to make varieties of porridges and tamales. The process was later used in the cuisine of the American South to prepare corn for grits and hominy.

 

Maize is a staple of Mexican cuisine. Masa (cornmeal treated with limewater) is the main ingredient for tortillas, atole and many other dishes of Central American food. It is the main ingredient of corn tortilla, tamales, pozole, atole and all the dishes based on them, like tacos, quesadillas, chilaquiles, enchiladas, tostadas and many more. In Mexico the fungus of maize, known as huitlacoche, is considered a delicacy.

 

Coarse maize meal is made into a thick porridge in many cultures: from the polenta of Italy, the angu of Brazil, the mămăligă of Romania, to cornmeal mush in the US (or hominy grits in the South) or the food called mieliepap in South Africa and sadza, nshima, ugali and other names in other parts of Africa. Introduced into Africa by the Portuguese in the 16th century, maize has become Africa's most important staple food crop. These are commonly eaten in the Southeastern United States, foods handed down from Native Americans, who called the dish sagamite.

 

Maize can also be harvested and consumed in the unripe state, when the kernels are fully grown but still soft. Unripe maize must usually be cooked to become palatable; this may be done by simply boiling or roasting the whole ears and eating the kernels right off the cob. Sweet corn, a genetic variety that is high in sugars and low in starch, is usually consumed in the unripe state. Such corn on the cob is a common dish in the United States, Canada, United Kingdom, Cyprus, some parts of South America, and the Balkans, but virtually unheard of in some European countries. Corn on the cob was hawked on the streets of early 19th-century New York City by poor, barefoot "Hot Corn Girls", who were thus the precursors of hot dog carts, churro wagons, and fruit stands seen on the streets of big cities today.

 

Within the United States, the usage of maize for human consumption constitutes only around 1/40th of the amount grown in the country. In the United States and Canada, maize is mostly grown to feed livestock, as forage, silage (made by fermentation of chopped green cornstalks), or grain. Maize meal is also a significant ingredient of some commercial animal food products.

 

NUTRITIONAL VALUE

Raw, yellow, sweet maize kernels are composed of 76% water, 19% carbohydrates, 3% protein, and 1% fat (table). In a 100-gram serving, maize kernels provide 86 calories and are a good source (10–19% of the Daily Value) of the B vitamins, thiamin, niacin (but see Pellagra warning below), pantothenic acid (B5) and folate (right table for raw, uncooked kernels, USDA Nutrient Database). In moderate amounts, they also supply dietary fiber and the essential minerals, magnesium and phosphorus whereas other nutrients are in low amounts (table).

 

Maize has suboptimal amounts of the essential amino acids tryptophan and lysine, which accounts for its lower status as a protein source. However, the proteins of beans and legumes complement those of maize.

 

FEED AND FODDER FOR LIVESTOCK

Maize is a major source of both grain feed and fodder for livestock. It is fed to the livestock in various ways. When it is used as a grain crop, the dried kernels are used as feed. They are often kept on the cob for storage in a corn crib, or they may be shelled off for storage in a grain bin. The farm that consumes the feed may produce it, purchase it on the market, or some of both. When the grain is used for feed, the rest of the plant (the corn stover) can be used later as fodder, bedding (litter), or soil amendment. When the whole maize plant (grain plus stalks and leaves) is used for fodder, it is usually chopped all at once and ensilaged, as digestibility and palatability are higher in the ensilaged form than in the dried form. Maize silage is one of the most valuable forages for ruminants. Before the advent of widespread ensilaging, it was traditional to gather the corn into shocks after harvesting, where it dried further. With or without a subsequent move to the cover of a barn, it was then stored for weeks to several months until fed to the livestock. Today ensilaging can occur not only in siloes but also in silage wrappers. However, in the tropics, maize can be harvested year-round and fed as green forage to the animals.

 

CHEMICALS

Starch from maize can also be made into plastics, fabrics, adhesives, and many other chemical products.

 

The corn steep liquor, a plentiful watery byproduct of maize wet milling process, is widely used in the biochemical industry and research as a culture medium to grow many kinds of microorganisms.

 

Chrysanthemin is found in purple corn and is used as a food coloring.

 

BIO-FUEL

"Feed maize" is being used increasingly for heating; specialized corn stoves (similar to wood stoves) are available and use either feed maize or wood pellets to generate heat. Maize cobs are also used as a biomass fuel source. Maize is relatively cheap and home-heating furnaces have been developed which use maize kernels as a fuel. They feature a large hopper that feeds the uniformly sized maize kernels (or wood pellets or cherry pits) into the fire.[citation needed]

 

Maize is increasingly used as a feedstock for the production of ethanol fuel. When considering where to construct an ethanol plant, one of the site selection criteria is to ensure there is locally available feedstock. Ethanol is mixed with gasoline to decrease the amount of pollutants emitted when used to fuel motor vehicles. High fuel prices in mid-2007 led to higher demand for ethanol, which in turn led to higher prices paid to farmers for maize. This led to the 2007 harvest being one of the most profitable maize crops in modern history for farmers. Because of the relationship between fuel and maize, prices paid for the crop now tend to track the price of oil.

 

The price of food is affected to a certain degree by the use of maize for biofuel production. The cost of transportation, production, and marketing are a large portion (80%) of the price of food in the United States. Higher energy costs affect these costs, especially transportation. The increase in food prices the consumer has been seeing is mainly due to the higher energy cost. The effect of biofuel production on other food crop prices is indirect. Use of maize for biofuel production increases the demand, and therefore price of maize. This, in turn, results in farm acreage being diverted from other food crops to maize production. This reduces the supply of the other food crops and increases their prices.

 

Maize is widely used in Germany as a feedstock for biogas plants. Here the maize is harvested, shredded then placed in silage clamps from which it is fed into the biogas plants. This process makes use of the whole plant rather than simply using the kernels as in the production of fuel ethanol.

 

A biomass gasification power plant in Strem near Güssing, Burgenland, Austria, began in 2005. Research is being done to make diesel out of the biogas by the Fischer Tropsch method.

 

Increasingly, ethanol is being used at low concentrations (10% or less) as an additive in gasoline (gasohol) for motor fuels to increase the octane rating, lower pollutants, and reduce petroleum use (what is nowadays also known as "biofuels" and has been generating an intense debate regarding the human beings' necessity of new sources of energy, on the one hand, and the need to maintain, in regions such as Latin America, the food habits and culture which has been the essence of civilizations such as the one originated in Mesoamerica; the entry, January 2008, of maize among the commercial agreements of NAFTA has increased this debate, considering the bad labor conditions of workers in the fields, and mainly the fact that NAFTA "opened the doors to the import of maize from the United States, where the farmers who grow it receive multimillion-dollar subsidies and other government supports. ... According to OXFAM UK, after NAFTA went into effect, the price of maize in Mexico fell 70% between 1994 and 2001. The number of farm jobs dropped as well: from 8.1 million in 1993 to 6.8 million in 2002. Many of those who found themselves without work were small-scale maize growers."). However, introduction in the northern latitudes of the US of tropical maize for biofuels, and not for human or animal consumption, may potentially alleviate this.

 

COMMODITY

Maize is bought and sold by investors and price speculators as a tradable commodity using corn futures contracts. These "futures" are traded on the Chicago Board of Trade (CBOT) under ticker symbol C. They are delivered every year in March, May, July, September, and December.

 

ORNAMENTAL AND OTHER USES

Some forms of the plant are occasionally grown for ornamental use in the garden. For this purpose, variegated and colored leaf forms as well as those with colorful ears are used.

 

Corncobs can be hollowed out and treated to make inexpensive smoking pipes, first manufactured in the United States in 1869.

 

An unusual use for maize is to create a "corn maze" (or "maize maze") as a tourist attraction. The idea of a maize maze was introduced by the American Maze Company who created a maze in Pennsylvania in 1993. Traditional mazes are most commonly grown using yew hedges, but these take several years to mature. The rapid growth of a field of maize allows a maze to be laid out using GPS at the start of a growing season and for the maize to grow tall enough to obstruct a visitor's line of sight by the start of the summer. In Canada and the US, these are popular in many farming communities.

 

Maize kernels can be used in place of sand in a sandboxlike enclosure for children's play.

 

Stigmas from female maize flowers, popularly called corn silk, are sold as herbal supplements.

 

Maize is used as a fish bait, called "dough balls". It is particularly popular in Europe for coarse fishing.

 

Additionally, feed corn is sometimes used by hunters to bait animals such as deer or wild hogs.

 

UNITED STATES USAGE BREAKDOWN

The breakdown of usage of the 12.1-billion-bushel (307-million-tonne) 2008 US maize crop was as follows, according to the World Agricultural Supply and Demand Estimates Report by the USDA.In the US since 2009/2010, maize feedstock use for ethanol production has somewhat exceeded direct use for livestock feed; maize use for fuel ethanol was 5,130 million bushels (130 million tonnes) in the 2013/2014 marketing year.A fraction of the maize feedstock dry matter used for ethanol production is usefully recovered as DDGS (dried distillers grains with solubles). In the 2010/2011 marketing year, about 29.1 million tonnes of DDGS were fed to US livestock and poultry. Because starch utilization in fermentation for ethanol production leaves other grain constituents more concentrated in the residue, the feed value per kg of DDGS, with regard to ruminant-metabolizable energy and protein, exceeds that of the grain. Feed value for monogastric animals, such as swine and poultry, is somewhat lower than for ruminants.

 

HAZARDS

PELLAGRA

When maize was first introduced into farming systems other than those used by traditional native-American peoples, it was generally welcomed with enthusiasm for its productivity. However, a widespread problem of malnutrition soon arose wherever maize was introduced as a staple food. This was a mystery, since these types of malnutrition were not normally seen among the indigenous Americans, for whom maize was the principal staple food.

 

It was eventually discovered that the indigenous Americans had learned to soak maize in alkali — water (the process now known as nixtamalization) — made with ashes and lime (calcium oxide) since at least 1200–1500 BC by Mesoamericans. They did this to liberate the corn hulls, but (unbeknownst to natives or colonists) it coincidentally liberates the B-vitamin niacin, the lack of which was the underlying cause of the condition known as pellagra.

 

Maize was introduced into the diet of non-indigenous Americans without the necessary cultural knowledge acquired over thousands of years in the Americas. In the late 19th century, pellagra reached epidemic proportions in parts of the southern US, as medical researchers debated two theories for its origin: the deficiency theory (which was eventually shown to be true) said that pellagra was due to a deficiency of some nutrient, and the germ theory said that pellagra was caused by a germ transmitted by stable flies. A third theory, promoted by the eugenicist Charles Davenport, held that people only contracted pellagra if they were susceptible to it due to certain "constitutional, inheritable" traits of the affected individual.

 

Once alkali processing and dietary variety were understood and applied, pellagra disappeared in the developed world. The development of high lysine maize and the promotion of a more balanced diet have also contributed to its demise. Pellagra still exists today in food-poor areas and refugee camps where people survive on donated maize.

 

ALLERGY

Maize contains lipid transfer protein, an indigestible protein that survives cooking. This protein has been linked to a rare and understudied allergy to maize in humans. The allergic reaction can cause skin rash, swelling or itching of mucous membranes, diarrhea, vomiting, asthma and, in severe cases, anaphylaxis. It is unclear how common this allergy is in the general population.

 

MYCOTOXINS

Fungicide application does not reduce fungal growth or mycotoxin dramatically, although it can be a part of a successful reduction strategy. Among the most common toxins are those produced by Aspergillus and Fusarium spp. The most common toxins are aflatoxins, fumonisins, zearalenone, and ochratoxin A. Bt maize discourages insect vectors and by so doing it dramatically reduces concentrations of fumonisins, significantly reduces aflatoxins, but only mildly reduces others.

 

ART

Maize has been an essential crop in the Andes since the pre-Columbian era. The Moche culture from Northern Peru made ceramics from earth, water, and fire. This pottery was a sacred substance, formed in significant shapes and used to represent important themes. Maize was represented anthropomorphically as well as naturally.

 

In the United States, maize ears along with tobacco leaves are carved into the capitals of columns in the United States Capitol building. Maize itself is sometimes used for temporary architectural detailing when the intent is to celebrate the fall season, local agricultural productivity and culture. Bundles of dried maize stalks are often displayed along with pumpkins, gourds and straw in autumnal displays outside homes and businesses. A well-known example of architectural use is the Corn Palace in Mitchell, South Dakota, which uses cobs and ears of colored maize to implement a mural design that is recycled annually. Another well-known example is the Field of Corn sculpture in Dublin, Ohio, where hundreds of concrete ears of corn stand in a grassy field.

 

A maize stalk with two ripe ears is depicted on the reverse of the Croatian 1 lipa coin, minted since 1993.

 

WIKIPEDIA

I spent the last few days preparing for my Advanced Cardiac Life Support recertification today. Pictured above is a potentially lethal rhythm, perhaps the most lethal of all heart rhythms. Torsades de pointes is a distinctive form of polymorphic ventricular tachycardia with a gradual change in the amplitude and twisting of the QRS complexes around the isoelectric line. The management of Torsades is quite different from that of garden-variety V tach. In particular, the use of group IA antidysrhythmic drugs, which tend to prolong the QT interval, can have disastrous consequences in Torsades. Rapid and accurate diagnosis, therefore, is critical.

 

The characteristics of Torsades are:

Rotation of the heart's electrical axis by at least 180º

Prolonged QT interval (LQTS)

Preceded by long and short RR-intervals

Triggered by an early premature ventricular contraction (R-on-T PVC)

 

Torsades can be caused by diarrhea, hypomagnesemia and hypokalemia. It is usually seen in the malnourished and alcoholic patient. In an otherwise stable patient, cardioversion is a last resort. Torsades tends to be paroxysmal in nature, and often occurs following cardioversion. Although Torsades frequently is self limiting, it may refract into ventricular fibrillation, requiring defibrillation.

 

To treat, remove causative factors. Predisposing conditions such as hypokalemia, hypomagnesemia, and bradycardia should be identified and corrected. Magnesium is the drug of choice for terminating Torsades. Magnesium can be given at 1-2 g IV initially in 30-60 seconds, which then can be repeated in 5 to 15 minutes. Alternatively, a continuous infusion can be started at a rate of 3-10 mg/min. Magnesium is effective even in patients with normal magnesium levels. Because of the danger of hypermagnesemia, the patient requires close monitoring.

 

Oh, how did I do on my recertification? I passed. I get to keep doing my job.

 

View Large and on White

 

Strobist: AB800 with Softlighter II camera right. AB800 open behind backdrop of white faux suede. Triggered by Cybersync.

☆Explore☆.

 

Description

Morelia viridis, the green tree python, is a species of python found in New Guinea, islands in Indonesia, and Cape York Peninsula in Australia.

 

Adults average 90-120 cm (3-4 feet) in length, with a maximum grow to about 213 cm (7 ft). The supralabial scales have thermoreceptive pits.

Completely arboreal with a striking green color in adults. The color pattern is vivid green with a broken vertebral stripe of white or dull yellow. Spots of the same color, or blue spots, may be scattered over the body. Cyanomorphs (blue morphs) are also known to occur. Juveniles are polymorphic, occurring in reddish, bright yellow and orange morphs

 

Adult Indian paradise flycatchers are 19–22 cm (7.5–8.7 in) long. Their heads are glossy black with a black crown and crest, their black bill round and sturdy, their eyes black. Female are rufous on the back with a greyish throat and underparts. Their wings are 86–92 mm (3.4–3.6 in) long. Young males look very much like females but have a black throat and blue-ringed eyes. As adults they develop up to 24 cm (9.4 in) long tail feathers with two central tail feathers growing up to 30 cm (12 in) long drooping streamers.

 

Young males are rufous and have short tails. They acquire long tails in their second or third year. Adult males are either predominantly bright rufous above or predominantly white. Some specimens show some degree of intermediacy between rufous and white. Long-tailed rufous birds are generally devoid of shaft streaks on the wing and tail feathers, while in white birds the shaft streaks, and sometimes the edges of the wing and tail feathers are black.

 

In the early 1960s, 680 long-tailed males were examined that are contained in collections of the British Museum of Natural History, Chicago Natural History Museum, Peabody Museum, Carnegie Museum, American Museum of Natural History, United States National Museum and Royal Ontario Museum. The specimens came from almost the entire range of the species, though some areas were poorly represented. The relative frequency of the rufous and white plumage types varies geographically. Rufous birds are rare in the extreme southeastern part of the species' range. Throughout the Indian area and, to a lesser extent, in China, asymmetrically patterned intermediates occur. Intermediates are rare or absent throughout the rest of the range of the species. In general, long-tailed males are

 

predominantly rufous with some white in wings and tail — collected in Turkestan, Kashmir, northern India, Punjab, Maharashtra, Sikkim and in Sri Lanka;

predominantly rufous with some white in wings — collected in Iran, Afghanistan, Baluchistan, Punjab, Kashmir, northern and central India, Rajasthan, Maharashtra, Bihar, Nepal;

predominantly rufous with some white in tail — collected in Punjab, northern and central India, Kolkata, Sri Lanka and in the Upper Yangtze Valley in China;

predominantly white with some rufous in tail and wings — collected in Kashmir, Maharashtra, Sichuan and North China;

predominantly white with some rufous in tail — collected in Maharashtra and Fuzhou, China;

predominantly white with back partly rufous — collected in Punjab and Chennai;

predominantly white with wings and tail irregularly blotched and washed with rufous in places — in the extreme southeastern edge of the range of the species : Alor Island and Sumba;

moulting from rufous into white plumage — collected in North Bihar.

Possible interpretations of this phenomenon are : males may be polymorphic for rufous and white plumage colour; rufous birds may be sub-adults; and there may even be two sympatric species distinguishable only in the mal

 

Papilio memnon, the great Mormon, is a large butterfly native to southern Asia that belongs to the swallowtail family. It is widely distributed and has thirteen subspecies. The female is polymorphic and with mimetic forms.

 

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Thanks to every friend ! Greetings ! And wish you every lucky !

 

2018.09.05

The abbey of Sant’ Antimo in Tuscany was, according to legend, founded by Charlemagne. What is for sure is that the monastery already existed in 842, as it is mentioned in writing in a charter by Louis le Pieux.

 

The church we can see today was built in the 1100s in the Burgundy Cistercian style, which demonstrates how intense and fertile the exchanges were between seemingly distant lands during all the Middle Ages: this church was influenced by Burgundy builders from France, while the basilica of Aime (of which I am simultaneously uploading pictures) was influenced by Lombard builders from Italy...!

 

Of course, neither “France” nor “Italy” had anything to do then with the two States we know today.

 

One example of fine Romanesque sculpture with these polymorphic monsters.

  

Innenpark. Bonifatius-Felsen.

Inner Park. Boniface Rock.

 

Asplenium trichomanes subsp. pachyrachis (Christ) Lovis et Reichst.

Willdenowia 10: 18. 1980.

 

Asplenium trichomanes sublus. pachyrachis Christ

Farnkr. Schweiz 1 (2): 92. 1900.

Asplenium csikii Kümmerle & András.

Magyar Bot. Lapok 17. 110. 1919 (nomen), 20. 3, fig. 1923

Asplenium pachyrhachis Landolt

Fl. Indicativa 268 2010.

 

Seestern-Braunschwarz-Streifenfarn, Dickstieliger Braunstieliger Streifenfarn, Dickstieliger Brauner Streifenfarn

Lobed Maidenhair Spleenwort

 

Anmerkung:

Die Asplenium trichomanes Gruppe, ist ein sehr polymorpher, taxonomisch kritischer Spezieskomplex! Die Evolutionsgeschichte und Beziehungen zwischen den Taxa in dieser Gruppe wurden intensiv untersucht. Allerdings sind morphologische Variation und die Verteilung dieser Taxa unzureichend bekannt, da sie in der lokalen Floren oder Checklisten häufig nicht vorkommen. Die Gründe für die Vernachlässigung der Taxa innerhalb der Asplenium trichomanes Gruppe sind der Mangel an diagnostischen morphologische Merkmalen, das häufige gemeinsame Auftreten an ihren Standorten, sowie die Hybridisierung unter den Taxa. Die Asplenium trichomanes Gruppe umfasst zytologisch und ökologisch unterschiedliche Taxa mit fast weltweiter Verbreitung, die offenbar noch aktiv in der Entwicklung sind (L. Ekrt & M. Štech, 2008).

 

Annotation:

The Asplenium trichomanes group is a very polymorphic, taxonomically critical species complex! The evolutionary history and relationships between the taxa in this group have been extensively studied. However, morphological variation and distribution of these taxa are poorly understood because they are often absent in local floras or checklists. The reasons for the neglect of the taxa within the Asplenium trichomanes group are the lack of diagnostic morphological features, the frequent common occurrence at their sites, as well as the hybridization among the taxa. The Asplenium trichomanes group includes cytologically and ecologically diverse taxa with almost worldwide distribution, which apparently are still active in development (L. Ekrt & M. Štech, 2008).

Photographed while walking at San Antonio Open Space Preserve, Los Altos, California

 

Please click on the photo or press the L key to view the larger size

 

This beautiful Red-tailed Hawk was perched on a horizontal branch, no more than 50 feet from a heavily-used trail that winds up the hillside from the parking area. Many hikers and runners passed this hawk in both directions without noticing the hawk. The hawk itself was constantly moving its head about as it was searching for prey.

 

Canon 7D Mark II. f/5.6 1/800 ISO 400

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From Wikipedia: The red-tailed hawk (Buteo jamaicensis) is a bird of prey that breeds throughout most of North America, from the interior of Alaska and northern Canada to as far south as Panama and the West Indies. It is one of the most common members within the genus of Buteo in North America or worldwide. The red-tailed hawk is one of three species colloquially known in the United States as the "chickenhawk," though it rarely preys on standard-sized chickens. The bird is sometimes also referred to as the red-tail for short, when the meaning is clear in context.

 

Red-tailed hawks can acclimate to all the biomes within their range, occurring on the edges of non-ideal habitats such as dense forests and sandy deserts. The red-tailed hawk occupies a wide range of habitats and altitudes including deserts, grasslands, coniferous and deciduous forests, agricultural fields and urban areas. Its latitudinal limits fall around the tree line in the Arctic and the species is absent from the high Arctic. It is legally protected in Canada, Mexico and the United States by the Migratory Bird Treaty Act.

 

The 14 recognized subspecies vary in appearance and range, varying most often in color, and in the west of North America, red-tails are particularly often strongly polymorphic, with individuals ranging from almost white to nearly all black. The subspecies Harlan's hawk (B. j. harlani) is sometimes considered a separate species (B. harlani). The red-tailed hawk is one of the largest members of the genus Buteo, typically weighing from 690 to 1,600 g (1.5 to 3.5 lb) and measuring 45–65 cm (18–26 in) in length, with a wingspan from 110–141 cm (3 ft 7 in–4 ft 8 in). This species displays sexual dimorphism in size, with females averaging about 25% heavier than males.

 

The diet of red-tailed hawks is highly variable and reflects their status as opportunistic generalist, but in North America, it is most often a predator of small mammals such as rodents. Prey that is terrestrial and diurnal is preferred so types such as ground squirrels are preferential where they naturally occur. Large numbers of birds and reptiles can occur in the diet in several areas and can even be the primary foods. Meanwhile, amphibians, fish and invertebrates can seem rare in the hawk’s regular diet; however, they are not infrequently taken by immature hawks.

 

Red-tailed hawks may survive on islands absent of native mammals on diets variously including invertebrates such as crabs, or lizards and birds. Like many Buteo, they hunt from a perch most often but can vary their hunting techniques where prey and habitat demand it. Because they are so common and easily trained as capable hunters, the majority of hawks captured for falconry in the United States are red-tails. Falconers are permitted to take only passage hawks (which have left the nest, are on their own, but are less than a year old) so as to not affect the breeding population. Adults, which may be breeding or rearing chicks, may not be taken for falconry purposes and it is illegal to do so. Passage red-tailed hawks are also preferred by falconers because these younger birds have not yet developed the adult behaviors which would make them more difficult to train.

 

Description:

Red-tailed hawk plumage can be variable, depending on the subspecies and the region. These color variations are morphs, and are not related to molting. The western North American population, B. j. calurus, is the most variable subspecies and has three main color morphs: light, dark, and intermediate or rufous. The dark and intermediate morphs constitute 10–20% of the population in the western United States but seem to constitute only 1-2% of B. j. calurus in western Canada. A whitish underbelly with a dark brown band across the belly, formed by horizontal streaks in feather patterning, is present in most color variations. This feature is variable in eastern hawks and generally absent in some light subspecies (i.e. B. j. fuertesi).

 

Most adult red-tails have a dark brown nape and upper head which gives them a somewhat hooded appearance, while the throat can variably present a lighter brown “necklace”. Especially in younger birds, the underside may be otherwise covered with dark brown spotting and some adults may too manifest this stippling. The back is usually a slightly darker brown than elsewhere with paler scapular feathers, ranging from tawny to white, forming a variable imperfect “V” on the back. The tail of most adults, which of course gives this species its name, is rufous brick-red above with a variably sized black subterminal band and generally appears light buff-orange from below. In comparison, the typical pale immatures (i.e. less than two years old) typically have a mildly paler headed and tend to show a darker back than adults with more apparent pale wing feather edges above (for descriptions of dark morph juveniles from B. j. calurus, which is also generally apt for description of rare dark morphs of other races, see under that subspecies description). In immature red-tailed hawks of all hues, the tail is a light brown above with numerous small dark brown bars of roughly equal width, but these tend to be much broader on dark morph birds.

 

Even in young red-tails, the tail may be a somewhat rufous tinge of brown. The bill is relatively short and dark, in the hooked shape characteristic of raptors, and the head can sometimes appear small in size against the thick body frame. The cere, the legs, and the feet of the red-tailed hawk are all yellow, as is the hue of bare parts in many accipitrids of different lineages. Immature birds can be readily identified at close range by their yellowish irises. As the bird attains full maturity over the course of 3–4 years, the iris slowly darkens into a reddish-brown hue, which is the adult eye-color in all races. Seen in flight, adults usually have dark brown along the lower edge of the wings, against a mostly pale wing, which bares light brownish barring. Individually, the underwing coverts can range from all dark to off-whitish (most often more heavily streaked with brown) which contrasts with a distinctive black patagium marking. The wing coloring of adults and immatures is similar but for typical pale morph immatures having somewhat heavier brownish markings.

 

Though the markings and hue vary across the subspecies, the basic appearance of the red-tailed hawk is relatively consistent. Overall, this species is blocky and broad in shape, often appearing (and being) heavier than other Buteos of similar length. They are the heaviest Buteos on average in eastern North America, albeit scarcely ahead of the larger winged rough-legged buzzard (Buteo lagopus), and second only in size in the west to the ferruginous hawk (Buteo regalis). Red-tailed hawks may be anywhere from the seventh to the ninth heaviest Buteo in the world depending on what figures are used. However, in the northwestern United States, ferruginous hawk females are 35% heavier than female red-tails from the same area. On average, western red-tailed hawks are relatively longer winged and lankier proportioned but are slightly less stocky, compact and heavy than eastern red-tailed hawks in North America. Eastern hawks may also have mildly larger talons and bills than western ones. Based on comparisons of morphology and function amongst all accipitrids, these features imply that western red-tails may need to vary their hunting more frequently to on the wing as the habitat diversifies to more open situations and presumably would hunt more variable and faster prey, whereas the birds of the east, which was historically well-wooded, are more dedicated perch hunters and can take somewhat larger prey but are likely more dedicated mammal hunters. In terms of size variation, red-tailed hawks run almost contrary to Bergmann's rule (i.e. that northern animals should be larger in relation than those closer to the Equator within a species) as one of the northernmost subspecies, B. j. alascensis, is the second smallest race based on linear dimensions and that two of the most southerly occurring races in the United States, B. j. fuertesi and B. j. umbrinus, respectively, are the largest proportioned of all red-tailed hawks. Red-tailed hawks tend have a relatively short but broad tails and thick, chunky wings. Although often described as long winged, the proportional size of the wings is quite small and red-tails have high wing loading for a buteonine hawk. For comparison, two other widespread Buteo hawks in North America were found to weigh: 30 g (1.1 oz) for every square centimeter of wing area in the rough-legged buzzard (Buteo lagopus) and 44 g (1.6 oz) per square cm in the red-shouldered hawk (Buteo lineatus). In contrast, the red-tailed hawk weighed considerably more for their wing area: 199 g (7.0 oz) per square cm.

 

As is the case with many raptors, the red-tailed hawk displays sexual dimorphism in size, as females are up to 25% larger than males. As is typical in large raptors, frequently reported mean body mass for Red-tailed Hawks are somewhat higher than expansive research reveals. Part of this weight variation is seasonal fluctuations, hawks tending to be heavier in winter than during migration or especially during the trying summer breeding season, and also due to clinal variation. Furthermore, immature hawks are usually lighter in mass than their adult counterparts despite averaging somewhat longer winged and tailed. Male red-tailed hawks may weigh from 690 to 1,300 g (1.52 to 2.87 lb) and females may weigh between 801 and 1,723 g (1.766 and 3.799 lb) (the lowest figure from a migrating female immature from Goshute Mountains, Nevada, the highest from a wintering female in Wisconsin). Some sources claim the largest females can weigh up to 2,000 g (4.4 lb) but whether this is in reference to wild hawks (as opposed to those in captivity or used for falconry) is not clear.[24] The largest known survey of body mass in red-tailed hawks is still credited to Craighead & Craighead (1956), who found 100 males to average 1,028 g (2.266 lb) and 108 females to average 1,244 g (2.743 lb). However, these figures were apparently taken from labels on museum specimens, apparently from natural history collections in Wisconsin and Pennsylvania, without note to the region, age or subspecies of the specimens. However, 16 sources ranging in sample size from the aforementioned 208 specimens to only four hawks in Puerto Rico (with 9 of the 16 studies of migrating red-tails), showed that males weigh a mean of 860.2 g (1.896 lb) and females weigh a mean of 1,036.2 g (2.284 lb), about 15% lighter than prior species-wide published weights. Within the continental United States, average weights of males can range from 840.8 g (1.854 lb) (for migrating males in Chelan County, Washington) to 1,031 g (2.273 lb) (for male hawks found dead in Massachusetts) and females ranged from 1,057.9 g (2.332 lb) (migrants in the Goshutes) to 1,373 g (3.027 lb) (for females diagnosed as B. j. borealis in western Kansas). Size variation in body mass reveals that the red-tailed hawks typically varies only a modest amount and that size differences are geographically inconsistent. Racial variation in average weights of great horned owls (Bubo virginianus) show that mean body mass is nearly twice (the heaviest race is about 36% heavier than the lightest known race on average) as variable as that of the hawk (where the heaviest race is only just over 18% heavier on average than the lightest). Also, great horned owls correspond well at the species level with Bergmann’s rule.

 

Male red-tailed hawks can reportedly measure 45 to 60 cm (18 to 24 in) in total length, females measuring 48 to 65 cm (19 to 26 in) long. The wingspan typically can range from 105 to 141 cm (3 ft 5 in to 4 ft 8 in), although the largest females may possible span up to 147 cm (4 ft 10 in). In the standard scientific method of measuring wing size, the wing chord is 325.1–444.5 mm (12.80–17.50 in) long. The tail measures 188 to 258.7 mm (7.40 to 10.19 in) in length. The exposed culmen was reported to range from 21.7 to 30.2 mm (0.85 to 1.19 in) and the tarsus averaged 74.7–95.8 mm (2.94–3.77 in) across the races. The middle toe (excluding talon) can range from 38.3 to 53.8 mm (1.51 to 2.12 in), with the hallux-claw (the talon of the rear toe, which has evolved to be the largest in accipitrids) measuring from 24.1 to 33.6 mm (0.95 to 1.32 in) in length.

 

Identification:

Although they overlap in range with most other American diurnal raptors, identifying most mature red-tailed hawks to species is relatively straightforward, particularly if viewing a typical adult at a reasonable distance. The red-tailed hawk is the only North American hawk with a rufous tail and a blackish patagium marking on the leading edge of its wing (which is obscured only on dark morph adults and Harlan’s hawks by similarly dark colored feathers).

 

Other larger adult Buteo in North America usually have obvious distinct markings that are absent in red-tails, whether the rufous-brown “beard” of Swainson's hawks (Buteo swainsonii) or the colorful rufous belly and shoulder markings and striking black-and-white mantle of red-shouldered hawks (also the small “windows” seen at the end of their primaries). In perched individuals, even as silhouettes, the shape of large Buteos may be distinctive, such as the wingtips overhanging the tail in several other species, but not in red-tails. North American Buteos range from the dainty, compact builds of much smaller Buteos, such as broad-winged hawk (Buteo platypterus) to the heavyset, neckless look of ferruginous hawks or the rough-legged buzzard which has a compact, smaller appearance than a red-tail in perched birds due to its small bill, short neck and much shorter tarsus, while the opposite effect occurs in flying rough-legs with their much bigger wing area.

 

In flight, most other large North American Buteo are distinctly longer and slenderer winged than red-tailed hawks, with the much paler ferruginous hawk having peculiarly slender wings in relation to its massive, chunky body. Swainson's hawks are distinctly darker on the wing and ferruginous hawks are much paler winged than typical red-tailed hawks. Pale morph adult ferruginous hawk can show mildly tawny-pink (but never truly rufous) upper tail, and like red-tails tend to have dark markings on underwing-coverts and can have a dark belly band but compared to red-tailed hawks have a distinctly broader head, their remiges are much whiter looking with very small dark primary tips, they lack the red-tail’s diagnostic patagial marks and usually (but not always) also lack the dark subterminal tail-band, and ferruginous have a totally feathered tarsus. With its whitish head, the ferruginous hawk is most similar to Krider's red-tailed hawks, especially in immature plumage, but the larger hawk has broader head and narrower wing shape and the ferruginous immatures are paler underneath and on their legs. Several species share a belly band with the typical red-tailed hawk but they vary from subtle (as in the ferruginous hawk) to solid blackish, the latter in most light-morph rough-legged buzzards. More difficult to identify among adult red-tails are its darkest variations, as most species of Buteo in North America also have dark morphs. Western dark morph red-tails (i.e. calurus) adults, however, retain the typical distinctive brick-red tail which other species lack, which may stand out even more against the otherwise all chocolate brown-black bird. Standard pale juveniles when perched show a whitish patch in the outer half of the upper surface of the wing which other juvenile Buteo lack. The most difficult to identify stages and plumage types are dark morph juveniles, Harlan’s hawk and some Krider’s hawks (the latter mainly with typical ferruginous hawks as aforementioned). Some darker juveniles are similar enough to other Buteo juveniles that it has been stated that they "cannot be identified to species with any confidence under various field conditions." However, field identification techniques have advanced in the last few decades and most experienced hawk-watchers can distinguish even the most vexingly plumaged immature hawks, especially as the wing shapes of each species becomes apparent after seeing many. Harlan’s hawks are most similar to dark morph rough-legged buzzards and dark morph ferruginous hawks. Wing shape is the most reliable identification tool for distinguishing the Harlan’s from these, but also the pale streaking on the breast of Harlan’s, which tends to be conspicuous in most individuals, and is lacking in the other hawks. Also dark morph ferruginous hawks do not have the dark subterminal band of a Harlan’s hawk but do bear a black undertail covert lacking in Harlan’s.

  

AB2A8602-3_fCAFlkr2

Yaacov Agam is an Israeli sculptor and experimental artist best known for his contributions to optical and kinetic art.

 

Great Agam's Fountain is one of refreshing places in huge futuristic kingdom of concrete, steel and glass.

 

This fountain was designed by Yaacov Agam, a key figure in the history and development of the genre known as kinetic art. Born in Palestine, Agam moved to Paris in the early 1950s where he has spent much of his life. The musical fountain seen here was created in 1977.

 

The fountain's pool is made of polymorphic mosaic surface. It is comprised of 66 vertical water jets shooting water up to 14 meters. The fountain, at night beautifuly illuminated, was further enhanced with the addition of five new triple tulip jets in 1991.

Cuculus canorus

[order] Cuculiformes | [family] Cuculidae | [latin] Cuculus canorus | [UK] Cuckoo | [FR] Coucou gris | [DE] Kuckuck | [ES] Cuco Europeo | [IT] Cuculo eurasiatico | [NL] Koekoek | [IRL] Cuach

 

Measurements

spanwidth min.: 54 cm

spanwidth max.: 60 cm

size min.: 32 cm

size max.: 36 cm

Breeding

incubation min.: 11 days

incubation max.: 12 days

fledging min.: 17 days

fledging max.: 17 days

broods 15

eggs min.: 1

eggs max.: 25

 

Status: Widespread summer visitor to Ireland from April to August.

 

Conservation Concern: Green-listed in Ireland. The European population is currently evaluated as secure.

 

Identification: Despite its obvious song, relatively infrequently seen. In flight, can be mistaken for a bird of prey such as Sparrowhawk, but has rapid wingbeats below the horizontal plane - ie. the wings are not raised above the body. Adult male Cuckoos are a uniform grey on the head, neck, back, wings and tail. The underparts are white with black barring. Adult females can appear in one of two forms. The so-called grey-morph resembles the adult male plumage, but has throat and breast barred black and white with yellowish wash. The rufous-morph has the grey replaced by rufous, with strong black barring on the wings, back and tail. Juvenile Cuckoos resemble the female rufous-morph, but are darker brown above.

 

Similar Species: Sparrowhawk

 

Call: The song is probably one of the most recognisable and well-known of all Irish bird species. The male gives a distinctive “wuck-oo”, which is occasionally doubled “wuck-uck-ooo”. The female has a distinctive bubbling “pupupupu”. The song period is late April to late June.

 

Diet: Mainly caterpillars and other insects.

 

Breeding: Widespread in Ireland, favouring open areas which hold their main Irish host species – Meadow Pipit. Has a remarkable breeding biology unlike any other Irish breeding species.

 

Wintering: Cuckoos winter in central and southern Africa.

 

To minimise the chance of being recognised and thus attacked by the birds they are trying to parasitize, female cuckoos have evolved different guises.

 

The common cuckoo (Cuculus canorus) lays its eggs in the nests of other birds. On hatching, the young cuckoo ejects the host's eggs and chicks from the nest, so the hosts end up raising a cuckoo chick rather than a brood of their own. To fight back, reed warblers (a common host across Europe) have a first line of defence: they attack, or ‘mob’, the female cuckoo, which reduces the chance that their nest is parasitized.

 

To deter the warbler from attacking, the colouring of the grey cuckoo mimics sparrow hawks, a common predator of reed warblers. However, other females are bright rufous (brownish-red). The presence of alternate colour morphs in the same species is rare in birds, but frequent among the females of parasitic cuckoo species. The new research shows that this is another cuckoo trick: cuckoos combat reed warbler mobbing by coming in different guises.

 

In the study, the researchers manipulated local frequencies of the more common grey colour cuckoo and the less common (in the United Kingdom) rufous colour cuckoo by placing models of the birds at neighbouring nests. They then recorded how the experience of watching their neighbours mob changed reed warbler responses to both cuckoos and a sparrow hawk at their own nest.

 

They found that reed warblers increased their mobbing, but only to the cuckoo morph that their neighbours had mobbed. Therefore, as one cuckoo morph increases in frequency, local host populations will become alerted specifically to that morph. This means the alternate morph will be more likely to slip past host defences and lay undetected. This is the first time that ‘social learning’ has been documented in the evolution of mimicry as well as the evolution of different observable characteristics - such as colour - in the same species (called polymorphism).

 

From the University of Cambridge “When mimicry becomes less effective, evolving to look completely different can be a successful trick. Our research shows that individuals assess disguises not only from personal experience, but also by observing others. However, because their learning is so specific, this social learning then selects for alternative cuckoo disguises and the arms race continues.”.

“It’s well known that cuckoos have evolved various egg types which mimic those of their hosts in order to combat rejection. This research shows that cuckoos have also evolved alternate female morphs to sneak through the hosts' defences. This explains why many species which use mimicry, such as the cuckoo, evolve different guises.”

 

Physical characteristics

Forests and woodlands, both coniferous and deciduous, second growth, open wooded areas, wooded steppe, scrub, heathland, also meadows, reedbeds. Lowlands and moorlands and hill country to 2 km.

 

Habitat

Forests and woodlands, both coniferous and deciduous, second growth, open wooded areas, wooded steppe, scrub, heathland, also meadows, reedbeds. Lowlands and moorlands and hill country to 2 km. Food and Feeding

 

Other details

Cuculus canorus is a widespread summer visitor to Europe, which accounts for less than half of its global breeding range. Its European breeding population is very large (>4,200,000 pairs), and was stable between 1970-1990. Although there were declines in many western populations-most notably France-during 1990-2000, most populations in the east, including key ones in Russia and Romania, were stable, and the species underwent only a slight decline overall

 

Feeding

Diet based on insects, mainly caterpillars, also dragonflies, mayflies, damselflies, crickets, and cicadas. Sometimes, spiders, snails, rarely fruit. Preys on eggs and nestling of small birds.

 

Conservation

This species has a large range, with an estimated global Extent of Occurrence of 10,000,000 km². It has a large global population, including an estimated 8,400,000-17,000,000 individuals in Europe (BirdLife International in prep.). Global population trends have not been quantified, but populations appear to be stable so the species is not believed to approach the thresholds for the population decline criterion of the IUCN Red List (i.e. declining more than 30% in ten years or three generations). For these reasons, the species is evaluated as Least Concern.

 

Breeding

May-Jun in NW Europe, Apr-May in Algeria, Apr-Jul in India and Myanmar. Brood-parasitic, hosts include many insectivorous songbird species, like: flycatchers, chats, warblers, pipits, wagtails and buntigs. Often mobbed by real or potential hosts near their nests. Eggs polymorphic in color and pattern, closely match those of host in color and pattern. Nestling period 17-18 days, evicts host's eggs and chicks.

  

Migration

Migratory in N of range, arriving in SW Britain mainly Apr - May, when occasionally recorded in small parties, and even in one flock of 50+ birds; also seasonal in hill country from Assam and Chin Hills to Shan States, where present Mar - Aug. Resident in tropical lowland areas of S Asia. Winter resident in sub-Saharan Africa and in Sri Lanka. W Palearctic populations migrate to Africa, where a Dutch-ringed juvenile found in Togo in Oct and a British-ringed juvenile found in Cameroon in Jan; migrants appear in N Senegal as early as late Jul through Oct; in W Africa nearly all records are in autumn ( Sept - Dec), birds apparently continuing on to C & S Africa. Race bangsi occurs on passage in W Africa, and winters S of equator from W Africa to L Tanganyika. Asian populations of nominate canorus and bakeri winter in India, SE Asia and Philippines, also in Africa, but the extent of migration of Asian birds to Africa is unknown; some subtelephonus migrate through Middle East and occur in winter from Uganda and E Zaire to Zimbabwe, Mozambique and Natal. Mainly a passage migrant in Middle East, though some breed in region. Migrants also appear on islands in W Indian Ocean ( Seychelles, Aldabra). Nominate canorus accidental in Iceland, Faeroes, Azores, Madeira, Canary Is and Cape Verde Is, rarely also Alaska and eastern N America; one record of canorus in Indonesia, off W Java in winter. Autumn migration starts in August and continues until October. The main passage through Egypt is in September and the first half of October, with a peak in the third week of September (Goodman & Meininger 1989). Southward movement through Africa lasts from September to December and is linked to the occurrence of rainfall and the growth of cover.

Banded Peacock.

Mariposa de Fátima.

 

This species has two polymorphic forms, one with white bands and one with yellow bands. Both morphs have a median band that runs through both the front and rear wings with seven small dots near the apex and the area behind the median of the front wing. There is no distinction between males and females other than the fact that females tend to have more diffuse border pattern elements.

 

Esta especie tiene dos formas polimórficas, una con bandas blancas y otra con bandas amarillas. Ambos morfos tienen una banda mediana que atraviesa tanto el ala delantera como la trasera con siete pequeños puntos cerca del ápice y el área posterior a la mediana del ala anterior. No hay distinción entre machos y hembras además del hecho de que las hembras tienden a tener elementos de patrón de bordes más difusos.

dead egos feeding other egos

I think this was a pregant female as she had a rather large belly. Photographed sunbathing on a stile along the coastal path at Saeford Haven, East Sussex.

 

The viviparous lizard or common lizard, Zootoca vivipara (formerly Lacerta vivipara), is a Eurasian lizard. It lives farther north than any other reptile species, and most populations are viviparous (giving birth to live young), rather than laying eggs as most other lizards do. It is the only species in the monotypic genus Zootoca.

 

Zootoca vivipara can be seen in a variety of different colors. Female Zootoca vivipara undergo color polymorphism (biology) more commonly than males. A female lizard's display differs in ventral coloration, ranging from pale yellow to bright orange and a mixed coloration. There have been many hypothesis for the genetic cause of this polymorphic coloration. These hypothesis test for coloration due to thermoregulation, predator avoidance, and social cues, specifically sexual reproduction. Through an experiment conducted by Vercken et al., color polymorphism in viviparous lizard is caused by social cues, rather than the other hypotheses. More specifically, the ventral coloration that is seen in female lizards is associated with patterns of sexual reproduction and sex allocation.

 

The length of the body is less than 12 cm (5 in) (excluding the tail). The tail is up to twice as long as the body, although it is often partially or wholly lost. The limbs are short, and the head is rather round. Males have more slender bodies than females. The neck and the tail are thick. The collar and other scales seem jagged.

 

The colour and patterning of this species is variable. The main colour is typically medium brown, but it can be also grey, olive brown or black. Females may have dark stripes on their flanks and down the middle of their backs. Sometimes females also have light-coloured stripes, or dark and light spots along the sides of their backs. Most males and some females have dark spots in their undersides. Males have brightly coloured undersides – typically yellow or orange, but more rarely red. Females have paler, whitish underparts. The throat is white, sometimes blue.

Illustration for a comparative ecophylogenetic analysis of local myrmecofaunas, based on r/K selection theory and intra / interspecific parabiosis / lestobiosis, particularly focused on allochthonous and invasive species.

 

[Pheidole Westwood 1839: 1,160+†7 spp]

 

Pheidole (from φειδωλός) is a hyperdiverse and polymorphic genus, abundant in the Neotropics and in many other areas of the world. While there are other genera with discrete worker castes, none have radiated to the extent of Pheidole. Wilson's 2003 treatment of the New World spp and other more recent taxophylogenetic studies have brought some clarity to defining the boundaries of many spp. Still, much work remains before it can be claimed that Pheidole has few new spp to be described and that the taxonomy of this group is definitively complete.

 

REFERENCES

 

E.M. Sarnat​ & al. 2015: Introduced Pheidole of the world.

E.P. Economo & al. 2015: Expansion and taxon cycles in Pheidole.

E.P. Economo & al. 2014: Global phylogenetics of Pheidole.

M.R. Pie​ & M.K. Tschá 2013: Evolution of ant worker morphology.

The skin's reaction to sunlight exposure most commonly consists of pink or red raised spots that are quite itchy. Although people can get the red welts anywhere, usually the arms but also the chest and legs, the skin on the face typically does not break out. The rash appears anywhere from a few hours to a few days after the sun-exposure. The spots slowly fade after you stay out of the sun for a few days (further sun exposure makes the rash worse). In more severe cases the skin can form blisters or large patches of dry, red skin. Other additional symptoms can include chills, headache, nausea, or a generally "sick" feeling.

 

p.s: i never scary with wrinkles... i just scary with my polymorphic light eruption...

i cant sleep all night when i got this :(

Butterflies are part of the class of insects in the order Lepidoptera, along with the moths. Adult butterflies have large, often brightly coloured wings, and conspicuous, fluttering flight. The group comprises the large superfamily Papilionoidea, along with two smaller groups, the skippers (superfamily Hesperioidea) and the moth-butterflies (superfamily Hedyloidea). Butterfly fossils date to the Palaeocene, about 56 million years ago. Butterflies are often polymorphic, and many species make use of camouflage, mimicry and aposematism to evade their predators.

 

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リコリス・ロンギチュバ ‘ピュア・ホワイト’

Lycoris longituba Y.C.Hsu et G.J.Fan, 1974 ‘Pure White’

(Imported at JAPAN, 1979 from China, By Mr. Kaneko, Japan.)

Very rare plants. 2n=16=6M+10T

 

Endemic to China (Jiangsu and Anhui). Very polymorphic species. Variable in flower color and shape. Plants with yellow flowers classified as a variety, "Lycoris longituba Y.C.Hsu et G.J.Fan, 1974". Very long tepatube and its fragrance are marked chracteristics of this species. Leaves lanceolate, to 63 cm long and to 4 cm wide, somewhat fleshy and pale green, appearing in early spring and wither up in May. Scape, 60-80 cm hight, appearing in July to August. Spathe, to 5 cm long. Pedicel 1.5-4.5 cm, tepaltube 4.5-6.0 cm. Tepals 7-9.5 cm long, . Stamens 6-7.5 cm, shorter than tepals. Style 7-9.5 cm, nearly equal to or slightly exceeding tepals. Before that it was far carrying out this plant in a scientific statement, it was indicated by U.S. an Advanced Horticulturalist Mr. Sam Coldwell from that it was a new species. The formal scientific statement of "Lycoris longituba" was announced by Y.C.Hsu & G.J.Fan in 1974. However, this plant was introduced into Japan and was already grown in the 1930s. It by misconception of scholar Dr. Inariyama (1948) of Japan "Lycoris x straminea Lindl., 1848" said. When it depended on research of scholar Dr. Kurita of Japan, it became clear that the individual of this plants is one of the variations of "Lycoris longituba Y.C.Hsu et G.J.Fan, 1974"

 

この植物は、学術記載をされるはるか以前から、アメリカの先進的園芸家・Mr. Sam Coldwell により新種ではないかと指摘をされていた。「Lycoris longituba」の正式な学術記載は Y.C.Hsu & G.J.Fan によって1974年になされた。しかし、既に1930年代にこの植物は日本に渡来して栽培されていた。日本の学者 Dr. Inariyama(1948)の誤認で「Lycoris × straminea Lindl., 1848」と言われていた。日本の学者 Dr. Kurita の研究に依ると、この画像の個体は「Lycoris longituba Y.C.Hsu et G.J.Fan, 1974」のバリエーションの一つだと判明した。

 

この個体はリコリス中でも最も大型の部類で、主に種子繁殖をし、非常に自家受粉しやすいが、その反面、分球はしずらく、1球が10球までになるには凡そ30年はかかった。ヒガンバナが30年で1000球に分けつする事と比較すると恐ろしいほどスローモーな分けつ力である。此の個体は、金子氏が40年前に中国から導入したヒガンバナ類の中から選抜した個体で、他に麦藁色の個体が明治期にもたらされていたが、在来の物より鑑賞価値は高い。春出葉型なので、短い期間しか葉が無い為、その間には良く日光に当て肥培をすることが肝心で有る。自然交雑種のナツズイセンの片親である事がDr. Kurita の研究で判明した。自然界でのリコリス・ロンギチュバは、白、ピンク、麦藁色など幅があり、黄花の物は別途変種扱いされている。此の個体自体は、蘂にやや色が乗るが、これほどに白い個体は少なく、殆どがやや濁っているのが殆どであり、選抜した意味は大きい。本種は大球性で他のリコリスよりもやや深く土中に球根が潜る。開花は、是から得られたナツズイセンよりも1ヶ月早く、7月中には開花する。

  

SONY NEX-7

Nikon Ai AF Micro Nikkor ED 200mm F4D (IF)

(Product Year : 1993)

Mais uma vez, muito obrigado ao Marcelo Cazani (Marcazani) pela identificação deste pássaro.

Once more, thank you very much Marcelo Cazani (Marcazani) for the identification of this bird.

 

A text, in english, from Wikipedia, the free encyclopedia:

See at en.wikipedia.org/wiki/American_Kestrel

Jump to: navigation, search

American Kestrel

Conservation status

 

Least Concern (IUCN 3.1)

Scientific classification

Kingdom: Animalia

Phylum: Chordata

Class: Aves

Order: Falconiformes

Family: Falconidae

Genus: Falco

Species: F. sparverius

Binomial name

Falco sparverius

Linnaeus, 1758

Synonyms

 

Cerchneis sparverius

Tinnunculus sparverius

 

The American Kestrel (Falco sparverius) is a small falcon. This bird was (and sometimes still is) colloquially known in North America as the "Sparrow Hawk". This name is misleading because it implies a connection with the Eurasian Sparrowhawk Accipiter nisus, which is unrelated; the latter is an accipiter rather than a falcon. Though both are diurnal raptors, they are only distantly related.

American Kestrels are widely distributed across the Americas. Their breeding range extends from central and western Alaska across northern Canada to Nova Scotia, and south throughout North America, into central Mexico, the Baja, and the Caribbean. They are local breeders in Central America and are widely distributed throughout South America.

Most of the birds breeding in Canada and the northern United States migrate south in the winter, although some males stay as year-round residents. It is a very rare vagrant to western Europe.

The American Kestrel is the smallest falcon in North America—about the size of an American Robin. Like all raptors, the American Kestrel is sexually dimorphic, although there is some overlap within the species. The female ranges in length from 23 to 28 centimeters (9-11 inches) with a wingspan of 53–61 centimeters (21–24 inches) and weighs an average of 120 grams (4.2 ounces). The length of the male varies between 20–25 centimeters (8–10 inches) with a wingspan ranging from 51–56 centimeters (20–22 inches) and weighing an average of 111 grams (3.9 ounces). These subtle differences are often difficult to discern in the field.

The coloration of the feathers, however, greatly varies between the sexes. Males have blue-grey secondary feathers on their wings, while the undersides are white with black barring. The back is rufous in coloration, with barring on the lower half. The belly and flanks are white with black spotting. The tail is also rufous, except for the outer rectrix set, which is white with a black subterminal band.

The back of the female American Kestrel is rufous with dark brown barring. The wings exhibit similar coloration and patterning to the back. The undersides of the females are white with rufous streaking. The tail of the female is noticeably different from the male, being rufous in color with numerous narrow dark brown or black bars. Juveniles exhibit coloration patterns similar to the adults.

In both sexes, the head is white with a bluish-grey top. There are also two narrow, vertical black facial markings on each side of the head; one below the eyes and one on the rear portion of the auriculars. Two black spots (ocelli) can be found on each side of the white or orangish nape. The wings are moderately long, fairly narrow, and taper to a point. While perched, the wingtips are noticeably shorter than the tail tip.

The American Kestrel has three basic vocalizations - the "klee" or "killy", the "whine", and the "chitter." The "klee" or "killy" is usually delivered as a rapid series - "killy, killy, killy, killy" when the kestrel is upset or excited. It is used at the apex of the dive display, during fights with other kestrels, and after unsuccessful hunting attempts.

The "whine" may last as long as one to two minutes and may be single or treble. The more intense the situation, the more likely the "whine" will move toward the treble extreme. "Whines" are given during courtship feeding and copulation. The treble whine is heard from breeding females and fledged hungry young.

The "chitter" is the most frequent vocalization in male - female interactions. Its volume and duration depends upon the stress or excitement of the situation. It is associated with friendly approaches and bodily contact between the sexes during breeding season. Occasionally a "chitter" follows a "whine."

Calling occurs throughout the day. Nestlings at two weeks can produce all three vocalizations. Female kestrels tend to have slightly lower pitched and harsher voices than males.

This bird is apparently not a true kestrel. mtDNA cytochrome b sequence analysis (Groombridge et al. 2002) indicates a Late Miocene split[1] between the ancestors of the American Kestrel, and those of the Common Kestrel and its closest relatives. The color pattern with its large areas of brown is reminiscent of kestrels, but the coloration of the head - notably the black ear patch, which is not found in any of the true kestrels - and the male's extensively gray wings are suggestive of a closer relationship with the hobbies, an informal grouping of falcons of usually average size.

Species such as the Merlin and the Aplomado Falcon are proposed as possible close relatives. Indeed, the Merlin is a highly polymorphic bird and although its grey tail and back are distinctive, certain morphs are the only birds that might conceivably be confused with American Kestrels. Conclusive evidence is lacking, and what can be said at present judging from the fairly noninclusive DNA sequence studies[2] is that the general relationships of the present species seem to lie with a number of rather basal "hobby" lineages, such as the Merlin and Aplomado Falcon mentioned already, or the Red-footed and Amur Falcons - or even the Peregrine Falcon lineage with its large species.

The American Kestrel is not very closely related to any of these groups, although it might be closer to the Aplomado Falcon (and its presumed close relatives, the Bat and Orange-breasted Falcons) than to any other living species (Wink et al. 1998) - an association that is also better supported by biogeography than a close relationship with the exclusively Old World true kestrels. It is nonetheless highly distinct in morphology from any of these and, interestingly, has a syrinx similar to the Peregrine and the hierofalcons[3].

In conclusion, until better evidence is available, it is best considered part of a radiation of falcon lineages that diversified around the North Atlantic at the end of the Miocene. Though several fossils of small falcons arte known from North America at roughly the correct time, the earliest testimony of the American Kestrel lineage is Pleistocene remains of the living species (Brodkorb 1964).

American Kestrels are found in a variety of habitats including parks, suburbs, open fields, forest edges and openings, alpine zones, grasslands, marshes, open areas on mountainsides, prairies, plains, deserts with giant cacti, and freeway and highway corridors.

In addition to requiring open space for hunting, American Kestrels seem to need perches for hunting from, cavities for nesting (either natural or man-made), and a sufficient food supply.

The American Kestrel is the only North American falcon to habitually hover with rapid wing beats, keeping its head motionless while scanning the ground for prey. The kestrel commonly perches along fences and powerlines. It glides with flat wings and wingtips curved upward. It occasionally soars in circles with its tail spread and its wings flat.

This falcon species is not long-lived. The oldest banded wild bird was 11 years and seven months old while a captive lived 17 years. A mortality rate average of 57 percent was found. First year mortality rates have declined since 1945 with a decrease in shooting. Major causes of death include collision with traffic, illegal shooting, and predation by other raptors, including the Red-tailed Hawk, Northern Goshawk, Cooper's Hawk, Peregrine Falcon, Barn Owl, and Great Horned Owl.

In summer, kestrels feed largely on grasshoppers, dragonflies, lizards, mice, and voles. They will also eat other small birds. Wintering birds feed primarily on rodents and birds. The birds characteristically hunt along roadsides from telephone wires, fence posts, trees or other convenient perches when not flying in search of food. When they are flying and looking for food they frequently hover with rapid wingbeats.

Because it feeds on both insects and vertebrates, the American Kestrel maintains fairly high population densities. It has a small breeding home range, from 1.75 square miles (4.5 km²) to 2 square miles (5.2 km²). Territory size has been estimated at 269 acres (1.1 km²) to 321 acres (1.3 km²) with much larger wintering home ranges.

Several hunting techniques are used by the American Kestrel. It will hover over one spot—when prey is sighted the kestrel will partly fold its wings and drop lower once or several times before striking. When the prey disappears the falcon will glide in a semicircle before turning back into the wind to hover again. It will also soar in circles, or figure eights, using the same stooping tactics as when hovering.

The kestrel commonly hunts from elevated perch sites, waiting for prey to move on the ground. The kestrel bobs its head and pumps its tail just before attacking.

Other prey capture techniques include direct pursuit, landing and flushing prey from the ground (especially for grasshoppers)and then taking them in flight, capturing flying insects from an elevated perch, and nest robbing including the burrows of Bank Swallows and the nests of Cliff Swallows. It is also an occasional bat catcher, taking bats from their tree roosts, or striking bats in flight from above or as the bats leave or enter caves. The kestrel will kill and cache food items.

The American Kestrel occasionally robs others of the same species. It has also been known to rob a shrike of its prey. Kestrels sometimes harass other hawks, and even Golden Eagles, in flight.

American Kestrels form pairs in which the bond is strong, tending toward permanence. Returning migrants commonly re-establish territories held the previous year. In one study[citation needed], a pair nested in the same tree for six consecutive years. Nesting occurs from late spring to late summer in North America, with incubation underway by the end of May[4]; in tropical South America the birds breed roughly from June onwards through to September or so[5].

Courtship begins shortly after the male establishes a territory. In early courtship, he may give the "dive display", a series of climbs and dives 33 to 66 feet (10 to 20 m) with 3-5 "klee" notes given near the peak of each climb. He may present the female with food during courtship feeding. He may entice her to the nest site by calling. He may "flutter-glide" toward her with quick and shallow wing beats while carrying food and she may also beg for food by flutter-gliding. The female initiates copulation by bowing with her tail in line with her body or slightly raised.

A cavity nester, American Kestrels will use holes in trees, rock cavities and crevices in cliffs, artificial nest boxes, or small spaces in buildings. The number of suitable breeding cavities limits this species' breeding density. The American Kestrel has adapted well to nest boxes. In one program, nest boxes were fixed to the backs of signs along a freeway thus allowing kestrels to breed in areas formerly devoid of nest sites. Pairs nesting in boxes on poles have much higher nesting success than pairs using boxes on trees. No nest is built inside. In nest boxes sawdust and wood shavings may be a suitable substrate for the eggs. Males and females defend the nest against intruders, with the male maintaining a small core territory and the female defending the nest cavity directly rather the surroundings[5].

Both sexes take turns incubating their eggs, a very rare situation among North American birds of prey where the female usually incubates exclusively. Correspondingly, both sexes develop bare oval patches on each side of their breasts where the warm bare skin can contact the eggs for warming. Eggs hatch 29 to 31 days after being laid.[6] There are from three to seven eggs laid, but four to six are average.[6] The eggs are typically short elliptical in shape, and are white or pinkish-white with an even covering of fine spots and flecks of brown shades, occasionally concentrating as a ring or a cap. They will renest if the first nest fails and have been reported to raise 2 broods per year in some of the southern states.

The young grow very quickly, becoming noisy between day 11 and 14 and assuming adult weight in about 2.5 weeks. The young fledge in 30 to 31 days.[7] [6]Early fledgling behavior varies. Broods typically stay together for a week or two. Some broods remain close to the nest area for a week or two while others travel throughout the parents' home range. Generally, young do not disperse more than 0.6 miles (one km) away from the nest area until two to four weeks old. Young disperse as hunting skills develop. Occasionally groups of older juveniles from various broods join together into flocks.

The American Kestrel can be double-brooded, particularly in the southern United States, in areas of abundant small mammals. Replacement clutches can be laid.

 

Um texto em português:

Falcão-americano ou quiriquiri (Falco sparverius) é um pequeno falcão (23-27 cm de comprimento e 85-140 g de peso). Tem uma ampla área de distribuição: desde o Alasca e Norte do Canadá até à ponta Sul da América do Sul (Terra do Fogo). Tem asas azul acizentadas. O dorso é avermelhado pontuado de preto. A cauda também é avermelhada possuindo uma larga lista preta. Na face possui 2 listas verticais que começam junto aos olhos e seguem para baixo. É um predador de pequeno tamanho, alimentando-se de insetos e microvertrebrados como roedores e pequenos pássaros; ocasionalmente caça morcegos.

Falcão é o nome genérico dado a várias aves da família Falconidae, mais estritamente aos animais classificados dentro do género Falco. O que diferencia os falcões das demais aves de rapina é o fato de terem evoluído no sentido de uma especialização no voo em velocidade (em oposição ao voo planado das águias e abutres e ao voo acrobático dos gaviões), facilitado pelas asas ponteagudas e finas, favorecendo a caça em espaços abertos – daí o fato dos falcões não serem aves de ambientes florestais, preferindo montanhas e penhascos, pradarias, estepes e desertos. Os falcões podem ser identificados, aliás, pelo fato de não planarem em correntes termais, como outras aves de rapina. O falcão-peregrino, especializado na caça de aves médias e grandes em voo, pode atingir 300 km/h em voo picado e é o animal mais rápido da terra. Diferentemente das águias e gaviões, que matam suas presas com os pés, os falcões utilizam as garras apenas para apreenderem a presa, matando-a depois com o bico por desconjuntamento das vértebras, para o que possuem um rebordo em forma de dente na mandíbula superior.

 

Na Idade Média, os falcões eram apreciados como animais de caça acessíveis apenas à elite.

(freshly eclosed from collected chrysalis)

 

The Great Mormon (Papilio memnon) is a large butterfly that belongs to the swallowtail family and is found in southern Asia. It is widely distributed and has thirteen subspecies. Furthermore, within these groupings, there are four male and many female forms (both tailed and tailless), the females being highly polymorphic and many of them being mimics of unpalatable butterflies. This species has been studied extensively for understanding the genetic basis for polymorphy and Batesian mimicry. As many as twenty-six female forms are reported.

 

Pu'er, Yunnan, China

 

see comments for caterpillar image…..

The Nikitin-Schyevchyenko IS series, (Istrebitel Skladnoi – folding fighter), were single seat polymorphic fighters designed and produced in the USSR from 1938.

 

A test pilot in Nikitins bureau decided to look in to the feasibility of an aircraft, that took off and landed as a bi plane but which had the ability for the lower wing to fold into the upper- offering increased speed and maneuverability as well as STOL capabilities. The first version was to be armed with 4 ShKas machine guns, when development continued this was changed to 2 ShKas and 2 Berezin UB, and the final version was intended to have tricycle landing gear. However, there were man design problems and the design afforded no more performance than fighters then entering production, namely the Yak 1, Mig 1, and Lagg-1.

 

Among other problems that the plane had, the lower wing became stick inside of the top wing on a few test flights and because the landing gear was mounted in this lower wing, belly landings had to be made.

Took a little bit of research as this one was perplexing. This is an Eastern Tiger Swallowtail butterfly female that is a "dark morph" of the species. (source)

 

Females of this species are bimorphic as they can be a yellow morph - making them look very similar to the males - or dark morph where the yellow banding is replaced with black. This happens quite a lot in nature (Eastern Screech owls, for example).

The song thrush is omnivorous, eating a wide range of invertebrates, especially earthworms and snails, as well as soft fruit and berries. Like its relative, the blackbird, the song thrush finds animal prey by sight, has a run-and-stop hunting technique on open ground, and will rummage through leaf-litter seeking potential food items.

 

Land snails are an especially important food item when drought or hard weather makes it hard to find other food. The thrush often uses a favourite stone as an "anvil" on which to break the shell of the snail before extracting the soft body and invariably wiping it on the ground before consumption. Young birds initially flick objects and attempt to play with them until they learn to use anvils as tools to smash snails. The nestlings are mainly fed on animal food such as worms, slugs, snails and insect larvae.

 

The grove Snail (Cepaea nemoralis) is regularly eaten by the song thrush, and its polymorphic shell patterns have been suggested as evolutionary responses to reduce predation;[34] however, song thrushes may not be the only selective force involved.

 

The song thrush was described by German ornithologist Christian Ludwig Brehm in 1831, and still bears its original scientific name, Turdus philomelos. The generic name, Turdus, is the Latin for thrush, and the specific epithet refers to a character in Greek mythology, Philomela, who had her tongue cut out, but was changed into a singing bird. Her name is derived from the Ancient Greek Φιλο philo- (loving), and μέλος melos (song). The dialect names throstle and mavis both mean thrush, being related to the German drossel and French mauvis respectively. Throstle dates back to at least the fourteenth century and was used by Chaucer in the Parliament of Fowls. Mavis is derived via Middle English mavys and Old French mauvis from Middle Breton milhuyt meaning "thrush." Mavis (Μαβής) can also mean "purple" in Greek.

For more information please visit en.wikipedia.org/wiki/Song_thrush

 

Mocow disorder.

Banded Peacock.

Mariposa de Fátima.

 

This species has two polymorphic forms, one with white bands and one with yellow bands. Both morphs have a median band that runs through both the front and rear wings with seven small dots near the apex and the area behind the median of the front wing. There is no distinction between males and females other than the fact that females tend to have more diffuse border pattern elements.

 

Esta especie tiene dos formas polimórficas, una con bandas blancas y otra con bandas amarillas. Ambos morfos tienen una banda mediana que atraviesa tanto el ala delantera como la trasera con siete pequeños puntos cerca del ápice y el área posterior a la mediana del ala anterior. No hay distinción entre machos y hembras además del hecho de que las hembras tienden a tener elementos de patrón de bordes más difusos.

The viviparous lizard or common lizard, Zootoca vivipara (formerly Lacerta vivipara), is a Eurasian lizard. It lives farther north than any other reptile species, and most populations are viviparous (giving birth to live young), rather than laying eggs as most other lizards do. It is the only species in the monotypic genus Zootoca.

 

Zootoca vivipara can be seen in a variety of different colors. Female Zootoca vivipara undergo color polymorphism (biology) more commonly than males. A female lizard's display differs in ventral coloration, ranging from pale yellow to bright orange and a mixed coloration. There have been many hypothesis for the genetic cause of this polymorphic coloration. These hypothesis test for coloration due to thermoregulation, predator avoidance, and social cues, specifically sexual reproduction. Through an experiment conducted by Vercken et al., color polymorphism in viviparous lizard is caused by social cues, rather than the other hypotheses. More specifically, the ventral coloration that is seen in female lizards is associated with patterns of sexual reproduction and sex allocation.

 

The length of the body is less than 12 cm (5 in) (excluding the tail). The tail is up to twice as long as the body, although it is often partially or wholly lost. The limbs are short, and the head is rather round. Males have more slender bodies than females. The neck and the tail are thick. The collar and other scales seem jagged.

 

The colour and patterning of this species is variable. The main colour is typically medium brown, but it can be also grey, olive brown or black. Females may have dark stripes on their flanks and down the middle of their backs. Sometimes females also have light-coloured stripes, or dark and light spots along the sides of their backs. Most males and some females have dark spots in their undersides. Males have brightly coloured undersides – typically yellow or orange, but more rarely red. Females have paler, whitish underparts. The throat is white, sometimes blue.

(Wikipedia)

  

In the Great Mormon butterfly, papilio agenor agenor, the female is polymorphic having several forms which differ in colour and the presence or absence of "tails".

I've noticed only 2 forms in the females which visit our garden, e.g. this one with no tails and a black abdomen. This may be the "agenor" form.

কালিম । Common Mormon (Female; Form: romulus) - Papilio polytes

 

This female form of the Common Mormon mimics the Common Rose very closely but lacks red markings on body. This is the commonest form wherever the Common Rose flies. The female of the Common Mormon is polymorphic. In South Asia, it has three forms or morphs. These are as follows: cyrus, stichius, romulus.

 

The Common Mormon (Papilio polytes) is a common species of swallowtail butterfly (family: papilionidae) widely distributed across Asia. Seen round the year throughout India from plains up to 2000m. This butterfly is known for the mimicry displayed by the numerous forms of its females.

 

Host Plant: Putush, পুটুশ ( Lantana camara "Feston Rose")

Evergreen shrubs and perennial herbs with green leaves and spiny branches that bear globular flowers, intermittently year round. Flowerheads with yellow flowers in the center and fuchsia to purple flowers at the circumference. Poisonous plants though leaves are in traditional medicinal use.

 

Lantana camara is an invasive species and has covered large areas in India, Australia and much of Africa. It is believed to be indigenous to the Lower Rio Grande Valley of Texas in the United States.

 

Family: Papilionidae

 

Damodor River Bank, Durgapur

Monsoon Images of Bengal, India

 

Sigma 150-600mm 5-6.3 DG HSM OS Contemporary plus Nikon D7000

Vitis (grapevine) is a genus of 81 accepted species of vining plants in the flowering plant family Vitaceae. The genus consists of species predominantly from the Northern Hemisphere. It is economically important as the source of grapes, both for direct consumption of the fruit and for fermentation to produce wine. The study and cultivation of grapevines is called viticulture.

 

Most cultivated Vitis varieties are wind-pollinated with hermaphroditic flowers containing both male and female reproductive structures, while wild species are dioecious. These flowers are grouped in bunches called inflorescences. In many species, such as Vitis vinifera, each successfully pollinated flower becomes a grape berry with the inflorescence turning into a cluster of grapes. While the flowers of the grapevines are usually very small, the berries are often large and brightly colored with sweet flavors that attract birds and other animals to disperse the seeds contained within the berries.

 

Grapevines usually only produce fruit on shoots that came from buds that were developed during the previous growing season. In viticulture, this is one of the principles behind pruning the previous year's growth (or "One year old wood") that includes shoots that have turned hard and woody during the winter (after harvest in commercial viticulture). These vines will be pruned either into a cane which will support 8 to 15 buds or to a smaller spur which holds 2 to 3 buds.

 

Description

Flower buds are formed late in the growing season and overwinter for blooming in spring of the next year. They produce leaf-opposed cymes. Vitis is distinguished from other genera of Vitaceae by having petals which remain joined at the tip and detach from the base to fall together as a calyptra or 'cap'. The flowers are mostly bisexual, pentamerous, with a hypogynous disk. The calyx is greatly reduced or nonexistent in most species and the petals are joined together at the tip into one unit but separated at the base. The fruit is a berry, ovoid in shape and juicy, with a two-celled ovary each containing two ovules, thus normally producing four seeds per flower (or fewer by way of aborted embryos).

 

Other parts of the vine include the tendrils which are leaf-opposed, branched in Vitis vinifera, and are used to support the climbing plant by twining onto surrounding structures such as branches or the trellising of a vine-training system.

 

In the wild, all species of Vitis are normally dioecious, but under domestication, variants with perfect flowers appear to have been selected.

 

The genus Vitis is divided into two subgenera, Euvitis Planch. have 38 chromosomes (n=19) with berries borne on clusters and Muscadinia Planch. 40 (n=20) with small clusters.

 

Wild grapes can resemble the single-seeded Menispermum canadense (moonseed), which is toxic.

 

Species

Most Vitis species are found mostly in the temperate regions of the Northern Hemisphere in North America and eastern Asia, exceptions being a few in the tropics and the wine grape Vitis vinifera which originated in southern Europe and southwestern Asia. Grape species occur in widely different geographical areas and show a great diversity of form.

 

Their growth makes leaf collection challenging and polymorphic leaves make identification of species difficult. Mature grapevines can grow up to 48 centimetres (19 inches) in diameter at breast height and reach the upper canopy of trees more than 35 metres (115 feet) in height.

 

Many species are sufficiently closely related to allow easy interbreeding and the resultant interspecific hybrids are invariably fertile and vigorous. Thus the concept of a species is less well defined and more likely represents the identification of different ecotypes of Vitis that have evolved in distinct geographical and environmental circumstances.

 

The exact number of species is not certain. Plants of the World Online states 81 species are accepted, but lists 84. More than 65 species in Asia are poorly defined. Approximately 25 species are known in North America and just one, V. vinifera has Eurasian origins; some of the more notable include:

 

Vitis aestivalis, the summer grape, native to the Eastern United States, especially the Southeastern United States

Vitis amurensis, native to the Asian continent, including parts of Siberia and China

Vitis arizonica, The Arizona grape is native to Arizona, Utah, Nevada, California, New Mexico, Texas, and Northern Mexico.

Vitis berlandieri, native to the southern North America, primarily Texas, New Mexico and Arkansas. Primarily known for good tolerance against soils with a high content of lime, which can cause chlorosis in many vines of American origin

Vitis californica, the California wild grape, or Northern California grape, or Pacific grape, is a wild grape species widespread across much of California as well as southwestern Oregon

Vitis coignetiae, the crimson glory vine, a species from East Asia grown as an ornamental plant for its crimson autumn foliage

Vitis labrusca L., the fox grapevine, sometimes used for winemaking and for jam. Native to the Eastern United States and Canada. The Concord grape was derived by a cross with this species

Vitis riparia, the riverbank grapevine, sometimes used for winemaking and for jam. Native to the entire Eastern United States and north to Quebec

Vitis rotundifolia (syn. Muscadinia rotundifolia), the muscadine, used for jams and wine. Native to the Southeastern United States from Delaware to the Gulf of Mexico

Vitis rupestris, the rock grapevine, used for breeding of Phylloxera resistant rootstock. Native to the Southern United States

Vitis vinifera, the European grapevine. Native to the Mediterranean and Central Asia.

Vitis vulpina, the frost grape, native to the Eastern United States, from Massachusetts to Florida, and west to Nebraska, Kansas, and Texas Treated by some as a synonym of V. riparia.

 

Plants of the World Online also includes:

Vitis acerifolia Raf.

Vitis amoena Z.H. Chen, Feng Chen & WW.Y. Xie

Vitis baihuashanensis M.S.Kang & D.Z.Lu

Vitis balansana Planch.

Vitis bashanica P.C.He

Vitis bellula (Rehder) W.T.Wang

Vitis betulifolia Diels & Gilg

Vitis biformis Rose

Vitis blancoi Munson

Vitis bloodworthiana Comeaux

Vitis bourgaeana Planch.

Vitis bryoniifolia Bunge

Vitis × champinii Planch.

Vitis chunganensis Hu

Vitis chungii F.P.Metcalf

Vitis cinerea (Engelm.) Millardet

Vitis davidi (Rom.Caill.) Foëx

Vitis × doaniana Munson ex Viala

Vitis erythrophylla W.T.Wang

Vitis fengqinensis C.L.Li

Vitis ficifolia Bunge

Vitis flavicosta Mickel & Beitel

Vitis flexuosa Thunb.

Vitis girdiana Munson

Vitis hancockii Hance

Vitis heyneana Schult.

Vitis hissarica Vassilcz.

Vitis hui W.C.Cheng

Vitis jaegeriana Comeaux

Vitis jinggangensis W.T.Wang

Vitis jinzhainensis X.S.Shen

Vitis kaihuaica Z.H.Chen, Feng Chen & W.Y Xie

Vitis kiusiana Momiy.

Vitis lanceolatifoliosa C.L.Li

Vitis longquanensis P.L.Chiu

Vitis luochengensis W.T.Wang

Vitis menghaiensis C.L.Li

Vitis mengziensis C.L.Li

Vitis metziana Miq.

Vitis monticola Buckley

Vitis mustangensis Buckley

Vitis nesbittiana Comeaux

Vitis × novae-angliae Fernald

Vitis novogranatensis Moldenke

Vitis nuristanica Vassilcz.

Vitis palmata Vahl

Vitis pedicellata M.A.Lawson

Vitis peninsularis M.E.Jones

Vitis piasezkii Maxim.

Vitis pilosonervia F.P.Metcalf

Vitis popenoei J.L.Fennell

Vitis pseudoreticulata W.T.Wang

Vitis quinlingensis P.C.He

Vitis retordii Rom.Caill. ex Planch.

Vitis romanetii Rom.Caill.

Vitis ruyuanensis C.L.Li

Vitis saccharifera Makino

Vitis shenxiensis C.L.Li

Vitis shizishanensis Z.Y.Ma, J.Wen, Q.Fu & X.Q.Liu

Vitis shuttleworthii House

Vitis silvestrii Pamp.

Vitis sinocinerea W.T.Wang

Vitis sinoternata W.T.Wang

Vitis tiliifolia Humb. & Bonpl. ex Schult.

Vitis tsoi Merr.

Vitis wenchowensis C.Ling

Vitis wenxianensis W.T.Wang

Vitis wilsoniae H.J.Veitch

Vitis wuhanensis C.L.Li

Vitis xunyangensis P.C.He

Vitis yunnanensis C.L.Li

Vitis zhejiang-adstricta P.L.Chiu

There are many cultivars of grapevines; most are cultivars of V. vinifera. One of them includes, Vitis 'Ornamental Grape'.

 

Hybrid grapes also exist, and these are primarily crosses between V. vinifera and one or more of V. labrusca, V. riparia or V. aestivalis. Hybrids tend to be less susceptible to frost and disease (notably phylloxera), but wine from some hybrids may have a little of the characteristic "foxy" taste of V. labrusca.

 

The Latin word Vitis is feminine,[19] and therefore adjectival species names take feminine forms, such as V. vinifera.

 

Ecology

Phylloxera is an American root aphid that devastated V. vinifera vineyards in Europe when accidentally introduced in the late 19th century. Attempts were made to breed in resistance from American species, but many winemakers and customers did not like the unusual flavour profile of the hybrid vines. However, V. vinifera grafts readily onto rootstocks of the American species and their hybrids with V. vinifera, and most commercial production of grapes now relies on such grafts.

 

Commercial distribution

According to the UN's Food and Agriculture Organization (FAO), 75,866 square kilometres of the world is dedicated to grapes. Approximately 71% of world grape production is used for wine, 27% as fresh fruit, and 2% as dried fruit. A portion of grape production goes to producing grape juice to be used as a sweetener for fruits canned "with no added sugar" and "100% natural". The area dedicated to vineyards is increasing by about 2% per year.

 

Domestic cultivation

Grapevines are widely cultivated by gardeners, and numerous suppliers cater specifically for this trade. The plants are valued for their decorative foliage, often colouring brightly in autumn; their ability to clothe walls, pergolas and arches, thus providing shade; and their fruits, which may be eaten as dessert or provide the basis for homemade wines. Popular varieties include:-

 

Buckland Sweetwater' (white dessert)

'Chardonnay' (white wine)

'Foster's Seedling' (white dessert)

'Grenache' (red wine)

'Muscat of Alexandria' (white dessert)

'Müller-Thurgau' (white wine)

'Phoenix' (white wine)

'Pinot noir' (red wine)

'Regent' (red wine)

'Schiava Grossa' (red dessert)

'Seyval blanc' (white wine)

'Tempranillo' (red wine)

 

The following varieties have gained the Royal Horticultural Society's Award of Garden Merit:-

'Boskoop Glory' (dessert/wine)

'Brant' (black dessert)

'Claret Cloak' or 'Frovit' (ornamental)

'New York Muscat' (black dessert)

'Purpurea' (ornamental)

 

Uses

The fruit of several Vitis species are grown commercially for consumption as fresh grapes and for fermentation into wine. Vitis vinifera is the most important such species.

 

The leaves of several species of grapevine are edible and are used in the production of dolmades and Vietnamese lot leaves.

 

Culture

The grapevine (typically Vitis vinifera) has been used as a symbol since ancient times. In Greek mythology, Dionysus (called Bacchus by the Romans) was god of the vintage and, therefore, a grapevine with bunches of the fruit are among his attributes. His attendants at the Bacchanalian festivals hence had the vine as an attribute, together with the thyrsus, the latter often entwined with vine branches. For the same reason, the Greek wine cup (cantharos) is commonly decorated with the vine and grapes, wine being drunk as a libation to the god.

 

The grapevine has a profound symbolic meaning in Jewish tradition and culture since antiquity. It is referenced 55 times in the Hebrew Bible (Old Testament), along with grapes and wine, which are also frequently mentioned (55 and 19, respectively). It is regarded as one of the Seven Species, and is employed several times in the Bible as a symbol of the Israelites as the chosen people. The grapevine has a prominent place in Jewish rituals: the wine was given a special blessing, "creator of the fruit of the vine", and the Kiddush blessing is recited over wine or grape juice on Shabbat and Jewish holidays. It is also employed in various parables and sayings in rabbinic literature. According to Josephus and the Mishnah, a golden vine was hung over the inner chamber of the Second Temple. The grapevine is featured on Hasmonean and Bar Kokhba revolt coinage, and as a decoration in mosaic floors of ancient synagogues.

 

In Christian iconography, the vine also frequently appears. It is mentioned several times in the New Testament. We have the parable of the kingdom of heaven likened to the father starting to engage laborers for his vineyard. The vine is used as symbol of Jesus Christ based on his own statement, "I am the true vine (John 15:1)." In that sense, a vine is placed as sole symbol on the tomb of Constantia, the sister of Constantine the Great, and elsewhere. In Byzantine art, the vine and grapes figure in early mosaics, and on the throne of Maximianus of Ravenna it is used as a decoration.

 

The vine and wheat ear have been frequently used as symbol of the blood and flesh of Christ, hence figuring as symbols (bread and wine) of the Eucharist and are found depicted on ostensories. Often the symbolic vine laden with grapes is found in ecclesiastical decorations with animals biting at the grapes. At times, the vine is used as symbol of temporal blessing.

 

In Mandaeism, uthras (angels or celestial beings) are often described as personified grapevines (gupna).

Siphonophorae ... is an order within Hydrozoa, which is a class of marine organisms within the phylum Cnidaria.

They are highly polymorphic and complex organisms. Although they may appear to be individual organisms, each specimen is in fact a colonial organism composed of medusoid and polypoid zooids that are morphologically and functionally specialized. Zooids are multicellular units that develop from a single fertilized egg and combine to create functional colonies able to reproduce, digest, float, maintain body positioning, and use jet propulsion to move. Most colonies are long, thin, transparent floaters living in the pelagic zone.

From animalia.bio/siphonophorae

P8310235

A very very variable in color (polymorphic) species in the stinkbug family from South America. I have a couple of other examples, but there seems to be no limit to the color variations this species has. How lovely and mysterious. Not sure what the research is on this thing, but it must be an interesting story. This is what you find when you dig around in the National Collection at the Smithsonian. Sadly all the specimens are old as there is little collecting going on these days.

~~~~~~~~~~{{{{{{0}}}}}}~~~~~~~~~~

 

All photographs are public domain, feel free to download and use as you wish.

 

Photography Information: Canon Mark II 5D, Zerene Stacker, Stackshot Sled, 65mm Canon MP-E 1-5X macro lens, Twin Macro Flash in Styrofoam Cooler, F5.0, ISO 100, Shutter Speed 200

 

Beauty is truth, truth beauty - that is all

Ye know on earth and all ye need to know

" Ode on a Grecian Urn"

John Keats

 

You can also follow us on Instagram account USGSBIML Want some Useful Links to the Techniques We Use? Well now here you go Citizen:

 

Art Photo Book: Bees: An Up-Close Look at Pollinators Around the World

www.qbookshop.com/products/216627/9780760347386/Bees.html...

 

Basic USGSBIML set up:

www.youtube.com/watch?v=S-_yvIsucOY

 

USGSBIML Photoshopping Technique: Note that we now have added using the burn tool at 50% opacity set to shadows to clean up the halos that bleed into the black background from "hot" color sections of the picture.

www.youtube.com/watch?v=Bdmx_8zqvN4

 

PDF of Basic USGSBIML Photography Set Up:

ftp://ftpext.usgs.gov/pub/er/md/laurel/Droege/How%20to%20Take%20MacroPhotographs%20of%20Insects%20BIML%20Lab2.pdf

 

Google Hangout Demonstration of Techniques:

plus.google.com/events/c5569losvskrv2nu606ltof8odo

or

www.youtube.com/watch?v=4c15neFttoU

 

Excellent Technical Form on Stacking:

www.photomacrography.net/

 

Contact information:

Sam Droege

sdroege@usgs.gov

301 497 5840

 

A very very variable in color (polymorphic) species in the stinkbug family from South America. I have a couple of other examples, but there seems to be no limit to the color variations this species has. How lovely and mysterious. Not sure what the research is on this thing, but it must be an interesting story. This is what you find when you dig around in the National Collection at the Smithsonian. Sadly all the specimens are old as there is little collecting going on these days.

~~~~~~~~~~{{{{{{0}}}}}}~~~~~~~~~~

 

All photographs are public domain, feel free to download and use as you wish.

 

Photography Information: Canon Mark II 5D, Zerene Stacker, Stackshot Sled, 65mm Canon MP-E 1-5X macro lens, Twin Macro Flash in Styrofoam Cooler, F5.0, ISO 100, Shutter Speed 200

 

Beauty is truth, truth beauty - that is all

Ye know on earth and all ye need to know

" Ode on a Grecian Urn"

John Keats

 

You can also follow us on Instagram account USGSBIML Want some Useful Links to the Techniques We Use? Well now here you go Citizen:

 

Art Photo Book: Bees: An Up-Close Look at Pollinators Around the World

www.qbookshop.com/products/216627/9780760347386/Bees.html...

 

Basic USGSBIML set up:

www.youtube.com/watch?v=S-_yvIsucOY

 

USGSBIML Photoshopping Technique: Note that we now have added using the burn tool at 50% opacity set to shadows to clean up the halos that bleed into the black background from "hot" color sections of the picture.

www.youtube.com/watch?v=Bdmx_8zqvN4

 

PDF of Basic USGSBIML Photography Set Up:

ftp://ftpext.usgs.gov/pub/er/md/laurel/Droege/How%20to%20Take%20MacroPhotographs%20of%20Insects%20BIML%20Lab2.pdf

 

Google Hangout Demonstration of Techniques:

plus.google.com/events/c5569losvskrv2nu606ltof8odo

or

www.youtube.com/watch?v=4c15neFttoU

 

Excellent Technical Form on Stacking:

www.photomacrography.net/

 

Contact information:

Sam Droege

sdroege@usgs.gov

301 497 5840