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Spherulitic obsidian from the Cenozoic of California, USA.

 

The black material is obsidian, a glassy-textured, felsic to intermediate volcanic rock. The whitish-gray areas are perlite devitrification spherulites composed of cristobalite (SiO2), a polymorph of quartz.

 

Locality: unrecorded/undisclosed site east of Cougar Butte, Medicine Lake Highlands, far-eastern Siskiyou County, northern California, USA

 

The 56 Full Sized Morphs Are:

01 Blaze a Trail | 02 Pearly King Morph | 03 The Messenger Morph | 04 The Power of Morphing Communication | 05 Morph Over, There's Room for Two! | 06 Morph into the Piñataverse | 07 Morpheus | 08 Apart Together | 09 London Parklife | 10 On Guard | 11 Mr Create | 12 Morph's Inspirational Dungarees | 13 Cactus Morph | 14 Forget-Me-Not | 15 Gingerbread Morph I 16 Totally Morphomatic! | 17 Dance-off Morph I 18 The Bard I 19 Mondrian Morph | 20 Morph Whizz Kidz Argonaut | 21 It's Raining Morphs! Halleujah! | 22 Messy Morph | 23 I Spy Morph | 24 Astromorph | 25 Make Your Mark | 26 Roll With It | 27 Morph and Friends Explore London | 28 Tartan Trailblazer | 29 London Collage | 30 Peace Love and Morph | 31 Midas Morph | 32 Freedom | 33 Good Vibes | 34 Tiger Morph | 35 Maximus Morpheus Londinium | 36 Chocks Away! | 37 Morph! It's the Wrong Trousers! | 38 Diverse-City | 39 Apples and Pears | 40 Morphlowers Please! | 41 Cyborg Morph | 42 Pride Morph | 43 The London Man | 44 Looking After the Ocean | 45 Rock Star! | 46 Wheelie | 47 Gentlemorph | 48 Polymorphism | 49 Whizz Bang! | 50 Stay Frosty | 51 Mmmmmmmoprh! | 52 Swashbuckler | 53 Morph Target | 54 Canary Morph | 55 Morph the Yeoman Guard | 56 Fish Ahoy!

 

The 23 Mini Morphs Are:

01 Neville | 02 Messy Morph | 03 Meta-MORPH-osis | 04 Morley the Morph - Ready to Board | 05 Near and Far | 06 Bright Ideas | 07 Creativity Rocks! | 08 Growing Together | 10 Many Hands Make Valence | 11 Mr. Tayo Shnubbub 'The Wellbeing Hero' | 12 Captain Compass I 13 Hands-On & Hands-Up | 14 This is Us | 15 The Adventures of Morph | 16 Our School | 17 Riverside Spirit | 18 Morpheby | 19 GRIT | 20 Happiness is an Inside Job | 21 Growing Together in Learning and in Faith | 22 Look for the Light I 23 Bringing Great Energy and Spirit to Make Things Happen

Princeton Molecular Biology Workshop

at the

Springside School, Philadelphia, PA

 

April 1, 2, 15, and 18, 2009

 

This photo is from

April 15 - Human VNTR Polymorphism

A mineral is a naturally-occurring, solid, inorganic, crystalline substance having a fairly definite chemical composition and having fairly definite physical properties. At its simplest, a mineral is a naturally-occurring solid chemical. Currently, there are over 6100 named and described minerals - about 200 of them are common and about 20 of them are very common. Mineral classification is based on anion chemistry. Major categories of minerals are: elements, sulfides, oxides, halides, carbonates, sulfates, phosphates, and silicates.

 

The carbonate minerals all contain one or more carbonate (CO3-2) anions.

 

Aragonite has the same chemistry as calcite - it is calcium carbonate (CaCO3). Why is it a different mineral? Aragonite has a different molecular structure - the atoms are packed differently. Different minerals having the same chemical formula are called polymorphs (another good example is graphite & diamond - both C).

 

The difference in atomic-level packing between calcite and aragonite can be seen at the level of mineral hand samples. Aragonite forms crystals in the orthorhombic class. Many aragonite crystals are acicular (needle-like). Many aragonites form pseudohexagonal crystals, the result of 6 orthorhombic prisms growing parallel to each other. The specimen seen here is a side view of a cyclic-twinned, pseudohexagonal aragonite mass.

 

Aragonite is slightly harder than calcite, at H=3.5 to 4, occurs in many colors, and easily bubbles in acid. Aragonite is a little bit heavier than calcite, due to closer packing of atoms.

 

Most modern seashells & coral skeletons are composed of the aragonite. Whitish-colored lime sand beaches in the world are aragonitic. Occasionally, "whitings" are seen in shallow, warm ocean environments. Whitings (cloudy, milky seawater) turn out to be loaded with tiny hair-like needles of aragonite.

 

In the rock record, aragonitic or aragonite-rich sediments convert to calcite over time. Cenozoic-aged carbonate sedimentary rocks are often aragonitic. Mesozoic- and Paleozoic-aged carbonates are almost always calcitic. Many ancient fossils have had their aragonitic shells dissolved away. Ancient shells that were originally calcitic are often still well preserved.

 

Locality: unrecorded / undisclosed, but possibly from Spain

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Photo gallery of aragonite:

www.mindat.org/gallery.php?min=307

 

This striking swallowtail actually has no tails at all, however like other similar mormon butterflies they originated in Southern Asia and Australia. Caterpillar host plants are most likely Rutaceae.

 

The origins of giving common English names to organisms, particularly butterflies for tropical species started in India around the mid 19th century ... The naming of Mormons evolved slowly. I think the first to get such a name was the Common Mormon (Papilio polytes), because it had three different females, a fact that could only have been observed in the field, and this they did in India. The name obviously reflected the ... Mormon sect in America, which as we know, practiced polygamy. Another school of thought as to why these 3 species are dubbed “Mormon”: The Scarlet Mormon (Papilio rumanzovia), Great Mormon (Papilio memnon) and Common Mormon (Papilio polytes) are all beautifully colored in black and white, often with red markings. The high contrast patterns along with their relatively large size cause members of these species to standout to visitors of the butterfly exhibit. Furthermore, Butterfly Wing visitors will think they are seeing more species than they actually are, because each of the Mormons is sexually dimorphic (males and females differ) as well as polymorphic (more than one wing pattern). These polymorphisms were first described nearly a century and a half ago.

 

Source: davesgarden.com/guides/bf/go/5058/

Common Banded Peacock (Papilio crino)

We figured that if we disguised two of us as hill giants (because that's all the polymorph spells we had), we could take some others of our party into the hall as our prisoners, make friends with the hill giants, and then ensorcel their leader to cease the awful war of aggression against the elves.

 

It all went well at first. Taranis and Kaela, the party's mage and psionic, were transformed into male hill giants. We left two of our best fighters to infliltrate the smoky rafters of the hall. We took our less agile companions in as prisoners, loosely bound.

 

The hill giant hazing ritual was going well. Fists were flying good-naturedly, and we were just about to be inducted into the hill giant group, where Taranis could get close enough to the leader. Unfortunately, Cerulean recognized Celog, and went for him. The elf flew into a panic at being approached by a 17-foot Storm giant. Yuri, who held no great love for old Cerulean, whipped out a powerful rod that transformed his bones to jelly, killing Cerulean instantly.

 

All hell broke loose after that. Taranis and Kaela quickly set off a few fireballs, killing most of the hill giants, but not denting the stone giants lurking at the other end of the room. Nosnra rallied his men into a retreat to their underground caves, and our party was left to wander the compound before it burned to the ground from all the fireballs.

 

Whow knows what will happen next time, on Bobbin's Angels?

 

The 56 Full Sized Morphs Are:

01 Blaze a Trail | 02 Pearly King Morph | 03 The Messenger Morph | 04 The Power of Morphing Communication | 05 Morph Over, There's Room for Two! | 06 Morph into the Piñataverse | 07 Morpheus | 08 Apart Together | 09 London Parklife | 10 On Guard | 11 Mr Create | 12 Morph's Inspirational Dungarees | 13 Cactus Morph | 14 Forget-Me-Not | 15 Gingerbread Morph I 16 Totally Morphomatic! | 17 Dance-off Morph I 18 The Bard I 19 Mondrian Morph | 20 Morph Whizz Kidz Argonaut | 21 It's Raining Morphs! Halleujah! | 22 Messy Morph | 23 I Spy Morph | 24 Astromorph | 25 Make Your Mark | 26 Roll With It | 27 Morph and Friends Explore London | 28 Tartan Trailblazer | 29 London Collage | 30 Peace Love and Morph | 31 Midas Morph | 32 Freedom | 33 Good Vibes | 34 Tiger Morph | 35 Maximus Morpheus Londinium | 36 Chocks Away! | 37 Morph! It's the Wrong Trousers! | 38 Diverse-City | 39 Apples and Pears | 40 Morphlowers Please! | 41 Cyborg Morph | 42 Pride Morph | 43 The London Man | 44 Looking After the Ocean | 45 Rock Star! | 46 Wheelie | 47 Gentlemorph | 48 Polymorphism | 49 Whizz Bang! | 50 Stay Frosty | 51 Mmmmmmmoprh! | 52 Swashbuckler | 53 Morph Target | 54 Canary Morph | 55 Morph the Yeoman Guard | 56 Fish Ahoy!

 

The 23 Mini Morphs Are:

01 Neville | 02 Messy Morph | 03 Meta-MORPH-osis | 04 Morley the Morph - Ready to Board | 05 Near and Far | 06 Bright Ideas | 07 Creativity Rocks! | 08 Growing Together | 10 Many Hands Make Valence | 11 Mr. Tayo Shnubbub 'The Wellbeing Hero' | 12 Captain Compass I 13 Hands-On & Hands-Up | 14 This is Us | 15 The Adventures of Morph | 16 Our School | 17 Riverside Spirit | 18 Morpheby | 19 GRIT | 20 Happiness is an Inside Job | 21 Growing Together in Learning and in Faith | 22 Look for the Light I 23 Bringing Great Energy and Spirit to Make Things Happen

The 56 Full Sized Morphs Are:

01 Blaze a Trail | 02 Pearly King Morph | 03 The Messenger Morph | 04 The Power of Morphing Communication | 05 Morph Over, There's Room for Two! | 06 Morph into the Piñataverse | 07 Morpheus | 08 Apart Together | 09 London Parklife | 10 On Guard | 11 Mr Create | 12 Morph's Inspirational Dungarees | 13 Cactus Morph | 14 Forget-Me-Not | 15 Gingerbread Morph I 16 Totally Morphomatic! | 17 Dance-off Morph I 18 The Bard I 19 Mondrian Morph | 20 Morph Whizz Kidz Argonaut | 21 It's Raining Morphs! Halleujah! | 22 Messy Morph | 23 I Spy Morph | 24 Astromorph | 25 Make Your Mark | 26 Roll With It | 27 Morph and Friends Explore London | 28 Tartan Trailblazer | 29 London Collage | 30 Peace Love and Morph | 31 Midas Morph | 32 Freedom | 33 Good Vibes | 34 Tiger Morph | 35 Maximus Morpheus Londinium | 36 Chocks Away! | 37 Morph! It's the Wrong Trousers! | 38 Diverse-City | 39 Apples and Pears | 40 Morphlowers Please! | 41 Cyborg Morph | 42 Pride Morph | 43 The London Man | 44 Looking After the Ocean | 45 Rock Star! | 46 Wheelie | 47 Gentlemorph | 48 Polymorphism | 49 Whizz Bang! | 50 Stay Frosty | 51 Mmmmmmmoprh! | 52 Swashbuckler | 53 Morph Target | 54 Canary Morph | 55 Morph the Yeoman Guard | 56 Fish Ahoy!

 

The 23 Mini Morphs Are:

01 Neville | 02 Messy Morph | 03 Meta-MORPH-osis | 04 Morley the Morph - Ready to Board | 05 Near and Far | 06 Bright Ideas | 07 Creativity Rocks! | 08 Growing Together | 10 Many Hands Make Valence | 11 Mr. Tayo Shnubbub 'The Wellbeing Hero' | 12 Captain Compass I 13 Hands-On & Hands-Up | 14 This is Us | 15 The Adventures of Morph | 16 Our School | 17 Riverside Spirit | 18 Morpheby | 19 GRIT | 20 Happiness is an Inside Job | 21 Growing Together in Learning and in Faith | 22 Look for the Light I 23 Bringing Great Energy and Spirit to Make Things Happen

The 56 Full Sized Morphs Are:

01 Blaze a Trail | 02 Pearly King Morph | 03 The Messenger Morph | 04 The Power of Morphing Communication | 05 Morph Over, There's Room for Two! | 06 Morph into the Piñataverse | 07 Morpheus | 08 Apart Together | 09 London Parklife | 10 On Guard | 11 Mr Create | 12 Morph's Inspirational Dungarees | 13 Cactus Morph | 14 Forget-Me-Not | 15 Gingerbread Morph I 16 Totally Morphomatic! | 17 Dance-off Morph I 18 The Bard I 19 Mondrian Morph | 20 Morph Whizz Kidz Argonaut | 21 It's Raining Morphs! Halleujah! | 22 Messy Morph | 23 I Spy Morph | 24 Astromorph | 25 Make Your Mark | 26 Roll With It | 27 Morph and Friends Explore London | 28 Tartan Trailblazer | 29 London Collage | 30 Peace Love and Morph | 31 Midas Morph | 32 Freedom | 33 Good Vibes | 34 Tiger Morph | 35 Maximus Morpheus Londinium | 36 Chocks Away! | 37 Morph! It's the Wrong Trousers! | 38 Diverse-City | 39 Apples and Pears | 40 Morphlowers Please! | 41 Cyborg Morph | 42 Pride Morph | 43 The London Man | 44 Looking After the Ocean | 45 Rock Star! | 46 Wheelie | 47 Gentlemorph | 48 Polymorphism | 49 Whizz Bang! | 50 Stay Frosty | 51 Mmmmmmmoprh! | 52 Swashbuckler | 53 Morph Target | 54 Canary Morph | 55 Morph the Yeoman Guard | 56 Fish Ahoy!

 

The 23 Mini Morphs Are:

01 Neville | 02 Messy Morph | 03 Meta-MORPH-osis | 04 Morley the Morph - Ready to Board | 05 Near and Far | 06 Bright Ideas | 07 Creativity Rocks! | 08 Growing Together | 10 Many Hands Make Valence | 11 Mr. Tayo Shnubbub 'The Wellbeing Hero' | 12 Captain Compass I 13 Hands-On & Hands-Up | 14 This is Us | 15 The Adventures of Morph | 16 Our School | 17 Riverside Spirit | 18 Morpheby | 19 GRIT | 20 Happiness is an Inside Job | 21 Growing Together in Learning and in Faith | 22 Look for the Light I 23 Bringing Great Energy and Spirit to Make Things Happen

Plethodon vehiculum, Clatsop County, Oregon. Found under a single board.

Teapotty! is an interactive electronic teapot I've constructed for the Chi-TEK tech tea party event at the V&A in Sept 11. Before and after the tea party, it will sit quietly on display in a cabinet for 3 months. www.mztek.org/programs/chi-tek/

 

Teapotty! is a teapot sitting on a servo which takes readings from a magnetometer which is influenced by neodymium magnets in a cup. The magnetometer takes the reading from the north position, plays a bit of something similar to the Tetley Tea tune and then moves to a new position - where the teacup moves to. BlinkM RGB LEDs indicate the new position of the teapot by changing colour based on the teapot's new position from 0-180 degrees. I made some polymorph diffuser covers for them & also added heart confetti to emphasise the feeling of heartwarming happiness a cup of tea can bring :-)

 

My videos & blog post are here: rainycatz.wordpress.com/2011/09/17/teapotty-electronic-te...

   

Femias inmaduras. Infuscans (esquerda) e androcroma (dereita).

 

"Infuscans (left) and androchrome (right) inmature females."

Fulgurite from Morocco.

 

Fulgurites are glassy to vesicular, tubular to branching masses caused by lightning striking the ground (often unconsolidated materials such as sand or soil). If the target materials are silica-rich (SiO2 - quartz), then the fulgurites will be composed of lechatelierite, a form of amorphous silica. The lechatelierite in older fulgurites will have converted to cristobalite or tridymite, which are crystalline polymorphs of quartz.

-----------------------------------

Info. from label:

 

Fulgurites form when a lightning bolt strikes the Earth's surface, fusing the material into a glass tube, sometimes with a hollow center. When lightning strikes sandy soil, the air and moisture in the sand are rapidly heated. Quartz sand melts at a temperature of roughly 1600° Celsius to 2000° Celsius, depending on the moisture content. The name comes from the Latin "fulgur", for lightning.

 

These are Type 1 fulgurites, which form in sand as tubular structures, sometimes with the central opening collapsed, and frequently coated with sand.

-----------------------------------

Locality: undisclosed site in the Sahara Dune Field, Morocco, northwestern Africa

-----------------------------------

Info. at:

en.wikipedia.org/wiki/Fulgurite

 

Philomachus pugnax

 

Ruff - Non-breeding male - Aswan, Egypt

 

These are about the same size as the Dunlin seen in the bay area, but are an old world species. It is a rare but regular migrant in Alaska, the Atlantic and Pacific coast and midwest. Normally breeds from Britain to Scandinavia to east Siberia in moist arctic tundra. Winters in sub-saharan Africa. This specimen was photographed on the west bank of the Nile in Aswan. It prefers freshwater habitats.

 

These birds are unique amongst shorebirds for their striking sexual dimorphism and polymorphism. They are also unique because of the grouselike lek display performed by the males. I would _LOVE_ to photograph these during one of the leks.

Princeton Molecular Biology Workshop

at the

Springside School, Philadelphia, PA

 

April 1, 2, 15, and 18, 2009

 

This photo is from

April 15 - Human VNTR Polymorphism

© Gwendolyn Lee | Facebook | Flickr | Instagram | Tumblr | Twitter

 

To book a shoot (live or promotional) or purchase prints, please send me an EMAIL. Most photos are available for purchase, but not all.

 

+61 401 922 140

 

Gwen.B.Lee@gmail.com

Princeton Molecular Biology Workshop

at the

Springside School, Philadelphia, PA

 

April 1, 2, 15, and 18, 2009

 

This photo is from

April 15 - Human VNTR Polymorphism

The concave shape was inspired by Steb1's small concave diffuser he created for his MT24 EX (Canon equivalent of Nikon's R1C1 macro flash system), you can see Steb1's equipment setup here - flic.kr/s/aHsjjHRWp5.

I molded some plastic around the end of the SB-R200 using polymorph which protuded enough to slot in a small plastic tube. The tube naturally bends into an oval shape when slotted into the plastic mold. A concave, oval shaped deodorant lid is placed in the end which has further diffusion by 2 layers of velum paper glued to the deodorant lid. Not ideal for lighting background but concentrates the light at the subject and is very portable and not clunky or oversized like previous designs.

Shortcuts to All 20 Morphs:-

 

01. Green Fingers (Laura-Kate Draws) | 02. The Starry Night (Glen Brooks) | 03. Timeless (Roy Meats) | 04. Meandering Morph (RP Roberts) | 05. Mr Create (Jenny Leonard) | 06. Not So Handy Man (Lei-Mai LeMaow) | 07. Morpheus (Jodie Silverman) | 08. Morph and Friends (Jessica Perrin) | 09. Metamorphosis (Donna Newman) | 10. Polymorphism (Sue Gutherie) | 11. Morph in the Jungle (Amanda Quellin) | 12. Astromorph (Megan Heather Smith-Evans) | 13. Tiger Morph (Sandra Russell) | 14. Mighty Morph (Steve Johnson) |15. Wildermorph (Jina Gelder) | 16. Flora (Lisa Kirkbride) | 17. Tesselate (Jim Edwards) | 18. Fish Ahoy (Ali Elly Design) | 19. A Taste of What's to Come (Emily Ward) | 20. Morph-Code (Glen Brooks)

Shortcuts to All 20 Morphs:-

 

01. Green Fingers (Laura-Kate Draws) | 02. The Starry Night (Glen Brooks) | 03. Timeless (Roy Meats) | 04. Meandering Morph (RP Roberts) | 05. Mr Create (Jenny Leonard) | 06. Not So Handy Man (Lei-Mai LeMaow) | 07. Morpheus (Jodie Silverman) | 08. Morph and Friends (Jessica Perrin) | 09. Metamorphosis (Donna Newman) | 10. Polymorphism (Sue Gutherie) | 11. Morph in the Jungle (Amanda Quellin) | 12. Astromorph (Megan Heather Smith-Evans) | 13. Tiger Morph (Sandra Russell) | 14. Mighty Morph (Steve Johnson) |15. Wildermorph (Jina Gelder) | 16. Flora (Lisa Kirkbride) | 17. Tesselate (Jim Edwards) | 18. Fish Ahoy (Ali Elly Design) | 19. A Taste of What's to Come (Emily Ward) | 20. Morph-Code (Glen Brooks)

The 56 Full Sized Morphs Are:

01 Blaze a Trail | 02 Pearly King Morph | 03 The Messenger Morph | 04 The Power of Morphing Communication | 05 Morph Over, There's Room for Two! | 06 Morph into the Piñataverse | 07 Morpheus | 08 Apart Together | 09 London Parklife | 10 On Guard | 11 Mr Create | 12 Morph's Inspirational Dungarees | 13 Cactus Morph | 14 Forget-Me-Not | 15 Gingerbread Morph I 16 Totally Morphomatic! | 17 Dance-off Morph I 18 The Bard I 19 Mondrian Morph | 20 Morph Whizz Kidz Argonaut | 21 It's Raining Morphs! Halleujah! | 22 Messy Morph | 23 I Spy Morph | 24 Astromorph | 25 Make Your Mark | 26 Roll With It | 27 Morph and Friends Explore London | 28 Tartan Trailblazer | 29 London Collage | 30 Peace Love and Morph | 31 Midas Morph | 32 Freedom | 33 Good Vibes | 34 Tiger Morph | 35 Maximus Morpheus Londinium | 36 Chocks Away! | 37 Morph! It's the Wrong Trousers! | 38 Diverse-City | 39 Apples and Pears | 40 Morphlowers Please! | 41 Cyborg Morph | 42 Pride Morph | 43 The London Man | 44 Looking After the Ocean | 45 Rock Star! | 46 Wheelie | 47 Gentlemorph | 48 Polymorphism | 49 Whizz Bang! | 50 Stay Frosty | 51 Mmmmmmmoprh! | 52 Swashbuckler | 53 Morph Target | 54 Canary Morph | 55 Morph the Yeoman Guard | 56 Fish Ahoy!

 

The 23 Mini Morphs Are:

01 Neville | 02 Messy Morph | 03 Meta-MORPH-osis | 04 Morley the Morph - Ready to Board | 05 Near and Far | 06 Bright Ideas | 07 Creativity Rocks! | 08 Growing Together | 10 Many Hands Make Valence | 11 Mr. Tayo Shnubbub 'The Wellbeing Hero' | 12 Captain Compass I 13 Hands-On & Hands-Up | 14 This is Us | 15 The Adventures of Morph | 16 Our School | 17 Riverside Spirit | 18 Morpheby | 19 GRIT | 20 Happiness is an Inside Job | 21 Growing Together in Learning and in Faith | 22 Look for the Light I 23 Bringing Great Energy and Spirit to Make Things Happen

Vug with aragonite crystals in dolostone in the Mississippian of Ohio, USA.

 

This rock has a small vug containing numerous clear to whitish needles of aragonite (CaCO3 - calcium carbonate). Aragonite is a less common polymorph of calcite and is rarely found in Ohio. The host rock is a glauconitic fossiliferous dolostone having crinoid stem columnals. The green spots in parts of the surrounding matrix are glauconite (click on the photo to zoom in & look around).

 

Stratigraphy: dolostone horizon below inferred Rushville Shale, upper Vinton Member, upper Logan Formation, ~middle Osagean Stage, upper Lower Mississippian

 

Locality: Trinway West 6 Outcrop - lower slopes of the eastern part of roadcut on the northwestern side of Rt. 16, 1.0 miles northeast of the Rt. 16-Old Riley Road intersection, northeast of the town of Frazeysburg & west of the town of Trinway, northwestern Muskingum County, Ohio, USA (~40° 08' 46.98" North latitude, 82° 04' 56.49" West longitude)

 

Smart DNA testing gives us an individualized report of our genetic susceptabilities to known polymorphisms (or mutations) that can have great impacts on our health. It gives us great insights into diet, lifestyle, supplements and functional pathology tests that ensure we optimize our potential.

Polymorph beads heated, mashed together and stretched and lit from within. Actual size about 3.5mm

Narcissus is a genus of predominantly spring flowering perennial plants of the amaryllis family, Amaryllidaceae. Various common names including daffodil, narcissus, and jonquil are used to describe all or some members of the genus. Narcissus has conspicuous flowers with six petal-like tepals surmounted by a cup- or trumpet-shaped corona. The flowers are generally white and yellow (also orange or pink in garden varieties), with either uniform or contrasting coloured tepals and corona.

 

Narcissus were well known in ancient civilisation, both medicinally and botanically, but formally described by Linnaeus in his Species Plantarum (1753). The genus is generally considered to have about ten sections with approximately 50 species. The number of species has varied, depending on how they are classified, due to similarity between species and hybridisation. The genus arose some time in the Late Oligocene to Early Miocene epochs, in the Iberian peninsula and adjacent areas of southwest Europe. The exact origin of the name Narcissus is unknown, but it is often linked to a Greek word (ancient Greek ναρκῶ narkō, "to make numb") and the myth of the youth of that name who fell in love with his own reflection. The English word "daffodil" appears to be derived from "asphodel", with which it was commonly compared.

 

The species are native to meadows and woods in southern Europe and North Africa with a centre of diversity in the Western Mediterranean, particularly the Iberian peninsula. Both wild and cultivated plants have naturalised widely, and were introduced into the Far East prior to the tenth century. Narcissi tend to be long-lived bulbs, which propagate by division, but are also insect-pollinated. Known pests, diseases and disorders include viruses, fungi, the larvae of flies, mites and nematodes. Some Narcissus species have become extinct, while others are threatened by increasing urbanisation and tourism.

 

Historical accounts suggest narcissi have been cultivated from the earliest times, but became increasingly popular in Europe after the 16th century and by the late 19th century were an important commercial crop centred primarily in the Netherlands. Today narcissi are popular as cut flowers and as ornamental plants in private and public gardens. The long history of breeding has resulted in thousands of different cultivars. For horticultural purposes, narcissi are classified into divisions, covering a wide range of shapes and colours. Like other members of their family, narcissi produce a number of different alkaloids, which provide some protection for the plant, but may be poisonous if accidentally ingested. This property has been exploited for medicinal use in traditional healing and has resulted in the production of galantamine for the treatment of Alzheimer's dementia. Long celebrated in art and literature, narcissi are associated with a number of themes in different cultures, ranging from death to good fortune, and as symbols of spring.

 

The daffodil is the national flower of Wales and the symbol of cancer charities in many countries. The appearance of the wild flowers in spring is associated with festivals in many places.

 

Narcissus is a genus of perennial herbaceous bulbiferous geophytes, which die back after flowering to an underground storage bulb. They regrow in the following year from brown-skinned ovoid bulbs with pronounced necks, and reach heights of 5–80 centimetres (2.0–31.5 in) depending on the species. Dwarf species such as N. asturiensis have a maximum height of 5–8 centimetres (2.0–3.1 in), while Narcissus tazetta may grow as tall as 80 centimetres (31 in).

 

The plants are scapose, having a single central leafless hollow flower stem (scape). Several green or blue-green, narrow, strap-shaped leaves arise from the bulb. The plant stem usually bears a solitary flower, but occasionally a cluster of flowers (umbel). The flowers, which are usually conspicuous and white or yellow, sometimes both or rarely green, consist of a perianth of three parts. Closest to the stem (proximal) is a floral tube above the ovary, then an outer ring composed of six tepals (undifferentiated sepals and petals), and a central disc to conical shaped corona. The flowers may hang down (pendant), or be erect. There are six pollen bearing stamens surrounding a central style. The ovary is inferior (below the floral parts) consisting of three chambers (trilocular). The fruit consists of a dry capsule that splits (dehisces) releasing numerous black seeds.

 

The bulb lies dormant after the leaves and flower stem die back and has contractile roots that pull it down further into the soil. The flower stem and leaves form in the bulb, to emerge the following season. Most species are dormant from summer to late winter, flowering in the spring, though a few species are autumn flowering.

 

The pale brown-skinned ovoid tunicate bulbs have a membranous tunic and a corky stem (base or basal) plate from which arise the adventitious root hairs in a ring around the edge, which grow up to 40 mm in length. Above the stem plate is the storage organ consisting of bulb scales, surrounding the previous flower stalk and the terminal bud. The scales are of two types, true storage organs and the bases of the foliage leaves. These have a thicker tip and a scar from where the leaf lamina became detached. The innermost leaf scale is semicircular only partly enveloping the flower stalk (semisheathed).(see Hanks Fig 1.3). The bulb may contain a number of branched bulb units, each with two to three true scales and two to three leaf bases. Each bulb unit has a life of about four years.

 

Once the leaves die back in summer, the roots also wither. After some years, the roots shorten pulling the bulbs deeper into the ground (contractile roots). The bulbs develop from the inside, pushing the older layers outwards which become brown and dry, forming an outer shell, the tunic or skin. Up to 60 layers have been counted in some wild species. While the plant appears dormant above the ground the flower stalk which will start to grow in the following spring, develops within the bulb surrounded by two to three deciduous leaves and their sheaths. The flower stem lies in the axil of the second true leaf.

 

The single leafless Plant stem stem or scape, appearing from early to late spring depending on the species, bears from 1 to 20 blooms. Stem shape depends on the species, some are highly compressed with a visible seam, while others are rounded. The stems are upright and located at the centre of the leaves. In a few species such as N. hedraeanthus the stem is oblique. The stem is hollow in the upper portion but towards the bulb is more solid and filled with a spongy material.

 

Narcissus plants have one to several basal leaf leaves which are linear, ligulate or strap-shaped (long and narrow), sometimes channelled adaxially to semiterete, and may (pedicellate) or may not (sessile) have a petiole stalk. The leaves are flat and broad to cylindrical at the base and arise from the bulb. The emerging plant generally has two leaves, but the mature plant usually three, rarely four, and they are covered with a cutin containing cuticle, giving them a waxy appearance. Leaf colour is light green to blue-green. In the mature plant, the leaves extend higher than the flower stem, but in some species, the leaves are low-hanging. The leaf base is encased in a colorless sheath. After flowering, the leaves turn yellow and die back once the seed pod (fruit) is ripe.

 

Jonquils usually have dark green, round, rush-like leaves.

 

The inflorescence is scapose, the single stem or scape bearing either a solitary flower or forming an umbel with up to 20 blooms. Species bearing a solitary flower include section Bulbocodium and most of section Pseudonarcissus. Umbellate species have a fleshy racemose inflorescence (unbranched, with short floral stalks) with 2 to 15 or 20 flowers, such as N. papyraceus (see illustration, left) and N. tazetta (see Table I). The flower arrangement on the inflorescence may be either with (pedicellate) or without (sessile) floral stalks.

 

Prior to opening, the flower buds are enveloped and protected in a thin dry papery or membranous (scarious) spathe. The spathe consists of a singular bract that is ribbed, and which remains wrapped around the base of the open flower. As the bud grows, the spathe splits longitudinally. Bracteoles are small or absent.

 

The flowers of Narcissus are hermaphroditic (bisexual), have three parts (tripartite), and are sometimes fragrant (see Fragrances). The flower symmetry is actinomorphic (radial) to slightly zygomorphic (bilateral) due to declinate-ascending stamens (curving downwards, then bent up at the tip). Narcissus flowers are characterised by their, usually conspicuous, corona (trumpet).

 

The three major floral parts (in all species except N. cavanillesii in which the corona is virtually absent - Table I: Section Tapeinanthus) are;

 

(i) the proximal floral tube (hypanthium),

(ii) the surrounding free tepals, and

(iii) the more distal corona (paraperigon, paraperigonium).

All three parts may be considered to be components of the perianth (perigon, perigonium). The perianth arises above the apex of the inferior ovary, its base forming the hypanthial floral tube.

 

The floral tube is formed by fusion of the basal segments of the tepals (proximally connate). Its shape is from an inverted cone (obconic) to funnel-shaped (funneliform) or cylindrical, and is surmounted by the more distal corona. Floral tubes can range from long and narrow sections Apodanthi and Jonquilla to rudimentary (N. cavanillesii).

 

Surrounding the floral tube and corona and reflexed (bent back) from the rest of the perianth are the six spreading tepals or floral leaves, in two whorls which may be distally ascending, reflexed (folded back), or lanceolate. Like many monocotyledons, the perianth is homochlamydeous, which is undifferentiated into separate calyx (sepals) and corolla (petals), but rather has six tepals. The three outer tepal segments may be considered sepals, and the three inner segments petals. The transition point between the floral tube and the corona is marked by the insertion of the free tepals on the fused perianth.

 

The corona, or paracorolla, is variously described as bell-shaped (funneliform, trumpet), bowl-shaped (cupular, crateriform, cup-shaped) or disc-shaped with margins that are often frilled, and is free from the stamens. Rarely is the corona a simple callose (hardened, thickened) ring. The corona is formed during floral development as a tubular outgrowth from stamens which fuse into a tubular structure, the anthers becoming reduced. At its base, the fragrances which attract pollinators are formed. All species produce nectar at the top of the ovary. Coronal morphology varies from the tiny pigmented disk of N. serotinus (see Table I) or the rudimentary structure in N. cavanillesii to the elongated trumpets of section Pseudonarcissus (trumpet daffodils, Table I).

 

While the perianth may point forwards, in some species such as N. cyclamineus it is folded back (reflexed, see illustration, left), while in some other species such as N. bulbocodium (Table I), it is reduced to a few barely visible pointed segments with a prominent corona.

 

The colour of the perianth is white, yellow or bicoloured, with the exception of the night flowering N. viridiflorus which is green. In addition the corona of N. poeticus has a red crenulate margin (see Table I). Flower diameter varies from 12 mm (N. bulbocodium) to over 125 mm (N. nobilis=N. pseudonarcissus subsp. nobilis).

 

Flower orientation varies from pendent or deflexed (hanging down) as in N. triandrus (see illustration, left), through declinate-ascendant as in N. alpestris = N. pseudonarcissus subsp. moschatus, horizontal (patent, spreading) such as N. gaditanus or N. poeticus, erect as in N. cavanillesii, N. serotinus and N. rupicola (Table I), or intermediate between these positions (erecto-patent).

 

The flowers of Narcissus demonstrate exceptional floral diversity and sexual polymorphism, primarily by corona size and floral tube length, associated with pollinator groups (see for instance Figs. 1 and 2 in Graham and Barrett). Barrett and Harder (2005) describe three separate floral patterns;

 

"Daffodil" form

"Paperwhite" form

"Triandrus" form.

The predominant patterns are the 'daffodil' and 'paperwhite' forms, while the "triandrus" form is less common. Each corresponds to a different group of pollinators (See Pollination).

 

The "daffodil" form, which includes sections Pseudonarcissus and Bulbocodium, has a relatively short, broad or highly funnelform tube (funnel-like), which grades into an elongated corona, which is large and funnelform, forming a broad, cylindrical or trumpet-shaped perianth. Section Pseudonarcissus consists of relatively large flowers with a corolla length of around 50 mm, generally solitary but rarely in inflorescences of 2–4 flowers. They have wide greenish floral tubes with funnel-shaped bright yellow coronas. The six tepals sometimes differ in colour from the corona and may be cream coloured to pale yellow.

 

The "paperwhite" form, including sections Jonquilla, Apodanthi and Narcissus, has a relatively long, narrow tube and a short, shallow, flaring corona. The flower is horizontal and fragrant.

 

The "triandrus" form is seen in only two species, N. albimarginatus (a Moroccan endemic) and N. triandrus. It combines features of both the "daffodil" and "paperwhite" forms, with a well-developed, long, narrow tube and an extended bell-shaped corona of almost equal length. The flowers are pendent.

 

Androecium

There are six stamens in one to two rows (whorls), with the filaments separate from the corona, attached at the throat or base of the tube (epipetalous), often of two separate lengths, straight or declinate-ascending (curving downwards, then upwards). The anthers are basifixed (attached at their base).

 

Gynoecium

The ovary is inferior (below the floral parts) and trilocular (three chambered) and there is a pistil with a minutely three lobed stigma and filiform (thread like) style, which is often exserted (extending beyond the tube).

 

Fruit

The fruit consists of dehiscent loculicidal capsules (splitting between the locules) that are ellipsoid to subglobose (almost spherical) in shape and are papery to leathery in texture.

 

Seeds

The fruit contains numerous subglobose seeds which are round and swollen with a hard coat, sometimes with an attached elaiosome. The testa is black and the pericarp dry.

 

Most species have 12 ovules and 36 seeds, although some species such as N. bulbocodium have more, up to a maximum of 60. Seeds take five to six weeks to mature. The seeds of sections Jonquilla and Bulbocodium are wedge-shaped and matte black, while those of other sections are ovate and glossy black. A gust of wind or contact with a passing animal is sufficient to disperse the mature seeds.

 

Chromosomes

Chromosome numbers include 2n=14, 22, 26, with numerous aneuploid and polyploid derivatives. The basic chromosome number is 7, with the exception of N. tazetta, N. elegans and N. broussonetii in which it is 10 or 11; this subgenus (Hermione) was in fact characterised by this characteristic. Polyploid species include N. papyraceus (4x=22) and N. dubius (6x=50).

 

Phytochemistry

Alkaloids

As with all Amarylidaceae genera, Narcissus contains unique isoquinoline alkaloids. The first alkaloid to be identified was lycorine, from N. pseudonarcissus in 1877. These are considered a protective adaptation and are utilised in the classification of species. Nearly 100 alkaloids have been identified in the genus, about a third of all known Amaryllidaceae alkaloids, although not all species have been tested. Of the nine alkaloid ring types identified in the family, Narcissus species most commonly demonstrate the presence of alkaloids from within the Lycorine (lycorine, galanthine, pluviine) and Homolycorine (homolycorine, lycorenine) groups. Hemanthamine, tazettine, narciclasine, montanine and galantamine alkaloids are also represented. The alkaloid profile of any plant varies with time, location, and developmental stage. Narcissus also contain fructans and low molecular weight glucomannan in the leaves and plant stems.

 

Fragrances

Fragrances are predominantly monoterpene isoprenoids, with a small amount of benzenoids, although N. jonquilla has both equally represented. Another exception is N. cuatrecasasii which produces mainly fatty acid derivatives. The basic monoterpene precursor is geranyl pyrophosphate, and the commonest monoterpenes are limonene, myrcene, and trans-β-ocimene. Most benzenoids are non-methoxylated, while a few species contain methoxylated forms (ethers), e.g. N. bujei. Other ingredient include indole, isopentenoids and very small amounts of sesquiterpenes. Fragrance patterns can be correlated with pollinators, and fall into three main groups (see Pollination).

 

The taxonomy of Narcissus is complex, and still not fully resolved. Known to the ancients, the genus name appears in Graeco-Roman literature, although their interest was as much medicinal as botanical. It is unclear which species the ancients were familiar with. Although frequently mentioned in Mediaeval and Renaissance texts it was not formally described till the work of Linnaeus in 1753. By 1789 it had been grouped into a family (Narcissi) but shortly thereafter this was renamed Amaryllideae, from which comes the modern placement within Amaryllidaceae, although for a while it was considered part of Liliaceae.

 

Many of the species now considered to be Narcissus were in separate genera during the nineteenth century, and the situation was further confused by the inclusion of many cultivated varieties. By 1875 the current circumscription was relatively settled. By 2004 phylogenetic studies had allowed the place of Narcissus within its fairly large family to be established, nested within a series of subfamilies (Amaryllidoideae) and tribes (Narcisseae). It shares its position in the latter tribe with Sternbergia.

 

The infrageneric classification has been even more complex and many schemes of subgenera, sections, subsections and series have been proposed, although all had certain similarities. Most authorities now consider there to be 10 – 11 sections based on phylogenetic evidence. The problems have largely arisen from the diversity of the wild species, frequent natural hybridisation and extensive cultivation with escape and subsequent naturalisation. The number of species has varied anywhere from 16 to nearly 160, but is probably around 50 – 60.

 

The genus appeared some time in the Late Oligocene to Early Miocene eras, around 24 million years ago, in the Iberian peninsula. While the exact origin of the word Narcissus is unknown it is frequently linked to its fragrance which was thought to be narcotic, and to the legend of the youth of that name who fell in love with his reflection. In the English language the common name Daffodil appears to be derived from the Asphodel with which it was commonly compared.

 

Early

Narcissus was first described by Theophrastus (Θεόφραστος, c 371 - c 287 BC) in his Historia Plantarum (Greek: Περὶ φυτῶν ἱστορία) as νάρκισσος, referring to N. poeticus, but comparing it to Asphodelus (ασφοδελωδες). Theophrastus' description was frequently referred to at length by later authors writing in Latin such as Pliny the Elder (Gaius Plinius Secundus, 23 AD – 79 AD) from whom came the Latin form narcissus (see also Culture). Pliny's account is from his Natural History (Latin: Naturalis Historia). Like his contemporaries, his interests were as much therapeutic as botanical. Another much-cited Greek authority was Dioscorides (Διοσκουρίδης, 40 AD – 90 AD) in his De Materia Medica (Greek: Περὶ ὕλης ἰατρικῆς). Both authors were to remain influential until at least the Renaissance, given that their descriptions went beyond the merely botanical, to the therapeutic (see also Antiquity).

 

An early European reference is found in the work of Albert Magnus (c. 1200 – 1280), who noted in his De vegetabilibus et plantis the similarity to the leek. William Turner in his A New Herball (1551) cites all three extensively in his description of the plant and its properties.It was to remain to Linnaeus in 1753 to formally describe and name Narcissus as a genus in his Species Plantarum, at which time there were six known species (N. poeticus, N. pseudonarcissus, N. bulbocodium, N. serotinus, N. jonquilla and N. tazetta).[1] At that time, Linnaeus loosely grouped it together with 50 other genera into his Hexandria monogynia.

 

Modern

It was de Jussieu in 1789 who first formally created a 'family' (Narcissi), as the seventh 'Ordo' (Order) of the third class (Stamina epigyna) of Monocots in which Narcissus and 15 other genera were placed. The use of the term Ordo at that time was closer to what we now understand as Family, rather than Order. The family has undergone much reorganisation since then, but in 1805 it was renamed after a different genus in the family, Amaryllis, as 'Amaryllideae' by Jaume St.-Hilaire and has retained that association since. Jaume St.-Hilaire divided the family into two unnamed sections and recognised five species of Narcissus, omitting N. serotinus.

 

De Candolle brought together Linnaeus' genera and Jussieau's families into a systematic taxonomy for the first time, but included Narcissus (together with Amaryllis) in the Liliaceae in his Flore française (1805-1815) rather than Amaryllidaceae, a family he had not yet recognised. Shortly thereafter he separated the 'Amaryllidées' from 'Liliacées' (1813), though attributing the term to Brown's 'Amaryllideae' in the latter's Prodromus (1810) rather than St.-Hilaire's 'Amaryllidées'. He also provided the text to the first four volumes of Redouté illustrations in the latter's Les liliacées between 1805 and 1808 (see illustration here of N. candidissimus).

 

Historically both wide and narrow interpretations of the genus have been proposed. In the nineteenth century genus splitting was common, favouring the narrow view. Haworth (1831) using a narrow view treated many species as separate genera, as did Salisbury (1866). These authors listed various species in related genera such as Queltia (hybrids), Ajax (=Pseudonarcissus) and Hermione (=Tazettae), sixteen in all in Haworth's classification. In contrast, Herbert (1837) took a very wide view reducing Harworth's sixteen genera to six. Herbert, treating the Amaryllidacea as an 'order' as was common then, considered the narcissi to be a suborder, the Narcisseae, the six genera being Corbularia, Ajax, Ganymedes, Queltia, Narcissus and Hermione and his relatively narrow circumscription of Narcissus having only three species. Later Spach (1846) took an even wider view bringing most of Harworth's genera into the genus Narcissus, but as separate subgenera. By the time that Baker (1875) wrote his monograph all of the genera with one exception were included as Narcissus. The exception was the monotypic group Tapeinanthus which various subsequent authors have chosen to either exclude (e.g. Cullen 1986) or include (e.g. Webb 1978, 1980). Today it is nearly always included.

 

The eventual position of Narcissus within the Amaryllidaceae family only became settled in the twenty-first century with the advent of phylogenetic analysis and the Angiosperm Phylogeny Group system. The genus Narcissus belongs to the Narcisseae tribe, one of 13 within the Amaryllidoideae subfamily of the Amaryllidaceae. It is one of two sister clades corresponding to genera in the Narcisseae, being distinguished from Sternbergia by the presence of a paraperigonium, and is monophyletic

 

The infrageneric phylogeny of Narcissus still remains relatively unsettled. The taxonomy has proved very complex and difficult to resolve, particularly for the Pseudonarcissus group. This is due to a number of factors, including the diversity of the wild species, the ease with which natural hybridisation occurs, and extensive cultivation and breeding accompanied by escape and naturalisation.

 

De Candolle, in the first systematic taxonomy of Narcissus, arranged the species into named groups, and those names (Faux-Narcisse or Pseudonarcissus, Poétiques, Tazettes, Bulbocodiens, Jonquilles) have largely endured for the various subdivisions since and bear his name. The evolution of classification was confused by including many unknown or garden varieties, until Baker (1875) made the important distinction of excluding all specimens except the wild species from his system. He then grouped all of the earlier related genera as sections under one genus, Narcissus, the exception being the monotypic Tapeinanthus. Consequently, the number of accepted species has varied widely.

 

A common modern classification system has been that of Fernandes (1951, 1968, 1975) based on cytology, as modified by Blanchard (1990) and Mathew (2002), although in some countries such as Germany, the system of Meyer (1966) was preferred. Fernandes described two subgenera based on basal chromosome number, Hermione, n = 5 (11) and Narcissus, n = 7 (13). He further subdivided these into ten sections (Apodanthi, Aurelia, Bulbocodii, Ganymedes, Jonquillae, Narcissus, Pseudonarcissi, Serotini, Tapeinanthus, Tazettae), as did Blanchard later.

 

In contrast to Fernandes, Webb's treatment of the genus for the Flora Europaea (1978, 1980) prioritised morphology over genetics, and abandoned the subgenera ranks. He also restored De Candolle's original nomenclature, and made a number of changes to section Jonquilla, merging the existing subsections, reducing Apodanthi to a subsection of Jonquilla, and moving N. viridiflorus from Jonquilla to a new monotypic section of its own (Chloranthi). Finally, he divided Pseudonarcissus into two subsections. Blanchard (1990), whose Narcissus: a guide to wild daffodils has been very influential, adopted a simple approach, restoring Apodanthae, and based largely on ten sections alone.

 

The Royal Horticultural Society (RHS) currently lists ten sections, based on Fernandes (1968), three of which are monotypic (contain only one species), while two others only containing two species. Most species are placed in Pseudonarcissus While infrageneric groupings within Narcissus have been relatively constant, their status (genera, subgenera, sections, subsections, series, species) has not. Some authors treat some sections as being further subdivided into subsections, e.g. Tazettae (3 subsections). These subdivisions correspond roughly to the popular names for narcissi types, e.g. Trumpet Daffodils, Tazettas, Pheasant's Eyes, Hoop Petticoats, Jonquils.

 

While Webb had simply divided the genus into sections, Mathew found this unsatisfactory, implying every section had equal status. He adapted both Fernandes and Webb to devise a more hierarchical scheme he believed better reflected the interrelationships within the genus. Mathew's scheme consists of three subgenera (Narcissus, Hermione and Corbularia). The first two subgenera were then divided into five and two sections respectively. He then further subdivided two of the sections (subgenus Narcissus section Jonquillae, and subgenus Hermione section Hermione) into three subsections each. Finally, he divided section Hermione subsection Hermione further into two series, Hermione and Albiflorae. While lacking a phylogenetic basis, the system is still in use in horticulture. For instance the Pacific Bulb Society uses his numbering system for classifying species.

 

The phylogenetic analysis of Graham and Barrett (2004) supported the infrageneric division of Narcissus into two clades corresponding to the subgenera Hermione and Narcissus, but does not support monophyly of all sections, with only Apodanthi demonstrating clear monophyly, corresponding to Clade III of Graham and Barrett, although some other clades corresponded approximately to known sections. These authors examined 36 taxa of the 65 listed then, and a later extended analysis by Rønsted et al. (2008) with five additional taxa confirmed this pattern.

 

A very large (375 accessions) molecular analysis by Zonneveld (2008) utilising nuclear DNA content sought to reduce some of the paraphyly identified by Graham and Barrett. This led to a revision of the sectional structure, shifting some species between sections, eliminating one section and creating two new ones. In subgenus Hermione, Aurelia was merged with Tazettae. In subgenus Narcissus section Jonquillae subsection Juncifolii was elevated to sectional rank, thus resolving the paraphyly in this section observed by Graham and Barrett in Clade II due to this anomalous subsection, the remaining species being in subsection Jonquillae, which was monophyletic. The relatively large section Pseudonarcissi was divided by splitting off a new section, Nevadensis (species from southern Spain) leaving species from France, northern Spain and Portugal in the parent section. At the same time Fernández-Casas (2008) proposed a new monotypic section Angustini to accommodate Narcissus deficiens, placing it within subgenus Hermione.

 

While Graham and Barrett (2004) had determined that subgenus Hermione was monophyletic, using a much larger accession Santos-Gally et al. (2011) did not. However the former had excluded species of hybrid origins, while the latter included both N. dubius and N. tortifolius. If these two species are excluded (forming a clade with subgenus Narcissus) then Hermione can be considered monophyletic, although as a section of Hermione, Tazettae is not monophyletic. They also confirmed the monophyly of Apodanthi.

 

Some so-called nothosections have been proposed, predominantly by Fernández-Casas, to accommodate natural ('ancient') hybrids (nothospecies).

 

Subgenera and sections

Showing revisions by Zonnefeld (2008)

 

subgenus Hermione (Haw.) Spach.

(Aurelia (Gay) Baker (monotypic) - merged with Tazettae (2008)

Serotini Parlatore (2 species)

Tazettae de Candolle (16 species) syn. Hermione (Salisbury) Sprengel, in Fernandes' scheme. Incorporating Aurelia (2008)

subgenus Narcissus L.

Apodanthi A. Fernandes (6 species)

Bulbocodium de Candolle (11 species)

Ganymedes (Haworth) Schultes f. (monotypic)

Jonquillae de Candolle (8 species)

Juncifolii (A. Fern.) Zonn. sect. nov. (2008)

Narcissus L. (2 species)

Nevadensis Zonn. sect. nov. (2008)

Pseudonarcissus de Candolle (36 species) Trumpet daffodils

Tapeinanthus (Herbert) Traub (monotypic)

 

Species

Estimates of the number of species in Narcissus have varied widely, from anywhere between 16 and nearly 160, even in the modern era. Linnaeus originally included six species in 1753. By the time of the 14th edition of the Systema Naturae in 1784, there were fourteen. The 1819 Encyclopaedia Londinensis lists sixteen (see illustration here of three species) and by 1831 Adrian Haworth had described 150 species.

 

Much of the variation lies in the definition of species, and whether closely related taxa are considered separate species or subspecies. Thus, a very wide view of each species, such as Webb's results in few species, while a very narrow view such as that of Fernandes results in a larger number. Another factor is the status of hybrids, given natural hybridisation, with a distinction between 'ancient hybrids' and 'recent hybrids'. The term 'ancient hybrid' refers to hybrids found growing over a large area, and therefore now considered as separate species, while 'recent hybrid' refers to solitary plants found amongst their parents, with a more restricted range.

 

In the twentieth century Fernandes (1951) accepted 22 species, on which were based the 27 species listed by Webb in the 1980 Flora Europaea. By 1968, Fernandes had accepted 63 species, and by 1990 Blanchard listed 65 species, and Erhardt 66 in 1993. In 2006 the Royal Horticultural Society's (RHS) International Daffodil Register and Classified List listed 87 species, while Zonneveld's genetic study (2008) resulted in only 36. As of September 2014, the World Checklist of Selected Plant Families accepts 52 species, along with at least 60 hybrids, while the RHS has 81 accepted names in its October 2014 list.

 

Evolution

Within the Narcisseae, Narcissus (western Mediterranean) diverged from Sternbergia (Eurasia) some time in the Late Oligocene to Early Miocene eras, around 29.3–18.1 Ma, with a best estimate of 23.6 Ma. Later the genus divided into the two subgenera (Hermione and Narcissus) between 27.4 and 16.1 Ma (21.4 Ma). The divisions between the sections of Hermione then took place during the Miocene period 19.9–7.8 Ma.

 

Narcissus appears to have arisen in the area of the Iberian peninsula, southern France and northwestern Italy, and within this area most sections of the genus appeared, with only a few taxa being dispersed to North Africa at a time when the African and West European platforms were closer together. Subgenus Hermione in turn arose in the southwestern mediterranean and north west Africa. However, these are reconstructions, the Amaryllidaceae lacking a fossil record.

 

Names and etymology

The derivation of the Latin narcissus (Greek: νάρκισσος) is unknown. It may be a loanword from another language; for instance, it is said to be related to the Sanskrit word nark, meaning 'hell'. It is frequently linked to the Greek myth of Narcissus described by Ovid in his Metamorphoses, who became so obsessed with his own reflection that as he knelt and gazed into a pool of water, he fell into the water and drowned. In some variations, he died of starvation and thirst. In both versions, the narcissus plant sprang from where he died. Although Ovid appeared to describe the plant we now know as Narcissus there is no evidence for this popular derivation, and the person's name may have come from the flower's name. The Poet's Narcissus (N. poeticus), which grows in Greece, has a fragrance that has been described as intoxicating. This explanation is largely discredited due to lack of proof. Pliny wrote that the plant 'narce narcissum dictum, non a fabuloso puero' ('named narcissus from narce, not from the legendary youth'), i.e. that it was named for its narcotic properties (ναρκάω narkao, 'I grow numb' in Greek), not from the legend. Furthermore, there were accounts of narcissi growing, such as in the legend of Persephone, long before the story of Narcissus appeared (see Greek culture). It has also been suggested that daffodils bending over streams evoked the image of the youth admiring his own reflection in the water.

 

Linnaeus used the Latin name for the plant in formally describing the genus, although Matthias de l'Obel had previously used the name in describing various species of Narcissi in his Icones stirpium of 1591, and other publications, as had Clusius in Rariorum stirpium (1576).

 

The plural form of the common name narcissus has caused some confusion. British English sources such as the Shorter Oxford English Dictionary give two alternate forms, narcissi and narcissuses. In contrast, in American English the Merriam-Webster Dictionary provides for a third form, narcissus, used for both singular and plural. The Oxford dictionaries only list this third form under American English, although the Cambridge Dictionary allows of all three in the same order. However, Garner's Modern American Usage states that narcissi is the commonest form, narcissuses being excessively sibilant. For similar reasons, Fowler prefers narcissi in British English usage. Neither support narcissus as a plural form. Common names such as narcissus do not capitalise the first letter in contrast to the person of that name and the Latin genus name.

 

The name Narcissus (feminine Narcissa) was not uncommon in Roman times, such as Tiberius Claudius Narcissus, a Roman official in Claudius' time, an early New Testament Christian in Rome and later bishops and saints.

 

Daffodil

The word daffodil was unknown in the English language before the sixteenth century. The name is derived from an earlier affodell, a variant of asphodel. In classical Greek literature the narcissus is frequently referred to as the asphodel, such as the meadows of the Elysian fields in Homer (see Antiquity). Asphodel in turn appears to be a loanword coming from French via Mediaeval Latin affodilus from Classical Latin asphodilus and ultimately the Greek asphodelos (Greek: ἀσφόδελος). The reason for the introduction of the initial d is not known, although a probable source is an etymological merging from the Dutch article de, as in de affodil, or English the, as th'affodil or t'affodil, hence daffodil, and in French de and affodil to form fleur d'aphrodille and daphrodille.

 

From at least the 16th century, daffadown dilly and daffydowndilly have appeared as playful synonyms of the name. In common parlance and in historical documents, the term daffodil may refer specifically to populations or specimens of the wild daffodil, N. pseudonarcissus. H. N. Ellacombe suggests this may be from Saffon Lilly, citing Prior in support, though admittedly conjectural.

 

Lady Wilkinson (1858), who provides an extensive discussion of the etymology of the various names for this plant, suggests a very different origin, namely the Old English word affodyle (that which cometh early), citing a 14th-century (but likely originally much earlier) manuscript in support of this theory, and which appears to describe a plant resembling the daffodil. Ellacombe provides further support for this from a fifteenth century English translation of Palladius that also refers to it.

 

Jonquil

The name jonquil is said to be a corruption via French from the Latin juncifolius meaning 'rush-leaf' (Juncaceae) and its use is generally restricted to those species and cultivars which have rush like leaves, e.g. N. juncifolius.

 

Other

A profusion of names have attached themselves in the English language, either to the genus as a whole or to individual species or groups of species such as sections. These include narcissus, jonquil, Lent lily, Lenten lily, lide lily, yellow lily, wort or wyrt, Julians, glens, Lent cocks, corn flower, bell rose, asphodel, Solomon's lily, gracy day, haverdrils, giggary, cowslip, and crow foot.

Die Gouldamadine (Chloebia gouldiae, auch Erythrura gouldiae) gehört zur Familie der Prachtfinken. Sie zählen zur Fauna Australiens und sind eine polymorphe Art. In der gleichen Population kommen meist zwei, gelegentlich sogar drei in der Färbung des Oberkopfes verschiedene Variationen nebeneinander vor. Sie verpaaren sich ohne Einschränkung untereinander, so dass sie nicht als Unterarten unterschieden werden.

 

Gouldamadinen sind in ihrer Heimat mittlerweile selten geworden und werden von der IUCN als gefährdet (vulnerable) eingestuft. Weltweit wird dieser farbenprächtig gefiederte Vogel, der auch in freier Wildbahn in drei unterschiedlichen Farbschlägen vorkommt, als Ziervogel gehalten.

Bei rot- und gelbköpfigen Vögeln setzt ein schwarzes Band rund um den Hinterkopf die Kopffärbung vom übrigen Gefieder ab. Bei allen Farbvarianten zeigt sich an Hinterkopf und Kehle außerdem ein hellblaues Band, das allmählich in das grüne Rückengefieder übergeht.

 

Das Gefieder der Flügeldecken ist bei Gouldamadinen ebenfalls grün gefärbt. Auffällig ist die lilafarbene Brust, die scharf gegen den gelb gefiederten Bauch abgesetzt ist. Dieser hellt sich in Richtung Schwanz fast bis zu einem Weiß auf. Der Bürzel sowie die obere Schwanzdecke sind hellblau. Hellblau ist auch der Lidring, der die dunkelbraunen Augen umgibt. Schnabel und Füße sind hornfarben.

 

Die Körperlänge der Vögel beträgt bei beiden Geschlechtern etwa elf Zentimeter. Vom Kopf bis zu den Enden der beiden mittleren Schwanzfedern beträgt ihre Länge zwischen 13 und 15 Zentimeter.

MQCOLL-JML04105

 

Pleurote pétaloïde

 

Pileipellis (à préciser avec photos)

Pileocystides en lamprocystides :

(65,5) 71,8 - 93,2 (95,1) × (5) 5,4 - 6,2 (6,6) µm

Q: (12,4) 12,6 - 15,5 (16,8) ; N: 8

Me : 81 × 5,7 µm ; Qe: 14,2

Trame lamellaire plus ou moins régulière divergente avec hyphes parfois emmêlées mesurant jusqu’à 68,8 x 11 µm et plus, et jusqu’à 0,4 µm de diam.

Boucles présentes à tous les niveaux

Basides clavées, à 4 stérigmates, hyalines, 18,4-34,8 x 5,4-6,8 µm

Stérigmates, 5,4-6,4 x 1,3-1,6 µm

Spores ellipsoïdes, lisses, à paroi mince, hyalines, inamyloïdes, uniguttulées, avec appendice hilaire distinct, 5,8-7,3 x 3,6-4,6 µm, Me : 6,4 x 4,3 µm, Q. : 1,5, N. : 26 spores

Appendice hilaire : jusqu’à 1,06 x 0,7 µm

Cheilocystides polymorphes, fusoïdes-ventrues, fusoïdes cylindriques, lécythiformes, capitées , rarement à col étranglé, parfois ramifiées, à paroi mince à un peu épaissie, avec masse muqueuse apicale brunâtre (gliosphex) souvent présente jusqu’à 8,7 µm de diam., 25,9-36,6 x 4,5-6,5 µm

Pleurocystides très rarement en lamprocystides, fusoïdes-ventrues, lancéolées, à paroi épaissie, hyalines, faiblement jaunâtres dans le KOH, brun rougeâtre dans le Melzer, 46-77 x 9,2-14,6 µm

 

Recherche et révision des travaux: R. Labbé

Étude microscopique, microphotographie et identification: J. Labrecque

 

Photo en studio:

www.flickr.com/photos/19369983@N06/43888097621/in/photoli...

 

Photo sur le terrain:

www.flickr.com/photos/19369983@N06/43029838874/in/datepos...

My Polymorph with some new & more colourful knobs (and less moodily lit).

Calcite is a carbonate mineral and the most stable polymorph of calcium carbonate (CaCO3). The other polymorphs are the minerals aragonite and vaterite. Aragonite will change to calcite at 380–470°C, and vaterite is even less stable.

This is the upright, tree-like form of this variable species. As with the coastal prostrate form, red and white flowered plants can be found growing intermixed. As there are usually no intermediates, the colour is presumably determined by two different alleles of a single gene. It is quite strange that the red/white polymorphism has been retained through the evolution of the prostrate versus the upright forms.

A phylogeny demonstrates that T. pallidum strains infecting baboons in Serengeti National Park and Lake Manyara National Park are genetically distinct from one another.Phylogenies were constructed using both Maximum Parsimony and Maximum Likelihood methods to analyze 25 polymorphisms in six concatenated regions of the Treponema genome. The phylogenies were congruent and a Maximum Parsimony tree was chosen for display, with bootstrap support displayed at all nodes that received greater than 50% using both methods.

As borboletas são insectos da ordem Lepidoptera classificados nas super-famílias Hesperioidea e Papilionoidea, que constituem o grupo informal Rhopalocera.

 

As borboletas têm dois pares de asas membranosas cobertas de escamas e peças bucais adaptadas a sucção. Distinguem-se das traças (mariposas) pelas antenas rectilíneas que terminam numa bola, pelos hábitos de vida diurnos, pela metamorfose que decorre dentro de uma crisálida rígida e pelo abdómen fino e alongado. Quando em repouso, as borboletas dobram as suas asas para cima.

 

As borboletas são importantes polinizadores de diversas espécies de plantas.

 

O ciclo de vida das borboletas engloba as seguintes etapas:

 

1) ovo→ fase pré-larval

2) larva→ chamada também de lagarta ou taturana,

3) pupa→ que se desenvolve dentro da crisálida (ou casulo)

4) imago→ fase adulta

_______________________

 

A butterfly is any of several groups of mainly day-flying insects of the order Lepidoptera, the butterflies and moths. Like other holometabolous insects, butterflies' life cycle consists of four parts, egg, larva, pupa and adult. Most species are diurnal. Butterflies have large, often brightly coloured wings, and conspicuous, fluttering flight. Butterflies comprise the true butterflies (superfamily Papilionoidea), the skippers (superfamily Hesperioidea) and the moth-butterflies (superfamily Hedyloidea). All the many other families within the Lepidoptera are referred to as moths.

 

Butterflies exhibit polymorphism, mimicry and aposematism. Some, like the Monarch, will migrate over long distances. Some butterflies have evolved symbiotic and parasitic relationships with social insects such as ants. Butterflies are important economically as agents of pollination. The caterpillars of some butterflies eat harmful insects. A few species are pests because in their larval stages they can damage domestic crops or trees. Culturally, butterflies are a popular motif in the visual and literary arts.

Dans le cadre de son exposition « Prototypes du Grand Napotakeu (2) » présentée au Cube jusqu’au 22 juillet, Jérôme Lefdup propose une sélection de ses vidéos réalisées au cours des trente dernières années. Ces vidéos sont autant des témoignages des phénomènes étranges observables dans Le Grand Napotakeu, mais aussi une ode à une de ses créatures les plus représentatives et déjà présente dans certaines pièces de son exposition au Cube : la Glute (et ses cousines Polyglutes, Multiglutes, Métaglutes, etc.).

 

La projection est suivie par la présentation et la mise en vente du tirage de tête (10 exemplaires) du livre en 5 volumes « Le Grand Napotakeu ». Les visiteurs munis de lecteur de QR-Codes pourront commander sur place les volumes de leur choix.

 

Une collation amicale après la projection vous convaincra que « les Glutes, c’est super ».

This is the upright, tree-like form of this variable species. As with the coastal prostrate form, red and white flowered plants can be found growing intermixed. As there are usually no intermediates, the colour is presumably determined by two different alleles of a single gene. It is quite strange that the red/white polymorphism has been retained through the evolution of the prostrate versus the upright forms.

A couple of weeks ago Museums and Public relations department had an away day at the Institute of Making.

 

Composite materials station.

Podarcis muralis (common wall lizard) is a species of lizard with a large distribution in Europe and well-established introduced populations in North America, where it is also called the European wall lizard. It can grow to about 20 cm (7.9 in) in total length. The animal has shown variation in the places it has been introduced to. Fossils have been found in a cave in Greece dating to the early part of the Holocene.

 

Description

The common wall lizard is a small, thin lizard whose small scales are highly variable in colour and pattern. Its coloration is generally brownish or greyish, and may occasionally be tinged with green. In some individuals, the row of spots along their backs may form a line, while others may have a reticulated pattern with dark spots on the side and scattered white spots that can be blue in the shoulder region. The tail is brown, grey or rust in colour, and may also have light bars on the sides. The belly region has six rows of larger rectangular scales that are generally reddish, pink, or orangish. Common wall lizards may also have dark markings on the throat. This lizard has six distinct morphological forms which are identified by the colouration of its throat and underbelly. Three of these are pure morphs consisting only of solid colours on their scales: white, red (orange) or yellow, and three other morphs are distinguished by a combination of colours: white-yellow, white-red and yellow-red.

 

Ecology

The common wall lizard prefers rocky environments, including urban settings, where it can scurry between rock, rubble, debris and buildings. In the southern part of its range it tends to occur in humid or semi-humid habitats, compared to drier habitats in the north.

 

Chemical communication

Common wall lizards rely both on visual and chemical signals to communicate with conspecifics. Male wall lizards are equipped with femoral glands, which produce a waxy secretion used for chemical signalling. Both the proteinic and lipophilic compounds in the secretions are known to carry socially relevant information.

 

Polymorphism

The six morphs of this lizard are primarily identifiable by colour and can be distinguished with the naked eye. However, they are commonly confirmed by digital photo and colour analysis. Source of colouration comes from different pigments within the lizard, with yellow colouration originating from carotenoids, and the expression of red/orange colour from pteridines. This is indicative that a phenomenon such as multiple alleles or epistasis may have a role in determining morphology. Morphs also differ in aspects other than colouration: differences between morphs are seen with femoral gland secretion, immune competence and body size. For males, colouration is visible on the underbelly, but in females of all the morphs there is less colouration seen in this region, indicating that sexual dimorphism occurs within this species. A study found sexual dimorphism in the digit ratios. Namely, they found a significant difference (after Bonferroni correction) of the 2D:4D ratio on both forepaws and the 2D:3D ratio on the left forepaw, with all ratios being larger in male animals. This study was conducted on a sample set of 18 male and 18 female museum specimens.

 

Variations between morphs

Aside from differences in colour, the morphs vary by length, survival rate, and immune resistance/response to infection.

 

Susceptibility to infection also varies between morphs, red and yellow-red morphs are the most susceptible to infection by the haemogregarine parasite, a common parasite for these lizards. White morphs are the most resistant to this parasite and the yellow morphs are at an intermediate value of immune resistance and intensity. Certain traits can also lead to a variation in snout-vent length in each of these morphs. For example, in orange morphs, sexual selection favours larger morphs which makes them, on average, larger than the other morphs.

 

The femoral gland secretions of males differ in chemical composition according to each respective morph. Secretions are used by males for intra-species communication, such as marking of territories and attracting potential mates. Individuals share the same organics within the secretions, but the concentrations of certain key compounds differ. For example, α-tocopherol is present in higher concentrations in the secretions of red morphs. This molecule allows for scents to remain in the environment longer by reducing the rate of oxidation in humid environments. This indicates that red morphs tend to be more territorial and maintain territories for longer periods of time compared to other morphs. This phenomenon is also seen when it comes to the age of lizards. Similarly, older, more territorial lizards have higher levels of α-tocopherol compared to younger individuals who are more prone to roaming regardless of morph. Again, indicating that the composition of the secretions relate to the function. Furanones are found in higher concentrations in white morphs, followed by yellow, then red morphs.

 

Among females, reproductive strategies differ by morph; yellow females are r strategists, producing many smaller eggs in their clutch which would allow for numerous offspring to be produced and proliferate in smaller populations with less competition. White females are K strategists, producing fewer, larger eggs which makes offspring more likely to survive in harsher, more competitive environments.[18] Red females can be r or K strategists based on the environment they are in. The existence of alternate strategies points to how morphs have adapted to different environments and hints to the maintenance of colour polymorphism as with fluctuations in environments each morph will eventually be the fittest and as such will not be selected out of the population.

 

The differences observed between morphs are unique evolutionary trade offs employed by each morph to promote survival within the different environments they face in light of limited ability to adapt perfectly to each pressure. Such a trade off is seen when comparing orange morphs to white morphs: morphs, on average, have larger body sizes compared to the white morphs but, in exchange, they are more prone to parasitic infection. In orange morphs, more emphasis is placed on being larger and having the ability to physically compete and ward of potential threats, but in white morphs, metabolic emphasis is placed on having a more hardy immune system to resist infection. Again, these trade offs can arise as there are selective differences in the environment each morph prefers. The differences in the relative proportions of the morphs with respect to location show that environmental pressures differ and some morphs' trade offs are more successful in specific environments than another's.

 

Distribution and status

The natural range spans much of the mainland Europe except from the north and very south and extends to Turkey. It occurs as introduced populations in southern Britain, where one such population in the seaside town of Ventnor on the Isle of Wight has become somewhat famous, and also in North America. There has been some scientific debate as to whether the populations in Southern England represent the northern edge of their native range.

 

North America

Podarcis muralis has been introduced in the United States and is spreading throughout the Cincinnati metropolitan area. It is commonly observed living in limestone outcrops, rock walls, and rubble along the Ohio River basin.

 

It is referred to locally in the Cincinnati/Northern Kentucky area as the "Lazarus lizard", as it was introduced to the area around 1950 by George Rau, a boy who was a member of the family who owned the Lazarus department store chain (now absorbed into Macy's). After he returned from a family vacation to northern Italy, he released about 10 of the reptiles near his Cincinnati home. Genetic testing has revealed that as few as only three of these lizards survived long enough to reproduce, meaning they were subject to an extreme genetic bottleneck. This prolific lizard has reproduced exponentially; it continues to expand its distribution range annually, and has established itself so well in southwest Ohio, it is now considered a naturalized species by the Ohio Department of Natural Resources and is protected under state law (it is illegal to harm, capture, or possess this animal without a proper licence).

 

Besides Ohio, P. muralis has also been introduced in other U.S. states. Populations occur in Kenton and Campbell counties in Kentucky, and in parts of Indiana.

 

The European wall lizard was also introduced to Vancouver Island, British Columbia, Canada in 1970, when a dozen individuals were released into the wild from a small private zoo

The 56 Full Sized Morphs Are:

01 Blaze a Trail | 02 Pearly King Morph | 03 The Messenger Morph | 04 The Power of Morphing Communication | 05 Morph Over, There's Room for Two! | 06 Morph into the Piñataverse | 07 Morpheus | 08 Apart Together | 09 London Parklife | 10 On Guard | 11 Mr Create | 12 Morph's Inspirational Dungarees | 13 Cactus Morph | 14 Forget-Me-Not | 15 Gingerbread Morph I 16 Totally Morphomatic! | 17 Dance-off Morph I 18 The Bard I 19 Mondrian Morph | 20 Morph Whizz Kidz Argonaut | 21 It's Raining Morphs! Halleujah! | 22 Messy Morph | 23 I Spy Morph | 24 Astromorph | 25 Make Your Mark | 26 Roll With It | 27 Morph and Friends Explore London | 28 Tartan Trailblazer | 29 London Collage | 30 Peace Love and Morph | 31 Midas Morph | 32 Freedom | 33 Good Vibes | 34 Tiger Morph | 35 Maximus Morpheus Londinium | 36 Chocks Away! | 37 Morph! It's the Wrong Trousers! | 38 Diverse-City | 39 Apples and Pears | 40 Morphlowers Please! | 41 Cyborg Morph | 42 Pride Morph | 43 The London Man | 44 Looking After the Ocean | 45 Rock Star! | 46 Wheelie | 47 Gentlemorph | 48 Polymorphism | 49 Whizz Bang! | 50 Stay Frosty | 51 Mmmmmmmoprh! | 52 Swashbuckler | 53 Morph Target | 54 Canary Morph | 55 Morph the Yeoman Guard | 56 Fish Ahoy!

 

The 23 Mini Morphs Are:

01 Neville | 02 Messy Morph | 03 Meta-MORPH-osis | 04 Morley the Morph - Ready to Board | 05 Near and Far | 06 Bright Ideas | 07 Creativity Rocks! | 08 Growing Together | 10 Many Hands Make Valence | 11 Mr. Tayo Shnubbub 'The Wellbeing Hero' | 12 Captain Compass I 13 Hands-On & Hands-Up | 14 This is Us | 15 The Adventures of Morph | 16 Our School | 17 Riverside Spirit | 18 Morpheby | 19 GRIT | 20 Happiness is an Inside Job | 21 Growing Together in Learning and in Faith | 22 Look for the Light I 23 Bringing Great Energy and Spirit to Make Things Happen

De la famille des Cactacées.

 

Collecteur : Vladimír Šorma

Espèce : Sulcorebutia steinbachii var. polymorpha

Localisation : Bolivie (Kayarani, Cochabamba)

Date : 1999

Notes : = LH730

 

Acheté sur eBay à kakteendieter en juillet 2012.

Obsidian in the Pleistocene of Wyoming, USA.

 

Obsidian is a glassy-textured, extrusive igneous rock. Glassy-textured rocks have no crystals at all. They form by very rapid cooling of lava or by cooling of high-viscosity lava. Most obsidians form by the latter. Obsidian can be felsic, intermediate, mafic, or alkaline in chemistry. Most are felsic to intermediate.

 

A famous locality in North America is Obsidian Cliff at Yellowstone, Wyoming. It is a Pleistocene-aged lava flow with the chemistry of rhyolite (= a light-colored, felsic, aphanitic, extrusive igneous rock). The cliff itself shows columnar jointing. The rocks principally range from aphyric rhyolitic obsidian to partially devitrified rhyolitic obsidian. Lithophysae are sometimes present. Extremely small, microscopic crystals are present - they can be seen in thin sections. Some samples are reported to have small olivine phenocrysts. Small clusters of crystals, composed of plagioclase feldspar, pyroxene, and olivine, are sometimes present.

 

Many of the whitish-colored spots and bands running through most Obsidian Cliff rock samples are areas of devitrification. Glass is unstable on geologic times scales and it slowly crystallizes. The light-colored spots and bands are now non-glassy. Spotted, partially devitrified obsidian is known by the rockhound term "snowflake obsidian" (see: www.flickr.com/photos/jsjgeology/16561606417). The spots are composed of silica (SiO2), but are not quartz. Rather, they are composed of a polymorph of quartz - cristobalite.

 

The subrounded cavities are lithophysae - they formed before the rock completely solidified. The original lava flow had some subspherical structures known as spherulites, composed of glassy to cryptocrystalline material (many felsic extrusive igneous rocks have these). Expanding gases in the spherulites destroyed this material, resulting in partially empty spaces.

 

Stratigraphy: Roaring Mountain Member, Plateau Rhyolite, Upper Pleistocene, ~59 ka

 

Locality: loose boulder near the base of Obsidian Cliff, Yellowstone National Park, northwestern Wyoming, USA

----------------------

Age & some lithologic info. from:

 

Wooton (2010) - Age and Petrogenesis of the Roaring Mountain Rhyolites, Yellowstone Volcanic Field, Wyoming. M.S. thesis. University of Nevada at Las Vegas. 296 pp.

 

As borboletas são insectos da ordem Lepidoptera classificados nas super-famílias Hesperioidea e Papilionoidea, que constituem o grupo informal Rhopalocera.

 

As borboletas têm dois pares de asas membranosas cobertas de escamas e peças bucais adaptadas a sucção. Distinguem-se das traças (mariposas) pelas antenas rectilíneas que terminam numa bola, pelos hábitos de vida diurnos, pela metamorfose que decorre dentro de uma crisálida rígida e pelo abdómen fino e alongado. Quando em repouso, as borboletas dobram as suas asas para cima.

 

As borboletas são importantes polinizadores de diversas espécies de plantas.

 

O ciclo de vida das borboletas engloba as seguintes etapas:

 

1) ovo→ fase pré-larval

2) larva→ chamada também de lagarta ou taturana,

3) pupa→ que se desenvolve dentro da crisálida (ou casulo)

4) imago→ fase adulta

_______________________

 

A butterfly is any of several groups of mainly day-flying insects of the order Lepidoptera, the butterflies and moths. Like other holometabolous insects, butterflies' life cycle consists of four parts, egg, larva, pupa and adult. Most species are diurnal. Butterflies have large, often brightly coloured wings, and conspicuous, fluttering flight. Butterflies comprise the true butterflies (superfamily Papilionoidea), the skippers (superfamily Hesperioidea) and the moth-butterflies (superfamily Hedyloidea). All the many other families within the Lepidoptera are referred to as moths.

 

Butterflies exhibit polymorphism, mimicry and aposematism. Some, like the Monarch, will migrate over long distances. Some butterflies have evolved symbiotic and parasitic relationships with social insects such as ants. Butterflies are important economically as agents of pollination. The caterpillars of some butterflies eat harmful insects. A few species are pests because in their larval stages they can damage domestic crops or trees. Culturally, butterflies are a popular motif in the visual and literary arts.

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