I can't think of a title for this one, although it does remind me of something, like my new lens. Feel free to add a tag of anything that may come to your mind when you look at this.

 

Testosterone

The chemical structure of testosterone.

A ball-and-stick model of testosterone.

Names

IUPAC name

17β-Hydroxyandrost-4-en-3-one

Systematic IUPAC name

(8R,9S,10R,13S,14S,17S)-17-Hydroxy-10,13-dimethyl-1,2,6,7,8,9,11,12,14,15,16,17-dodecahydrocyclopenta[a]phenanthren-3-one

Other names

Androst-4-en-17β-ol-3-one

Identifiers

CAS Number

58-22-0 ☑

3D model (JSmol)

Interactive image

ChEBI

CHEBI:17347 ☑

ChEMBL

ChEMBL386630 ☑

ChemSpider

5791 ☑

DrugBank

DB00624 ☑

ECHA InfoCard100.000.336

KEGG

D00075 ☑

PubChem CID

6013

UNII

3XMK78S47O ☑

InChI[show]

SMILES[show]

Properties

Chemical formula

C19H28O2

Molar mass288.431 g·mol−1

Melting point155 °C

Pharmacology

ATC code

G03BA03 (WHO)

License data

EU EMA: by INN

Routes of

administration

Transdermal (gel, cream, solution, patch), by mouth (as testosterone undecanoate), in the cheek, intranasal (gel), intramuscular injection (as esters), subcutaneous pellets

Pharmacokinetics:

Bioavailability

Oral: very low (due to extensive first pass metabolism)

Protein binding

97.0–99.5% (to SHBG and albumin)[1]

Metabolism

Liver (mainly reduction and conjugation)

Biological half-life

2–4 hours[citation needed]

Excretion

Urine (90%), feces (6%)

Except where otherwise noted, data are given for materials in their standard state (at 25 °C [77 °F], 100 kPa).

☑ verify (what is ☑☒ ?)

Infobox references

Testosterone is the primary male sex hormone and an anabolic steroid. In male humans, testosterone plays a key role in the development of male reproductive tissues such as testes and prostate, as well as promoting secondary sexual characteristics such as increased muscle and bone mass, and the growth of body hair.[2] In addition, testosterone is involved in health and well-being,[3] and the prevention of osteoporosis.[4] Insufficient levels of testosterone in men may lead to abnormalities including frailty and bone loss.

 

Testosterone is a steroid from the androstane class containing a keto and hydroxyl groups at the three and seventeen positions respectively. It is biosynthesized in several steps from cholesterol and is converted in the liver to inactive metabolites.[5] It exerts its action through binding to and activation of the androgen receptor.[5] In humans and most other vertebrates, testosterone is secreted primarily by the testicles of males and, to a lesser extent, the ovaries of females. On average, in adult males, levels of testosterone are about 7 to 8 times as great as in adult females.[6] As the metabolism of testosterone in males is more pronounced, the daily production is about 20 times greater in men.[7][8] Females are also more sensitive to the hormone.[9]

 

In addition to its role as a natural hormone, testosterone is used as a medication, for instance in the treatment of low testosterone levels in men and breast cancer in women.[10] Since testosterone levels decrease as men age, testosterone is sometimes used in older men to counteract this deficiency. It is also used illicitly to enhance physique and performance, for instance in athletes.

  

Contents

1Biological effects

1.1Before birth

1.2Early infancy

1.3Before puberty

1.4Pubertal

1.5Adult

1.6Aggression and criminality

1.7Brain

2Medical use

3Biological activity

3.1Steroid hormone activity

3.2Neurosteroid activity

4Biochemistry

4.1Biosynthesis

4.2Distribution

4.3Metabolism

4.4Levels

5Measurement

6History

7Other animals

8See also

9References

10Further reading

Biological effects[edit]

In general, androgens such as testosterone promote protein synthesis and thus growth of tissues with androgen receptors.[11] Testosterone can be described as having virilising and anabolic effects (though these categorical descriptions are somewhat arbitrary, as there is a great deal of mutual overlap between them).[12]

 

Anabolic effects include growth of muscle mass and strength, increased bone density and strength, and stimulation of linear growth and bone maturation.

Androgenic effects include maturation of the sex organs, particularly the penis and the formation of the scrotum in the fetus, and after birth (usually at puberty) a deepening of the voice, growth of facial hair (such as the beard) and axillary (underarm) hair. Many of these fall into the category of male secondary sex characteristics.

Testosterone effects can also be classified by the age of usual occurrence. For postnatal effects in both males and females, these are mostly dependent on the levels and duration of circulating free testosterone.

 

Before birth[edit]

Effects before birth are divided into two categories, classified in relation to the stages of development.

 

The first period occurs between 4 and 6 weeks of the gestation. Examples include genital virilisation such as midline fusion, phallic urethra, scrotal thinning and rugation, and phallic enlargement; although the role of testosterone is far smaller than that of dihydrotestosterone. There is also development of the prostate gland and seminal vesicles.

 

During the second trimester, androgen level is associated with sex formation.[13] This period affects the femininization or masculinization of the fetus and can be a better predictor of feminine or masculine behaviours such as sex typed behaviour than an adult's own levels. A mother's testosterone level during pregnancy is correlated with her daughter's sex-typical behavior as an adult, and the correlation is even stronger than with the daughter's own adult testosterone level.[14]

 

Early infancy[edit]

Early infancy androgen effects are the least understood. In the first weeks of life for male infants, testosterone levels rise. The levels remain in a pubertal range for a few months, but usually reach the barely detectable levels of childhood by 4–7 months of age.[15][16] The function of this rise in humans is unknown. It has been theorized that brain masculinization is occurring since no significant changes have been identified in other parts of the body.[17] The male brain is masculinized by the aromatization of testosterone into estrogen, which crosses the blood–brain barrier and enters the male brain, whereas female fetuses have α-fetoprotein, which binds the estrogen so that female brains are not affected.[18]

 

Before puberty[edit]

Before puberty effects of rising androgen levels occur in both boys and girls. These include adult-type body odor, increased oiliness of skin and hair, acne, pubarche (appearance of pubic hair), axillary hair (armpit hair), growth spurt, accelerated bone maturation, and facial hair.[19]

 

Pubertal[edit]

Pubertal effects begin to occur when androgen has been higher than normal adult female levels for months or years. In males, these are usual late pubertal effects, and occur in women after prolonged periods of heightened levels of free testosterone in the blood. The effects include:[19][20]

 

Growth of spermatogenic tissue in testicles, male fertility, penis or clitoris enlargement, increased libido and frequency of erection or clitoral engorgement occurs. Growth of jaw, brow, chin, and nose and remodeling of facial bone contours, in conjunction with human growth hormone occurs.[21] Completion of bone maturation and termination of growth. This occurs indirectly via estradiol metabolites and hence more gradually in men than women. Increased muscle strength and mass, shoulders become broader and rib cage expands, deepening of voice, growth of the Adam's apple. Enlargement of sebaceous glands. This might cause acne, subcutaneous fat in face decreases. Pubic hair extends to thighs and up toward umbilicus, development of facial hair (sideburns, beard, moustache), loss of scalp hair (androgenetic alopecia), increase in chest hair, periareolar hair, perianal hair, leg hair, armpit hair.

 

Adult[edit]

Testosterone is necessary for normal sperm development. It activates genes in Sertoli cells, which promote differentiation of spermatogonia. It regulates acute HPA (hypothalamic–pituitary–adrenal axis) response under dominance challenge.[22] Androgen including testosterone enhances muscle growth. Testosterone also regulates the population of thromboxane A2 receptors on megakaryocytes and platelets and hence platelet aggregation in humans.[23][24]

 

Adult testosterone effects are more clearly demonstrable in males than in females, but are likely important to both sexes. Some of these effects may decline as testosterone levels might decrease in the later decades of adult life.[25]

 

Health risks[edit]

Testosterone does not appear to increase the risk of developing prostate cancer. In people who have undergone testosterone deprivation therapy, testosterone increases beyond the castrate level have been shown to increase the rate of spread of an existing prostate cancer.[26][27][28]

 

Conflicting results have been obtained concerning the importance of testosterone in maintaining cardiovascular health.[29][30] Nevertheless, maintaining normal testosterone levels in elderly men has been shown to improve many parameters that are thought to reduce cardiovascular disease risk, such as increased lean body mass, decreased visceral fat mass, decreased total cholesterol, and glycemic control.[31]

 

High androgen levels are associated with menstrual cycle irregularities in both clinical populations and healthy women.[32]

 

Sexual arousal[edit]

See also: Hormones and sexual arousal

When testosterone and endorphins in ejaculated semen meet the cervical wall after sexual intercourse, females receive a spike in testosterone, endorphin, and oxytocin levels, and males after orgasm during copulation experience an increase in endorphins and a marked increase in oxytocin levels. This adds to the hospitable physiological environment in the female internal reproductive tract for conceiving, and later for nurturing the conceptus in the pre-embryonic stages, and stimulates feelings of love, desire, and paternal care in the male (this is the only time male oxytocin levels rival a female's).[citation needed]

 

Testosterone levels follow a nyctohemeral rhythm that peaks early each day, regardless of sexual activity.[33]

 

There are positive correlations between positive orgasm experience in women and testosterone levels where relaxation was a key perception of the experience. There is no correlation between testosterone and men's perceptions of their orgasm experience, and also no correlation between higher testosterone levels and greater sexual assertiveness in either sex.[34]

 

Sexual arousal and masturbation in women produce small increases in testosterone concentrations.[35] The plasma levels of various steroids significantly increase after masturbation in men and the testosterone levels correlate to those levels.[36]

 

Mammalian studies[edit]

Studies conducted in rats have indicated that their degree of sexual arousal is sensitive to reductions in testosterone. When testosterone-deprived rats were given medium levels of testosterone, their sexual behaviors (copulation, partner preference, etc.) resumed, but not when given low amounts of the same hormone. Therefore, these mammals may provide a model for studying clinical populations among humans suffering from sexual arousal deficits such as hypoactive sexual desire disorder.[37]

 

In every mammalian species examined demonstrated a marked increase in a male's testosterone level upon encountering a novel female. The reflexive testosterone increases in male mice is related to the male's initial level of sexual arousal.[38]

 

In non-human primates, it may be that testosterone in puberty stimulates sexual arousal, which allows the primate to increasingly seek out sexual experiences with females and thus creates a sexual preference for females.[39] Some research has also indicated that if testosterone is eliminated in an adult male human or other adult male primate's system, its sexual motivation decreases, but there is no corresponding decrease in ability to engage in sexual activity (mounting, ejaculating, etc.).[39]

 

In accordance with sperm competition theory, testosterone levels are shown to increase as a response to previously neutral stimuli when conditioned to become sexual in male rats.[40] This reaction engages penile reflexes (such as erection and ejaculation) that aid in sperm competition when more than one male is present in mating encounters, allowing for more production of successful sperm and a higher chance of reproduction.

 

Males[edit]

In men, higher levels of testosterone are associated with periods of sexual activity.[41][42]

 

Men who watch a sexually explicit movie have an average increase of 35% in testosterone, peaking at 60–90 minutes after the end of the film, but no increase is seen in men who watch sexually neutral films.[43] Men who watch sexually explicit films also report increased motivation, competitiveness, and decreased exhaustion.[44] A link has also been found between relaxation following sexual arousal and testosterone levels.[45]

 

Men's levels of testosterone, a hormone known to affect men's mating behaviour, changes depending on whether they are exposed to an ovulating or nonovulating woman's body odour. Men who are exposed to scents of ovulating women maintained a stable testosterone level that was higher than the testosterone level of men exposed to nonovulation cues. Testosterone levels and sexual arousal in men are heavily aware of hormone cycles in females.[46] This may be linked to the ovulatory shift hypothesis,[47] where males are adapted to respond to the ovulation cycles of females by sensing when they are most fertile and whereby females look for preferred male mates when they are the most fertile; both actions may be driven by hormones.

 

Females[edit]

Androgens may modulate the physiology of vaginal tissue and contribute to female genital sexual arousal.[48] Women's level of testosterone is higher when measured pre-intercourse vs pre-cuddling, as well as post-intercourse vs post-cuddling.[49] There is a time lag effect when testosterone is administered, on genital arousal in women. In addition, a continuous increase in vaginal sexual arousal may result in higher genital sensations and sexual appetitive behaviors.[50]

 

When females have a higher baseline level of testosterone, they have higher increases in sexual arousal levels but smaller increases in testosterone, indicating a ceiling effect on testosterone levels in females. Sexual thoughts also change the level of testosterone but not level of cortisol in the female body, and hormonal contraceptives may affect the variation in testosterone response to sexual thoughts.[51]

 

Testosterone may prove to be an effective treatment in female sexual arousal disorders,[52] and is available as a dermal patch. There is no FDA approved androgen preparation for the treatment of androgen insufficiency; however, it has been used off-label to treat low libido and sexual dysfunction in older women. Testosterone may be a treatment for postmenopausal women as long as they are effectively estrogenized.[52]

 

Romantic relationships[edit]

Falling in love decreases men's testosterone levels while increasing women's testosterone levels. There has been speculation that these changes in testosterone result in the temporary reduction of differences in behavior between the sexes.[53] However, it is suggested that after the "honeymoon phase" ends—about four years into a relationship—this change in testosterone levels is no longer apparent.[53] Men who produce less testosterone are more likely to be in a relationship[54] or married,[55] and men who produce more testosterone are more likely to divorce;[55] however, causality cannot be determined in this correlation. Marriage or commitment could cause a decrease in testosterone levels.[56] Single men who have not had relationship experience have lower testosterone levels than single men with experience. It is suggested that these single men with prior experience are in a more competitive state than their non-experienced counterparts.[57] Married men who engage in bond-maintenance activities such as spending the day with their spouse/and or child have no different testosterone levels compared to times when they do not engage in such activities. Collectively, these results suggest that the presence of competitive activities rather than bond-maintenance activities are more relevant to changes in testosterone levels.[58]

 

Men who produce more testosterone are more likely to engage in extramarital sex.[55] Testosterone levels do not rely on physical presence of a partner; testosterone levels of men engaging in same-city and long-distance relationships are similar.[54] Physical presence may be required for women who are in relationships for the testosterone–partner interaction, where same-city partnered women have lower testosterone levels than long-distance partnered women.[59]

 

Fatherhood[edit]

Fatherhood decreases testosterone levels in men, suggesting that the emotions and behavior tied to decreased testosterone promote paternal care. In humans and other species that utilize allomaternal care, paternal investment in offspring is beneficial to said offspring's survival because it allows the parental dyad to raise multiple children simultaneously. This increases the reproductive fitness of the parents, because their offspring are more likely to survive and reproduce. Paternal care increases offspring survival due to increased access to higher quality food and reduced physical and immunological threats.[60] This is particularly beneficial for humans since offspring are dependent on parents for extended periods of time and mothers have relatively short inter-birth intervals.[61] While extent of paternal care varies between cultures, higher investment in direct child care has been seen to be correlated with lower average testosterone levels as well as temporary fluctuations.[62] For instance, fluctuation in testosterone levels when a child is in distress has been found to be indicative of fathering styles. If a father's testosterone levels decrease in response to hearing their baby cry, it is an indication of empathizing with the baby. This is associated with increased nurturing behavior and better outcomes for the infant.[63]

 

Motivation[edit]

Testosterone levels play a major role in risk-taking during financial decisions.[64][65]

 

Aggression and criminality [edit]

See also: Aggression § Testosterone, and Biosocial criminology

Most studies support a link between adult criminality and testosterone, although the relationship is modest if examined separately for each sex. Nearly all studies of juvenile delinquency and testosterone are not significant. Most studies have also found testosterone to be associated with behaviors or personality traits linked with criminality such as antisocial behavior and alcoholism. Many studies have also been done on the relationship between more general aggressive behavior/feelings and testosterone. About half the studies have found a relationship and about half no relationship.[66]

 

Testosterone is only one of many factors that influence aggression and the effects of previous experience and environmental stimuli have been found to correlate more strongly. A few studies indicate that the testosterone derivative estradiol (one form of estrogen) might play an important role in male aggression.[66][67][68][69] Studies have also found that testosterone facilitates aggression by modulating vasopressin receptors in the hypothalamus.[70]

 

The sexual hormone can encourage fair behavior. For the study, subjects took part in a behavioral experiment where the distribution of a real amount of money was decided. The rules allowed both fair and unfair offers. The negotiating partner could subsequently accept or decline the offer. The fairer the offer, the less probable a refusal by the negotiating partner. If no agreement was reached, neither party earned anything. Test subjects with an artificially enhanced testosterone level generally made better, fairer offers than those who received placebos, thus reducing the risk of a rejection of their offer to a minimum. Two later studies have empirically confirmed these results.[71][72][73] However men with high testosterone were significantly 27% less generous in an ultimatum game.[74] The Annual NY Academy of Sciences has also found anabolic steroid use which increase testosterone to be higher in teenagers, and this was associated with increased violence.[75] Studies have also found administered testosterone to increase verbal aggression and anger in some participants.[76]

 

Testosterone is significantly correlated with aggression and competitive behaviour and is directly facilitated by the latter. There are two theories on the role of testosterone in aggression and competition.[77] The first one is the challenge hypothesis which states that testosterone would increase during puberty thus facilitating reproductive and competitive behaviour which would include aggression.[77] Thus it is the challenge of competition among males of the species that facilitates aggression and violence.[77] Studies conducted have found direct correlation between testosterone and dominance especially among the most violent criminals in prison who had the highest testosterone levels.[77] The same research also found fathers (those outside competitive environments) had the lowest testosterone levels compared to other males.[77]

 

The second theory is similar and is known as "evolutionary neuroandrogenic (ENA) theory of male aggression".[78][79] Testosterone and other androgens have evolved to masculinize a brain in order to be competitive even to the point of risking harm to the person and others. By doing so, individuals with masculinized brains as a result of pre-natal and adult life testosterone and androgens enhance their resource acquiring abilities in order to survive, attract and copulate with mates as much as possible.[78] The masculinization of the brain is not just mediated by testosterone levels at the adult stage, but also testosterone exposure in the womb as a fetus. Higher pre-natal testosterone indicated by a low digit ratio as well as adult testosterone levels increased risk of fouls or aggression among male players in a soccer game.[80] Studies have also found higher pre-natal testosterone or lower digit ratio to be correlated with higher aggression in males.[81][82][83][84][85]

 

The rise in testosterone levels during competition predicted aggression in males but not in females.[86] Subjects who interacted with hand guns and an experimental game showed rise in testosterone and aggression.[87] Natural selection might have evolved males to be more sensitive to competitive and status challenge situations and that the interacting roles of testosterone are the essential ingredient for aggressive behaviour in these situations.[88] Testosterone produces aggression by activating subcortical areas in the brain, which may also be inhibited or suppressed by social norms or familial situations while still manifesting in diverse intensities and ways through thoughts, anger, verbal aggression, competition, dominance and physical violence.[89] Testosterone mediates attraction to cruel and violent cues in men by promoting extended viewing of violent stimuli.[90] Testosterone specific structural brain characteristic can predict aggressive behaviour in individuals.[91]

 

Estradiol is known to correlate with aggression in male mice.[92] Moreover, the conversion of testosterone to estradiol regulates male aggression in sparrows during breeding season.[93] Rats who were given anabolic steroids that increase testosterone were also more physically aggressive to provocation as a result of "threat sensitivity".[94]

 

Brain[edit]

The brain is also affected by this sexual differentiation;[13] the enzyme aromatase converts testosterone into estradiol that is responsible for masculinization of the brain in male mice. In humans, masculinization of the fetal brain appears, by observation of gender preference in patients with congenital diseases of androgen formation or androgen receptor function, to be associated with functional androgen receptors.[95]

 

There are some differences between a male and female brain (possibly the result of different testosterone levels), one of them being size: the male human brain is, on average, larger.[96] Men were found to have a total myelinated fiber length of 176 000 km at the age of 20, whereas in women the total length was 149 000 km (approx. 15% less).[97]

 

No immediate short term effects on mood or behavior were found from the administration of supraphysiologic doses of testosterone for 10 weeks on 43 healthy men.[98] A correlation between testosterone and risk tolerance in career choice exists among women.[64][99]

 

Attention, memory, and spatial ability are key cognitive functions affected by testosterone in humans. Preliminary evidence suggests that low testosterone levels may be a risk factor for cognitive decline and possibly for dementia of the Alzheimer's type,[100][101][102][103] a key argument in life extension medicine for the use of testosterone in anti-aging therapies. Much of the literature, however, suggests a curvilinear or even quadratic relationship between spatial performance and circulating testosterone,[104] where both hypo- and hypersecretion (deficient- and excessive-secretion) of circulating androgens have negative effects on cognition.

 

Medical use[edit]

Main article: Testosterone (medication)

Testosterone is used as a medication for the treatment of males with too little or no natural testosterone production, certain forms of breast cancer,[10] and gender dysphoria in transgender men. This is known as hormone replacement therapy (HRT) or testosterone replacement therapy (TRT), which maintains serum testosterone levels in the normal range. Decline of testosterone production with age has led to interest in androgen replacement therapy.[105] It is unclear if the use of testosterone for low levels due to aging is beneficial or harmful.[106]

 

Testosterone is included in the World Health Organization's list of essential medicines, which are the most important medications needed in a basic health system.[107] It is available as a generic medication.[10] The price depends on the form of testosterone used.[108] It can be administered as a cream or transdermal patch that is applied to the skin, by injection into a muscle, as a tablet that is placed in the cheek, or by ingestion.[10]

 

Common side effects from testosterone medication include acne, swelling, and breast enlargement in males.[10] Serious side effects may include liver toxicity, heart disease, and behavioral changes.[10] Women and children who are exposed may develop virilization.[10] It is recommended that individuals with prostate cancer not use the medication.[10] It can cause harm if used during pregnancy or breastfeeding.[10]

 

Biological activity[edit]

Steroid hormone activity[edit]

The effects of testosterone in humans and other vertebrates occur by way of multiple mechanisms: by activation of the androgen receptor (directly or as DHT), and by conversion to estradiol and activation of certain estrogen receptors.[109][110] Androgens such as testosterone have also been found to bind to and activate membrane androgen receptors.[111][112][113]

 

Free testosterone (T) is transported into the cytoplasm of target tissue cells, where it can bind to the androgen receptor, or can be reduced to 5α-dihydrotestosterone (DHT) by the cytoplasmic enzyme 5α-reductase. DHT binds to the same androgen receptor even more strongly than testosterone, so that its androgenic potency is about 5 times that of T.[114] The T-receptor or DHT-receptor complex undergoes a structural change that allows it to move into the cell nucleus and bind directly to specific nucleotide sequences of the chromosomal DNA. The areas of binding are called hormone response elements (HREs), and influence transcriptional activity of certain genes, producing the androgen effects.

 

Androgen receptors occur in many different vertebrate body system tissues, and both males and females respond similarly to similar levels. Greatly differing amounts of testosterone prenatally, at puberty, and throughout life account for a share of biological differences between males and females.

 

The bones and the brain are two important tissues in humans where the primary effect of testosterone is by way of aromatization to estradiol. In the bones, estradiol accelerates ossification of cartilage into bone, leading to closure of the epiphyses and conclusion of growth. In the central nervous system, testosterone is aromatized to estradiol. Estradiol rather than testosterone serves as the most important feedback signal to the hypothalamus (especially affecting LH secretion).[115] In many mammals, prenatal or perinatal "masculinization" of the sexually dimorphic areas of the brain by estradiol derived from testosterone programs later male sexual behavior.[116]

 

Neurosteroid activity[edit]

Testosterone, via its active metabolite 3α-androstanediol, is a potent positive allosteric modulator of the GABAA receptor.[117]

 

Testosterone has been found to act as an antagonist of the TrkA and p75NTR, receptors for the neurotrophin nerve growth factor (NGF), with high affinity (around 5 nM).[118][119][120] In contrast to testosterone, DHEA and DHEA sulfate have been found to act as high-affinity agonists of these receptors.[118][119][120]

 

Testosterone is an antagonist of the sigma σ1 receptor (Ki = 1,014 or 201 nM).[121] However, the concentrations of testosterone required for binding the receptor are far above even total circulating concentrations of testosterone in adult males (which range between 10 and 35 nM).[122]

 

Biochemistry[edit]

 

Human steroidogenesis, showing testosterone near bottom.[123]

Biosynthesis[edit]

Like other steroid hormones, testosterone is derived from cholesterol (see figure).[124] The first step in the biosynthesis involves the oxidative cleavage of the side-chain of cholesterol by cholesterol side-chain cleavage enzyme (P450scc, CYP11A1), a mitochondrial cytochrome P450 oxidase with the loss of six carbon atoms to give pregnenolone. In the next step, two additional carbon atoms are removed by the CYP17A1 (17α-hydroxylase/17,20-lyase) enzyme in the endoplasmic reticulum to yield a variety of C19 steroids.[125] In addition, the 3β-hydroxyl group is oxidized by 3β-hydroxysteroid dehydrogenase to produce androstenedione. In the final and rate limiting step, the C17 keto group androstenedione is reduced by 17β-hydroxysteroid dehydrogenase to yield testosterone.

 

The largest amounts of testosterone (>95%) are produced by the testes in men,[2] while the adrenal glands account for most of the remainder. Testosterone is also synthesized in far smaller total quantities in women by the adrenal glands, thecal cells of the ovaries, and, during pregnancy, by the placenta.[126] In the testes, testosterone is produced by the Leydig cells.[127] The male generative glands also contain Sertoli cells, which require testosterone for spermatogenesis. Like most hormones, testosterone is supplied to target tissues in the blood where much of it is transported bound to a specific plasma protein, sex hormone-binding globulin (SHBG).

 

Regulation[edit]

 

Hypothalamic–pituitary–testicular axis

In males, testosterone is synthesized primarily in Leydig cells. The number of Leydig cells in turn is regulated by luteinizing hormone (LH) and follicle-stimulating hormone (FSH). In addition, the amount of testosterone produced by existing Leydig cells is under the control of LH, which regulates the expression of 17β-hydroxysteroid dehydrogenase.[128]

 

The amount of testosterone synthesized is regulated by the hypothalamic–pituitary–testicular axis (see figure to the right).[129] When testosterone levels are low, gonadotropin-releasing hormone (GnRH) is released by the hypothalamus, which in turn stimulates the pituitary gland to release FSH and LH. These latter two hormones stimulate the testis to synthesize testosterone. Finally, increasing levels of testosterone through a negative feedback loop act on the hypothalamus and pituitary to inhibit the release of GnRH and FSH/LH, respectively.

 

Factors affecting testosterone levels may include:

 

Age: Testosterone levels gradually reduce as men age.[130][131] This effect is sometimes referred to as andropause or late-onset hypogonadism.[132]

Exercise: Resistance training increases testosterone levels,[133] however, in older men, that increase can be avoided by protein ingestion.[134] Endurance training in men may lead to lower testosterone levels.[135]

Nutrients: Vitamin A deficiency may lead to sub-optimal plasma testosterone levels.[136] The secosteroid vitamin D in levels of 400–1000 IU/d (10–25 µg/d) raises testosterone levels.[137] Zinc deficiency lowers testosterone levels[138] but over-supplementation has no effect on serum testosterone.[139]

Weight loss: Reduction in weight may result in an increase in testosterone levels. Fat cells synthesize the enzyme aromatase, which converts testosterone, the male sex hormone, into estradiol, the female sex hormone.[140] However no clear association between body mass index and testosterone levels has been found.[141]

Miscellaneous: Sleep: (REM sleep) increases nocturnal testosterone levels.[142] Behavior: Dominance challenges can, in some cases, stimulate increased testosterone release in men.[143] Drugs: Natural or man-made antiandrogens including spearmint tea reduce testosterone levels.[144][145][146] Licorice can decrease the production of testosterone and this effect is greater in females.[147]

Distribution[edit]

The plasma protein binding of testosterone is 98.0 to 98.5%, with 1.5 to 2.0% free or unbound.[148] It is bound 65% to sex hormone-binding globulin (SHBG) and 33% bound weakly to albumin.[149]

 

Plasma protein binding of testosterone and dihydrotestosterone show

Metabolism[edit]

vte Testosterone metabolism in humans

Testosterone structures

The image above contains clickable linksTestosterone metabolism in humans. Conjugation (sulfation and glucuronidation) occurs both with testosterone and with all of the other steroids that have one or more available hydroxyl (-OH) groups in this diagram.

Both testosterone and 5α-DHT are metabolized mainly in the liver.[1][151] Approximately 50% of testosterone is metabolized via conjugation into testosterone glucuronide and to a lesser extent testosterone sulfate by glucuronosyltransferases and sulfotransferases, respectively.[1] An additional 40% of testosterone is metabolized in equal proportions into the 17-ketosteroids androsterone and etiocholanolone via the combined actions of 5α- and 5β-reductases, 3α-hydroxysteroid dehydrogenase, and 17β-HSD, in that order.[1][151][152] Androsterone and etiocholanolone are then glucuronidated and to a lesser extent sulfated similarly to testosterone.[1][151] The conjugates of testosterone and its hepatic metabolites are released from the liver into circulation and excreted in the urine and bile.[1][151][152] Only a small fraction (2%) of testosterone is excreted unchanged in the urine.[151]

 

In the hepatic 17-ketosteroid pathway of testosterone metabolism, testosterone is converted in the liver by 5α-reductase and 5β-reductase into 5α-DHT and the inactive 5β-DHT, respectively.[1][151] Then, 5α-DHT and 5β-DHT are converted by 3α-HSD into 3α-androstanediol and 3α-etiocholanediol, respectively.[1][151] Subsequently, 3α-androstanediol and 3α-etiocholanediol are converted by 17β-HSD into androsterone and etiocholanolone, which is followed by their conjugation and excretion.[1][151] 3β-Androstanediol and 3β-etiocholanediol can also be formed in this pathway when 5α-DHT and 5β-DHT are acted upon by 3β-HSD instead of 3α-HSD, respectively, and they can then be transformed into epiandrosterone and epietiocholanolone, respectively.[153][154] A small portion of approximately 3% of testosterone is reversibly converted in the liver into androstenedione by 17β-HSD.[152]

 

In addition to conjugation and the 17-ketosteroid pathway, testosterone can also be hydroxylated and oxidized in the liver by cytochrome P450 enzymes, including CYP3A4, CYP3A5, CYP2C9, CYP2C19, and CYP2D6.[155] 6β-Hydroxylation and to a lesser extent 16β-hydroxylation are the major transformations.[155] The 6β-hydroxylation of testosterone is catalyzed mainly by CYP3A4 and to a lesser extent CYP3A5 and is responsible for 75 to 80% of cytochrome P450-mediated testosterone metabolism.[155] In addition to 6β- and 16β-hydroxytestosterone, 1β-, 2α/β-, 11β-, and 15β-hydroxytestosterone are also formed as minor metabolites.[155][156] Certain cytochrome P450 enzymes such as CYP2C9 and CYP2C19 can also oxidize testosterone at the C17 position to form androstenedione.[155]

 

Two of the immediate metabolites of testosterone, 5α-DHT and estradiol, are biologically important and can be formed both in the liver and in extrahepatic tissues.[151] Approximately 5 to 7% of testosterone is converted by 5α-reductase into 5α-DHT, with circulating levels of 5α-DHT about 10% of those of testosterone, and approximately 0.3% of testosterone is converted into estradiol by aromatase.[2][151][157][158] 5α-Reductase is highly expressed in the male reproductive organs (including the prostate gland, seminal vesicles, and epididymides),[159] skin, hair follicles, and brain[160] and aromatase is highly expressed in adipose tissue, bone, and the brain.[161][162] As much as 90% of testosterone is converted into 5α-DHT in so-called androgenic tissues with high 5α-reductase expression,[152] and due to the several-fold greater potency of 5α-DHT as an AR agonist relative to testosterone,[163] it has been estimated that the effects of testosterone are potentiated 2- to 3-fold in such tissues.[164]

 

Levels[edit]

Total levels of testosterone in the body are 264 to 916 ng/dL in men age 19 to 39 years,[165] while mean testosterone levels in adult men have been reported as 630 ng/dL.[166] Levels of testosterone in men decline with age.[165] In women, mean levels of total testosterone have been reported to be 32.6 ng/dL.[167][168] In women with hyperandrogenism, mean levels of total testosterone have been reported to be 62.1 ng/dL.[167][168]

 

Testosterone levels in males and females show

Total testosterone levels in males throughout life show

 

Reference ranges for blood tests, showing adult male testosterone levels in light blue at center-left.

Measurement[edit]

Testosterone’s bioavailable concentration is commonly determined using the Vermeulen calculation or more precisely using the modified Vermeulen method,[174][175] which considers the dimeric form of sex-hormone-binding-globulin.[176]

 

Both methods use chemical equilibrium to derive the concentration of bioavailable testosterone: in circulation testosterone has two major binding partners, albumin (weakly bound) and sex-hormone-binding-globulin (strongly bound). These methods are described in detail in the accompanying figure.

  

Dimeric sex-hormone-binding-globulin with its testosterone ligands

  

Two methods for determining concentration of bioavailable testosterone.

History[edit]

A testicular action was linked to circulating blood fractions – now understood to be a family of androgenic hormones – in the early work on castration and testicular transplantation in fowl by Arnold Adolph Berthold (1803–1861).[177] Research on the action of testosterone received a brief boost in 1889, when the Harvard professor Charles-Édouard Brown-Séquard (1817–1894), then in Paris, self-injected subcutaneously a "rejuvenating elixir" consisting of an extract of dog and guinea pig testicle. He reported in The Lancet that his vigor and feeling of well-being were markedly restored but the effects were transient,[178] and Brown-Séquard's hopes for the compound were dashed. Suffering the ridicule of his colleagues, he abandoned his work on the mechanisms and effects of androgens in human beings.

 

In 1927, the University of Chicago's Professor of Physiologic Chemistry, Fred C. Koch, established easy access to a large source of bovine testicles — the Chicago stockyards — and recruited students willing to endure the tedious work of extracting their isolates. In that year, Koch and his student, Lemuel McGee, derived 20 mg of a substance from a supply of 40 pounds of bovine testicles that, when administered to castrated roosters, pigs and rats, remasculinized them.[179] The group of Ernst Laqueur at the University of Amsterdam purified testosterone from bovine testicles in a similar manner in 1934, but isolation of the hormone from animal tissues in amounts permitting serious study in humans was not feasible until three European pharmaceutical giants—Schering (Berlin, Germany), Organon (Oss, Netherlands) and Ciba (Basel, Switzerland)—began full-scale steroid research and development programs in the 1930s.

  

Nobel Prize winner, Leopold Ruzicka of Ciba, a pharmaceutical industry giant that synthesized testosterone.

The Organon group in the Netherlands were the first to isolate the hormone, identified in a May 1935 paper "On Crystalline Male Hormone from Testicles (Testosterone)".[180] They named the hormone testosterone, from the stems of testicle and sterol, and the suffix of ketone. The structure was worked out by Schering's Adolf Butenandt, at the Chemisches Institut of Technical University in Gdańsk.[181][182]

 

The chemical synthesis of testosterone from cholesterol was achieved in August that year by Butenandt and Hanisch.[183] Only a week later, the Ciba group in Zurich, Leopold Ruzicka (1887–1976) and A. Wettstein, published their synthesis of testosterone.[184] These independent partial syntheses of testosterone from a cholesterol base earned both Butenandt and Ruzicka the joint 1939 Nobel Prize in Chemistry.[182][185] Testosterone was identified as 17β-hydroxyandrost-4-en-3-one (C19H28O2), a solid polycyclic alcohol with a hydroxyl group at the 17th carbon atom. This also made it obvious that additional modifications on the synthesized testosterone could be made, i.e., esterification and alkylation.

 

The partial synthesis in the 1930s of abundant, potent testosterone esters permitted the characterization of the hormone's effects, so that Kochakian and Murlin (1936) were able to show that testosterone raised nitrogen retention (a mechanism central to anabolism) in the dog, after which Allan Kenyon's group[186] was able to demonstrate both anabolic and androgenic effects of testosterone propionate in eunuchoidal men, boys, and women. The period of the early 1930s to the 1950s has been called "The Golden Age of Steroid Chemistry",[187] and work during this period progressed quickly. Research in this golden age proved that this newly synthesized compound—testosterone—or rather family of compounds (for many derivatives were developed from 1940 to 1960), was a potent multiplier of muscle, strength, and well-being.[188]

 

Other animals[edit]

Testosterone is observed in most vertebrates. Testosterone and the classical nuclear androgen receptor first appeared in gnathostomes (jawed vertebrates).[189] Agnathans (jawless vertebrates) such as lampreys do not produce testosterone but instead use androstenedione as a male sex hormone.[190] Fish make a slightly different form called 11-ketotestosterone.[191] Its counterpart in insects is ecdysone.[192] The presence of these ubiquitous steroids in a wide range of animals suggest that sex hormones have an ancient evolutionary history.[193]

 

en.wikipedia.org/wiki/Testosterone

Found the cute little gourd-bud suggestive, and the leaf in the background reminded of the proverbial (fig) leaf ..

Thank you very much for visiting, your faves and comments. Cheers.

 

Echidnas /ɨˈkɪdnə/, sometimes known as spiny anteaters, belong to the family Tachyglossidae in the monotreme order of egg-laying mammals. The four extant species, together with the platypus, are the only surviving members of that order and are the only extant mammals that lay eggs. Their diet consists of ants and termites, but they are not closely related to the true anteaters of the Americas. They live in Australia and New Guinea.

Description: Echidnas are small, solitary mammals covered with coarse hair and spines. Superficially, they resemble the anteaters of South America and other spiny mammals such as hedgehogs and porcupines. They are usually black or brown in colour. There have been several reports of 'albino' echidnas, their eyes pink and their spines white. They have elongated and slender snouts that function as both mouth and nose. Like the platypus, they are equipped with electrosensors, but while the platypus has 40,000 electroreceptors on its bill, the long-billed echidna has only 2,000, and the short-billed echidna, which lives in a drier environment, has no more than 400 located at the tip of its snout. They have very short, strong limbs with large claws, and are powerful diggers. Echidnas have tiny mouths and toothless jaws. The echidna feeds by tearing open soft logs, anthills and the like, and using its long, sticky tongue, which protrudes from its snout, to collect prey.

Diet: The short-beaked echidna's diet consists largely of ants and termites, while the Zaglossus species typically eats worms and insect larvae. They have no teeth, and break down their food by grinding it between the bottoms of their mouths and their tongues. The echidnas' ears are slits on the sides of their heads that are usually unseen due to the fact that they are blanketed by their spines. The external ear is called the "pinna", which is created by a large cartilaginous funnel, deep in the muscle.

Habitat: Echidnas do not enjoy extreme temperatures, and use caves and rock crevasses to hide from harsh weather conditions. Echidnas can be found in forests, woodlands, snuggled under vegetation, roots or piles of debris. They sometimes hide in other animal burrows, such as rabbits and wombats. Echidnas have large territories causing their areas to overlap. In addition, echidnas are decent swimmers. When swimming, echidnas expose their snout and some of their spines. They are known to journey their way to water in order to groom and bathe themselves. Long-beaked echidnas have sharp, tiny spines on their tongues that help capture their prey. The echidnas feces are 7 centimeters long and are cylindrical in shape; they are usually broken and unrounded. When eating they consume large amounts of dirt and ant hill material, which makes up majority of their feces.

Classification: Echidnas and the platypus are the only egg-laying mammals, known as monotremes. The neocortex makes up half of the echidna's brain, compared to 80% of a human brain. Due to their low metabolism and accompanying stress resistance, echidnas are long-lived for their size; the longest recorded lifespan for a captive echidna is 50 years, with anecdotal accounts of wild individuals reaching 45 years. Contrary to previous research, the echidna does enter REM sleep, but only when the ambient temperature is around 25 °C (77 °F). At temperatures of 15 °C (59 °F) and 28 °C (~82 °F), REM sleep is suppressed. The average lifespan of an echidna in the wild is estimated around 16 years. When fully grown a female can weigh up to 4.5 kilograms and a male can weigh up to 6 kilograms. You can determine the echidnas' sex by their size; males are 25% larger than females. As well, the reproductive organs differ even though both sexes have a single opening they use to urinate, release their faeces and use to mate. Male echidnas have non-venomous spurs on the hind feet.

Reproduction: The female lays a single soft-shelled, leathery egg 22 days after mating, and deposits it directly into her pouch. Hatching takes place after 10 days; the young echidna then sucks milk from the pores of the two milk patches (monotremes have no nipples) and remains in the pouch for 45 to 55 days, at which time it starts to develop spines. The mother digs a nursery burrow and deposits the young, returning every five days to suckle it until it is weaned at seven months. Male echidnas have a four-headed penis. During mating, the heads on one side "shut down" and do not grow in size; the other two are used to release semen into the female's two-branched reproductive tract. Each time it has sex, it alternates heads in sets of two. When not in use, the penis is retracted inside a preputial sac in the cloaca. The male echidna's penis is 7 centimeters long when erect, and its shaft is covered with penile spines. These may be used to induce ovulation in the female. It is a challenge to study the echidna in their natural habitat and they show no interest in mating while in captivity. Therefore, no one has ever seen an echidna ejaculate. There have been previous attempts, trying to force the echidna to ejaculate through the use of electrically stimulated ejaculation in order to obtain semen samples but has only resulted in the penis swelling. Breeding season begins in late June and extends through September. Males will form lines up to ten individuals long, the youngest echidna trailing last, that follow the female and attempt to mate. During a mating season an echidna may switch between lines. This is known as the "train" system. Two weeks after mating, a single fertilized egg, weighing 380 milligrams and being about 1.4 centimeters long, is implanted in a rear-facing pouch that has developed on the female, where it is held for ten days before hatching. The young echidna, called a puggle, is then held in the pouch for two to three months before being expelled. Puggles will stay within their mother's den for up to a year before leaving.

(Source: Wikipedia)

 

© Chris Burns 2014

__________________________________________

 

All rights reserved.

This image may not be copied, reproduced, distributed, republished, downloaded, displayed, posted or transmitted in any form or by any means, including electronic, mechanical, photocopying and recording without my written consent.

In the almost thirty years since I took this picture in June 1989, the

stylisation of penile graffiti has increased so much that this seems

positively draughtsman-like.

 

I took the picture because I had not seen penises in local graffiti

before. Nowadays, they show up everywhere. After almost three

decades, I still don't understand why.

 

Plus-X in Bessa (6x9 cm).

How Do Penis Enlargers Function? The cavernosum 's what virtually just about every penis enlarger targets to varying degrees, even surgery. Lots of prospective shoppers of penile elongation devices and techniques wish to know what the specific product impacts; well, this really is it. Where

www.rebelmouse.com/jgncemburu/how-penis-pump-work-1440530...

Accelerate & Super Charge Your Results Almost Overnight With A Brand-New "10 Second Ritual" That Supports Enhanced Energy, Vitality and Rock-Hard Sexual Power!

 

click here to know more

 

bit.ly/3v9u8yT

   

pearlypenilepapulestube.com/pearly-penile-papules-removal...

pearlypenilepapulestube.com/natural-and-effective-pearly-...

A stallion is a male horse that has not been gelded (castrated). Stallions follow the conformation and phenotype of their breed, but within that standard, the presence of hormones such as testosterone may give stallions a thicker, "cresty" neck, as well as a somewhat more muscular physique as compared to female horses, known as mares, and castrated males, called geldings.

 

Temperament varies widely based on genetics, and training, but because of their instincts as herd animals, they may be prone to aggressive behavior, particularly toward other stallions, and thus require careful management by knowledgeable handlers. With proper training and management, stallions are effective equine athletes at the highest levels of many disciplines, including horse racing, horse shows, and international Olympic competition.

 

"Stallion" is also used to refer to males of other equids, including zebras and donkeys.

 

Herd behavior

Fillies usually soon join a different band with a dominant stallion different from the one that sired them. Colts or young stallions without mares of their own usually form small, all-male, "bachelor bands" in the wild. Living in a group gives these stallions the social and protective benefits of living in a herd. A bachelor herd may also contain older stallions who have lost their herd in a challenge.

 

Reproductive anatomy

The external genitalia comprise:

 

the testes, which are suspended horizontally within the scrotum. The testes of an average stallion are ovoids 8 to 12 cm (3.1 to 4.7 in) long, 6 to 7 cm (2.4 to 2.8 in) high by 5 cm (2.0 in) wide;

 

the penis, within the "penile sheath". Stallions have a vascular penis. When non-erect, it is quite flaccid and contained within the sheath. The retractor penis muscle is relatively underdeveloped. Erection and protrusion take place gradually, by the increasing tumescence of the erectile vascular tissue in the corpus cavernosum penis. When not erect, the penis is housed within the prepuce, 50 cm (20 in) long and 2.5 to 6 cm (0.98 to 2.36 in) in diameter with the distal end 15 to 20 cm (5.9 to 7.9 in). The retractor muscle contracts to retract the penis into the sheath and relaxes to allow the penis to extend from the sheath. When erect, the penis doubles in length and thickness and the glans increases by three to four times. The urethra opens within the urethral fossa, a small pouch at the distal end of the glans. A structure called the urethral process projects beyond the glans.

 

The internal genitalia comprise the accessory sex glands, which include the vesicular glands, the prostate gland and the bulbourethral glands. These contribute fluid to the semen at ejaculation, but are not strictly necessary for fertility.

 

Management and handling of domesticated stallions

Domesticated stallions are trained and managed in a variety of ways, depending on the region of the world, the owner's philosophy, and the individual stallion's temperament. In all cases, however, stallions have an inborn tendency to attempt to dominate both other horses and human handlers, and will be affected to some degree by proximity to other horses, especially mares in heat. They must be trained to behave with respect toward humans at all times or else their natural aggressiveness, particularly a tendency to bite, may pose a danger of serious injury.

 

The advantage of natural types of management is that the stallion is allowed to behave "like a horse" and may exhibit fewer stable vices. In a harem model, the mares may "cycle" or achieve estrus more readily. Proponents of natural management also assert that mares are more likely to "settle" (become pregnant) in a natural herd setting. Some stallion managers keep a stallion with a mare herd year-round, others will only turn a stallion out with mares during the breeding season.

 

In some places, young domesticated stallions are allowed to live separately in a "bachelor herd" while growing up, kept out of sight, sound or smell of mares. A Swiss study demonstrated that even mature breeding stallions kept well away from other horses could live peacefully together in a herd setting if proper precautions were taken while the initial herd hierarchy was established.

 

As an example, in the New Forest, England, breeding stallions run out on the open Forest for about two to three months each year with the mares and youngstock. On being taken off the Forest, many of them stay together in bachelor herds for most of the rest of the year. New Forest stallions, when not in their breeding work, take part on the annual round-ups, working alongside mares and geldings, and compete successfully in many disciplines.

 

There are drawbacks to natural management, however. One is that the breeding date, and hence foaling date, of any given mare will be uncertain. Another problem is the risk of injury to the stallion or mare in the process of natural breeding, or the risk of injury while a hierarchy is established within an all-male herd. Some stallions become very anxious or temperamental in a herd setting and may lose considerable weight, sometimes to the point of a health risk. Some may become highly protective of their mares and thus more aggressive and dangerous to handle. There is also a greater risk that the stallion may escape from a pasture or be stolen. Stallions may break down fences between adjoining fields to fight another stallion or mate with the "wrong" herd of mares, thus putting the pedigree of ensuing foals in question.

 

Complete isolation has significant drawbacks; stallions may develop additional behavior problems with aggression due to frustration and pent-up energy. As a general rule, a stallion that has been isolated from the time of weaning or sexual maturity will have a more difficult time adapting to a herd environment than one allowed to live close to other animals. As horses are instinctively social creatures, even stallions are believed to benefit from being allowed social interaction with other horses, though proper management and cautions are needed.

 

Properly trained stallions can live and work close to mares and to one another. Examples include the Lipizzan stallions of the Spanish Riding School in Vienna, Austria, where the entire group of stallions live part-time in a bachelor herd as young colts, then are stabled, train, perform, and travel worldwide as adults with few if any management problems. Even stallions who are unfamiliar with each other can work safely in reasonable proximity if properly trained; the vast majority of Thoroughbred horses on the racetrack are stallions, as are many equine athletes in other forms of competition. Stallions are often shown together in the same ring at horse shows, particularly in halter classes where their conformation is evaluated. In horse show performance competition, stallions and mares often compete in the same arena with one another, particularly in Western and English "pleasure"-type classes where horses are worked as a group. Overall, stallions can be trained to keep focused on work and may be brilliant performers if properly handled.

 

A breeding stallion is more apt to present challenging behavior to a human handler than one who has not bred mares, and stallions may be more difficult to handle in spring and summer, during the breeding season, than during the fall and winter. Some stallions are used for both equestrian uses and for breeding at the same general time of year. Though compromises may need to be made in expectations for both athletic performance and fertility rate, well-trained stallions with good temperaments can be taught that breeding behavior is only allowed in a certain area, or with certain cues, equipment, or with a particular handler.

 

Geldings

If a stallion is not to be used for breeding, castrating (gelding) the male horse will allow it to live full-time in a herd with both males and females, reduce aggressive or disruptive behavior, and allow the horse to be around other animals without being seriously distracted. If a horse is not to be used for breeding, it can be gelded prior to reaching sexual maturity. A horse gelded young may grow taller and behave better if this is done. Older stallions that are sterile or otherwise no longer used for breeding may also be gelded and will exhibit calmer behavior, even if previously used for breeding. However, they are more likely to continue stallion-like behaviors than horses gelded at a younger age, especially if they have been used as a breeding stallion. Modern surgical techniques allow castration to be performed on a horse of almost any age with relatively few risks.

 

In most cases, particularly in modern industrialized cultures, a male horse that is not of sufficient quality to be used for breeding will have a happier life without having to deal with the instinctive, hormone-driven behaviors that come with being left intact. Geldings are safer to handle and present fewer management problems.

 

Some in the animal rights community maintains that castration is mutilation and damaging to the animal's psyche.

 

Ridglings

A ridgling or "rig" is a cryptorchid, a stallion which has one or both testicles undescended. If both testicles are not descended, the horse may appear to be a gelding, but will still behave like a stallion. A gelding that displays stallion-like behaviors is sometimes called a "false rig". In many cases, ridglings are infertile, or have fertility levels that are significantly reduced. The condition is most easily corrected by gelding the horse. A more complex and costly surgical procedure can sometimes correct the condition and restore the animal's fertility, though it is only cost-effective for a horse that has very high potential as a breeding stallion. This surgery generally removes the non-descended testicle, leaving the descended testicle, and creating a horse known as a monorchid stallion. Keeping cryptorchids or surgically-created monorchids as breeding stallions is controversial, as the condition is at least partially genetic and some handlers claim that cryptorchids tend to have greater levels of behavioral problems than normal stallions.

pearlypenilepapulestube.com/pearly-penile-papules-removal...

Sloane Square & Saatchi

Designed to be used in the bath or shower, the Bathmate HYDROMAX7 WIDE BOY Penis Pump utilises the impressive power of water therapy to help enhance erections. With 35% more power than the already-legendary HYDRO7, this is one routine step you won’t want to skip.

 

As a safe, penile rigidity device, the Bathmate HYDROMAX7 WIDE BOY may help to achieve harder erections for longer, over time.

 

Bathmate HYDROMAX7 WIDE BOY Penis Pump

pearlypenilepapulestube.com/pearly-penile-papules-removal...

Pearly penile papules (bumps on penile shaft) are tiny bumps that ring the rim of the penis head. They are typically flesh-colored or a little lighter and have a smooth dome shape. Interestingly, they also seem to form in organized rows, like a string of pearls. There are no harmful effects other than some possible sensitivity that may be uncomfortable during sex or other activities.

  

pearlypenilepapulestube.com/how-to-remove-pearly-penile-p...

this art is called 'chabi bharna' where he rolls up his penis to a stick and swirls the stick around to show his penile prowess ;)

Cock and ball torture (CBT), penis torture or dick torture is a sexual activity involving application of pain or constriction to the penis or testicles. This may involve directly painful activities, such as genital piercing, wax play, genital spanking, squeezing, ball-busting, genital flogging, urethral play, tickle torture, erotic electrostimulation, kneeing or kicking.[1] The recipient of such activities may receive direct physical pleasure via masochism, or emotional pleasure through erotic humiliation, or knowledge that the play is pleasing to a sadistic dominant. Many of these practices carry significant health risks.[2]

 

Contents

Devices and practices

  

Top left: Erotic electrostimulation. Top right: Trampling the penis. Bottom left: Wax play. Bottom right: Chastity piercing.

Similar to many other sexual activities, CBT can be performed using toys and devices to make the penis and testicles more easily accessible for attack, or for foreplay purposes.[3][4]

 

Ball stretcher

 

A metal ball stretcher and cock ring, which forces penis' erection.

A ball stretcher is a sex toy that is used to elongate the scrotum and provide a feeling of weight pulling the testicles away from the body. This can be particularly enjoyable for the wearer as it can make an orgasm more intense, as testicles are prevented from moving up. Intended to make one's testicles permanently hang much lower than before (if used regularly for extended periods of time), this sex toy can be potentially harmful to the male genitals as the circulation of blood can be easily cut off if over-tightened.

 

While leather stretchers are most common, other models consist of an assortment of steel rings that fastens with screws, causing additional but only mildly uncomfortable weight to the wearer's testicles. The length of the stretcher may vary from 1-4 inches.[5] A more dangerous type of ball stretcher can be home-made simply by wrapping rope or string around one's scrotum until it is eventually stretched to the desired length.

 

Ball crusher

A ball crusher is a device made from either metal or often clear acrylic that squeezes the testicles slowly by turning a nut or screw. How tight it is clamped depends on the pain tolerance of the person it is used on. A ball crusher is often combined with bondage, either with a partner or by oneself.

 

Parachute

A parachute is a small collar, usually made from leather, which fastens around the scrotum, and from which weights can be hung. It is conical in shape, with three or four short chains hanging beneath, to which weights can be attached.

 

Used as part of cock and ball torture within a BDSM relationship, the parachute provides a constant drag, and a squeezing effect on the testicles. Moderate weights of 3–5 kg can be suspended, especially during bondage, though occasionally much heavier weights are used. Smaller weights can be used when the male wearing it is free to move; the swinging effect of the weight can restrict sudden movements, as well as providing a visual stimulus for the dominant partner.[citation needed]

 

Humbler

 

Man with humbler

A humbler is a BDSM physical restraint device used to restrict the movement of a submissive male participant in a BDSM scene.

 

The humbler consists of a testicle cuff device that clamps around the base of the scrotum, mounted in the center of a bar that passes behind the thighs at the base of the buttocks. This forces the wearer to keep his legs folded forward, as any attempt to straighten the legs even slightly pulls hard on the scrotum, causing anything from considerable discomfort to extreme pain.

 

Testicle cuffs

A testicle cuff is a ring-shaped device around the scrotum between the body and the testicles which when closed does not allow the testicles to slide through it. A common type has two connected cuffs, one around the scrotum and the other around the base of the penis. They are just one of many devices to restrain the male genitalia. A standard padlock, which cannot be removed without its key, may also be locked around the scrotum.

 

Some passive men enjoy the feeling of being "owned", while dominant individuals enjoy the sense of "owning" their partners. Requiring such a man wear testicle cuffs symbolizes that his sexual organs belong to his partner, who may be either male or female. There is a level of humiliation involved, by which they find sexual arousal. The cuffs may even form part of a sexual fetish of the wearer or his partner.[citation needed]

 

However, these are extreme uses of testicle cuffs. More conventionally, the device pulls down the testicles and keeps them there during stimulation, which has a number of benefits:

 

Making the penis appear longer. Pulling the testicles down and away from the base of the penis stretches the skin over the base of the penis and pubic bone, exposing the additional inch or so of penile shaft that is normally hidden from view.

Improving sexual arousal. While some men may be aroused by the feeling of being "owned", the physical feeling of stretching the ligaments that suspend the testicles has an effect similar to the more common practice of stretching one's legs and pointing the toes.

Preventing the testicles from lifting up so far that they become lodged under the skin immediately adjacent to the base of the penis, a condition which can be very uncomfortable, especially if the testicle is then squashed by the slap of skin during thrusting in sexual intercourse.

Delaying or intensifying ejaculation by preventing the testicles from rising normally to the "point of no return". It is much harder to reach an orgasm.

Cock harness

A cock harness is a penile sex toy designed to be worn around the penis and scrotum. Its function is similar to that of a cock ring. These devices are often associated with BDSM activities. The Gates of Hell is a male chastity device made up of multiple cock rings that can be used for CBT.[6] Kali's Teeth is a metal bracelet with interior spikes that closes around the penis and can be used for preventing or punishing erections.[7]

 

Ball busting

 

Ball busting: A woman kicking male submissive's testicles at Folsom Street Fair, USA

"Ball busting" is the practice of kicking or kneeing men in the testicles. It carries several medical risks, including the danger of testicular rupture from blunt trauma.

 

CBT in Japan

 

Tamakeri (玉蹴り) (lit. ball kicking) is a sexual fetish and subgenre of BDSM within which a man's testicles are abused. The genre is also referred to as ballbusting ("bb" for short).[8][9] Tamakeri is the Japanese term, but it is used by many non-Japanese people to describe media where Asian people—mainly women—are participating in it. The dynamics of tamakeri consist of a masochist having their testicles hurt by a sadist.[10] The fetish is popular among heterosexual and homosexual men and women.[11]

 

Denkianma (電気按摩) (lit. "electric massage") is a popular Japanese prank played between two people where one person puts their foot into the genital area of the other and shakes it in a vibrating motion.[12][13] Often this is done by grabbing the other person's feet, raising them, and then placing one's own foot on their crotch and vibrating it. This is often done between school aged boys as a prank similar to kancho and could be seen by a western audience as a type of bullying. In 2006 the crisps company Doritos released a new flavour of potato chip called "Denkianma".[14]

 

Safety

 

This section needs attention from an expert in health. The specific problem is: concerns medical topics, but lacks appropriate citations.

Loss of blood flow is one of the biggest risks in cock and ball torture, which can be seen with loss of color and edemas.[15] Bondage in which the testicles are tied to something else is especially dangerous, increasing the risk of the testicles getting damaged through excessive tension or pulling.

 

The most serious injuries are testicular rupture, testicular torsion and testicular avulsion, which are medical emergencies that require urgent medical attention.[16]

Lumps on penile sheath of black & white horse. We were concerned they were haemangiosarcomas.

The cat (Felis catus), commonly referred to as the domestic cat or house cat, is the only domesticated species in the family Felidae. Recent advances in archaeology and genetics have shown that the domestication of the cat occurred in the Near East around 7500 BC. It is commonly kept as a house pet and farm cat, but also ranges freely as a feral cat avoiding human contact. It is valued by humans for companionship and its ability to kill vermin. Because of its retractable claws it is adapted to killing small prey like mice and rats. It has a strong flexible body, quick reflexes, sharp teeth, and its night vision and sense of smell are well developed. It is a social species, but a solitary hunter and a crepuscular predator. Cat communication includes vocalizations like meowing, purring, trilling, hissing, growling, and grunting as well as cat body language. It can hear sounds too faint or too high in frequency for human ears, such as those made by small mammals. It also secretes and perceives pheromones.

 

Female domestic cats can have kittens from spring to late autumn in temperate zones and throughout the year in equatorial regions, with litter sizes often ranging from two to five kittens. Domestic cats are bred and shown at events as registered pedigreed cats, a hobby known as cat fancy. Animal population control of cats may be achieved by spaying and neutering, but their proliferation and the abandonment of pets has resulted in large numbers of feral cats worldwide, contributing to the extinction of bird, mammal and reptile species.

 

As of 2017, the domestic cat was the second most popular pet in the United States, with 95.6 million cats owned and around 42 million households owning at least one cat. In the United Kingdom, 26% of adults have a cat, with an estimated population of 10.9 million pet cats as of 2020. As of 2021, there were an estimated 220 million owned and 480 million stray cats in the world.

 

Etymology and naming

The origin of the English word cat, Old English catt, is thought to be the Late Latin word cattus, which was first used at the beginning of the 6th century. The Late Latin word may be derived from an unidentified African language. The Nubian word kaddîska 'wildcat' and Nobiin kadīs are possible sources or cognates. The Nubian word may be a loan from Arabic قَطّ‎ qaṭṭ ~ قِطّ qiṭṭ.

 

The forms might also have derived from an ancient Germanic word that was imported into Latin and then into Greek, Syriac, and Arabic. The word may be derived from Germanic and Northern European languages, and ultimately be borrowed from Uralic, cf. Northern Sámi gáđfi, 'female stoat', and Hungarian hölgy, 'lady, female stoat'; from Proto-Uralic *käďwä, 'female (of a furred animal)'.

 

The English puss, extended as pussy and pussycat, is attested from the 16th century and may have been introduced from Dutch poes or from Low German puuskatte, related to Swedish kattepus, or Norwegian pus, pusekatt. Similar forms exist in Lithuanian puižė and Irish puisín or puiscín. The etymology of this word is unknown, but it may have arisen from a sound used to attract a cat.

 

A male cat is called a tom or tomcat (or a gib, if neutered). A female is called a queen or a molly, if spayed, especially in a cat-breeding context. A juvenile cat is referred to as a kitten. In Early Modern English, the word kitten was interchangeable with the now-obsolete word catling.

 

A group of cats can be referred to as a clowder or a glaring.

 

Taxonomy

The scientific name Felis catus was proposed by Carl Linnaeus in 1758 for a domestic cat. Felis catus domesticus was proposed by Johann Christian Polycarp Erxleben in 1777. Felis daemon proposed by Konstantin Satunin in 1904 was a black cat from the Transcaucasus, later identified as a domestic cat.

 

In 2003, the International Commission on Zoological Nomenclature ruled that the domestic cat is a distinct species, namely Felis catus. In 2007, it was considered a subspecies, F. silvestris catus, of the European wildcat (F. silvestris) following results of phylogenetic research. In 2017, the IUCN Cat Classification Taskforce followed the recommendation of the ICZN in regarding the domestic cat as a distinct species, Felis catus.

 

Evolution

Main article: Cat evolution

The domestic cat is a member of the Felidae, a family that had a common ancestor about 10 to 15 million years ago. The evolutionary radiation of the Felidae began in Asia during the Miocene around 8.38 to 14.45 million years ago. Analysis of mitochondrial DNA of all Felidae species indicates a radiation at 6.46 to 16.76 million years ago. The genus Felis genetically diverged from other Felidae around 6 to 7 million years ago. Results of phylogenetic research shows that the wild members of this genus evolved through sympatric or parapatric speciation, whereas the domestic cat evolved through artificial selection. The domestic cat and its closest wild ancestor are diploid and both possess 38 chromosomes and roughly 20,000 genes.

 

Domestication

See also: Domestication of the cat and Cats in ancient Egypt

It was long thought that the domestication of the cat began in ancient Egypt, where cats were venerated from around 3100 BC, However, the earliest known indication for the taming of an African wildcat was excavated close by a human Neolithic grave in Shillourokambos, southern Cyprus, dating to about 7500–7200 BC. Since there is no evidence of native mammalian fauna on Cyprus, the inhabitants of this Neolithic village most likely brought the cat and other wild mammals to the island from the Middle Eastern mainland. Scientists therefore assume that African wildcats were attracted to early human settlements in the Fertile Crescent by rodents, in particular the house mouse (Mus musculus), and were tamed by Neolithic farmers. This mutual relationship between early farmers and tamed cats lasted thousands of years. As agricultural practices spread, so did tame and domesticated cats. Wildcats of Egypt contributed to the maternal gene pool of the domestic cat at a later time.

The earliest known evidence for the occurrence of the domestic cat in Greece dates to around 1200 BC. Greek, Phoenician, Carthaginian and Etruscan traders introduced domestic cats to southern Europe. During the Roman Empire they were introduced to Corsica and Sardinia before the beginning of the 1st millennium. By the 5th century BC, they were familiar animals around settlements in Magna Graecia and Etruria. By the end of the Western Roman Empire in the 5th century, the Egyptian domestic cat lineage had arrived in a Baltic Sea port in northern Germany.

 

The leopard cat (Prionailurus bengalensis) was tamed independently in China around 5500 BC. This line of partially domesticated cats leaves no trace in the domestic cat populations of today.

 

During domestication, cats have undergone only minor changes in anatomy and behavior, and they are still capable of surviving in the wild. Several natural behaviors and characteristics of wildcats may have pre-adapted them for domestication as pets. These traits include their small size, social nature, obvious body language, love of play, and high intelligence. Captive Leopardus cats may also display affectionate behavior toward humans but were not domesticated. House cats often mate with feral cats. Hybridisation between domestic and other Felinae species is also possible, producing hybrids such as the Kellas cat in Scotland.

 

Development of cat breeds started in the mid 19th century. An analysis of the domestic cat genome revealed that the ancestral wildcat genome was significantly altered in the process of domestication, as specific mutations were selected to develop cat breeds. Most breeds are founded on random-bred domestic cats. Genetic diversity of these breeds varies between regions, and is lowest in purebred populations, which show more than 20 deleterious genetic disorders.

 

Characteristics

Main article: Cat anatomy

Size

The domestic cat has a smaller skull and shorter bones than the European wildcat. It averages about 46 cm (18 in) in head-to-body length and 23–25 cm (9.1–9.8 in) in height, with about 30 cm (12 in) long tails. Males are larger than females. Adult domestic cats typically weigh 4–5 kg (8.8–11.0 lb).

 

Skeleton

Cats have seven cervical vertebrae (as do most mammals); 13 thoracic vertebrae (humans have 12); seven lumbar vertebrae (humans have five); three sacral vertebrae (as do most mammals, but humans have five); and a variable number of caudal vertebrae in the tail (humans have only three to five vestigial caudal vertebrae, fused into an internal coccyx).  The extra lumbar and thoracic vertebrae account for the cat's spinal mobility and flexibility. Attached to the spine are 13 ribs, the shoulder, and the pelvis.  Unlike human arms, cat forelimbs are attached to the shoulder by free-floating clavicle bones which allow them to pass their body through any space into which they can fit their head.

 

Skull

The cat skull is unusual among mammals in having very large eye sockets and a powerful specialized jaw.  Within the jaw, cats have teeth adapted for killing prey and tearing meat. When it overpowers its prey, a cat delivers a lethal neck bite with its two long canine teeth, inserting them between two of the prey's vertebrae and severing its spinal cord, causing irreversible paralysis and death. Compared to other felines, domestic cats have narrowly spaced canine teeth relative to the size of their jaw, which is an adaptation to their preferred prey of small rodents, which have small vertebrae.

 

The premolar and first molar together compose the carnassial pair on each side of the mouth, which efficiently shears meat into small pieces, like a pair of scissors. These are vital in feeding, since cats' small molars cannot chew food effectively, and cats are largely incapable of mastication.:  Cats tend to have better teeth than most humans, with decay generally less likely because of a thicker protective layer of enamel, a less damaging saliva, less retention of food particles between teeth, and a diet mostly devoid of sugar. Nonetheless, they are subject to occasional tooth loss and infection.

 

Claws

Cats have protractible and retractable claws. In their normal, relaxed position, the claws are sheathed with the skin and fur around the paw's toe pads. This keeps the claws sharp by preventing wear from contact with the ground and allows for the silent stalking of prey. The claws on the forefeet are typically sharper than those on the hindfeet. Cats can voluntarily extend their claws on one or more paws. They may extend their claws in hunting or self-defense, climbing, kneading, or for extra traction on soft surfaces. Cats shed the outside layer of their claw sheaths when scratching rough surfaces.

 

Most cats have five claws on their front paws and four on their rear paws. The dewclaw is proximal to the other claws. More proximally is a protrusion which appears to be a sixth "finger". This special feature of the front paws on the inside of the wrists has no function in normal walking but is thought to be an antiskidding device used while jumping. Some cat breeds are prone to having extra digits ("polydactyly"). Polydactylous cats occur along North America's northeast coast and in Great Britain.

 

Ambulation

The cat is digitigrade. It walks on the toes, with the bones of the feet making up the lower part of the visible leg. Unlike most mammals, it uses a "pacing" gait and moves both legs on one side of the body before the legs on the other side. It registers directly by placing each hind paw close to the track of the corresponding fore paw, minimizing noise and visible tracks. This also provides sure footing for hind paws when navigating rough terrain. As it speeds up from walking to trotting, its gait changes to a "diagonal" gait: The diagonally opposite hind and fore legs move simultaneously.

 

Balance

Cats are generally fond of sitting in high places or perching. A higher place may serve as a concealed site from which to hunt; domestic cats strike prey by pouncing from a perch such as a tree branch. Another possible explanation is that height gives the cat a better observation point, allowing it to survey its territory. A cat falling from heights of up to 3 m (9.8 ft) can right itself and land on its paws.

 

During a fall from a high place, a cat reflexively twists its body and rights itself to land on its feet using its acute sense of balance and flexibility. This reflex is known as the cat righting reflex. A cat always rights itself in the same way during a fall, if it has enough time to do so, which is the case in falls of 90 cm (3.0 ft) or more. How cats are able to right themselves when falling has been investigated as the "falling cat problem".

 

Coats

Main article: Cat coat genetics

The cat family (Felidae) can pass down many colors and patterns to their offspring. The domestic cat genes MC1R and ASIP allow for the variety of color in coats. The feline ASIP gene consists of three coding exons. Three novel microsatellite markers linked to ASIP were isolated from a domestic cat BAC clone containing this gene and were used to perform linkage analysis in a pedigree of 89 domestic cats that segregated for melanism.[citation needed]

 

Senses

Main article: Cat senses

Vision

A cat's nictitating membrane shown as it blinks

Cats have excellent night vision and can see at only one-sixth the light level required for human vision.  This is partly the result of cat eyes having a tapetum lucidum, which reflects any light that passes through the retina back into the eye, thereby increasing the eye's sensitivity to dim light. Large pupils are an adaptation to dim light. The domestic cat has slit pupils, which allow it to focus bright light without chromatic aberration. At low light, a cat's pupils expand to cover most of the exposed surface of its eyes. The domestic cat has rather poor color vision and only two types of cone cells, optimized for sensitivity to blue and yellowish green; its ability to distinguish between red and green is limited. A response to middle wavelengths from a system other than the rod cells might be due to a third type of cone. This appears to be an adaptation to low light levels rather than representing true trichromatic vision. Cats also have a nictitating membrane, allowing them to blink without hindering their vision.

 

Hearing

The domestic cat's hearing is most acute in the range of 500 Hz to 32 kHz. It can detect an extremely broad range of frequencies ranging from 55 Hz to 79 kHz, whereas humans can only detect frequencies between 20 Hz and 20 kHz. It can hear a range of 10.5 octaves, while humans and dogs can hear ranges of about 9 octaves. Its hearing sensitivity is enhanced by its large movable outer ears, the pinnae, which amplify sounds and help detect the location of a noise. It can detect ultrasound, which enables it to detect ultrasonic calls made by rodent prey. Recent research has shown that cats have socio-spatial cognitive abilities to create mental maps of owners' locations based on hearing owners' voices.

 

Smell

Cats have an acute sense of smell, due in part to their well-developed olfactory bulb and a large surface of olfactory mucosa, about 5.8 cm2 (0.90 in2) in area, which is about twice that of humans. Cats and many other animals have a Jacobson's organ in their mouths that is used in the behavioral process of flehmening. It allows them to sense certain aromas in a way that humans cannot. Cats are sensitive to pheromones such as 3-mercapto-3-methylbutan-1-ol, which they use to communicate through urine spraying and marking with scent glands. Many cats also respond strongly to plants that contain nepetalactone, especially catnip, as they can detect that substance at less than one part per billion. About 70–80% of cats are affected by nepetalactone. This response is also produced by other plants, such as silver vine (Actinidia polygama) and the herb valerian; it may be caused by the smell of these plants mimicking a pheromone and stimulating cats' social or sexual behaviors.

 

Taste

Cats have relatively few taste buds compared to humans (470 or so versus more than 9,000 on the human tongue). Domestic and wild cats share a taste receptor gene mutation that keeps their sweet taste buds from binding to sugary molecules, leaving them with no ability to taste sweetness. They, however, possess taste bud receptors specialized for acids, amino acids like protein, and bitter tastes. Their taste buds possess the receptors needed to detect umami. However, these receptors contain molecular changes that make the cat taste of umami different from that of humans. In humans, they detect the amino acids of glutamic acid and aspartic acid, but in cats they instead detect nucleotides, in this case inosine monophosphate and l-Histidine. These nucleotides are particularly enriched in tuna. This has been argued is why cats find tuna so palatable: as put by researchers into cat taste, "the specific combination of the high IMP and free l-Histidine contents of tuna" .. "produces a strong umami taste synergy that is highly preferred by cats". One of the researchers involved in this research has further claimed, "I think umami is as important for cats as sweet is for humans".[87]

 

Cats also have a distinct temperature preference for their food, preferring food with a temperature around 38 °C (100 °F) which is similar to that of a fresh kill; some cats reject cold food (which would signal to the cat that the "prey" item is long dead and therefore possibly toxic or decomposing).

 

Whiskers

To aid with navigation and sensation, cats have dozens of movable whiskers (vibrissae) over their body, especially their faces. These provide information on the width of gaps and on the location of objects in the dark, both by touching objects directly and by sensing air currents; they also trigger protective blink reflexes to protect the eyes from damage.: 47 

 

Behavior

See also: Cat behavior

Outdoor cats are active both day and night, although they tend to be slightly more active at night.[88] Domestic cats spend the majority of their time in the vicinity of their homes but can range many hundreds of meters from this central point. They establish territories that vary considerably in size, in one study ranging 7–28 ha (17–69 acres). The timing of cats' activity is quite flexible and varied but being low-light predators, they are generally crepuscular, which means they tend to be more active near dawn and dusk. However, house cats' behavior is also influenced by human activity and they may adapt to their owners' sleeping patterns to some extent.

 

Cats conserve energy by sleeping more than most animals, especially as they grow older. The daily duration of sleep varies, usually between 12 and 16 hours, with 13 and 14 being the average. Some cats can sleep as much as 20 hours. The term "cat nap" for a short rest refers to the cat's tendency to fall asleep (lightly) for a brief period. While asleep, cats experience short periods of rapid eye movement sleep often accompanied by muscle twitches, which suggests they are dreaming.

 

Sociability

The social behavior of the domestic cat ranges from widely dispersed individuals to feral cat colonies that gather around a food source, based on groups of co-operating females. Within such groups, one cat is usually dominant over the others. Each cat in a colony holds a distinct territory, with sexually active males having the largest territories, which are about 10 times larger than those of female cats and may overlap with several females' territories. These territories are marked by urine spraying, by rubbing objects at head height with secretions from facial glands, and by defecation. Between these territories are neutral areas where cats watch and greet one another without territorial conflicts. Outside these neutral areas, territory holders usually chase away stranger cats, at first by staring, hissing, and growling and, if that does not work, by short but noisy and violent attacks. Despite this colonial organization, cats do not have a social survival strategy or a herd behavior, and always hunt alone.

 

Life in proximity to humans and other domestic animals has led to a symbiotic social adaptation in cats, and cats may express great affection toward humans or other animals. Ethologically, a cat's human keeper functions as if a mother surrogate. Adult cats live their lives in a kind of extended kittenhood, a form of behavioral neoteny. Their high-pitched sounds may mimic the cries of a hungry human infant, making them particularly difficult for humans to ignore. Some pet cats are poorly socialized. In particular, older cats show aggressiveness toward newly arrived kittens, which include biting and scratching; this type of behavior is known as feline asocial aggression.

 

Redirected aggression is a common form of aggression which can occur in multiple cat households. In redirected aggression there is usually something that agitates the cat: this could be a sight, sound, or another source of stimuli which causes a heightened level of anxiety or arousal. If the cat cannot attack the stimuli, it may direct anger elsewhere by attacking or directing aggression to the nearest cat, dog, human or other being.

 

Domestic cats' scent rubbing behavior toward humans or other cats is thought to be a feline means for social bonding.

 

Communication

Main article: Cat communication

Domestic cats use many vocalizations for communication, including purring, trilling, hissing, growling/snarling, grunting, and several different forms of meowing. Their body language, including position of ears and tail, relaxation of the whole body, and kneading of the paws, are all indicators of mood. The tail and ears are particularly important social signal mechanisms in cats. A raised tail indicates a friendly greeting, and flattened ears indicate hostility. Tail-raising also indicates the cat's position in the group's social hierarchy, with dominant individuals raising their tails less often than subordinate ones. Feral cats are generally silent.: 208  Nose-to-nose touching is also a common greeting and may be followed by social grooming, which is solicited by one of the cats raising and tilting its head.

 

Purring may have developed as an evolutionary advantage as a signaling mechanism of reassurance between mother cats and nursing kittens, who are thought to use it as a care-soliciting signal. Post-nursing cats also often purr as a sign of contentment: when being petted, becoming relaxed, or eating. Even though purring is popularly interpreted as indicative of pleasure, it has been recorded in a wide variety of circumstances, most of which involve physical contact between the cat and another, presumably trusted individual. Some cats have been observed to purr continuously when chronically ill or in apparent pain.

 

The exact mechanism by which cats purr has long been elusive, but it has been proposed that purring is generated via a series of sudden build-ups and releases of pressure as the glottis is opened and closed, which causes the vocal folds to separate forcefully. The laryngeal muscles in control of the glottis are thought to be driven by a neural oscillator which generates a cycle of contraction and release every 30–40 milliseconds (giving a frequency of 33 to 25 Hz).

 

Domestic cats observed in a rescue facility have total of 276 distinct facial expressions based on 26 different facial movements; each facial expression corresponds to different social functions that are likely influenced by domestication.

 

Grooming

Cats are known for spending considerable amounts of time licking their coats to keep them clean. The cat's tongue has backward-facing spines about 500 μm long, which are called papillae. These contain keratin which makes them rigid so the papillae act like a hairbrush. Some cats, particularly longhaired cats, occasionally regurgitate hairballs of fur that have collected in their stomachs from grooming. These clumps of fur are usually sausage-shaped and about 2–3 cm (0.79–1.18 in) long. Hairballs can be prevented with remedies that ease elimination of the hair through the gut, as well as regular grooming of the coat with a comb or stiff brush.

 

Fighting

Among domestic cats, males are more likely to fight than females. Among feral cats, the most common reason for cat fighting is competition between two males to mate with a female. In such cases, most fights are won by the heavier male. Another common reason for fighting in domestic cats is the difficulty of establishing territories within a small home. Female cats also fight over territory or to defend their kittens. Neutering will decrease or eliminate this behavior in many cases, suggesting that the behavior is linked to sex hormones.

 

When cats become aggressive, they try to make themselves appear larger and more threatening by raising their fur, arching their backs, turning sideways and hissing or spitting. Often, the ears are pointed down and back to avoid damage to the inner ear and potentially listen for any changes behind them while focused forward. Cats may also vocalize loudly and bare their teeth in an effort to further intimidate their opponents. Fights usually consist of grappling and delivering powerful slaps to the face and body with the forepaws as well as bites. Cats also throw themselves to the ground in a defensive posture to rake their opponent's belly with their powerful hind legs.

 

Serious damage is rare, as the fights are usually short in duration, with the loser running away with little more than a few scratches to the face and ears. Fights for mating rights are typically more severe and injuries may include deep puncture wounds and lacerations. Normally, serious injuries from fighting are limited to infections of scratches and bites, though these can occasionally kill cats if untreated. In addition, bites are probably the main route of transmission of feline immunodeficiency virus. Sexually active males are usually involved in many fights during their lives, and often have decidedly battered faces with obvious scars and cuts to their ears and nose. Cats are willing to threaten animals larger than them to defend their territory, such as dogs and foxes.

 

Hunting and feeding

See also: Cat food

The shape and structure of cats' cheeks is insufficient to allow them to take in liquids using suction. Therefore, when drinking they lap with the tongue to draw liquid upward into their mouths. Lapping at a rate of four times a second, the cat touches the smooth tip of its tongue to the surface of the water, and quickly retracts it like a corkscrew, drawing water upward.

 

Feral cats and free-fed house cats consume several small meals in a day. The frequency and size of meals varies between individuals. They select food based on its temperature, smell and texture; they dislike chilled foods and respond most strongly to moist foods rich in amino acids, which are similar to meat. Cats reject novel flavors (a response termed neophobia) and learn quickly to avoid foods that have tasted unpleasant in the past. It is also a common misconception that cats like milk/cream, as they tend to avoid sweet food and milk. Most adult cats are lactose intolerant; the sugar in milk is not easily digested and may cause soft stools or diarrhea. Some also develop odd eating habits and like to eat or chew on things like wool, plastic, cables, paper, string, aluminum foil, or even coal. This condition, pica, can threaten their health, depending on the amount and toxicity of the items eaten.

 

Cats hunt small prey, primarily birds and rodents, and are often used as a form of pest control. Other common small creatures such as lizards and snakes may also become prey. Cats use two hunting strategies, either stalking prey actively, or waiting in ambush until an animal comes close enough to be captured. The strategy used depends on the prey species in the area, with cats waiting in ambush outside burrows, but tending to actively stalk birds.: 153  Domestic cats are a major predator of wildlife in the United States, killing an estimated 1.3 to 4.0 billion birds and 6.3 to 22.3 billion mammals annually.

 

Certain species appear more susceptible than others; in one English village, for example, 30% of house sparrow mortality was linked to the domestic cat. In the recovery of ringed robins (Erithacus rubecula) and dunnocks (Prunella modularis) in Britain, 31% of deaths were a result of cat predation. In parts of North America, the presence of larger carnivores such as coyotes which prey on cats and other small predators reduces the effect of predation by cats and other small predators such as opossums and raccoons on bird numbers and variety.

 

Perhaps the best-known element of cats' hunting behavior, which is commonly misunderstood and often appalls cat owners because it looks like torture, is that cats often appear to "play" with prey by releasing and recapturing it. This cat and mouse behavior is due to an instinctive imperative to ensure that the prey is weak enough to be killed without endangering the cat.

 

Another poorly understood element of cat hunting behavior is the presentation of prey to human guardians. One explanation is that cats adopt humans into their social group and share excess kill with others in the group according to the dominance hierarchy, in which humans are reacted to as if they are at or near the top. Another explanation is that they attempt to teach their guardians to hunt or to help their human as if feeding "an elderly cat, or an inept kitten". This hypothesis is inconsistent with the fact that male cats also bring home prey, despite males having negligible involvement in raising kittens.:

 

Play

Main article: Cat play and toys

Domestic cats, especially young kittens, are known for their love of play. This behavior mimics hunting and is important in helping kittens learn to stalk, capture, and kill prey. Cats also engage in play fighting, with each other and with humans. This behavior may be a way for cats to practice the skills needed for real combat, and might also reduce any fear they associate with launching attacks on other animals.

 

Cats also tend to play with toys more when they are hungry. Owing to the close similarity between play and hunting, cats prefer to play with objects that resemble prey, such as small furry toys that move rapidly, but rapidly lose interest. They become habituated to a toy they have played with before. String is often used as a toy, but if it is eaten, it can become caught at the base of the cat's tongue and then move into the intestines, a medical emergency which can cause serious illness, even death. Owing to the risks posed by cats eating string, it is sometimes replaced with a laser pointer's dot, which cats may chase.

 

Reproduction

See also: Kitten

The cat secretes and perceives pheromones. Female cats, called queens, are polyestrous with several estrus cycles during a year, lasting usually 21 days. They are usually ready to mate between early February and August in northern temperate zones and throughout the year in equatorial regions.

 

Several males, called tomcats, are attracted to a female in heat. They fight over her, and the victor wins the right to mate. At first, the female rejects the male, but eventually, the female allows the male to mate. The female utters a loud yowl as the male pulls out of her because a male cat's penis has a band of about 120–150 backward-pointing penile spines, which are about 1 mm (0.039 in) long; upon withdrawal of the penis, the spines may provide the female with increased sexual stimulation, which acts to induce ovulation.

 

After mating, the female cleans her vulva thoroughly. If a male attempts to mate with her at this point, the female attacks him. After about 20 to 30 minutes, once the female is finished grooming, the cycle will repeat. Because ovulation is not always triggered by a single mating, females may not be impregnated by the first male with which they mate. Furthermore, cats are superfecund; that is, a female may mate with more than one male when she is in heat, with the result that different kittens in a litter may have different fathers.

 

The morula forms 124 hours after conception. At 148 hours, early blastocysts form. At 10–12 days, implantation occurs. The gestation of queens lasts between 64 and 67 days, with an average of 65 days.

 

Data on the reproductive capacity of more than 2,300 free-ranging queens were collected during a study between May 1998 and October 2000. They had one to six kittens per litter, with an average of three kittens. They produced a mean of 1.4 litters per year, but a maximum of three litters in a year. Of 169 kittens, 127 died before they were six months old due to a trauma caused in most cases by dog attacks and road accidents. The first litter is usually smaller than subsequent litters. Kittens are weaned between six and seven weeks of age. Queens normally reach sexual maturity at 5–10 months, and males at 5–7 months. This varies depending on breed. Kittens reach puberty at the age of 9–10 months.

 

Cats are ready to go to new homes at about 12 weeks of age, when they are ready to leave their mother. They can be surgically sterilized (spayed or castrated) as early as seven weeks to limit unwanted reproduction. This surgery also prevents undesirable sex-related behavior, such as aggression, territory marking (spraying urine) in males and yowling (calling) in females. Traditionally, this surgery was performed at around six to nine months of age, but it is increasingly being performed before puberty, at about three to six months. In the United States, about 80% of household cats are neutered.

 

Lifespan and health

Main articles: Cat health and Aging in cats

The average lifespan of pet cats has risen in recent decades. In the early 1980s, it was about seven years,: 33  rising to 9.4 years in 1995: 33  and an average of about 13 years as of 2014 and 2023. Some cats have been reported as surviving into their 30s, with the oldest known cat dying at a verified age of 38.

 

Neutering increases life expectancy: one study found castrated male cats live twice as long as intact males, while spayed female cats live 62% longer than intact females.: 35  Having a cat neutered confers health benefits, because castrated males cannot develop testicular cancer, spayed females cannot develop uterine or ovarian cancer, and both have a reduced risk of mammary cancer.

 

Disease

Main article: List of feline diseases

About 250 heritable genetic disorders have been identified in cats, many similar to human inborn errors of metabolism. The high level of similarity among the metabolism of mammals allows many of these feline diseases to be diagnosed using genetic tests that were originally developed for use in humans, as well as the use of cats as animal models in the study of the human diseases. Diseases affecting domestic cats include acute infections, parasitic infestations, injuries, and chronic diseases such as kidney disease, thyroid disease, and arthritis. Vaccinations are available for many infectious diseases, as are treatments to eliminate parasites such as worms, ticks, and fleas.

 

Ecology

Habitats

The domestic cat is a cosmopolitan species and occurs across much of the world. It is adaptable and now present on all continents except Antarctica, and on 118 of the 131 main groups of islands, even on the isolated Kerguelen Islands. Due to its ability to thrive in almost any terrestrial habitat, it is among the world's most invasive species. It lives on small islands with no human inhabitants. Feral cats can live in forests, grasslands, tundra, coastal areas, agricultural land, scrublands, urban areas, and wetlands.

 

The unwantedness that leads to the domestic cat being treated as an invasive species is twofold. On one hand, as it is little altered from the wildcat, it can readily interbreed with the wildcat. This hybridization poses a danger to the genetic distinctiveness of some wildcat populations, particularly in Scotland and Hungary, possibly also the Iberian Peninsula, and where protected natural areas are close to human-dominated landscapes, such as Kruger National Park in South Africa. However, its introduction to places where no native felines are present also contributes to the decline of native species.

 

Ferality

Main article: Feral cat

Feral cats are domestic cats that were born in or have reverted to a wild state. They are unfamiliar with and wary of humans and roam freely in urban and rural areas. The numbers of feral cats is not known, but estimates of the United States feral population range from 25 to 60 million. Feral cats may live alone, but most are found in large colonies, which occupy a specific territory and are usually associated with a source of food. Famous feral cat colonies are found in Rome around the Colosseum and Forum Romanum, with cats at some of these sites being fed and given medical attention by volunteers.

 

Public attitudes toward feral cats vary widely, from seeing them as free-ranging pets to regarding them as vermin.

 

Some feral cats can be successfully socialized and 're-tamed' for adoption; young cats, especially kittens and cats that have had prior experience and contact with humans are the most receptive to these efforts.

 

Impact on wildlife

Main article: Cat predation on wildlife

On islands, birds can contribute as much as 60% of a cat's diet. In nearly all cases, the cat cannot be identified as the sole cause for reducing the numbers of island birds, and in some instances, eradication of cats has caused a "mesopredator release" effect; where the suppression of top carnivores creates an abundance of smaller predators that cause a severe decline in their shared prey. Domestic cats are a contributing factor to the decline of many species, a factor that has ultimately led, in some cases, to extinction. The South Island piopio, Chatham rail, and the New Zealand merganser are a few from a long list, with the most extreme case being the flightless Lyall's wren, which was driven to extinction only a few years after its discovery. One feral cat in New Zealand killed 102 New Zealand lesser short-tailed bats in seven days. In the US, feral and free-ranging domestic cats kill an estimated 6.3 – 22.3 billion mammals annually.

 

In Australia, the impact of cats on mammal populations is even greater than the impact of habitat loss. More than one million reptiles are killed by feral cats each day, representing 258 species. Cats have contributed to the extinction of the Navassa curly-tailed lizard and Chioninia coctei.

 

Interaction with humans

Main article: Human interaction with cats

Cats are common pets throughout the world, and their worldwide population as of 2007 exceeded 500 million. As of 2017, the domestic cat was the second most popular pet in the United States, with 95.6 million cats owned and around 42 million households owning at least one cat. In the United Kingdom, 26% of adults have a cat, with an estimated population of 10.9 million pet cats as of 2020. As of 2021, there were an estimated 220 million owned and 480 million stray cats in the world.

 

Cats have been used for millennia to control rodents, notably around grain stores and aboard ships, and both uses extend to the present day.

 

As well as being kept as pets, cats are also used in the international fur trade and leather industries for making coats, hats, blankets, stuffed toys, shoes, gloves, and musical instruments. About 24 cats are needed to make a cat-fur coat. This use has been outlawed in the United States since 2000 and in the European Union (as well as the United Kingdom) since 2007.

 

Cat pelts have been used for superstitious purposes as part of the practice of witchcraft, and are still made into blankets in Switzerland as traditional medicine thought to cure rheumatism.

 

A few attempts to build a cat census have been made over the years, both through associations or national and international organizations (such as that of the Canadian Federation of Humane Societies) and over the Internet, but such a task does not seem simple to achieve. General estimates for the global population of domestic cats range widely from anywhere between 200 million to 600 million. Walter Chandoha made his career photographing cats after his 1949 images of Loco, an especially charming stray taken in, were published around the world. He is reported to have photographed 90,000 cats during his career and maintained an archive of 225,000 images that he drew from for publications during his lifetime.

 

Shows

Main article: Cat show

A cat show is a judged event in which the owners of cats compete to win titles in various cat-registering organizations by entering their cats to be judged after a breed standard. It is often required that a cat must be healthy and vaccinated in order to participate in a cat show. Both pedigreed and non-purebred companion ("moggy") cats are admissible, although the rules differ depending on the organization. Competing cats are compared to the applicable breed standard, and assessed for temperament.

 

Infection

Main article: Feline zoonosis

Cats can be infected or infested with viruses, bacteria, fungus, protozoans, arthropods or worms that can transmit diseases to humans. In some cases, the cat exhibits no symptoms of the disease. The same disease can then become evident in a human. The likelihood that a person will become diseased depends on the age and immune status of the person. Humans who have cats living in their home or in close association are more likely to become infected. Others might also acquire infections from cat feces and parasites exiting the cat's body. Some of the infections of most concern include salmonella, cat-scratch disease and toxoplasmosis.

 

History and mythology

Main articles: Cultural depictions of cats and Cats in ancient Egypt

In ancient Egypt, cats were worshipped, and the goddess Bastet often depicted in cat form, sometimes taking on the war-like aspect of a lioness. The Greek historian Herodotus reported that killing a cat was forbidden, and when a household cat died, the entire family mourned and shaved their eyebrows. Families took their dead cats to the sacred city of Bubastis, where they were embalmed and buried in sacred repositories. Herodotus expressed astonishment at the domestic cats in Egypt, because he had only ever seen wildcats.

 

Ancient Greeks and Romans kept weasels as pets, which were seen as the ideal rodent-killers. The earliest unmistakable evidence of the Greeks having domestic cats comes from two coins from Magna Graecia dating to the mid-fifth century BC showing Iokastos and Phalanthos, the legendary founders of Rhegion and Taras respectively, playing with their pet cats. The usual ancient Greek word for 'cat' was ailouros, meaning 'thing with the waving tail'. Cats are rarely mentioned in ancient Greek literature. Aristotle remarked in his History of Animals that "female cats are naturally lecherous." The Greeks later syncretized their own goddess Artemis with the Egyptian goddess Bastet, adopting Bastet's associations with cats and ascribing them to Artemis. In Ovid's Metamorphoses, when the deities flee to Egypt and take animal forms, the goddess Diana turns into a cat.

 

Cats eventually displaced weasels as the pest control of choice because they were more pleasant to have around the house and were more enthusiastic hunters of mice. During the Middle Ages, many of Artemis's associations with cats were grafted onto the Virgin Mary. Cats are often shown in icons of Annunciation and of the Holy Family and, according to Italian folklore, on the same night that Mary gave birth to Jesus, a cat in Bethlehem gave birth to a kitten. Domestic cats were spread throughout much of the rest of the world during the Age of Discovery, as ships' cats were carried on sailing ships to control shipboard rodents and as good-luck charms.

 

Several ancient religions believed cats are exalted souls, companions or guides for humans, that are all-knowing but mute so they cannot influence decisions made by humans. In Japan, the maneki neko cat is a symbol of good fortune. In Norse mythology, Freyja, the goddess of love, beauty, and fertility, is depicted as riding a chariot drawn by cats. In Jewish legend, the first cat was living in the house of the first man Adam as a pet that got rid of mice. The cat was once partnering with the first dog before the latter broke an oath they had made which resulted in enmity between the descendants of these two animals. It is also written that neither cats nor foxes are represented in the water, while every other animal has an incarnation species in the water. Although no species are sacred in Islam, cats are revered by Muslims. Some Western writers have stated Muhammad had a favorite cat, Muezza. He is reported to have loved cats so much, "he would do without his cloak rather than disturb one that was sleeping on it". The story has no origin in early Muslim writers, and seems to confuse a story of a later Sufi saint, Ahmed ar-Rifa'i, centuries after Muhammad. One of the companions of Muhammad was known as Abu Hurayrah ("father of the kitten"), in reference to his documented affection to cats.

 

Superstitions and rituals

Many cultures have negative superstitions about cats. An example would be the belief that encountering a black cat ("crossing one's path") leads to bad luck, or that cats are witches' familiars used to augment a witch's powers and skills. The killing of cats in Medieval Ypres, Belgium, is commemorated in the innocuous present-day Kattenstoet (cat parade). In mid-16th century France, cats would be burnt alive as a form of entertainment, particularly during midsummer festivals. According to Norman Davies, the assembled people "shrieked with laughter as the animals, howling with pain, were singed, roasted, and finally carbonized". The remaining ashes were sometimes taken back home by the people for good luck.

 

According to a myth in many cultures, cats have multiple lives. In many countries, they are believed to have nine lives, but in Italy, Germany, Greece, Brazil and some Spanish-speaking regions, they are said to have seven lives, while in Arabic traditions, the number of lives is six. An early mention of the myth can be found in John Heywood's The Proverbs of John Heywood (1546)

 

Husband, (quoth she), ye studie, be merrie now,

And even as ye thinke now, so come to yow.

Nay not so, (quoth he), for my thought to tell right,

I thinke how you lay groning, wife, all last night.

Husband, a groning horse and a groning wife

Never faile their master, (quoth she), for my life.

No wife, a woman hath nine lives like a cat.

 

The myth is attributed to the natural suppleness and swiftness cats exhibit to escape life-threatening situations. Also lending credence to this myth is the fact that falling cats often land on their feet, using an instinctive righting reflex to twist their bodies around. Nonetheless, cats can still be injured or killed by a high fall.

Dear You Tube

 

I am deeply saddened at your mindset and your regressive attitude as a You Tuber I follow You Tube guidelines.

I am a Muslim photographer an Indian by birth and my main concern has been promoting Hindu Muslim Amity through my documentation of diverse cultures that unify India a land of peace hope humanity.

I shoot Sufis Shias Naga Sadhus Aghoris Tantrics Hijras but no porn

 

You removed one one of my videos NagaSadhus Pulling a car with a penis an integral part of NagaSadhu rituals I do not mind the stance you take but to add that as a strike to my profile and character I think is biased and racist on your part..

 

Please remove my videos if you feel they are not proper as per your Victorian mindset but don't term as pornography it is an insult to my Hindu culture and all that it stands for.

 

I am deeply hurt and I think your You Tube curator has certainly lost it ask him to see Nat Geo and NagaSadhus and penile yoga and Ling Kriya.

You can age revise such videos but removing them makes me feel you don't really know the ethnicity of a religious Hindu Order who live naked the NagaSadhus of India.

I have shot 4 Kumbh Melas and here it I'd NagaSadhus of the JunaAkhara that attract thousands of photographers film makers from the West so I feel you demean my efforts as a photographer and a man of Peace.

  

As ever

Firoze Shakir

in pain as a You Tube Creator

 

www.youtube.com/user/photographerno1/community?view_as=su...

  

Message from You Tube

 

Hi Firoze Shakir,

 

As you may know, our Community Guidelines describe which content we allow – and don’t allow – on YouTube. Your video "Ling Kriya Naga Sadhu Pulling the Car With His Penis" was flagged for review. Upon review, we’ve determined that it violates our guidelines. We’ve removed it from YouTube and assigned a Community Guidelines strike, or temporary penalty, to your account.

 

Video content restrictions

If a video contains nudity, pornography, or other sexually provocative content, it's less likely to be allowed on YouTube. YouTube reviews content on a case by case basis and will make limited exceptions for appropriate educational, documentary, artistic, and scientific contexts. In such cases, we may apply an age restriction so that only viewers over a certain age can view the content. Learn more here.

The impact of strikes

This is the first strike applied to your account. We understand that users seldom intend to violate our policies. That’s why strikes don’t last forever – this strike will expire in three months. However, it’s important to remember that additional strikes could prevent you from posting content to YouTube or even lead to your account being terminated.

How you can respond

If you believe this was a mistake, we’d like to hear from you. Please follow both of these steps as simply deleting the video won’t resolve the strike on your account.

The next time you sign in you will be asked to acknowledge this strike on your account.

If you would like to appeal this strike, please submit this form. Our team will thoroughly review your appeal and will contact you again very soon.

We value your opinions and feedback. Please take a few minutes to fill out our survey.

 

Sincerely,

- The YouTube Team

 

You tube final reply

  

Dear Firoze Shakir:

Thank you for submitting your video appeal to YouTube.

 

After further review of the content, we've determined that your video does violate our Community Guidelines and have upheld our original decision. We appreciate your understanding.

 

Sincerely,

 

— The YouTube Team

 

pearlypenilepapulestube.com/how-to-remove-pearly-penile-p...

pearlypenilepapulestube.com/pearly-penile-papules-removal...

The Icelandic Phallological Museum (Icelandic: Hið Íslenzka Reðasafn), located in Reykjavík, Iceland, houses the world's largest display of penises and penile parts. The collection of 280 specimens from 93 species of animals includes 55 penises taken from whales, 36 from seals and 118 from land mammals, allegedly including elves and trolls. In July 2011, the museum obtained its first human penis, one of four promised by would-be donors. Its detachment from the donor's body did not go according to plan and it was reduced to a greyish-brown shrivelled mass pickled in a jar of formaldehyde. The museum continues to search for "a younger and a bigger and better one.

Wiki

pearlypenilepapulestube.com/%e2%98%85%e2%98%85%e2%98%85pe...

pearlypenilepapulestube.com/%e2%9d%84pearly-papules-male-...

Adult males may make several displays when showing dominance to their group or rival male. One such display is a “run display,” where the males will leap through the branches making as much noise as possible to make their presence known. Another display has the male standing on all four limbs with his tail curled in an effort to show his size and strength. Another display is climbing the vantage point and defecate or penile erection (studies suggested that a male may display to another male in the closed position as a means of exerting and establishing dominance)!

 

Capped langurs are not known to be territorial animals. Territorial ranges often overlap with multiple groups, and they do very little to defend it from intruders. The one exception to this is when a rogue male (guess it was me here :D) tries to steal a female from a group to start a group of his own. Dominant males are highly protective of their mating partners and offspring.

 

The average group of capped langurs consists of around nine individuals featuring a single dominant male, a few adult females, and their offspring. Other group types include all-male bands as well as loner males. Some groups with multiple adult males and females have been observed, but they are rare.

pearlypenilepapulestube.com/penile-pearly-papule-treatmen...

Sparks Steak House is one of the most popular steak joints in all of New York City. It was started in 1966 as Sparks Pub by two brothers, Pat and Mike Cetta. Based upon the success of their venture they decide to move it from its original home on East 18th Street up to Midtown where they have now been located since 1977. Sparks is continually rated as one of New York City’s finest steakhouses and is also well known for its lamb and veal chops.

 

Our interest here, however, does not lie in prime rib or extravagant wines, but an incident which went down here on December 16, 1985. It was here outside of Sparks at approximately 5pm that the head of the Gambino crime family, Big Paul Costellano was murdered in broad daylight as per a hit put out by John Gotti. This incident instantly elevated Gotti to the ranks of mafia chieftain.

 

To explain how this incident went down let’s begin by exploring the characters involved in our tale. First there is the former head of the Gambino family, Paul Castellano. Castellano lived his life by a series of contradictions. He attempted to legitimize many of the mobs activities and expressly forbid activities such as selling drugs. Those caught doing such activities were executed; however Castellano was known to have sold on occasion himself. Castellano also did not believe in the world of “excess” and often dressed down in public to emphasize this fact. In turn, he built a 3.5 acre, 17-room estate on Staten Island that he had constructed to mimic the White House. Lastly, Castalleno was a dedicated family man but he was openly having an affair with his maid, Gloria Olarte.

 

Gloria was an immigrant who most described as rather ordinary looking and whom spoke almost no English. Castellano was so infatuated with Gloria though that he often brought her with on family vacations and he even bought her a brand new sports car, despite the fact that she didn’t drive or have a valid license. It was also suspected by members of Castellano’s crew that he had secretly had a penile implant to help consummate his affair and to remedy impotence brought on by his diabetes. Ironically it was Gloria who attempted to frame him in 1983 and allowed the FBI to tap phones within his version of the White House. From these taps officials were able to gather sufficient evidence to charge Castellano with 24 counts of murder in 1984. These charges were still outstanding and pending upon his assassination the following year.

 

Last in our tale is the conspirator of the whole incident, Mr. John Joseph Gotti, Jr. Gotti in many ways was the antithesis of Castellano. Whereas Castallano outright forbid the selling of drugs. Gotti was prone to either turn the other cheek if not engage in the activity on occasion himself. Also, Gotti was a man who liked to flaunt the extravagances to which he was privy. Despite only living on a reported annual salary of $100,000 as a salesman of plumbing supplies, Gotti was often seen sporting new $1,000 suits and wearing monogrammed socks. In truth it was estimated that Gotti brought in approximately $10-$12 million of cash annually. He took his arrogance even further when he would catch police officers watching him while on stakeouts. He would be so bold as to walk directly up to the vehicles and tell the officers, “Naughty, naughty.”

 

It was probably in part due to Gotti’s arrogance that led him to plot the murder of the head of the Gambino family. Gotti, along with other members of Castellano’s family, had become increasingly upset with him over numerous affairs including the contradictions to which Castellano had long been running the Gambino family. Gotti, in fact largely expected that Castellano was planning action versus him for selling drugs. Thus Gotti concocted a plan to take matters into his own hands.

 

On the afternoon of December 16, 1985, four men all dressed in trench coats and hats approached a vehicle which was unloading passengers outside the front entrance of Sparks Steak House. All men appeared identical (a detail not lost in Gotti’s plans) and even simultaneously drew their hidden weapons to open fire on the unsuspecting mobster boss along with his number two man, Tommy Bilotti. Both men were killed instantly and the gunmen casually away from the incident towards Second Avenue. One of the men was seen speaking into a walkie-talkie just prior to being picked up by their getaway vehicle (which had been parked one block over on 47th Street) and driving off.

 

While police largely suspected Gotti for this crime they were never able to pin the charges upon him. They did bring federal charges versus him for violation of the Racketeer Influenced and Corrupt Organizations Act (RICO) for which he went to trial in August 1986. The jury ultimately ended up finding him non-guilty though, although it was later discovered that Gotti had actually bribed the jury foreman (who was later convicted himself on bribery charges).

 

Needing a new plan, federal agents began to track Gotti’s whereabouts to the Ravenite Social Club where he often frequented. Through the usage of bug devices planted in the apartment above the Ravenite, the Feds gathered audio evidence which implicated Gotti on thirteen charges, including five murders. He was arrested within the Ravenite Club in December 1990 and found guilty on all accounts on April 2, 1992. For his crimes, Gotti was sentenced to life in prison. He served his time in a federal prison in Springfield, MO where he would pass away at the age of 61 in 2002.

 

For more history regarding this site, including how you can visit this locale via one of our MP3 audio walking tours, check out our site here: iwalkedaudiotours.com/2012/10/iwalked-new-york-citys-spar...

The cat (Felis catus), commonly referred to as the domestic cat or house cat, is the only domesticated species in the family Felidae. Recent advances in archaeology and genetics have shown that the domestication of the cat occurred in the Near East around 7500 BC. It is commonly kept as a house pet and farm cat, but also ranges freely as a feral cat avoiding human contact. It is valued by humans for companionship and its ability to kill vermin. Because of its retractable claws it is adapted to killing small prey like mice and rats. It has a strong flexible body, quick reflexes, sharp teeth, and its night vision and sense of smell are well developed. It is a social species, but a solitary hunter and a crepuscular predator. Cat communication includes vocalizations like meowing, purring, trilling, hissing, growling, and grunting as well as cat body language. It can hear sounds too faint or too high in frequency for human ears, such as those made by small mammals. It also secretes and perceives pheromones.

 

Female domestic cats can have kittens from spring to late autumn in temperate zones and throughout the year in equatorial regions, with litter sizes often ranging from two to five kittens. Domestic cats are bred and shown at events as registered pedigreed cats, a hobby known as cat fancy. Animal population control of cats may be achieved by spaying and neutering, but their proliferation and the abandonment of pets has resulted in large numbers of feral cats worldwide, contributing to the extinction of bird, mammal and reptile species.

 

As of 2017, the domestic cat was the second most popular pet in the United States, with 95.6 million cats owned and around 42 million households owning at least one cat. In the United Kingdom, 26% of adults have a cat, with an estimated population of 10.9 million pet cats as of 2020. As of 2021, there were an estimated 220 million owned and 480 million stray cats in the world.

 

Etymology and naming

The origin of the English word cat, Old English catt, is thought to be the Late Latin word cattus, which was first used at the beginning of the 6th century. The Late Latin word may be derived from an unidentified African language. The Nubian word kaddîska 'wildcat' and Nobiin kadīs are possible sources or cognates. The Nubian word may be a loan from Arabic قَطّ‎ qaṭṭ ~ قِطّ qiṭṭ.

 

The forms might also have derived from an ancient Germanic word that was imported into Latin and then into Greek, Syriac, and Arabic. The word may be derived from Germanic and Northern European languages, and ultimately be borrowed from Uralic, cf. Northern Sámi gáđfi, 'female stoat', and Hungarian hölgy, 'lady, female stoat'; from Proto-Uralic *käďwä, 'female (of a furred animal)'.

 

The English puss, extended as pussy and pussycat, is attested from the 16th century and may have been introduced from Dutch poes or from Low German puuskatte, related to Swedish kattepus, or Norwegian pus, pusekatt. Similar forms exist in Lithuanian puižė and Irish puisín or puiscín. The etymology of this word is unknown, but it may have arisen from a sound used to attract a cat.

 

A male cat is called a tom or tomcat (or a gib, if neutered). A female is called a queen or a molly, if spayed, especially in a cat-breeding context. A juvenile cat is referred to as a kitten. In Early Modern English, the word kitten was interchangeable with the now-obsolete word catling.

 

A group of cats can be referred to as a clowder or a glaring.

 

Taxonomy

The scientific name Felis catus was proposed by Carl Linnaeus in 1758 for a domestic cat. Felis catus domesticus was proposed by Johann Christian Polycarp Erxleben in 1777. Felis daemon proposed by Konstantin Satunin in 1904 was a black cat from the Transcaucasus, later identified as a domestic cat.

 

In 2003, the International Commission on Zoological Nomenclature ruled that the domestic cat is a distinct species, namely Felis catus. In 2007, it was considered a subspecies, F. silvestris catus, of the European wildcat (F. silvestris) following results of phylogenetic research. In 2017, the IUCN Cat Classification Taskforce followed the recommendation of the ICZN in regarding the domestic cat as a distinct species, Felis catus.

 

Evolution

Main article: Cat evolution

The domestic cat is a member of the Felidae, a family that had a common ancestor about 10 to 15 million years ago. The evolutionary radiation of the Felidae began in Asia during the Miocene around 8.38 to 14.45 million years ago. Analysis of mitochondrial DNA of all Felidae species indicates a radiation at 6.46 to 16.76 million years ago. The genus Felis genetically diverged from other Felidae around 6 to 7 million years ago. Results of phylogenetic research shows that the wild members of this genus evolved through sympatric or parapatric speciation, whereas the domestic cat evolved through artificial selection. The domestic cat and its closest wild ancestor are diploid and both possess 38 chromosomes and roughly 20,000 genes.

 

Domestication

See also: Domestication of the cat and Cats in ancient Egypt

It was long thought that the domestication of the cat began in ancient Egypt, where cats were venerated from around 3100 BC, However, the earliest known indication for the taming of an African wildcat was excavated close by a human Neolithic grave in Shillourokambos, southern Cyprus, dating to about 7500–7200 BC. Since there is no evidence of native mammalian fauna on Cyprus, the inhabitants of this Neolithic village most likely brought the cat and other wild mammals to the island from the Middle Eastern mainland. Scientists therefore assume that African wildcats were attracted to early human settlements in the Fertile Crescent by rodents, in particular the house mouse (Mus musculus), and were tamed by Neolithic farmers. This mutual relationship between early farmers and tamed cats lasted thousands of years. As agricultural practices spread, so did tame and domesticated cats. Wildcats of Egypt contributed to the maternal gene pool of the domestic cat at a later time.

The earliest known evidence for the occurrence of the domestic cat in Greece dates to around 1200 BC. Greek, Phoenician, Carthaginian and Etruscan traders introduced domestic cats to southern Europe. During the Roman Empire they were introduced to Corsica and Sardinia before the beginning of the 1st millennium. By the 5th century BC, they were familiar animals around settlements in Magna Graecia and Etruria. By the end of the Western Roman Empire in the 5th century, the Egyptian domestic cat lineage had arrived in a Baltic Sea port in northern Germany.

 

The leopard cat (Prionailurus bengalensis) was tamed independently in China around 5500 BC. This line of partially domesticated cats leaves no trace in the domestic cat populations of today.

 

During domestication, cats have undergone only minor changes in anatomy and behavior, and they are still capable of surviving in the wild. Several natural behaviors and characteristics of wildcats may have pre-adapted them for domestication as pets. These traits include their small size, social nature, obvious body language, love of play, and high intelligence. Captive Leopardus cats may also display affectionate behavior toward humans but were not domesticated. House cats often mate with feral cats. Hybridisation between domestic and other Felinae species is also possible, producing hybrids such as the Kellas cat in Scotland.

 

Development of cat breeds started in the mid 19th century. An analysis of the domestic cat genome revealed that the ancestral wildcat genome was significantly altered in the process of domestication, as specific mutations were selected to develop cat breeds. Most breeds are founded on random-bred domestic cats. Genetic diversity of these breeds varies between regions, and is lowest in purebred populations, which show more than 20 deleterious genetic disorders.

 

Characteristics

Main article: Cat anatomy

Size

The domestic cat has a smaller skull and shorter bones than the European wildcat. It averages about 46 cm (18 in) in head-to-body length and 23–25 cm (9.1–9.8 in) in height, with about 30 cm (12 in) long tails. Males are larger than females. Adult domestic cats typically weigh 4–5 kg (8.8–11.0 lb).

 

Skeleton

Cats have seven cervical vertebrae (as do most mammals); 13 thoracic vertebrae (humans have 12); seven lumbar vertebrae (humans have five); three sacral vertebrae (as do most mammals, but humans have five); and a variable number of caudal vertebrae in the tail (humans have only three to five vestigial caudal vertebrae, fused into an internal coccyx).  The extra lumbar and thoracic vertebrae account for the cat's spinal mobility and flexibility. Attached to the spine are 13 ribs, the shoulder, and the pelvis.  Unlike human arms, cat forelimbs are attached to the shoulder by free-floating clavicle bones which allow them to pass their body through any space into which they can fit their head.

 

Skull

The cat skull is unusual among mammals in having very large eye sockets and a powerful specialized jaw.  Within the jaw, cats have teeth adapted for killing prey and tearing meat. When it overpowers its prey, a cat delivers a lethal neck bite with its two long canine teeth, inserting them between two of the prey's vertebrae and severing its spinal cord, causing irreversible paralysis and death. Compared to other felines, domestic cats have narrowly spaced canine teeth relative to the size of their jaw, which is an adaptation to their preferred prey of small rodents, which have small vertebrae.

 

The premolar and first molar together compose the carnassial pair on each side of the mouth, which efficiently shears meat into small pieces, like a pair of scissors. These are vital in feeding, since cats' small molars cannot chew food effectively, and cats are largely incapable of mastication.:  Cats tend to have better teeth than most humans, with decay generally less likely because of a thicker protective layer of enamel, a less damaging saliva, less retention of food particles between teeth, and a diet mostly devoid of sugar. Nonetheless, they are subject to occasional tooth loss and infection.

 

Claws

Cats have protractible and retractable claws. In their normal, relaxed position, the claws are sheathed with the skin and fur around the paw's toe pads. This keeps the claws sharp by preventing wear from contact with the ground and allows for the silent stalking of prey. The claws on the forefeet are typically sharper than those on the hindfeet. Cats can voluntarily extend their claws on one or more paws. They may extend their claws in hunting or self-defense, climbing, kneading, or for extra traction on soft surfaces. Cats shed the outside layer of their claw sheaths when scratching rough surfaces.

 

Most cats have five claws on their front paws and four on their rear paws. The dewclaw is proximal to the other claws. More proximally is a protrusion which appears to be a sixth "finger". This special feature of the front paws on the inside of the wrists has no function in normal walking but is thought to be an antiskidding device used while jumping. Some cat breeds are prone to having extra digits ("polydactyly"). Polydactylous cats occur along North America's northeast coast and in Great Britain.

 

Ambulation

The cat is digitigrade. It walks on the toes, with the bones of the feet making up the lower part of the visible leg. Unlike most mammals, it uses a "pacing" gait and moves both legs on one side of the body before the legs on the other side. It registers directly by placing each hind paw close to the track of the corresponding fore paw, minimizing noise and visible tracks. This also provides sure footing for hind paws when navigating rough terrain. As it speeds up from walking to trotting, its gait changes to a "diagonal" gait: The diagonally opposite hind and fore legs move simultaneously.

 

Balance

Cats are generally fond of sitting in high places or perching. A higher place may serve as a concealed site from which to hunt; domestic cats strike prey by pouncing from a perch such as a tree branch. Another possible explanation is that height gives the cat a better observation point, allowing it to survey its territory. A cat falling from heights of up to 3 m (9.8 ft) can right itself and land on its paws.

 

During a fall from a high place, a cat reflexively twists its body and rights itself to land on its feet using its acute sense of balance and flexibility. This reflex is known as the cat righting reflex. A cat always rights itself in the same way during a fall, if it has enough time to do so, which is the case in falls of 90 cm (3.0 ft) or more. How cats are able to right themselves when falling has been investigated as the "falling cat problem".

 

Coats

Main article: Cat coat genetics

The cat family (Felidae) can pass down many colors and patterns to their offspring. The domestic cat genes MC1R and ASIP allow for the variety of color in coats. The feline ASIP gene consists of three coding exons. Three novel microsatellite markers linked to ASIP were isolated from a domestic cat BAC clone containing this gene and were used to perform linkage analysis in a pedigree of 89 domestic cats that segregated for melanism.[citation needed]

 

Senses

Main article: Cat senses

Vision

A cat's nictitating membrane shown as it blinks

Cats have excellent night vision and can see at only one-sixth the light level required for human vision.  This is partly the result of cat eyes having a tapetum lucidum, which reflects any light that passes through the retina back into the eye, thereby increasing the eye's sensitivity to dim light. Large pupils are an adaptation to dim light. The domestic cat has slit pupils, which allow it to focus bright light without chromatic aberration. At low light, a cat's pupils expand to cover most of the exposed surface of its eyes. The domestic cat has rather poor color vision and only two types of cone cells, optimized for sensitivity to blue and yellowish green; its ability to distinguish between red and green is limited. A response to middle wavelengths from a system other than the rod cells might be due to a third type of cone. This appears to be an adaptation to low light levels rather than representing true trichromatic vision. Cats also have a nictitating membrane, allowing them to blink without hindering their vision.

 

Hearing

The domestic cat's hearing is most acute in the range of 500 Hz to 32 kHz. It can detect an extremely broad range of frequencies ranging from 55 Hz to 79 kHz, whereas humans can only detect frequencies between 20 Hz and 20 kHz. It can hear a range of 10.5 octaves, while humans and dogs can hear ranges of about 9 octaves. Its hearing sensitivity is enhanced by its large movable outer ears, the pinnae, which amplify sounds and help detect the location of a noise. It can detect ultrasound, which enables it to detect ultrasonic calls made by rodent prey. Recent research has shown that cats have socio-spatial cognitive abilities to create mental maps of owners' locations based on hearing owners' voices.

 

Smell

Cats have an acute sense of smell, due in part to their well-developed olfactory bulb and a large surface of olfactory mucosa, about 5.8 cm2 (0.90 in2) in area, which is about twice that of humans. Cats and many other animals have a Jacobson's organ in their mouths that is used in the behavioral process of flehmening. It allows them to sense certain aromas in a way that humans cannot. Cats are sensitive to pheromones such as 3-mercapto-3-methylbutan-1-ol, which they use to communicate through urine spraying and marking with scent glands. Many cats also respond strongly to plants that contain nepetalactone, especially catnip, as they can detect that substance at less than one part per billion. About 70–80% of cats are affected by nepetalactone. This response is also produced by other plants, such as silver vine (Actinidia polygama) and the herb valerian; it may be caused by the smell of these plants mimicking a pheromone and stimulating cats' social or sexual behaviors.

 

Taste

Cats have relatively few taste buds compared to humans (470 or so versus more than 9,000 on the human tongue). Domestic and wild cats share a taste receptor gene mutation that keeps their sweet taste buds from binding to sugary molecules, leaving them with no ability to taste sweetness. They, however, possess taste bud receptors specialized for acids, amino acids like protein, and bitter tastes. Their taste buds possess the receptors needed to detect umami. However, these receptors contain molecular changes that make the cat taste of umami different from that of humans. In humans, they detect the amino acids of glutamic acid and aspartic acid, but in cats they instead detect nucleotides, in this case inosine monophosphate and l-Histidine. These nucleotides are particularly enriched in tuna. This has been argued is why cats find tuna so palatable: as put by researchers into cat taste, "the specific combination of the high IMP and free l-Histidine contents of tuna" .. "produces a strong umami taste synergy that is highly preferred by cats". One of the researchers involved in this research has further claimed, "I think umami is as important for cats as sweet is for humans".[87]

 

Cats also have a distinct temperature preference for their food, preferring food with a temperature around 38 °C (100 °F) which is similar to that of a fresh kill; some cats reject cold food (which would signal to the cat that the "prey" item is long dead and therefore possibly toxic or decomposing).

 

Whiskers

To aid with navigation and sensation, cats have dozens of movable whiskers (vibrissae) over their body, especially their faces. These provide information on the width of gaps and on the location of objects in the dark, both by touching objects directly and by sensing air currents; they also trigger protective blink reflexes to protect the eyes from damage.: 47 

 

Behavior

See also: Cat behavior

Outdoor cats are active both day and night, although they tend to be slightly more active at night.[88] Domestic cats spend the majority of their time in the vicinity of their homes but can range many hundreds of meters from this central point. They establish territories that vary considerably in size, in one study ranging 7–28 ha (17–69 acres). The timing of cats' activity is quite flexible and varied but being low-light predators, they are generally crepuscular, which means they tend to be more active near dawn and dusk. However, house cats' behavior is also influenced by human activity and they may adapt to their owners' sleeping patterns to some extent.

 

Cats conserve energy by sleeping more than most animals, especially as they grow older. The daily duration of sleep varies, usually between 12 and 16 hours, with 13 and 14 being the average. Some cats can sleep as much as 20 hours. The term "cat nap" for a short rest refers to the cat's tendency to fall asleep (lightly) for a brief period. While asleep, cats experience short periods of rapid eye movement sleep often accompanied by muscle twitches, which suggests they are dreaming.

 

Sociability

The social behavior of the domestic cat ranges from widely dispersed individuals to feral cat colonies that gather around a food source, based on groups of co-operating females. Within such groups, one cat is usually dominant over the others. Each cat in a colony holds a distinct territory, with sexually active males having the largest territories, which are about 10 times larger than those of female cats and may overlap with several females' territories. These territories are marked by urine spraying, by rubbing objects at head height with secretions from facial glands, and by defecation. Between these territories are neutral areas where cats watch and greet one another without territorial conflicts. Outside these neutral areas, territory holders usually chase away stranger cats, at first by staring, hissing, and growling and, if that does not work, by short but noisy and violent attacks. Despite this colonial organization, cats do not have a social survival strategy or a herd behavior, and always hunt alone.

 

Life in proximity to humans and other domestic animals has led to a symbiotic social adaptation in cats, and cats may express great affection toward humans or other animals. Ethologically, a cat's human keeper functions as if a mother surrogate. Adult cats live their lives in a kind of extended kittenhood, a form of behavioral neoteny. Their high-pitched sounds may mimic the cries of a hungry human infant, making them particularly difficult for humans to ignore. Some pet cats are poorly socialized. In particular, older cats show aggressiveness toward newly arrived kittens, which include biting and scratching; this type of behavior is known as feline asocial aggression.

 

Redirected aggression is a common form of aggression which can occur in multiple cat households. In redirected aggression there is usually something that agitates the cat: this could be a sight, sound, or another source of stimuli which causes a heightened level of anxiety or arousal. If the cat cannot attack the stimuli, it may direct anger elsewhere by attacking or directing aggression to the nearest cat, dog, human or other being.

 

Domestic cats' scent rubbing behavior toward humans or other cats is thought to be a feline means for social bonding.

 

Communication

Main article: Cat communication

Domestic cats use many vocalizations for communication, including purring, trilling, hissing, growling/snarling, grunting, and several different forms of meowing. Their body language, including position of ears and tail, relaxation of the whole body, and kneading of the paws, are all indicators of mood. The tail and ears are particularly important social signal mechanisms in cats. A raised tail indicates a friendly greeting, and flattened ears indicate hostility. Tail-raising also indicates the cat's position in the group's social hierarchy, with dominant individuals raising their tails less often than subordinate ones. Feral cats are generally silent.: 208  Nose-to-nose touching is also a common greeting and may be followed by social grooming, which is solicited by one of the cats raising and tilting its head.

 

Purring may have developed as an evolutionary advantage as a signaling mechanism of reassurance between mother cats and nursing kittens, who are thought to use it as a care-soliciting signal. Post-nursing cats also often purr as a sign of contentment: when being petted, becoming relaxed, or eating. Even though purring is popularly interpreted as indicative of pleasure, it has been recorded in a wide variety of circumstances, most of which involve physical contact between the cat and another, presumably trusted individual. Some cats have been observed to purr continuously when chronically ill or in apparent pain.

 

The exact mechanism by which cats purr has long been elusive, but it has been proposed that purring is generated via a series of sudden build-ups and releases of pressure as the glottis is opened and closed, which causes the vocal folds to separate forcefully. The laryngeal muscles in control of the glottis are thought to be driven by a neural oscillator which generates a cycle of contraction and release every 30–40 milliseconds (giving a frequency of 33 to 25 Hz).

 

Domestic cats observed in a rescue facility have total of 276 distinct facial expressions based on 26 different facial movements; each facial expression corresponds to different social functions that are likely influenced by domestication.

 

Grooming

Cats are known for spending considerable amounts of time licking their coats to keep them clean. The cat's tongue has backward-facing spines about 500 μm long, which are called papillae. These contain keratin which makes them rigid so the papillae act like a hairbrush. Some cats, particularly longhaired cats, occasionally regurgitate hairballs of fur that have collected in their stomachs from grooming. These clumps of fur are usually sausage-shaped and about 2–3 cm (0.79–1.18 in) long. Hairballs can be prevented with remedies that ease elimination of the hair through the gut, as well as regular grooming of the coat with a comb or stiff brush.

 

Fighting

Among domestic cats, males are more likely to fight than females. Among feral cats, the most common reason for cat fighting is competition between two males to mate with a female. In such cases, most fights are won by the heavier male. Another common reason for fighting in domestic cats is the difficulty of establishing territories within a small home. Female cats also fight over territory or to defend their kittens. Neutering will decrease or eliminate this behavior in many cases, suggesting that the behavior is linked to sex hormones.

 

When cats become aggressive, they try to make themselves appear larger and more threatening by raising their fur, arching their backs, turning sideways and hissing or spitting. Often, the ears are pointed down and back to avoid damage to the inner ear and potentially listen for any changes behind them while focused forward. Cats may also vocalize loudly and bare their teeth in an effort to further intimidate their opponents. Fights usually consist of grappling and delivering powerful slaps to the face and body with the forepaws as well as bites. Cats also throw themselves to the ground in a defensive posture to rake their opponent's belly with their powerful hind legs.

 

Serious damage is rare, as the fights are usually short in duration, with the loser running away with little more than a few scratches to the face and ears. Fights for mating rights are typically more severe and injuries may include deep puncture wounds and lacerations. Normally, serious injuries from fighting are limited to infections of scratches and bites, though these can occasionally kill cats if untreated. In addition, bites are probably the main route of transmission of feline immunodeficiency virus. Sexually active males are usually involved in many fights during their lives, and often have decidedly battered faces with obvious scars and cuts to their ears and nose. Cats are willing to threaten animals larger than them to defend their territory, such as dogs and foxes.

 

Hunting and feeding

See also: Cat food

The shape and structure of cats' cheeks is insufficient to allow them to take in liquids using suction. Therefore, when drinking they lap with the tongue to draw liquid upward into their mouths. Lapping at a rate of four times a second, the cat touches the smooth tip of its tongue to the surface of the water, and quickly retracts it like a corkscrew, drawing water upward.

 

Feral cats and free-fed house cats consume several small meals in a day. The frequency and size of meals varies between individuals. They select food based on its temperature, smell and texture; they dislike chilled foods and respond most strongly to moist foods rich in amino acids, which are similar to meat. Cats reject novel flavors (a response termed neophobia) and learn quickly to avoid foods that have tasted unpleasant in the past. It is also a common misconception that cats like milk/cream, as they tend to avoid sweet food and milk. Most adult cats are lactose intolerant; the sugar in milk is not easily digested and may cause soft stools or diarrhea. Some also develop odd eating habits and like to eat or chew on things like wool, plastic, cables, paper, string, aluminum foil, or even coal. This condition, pica, can threaten their health, depending on the amount and toxicity of the items eaten.

 

Cats hunt small prey, primarily birds and rodents, and are often used as a form of pest control. Other common small creatures such as lizards and snakes may also become prey. Cats use two hunting strategies, either stalking prey actively, or waiting in ambush until an animal comes close enough to be captured. The strategy used depends on the prey species in the area, with cats waiting in ambush outside burrows, but tending to actively stalk birds.: 153  Domestic cats are a major predator of wildlife in the United States, killing an estimated 1.3 to 4.0 billion birds and 6.3 to 22.3 billion mammals annually.

 

Certain species appear more susceptible than others; in one English village, for example, 30% of house sparrow mortality was linked to the domestic cat. In the recovery of ringed robins (Erithacus rubecula) and dunnocks (Prunella modularis) in Britain, 31% of deaths were a result of cat predation. In parts of North America, the presence of larger carnivores such as coyotes which prey on cats and other small predators reduces the effect of predation by cats and other small predators such as opossums and raccoons on bird numbers and variety.

 

Perhaps the best-known element of cats' hunting behavior, which is commonly misunderstood and often appalls cat owners because it looks like torture, is that cats often appear to "play" with prey by releasing and recapturing it. This cat and mouse behavior is due to an instinctive imperative to ensure that the prey is weak enough to be killed without endangering the cat.

 

Another poorly understood element of cat hunting behavior is the presentation of prey to human guardians. One explanation is that cats adopt humans into their social group and share excess kill with others in the group according to the dominance hierarchy, in which humans are reacted to as if they are at or near the top. Another explanation is that they attempt to teach their guardians to hunt or to help their human as if feeding "an elderly cat, or an inept kitten". This hypothesis is inconsistent with the fact that male cats also bring home prey, despite males having negligible involvement in raising kittens.:

 

Play

Main article: Cat play and toys

Domestic cats, especially young kittens, are known for their love of play. This behavior mimics hunting and is important in helping kittens learn to stalk, capture, and kill prey. Cats also engage in play fighting, with each other and with humans. This behavior may be a way for cats to practice the skills needed for real combat, and might also reduce any fear they associate with launching attacks on other animals.

 

Cats also tend to play with toys more when they are hungry. Owing to the close similarity between play and hunting, cats prefer to play with objects that resemble prey, such as small furry toys that move rapidly, but rapidly lose interest. They become habituated to a toy they have played with before. String is often used as a toy, but if it is eaten, it can become caught at the base of the cat's tongue and then move into the intestines, a medical emergency which can cause serious illness, even death. Owing to the risks posed by cats eating string, it is sometimes replaced with a laser pointer's dot, which cats may chase.

 

Reproduction

See also: Kitten

The cat secretes and perceives pheromones. Female cats, called queens, are polyestrous with several estrus cycles during a year, lasting usually 21 days. They are usually ready to mate between early February and August in northern temperate zones and throughout the year in equatorial regions.

 

Several males, called tomcats, are attracted to a female in heat. They fight over her, and the victor wins the right to mate. At first, the female rejects the male, but eventually, the female allows the male to mate. The female utters a loud yowl as the male pulls out of her because a male cat's penis has a band of about 120–150 backward-pointing penile spines, which are about 1 mm (0.039 in) long; upon withdrawal of the penis, the spines may provide the female with increased sexual stimulation, which acts to induce ovulation.

 

After mating, the female cleans her vulva thoroughly. If a male attempts to mate with her at this point, the female attacks him. After about 20 to 30 minutes, once the female is finished grooming, the cycle will repeat. Because ovulation is not always triggered by a single mating, females may not be impregnated by the first male with which they mate. Furthermore, cats are superfecund; that is, a female may mate with more than one male when she is in heat, with the result that different kittens in a litter may have different fathers.

 

The morula forms 124 hours after conception. At 148 hours, early blastocysts form. At 10–12 days, implantation occurs. The gestation of queens lasts between 64 and 67 days, with an average of 65 days.

 

Data on the reproductive capacity of more than 2,300 free-ranging queens were collected during a study between May 1998 and October 2000. They had one to six kittens per litter, with an average of three kittens. They produced a mean of 1.4 litters per year, but a maximum of three litters in a year. Of 169 kittens, 127 died before they were six months old due to a trauma caused in most cases by dog attacks and road accidents. The first litter is usually smaller than subsequent litters. Kittens are weaned between six and seven weeks of age. Queens normally reach sexual maturity at 5–10 months, and males at 5–7 months. This varies depending on breed. Kittens reach puberty at the age of 9–10 months.

 

Cats are ready to go to new homes at about 12 weeks of age, when they are ready to leave their mother. They can be surgically sterilized (spayed or castrated) as early as seven weeks to limit unwanted reproduction. This surgery also prevents undesirable sex-related behavior, such as aggression, territory marking (spraying urine) in males and yowling (calling) in females. Traditionally, this surgery was performed at around six to nine months of age, but it is increasingly being performed before puberty, at about three to six months. In the United States, about 80% of household cats are neutered.

 

Lifespan and health

Main articles: Cat health and Aging in cats

The average lifespan of pet cats has risen in recent decades. In the early 1980s, it was about seven years,: 33  rising to 9.4 years in 1995: 33  and an average of about 13 years as of 2014 and 2023. Some cats have been reported as surviving into their 30s, with the oldest known cat dying at a verified age of 38.

 

Neutering increases life expectancy: one study found castrated male cats live twice as long as intact males, while spayed female cats live 62% longer than intact females.: 35  Having a cat neutered confers health benefits, because castrated males cannot develop testicular cancer, spayed females cannot develop uterine or ovarian cancer, and both have a reduced risk of mammary cancer.

 

Disease

Main article: List of feline diseases

About 250 heritable genetic disorders have been identified in cats, many similar to human inborn errors of metabolism. The high level of similarity among the metabolism of mammals allows many of these feline diseases to be diagnosed using genetic tests that were originally developed for use in humans, as well as the use of cats as animal models in the study of the human diseases. Diseases affecting domestic cats include acute infections, parasitic infestations, injuries, and chronic diseases such as kidney disease, thyroid disease, and arthritis. Vaccinations are available for many infectious diseases, as are treatments to eliminate parasites such as worms, ticks, and fleas.

 

Ecology

Habitats

The domestic cat is a cosmopolitan species and occurs across much of the world. It is adaptable and now present on all continents except Antarctica, and on 118 of the 131 main groups of islands, even on the isolated Kerguelen Islands. Due to its ability to thrive in almost any terrestrial habitat, it is among the world's most invasive species. It lives on small islands with no human inhabitants. Feral cats can live in forests, grasslands, tundra, coastal areas, agricultural land, scrublands, urban areas, and wetlands.

 

The unwantedness that leads to the domestic cat being treated as an invasive species is twofold. On one hand, as it is little altered from the wildcat, it can readily interbreed with the wildcat. This hybridization poses a danger to the genetic distinctiveness of some wildcat populations, particularly in Scotland and Hungary, possibly also the Iberian Peninsula, and where protected natural areas are close to human-dominated landscapes, such as Kruger National Park in South Africa. However, its introduction to places where no native felines are present also contributes to the decline of native species.

 

Ferality

Main article: Feral cat

Feral cats are domestic cats that were born in or have reverted to a wild state. They are unfamiliar with and wary of humans and roam freely in urban and rural areas. The numbers of feral cats is not known, but estimates of the United States feral population range from 25 to 60 million. Feral cats may live alone, but most are found in large colonies, which occupy a specific territory and are usually associated with a source of food. Famous feral cat colonies are found in Rome around the Colosseum and Forum Romanum, with cats at some of these sites being fed and given medical attention by volunteers.

 

Public attitudes toward feral cats vary widely, from seeing them as free-ranging pets to regarding them as vermin.

 

Some feral cats can be successfully socialized and 're-tamed' for adoption; young cats, especially kittens and cats that have had prior experience and contact with humans are the most receptive to these efforts.

 

Impact on wildlife

Main article: Cat predation on wildlife

On islands, birds can contribute as much as 60% of a cat's diet. In nearly all cases, the cat cannot be identified as the sole cause for reducing the numbers of island birds, and in some instances, eradication of cats has caused a "mesopredator release" effect; where the suppression of top carnivores creates an abundance of smaller predators that cause a severe decline in their shared prey. Domestic cats are a contributing factor to the decline of many species, a factor that has ultimately led, in some cases, to extinction. The South Island piopio, Chatham rail, and the New Zealand merganser are a few from a long list, with the most extreme case being the flightless Lyall's wren, which was driven to extinction only a few years after its discovery. One feral cat in New Zealand killed 102 New Zealand lesser short-tailed bats in seven days. In the US, feral and free-ranging domestic cats kill an estimated 6.3 – 22.3 billion mammals annually.

 

In Australia, the impact of cats on mammal populations is even greater than the impact of habitat loss. More than one million reptiles are killed by feral cats each day, representing 258 species. Cats have contributed to the extinction of the Navassa curly-tailed lizard and Chioninia coctei.

 

Interaction with humans

Main article: Human interaction with cats

Cats are common pets throughout the world, and their worldwide population as of 2007 exceeded 500 million. As of 2017, the domestic cat was the second most popular pet in the United States, with 95.6 million cats owned and around 42 million households owning at least one cat. In the United Kingdom, 26% of adults have a cat, with an estimated population of 10.9 million pet cats as of 2020. As of 2021, there were an estimated 220 million owned and 480 million stray cats in the world.

 

Cats have been used for millennia to control rodents, notably around grain stores and aboard ships, and both uses extend to the present day.

 

As well as being kept as pets, cats are also used in the international fur trade and leather industries for making coats, hats, blankets, stuffed toys, shoes, gloves, and musical instruments. About 24 cats are needed to make a cat-fur coat. This use has been outlawed in the United States since 2000 and in the European Union (as well as the United Kingdom) since 2007.

 

Cat pelts have been used for superstitious purposes as part of the practice of witchcraft, and are still made into blankets in Switzerland as traditional medicine thought to cure rheumatism.

 

A few attempts to build a cat census have been made over the years, both through associations or national and international organizations (such as that of the Canadian Federation of Humane Societies) and over the Internet, but such a task does not seem simple to achieve. General estimates for the global population of domestic cats range widely from anywhere between 200 million to 600 million. Walter Chandoha made his career photographing cats after his 1949 images of Loco, an especially charming stray taken in, were published around the world. He is reported to have photographed 90,000 cats during his career and maintained an archive of 225,000 images that he drew from for publications during his lifetime.

 

Shows

Main article: Cat show

A cat show is a judged event in which the owners of cats compete to win titles in various cat-registering organizations by entering their cats to be judged after a breed standard. It is often required that a cat must be healthy and vaccinated in order to participate in a cat show. Both pedigreed and non-purebred companion ("moggy") cats are admissible, although the rules differ depending on the organization. Competing cats are compared to the applicable breed standard, and assessed for temperament.

 

Infection

Main article: Feline zoonosis

Cats can be infected or infested with viruses, bacteria, fungus, protozoans, arthropods or worms that can transmit diseases to humans. In some cases, the cat exhibits no symptoms of the disease. The same disease can then become evident in a human. The likelihood that a person will become diseased depends on the age and immune status of the person. Humans who have cats living in their home or in close association are more likely to become infected. Others might also acquire infections from cat feces and parasites exiting the cat's body. Some of the infections of most concern include salmonella, cat-scratch disease and toxoplasmosis.

 

History and mythology

Main articles: Cultural depictions of cats and Cats in ancient Egypt

In ancient Egypt, cats were worshipped, and the goddess Bastet often depicted in cat form, sometimes taking on the war-like aspect of a lioness. The Greek historian Herodotus reported that killing a cat was forbidden, and when a household cat died, the entire family mourned and shaved their eyebrows. Families took their dead cats to the sacred city of Bubastis, where they were embalmed and buried in sacred repositories. Herodotus expressed astonishment at the domestic cats in Egypt, because he had only ever seen wildcats.

 

Ancient Greeks and Romans kept weasels as pets, which were seen as the ideal rodent-killers. The earliest unmistakable evidence of the Greeks having domestic cats comes from two coins from Magna Graecia dating to the mid-fifth century BC showing Iokastos and Phalanthos, the legendary founders of Rhegion and Taras respectively, playing with their pet cats. The usual ancient Greek word for 'cat' was ailouros, meaning 'thing with the waving tail'. Cats are rarely mentioned in ancient Greek literature. Aristotle remarked in his History of Animals that "female cats are naturally lecherous." The Greeks later syncretized their own goddess Artemis with the Egyptian goddess Bastet, adopting Bastet's associations with cats and ascribing them to Artemis. In Ovid's Metamorphoses, when the deities flee to Egypt and take animal forms, the goddess Diana turns into a cat.

 

Cats eventually displaced weasels as the pest control of choice because they were more pleasant to have around the house and were more enthusiastic hunters of mice. During the Middle Ages, many of Artemis's associations with cats were grafted onto the Virgin Mary. Cats are often shown in icons of Annunciation and of the Holy Family and, according to Italian folklore, on the same night that Mary gave birth to Jesus, a cat in Bethlehem gave birth to a kitten. Domestic cats were spread throughout much of the rest of the world during the Age of Discovery, as ships' cats were carried on sailing ships to control shipboard rodents and as good-luck charms.

 

Several ancient religions believed cats are exalted souls, companions or guides for humans, that are all-knowing but mute so they cannot influence decisions made by humans. In Japan, the maneki neko cat is a symbol of good fortune. In Norse mythology, Freyja, the goddess of love, beauty, and fertility, is depicted as riding a chariot drawn by cats. In Jewish legend, the first cat was living in the house of the first man Adam as a pet that got rid of mice. The cat was once partnering with the first dog before the latter broke an oath they had made which resulted in enmity between the descendants of these two animals. It is also written that neither cats nor foxes are represented in the water, while every other animal has an incarnation species in the water. Although no species are sacred in Islam, cats are revered by Muslims. Some Western writers have stated Muhammad had a favorite cat, Muezza. He is reported to have loved cats so much, "he would do without his cloak rather than disturb one that was sleeping on it". The story has no origin in early Muslim writers, and seems to confuse a story of a later Sufi saint, Ahmed ar-Rifa'i, centuries after Muhammad. One of the companions of Muhammad was known as Abu Hurayrah ("father of the kitten"), in reference to his documented affection to cats.

 

Superstitions and rituals

Many cultures have negative superstitions about cats. An example would be the belief that encountering a black cat ("crossing one's path") leads to bad luck, or that cats are witches' familiars used to augment a witch's powers and skills. The killing of cats in Medieval Ypres, Belgium, is commemorated in the innocuous present-day Kattenstoet (cat parade). In mid-16th century France, cats would be burnt alive as a form of entertainment, particularly during midsummer festivals. According to Norman Davies, the assembled people "shrieked with laughter as the animals, howling with pain, were singed, roasted, and finally carbonized". The remaining ashes were sometimes taken back home by the people for good luck.

 

According to a myth in many cultures, cats have multiple lives. In many countries, they are believed to have nine lives, but in Italy, Germany, Greece, Brazil and some Spanish-speaking regions, they are said to have seven lives, while in Arabic traditions, the number of lives is six. An early mention of the myth can be found in John Heywood's The Proverbs of John Heywood (1546)

 

Husband, (quoth she), ye studie, be merrie now,

And even as ye thinke now, so come to yow.

Nay not so, (quoth he), for my thought to tell right,

I thinke how you lay groning, wife, all last night.

Husband, a groning horse and a groning wife

Never faile their master, (quoth she), for my life.

No wife, a woman hath nine lives like a cat.

 

The myth is attributed to the natural suppleness and swiftness cats exhibit to escape life-threatening situations. Also lending credence to this myth is the fact that falling cats often land on their feet, using an instinctive righting reflex to twist their bodies around. Nonetheless, cats can still be injured or killed by a high fall.

I really wish I were uncut--I was born at the absolute height of America's circumcision craze...roughly 95%. Now it's down to roughly half.

 

It's a trickier decision now, since it's pretty clear that the Langerhans cells in foreskin promote HIV infection in uncut men. If I'd been uncut, I might be dead, though I came out late enough that I've had very little unsafe sex.

 

Growing up, any time the topic came up, my dad would just screw up his face in disgust and say, "yucckkkk." In the ultra-sanitary 1960s suburban world, being uncircumcised was synonymous with smegma...I think we all assumed uncut weenies were just filthy 24x7.

 

I also never knew (until recently) that 2/3 of the penile nerve endings are in the foreskin, nor did I understand how perfect the mechanics of foreskin rolling/unrolling is for sexual gratification. It's an amazing piece of engineering. I've never known an uncut guy to need to jackhammer his johnson to get off like a lot of cut guys do. No wonder... they have 3 times as many pleasure nerve endings all rubbing against each other.

 

For a while I tried the "restoration" techniques...even though you don't get the nerve endings back, there is some benefit to having the coverage and the gliding action. But it's a real hassle, and I threw in the towel.

 

That said, I still have a huge fetish for natural men, as well as a huge dose of envy. Like Todd's bodybuilder collection,

www.flickr.com/photos/tlcub4bear/676209427/in/set-7215760...

I have a huge collection of foreskin photos. It's less about porn and more about my sheer admiration of their beauty.

this art is called 'chabi bharna' where he rolls up his penis to a stick and swirls the stick around to show his penile prowess ;)

sorry my black tea

as facebook update

on your laid back

gaming gracelessness

I did spill

nothing personal

I blog When

I have Time

to Kill

At Facebook

I am definitely

not into groups

events or Fucked

Farmville

I am a blogger

with god given

creativity

some skill

unlike jack

I dont hump jill

a pail of water

to make love

on mountain

of a molehill

dharma

karma

as god wills

I dont need Viagra

or penile enlargement pills

I blog freely

wthout selling

my pictures

for dollar bills

a pedestrian poet

my earthly

bank balance

totally nil

a hard core

hindustani

within her

heart beat

hai mera dil

  

Good Luck Irani Hotel

Near Mehboob Studio

Many years back after we left our home at De Monte street , with our eldest son on the verge of getting married we came to stay on rent at Pitru Chayya next to Mehboob Studio, we stayed here for about 11 months and Good Luck became my breakfast joint , I came here to have keema brun , black tea ..and I was just a wall away from Mehboob Studio..We left this place soon after to stay another few months in a rustic Bandra Back Lane on rent a place called White House.. finally 10 years back we came to our present residence at Bandra Reclamation thanks to the support from friends.. this was our Good Luck home ,, and we never forgot Good Luck Irani Restaurant

pearlypenilepapulestube.com/pearly-penile-papules-removal...

pearlypenilepapulestube.com/pearly-penile-papule-removal-2/

pearlypenilepapulestube.com/pearly-penile-papules-home-tr...

The cat (Felis catus), commonly referred to as the domestic cat or house cat, is the only domesticated species in the family Felidae. Recent advances in archaeology and genetics have shown that the domestication of the cat occurred in the Near East around 7500 BC. It is commonly kept as a house pet and farm cat, but also ranges freely as a feral cat avoiding human contact. It is valued by humans for companionship and its ability to kill vermin. Because of its retractable claws it is adapted to killing small prey like mice and rats. It has a strong flexible body, quick reflexes, sharp teeth, and its night vision and sense of smell are well developed. It is a social species, but a solitary hunter and a crepuscular predator. Cat communication includes vocalizations like meowing, purring, trilling, hissing, growling, and grunting as well as cat body language. It can hear sounds too faint or too high in frequency for human ears, such as those made by small mammals. It also secretes and perceives pheromones.

 

Female domestic cats can have kittens from spring to late autumn in temperate zones and throughout the year in equatorial regions, with litter sizes often ranging from two to five kittens. Domestic cats are bred and shown at events as registered pedigreed cats, a hobby known as cat fancy. Animal population control of cats may be achieved by spaying and neutering, but their proliferation and the abandonment of pets has resulted in large numbers of feral cats worldwide, contributing to the extinction of bird, mammal and reptile species.

 

As of 2017, the domestic cat was the second most popular pet in the United States, with 95.6 million cats owned and around 42 million households owning at least one cat. In the United Kingdom, 26% of adults have a cat, with an estimated population of 10.9 million pet cats as of 2020. As of 2021, there were an estimated 220 million owned and 480 million stray cats in the world.

 

Etymology and naming

The origin of the English word cat, Old English catt, is thought to be the Late Latin word cattus, which was first used at the beginning of the 6th century. The Late Latin word may be derived from an unidentified African language. The Nubian word kaddîska 'wildcat' and Nobiin kadīs are possible sources or cognates. The Nubian word may be a loan from Arabic قَطّ‎ qaṭṭ ~ قِطّ qiṭṭ.

 

The forms might also have derived from an ancient Germanic word that was imported into Latin and then into Greek, Syriac, and Arabic. The word may be derived from Germanic and Northern European languages, and ultimately be borrowed from Uralic, cf. Northern Sámi gáđfi, 'female stoat', and Hungarian hölgy, 'lady, female stoat'; from Proto-Uralic *käďwä, 'female (of a furred animal)'.

 

The English puss, extended as pussy and pussycat, is attested from the 16th century and may have been introduced from Dutch poes or from Low German puuskatte, related to Swedish kattepus, or Norwegian pus, pusekatt. Similar forms exist in Lithuanian puižė and Irish puisín or puiscín. The etymology of this word is unknown, but it may have arisen from a sound used to attract a cat.

 

A male cat is called a tom or tomcat (or a gib, if neutered). A female is called a queen or a molly, if spayed, especially in a cat-breeding context. A juvenile cat is referred to as a kitten. In Early Modern English, the word kitten was interchangeable with the now-obsolete word catling.

 

A group of cats can be referred to as a clowder or a glaring.

 

Taxonomy

The scientific name Felis catus was proposed by Carl Linnaeus in 1758 for a domestic cat. Felis catus domesticus was proposed by Johann Christian Polycarp Erxleben in 1777. Felis daemon proposed by Konstantin Satunin in 1904 was a black cat from the Transcaucasus, later identified as a domestic cat.

 

In 2003, the International Commission on Zoological Nomenclature ruled that the domestic cat is a distinct species, namely Felis catus. In 2007, it was considered a subspecies, F. silvestris catus, of the European wildcat (F. silvestris) following results of phylogenetic research. In 2017, the IUCN Cat Classification Taskforce followed the recommendation of the ICZN in regarding the domestic cat as a distinct species, Felis catus.

 

Evolution

Main article: Cat evolution

The domestic cat is a member of the Felidae, a family that had a common ancestor about 10 to 15 million years ago. The evolutionary radiation of the Felidae began in Asia during the Miocene around 8.38 to 14.45 million years ago. Analysis of mitochondrial DNA of all Felidae species indicates a radiation at 6.46 to 16.76 million years ago. The genus Felis genetically diverged from other Felidae around 6 to 7 million years ago. Results of phylogenetic research shows that the wild members of this genus evolved through sympatric or parapatric speciation, whereas the domestic cat evolved through artificial selection. The domestic cat and its closest wild ancestor are diploid and both possess 38 chromosomes and roughly 20,000 genes.

 

Domestication

See also: Domestication of the cat and Cats in ancient Egypt

It was long thought that the domestication of the cat began in ancient Egypt, where cats were venerated from around 3100 BC, However, the earliest known indication for the taming of an African wildcat was excavated close by a human Neolithic grave in Shillourokambos, southern Cyprus, dating to about 7500–7200 BC. Since there is no evidence of native mammalian fauna on Cyprus, the inhabitants of this Neolithic village most likely brought the cat and other wild mammals to the island from the Middle Eastern mainland. Scientists therefore assume that African wildcats were attracted to early human settlements in the Fertile Crescent by rodents, in particular the house mouse (Mus musculus), and were tamed by Neolithic farmers. This mutual relationship between early farmers and tamed cats lasted thousands of years. As agricultural practices spread, so did tame and domesticated cats. Wildcats of Egypt contributed to the maternal gene pool of the domestic cat at a later time.

The earliest known evidence for the occurrence of the domestic cat in Greece dates to around 1200 BC. Greek, Phoenician, Carthaginian and Etruscan traders introduced domestic cats to southern Europe. During the Roman Empire they were introduced to Corsica and Sardinia before the beginning of the 1st millennium. By the 5th century BC, they were familiar animals around settlements in Magna Graecia and Etruria. By the end of the Western Roman Empire in the 5th century, the Egyptian domestic cat lineage had arrived in a Baltic Sea port in northern Germany.

 

The leopard cat (Prionailurus bengalensis) was tamed independently in China around 5500 BC. This line of partially domesticated cats leaves no trace in the domestic cat populations of today.

 

During domestication, cats have undergone only minor changes in anatomy and behavior, and they are still capable of surviving in the wild. Several natural behaviors and characteristics of wildcats may have pre-adapted them for domestication as pets. These traits include their small size, social nature, obvious body language, love of play, and high intelligence. Captive Leopardus cats may also display affectionate behavior toward humans but were not domesticated. House cats often mate with feral cats. Hybridisation between domestic and other Felinae species is also possible, producing hybrids such as the Kellas cat in Scotland.

 

Development of cat breeds started in the mid 19th century. An analysis of the domestic cat genome revealed that the ancestral wildcat genome was significantly altered in the process of domestication, as specific mutations were selected to develop cat breeds. Most breeds are founded on random-bred domestic cats. Genetic diversity of these breeds varies between regions, and is lowest in purebred populations, which show more than 20 deleterious genetic disorders.

 

Characteristics

Main article: Cat anatomy

Size

The domestic cat has a smaller skull and shorter bones than the European wildcat. It averages about 46 cm (18 in) in head-to-body length and 23–25 cm (9.1–9.8 in) in height, with about 30 cm (12 in) long tails. Males are larger than females. Adult domestic cats typically weigh 4–5 kg (8.8–11.0 lb).

 

Skeleton

Cats have seven cervical vertebrae (as do most mammals); 13 thoracic vertebrae (humans have 12); seven lumbar vertebrae (humans have five); three sacral vertebrae (as do most mammals, but humans have five); and a variable number of caudal vertebrae in the tail (humans have only three to five vestigial caudal vertebrae, fused into an internal coccyx).  The extra lumbar and thoracic vertebrae account for the cat's spinal mobility and flexibility. Attached to the spine are 13 ribs, the shoulder, and the pelvis.  Unlike human arms, cat forelimbs are attached to the shoulder by free-floating clavicle bones which allow them to pass their body through any space into which they can fit their head.

 

Skull

The cat skull is unusual among mammals in having very large eye sockets and a powerful specialized jaw.  Within the jaw, cats have teeth adapted for killing prey and tearing meat. When it overpowers its prey, a cat delivers a lethal neck bite with its two long canine teeth, inserting them between two of the prey's vertebrae and severing its spinal cord, causing irreversible paralysis and death. Compared to other felines, domestic cats have narrowly spaced canine teeth relative to the size of their jaw, which is an adaptation to their preferred prey of small rodents, which have small vertebrae.

 

The premolar and first molar together compose the carnassial pair on each side of the mouth, which efficiently shears meat into small pieces, like a pair of scissors. These are vital in feeding, since cats' small molars cannot chew food effectively, and cats are largely incapable of mastication.:  Cats tend to have better teeth than most humans, with decay generally less likely because of a thicker protective layer of enamel, a less damaging saliva, less retention of food particles between teeth, and a diet mostly devoid of sugar. Nonetheless, they are subject to occasional tooth loss and infection.

 

Claws

Cats have protractible and retractable claws. In their normal, relaxed position, the claws are sheathed with the skin and fur around the paw's toe pads. This keeps the claws sharp by preventing wear from contact with the ground and allows for the silent stalking of prey. The claws on the forefeet are typically sharper than those on the hindfeet. Cats can voluntarily extend their claws on one or more paws. They may extend their claws in hunting or self-defense, climbing, kneading, or for extra traction on soft surfaces. Cats shed the outside layer of their claw sheaths when scratching rough surfaces.

 

Most cats have five claws on their front paws and four on their rear paws. The dewclaw is proximal to the other claws. More proximally is a protrusion which appears to be a sixth "finger". This special feature of the front paws on the inside of the wrists has no function in normal walking but is thought to be an antiskidding device used while jumping. Some cat breeds are prone to having extra digits ("polydactyly"). Polydactylous cats occur along North America's northeast coast and in Great Britain.

 

Ambulation

The cat is digitigrade. It walks on the toes, with the bones of the feet making up the lower part of the visible leg. Unlike most mammals, it uses a "pacing" gait and moves both legs on one side of the body before the legs on the other side. It registers directly by placing each hind paw close to the track of the corresponding fore paw, minimizing noise and visible tracks. This also provides sure footing for hind paws when navigating rough terrain. As it speeds up from walking to trotting, its gait changes to a "diagonal" gait: The diagonally opposite hind and fore legs move simultaneously.

 

Balance

Cats are generally fond of sitting in high places or perching. A higher place may serve as a concealed site from which to hunt; domestic cats strike prey by pouncing from a perch such as a tree branch. Another possible explanation is that height gives the cat a better observation point, allowing it to survey its territory. A cat falling from heights of up to 3 m (9.8 ft) can right itself and land on its paws.

 

During a fall from a high place, a cat reflexively twists its body and rights itself to land on its feet using its acute sense of balance and flexibility. This reflex is known as the cat righting reflex. A cat always rights itself in the same way during a fall, if it has enough time to do so, which is the case in falls of 90 cm (3.0 ft) or more. How cats are able to right themselves when falling has been investigated as the "falling cat problem".

 

Coats

Main article: Cat coat genetics

The cat family (Felidae) can pass down many colors and patterns to their offspring. The domestic cat genes MC1R and ASIP allow for the variety of color in coats. The feline ASIP gene consists of three coding exons. Three novel microsatellite markers linked to ASIP were isolated from a domestic cat BAC clone containing this gene and were used to perform linkage analysis in a pedigree of 89 domestic cats that segregated for melanism.[citation needed]

 

Senses

Main article: Cat senses

Vision

A cat's nictitating membrane shown as it blinks

Cats have excellent night vision and can see at only one-sixth the light level required for human vision.  This is partly the result of cat eyes having a tapetum lucidum, which reflects any light that passes through the retina back into the eye, thereby increasing the eye's sensitivity to dim light. Large pupils are an adaptation to dim light. The domestic cat has slit pupils, which allow it to focus bright light without chromatic aberration. At low light, a cat's pupils expand to cover most of the exposed surface of its eyes. The domestic cat has rather poor color vision and only two types of cone cells, optimized for sensitivity to blue and yellowish green; its ability to distinguish between red and green is limited. A response to middle wavelengths from a system other than the rod cells might be due to a third type of cone. This appears to be an adaptation to low light levels rather than representing true trichromatic vision. Cats also have a nictitating membrane, allowing them to blink without hindering their vision.

 

Hearing

The domestic cat's hearing is most acute in the range of 500 Hz to 32 kHz. It can detect an extremely broad range of frequencies ranging from 55 Hz to 79 kHz, whereas humans can only detect frequencies between 20 Hz and 20 kHz. It can hear a range of 10.5 octaves, while humans and dogs can hear ranges of about 9 octaves. Its hearing sensitivity is enhanced by its large movable outer ears, the pinnae, which amplify sounds and help detect the location of a noise. It can detect ultrasound, which enables it to detect ultrasonic calls made by rodent prey. Recent research has shown that cats have socio-spatial cognitive abilities to create mental maps of owners' locations based on hearing owners' voices.

 

Smell

Cats have an acute sense of smell, due in part to their well-developed olfactory bulb and a large surface of olfactory mucosa, about 5.8 cm2 (0.90 in2) in area, which is about twice that of humans. Cats and many other animals have a Jacobson's organ in their mouths that is used in the behavioral process of flehmening. It allows them to sense certain aromas in a way that humans cannot. Cats are sensitive to pheromones such as 3-mercapto-3-methylbutan-1-ol, which they use to communicate through urine spraying and marking with scent glands. Many cats also respond strongly to plants that contain nepetalactone, especially catnip, as they can detect that substance at less than one part per billion. About 70–80% of cats are affected by nepetalactone. This response is also produced by other plants, such as silver vine (Actinidia polygama) and the herb valerian; it may be caused by the smell of these plants mimicking a pheromone and stimulating cats' social or sexual behaviors.

 

Taste

Cats have relatively few taste buds compared to humans (470 or so versus more than 9,000 on the human tongue). Domestic and wild cats share a taste receptor gene mutation that keeps their sweet taste buds from binding to sugary molecules, leaving them with no ability to taste sweetness. They, however, possess taste bud receptors specialized for acids, amino acids like protein, and bitter tastes. Their taste buds possess the receptors needed to detect umami. However, these receptors contain molecular changes that make the cat taste of umami different from that of humans. In humans, they detect the amino acids of glutamic acid and aspartic acid, but in cats they instead detect nucleotides, in this case inosine monophosphate and l-Histidine. These nucleotides are particularly enriched in tuna. This has been argued is why cats find tuna so palatable: as put by researchers into cat taste, "the specific combination of the high IMP and free l-Histidine contents of tuna" .. "produces a strong umami taste synergy that is highly preferred by cats". One of the researchers involved in this research has further claimed, "I think umami is as important for cats as sweet is for humans".[87]

 

Cats also have a distinct temperature preference for their food, preferring food with a temperature around 38 °C (100 °F) which is similar to that of a fresh kill; some cats reject cold food (which would signal to the cat that the "prey" item is long dead and therefore possibly toxic or decomposing).

 

Whiskers

To aid with navigation and sensation, cats have dozens of movable whiskers (vibrissae) over their body, especially their faces. These provide information on the width of gaps and on the location of objects in the dark, both by touching objects directly and by sensing air currents; they also trigger protective blink reflexes to protect the eyes from damage.: 47 

 

Behavior

See also: Cat behavior

Outdoor cats are active both day and night, although they tend to be slightly more active at night.[88] Domestic cats spend the majority of their time in the vicinity of their homes but can range many hundreds of meters from this central point. They establish territories that vary considerably in size, in one study ranging 7–28 ha (17–69 acres). The timing of cats' activity is quite flexible and varied but being low-light predators, they are generally crepuscular, which means they tend to be more active near dawn and dusk. However, house cats' behavior is also influenced by human activity and they may adapt to their owners' sleeping patterns to some extent.

 

Cats conserve energy by sleeping more than most animals, especially as they grow older. The daily duration of sleep varies, usually between 12 and 16 hours, with 13 and 14 being the average. Some cats can sleep as much as 20 hours. The term "cat nap" for a short rest refers to the cat's tendency to fall asleep (lightly) for a brief period. While asleep, cats experience short periods of rapid eye movement sleep often accompanied by muscle twitches, which suggests they are dreaming.

 

Sociability

The social behavior of the domestic cat ranges from widely dispersed individuals to feral cat colonies that gather around a food source, based on groups of co-operating females. Within such groups, one cat is usually dominant over the others. Each cat in a colony holds a distinct territory, with sexually active males having the largest territories, which are about 10 times larger than those of female cats and may overlap with several females' territories. These territories are marked by urine spraying, by rubbing objects at head height with secretions from facial glands, and by defecation. Between these territories are neutral areas where cats watch and greet one another without territorial conflicts. Outside these neutral areas, territory holders usually chase away stranger cats, at first by staring, hissing, and growling and, if that does not work, by short but noisy and violent attacks. Despite this colonial organization, cats do not have a social survival strategy or a herd behavior, and always hunt alone.

 

Life in proximity to humans and other domestic animals has led to a symbiotic social adaptation in cats, and cats may express great affection toward humans or other animals. Ethologically, a cat's human keeper functions as if a mother surrogate. Adult cats live their lives in a kind of extended kittenhood, a form of behavioral neoteny. Their high-pitched sounds may mimic the cries of a hungry human infant, making them particularly difficult for humans to ignore. Some pet cats are poorly socialized. In particular, older cats show aggressiveness toward newly arrived kittens, which include biting and scratching; this type of behavior is known as feline asocial aggression.

 

Redirected aggression is a common form of aggression which can occur in multiple cat households. In redirected aggression there is usually something that agitates the cat: this could be a sight, sound, or another source of stimuli which causes a heightened level of anxiety or arousal. If the cat cannot attack the stimuli, it may direct anger elsewhere by attacking or directing aggression to the nearest cat, dog, human or other being.

 

Domestic cats' scent rubbing behavior toward humans or other cats is thought to be a feline means for social bonding.

 

Communication

Main article: Cat communication

Domestic cats use many vocalizations for communication, including purring, trilling, hissing, growling/snarling, grunting, and several different forms of meowing. Their body language, including position of ears and tail, relaxation of the whole body, and kneading of the paws, are all indicators of mood. The tail and ears are particularly important social signal mechanisms in cats. A raised tail indicates a friendly greeting, and flattened ears indicate hostility. Tail-raising also indicates the cat's position in the group's social hierarchy, with dominant individuals raising their tails less often than subordinate ones. Feral cats are generally silent.: 208  Nose-to-nose touching is also a common greeting and may be followed by social grooming, which is solicited by one of the cats raising and tilting its head.

 

Purring may have developed as an evolutionary advantage as a signaling mechanism of reassurance between mother cats and nursing kittens, who are thought to use it as a care-soliciting signal. Post-nursing cats also often purr as a sign of contentment: when being petted, becoming relaxed, or eating. Even though purring is popularly interpreted as indicative of pleasure, it has been recorded in a wide variety of circumstances, most of which involve physical contact between the cat and another, presumably trusted individual. Some cats have been observed to purr continuously when chronically ill or in apparent pain.

 

The exact mechanism by which cats purr has long been elusive, but it has been proposed that purring is generated via a series of sudden build-ups and releases of pressure as the glottis is opened and closed, which causes the vocal folds to separate forcefully. The laryngeal muscles in control of the glottis are thought to be driven by a neural oscillator which generates a cycle of contraction and release every 30–40 milliseconds (giving a frequency of 33 to 25 Hz).

 

Domestic cats observed in a rescue facility have total of 276 distinct facial expressions based on 26 different facial movements; each facial expression corresponds to different social functions that are likely influenced by domestication.

 

Grooming

Cats are known for spending considerable amounts of time licking their coats to keep them clean. The cat's tongue has backward-facing spines about 500 μm long, which are called papillae. These contain keratin which makes them rigid so the papillae act like a hairbrush. Some cats, particularly longhaired cats, occasionally regurgitate hairballs of fur that have collected in their stomachs from grooming. These clumps of fur are usually sausage-shaped and about 2–3 cm (0.79–1.18 in) long. Hairballs can be prevented with remedies that ease elimination of the hair through the gut, as well as regular grooming of the coat with a comb or stiff brush.

 

Fighting

Among domestic cats, males are more likely to fight than females. Among feral cats, the most common reason for cat fighting is competition between two males to mate with a female. In such cases, most fights are won by the heavier male. Another common reason for fighting in domestic cats is the difficulty of establishing territories within a small home. Female cats also fight over territory or to defend their kittens. Neutering will decrease or eliminate this behavior in many cases, suggesting that the behavior is linked to sex hormones.

 

When cats become aggressive, they try to make themselves appear larger and more threatening by raising their fur, arching their backs, turning sideways and hissing or spitting. Often, the ears are pointed down and back to avoid damage to the inner ear and potentially listen for any changes behind them while focused forward. Cats may also vocalize loudly and bare their teeth in an effort to further intimidate their opponents. Fights usually consist of grappling and delivering powerful slaps to the face and body with the forepaws as well as bites. Cats also throw themselves to the ground in a defensive posture to rake their opponent's belly with their powerful hind legs.

 

Serious damage is rare, as the fights are usually short in duration, with the loser running away with little more than a few scratches to the face and ears. Fights for mating rights are typically more severe and injuries may include deep puncture wounds and lacerations. Normally, serious injuries from fighting are limited to infections of scratches and bites, though these can occasionally kill cats if untreated. In addition, bites are probably the main route of transmission of feline immunodeficiency virus. Sexually active males are usually involved in many fights during their lives, and often have decidedly battered faces with obvious scars and cuts to their ears and nose. Cats are willing to threaten animals larger than them to defend their territory, such as dogs and foxes.

 

Hunting and feeding

See also: Cat food

The shape and structure of cats' cheeks is insufficient to allow them to take in liquids using suction. Therefore, when drinking they lap with the tongue to draw liquid upward into their mouths. Lapping at a rate of four times a second, the cat touches the smooth tip of its tongue to the surface of the water, and quickly retracts it like a corkscrew, drawing water upward.

 

Feral cats and free-fed house cats consume several small meals in a day. The frequency and size of meals varies between individuals. They select food based on its temperature, smell and texture; they dislike chilled foods and respond most strongly to moist foods rich in amino acids, which are similar to meat. Cats reject novel flavors (a response termed neophobia) and learn quickly to avoid foods that have tasted unpleasant in the past. It is also a common misconception that cats like milk/cream, as they tend to avoid sweet food and milk. Most adult cats are lactose intolerant; the sugar in milk is not easily digested and may cause soft stools or diarrhea. Some also develop odd eating habits and like to eat or chew on things like wool, plastic, cables, paper, string, aluminum foil, or even coal. This condition, pica, can threaten their health, depending on the amount and toxicity of the items eaten.

 

Cats hunt small prey, primarily birds and rodents, and are often used as a form of pest control. Other common small creatures such as lizards and snakes may also become prey. Cats use two hunting strategies, either stalking prey actively, or waiting in ambush until an animal comes close enough to be captured. The strategy used depends on the prey species in the area, with cats waiting in ambush outside burrows, but tending to actively stalk birds.: 153  Domestic cats are a major predator of wildlife in the United States, killing an estimated 1.3 to 4.0 billion birds and 6.3 to 22.3 billion mammals annually.

 

Certain species appear more susceptible than others; in one English village, for example, 30% of house sparrow mortality was linked to the domestic cat. In the recovery of ringed robins (Erithacus rubecula) and dunnocks (Prunella modularis) in Britain, 31% of deaths were a result of cat predation. In parts of North America, the presence of larger carnivores such as coyotes which prey on cats and other small predators reduces the effect of predation by cats and other small predators such as opossums and raccoons on bird numbers and variety.

 

Perhaps the best-known element of cats' hunting behavior, which is commonly misunderstood and often appalls cat owners because it looks like torture, is that cats often appear to "play" with prey by releasing and recapturing it. This cat and mouse behavior is due to an instinctive imperative to ensure that the prey is weak enough to be killed without endangering the cat.

 

Another poorly understood element of cat hunting behavior is the presentation of prey to human guardians. One explanation is that cats adopt humans into their social group and share excess kill with others in the group according to the dominance hierarchy, in which humans are reacted to as if they are at or near the top. Another explanation is that they attempt to teach their guardians to hunt or to help their human as if feeding "an elderly cat, or an inept kitten". This hypothesis is inconsistent with the fact that male cats also bring home prey, despite males having negligible involvement in raising kittens.:

 

Play

Main article: Cat play and toys

Domestic cats, especially young kittens, are known for their love of play. This behavior mimics hunting and is important in helping kittens learn to stalk, capture, and kill prey. Cats also engage in play fighting, with each other and with humans. This behavior may be a way for cats to practice the skills needed for real combat, and might also reduce any fear they associate with launching attacks on other animals.

 

Cats also tend to play with toys more when they are hungry. Owing to the close similarity between play and hunting, cats prefer to play with objects that resemble prey, such as small furry toys that move rapidly, but rapidly lose interest. They become habituated to a toy they have played with before. String is often used as a toy, but if it is eaten, it can become caught at the base of the cat's tongue and then move into the intestines, a medical emergency which can cause serious illness, even death. Owing to the risks posed by cats eating string, it is sometimes replaced with a laser pointer's dot, which cats may chase.

 

Reproduction

See also: Kitten

The cat secretes and perceives pheromones. Female cats, called queens, are polyestrous with several estrus cycles during a year, lasting usually 21 days. They are usually ready to mate between early February and August in northern temperate zones and throughout the year in equatorial regions.

 

Several males, called tomcats, are attracted to a female in heat. They fight over her, and the victor wins the right to mate. At first, the female rejects the male, but eventually, the female allows the male to mate. The female utters a loud yowl as the male pulls out of her because a male cat's penis has a band of about 120–150 backward-pointing penile spines, which are about 1 mm (0.039 in) long; upon withdrawal of the penis, the spines may provide the female with increased sexual stimulation, which acts to induce ovulation.

 

After mating, the female cleans her vulva thoroughly. If a male attempts to mate with her at this point, the female attacks him. After about 20 to 30 minutes, once the female is finished grooming, the cycle will repeat. Because ovulation is not always triggered by a single mating, females may not be impregnated by the first male with which they mate. Furthermore, cats are superfecund; that is, a female may mate with more than one male when she is in heat, with the result that different kittens in a litter may have different fathers.

 

The morula forms 124 hours after conception. At 148 hours, early blastocysts form. At 10–12 days, implantation occurs. The gestation of queens lasts between 64 and 67 days, with an average of 65 days.

 

Data on the reproductive capacity of more than 2,300 free-ranging queens were collected during a study between May 1998 and October 2000. They had one to six kittens per litter, with an average of three kittens. They produced a mean of 1.4 litters per year, but a maximum of three litters in a year. Of 169 kittens, 127 died before they were six months old due to a trauma caused in most cases by dog attacks and road accidents. The first litter is usually smaller than subsequent litters. Kittens are weaned between six and seven weeks of age. Queens normally reach sexual maturity at 5–10 months, and males at 5–7 months. This varies depending on breed. Kittens reach puberty at the age of 9–10 months.

 

Cats are ready to go to new homes at about 12 weeks of age, when they are ready to leave their mother. They can be surgically sterilized (spayed or castrated) as early as seven weeks to limit unwanted reproduction. This surgery also prevents undesirable sex-related behavior, such as aggression, territory marking (spraying urine) in males and yowling (calling) in females. Traditionally, this surgery was performed at around six to nine months of age, but it is increasingly being performed before puberty, at about three to six months. In the United States, about 80% of household cats are neutered.

 

Lifespan and health

Main articles: Cat health and Aging in cats

The average lifespan of pet cats has risen in recent decades. In the early 1980s, it was about seven years,: 33  rising to 9.4 years in 1995: 33  and an average of about 13 years as of 2014 and 2023. Some cats have been reported as surviving into their 30s, with the oldest known cat dying at a verified age of 38.

 

Neutering increases life expectancy: one study found castrated male cats live twice as long as intact males, while spayed female cats live 62% longer than intact females.: 35  Having a cat neutered confers health benefits, because castrated males cannot develop testicular cancer, spayed females cannot develop uterine or ovarian cancer, and both have a reduced risk of mammary cancer.

 

Disease

Main article: List of feline diseases

About 250 heritable genetic disorders have been identified in cats, many similar to human inborn errors of metabolism. The high level of similarity among the metabolism of mammals allows many of these feline diseases to be diagnosed using genetic tests that were originally developed for use in humans, as well as the use of cats as animal models in the study of the human diseases. Diseases affecting domestic cats include acute infections, parasitic infestations, injuries, and chronic diseases such as kidney disease, thyroid disease, and arthritis. Vaccinations are available for many infectious diseases, as are treatments to eliminate parasites such as worms, ticks, and fleas.

 

Ecology

Habitats

The domestic cat is a cosmopolitan species and occurs across much of the world. It is adaptable and now present on all continents except Antarctica, and on 118 of the 131 main groups of islands, even on the isolated Kerguelen Islands. Due to its ability to thrive in almost any terrestrial habitat, it is among the world's most invasive species. It lives on small islands with no human inhabitants. Feral cats can live in forests, grasslands, tundra, coastal areas, agricultural land, scrublands, urban areas, and wetlands.

 

The unwantedness that leads to the domestic cat being treated as an invasive species is twofold. On one hand, as it is little altered from the wildcat, it can readily interbreed with the wildcat. This hybridization poses a danger to the genetic distinctiveness of some wildcat populations, particularly in Scotland and Hungary, possibly also the Iberian Peninsula, and where protected natural areas are close to human-dominated landscapes, such as Kruger National Park in South Africa. However, its introduction to places where no native felines are present also contributes to the decline of native species.

 

Ferality

Main article: Feral cat

Feral cats are domestic cats that were born in or have reverted to a wild state. They are unfamiliar with and wary of humans and roam freely in urban and rural areas. The numbers of feral cats is not known, but estimates of the United States feral population range from 25 to 60 million. Feral cats may live alone, but most are found in large colonies, which occupy a specific territory and are usually associated with a source of food. Famous feral cat colonies are found in Rome around the Colosseum and Forum Romanum, with cats at some of these sites being fed and given medical attention by volunteers.

 

Public attitudes toward feral cats vary widely, from seeing them as free-ranging pets to regarding them as vermin.

 

Some feral cats can be successfully socialized and 're-tamed' for adoption; young cats, especially kittens and cats that have had prior experience and contact with humans are the most receptive to these efforts.

 

Impact on wildlife

Main article: Cat predation on wildlife

On islands, birds can contribute as much as 60% of a cat's diet. In nearly all cases, the cat cannot be identified as the sole cause for reducing the numbers of island birds, and in some instances, eradication of cats has caused a "mesopredator release" effect; where the suppression of top carnivores creates an abundance of smaller predators that cause a severe decline in their shared prey. Domestic cats are a contributing factor to the decline of many species, a factor that has ultimately led, in some cases, to extinction. The South Island piopio, Chatham rail, and the New Zealand merganser are a few from a long list, with the most extreme case being the flightless Lyall's wren, which was driven to extinction only a few years after its discovery. One feral cat in New Zealand killed 102 New Zealand lesser short-tailed bats in seven days. In the US, feral and free-ranging domestic cats kill an estimated 6.3 – 22.3 billion mammals annually.

 

In Australia, the impact of cats on mammal populations is even greater than the impact of habitat loss. More than one million reptiles are killed by feral cats each day, representing 258 species. Cats have contributed to the extinction of the Navassa curly-tailed lizard and Chioninia coctei.

 

Interaction with humans

Main article: Human interaction with cats

Cats are common pets throughout the world, and their worldwide population as of 2007 exceeded 500 million. As of 2017, the domestic cat was the second most popular pet in the United States, with 95.6 million cats owned and around 42 million households owning at least one cat. In the United Kingdom, 26% of adults have a cat, with an estimated population of 10.9 million pet cats as of 2020. As of 2021, there were an estimated 220 million owned and 480 million stray cats in the world.

 

Cats have been used for millennia to control rodents, notably around grain stores and aboard ships, and both uses extend to the present day.

 

As well as being kept as pets, cats are also used in the international fur trade and leather industries for making coats, hats, blankets, stuffed toys, shoes, gloves, and musical instruments. About 24 cats are needed to make a cat-fur coat. This use has been outlawed in the United States since 2000 and in the European Union (as well as the United Kingdom) since 2007.

 

Cat pelts have been used for superstitious purposes as part of the practice of witchcraft, and are still made into blankets in Switzerland as traditional medicine thought to cure rheumatism.

 

A few attempts to build a cat census have been made over the years, both through associations or national and international organizations (such as that of the Canadian Federation of Humane Societies) and over the Internet, but such a task does not seem simple to achieve. General estimates for the global population of domestic cats range widely from anywhere between 200 million to 600 million. Walter Chandoha made his career photographing cats after his 1949 images of Loco, an especially charming stray taken in, were published around the world. He is reported to have photographed 90,000 cats during his career and maintained an archive of 225,000 images that he drew from for publications during his lifetime.

 

Shows

Main article: Cat show

A cat show is a judged event in which the owners of cats compete to win titles in various cat-registering organizations by entering their cats to be judged after a breed standard. It is often required that a cat must be healthy and vaccinated in order to participate in a cat show. Both pedigreed and non-purebred companion ("moggy") cats are admissible, although the rules differ depending on the organization. Competing cats are compared to the applicable breed standard, and assessed for temperament.

 

Infection

Main article: Feline zoonosis

Cats can be infected or infested with viruses, bacteria, fungus, protozoans, arthropods or worms that can transmit diseases to humans. In some cases, the cat exhibits no symptoms of the disease. The same disease can then become evident in a human. The likelihood that a person will become diseased depends on the age and immune status of the person. Humans who have cats living in their home or in close association are more likely to become infected. Others might also acquire infections from cat feces and parasites exiting the cat's body. Some of the infections of most concern include salmonella, cat-scratch disease and toxoplasmosis.

 

History and mythology

Main articles: Cultural depictions of cats and Cats in ancient Egypt

In ancient Egypt, cats were worshipped, and the goddess Bastet often depicted in cat form, sometimes taking on the war-like aspect of a lioness. The Greek historian Herodotus reported that killing a cat was forbidden, and when a household cat died, the entire family mourned and shaved their eyebrows. Families took their dead cats to the sacred city of Bubastis, where they were embalmed and buried in sacred repositories. Herodotus expressed astonishment at the domestic cats in Egypt, because he had only ever seen wildcats.

 

Ancient Greeks and Romans kept weasels as pets, which were seen as the ideal rodent-killers. The earliest unmistakable evidence of the Greeks having domestic cats comes from two coins from Magna Graecia dating to the mid-fifth century BC showing Iokastos and Phalanthos, the legendary founders of Rhegion and Taras respectively, playing with their pet cats. The usual ancient Greek word for 'cat' was ailouros, meaning 'thing with the waving tail'. Cats are rarely mentioned in ancient Greek literature. Aristotle remarked in his History of Animals that "female cats are naturally lecherous." The Greeks later syncretized their own goddess Artemis with the Egyptian goddess Bastet, adopting Bastet's associations with cats and ascribing them to Artemis. In Ovid's Metamorphoses, when the deities flee to Egypt and take animal forms, the goddess Diana turns into a cat.

 

Cats eventually displaced weasels as the pest control of choice because they were more pleasant to have around the house and were more enthusiastic hunters of mice. During the Middle Ages, many of Artemis's associations with cats were grafted onto the Virgin Mary. Cats are often shown in icons of Annunciation and of the Holy Family and, according to Italian folklore, on the same night that Mary gave birth to Jesus, a cat in Bethlehem gave birth to a kitten. Domestic cats were spread throughout much of the rest of the world during the Age of Discovery, as ships' cats were carried on sailing ships to control shipboard rodents and as good-luck charms.

 

Several ancient religions believed cats are exalted souls, companions or guides for humans, that are all-knowing but mute so they cannot influence decisions made by humans. In Japan, the maneki neko cat is a symbol of good fortune. In Norse mythology, Freyja, the goddess of love, beauty, and fertility, is depicted as riding a chariot drawn by cats. In Jewish legend, the first cat was living in the house of the first man Adam as a pet that got rid of mice. The cat was once partnering with the first dog before the latter broke an oath they had made which resulted in enmity between the descendants of these two animals. It is also written that neither cats nor foxes are represented in the water, while every other animal has an incarnation species in the water. Although no species are sacred in Islam, cats are revered by Muslims. Some Western writers have stated Muhammad had a favorite cat, Muezza. He is reported to have loved cats so much, "he would do without his cloak rather than disturb one that was sleeping on it". The story has no origin in early Muslim writers, and seems to confuse a story of a later Sufi saint, Ahmed ar-Rifa'i, centuries after Muhammad. One of the companions of Muhammad was known as Abu Hurayrah ("father of the kitten"), in reference to his documented affection to cats.

 

Superstitions and rituals

Many cultures have negative superstitions about cats. An example would be the belief that encountering a black cat ("crossing one's path") leads to bad luck, or that cats are witches' familiars used to augment a witch's powers and skills. The killing of cats in Medieval Ypres, Belgium, is commemorated in the innocuous present-day Kattenstoet (cat parade). In mid-16th century France, cats would be burnt alive as a form of entertainment, particularly during midsummer festivals. According to Norman Davies, the assembled people "shrieked with laughter as the animals, howling with pain, were singed, roasted, and finally carbonized". The remaining ashes were sometimes taken back home by the people for good luck.

 

According to a myth in many cultures, cats have multiple lives. In many countries, they are believed to have nine lives, but in Italy, Germany, Greece, Brazil and some Spanish-speaking regions, they are said to have seven lives, while in Arabic traditions, the number of lives is six. An early mention of the myth can be found in John Heywood's The Proverbs of John Heywood (1546)

 

Husband, (quoth she), ye studie, be merrie now,

And even as ye thinke now, so come to yow.

Nay not so, (quoth he), for my thought to tell right,

I thinke how you lay groning, wife, all last night.

Husband, a groning horse and a groning wife

Never faile their master, (quoth she), for my life.

No wife, a woman hath nine lives like a cat.

 

The myth is attributed to the natural suppleness and swiftness cats exhibit to escape life-threatening situations. Also lending credence to this myth is the fact that falling cats often land on their feet, using an instinctive righting reflex to twist their bodies around. Nonetheless, cats can still be injured or killed by a high fall.