Jan 5 005/366

 

Rassi is a very social cat - He loves attention!!

(I stuck my hand in the picture so I could use it for 365, if I needed to, LOL!!)

 

The hole in the soil was made by voles intent on stripping the covering from my car's central controller (allows the roof to fold down and rise, also the windows to go up and down!). Only cost me £500 to replace ;'-0

 

Anyway ... the wasp appeared to be digging in a new direction ... wonder if it was nesting? Strange!

Purbeck mason wasp (Pseudepipona herrichii) excavating nest burrow on heathland. Dorset, UK.

 

photo.domgreves.com

A diptych of a couple of rather distant shots of a Great White Egret being mobbed by a Black-headed Gull at Moor Green Lakes. I think the egret was minding its own business, but I guess the gull's instinct is to view it as a potential predator.

Model: Caleb Seguin

 

www.facebook.com/jessiehymaphotography

Tophill Low Nature Reserve

Nuthatch strategically making it's way to the feeders. Cromwell Bottom Nature Reserve.

Museum of Modern Art De Pont, Tilburg, The Netherlands.

 

Part of the Museum Behaviour series bit.ly/H0skbp

 

website | maasvlakte book | portfolio book

Farne Islands.

I have a lot of shots to post. I have been very busy, and then there are the photos I helped escape the house-clearance people from Mum's.

 

So, back to the matter in hand: Ospringe.

 

Ospringe is one of the most easily identifiable churches in Kent, with its unusual saddleback tower, but it is well seen, as you can see the tower before the turn off to Faversham. It looks fabulous.

 

Ospringe was a small village, but now is part of the urban sprawl of Faversham as it spreads to the south of the old A2.

 

You turn down a tight junction, then along a narrow road with cars parked on either side, until you break into open country, and the church is on a bend in the road.

 

I was last here on winter about a decade ago, it was a bitterly cold day and the planned Christmas Tree festival had been delayed a week due to bad weather the weekend before.

 

I cam here on the off-chance, and I was met by a volunteer come to clean the church, but no one with a key.

 

The vicar arrived, and after explaining again about the project, he reluctantly let me in, but warned he would not be here long.

 

Last time here, i took 7 shots, and none of details, so I made busy with the nifty fifty.....

 

John Vigar says this is a church hard to gain access too, maybe I have been lucky, but worth seeking out if you're passing.

 

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A pretty church whose thirteenth century origins seem lost beneath a Victorian veneer – yet inside all become clear. The north wall is thickened to take the rood loft staircase (see also Challock) but there is a medieval stair in the south side too, just to confuse. The font is a lovely twelfth century piece supported by the familiar five columns. Much of the glass is by Thomas Willement and displays his signature TW, which can also be seen in the Alpha emblem in the top of the striking east window. The chancel is a riot of Victoriana of grand design – constructed in several campaigns, the reredos and flooring definitely by different hands. Old photos show that the whole church was once stencilled, but now that the nave is relatively plain, the chancel is once more the focus of attention. The south chapel has a rather nice 19th century roof structure and must once have been a grand family chapel. All in all a lovely church full of interest and one which should be more accessible and better known.

 

www.kentchurches.info/church.asp?p=Ospringe

 

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OSPRINGE

LIES the next parish north westward from Sheldwich. It is usually written in antient records Ospringes, and takes its name from the spring or fresh stream which rises in it.

 

The town of Ospringe, as it is called, is a franchise separate from the hundred of Faversham, having a constable of its own, but the rest of the parish is within the jurisdiction of that hundred.

 

The borough of Chetham, in this parish, was given to the abbey of Faversham by Richard de Lucy, and confirmed to it by king Henry II. king John, and king Henry III. (fn. 1) It still continues an appendage to the manor of Faversham, at which a borsholder is chosen yearly for this borough, and extends over Beacon farm on the south side of the London road, at the 45th mile stone in Ospringe and Stone, and very little besides. There is another small borough in this parish, called the borough of Brimstone, for which a borsholder is elected annually at the same manor. It extends over the Red Lion inn, in Ospringe-street, and some land, an house and oast behind the bowling-green, northward of it.

 

The parish of Ospringe is of large extent, being near five miles from north to south, though it is not much more than two miles in breadth. The village, or town of Ospringe, as it was formerly called, and now usually Ospringe-street, stands on the high London road, between the 46th and 47th mile-stone, but the north side of the street, as well as of that road, from the summit of Judde hill, as far eastward as the 47th mile stone, is within Faversham parish, the liberties of which town begin from the rivulet in Ospringe, and extend eastward, including the late Mr. Lypeatt's new-built house. Thus that parish intervenes, and entirely separates from the rest of it that part of Ospringe parish, at the northern boundaries of it, in which are the storekeeper's house, part of the offices, &c. and some of the royal powder mills, and in the town of Faversham, that parish again intervening, there is a small part of Weststreet within this parish. The grand valley, called Newnham bottom, through which the high road leads to Maidstone, lies at the western boundary of the parish, on the summit of the hill eastward of it is Juddehouse, built after a design of Inigo Jones, a fine situation, having a most beautiful prospect eastward, over a most fertile extent of country, to the Boughton hills, and the channel north eastward of it, but the large tract of woodland, of many hundred acres, which reach up close to the gardens at the back of it, render it rather an unhealthy situation. About a quarter of a mile eastward of Ospringe-street is a good house, called from the antient oratory or chapel formerly adjoining to it, but pulled down within these few years, chapelhouse. This oratory was dedicated to St. Nicholas, and erected for a priest to say mass in it, for the safety and good success of passengers, who left their acknowledgments for his pains in it. It belonged lately to Mr. John Simmons, whose son sold it to Isaac Rutton, esq. and he alienated the house to Mr. Neame, the present owner; but on a part of the land adjoining he built an elegant villa, naming it Ospringe Place, in which he now resides.

 

In Ospringe-street there is a tolerable inn, and the remains of the Maison Dieu on each side of the high road close to the small rivulet which crosses the street. This stream rises at Westbrook, at a small distance southward of the hamlet of Whitehill, at the back of which it runs, and at about a mile and an half distance, passing by Ospringe church, and the mansion of Queen-court, now a respectable farm-house, it turns a mill, erected some years ago for the manufacturing of madder, though now used for the grinding corn, and having crossed Ospringe-street, it turns a gunpowder mill not far from it, occupied by government, but belonging to St. John's college, in Cambridge, and having supplied the storekeeper's gardens, it afterwards turns a corn-mill, close to the west side of Faversham town, after which it supplies the rest of the government mills and works, and runs from thence into Faversham creek, to which it is a very necessary and beneficial back water. There is a nailbourne, or temporary land spring, such as are not unusual in the parts of this county eastward of Sittingborne, which run but once perhaps in several years, their failing and continuance having no certain periods, the breaking forth of them being held by the common people to be a forerunner of scarcity and dearness of corn and victuals. This at Ospringe, when it breaks out, rises about half a mile southward of Whitehill, near Kennaways, in the road to Stalisfield, and joining the above-mentioned rivulet, which it considerably increases, flows with it into Faversham creek. In February, 1674, it began to run, but stopped before Michaelmas. It broke forth in February, 1712, and run with such violence along the high road, that trenches were cut through the lands adjoining to carry the water off, but it stopped again before Michaelmas. It had continued dry till it broke out afresh in 1753, and continued to run till summer 1778, when it stopped, and has continued dry ever since.

 

About a mile southward of Ospringe-street is the hamlet of Whitehill, mentioned before, situated in the vale through which the rivulet takes its course. There are two houses of some account in it, formerly owned by the family of Drayton, who had resided in this parish for many years. Robert Drayton resided here anno 7 Edward IV. in which year he died, and was buried in the church-yard of Ospringe, being then possessed, as appears by his will, of a house called Smythes, with its lands and appurtenances, at Whitehill. After this family had become extinct here, one of these houses came into the possession of Ruck, and escheated, for want of lawful heirs, to the lord of the manor, and now as such belongs to the earl of Guildford, but Mr. James Foord resides in it. The other, after the Draytons were become extinct here, came into the name of Wreight, one of whom, Henry Wreight, gent. died possessed of it in 1695, and was buried in Faversham church. His son of the same name resided here, and died in 1773, and his grandson Henry Wreight, gent. of Faversham, sold it to John Montresor of Belmont, esq. who now owns it, but John Smith esq. resides in it. About a mile westward on the hill, near Hanslets Fostall and the parsonage, is a new-erected house, called the Oaks, built not many years since, on the scite of an antient one, called Nicholas, formerly belonging to the Draytons, by Mr. John Toker, who resides in it; the woodgrounds in the upland parts of this parish are very extensive, and contain many hundred acres. The soil of this parish, from its large extent, is various, to the north and north-east of the church the lands are level and very fertile, being a fine rich loam, but as they extend southward to the uplands, the soil becomes more and more barren, much of it chalky, and the rest a cludgy red earth, stiff tillage land, and very stony. A fair is held in Ospringe-street on the 29th of May.

 

¶Much has already been said in the former parts of these volumes, of the different opinions of learned men where the Roman station, called in the second iter of Antonine Durolevum, ought to be placed. Most of the copies of Antonine make the distance from the last station Durobrovis, which is allowed by all to be Rochester, to the station of Durolevum, to be xiii or xvi miles, though the Peutongerian tables make it only vii. If the number xvi is right, no place bids so fair for it as Judde-hill, in this parish, which then would have every probable circumstance in favor of it. The Romans undoubtedly had some strong military post on this hill, on the summit of which there are the remains of a very deep and broad ditch, the south and east sides are still entire, as is a small part of the north side at the eastern corners of it, the remaining part of the north side was filled up not many years since. The west side has nothing left of it; close within the southern part of it is a high mount of earth thrown up to a considerable height above the ground round it, the scite of Judde house, and the gardens are contained within it. The form of it seems to have been a square, with the corners rounded, and to have contained between three and four acres of ground within its area, the common people call it king Stephen's castle, but it is certainly of a much older date. At a small distance from it, on the opposite, or north side of the high road, there are several breast works cast up across the field facing the west. At the bottom of the hill, in the next field to this, are the ruins of Stone chapel, in which numbers of Roman bricks are interspersed among the flints, and in the midst of the south wall of it, there is a separate piece of a Roman building, about a rod in length, and near three feet high, composed of two rows of Roman tiles, of about fourteen inches square each, and on them are laid small stones hewed, but of no regular size or shape, for about a foot high, and then tiles again, and so on alternately.

  

THIS PARISH is within the ECCLESIASTICAL JURISDICTION of the diocese of Canterbury, and deanry of Ospringe

 

The church stands within the jurisdiction of the town of Ospringe, about half a mile southward from Ospringe-street. It is dedicated to St. Peter and St. Paul. It is an antient building, consisting of three isles and a chancel. The steeple was formerly at the west end, and was built circular of flints, supposed to be Danish, with a shingled spire on it, of upwards of fifty feet high, in which were four bells; but in ringing them on Oct. 11, 1695, on king William's return from Flanders, it suddenly fell to the ground, providentially no one was hurt by it. There are no remains left of any painted glass in the windows of this church, though there was formerly much in most of them; particularly, in the window of the north isle was once the figure of a mitred bishop, on the rack, with a knife on the table by him, and of another person tied to a tree, and wounded with arrows. In another was a label to the memory of Robert Seton, and of a woman kneeling; and there was not many years ago remaining in the east window, at the end of the south isle, forming a kind of chancel, the effigies of a knight in his tabard of arms, with spurs on his heels, in a kneeling posture, looking up to a crucisix, painted just above him, of which there remained only the lower part. The knight's arms, Azure, three harts heads, caboshed, or, were thrown under him, and at a little distance some part of his crest, An hart's head, attired full, or, with a crown about his neck, azure, and underneath, Pray for the soul of Thomas Hart. This Sir Thomas Hart was possessed of an estate in this parish, which he purchased of Norwood. The Greenstreets, of Selling, lately claimed this chancel, and several of them lie buried in it. There was a chapel, dedicated to St. Thomas, in this church.

 

In the east part of the church-yard there was once a chapel, said to have been built by Sir John Denton, of Denton, in this parish and Easling, the foundations of which are still visible.

 

It appears by the Testa de Nevil, taken in the reign of king Henry III. that the church of Ospringe was in the king's gift, and was afterwards given by king John to John de Burgo, who then held it, and that it was worth forty marcs. After which, in the 8th year of Richard II. anno 1384, it was become appropriated to the abbot of Pontiniac, and was valued at 13l. 6s. 8d. at which time there was a vicarage here of his patronage likewise. It afterwards became part of the possessions of the hospital or Maison Dieu, in Ospringestreet, but by what means, or when, I have not found, and it continued so till the escheat of the hospital anno 20 Edward IV. after which, the parsonage appropriate of this church of Ospringe, together with the advowson of the vicarage, was by means of Fisher, bishop of Rochester, obtained of Henry VIII. in manner as has been already mentioned, for St. John's college, in Cambridge, the master and fellows of which are at this time entitled to them, the parsonage being let by them on a beneficial lease; but the advowson of the vicarage they retain in their own hands.

 

The lessee of this parsonage, in the reign of queen Elizabeth, was Robert Streynsham, esq. who rebuilt the house and offices belonging to it, and afterwards resided in it. He had been fellow of All Souls college, LL. B. and secretary to the earl of Pembroke. He lies buried in this church, and bore for his arms, Or, a pale dancette, gules. He left two daughters and coheirs, of whom, Audrey, the eldest, carried her interest in it in marriage to Edward Master, esq. eldest son of James Master, esq. of East Langdon, who was first of Sandwich, and afterwards built a seat for himself and his posterity at East Langdon. He was twice married, and had fourteen children; at length worn out with age, he betook himself hither to his eldest son Edward, and dying in 1631, æt. 84, was buried in this church. Edward Master, the son, resided here, and was afterwards knighted, and on his father's death in 1631 removed to that seat, in whose descendants it continued till it was at length alienated to Buller, of Cornwall, whose son sold his interest in to Markham, as he did to Mr. Robert Lyddel, merchant, of London, brother of Sir Henry Lyddel, who in 1751 assigned his interest in it to Ralph Terrey, yeoman, of Knolton, whose son Mr. Michael Terrey, of Ospringe, devised it to his only daughter and heir Olive, who married Nathaniel Marsh, esq. of Boughton Blean, and the heirs of his son Terrey Marsh, esq. late of that parish, are the present lessees of it.

 

The vicarage of Ospringe is valued in the king's books at ten pounds, and the yearly tenths at one pound.

 

In 1640 it was valued at sixty pounds, when there were communicants here 226.

 

The vicarage is endowed with all vicarial tithes, woad only excepted, and also with those of hay, saintfoin, clover, and coppice woods. There are about twenty-seven acres of glebe-land belonging to it. The vicarage-house is situated in the valley, at a small distance eastward from the church, and the parsonagehouse near a mile southward of that.

 

Ospringe was formerly the head of a rural deanry, of which institution it will be necessary to give some account here.

 

The office of rural dean was not unknown to our Saxon ancestors, as appears by the laws of king Edward the Confessor; they were called both Archipresbiteri and Decani Temporarii, to distinguish them from the deans of cathedrals, who were Decani Perpetui. Besides these, there were in the greater monasteries, especially those of the Benedictine order, such officers called deans, and there are deans still remaining in several of the colleges of the universities, who take care of the studies and exercises of the youth, and are a check on the morals and behaviour of such as are members under them.

 

¶The antient exercise of jurisdiction in the church seems to have been instituted in conformity to like subordinations in the state. Thus the dioceses within this realm seem to have been divided into archdeaconries and rural deanries, to make them correspond to the like division of the kingdom into counties and hundreds; hence the former, whose courts were to answer those of the county, had the county usually for their district, and took their title from thence, and the names of the latter from the hundred, or chief place of it, wherein they acted; and as in the state every hundred was at first divided into ten tithings or fribourghs, and every tithing was made up of ten families, both which kept their original names, notwithstanding the increase of villages and people; so in the church the name of deanry continued, notwithstanding the increase of persons and churches, and the districts of them were contracted and enlarged from time to time, at the discretion of the bishop, the rural dean of Ospringe having jurisdiction over the whole deanry of it, consisting of twenty-six parishes. He had a seal of office, which being temporary, it had only the name of the office, and not, as other seals of jurisdiction, the name of the person also, engraved on it. The seal belonging to this deanry had on it, the Virgin Mary crowned, with the sceptre in her left hand, and her child, with a glory round his head, in her right, and round the margin, Sigillu Decani Decanatus de Ospreng. He was in antient times called the dean of the bishop, because appointed by him, and had alone the inspection of the lives and manners of the clergy and people within the district under him, and was to report the same to the bishop; to which end, that he might have a thorough knowledge of the state and condition of his respective deanry, he had a power to convene rural chapters, which were made up of the instituted clergy, or their curates as proxies of them, and the dean as president of them, where the clergy brought information of all irregularities committed within their respective parishes. Those upon ordinary occasions were held at first every three weeks, in imitation of the courts of manors, held from three weeks to three weeks, and afterwards each month, and from thence were called Kalendæ, but their more solemn and principal chapters were assembled once a quarter, where maters of greater import were transacted, and a fuller attendance given. They were at first held in any one church within the district, where the minister of the place was to procure and provide entertainment and procurations for the dean and his immediate officers, and they were afterwards held only in the larger or more eminent parishes. The part of their office of inspecting and reporting the manners of the clergy and people, rendered them necessary attendants on the episcopal synod or general visitation, in which they were the standing representatives of the rest of the clergy within their division, and they were there to deliver information of abuses committed within their knowledge, and consult for the reformation of them; for which they were to have their expences, called from hence synodals, allowed them by those whom they represented, according to the time of their attendance. That part of their office, of being convened to provincial and episcopal synods, was transferred to two proctors, or representatives of the parochial clergy in each diocese; and that of information of scandals and offences, has devolved on the churchwardens of the respective parishes. Besides this another principal part of the duty of a rural dean was to execute all processes of the bishop, or of the officers and ministers under his authority; but by the constitution of the pope's legate, Otho, the archdeacon, in the reign of Henry III. was required to be frequently present at them, who being superior to the rural dean, did in effect take the presidency out of his hands; and these chapters were afterwards often held by the archdeacon's officials, from which may be dated the decay of rural deanries, for the rural dean was not only discouraged by this, but the archdeacon and his official, as might naturally be supposed he would, drew the business usually transacted there to his own visitation, or chapter, as it might be termed. By which intersering of the archdeacon and his officials, it happened that in the age next before the reformation, the jurisdiction of rural deans declined almost to nothing, and at the reformation nothing was done for their restoration by the legislative power, so that they became extinct in most deanries, nor did this of Ospringe survive the earliest decline of them. (fn. 16) Where they still continue, they have only the name and shadow left, and what little remains of this dignity and jurisdiction, de pends greatly on the custom of places, and the pleasure of diocesans.

 

In the 31st year of Edward I. Richard Christian, dean of Ospringe, being sent to execute some citations of the archbishop at Selling, was set upon by the people there, who placed him with his face to his horse's tail, which they made him hold in his hand for a bridle, in which posture they led him through the village, with songs, shouts, and dances, and afterwards having cut off the tail, ears, and lips of the beast, they threw the dean into the dirt, to his great disgrace; for which, the king directed his writ to the sheriff, to make enquiry by inquisition of a jury concerning it.

 

www.british-history.ac.uk/survey-kent/vol6/pp499-531

Behaving to type and following a leader. Sheep are sheared at this time of year to avoid heatstroke and combat parasites. I’m sure ewe knew that already.

Fish, any of approximately 34,000 species of vertebrate animals (phylum Chordata) found in the fresh and salt waters of the world. Living species range from the primitive jawless lampreys and hagfishes through the cartilaginous sharks, skates, and rays to the abundant and diverse bony fishes. Most fish species are cold-blooded; however, one species, the opah (Lampris guttatus), is warm-blooded.

 

The term fish is applied to a variety of vertebrates of several evolutionary lines. It describes a life-form rather than a taxonomic group. As members of the phylum Chordata, fish share certain features with other vertebrates. These features are gill slits at some point in the life cycle, a notochord, or skeletal supporting rod, a dorsal hollow nerve cord, and a tail. Living fishes represent some five classes, which are as distinct from one another as are the four classes of familiar air-breathing animals—amphibians, reptiles, birds, and mammals. For example, the jawless fishes (Agnatha) have gills in pouches and lack limb girdles. Extant agnathans are the lampreys and the hagfishes. As the name implies, the skeletons of fishes of the class Chondrichthyes (from chondr, “cartilage,” and ichthyes, “fish”) are made entirely of cartilage. Modern fish of this class lack a swim bladder, and their scales and teeth are made up of the same placoid material. Sharks, skates, and rays are examples of cartilaginous fishes. The bony fishes are by far the largest class. Examples range from the tiny seahorse to the 450-kg (1,000-pound) blue marlin, from the flattened soles and flounders to the boxy puffers and ocean sunfishes. Unlike the scales of the cartilaginous fishes, those of bony fishes, when present, grow throughout life and are made up of thin overlapping plates of bone. Bony fishes also have an operculum that covers the gill slits.

 

The study of fishes, the science of ichthyology, is of broad importance. Fishes are of interest to humans for many reasons, the most important being their relationship with and dependence on the environment. A more obvious reason for interest in fishes is their role as a moderate but important part of the world’s food supply. This resource, once thought unlimited, is now realized to be finite and in delicate balance with the biological, chemical, and physical factors of the aquatic environment. Overfishing, pollution, and alteration of the environment are the chief enemies of proper fisheries management, both in fresh waters and in the ocean. (For a detailed discussion of the technology and economics of fisheries, see commercial fishing.) Another practical reason for studying fishes is their use in disease control. As predators on mosquito larvae, they help curb malaria and other mosquito-borne diseases.

 

Fishes are valuable laboratory animals in many aspects of medical and biological research. For example, the readiness of many fishes to acclimate to captivity has allowed biologists to study behaviour, physiology, and even ecology under relatively natural conditions. Fishes have been especially important in the study of animal behaviour, where research on fishes has provided a broad base for the understanding of the more flexible behaviour of the higher vertebrates. The zebra fish is used as a model in studies of gene expression.

 

There are aesthetic and recreational reasons for an interest in fishes. Millions of people keep live fishes in home aquariums for the simple pleasure of observing the beauty and behaviour of animals otherwise unfamiliar to them. Aquarium fishes provide a personal challenge to many aquarists, allowing them to test their ability to keep a small section of the natural environment in their homes. Sportfishing is another way of enjoying the natural environment, also indulged in by millions of people every year. Interest in aquarium fishes and sportfishing supports multimillion-dollar industries throughout the world.

 

Fishes have been in existence for more than 450 million years, during which time they have evolved repeatedly to fit into almost every conceivable type of aquatic habitat. In a sense, land vertebrates are simply highly modified fishes: when fishes colonized the land habitat, they became tetrapod (four-legged) land vertebrates. The popular conception of a fish as a slippery, streamlined aquatic animal that possesses fins and breathes by gills applies to many fishes, but far more fishes deviate from that conception than conform to it. For example, the body is elongate in many forms and greatly shortened in others; the body is flattened in some (principally in bottom-dwelling fishes) and laterally compressed in many others; the fins may be elaborately extended, forming intricate shapes, or they may be reduced or even lost; and the positions of the mouth, eyes, nostrils, and gill openings vary widely. Air breathers have appeared in several evolutionary lines.

 

Many fishes are cryptically coloured and shaped, closely matching their respective environments; others are among the most brilliantly coloured of all organisms, with a wide range of hues, often of striking intensity, on a single individual. The brilliance of pigments may be enhanced by the surface structure of the fish, so that it almost seems to glow. A number of unrelated fishes have actual light-producing organs. Many fishes are able to alter their coloration—some for the purpose of camouflage, others for the enhancement of behavioral signals.

 

Fishes range in adult length from less than 10 mm (0.4 inch) to more than 20 metres (60 feet) and in weight from about 1.5 grams (less than 0.06 ounce) to many thousands of kilograms. Some live in shallow thermal springs at temperatures slightly above 42 °C (100 °F), others in cold Arctic seas a few degrees below 0 °C (32 °F) or in cold deep waters more than 4,000 metres (13,100 feet) beneath the ocean surface. The structural and, especially, the physiological adaptations for life at such extremes are relatively poorly known and provide the scientifically curious with great incentive for study.

 

Almost all natural bodies of water bear fish life, the exceptions being very hot thermal ponds and extremely salt-alkaline lakes, such as the Dead Sea in Asia and the Great Salt Lake in North America. The present distribution of fishes is a result of the geological history and development of Earth as well as the ability of fishes to undergo evolutionary change and to adapt to the available habitats. Fishes may be seen to be distributed according to habitat and according to geographical area. Major habitat differences are marine and freshwater. For the most part, the fishes in a marine habitat differ from those in a freshwater habitat, even in adjacent areas, but some, such as the salmon, migrate from one to the other. The freshwater habitats may be seen to be of many kinds. Fishes found in mountain torrents, Arctic lakes, tropical lakes, temperate streams, and tropical rivers will all differ from each other, both in obvious gross structure and in physiological attributes. Even in closely adjacent habitats where, for example, a tropical mountain torrent enters a lowland stream, the fish fauna will differ. The marine habitats can be divided into deep ocean floors (benthic), mid-water oceanic (bathypelagic), surface oceanic (pelagic), rocky coast, sandy coast, muddy shores, bays, estuaries, and others. Also, for example, rocky coastal shores in tropical and temperate regions will have different fish faunas, even when such habitats occur along the same coastline.

 

Although much is known about the present geographical distribution of fishes, far less is known about how that distribution came about. Many parts of the fish fauna of the fresh waters of North America and Eurasia are related and undoubtedly have a common origin. The faunas of Africa and South America are related, extremely old, and probably an expression of the drifting apart of the two continents. The fauna of southern Asia is related to that of Central Asia, and some of it appears to have entered Africa. The extremely large shore-fish faunas of the Indian and tropical Pacific oceans comprise a related complex, but the tropical shore fauna of the Atlantic, although containing Indo-Pacific components, is relatively limited and probably younger. The Arctic and Antarctic marine faunas are quite different from each other. The shore fauna of the North Pacific is quite distinct, and that of the North Atlantic more limited and probably younger. Pelagic oceanic fishes, especially those in deep waters, are similar the world over, showing little geographical isolation in terms of family groups. The deep oceanic habitat is very much the same throughout the world, but species differences do exist, showing geographical areas determined by oceanic currents and water masses.

 

All aspects of the life of a fish are closely correlated with adaptation to the total environment, physical, chemical, and biological. In studies, all the interdependent aspects of fish, such as behaviour, locomotion, reproduction, and physical and physiological characteristics, must be taken into account.

 

Correlated with their adaptation to an extremely wide variety of habitats is the extremely wide variety of life cycles that fishes display. The great majority hatch from relatively small eggs a few days to several weeks or more after the eggs are scattered in the water. Newly hatched young are still partially undeveloped and are called larvae until body structures such as fins, skeleton, and some organs are fully formed. Larval life is often very short, usually less than a few weeks, but it can be very long, some lampreys continuing as larvae for at least five years. Young and larval fishes, before reaching sexual maturity, must grow considerably, and their small size and other factors often dictate that they live in a habitat different than that of the adults. For example, most tropical marine shore fishes have pelagic larvae. Larval food also is different, and larval fishes often live in shallow waters, where they may be less exposed to predators.

 

After a fish reaches adult size, the length of its life is subject to many factors, such as innate rates of aging, predation pressure, and the nature of the local climate. The longevity of a species in the protected environment of an aquarium may have nothing to do with how long members of that species live in the wild. Many small fishes live only one to three years at the most. In some species, however, individuals may live as long as 10 or 20 or even 100 years.

 

Fish behaviour is a complicated and varied subject. As in almost all animals with a central nervous system, the nature of a response of an individual fish to stimuli from its environment depends upon the inherited characteristics of its nervous system, on what it has learned from past experience, and on the nature of the stimuli. Compared with the variety of human responses, however, that of a fish is stereotyped, not subject to much modification by “thought” or learning, and investigators must guard against anthropomorphic interpretations of fish behaviour.

 

Fishes perceive the world around them by the usual senses of sight, smell, hearing, touch, and taste and by special lateral line water-current detectors. In the few fishes that generate electric fields, a process that might best be called electrolocation aids in perception. One or another of these senses often is emphasized at the expense of others, depending upon the fish’s other adaptations. In fishes with large eyes, the sense of smell may be reduced; others, with small eyes, hunt and feed primarily by smell (such as some eels).

 

Specialized behaviour is primarily concerned with the three most important activities in the fish’s life: feeding, reproduction, and escape from enemies. Schooling behaviour of sardines on the high seas, for instance, is largely a protective device to avoid enemies, but it is also associated with and modified by their breeding and feeding requirements. Predatory fishes are often solitary, lying in wait to dart suddenly after their prey, a kind of locomotion impossible for beaked parrot fishes, which feed on coral, swimming in small groups from one coral head to the next. In addition, some predatory fishes that inhabit pelagic environments, such as tunas, often school.

 

Sleep in fishes, all of which lack true eyelids, consists of a seemingly listless state in which the fish maintains its balance but moves slowly. If attacked or disturbed, most can dart away. A few kinds of fishes lie on the bottom to sleep. Most catfishes, some loaches, and some eels and electric fishes are strictly nocturnal, being active and hunting for food during the night and retiring during the day to holes, thick vegetation, or other protective parts of the environment.

 

Communication between members of a species or between members of two or more species often is extremely important, especially in breeding behaviour (see below Reproduction). The mode of communication may be visual, as between the small so-called cleaner fish and a large fish of a very different species. The larger fish often allows the cleaner to enter its mouth to remove gill parasites. The cleaner is recognized by its distinctive colour and actions and therefore is not eaten, even if the larger fish is normally a predator. Communication is often chemical, signals being sent by specific chemicals called pheromones.

 

Many fishes have a streamlined body and swim freely in open water. Fish locomotion is closely correlated with habitat and ecological niche (the general position of the animal to its environment).

 

Many fishes in both marine and fresh waters swim at the surface and have mouths adapted to feed best (and sometimes only) at the surface. Often such fishes are long and slender, able to dart at surface insects or at other surface fishes and in turn to dart away from predators; needlefishes, halfbeaks, and topminnows (such as killifish and mosquito fish) are good examples. Oceanic flying fishes escape their predators by gathering speed above the water surface, with the lower lobe of the tail providing thrust in the water. They then glide hundreds of yards on enlarged, winglike pectoral and pelvic fins. South American freshwater flying fishes escape their enemies by jumping and propelling their strongly keeled bodies out of the water.

 

So-called mid-water swimmers, the most common type of fish, are of many kinds and live in many habitats. The powerful fusiform tunas and the trouts, for example, are adapted for strong, fast swimming, the tunas to capture prey speedily in the open ocean and the trouts to cope with the swift currents of streams and rivers. The trout body form is well adapted to many habitats. Fishes that live in relatively quiet waters such as bays or lake shores or slow rivers usually are not strong, fast swimmers but are capable of short, quick bursts of speed to escape a predator. Many of these fishes have their sides flattened, examples being the sunfish and the freshwater angelfish of aquarists. Fish associated with the bottom or substrate usually are slow swimmers. Open-water plankton-feeding fishes almost always remain fusiform and are capable of rapid, strong movement (for example, sardines and herrings of the open ocean and also many small minnows of streams and lakes).

 

Bottom-living fishes are of many kinds and have undergone many types of modification of their body shape and swimming habits. Rays, which evolved from strong-swimming mid-water sharks, usually stay close to the bottom and move by undulating their large pectoral fins. Flounders live in a similar habitat and move over the bottom by undulating the entire body. Many bottom fishes dart from place to place, resting on the bottom between movements, a motion common in gobies. One goby relative, the mudskipper, has taken to living at the edge of pools along the shore of muddy mangrove swamps. It escapes its enemies by flipping rapidly over the mud, out of the water. Some catfishes, synbranchid eels, the so-called climbing perch, and a few other fishes venture out over damp ground to find more promising waters than those that they left. They move by wriggling their bodies, sometimes using strong pectoral fins; most have accessory air-breathing organs. Many bottom-dwelling fishes live in mud holes or rocky crevices. Marine eels and gobies commonly are found in such habitats and for the most part venture far beyond their cavelike homes. Some bottom dwellers, such as the clingfishes (Gobiesocidae), have developed powerful adhesive disks that enable them to remain in place on the substrate in areas such as rocky coasts, where the action of the waves is great.

 

The methods of reproduction in fishes are varied, but most fishes lay a large number of small eggs, fertilized and scattered outside of the body. The eggs of pelagic fishes usually remain suspended in the open water. Many shore and freshwater fishes lay eggs on the bottom or among plants. Some have adhesive eggs. The mortality of the young and especially of the eggs is very high, and often only a few individuals grow to maturity out of hundreds, thousands, and in some cases millions of eggs laid.

 

Males produce sperm, usually as a milky white substance called milt, in two (sometimes one) testes within the body cavity. In bony fishes a sperm duct leads from each testis to a urogenital opening behind the vent or anus. In sharks and rays and in cyclostomes the duct leads to a cloaca. Sometimes the pelvic fins are modified to help transmit the milt to the eggs at the female’s vent or on the substrate where the female has placed them. Sometimes accessory organs are used to fertilize females internally—for example, the claspers of many sharks and rays.

 

In the females the eggs are formed in two ovaries (sometimes only one) and pass through the ovaries to the urogenital opening and to the outside. In some fishes the eggs are fertilized internally but are shed before development takes place. Members of about a dozen families each of bony fishes (teleosts) and sharks bear live young. Many skates and rays also bear live young. In some bony fishes the eggs simply develop within the female, the young emerging when the eggs hatch (ovoviviparous). Others develop within the ovary and are nourished by ovarian tissues after hatching (viviparous). There are also other methods utilized by fishes to nourish young within the female. In all live-bearers the young are born at a relatively large size and are few in number. In one family of primarily marine fishes, the surfperches from the Pacific coast of North America, Japan, and Korea, the males of at least one species are born sexually mature, although they are not fully grown.

 

Some fishes are hermaphroditic—an individual producing both sperm and eggs, usually at different stages of its life. Self-fertilization, however, is probably rare.

 

Successful reproduction and, in many cases, defense of the eggs and the young are assured by rather stereotypical but often elaborate courtship and parental behaviour, either by the male or the female or both. Some fishes prepare nests by hollowing out depressions in the sand bottom (cichlids, for example), build nests with plant materials and sticky threads excreted by the kidneys (sticklebacks), or blow a cluster of mucus-covered bubbles at the water surface (gouramis). The eggs are laid in these structures. Some varieties of cichlids and catfishes incubate eggs in their mouths.

 

Some fishes, such as salmon, undergo long migrations from the ocean and up large rivers to spawn in the gravel beds where they themselves hatched (anadromous fishes). Some, such as the freshwater eels (family Anguillidae), live and grow to maturity in fresh water and migrate to the sea to spawn (catadromous fishes). Other fishes undertake shorter migrations from lakes into streams, within the ocean, or enter spawning habitats that they do not ordinarily occupy in other ways.

 

The basic structure and function of the fish body are similar to those of all other vertebrates. The usual four types of tissues are present: surface or epithelial, connective (bone, cartilage, and fibrous tissues, as well as their derivative, blood), nerve, and muscle tissues. In addition, the fish’s organs and organ systems parallel those of other vertebrates.

 

The typical fish body is streamlined and spindle-shaped, with an anterior head, a gill apparatus, and a heart, the latter lying in the midline just below the gill chamber. The body cavity, containing the vital organs, is situated behind the head in the lower anterior part of the body. The anus usually marks the posterior termination of the body cavity and most often occurs just in front of the base of the anal fin. The spinal cord and vertebral column continue from the posterior part of the head to the base of the tail fin, passing dorsal to the body cavity and through the caudal (tail) region behind the body cavity. Most of the body is of muscular tissue, a high proportion of which is necessitated by swimming. In the course of evolution this basic body plan has been modified repeatedly into the many varieties of fish shapes that exist today.

 

The skeleton forms an integral part of the fish’s locomotion system, as well as serving to protect vital parts. The internal skeleton consists of the skull bones (except for the roofing bones of the head, which are really part of the external skeleton), the vertebral column, and the fin supports (fin rays). The fin supports are derived from the external skeleton but will be treated here because of their close functional relationship to the internal skeleton. The internal skeleton of cyclostomes, sharks, and rays is of cartilage; that of many fossil groups and some primitive living fishes is mostly of cartilage but may include some bone. In place of the vertebral column, the earliest vertebrates had a fully developed notochord, a flexible stiff rod of viscous cells surrounded by a strong fibrous sheath. During the evolution of modern fishes the rod was replaced in part by cartilage and then by ossified cartilage. Sharks and rays retain a cartilaginous vertebral column; bony fishes have spool-shaped vertebrae that in the more primitive living forms only partially replace the notochord. The skull, including the gill arches and jaws of bony fishes, is fully, or at least partially, ossified. That of sharks and rays remains cartilaginous, at times partially replaced by calcium deposits but never by true bone.

 

The supportive elements of the fins (basal or radial bones or both) have changed greatly during fish evolution. Some of these changes are described in the section below (Evolution and paleontology). Most fishes possess a single dorsal fin on the midline of the back. Many have two and a few have three dorsal fins. The other fins are the single tail and anal fins and paired pelvic and pectoral fins. A small fin, the adipose fin, with hairlike fin rays, occurs in many of the relatively primitive teleosts (such as trout) on the back near the base of the caudal fin.

 

The skin of a fish must serve many functions. It aids in maintaining the osmotic balance, provides physical protection for the body, is the site of coloration, contains sensory receptors, and, in some fishes, functions in respiration. Mucous glands, which aid in maintaining the water balance and offer protection from bacteria, are extremely numerous in fish skin, especially in cyclostomes and teleosts. Since mucous glands are present in the modern lampreys, it is reasonable to assume that they were present in primitive fishes, such as the ancient Silurian and Devonian agnathans. Protection from abrasion and predation is another function of the fish skin, and dermal (skin) bone arose early in fish evolution in response to this need. It is thought that bone first evolved in skin and only later invaded the cartilaginous areas of the fish’s body, to provide additional support and protection. There is some argument as to which came first, cartilage or bone, and fossil evidence does not settle the question. In any event, dermal bone has played an important part in fish evolution and has different characteristics in different groups of fishes. Several groups are characterized at least in part by the kind of bony scales they possess.

 

Scales have played an important part in the evolution of fishes. Primitive fishes usually had thick bony plates or thick scales in several layers of bone, enamel, and related substances. Modern teleost fishes have scales of bone, which, while still protective, allow much more freedom of motion in the body. A few modern teleosts (some catfishes, sticklebacks, and others) have secondarily acquired bony plates in the skin. Modern and early sharks possessed placoid scales, a relatively primitive type of scale with a toothlike structure, consisting of an outside layer of enamel-like substance (vitrodentine), an inner layer of dentine, and a pulp cavity containing nerves and blood vessels. Primitive bony fishes had thick scales of either the ganoid or the cosmoid type. Cosmoid scales have a hard, enamel-like outer layer, an inner layer of cosmine (a form of dentine), and then a layer of vascular bone (isopedine). In ganoid scales the hard outer layer is different chemically and is called ganoin. Under this is a cosminelike layer and then a vascular bony layer. The thin, translucent bony scales of modern fishes, called cycloid and ctenoid (the latter distinguished by serrations at the edges), lack enameloid and dentine layers.

 

Skin has several other functions in fishes. It is well supplied with nerve endings and presumably receives tactile, thermal, and pain stimuli. Skin is also well supplied with blood vessels. Some fishes breathe in part through the skin, by the exchange of oxygen and carbon dioxide between the surrounding water and numerous small blood vessels near the skin surface.

 

Skin serves as protection through the control of coloration. Fishes exhibit an almost limitless range of colours. The colours often blend closely with the surroundings, effectively hiding the animal. Many fishes use bright colours for territorial advertisement or as recognition marks for other members of their own species, or sometimes for members of other species. Many fishes can change their colour to a greater or lesser degree, by movement of pigment within the pigment cells (chromatophores). Black pigment cells (melanophores), of almost universal occurrence in fishes, are often juxtaposed with other pigment cells. When placed beneath iridocytes or leucophores (bearing the silvery or white pigment guanine), melanophores produce structural colours of blue and green. These colours are often extremely intense, because they are formed by refraction of light through the needlelike crystals of guanine. The blue and green refracted colours are often relatively pure, lacking the red and yellow rays, which have been absorbed by the black pigment (melanin) of the melanophores. Yellow, orange, and red colours are produced by erythrophores, cells containing the appropriate carotenoid pigments. Other colours are produced by combinations of melanophores, erythrophores, and iridocytes.

 

The major portion of the body of most fishes consists of muscles. Most of the mass is trunk musculature, the fin muscles usually being relatively small. The caudal fin is usually the most powerful fin, being moved by the trunk musculature. The body musculature is usually arranged in rows of chevron-shaped segments on each side. Contractions of these segments, each attached to adjacent vertebrae and vertebral processes, bends the body on the vertebral joint, producing successive undulations of the body, passing from the head to the tail, and producing driving strokes of the tail. It is the latter that provides the strong forward movement for most fishes.

 

The digestive system, in a functional sense, starts at the mouth, with the teeth used to capture prey or collect plant foods. Mouth shape and tooth structure vary greatly in fishes, depending on the kind of food normally eaten. Most fishes are predacious, feeding on small invertebrates or other fishes and have simple conical teeth on the jaws, on at least some of the bones of the roof of the mouth, and on special gill arch structures just in front of the esophagus. The latter are throat teeth. Most predacious fishes swallow their prey whole, and the teeth are used for grasping and holding prey, for orienting prey to be swallowed (head first) and for working the prey toward the esophagus. There are a variety of tooth types in fishes. Some fishes, such as sharks and piranhas, have cutting teeth for biting chunks out of their victims. A shark’s tooth, although superficially like that of a piranha, appears in many respects to be a modified scale, while that of the piranha is like that of other bony fishes, consisting of dentine and enamel. Parrot fishes have beaklike mouths with short incisor-like teeth for breaking off coral and have heavy pavementlike throat teeth for crushing the coral. Some catfishes have small brushlike teeth, arranged in rows on the jaws, for scraping plant and animal growth from rocks. Many fishes (such as the Cyprinidae or minnows) have no jaw teeth at all but have very strong throat teeth.

 

Some fishes gather planktonic food by straining it from their gill cavities with numerous elongate stiff rods (gill rakers) anchored by one end to the gill bars. The food collected on these rods is passed to the throat, where it is swallowed. Most fishes have only short gill rakers that help keep food particles from escaping out the mouth cavity into the gill chamber.

 

Once reaching the throat, food enters a short, often greatly distensible esophagus, a simple tube with a muscular wall leading into a stomach. The stomach varies greatly in fishes, depending upon the diet. In most predacious fishes it is a simple straight or curved tube or pouch with a muscular wall and a glandular lining. Food is largely digested there and leaves the stomach in liquid form.

 

Between the stomach and the intestine, ducts enter the digestive tube from the liver and pancreas. The liver is a large, clearly defined organ. The pancreas may be embedded in it, diffused through it, or broken into small parts spread along some of the intestine. The junction between the stomach and the intestine is marked by a muscular valve. Pyloric ceca (blind sacs) occur in some fishes at this junction and have a digestive or absorptive function or both.

 

The intestine itself is quite variable in length, depending upon the fish’s diet. It is short in predacious forms, sometimes no longer than the body cavity, but long in herbivorous forms, being coiled and several times longer than the entire length of the fish in some species of South American catfishes. The intestine is primarily an organ for absorbing nutrients into the bloodstream. The larger its internal surface, the greater its absorptive efficiency, and a spiral valve is one method of increasing its absorption surface.

 

Sharks, rays, chimaeras, lungfishes, surviving chondrosteans, holosteans, and even a few of the more primitive teleosts have a spiral valve or at least traces of it in the intestine. Most modern teleosts have increased the area of the intestinal walls by having numerous folds and villi (fingerlike projections) somewhat like those in humans. Undigested substances are passed to the exterior through the anus in most teleost fishes. In lungfishes, sharks, and rays, it is first passed through the cloaca, a common cavity receiving the intestinal opening and the ducts from the urogenital system.

 

Oxygen and carbon dioxide dissolve in water, and most fishes exchange dissolved oxygen and carbon dioxide in water by means of the gills. The gills lie behind and to the side of the mouth cavity and consist of fleshy filaments supported by the gill arches and filled with blood vessels, which give gills a bright red colour. Water taken in continuously through the mouth passes backward between the gill bars and over the gill filaments, where the exchange of gases takes place. The gills are protected by a gill cover in teleosts and many other fishes but by flaps of skin in sharks, rays, and some of the older fossil fish groups. The blood capillaries in the gill filaments are close to the gill surface to take up oxygen from the water and to give up excess carbon dioxide to the water.

 

Most modern fishes have a hydrostatic (ballast) organ, called the swim bladder, that lies in the body cavity just below the kidney and above the stomach and intestine. It originated as a diverticulum of the digestive canal. In advanced teleosts, especially the acanthopterygians, the bladder has lost its connection with the digestive tract, a condition called physoclistic. The connection has been retained (physostomous) by many relatively primitive teleosts. In several unrelated lines of fishes, the bladder has become specialized as a lung or, at least, as a highly vascularized accessory breathing organ. Some fishes with such accessory organs are obligate air breathers and will drown if denied access to the surface, even in well-oxygenated water. Fishes with a hydrostatic form of swim bladder can control their depth by regulating the amount of gas in the bladder. The gas, mostly oxygen, is secreted into the bladder by special glands, rendering the fish more buoyant; the gas is absorbed into the bloodstream by another special organ, reducing the overall buoyancy and allowing the fish to sink. Some deep-sea fishes may have oils, rather than gas, in the bladder. Other deep-sea and some bottom-living forms have much-reduced swim bladders or have lost the organ entirely.

 

The swim bladder of fishes follows the same developmental pattern as the lungs of land vertebrates. There is no doubt that the two structures have the same historical origin in primitive fishes. More or less intermediate forms still survive among the more primitive types of fishes, such as the lungfishes Lepidosiren and Protopterus.

 

The circulatory, or blood vascular, system consists of the heart, the arteries, the capillaries, and the veins. It is in the capillaries that the interchange of oxygen, carbon dioxide, nutrients, and other substances such as hormones and waste products takes place. The capillaries lead to the veins, which return the venous blood with its waste products to the heart, kidneys, and gills. There are two kinds of capillary beds: those in the gills and those in the rest of the body. The heart, a folded continuous muscular tube with three or four saclike enlargements, undergoes rhythmic contractions and receives venous blood in a sinus venosus. It passes the blood to an auricle and then into a thick muscular pump, the ventricle. From the ventricle the blood goes to a bulbous structure at the base of a ventral aorta just below the gills. The blood passes to the afferent (receiving) arteries of the gill arches and then to the gill capillaries. There waste gases are given off to the environment, and oxygen is absorbed. The oxygenated blood enters efferent (exuant) arteries of the gill arches and then flows into the dorsal aorta. From there blood is distributed to the tissues and organs of the body. One-way valves prevent backflow. The circulation of fishes thus differs from that of the reptiles, birds, and mammals in that oxygenated blood is not returned to the heart prior to distribution to the other parts of the body.

 

The primary excretory organ in fishes, as in other vertebrates, is the kidney. In fishes some excretion also takes place in the digestive tract, skin, and especially the gills (where ammonia is given off). Compared with land vertebrates, fishes have a special problem in maintaining their internal environment at a constant concentration of water and dissolved substances, such as salts. Proper balance of the internal environment (homeostasis) of a fish is in a great part maintained by the excretory system, especially the kidney.

 

The kidney, gills, and skin play an important role in maintaining a fish’s internal environment and checking the effects of osmosis. Marine fishes live in an environment in which the water around them has a greater concentration of salts than they can have inside their body and still maintain life. Freshwater fishes, on the other hand, live in water with a much lower concentration of salts than they require inside their bodies. Osmosis tends to promote the loss of water from the body of a marine fish and absorption of water by that of a freshwater fish. Mucus in the skin tends to slow the process but is not a sufficient barrier to prevent the movement of fluids through the permeable skin. When solutions on two sides of a permeable membrane have different concentrations of dissolved substances, water will pass through the membrane into the more concentrated solution, while the dissolved chemicals move into the area of lower concentration (diffusion).

 

The kidney of freshwater fishes is often larger in relation to body weight than that of marine fishes. In both groups the kidney excretes wastes from the body, but the kidney of freshwater fishes also excretes large amounts of water, counteracting the water absorbed through the skin. Freshwater fishes tend to lose salt to the environment and must replace it. They get some salt from their food, but the gills and skin inside the mouth actively absorb salt from water passed through the mouth. This absorption is performed by special cells capable of moving salts against the diffusion gradient. Freshwater fishes drink very little water and take in little water with their food.

 

Marine fishes must conserve water, and therefore their kidneys excrete little water. To maintain their water balance, marine fishes drink large quantities of seawater, retaining most of the water and excreting the salt. Most nitrogenous waste in marine fishes appears to be secreted by the gills as ammonia. Marine fishes can excrete salt by clusters of special cells (chloride cells) in the gills.

 

There are several teleosts—for example, the salmon—that travel between fresh water and seawater and must adjust to the reversal of osmotic gradients. They adjust their physiological processes by spending time (often surprisingly little time) in the intermediate brackish environment.

 

Marine hagfishes, sharks, and rays have osmotic concentrations in their blood about equal to that of seawater and so do not have to drink water nor perform much physiological work to maintain their osmotic balance. In sharks and rays the osmotic concentration is kept high by retention of urea in the blood. Freshwater sharks have a lowered concentration of urea in the blood.

 

Endocrine glands secrete their products into the bloodstream and body tissues and, along with the central nervous system, control and regulate many kinds of body functions. Cyclostomes have a well-developed endocrine system, and presumably it was well developed in the early Agnatha, ancestral to modern fishes. Although the endocrine system in fishes is similar to that of higher vertebrates, there are numerous differences in detail. The pituitary, the thyroid, the suprarenals, the adrenals, the pancreatic islets, the sex glands (ovaries and testes), the inner wall of the intestine, and the bodies of the ultimobranchial gland make up the endocrine system in fishes. There are some others whose function is not well understood. These organs regulate sexual activity and reproduction, growth, osmotic pressure, general metabolic activities such as the storage of fat and the utilization of foodstuffs, blood pressure, and certain aspects of skin colour. Many of these activities are also controlled in part by the central nervous system, which works with the endocrine system in maintaining the life of a fish. Some parts of the endocrine system are developmentally, and undoubtedly evolutionarily, derived from the nervous system.

 

As in all vertebrates, the nervous system of fishes is the primary mechanism coordinating body activities, as well as integrating these activities in the appropriate manner with stimuli from the environment. The central nervous system, consisting of the brain and spinal cord, is the primary integrating mechanism. The peripheral nervous system, consisting of nerves that connect the brain and spinal cord to various body organs, carries sensory information from special receptor organs such as the eyes, internal ears, nares (sense of smell), taste glands, and others to the integrating centres of the brain and spinal cord. The peripheral nervous system also carries information via different nerve cells from the integrating centres of the brain and spinal cord. This coded information is carried to the various organs and body systems, such as the skeletal muscular system, for appropriate action in response to the original external or internal stimulus. Another branch of the nervous system, the autonomic nervous system, helps to coordinate the activities of many glands and organs and is itself closely connected to the integrating centres of the brain.

 

The brain of the fish is divided into several anatomical and functional parts, all closely interconnected but each serving as the primary centre of integrating particular kinds of responses and activities. Several of these centres or parts are primarily associated with one type of sensory perception, such as sight, hearing, or smell (olfaction).

 

The sense of smell is important in almost all fishes. Certain eels with tiny eyes depend mostly on smell for location of food. The olfactory, or nasal, organ of fishes is located on the dorsal surface of the snout. The lining of the nasal organ has special sensory cells that perceive chemicals dissolved in the water, such as substances from food material, and send sensory information to the brain by way of the first cranial nerve. Odour also serves as an alarm system. Many fishes, especially various species of freshwater minnows, react with alarm to a chemical released from the skin of an injured member of their own species.

 

Many fishes have a well-developed sense of taste, and tiny pitlike taste buds or organs are located not only within their mouth cavities but also over their heads and parts of their body. Catfishes, which often have poor vision, have barbels (“whiskers”) that serve as supplementary taste organs, those around the mouth being actively used to search out food on the bottom. Some species of naturally blind cave fishes are especially well supplied with taste buds, which often cover most of their body surface.

 

Sight is extremely important in most fishes. The eye of a fish is basically like that of all other vertebrates, but the eyes of fishes are extremely varied in structure and adaptation. In general, fishes living in dark and dim water habitats have large eyes, unless they have specialized in some compensatory way so that another sense (such as smell) is dominant, in which case the eyes will often be reduced. Fishes living in brightly lighted shallow waters often will have relatively small but efficient eyes. Cyclostomes have somewhat less elaborate eyes than other fishes, with skin stretched over the eyeball perhaps making their vision somewhat less effective. Most fishes have a spherical lens and accommodate their vision to far or near subjects by moving the lens within the eyeball. A few sharks accommodate by changing the shape of the lens, as in land vertebrates. Those fishes that are heavily dependent upon the eyes have especially strong muscles for accommodation. Most fishes see well, despite the restrictions imposed by frequent turbidity of the water and by light refraction.

 

Fossil evidence suggests that colour vision evolved in fishes more than 300 million years ago, but not all living fishes have retained this ability. Experimental evidence indicates that many shallow-water fishes, if not all, have colour vision and see some colours especially well, but some bottom-dwelling shore fishes live in areas where the water is sufficiently deep to filter out most if not all colours, and these fishes apparently never see colours. When tested in shallow water, they apparently are unable to respond to colour differences.

 

Sound perception and balance are intimately associated senses in a fish. The organs of hearing are entirely internal, located within the skull, on each side of the brain and somewhat behind the eyes. Sound waves, especially those of low frequencies, travel readily through water and impinge directly upon the bones and fluids of the head and body, to be transmitted to the hearing organs. Fishes readily respond to sound; for example, a trout conditioned to escape by the approach of fishermen will take flight upon perceiving footsteps on a stream bank even if it cannot see a fisherman. Compared with humans, however, the range of sound frequencies heard by fishes is greatly restricted. Many fishes communicate with each other by producing sounds in their swim bladders, in their throats by rasping their teeth, and in other ways.

 

A fish or other vertebrate seldom has to rely on a single type of sensory information to determine the nature of the environment around it. A catfish uses taste and touch when examining a food object with its oral barbels. Like most other animals, fishes have many touch receptors over their body surface. Pain and temperature receptors also are present in fishes and presumably produce the same kind of information to a fish as to humans. Fishes react in a negative fashion to stimuli that would be painful to human beings, suggesting that they feel a sensation of pain.

 

An important sensory system in fishes that is absent in other vertebrates (except some amphibians) is the lateral line system. This consists of a series of heavily innervated small canals located in the skin and bone around the eyes, along the lower jaw, over the head, and down the mid-side of the body, where it is associated with the scales. Intermittently along these canals are located tiny sensory organs (pit organs) that apparently detect changes in pressure. The system allows a fish to sense changes in water currents and pressure, thereby helping the fish to orient itself to the various changes that occur in the physical environment.

  

Wonderful to watch. Captured in Dorset.

Cromwell Bottom Nature Reserve

I watched a Fries goby dance around the mud in 25m of water in Loch Fyne creating this pattern in an attempt to attract a mate. Its the first time I noticed this unusual behaviour. There is a slender seapen in the bottom right hand corner and at one oclock to act as a sense of scale. The image is a plan view the central bit is slightly raised. Unfortunately the fish noticed I was watching and soon disappeared into a hole

 

Greater Manchester Police has praised the behaviour of visitors to Manchester during a weekend of sport in the city.

 

On Friday 20 May 2016, the Great City Games saw a number of athletes compete in various events on a purpose-built athletics arena in Albert Square and track on Deansgate.

 

The following day (21 May 2016) saw Manchester United beat Crystal Palace 2-1 to win the FA Cup at Wembley Stadium, with a number of fans watching the match in public venues throughout Greater Manchester.

 

The weekend extravaganza concluded on Sunday 22 May 2016 with over 30,000 lining up to take part in the Great Manchester Run before England defeated Turkey 2-1 at the Etihad Stadium in a UEFA Euro 2016 warm-up match in the evening.

 

The events saw tens of thousands of visitors to the city centre, creating a buzzing and carnival-like atmosphere.

 

Assistant Chief Constable John O’Hare said: “This has been a fantastic weekend for Manchester and the atmosphere in the city has been superb from start to finish.

 

“It was great to see so many pictures of smiling faces and people having a good time and I hope everyone who has visited the city this weekend will be going away with some great memories.

 

“I would like to thank everyone who has played a key role in ensuring that the weekend has been successful.”

 

For more information about Policing in Greater Manchester please visit www.gmp.police.uk

 

To report crime call police on 101 the national non-emergency number.

 

You can also call anonymously with information about crime to Crimestoppers on 0800 555 111. Crimestoppers is an independent charity who will not want your name, just your information. Your call will not be traced or recorded and you do not have to go to court or give a statement.

 

Squid are cephalopods in the superorder Decapodiformes with elongated bodies, large eyes, eight arms and two tentacles. Like all other cephalopods, squid have a distinct head, bilateral symmetry, and a mantle. They are mainly soft-bodied, like octopuses, but have a small internal skeleton in the form of a rod-like gladius or pen, made of chitin.

 

Squid diverged from other cephalopods during the Jurassic and occupy a similar role to teleost fish as open water predators of similar size and behaviour. They play an important role in the open water food web. The two long tentacles are used to grab prey and the eight arms to hold and control it. The beak then cuts the food into suitable size chunks for swallowing. Squid are rapid swimmers, moving by jet propulsion, and largely locate their prey by sight. They are among the most intelligent of invertebrates, with groups of Humboldt squid having been observed hunting cooperatively. They are preyed on by sharks, other fish, sea birds, seals and cetaceans, particularly sperm whales.

 

Squid can change colour for camouflage and signalling. Some species are bioluminescent, using their light for counter-illumination camouflage, while many species can eject a cloud of ink to distract predators.

 

Squid are used for human consumption with commercial fisheries in Japan, the Mediterranean, the southwestern Atlantic, the eastern Pacific and elsewhere. They are used in cuisines around the world, often known as "calamari". Squid have featured in literature since classical times, especially in tales of giant squid and sea monsters.

 

TAXONOMY AND PHYLOGENY

Squid are members of the class Cephalopoda, subclass Coleoidea. The squid orders Myopsida and Oegopsida are in the superorder Decapodiformes (from the Greek for "ten-legged"). Two other orders of decapodiform cephalopods are also called squid, although they are taxonomically distinct from squids and differ recognizably in their gross anatomical features. They are the bobtail squid of order Sepiolida and the ram's horn squid of the monotypic order Spirulida. The vampire squid, however, is more closely related to the octopuses than to any squid.

 

The cladogram, not fully resolved, is based on Sanchez et al, 2018. Their molecular phylogeny used mitochondrial and nuclear DNA marker sequences; they comment that a robust phylogeny "has proven very challenging to obtain". If it is accepted that Sepiidae cuttlefish are a kind of squid, then the squids, excluding the vampire squid, form a clade as illustrated. Orders are shown in boldface; all the families not included in those orders, except Sepiadariidae and Sepiidae are in the paraphyletic order "Sepiida", are in the paraphyletic order "Oegopsida".

 

EVOLUTION

Crown coleoids (the ancestors of octopuses and squid) diverged at the end of the Paleozoic, in the Permian. Squid diverged during the Jurassic, but many squid families appeared in or after the Cretaceous. Both the coleoids and the teleost fish were involved in much adaptive radiation at this time, and the two modern groups resemble each other in size, ecology, habitat, morphology and behaviour, however some fish moved into fresh water while the coleoids remained in marine environments.

 

The ancestral coleoid was probably nautiloid-like with a strait septate shell that became immersed in the mantle and was used for buoyancy control. Four lines diverged from this, Spirulida (with one living member), the cuttlefishes, the squids and the octopuses. Squid have differentiated from the ancestral mollusc such that the body plan has been condensed antero-posteriorly and extended dorso-ventrally. What may have been the foot of the ancestor is modified into a complex set of appendages around the mouth. The sense organs are highly developed and include advanced eyes similar to those of vertebrates.

 

The ancestral shell has been lost, with only an internal gladius, or pen, remaining. The pen, made of a chitin-like material, is a feather-shaped internal structure that supports the squid's mantle and serves as a site for muscle attachment. The cuttlebone or sepion of the Sepiidae is calcareous and appears to have evolved afresh in the Tertiary

 

DESCIPTION

Squid are soft-bodied molluscs whose forms evolved to adopt an active predatory lifestyle. The head and foot of the squid are at one end of a long body, and this end is functionally anterior, leading the animal as it moves through the water. A set of eight arms and two distinctive tentacles surround the mouth; each appendage takes the form of a muscular hydrostat and is flexible and prehensile, usually bearing disc-like suckers.

 

The suckers may lie directly on the arm or be stalked. Their rims are stiffened with chitin and may contain minute toothlike denticles. These features, as well as strong musculature, and a small ganglion beneath each sucker to allow individual control, provide a very powerful adhesion to grip prey. Hooks are present on the arms and tentacles in some species, but their function is unclear. The two tentacles are much longer than the arms and are retractile. Suckers are limited to the spatulate tip of the tentacle, known as the manus.

 

In the mature male, the outer half of one of the left arms is hectocotylised – and ends in a copulatory pad rather than suckers. This is used for depositing a spermatophore inside the mantle cavity of a female. A ventral part of the foot has been converted into a funnel through which water exits the mantle cavity.

 

The main body mass is enclosed in the mantle, which has a swimming fin along each side. These fins are not the main source of locomotion in most species. The mantle wall is heavily muscled and internal. The visceral mass, which is covered by a thin, membranous epidermis, forms a cone-shaped posterior region known as the "visceral hump". The mollusc shell is reduced to an internal, longitudinal chitinous "pen" in the functionally dorsal part of the animal; the pen acts to stiffen the squid and provides attachments for muscles.

 

On the functionally ventral part of the body is an opening to the mantle cavity, which contains the gills (ctenidia) and openings from the excretory, digestive and reproductive systems. An inhalant siphon behind the funnel draws water into the mantel cavity via a valve. The squid uses the funnel for locomotion via precise jet propulsion. In this form of locomotion, water is sucked into the mantle cavity and expelled out of the funnel in a fast, strong jet. The direction of travel is varied by the orientation of the funnel. Squid are strong swimmers and certain species can "fly" for short distances out of the water.

 

CAMOUFLAGE

Squid make use of different kinds of camouflage, namely active camouflage for background matching (in shallow water) and counter-illumination. This helps to protect them from their predators and allows them to approach their prey.

 

The skin is covered in controllable chromatophores of different colours, enabling the squid to match its coloration to its surroundings. The play of colours may in addition distract prey from the squid's approaching tentacles. The skin also contains light reflectors called iridophores and leucophores that, when activated, in milliseconds create changeable skin patterns of polarized light. Such skin camouflage may serve various functions, such as communication with nearby squid, prey detection, navigation, and orientation during hunting or seeking shelter. Neural control of the iridophores enabling rapid changes in skin iridescence appears to be regulated by a cholinergic process affecting reflectin proteins.

 

Some mesopelagic squid such as the firefly squid (Watasenia scintillans) and the midwater squid (Abralia veranyi) use counter-illumination camouflage, generating light to match the downwelling light from the ocean surface. This creates the effect of countershading, making the underside lighter than the upperside.

 

Counter-illumination is also used by the Hawaiian bobtail squid (Euprymna scolopes), which has symbiotic bacteria (Aliivibrio fischeri) that produce light to help the squid avoid nocturnal predators. This light shines through the squid's skin on its underside and is generated by a large and complex two-lobed light organ inside the squid's mantle cavity. From there, it escapes downwards, some of it travelling directly, some coming off a reflector at the top of the organ (dorsal side). Below there is a kind of iris, which has branches (diverticula) of its ink sac, with a lens below that; both the reflector and lens are derived from mesoderm. The squid controls light production by changing the shape of its iris or adjusting the strength of yellow filters on its underside, which presumably change the balance of wavelengths emitted. Light production shows a correlation with intensity of down-welling light, but it is about one third as bright; the squid can track repeated changes in brightness. Because the Hawaiian bobtail squid hides in sand during the day to avoid predators, it does not use counter-illumination during daylight

 

PREDATOR DISTRACTION WITH INK

Squid distract attacking predators by ejecting a cloud of ink, giving themselves an opportunity to escape. The ink gland and its associated ink sac empties into the rectum close to the anus, allowing the squid to rapidly discharge black ink into the mantle cavity and surrounding water. The ink is a suspension of melanin particles and quickly disperses to form a dark cloud that obscures the escape manoeuvres of the squid. Predatory fish may also be deterred by the alkaloid nature of the discharge which may interfere with their chemoreceptors.

 

NERVOUS SYSTEM AND SENSE ORGANS

Cephalopods have the most highly developed nervous systems among invertebrates. Squids have a complex brain in the form of a nerve ring encircling the oesophagus, enclosed in a cartilaginous cranium. Paired cerebral ganglia above the oesophagus receive sensory information from the eyes and statocysts, and further ganglia below control the muscles of the mouth, foot, mantle and viscera. Giant axons up to 1 mm in diameter convey nerve messages with great rapidity to the circular muscles of the mantle wall, allowing a synchronous, powerful contraction and maximum speed in the jet propulsion system.

 

The paired eyes, on either side of the head, are housed in capsules fused to the cranium. Their structure is very similar to that of a fish eye, with a globular lens that has a depth of focus from 3 cm to infinity. The image is focused by changing the position of the lens, as in a camera or telescope, rather than changing the shape of the lens, as in the human eye. Squid adjust to changes in light intensity by expanding and contracting the slit-shaped pupil. Deep sea squids in the family Histioteuthidae have eyes of two different types and orientation. The large left eye is tubular in shape and looks upwards, presumably searching for the silhouettes of animals higher in the water column. The normally-shaped right eye points forwards and downwards to detect prey.

 

The statocysts are involved in maintaining balance and are analogous to the inner ear of fish. They are housed in cartilaginous capsules on either side of the cranium. They provide the squid with information on its body position in relation to gravity, its orientation, acceleration and rotation, and are able to perceive incoming vibrations. Without the statocysts, the squid cannot maintain equilibrium. Squid appear to have limited hearing, but the head and arms bear lines of hair-cells that are weakly sensitive to water movements and changes in pressure, and are analogous in function to the lateral line system of fish.

 

REPRODUCTIVE SYSTEM

The sexes are separate in squid, there being a single gonad in the posterior part of the body with fertilisation being external, and usually taking place in the mantle cavity of the female. The male has a testis from which sperm pass into a single gonoduct where they are rolled together into a long bundle, or spermatophore. The gonoduct is elongated into a "penis" that extends into the mantle cavity and through which spermatophores are ejected. In shallow water species, the penis is short, and the spermatophore is removed from the mantle cavity by a tentacle of the male, which is specially adapted for the purpose and known as a hectocotylus, and placed inside the mantle cavity of the female during mating.The female has a large translucent ovary, situated towards the posterior of the visceral mass. From here, eggs travel along the gonocoel, where there are a pair of white nidamental glands, which lie anterior to the gills. Also present are red-spotted accessory nidamental glands containing symbiotic bacteria; both organs are associated with nutrient manufacture and forming shells for the eggs. The gonocoel enters the mantle cavity at the gonopore, and in some species, receptacles for storing spermatophores are located nearby, in the mantle wall. In shallow-water species of the continental shelf and epipelagic or mesopelagic zones, it is frequently one or both of arm pair IV of males that are modified into hectocotyli. However, most deep-sea squid lack hectocotyl arms and have longer penises; Ancistrocheiridae and Cranchiinae are exceptions. Giant squid of the genus Architeuthis are unusual in that they possess both a large penis and modified arm tips, although whether the latter are used for spermatophore transfer is uncertain. Penis elongation has been observed in the deep-water species Onykia ingens; when erect, the penis may be as long as the mantle, head, and arms combined. As such, deep-water squid have the greatest known penis length relative to body size of all mobile animals, second in the entire animal kingdom only to certain sessile barnacles.

 

DIGESTIVE SYSTEM

Like all cephalopods, squids are predators and have complex digestive systems. The mouth is equipped with a sharp, horny beak mainly made of chitin and cross-linked proteins, which is used to kill and tear prey into manageable pieces. The beak is very robust, but does not contain minerals, unlike the teeth and jaws of many other organisms; the cross-linked proteins are histidine- and glycine-rich and give the beak a stiffness and hardness greater than most equivalent synthetic organic materials. The stomachs of captured whales often have indigestible squid beaks inside. The mouth contains the radula, the rough tongue common to all molluscs except bivalvia, which is equipped with multiple rows of teeth.[6] In some species, toxic saliva helps to control large prey; when subdued, the food can be torn in pieces by the beak, moved to the oesophagus by the radula, and swallowed.

 

The food bolus is moved along the gut by waves of muscular contractions (peristalsis). The long oesophagus leads to a muscular stomach roughly in the middle of the visceral mass. The digestive gland, which is equivalent to a vertebrate liver, diverticulates here, as does the pancreas, and both of these empty into the caecum, a pouch-shaped sac where most of the absorption of nutrients takes place. Indigestible food can be passed directly from the stomach to the rectum where it joins the flow from the caecum and is voided through the anus into the mantle cavity. Cephalopods are short-lived, and in mature squid, priority is given to reproduction; the female Onychoteuthis banksii for example, sheds its feeding tentacles on reaching maturity, and becomes flaccid and weak after spawning.

 

CARDIOVASCULAR AND EXCRETORY SYSTEMS

The squid mantle cavity is a seawater-filled sac containing three hearts and other organs supporting circulation, respiration, and excretion. Squid have a main systemic heart that pumps blood around the body as part of the general circulatory system, and two branchial hearts. The systemic heart consists of three chambers, a lower ventricle and two upper atria, all of which can contract to propel the blood. The branchial hearts pump blood specifically to the gills for oxygenation, before returning it to the systemic heart. The blood contains the copper-rich protein hemocyanin, which is used for oxygen transport at low ocean temperatures and low oxygen concentrations, and makes the oxygenated blood a deep, blue color. As systemic blood returns via two vena cavae to the branchial hearts, excretion of urine, carbon dioxide, and waste solutes occurs through outpockets (called nephridial appendages) in the vena cavae walls that enable gas exchange and excretion via the mantle cavity seawater.

 

BUOYANCY

Unlike nautiloids which have gas-filled chambers inside their shells which provide buoyancy, and octopuses which live near and rest on the seabed and do not require to be buoyant, many squid have a fluid-filled receptacle, equivalent to the swim bladder of a fish, in the coelom or connective tissue. This reservoir acts as a chemical buoyancy chamber, with the heavy metallic cations typical of seawater replaced by low molecular-weight ammonium ions, a product of excretion. The small difference in density provides a small contribution to buoyancy per unit volume, so the mechanism requires a large buoyancy chamber to be effective. Since the chamber is filled with liquid, it has the advantage over a swim bladder of not changing significantly in volume with pressure. Glass squids in the family Cranchiidae for example, have an enormous transparent coelom containing ammonium ions and occupying about two-thirds the volume of the animal, allowing it to float at the required depth. About half of the 28 families of squid use this mechanism to solve their buoyancy issues.

 

LARGEST AND SMALLEST

The majority of squid are no more than 60 cm long, although the giant squid may reach 13 m. The smallest species are probably the benthic pygmy squids Idiosepius, which grow to a mantle length of 10 to 18 mm, and have short bodies and stubby arms.

 

In 1978, sharp, curved claws on the suction cups of squid tentacles cut up the rubber coating on the hull of the USS Stein. The size suggested the largest squid known at the time.

 

In 2003, a large specimen of an abundant but poorly understood species, Mesonychoteuthis hamiltoni (the colossal squid), was discovered. This species may grow to 10 m in length, making it the largest invertebrate. In February 2007, a New Zealand fishing vessel caught the largest squid ever documented, weighing 495 kg and measuring around 10 m off the coast of Antarctica. Dissection showed that the eyes, used to detect prey in the deep Southern Ocean, exceeded the size of footballs; these may be among the largest eyes ever to exist in the animal kingdom.

 

DEVELOPMENT

The eggs of squid are large for a mollusc, containing a large amount of yolk to nourish the embryo as it develops directly, without an intervening veliger larval stage. The embryo grows as a disc of cells on top of the yolk. During the gastrulation stage, the margins of the disc grow to surround the yolk, forming a yolk sac, which eventually forms part of the animal's gut. The dorsal side of the disc grows upwards and forms the embryo, with a shell gland on its dorsal surface, gills, mantle and eyes. The arms and funnel develop as part of the foot on the ventral side of the disc. The arms later migrate upwards, coming to form a ring around the funnel and mouth. The yolk is gradually absorbed as the embryo grows. Some juvenile squid live higher in the water column than do adults. Squids tend to be short-lived; Loligo for example lives from one to three years according to species, typically dying soon after spawning.

 

n a well-studied bioluminescent species, the Hawaiian bobtail squid, a special light organ in the squid's mantle is rapidly colonized with Aliivibrio fischeri bacteria within hours of hatching. This light-organ colonization requires this particular bacterial species for a symbiotic relationship; no colonization occurs in the absence of A. fischeri. Colonization occurs in a horizontal manner, such that the hosts acquires its bacterial partners from the environment. The symbiosis is obligate for the squid, but facultative for the bacteria. Once the bacteria enter the squid, they colonize interior epithelial cells in the light organ, living in crypts with complex microvilli protrusions. The bacteria also interact with hemocytes, macrophage-like blood cells that migrate between epithelial cells, but the mechanism and function of this process is not well understood. Bioluminescence reaches its highest levels during the early evening hours and bottoms out before dawn; this occurs because at the end of each day, the contents of the squid's crypts are expelled into the surrounding environment. About 95% of the bacteria are voided each morning before the bacterial population builds up again by nightfall.

 

BEHAVIOUR

LOCOMOTION

Squid can move about in several different ways. Slow movement is achieved by a gentle undulation of the muscular lateral fins on either side of the trunk which drives the animal forward. A more common means of locomotion providing sustained movement is achieved using jetting, during which contraction of the muscular wall of the mantle cavity provides jet propulsion.

 

Slow jetting is used for ordinary locomotion, and ventilation of the gills is achieved at the same time. The circular muscles in the mantle wall contract; this causes the inhalant valve to close, the exhalant valve to open and the mantle edge to lock tightly around the head. Water is forced out through the funnel which is pointed in the opposite direction to the required direction of travel. The inhalant phase is initiated by the relaxation of the circular muscles causes them to stretch, the connective tissue in the mantle wall recoils elastically, the mantle cavity expands causing the inhalant valve to open, the exhalant valve to close and water to flow into the cavity. This cycle of exhalation and inhalation is repeated to provide continuous locomotion.

 

Fast jetting is an escape response. In this form of locomotion, radial muscles in the mantle wall are involved as well as circular ones, making it possible to hyper-inflate the mantle cavity with a larger volume of water than during slow jetting. On contraction, water flows out with great force, the funnel always being pointed anteriorly, and travel is backwards. During this means of locomotion, some squid exit the water in a similar way to flying fish, gliding through the air for up to 50 m, and occasionally ending up on the decks of ships.

 

FEEDING

Squid are carnivores, and, with their strong arms and suckers, can overwhelm relatively large animals efficiently. Prey is identified by sight or by touch, grabbed by the tentacles which can be shot out with great rapidity, brought back to within reach of the arms, and held by the hooks and suckers on their surface. In some species, the squid's saliva contains toxins which act to subdue the prey. These are injected into its bloodstream when the prey is bitten, along with vasodilators and chemicals to stimulate the heart, and quickly circulate to all parts of its body. The deep sea squid Taningia danae has been filmed releasing blinding flashes of light from large photophores on its arms to illuminate and disorientate potential prey.

 

Although squid can catch large prey, the mouth is relatively small, and the food must be cut into pieces by the chitinous beak with its powerful muscles before being swallowed. The radula is located in the buccal cavity and has multiple rows of tiny teeth that draw the food backwards and grind it in pieces. The deep sea squid Mastigoteuthis has the whole length of its whip-like tentacles covered with tiny suckers; it probably catches small organisms in the same way that flypaper traps flies. The tentacles of some bathypelagic squids bear photophores which may bring food within its reach by attracting prey.

 

Squid are among the most intelligent invertebrates. For example, groups of Humboldt squid hunt cooperatively, spiralling up through the water at night and coordinating their vertical and horizontal movements while foraging.

 

REPRODUCTION

Courtship in squid takes place in the open water and involves the male selecting a female, the female responding, and the transfer by the male of spermatophores to the female. In many instances, the male may display to identify himself to the female and drive off any potential competitors.[46] Elaborate changes in body patterning take place in some species in both agonistic and courtship behaviour. The Caribbean reef squid (Sepioteuthis sepioidea), for example, employs a complex array of colour changes during courtship and social interactions and has a range of about 16 body patterns in its repertoire.

 

The pair adopt a head-to-head position, and "jaw locking" may take place, in a similar manner to that adopted by some cichlid fish. The heterodactylus of the male is used to transfer the spermatophore and deposit it in the female's mantle cavity in the position appropriate for the species; this may be adjacent to the gonopore or in a seminal receptacle.

 

The sperm may be used immediately or may be stored. As the eggs pass down the oviduct, they are wrapped in a gelatinous coating, before continuing to the mantle cavity, where they are fertilised. In Loligo, further coatings are added by the nidimental glands in the walls of the cavity and the eggs leave through a funnel formed by the arms. The female attaches them to the substrate in strings or groups, the coating layers swelling and hardening after contact with sea water. Loligo sometimes forms breeding aggregations which may create a "community pile" of egg strings. Some pelagic and deep sea squid do not attach their egg masses, which float freely.

 

ECOLOGY

Squid mostly have an annual life cycle, growing fast and dying soon after spawning. The diet changes as they grow but mostly consists of large zooplankton and small nekton. In Antarctica for example, krill is the main constituent of the diet, with other food items being amphipods, other small crustaceans, and large arrow worms. Fish are also eaten, and some squid are cannibalistic.

 

As well as occupying a key role in the food chain, squid are an important prey for predators including sharks, sea birds, seals and whales. Juvenile squid provide part of the diet for worms and small fish. When researchers studied the contents of the stomachs of elephant seals in South Georgia, they found 96% squid by weight. In a single day, a sperm whale can eat 700 to 800 squid, and a Risso's dolphin entangled in a net in the Mediterranean was found to have eaten angel clubhook squid, umbrella squid, reverse jewel squid and European flying squid, all identifiable from their indigestible beaks. Ornithoteuthis volatilis, a common squid from the tropical Indo-Pacific, is predated by yellowfin tuna, longnose lancetfish, common dolphinfish and swordfish, the tiger shark, the scalloped hammerhead shark and the smooth hammerhead shark. Sperm whales also hunt this species extensively as does the brown fur seal. In the Southern Ocean, penguins and wandering albatrosses are major predators of Gonatus antarcticus.

 

HUMAN USES

IN LITERATUR AND ART

Giant squid have featured as monsters of the deep since classical times. Giant squid were described by Aristotle (4th century BC) in his History of Animals and Pliny the Elder (1st century AD) in his Natural History. The Gorgon of Greek mythology may have been inspired by squid or octopus, the animal itself representing the severed head of Medusa, the beak as the protruding tongue and fangs, and its tentacles as the snakes. The six-headed sea monster of the Odyssey, Scylla, may have had a similar origin. The Nordic legend of the kraken may also have derived from sightings of large cephalopods.

 

In literature, H. G. Wells' short story "The Sea Raiders" featured a man-eating squid species Haploteuthis ferox.[59] The science fiction writer Jules Verne told a tale of a kraken-like monster in his 1870 novel Twenty Thousand Leagues Under the Sea.

 

AS FOOD

Squid form a major food resource and are used in cuisines around the world, notably in Japan where it is eaten as ika sōmen, sliced into vermicelli-like strips; as sashimi; and as tempura. Three species of Loligo are used in large quantities, L. vulgaris in the Mediterranean (known as Calamar in Spanish, Calamaro in Italian); L. forbesii in the Northeast Atlantic; and L. pealei on the American East Coast. Among the Ommastrephidae, Todarodes pacificus is the main commercial species, harvested in large quantities across the North Pacific in Canada, Japan and China.

 

In English-speaking countries, squid as food is often called calamari, adopted from Italian into English in the 17th century. Squid are found abundantly in certain areas, and provide large catches for fisheries. The body can be stuffed whole, cut into flat pieces, or sliced into rings. The arms, tentacles, and ink are also edible; the only parts not eaten are the beak and gladius (pen). Squid is a good food source for zinc and manganese, and high in copper, selenium, vitamin B12, and riboflavin.

 

COMMERCIAL FISHING

According to the FAO, the cephalopod catch for 2002 was 3,173,272 tonnes. Of this, 2,189,206 tonnes, or 75.8 percent, was squid. The following table lists squid species fishery catches that exceeded 10,000 tonnes in 2002.

 

IN BIOMIMICRY

Prototype chromatophores that mimic the squid's adaptive camouflage, have been made by Bristol University researchers using an electroactive dielectric elastomer, a flexible "smart" material that changes its colour and texture in response to electrical signals. The researchers state that their goal is to create an artificial skin that provides rapid active camouflage.

 

The squid giant axon inspired Otto Schmitt to develop a comparator circuit with hysteresis now called the Schmitt trigger, replicating the axon's propagation of nerve impulses.

 

WIKIPEDIA

There were lots of bees and wasps on the sunny side of a fountain. I presume they were eating the algae that was growing at the edge of the water rather than drinking the water. Thanks for your comments - so they are drinking the water then.

 

August 2016

 

All rights reserved. This photo is not authorized for use on your blogs, pin boards, websites or use in any other way without specific written permission.

Nuthatch checking it's surroundings after stashing seeds underneath this log.

Taken late in the day and a stop under exposed so quality not the best.

Latest from Fattercrombie & Fitch: crimson turtleneck and light beige trousers.

 

[Uploaded using Charlie 1.60]

Fair question! I suppose I'm looking at a ruddy turnstone. Sure, I've seen them before. They were all wild, living off their wits, spread out.

 

Here there are flocks. Not just flocks. Flocks of habituated urban birds scrabbling for crumbs; scraps of a livelihood. I'm cautious of them about my feet they are so ingenuous.

 

I can't say I like this experience. I don't suppose these birds were trained for this lifestyle. Instead they've likely evolved behaviours; adapting to the economy of scrounging in a tourist-popular fishing port. They don't add to the scene in the way they do on a wild coast.

 

Oh well, this is another thing to note down and move on. I have St Ives for a short while and I'm not keen to waste it on what were once shorebirds.

  

Jay

Greater Manchester Police has praised the behaviour of visitors to Manchester during a weekend of sport in the city.

 

On Friday 20 May 2016, the Great City Games saw a number of athletes compete in various events on a purpose-built athletics arena in Albert Square and track on Deansgate.

 

The following day (21 May 2016) saw Manchester United beat Crystal Palace 2-1 to win the FA Cup at Wembley Stadium, with a number of fans watching the match in public venues throughout Greater Manchester.

 

The weekend extravaganza concluded on Sunday 22 May 2016 with over 30,000 lining up to take part in the Great Manchester Run before England defeated Turkey 2-1 at the Etihad Stadium in a UEFA Euro 2016 warm-up match in the evening.

 

The events saw tens of thousands of visitors to the city centre, creating a buzzing and carnival-like atmosphere.

 

Assistant Chief Constable John O’Hare said: “This has been a fantastic weekend for Manchester and the atmosphere in the city has been superb from start to finish.

 

“It was great to see so many pictures of smiling faces and people having a good time and I hope everyone who has visited the city this weekend will be going away with some great memories.

 

“I would like to thank everyone who has played a key role in ensuring that the weekend has been successful.”

 

For more information about Policing in Greater Manchester please visit www.gmp.police.uk

 

To report crime call police on 101 the national non-emergency number.

 

You can also call anonymously with information about crime to Crimestoppers on 0800 555 111. Crimestoppers is an independent charity who will not want your name, just your information. Your call will not be traced or recorded and you do not have to go to court or give a statement.

 

Thats the reaction after an orgasm :-)

Sometimes it's better not to know what happens in the background

Just out of shot a parent as just resurfaced with a fish, the race is on.

Name: Bernard Andersen

Arrested for: not given

Arrested at: North Shields Police Station

Arrested on: 15 November 1905

Tyne and Wear Archives ref: DX1388-1-81-Bernard Anderson

 

The Shields Daily News for 15 November 1905 reports:

 

“THREE MONTHS’ IMPRISONMENT.

 

Bernard Andersen, a Swede, was sent to prison for three months for indecent conduct.”

 

These images are a selection from an album of photographs of prisoners brought before the North Shields Police Court between 1902 and 1916 in the collection of Tyne & Wear Archives (TWA ref DX1388/1).

 

(Copyright) We're happy for you to share this digital image within the spirit of The Commons. Please cite 'Tyne & Wear Archives & Museums' when reusing. Certain restrictions on high quality reproductions and commercial use of the original physical version apply though; if you're unsure please email archives@twmuseums.org.uk.

The adult falcon looked in 2 sides of the large windows in different areas (top and bottom). It's almost looking for a pigeon to grab which is very unusual for them as they are ariel predators. Be interesting to know. I've never seen this before. Deliberate attempts to look where Feral Pigeons are nesting

Heute in ZOOM Erlebniswelt Gelsenkirchen.

You cant leave anything lying around with these guys around! Thanks to kevinmboots77 for the idea.

By now it was raining quite hard, so I grabbed a couple of shots from the other side of the road before going in.

 

Hello I said to the churchwarden, we've come to photograph the church.

 

Oh I don't know if that's possible, you might be journalists. What do you want the pictures for?

 

I explained about the website and liking churches and that we had come from Dover to see this church. I gave he my Moo card, and she said it was OK. And then would not shut up, she told us all about the history of the church, the town, businesses. All nice, but I wanted to snap the church.

 

In the end, Jools took over and I set about snapping. And very fine it is too.

 

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A huge church that is accessed most days through the coffee shop next door. Saxon in its origins it was extended many times as befits a market town on the main London-Dover road. Today it consists of nave and chancel with aisles and chapels. Much remodelled in the nineteenth century (by Blomfield) and again late in the 20th it may lack atmosphere, but it certainly does not lack appeal.... or its part in national history. The body of Henry V rested here overnight on its journey back from France in 1422. Behind the painting in a medieval vestry, itself a rarity. There are lots of brasses and a large monument to John Spilman who introduced papermaking here in the seventeenth century. In the south chapel is a huge wall painting of St. George - the largest medieval painting in Kent.

 

www.kentchurches.info/church.asp?p=Dartford+1

 

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DARTFORD

LIES the next parish eastward from Crayford, on the high road from London to Dover, about fifteen miles from the former. It was called in Saxon Derentford, in Latin Derenti Vadum, signifying the forde or passage over the river Derent. (fn. 1) In Domesday it is written Tarentefort.

 

THIS PARISH takes within its bounds almost the whole both of Dartford-heath and the Brent. It contains about 4300 acres of land. The town has about four hundred houses and about two thousand five hundred inhabitants. The upland parts of the parish are but thin and gravelly, the crops of which are greatly increased by the culture of turnips; the vallies are a sertile and rich loam, the northern part of the parish is marsh land, which reaches to the Thames, containing about eight hundred acres, none of which is ever ploughed. The town of Dartford is situated in a valley, between two hills, which rise suddenly and sleep at each end of it. On that at the western extremity are chalk pits, which have been worked underneath to a considerable extent, and have rather a fearful and dangerous appearance to travellers; the opposite hill is a deep sandy loam. Dartford is at present a handsome and wealthy town, still increasing in size and inhabitants, the principal street of which is the great thoroughfare from London to Dover, on which there are built several good inns. From this street southward branches off the high road through Farningham to Sevenoke, in which stands Horseman's-place, now used, with the gardens, by a public gardener; northward from the high street is the Water-lane (so called from the little stream, the Cranford, which rises about a mile and a half southward of the town, at Hawley, which runs through it) and leads to the wharss at the water side, not far distant from which stands the Place-house, formerly the priory, with the buildings belonging to it, now used as a farm house and offices, adjoining to which is a piece of land, inclosed with a wall, formerly belonging to the priory, exceeding rich, which has been for many years been made use of as a public garden ground. The artichokes growing in it are noted for being the largest and best flavoured of any brought to the London markets, and are called, for distinction sake, the Dartford artichoke.

 

There is a good market for corn and provisions here on a Saturday, weekly; and a fair yearly, on the 2d and 3d of August. The old market house and shambles stood very inconveniently in the middle of the high street, but they were removed some years ago, and the present market place and shambles were built more commodiously elsewhere, by public subscription, to the great embellishment of the town, and the satisfaction of all travellers; at the same time the old uneasy pavement through the town was removed, and a new road of gravel made in its room, with a handsome footway of curbed stone on each side; near the east end of it stands the church, almost adjoining to the river Darent, which here crosses the high road under a handsome bridge. In king Edward III.'s reign there appears to have been no bridge here, the passage or ferry over the Darent at this place being valued among the rents of the manor; however, there was one built before the end of king Henry VI.'s reign, but it was one most narrow, steep, and dangerous for travellers, which continued so till not many years since it was altered to its present more commodious state, at the public charge of the county. A little below this bridge, the Darent becomes navigable for barges; and at the distance of about two miles, receiving the Cray into its channel, at a like distance empties itself into the Thames. On this creek there was formerly a considerable fishery, as appears by the records before mentioned; for so late as king James I.'s reign, the royal manor of Dartford received for the fishery six salmons yearly, a kind of fish now unknown here; and the manor of Dartford priory received a yearly rent of fifty pounds for a fishery likewise here at the same time; but no fishery at this time exists, nor has for many years past.

 

The trade and manufacture carried on by the several mills on the Darent contribute much to the flourishing state this town is in at present; for besides the powder-mills, first erected by Sir John Spilman as a paper mill, as before mentioned, situated a quarter of a mile above the town; there is a paper mill at a small distance below it, where there was one so early as 1590, erected by one Geoffry Box of Liege, for the cutting of iron bars into rods, being the first supposed to be erected for this purpose in England, and for the more easy converting of that metal to different uses; lower down, at the east end of the town, are two corn mills, and farther below bridge the ruins of the mill, employed as a cotton manufactory, which was burned down in 1796, and now lies in ruins. It was before made use of as a sawing mill, and before that as a brasel mill, for the slitting of iron bars into rods, nails, &c. being first erected for that purpose by John Browne, soon after the death of king Charles I. Near this is the public wharf, to which hoys and barges come up from the Thames. To this wharf is brought the produce of the woods in this neighbourhood, which are of considerable extent, and the manufactures, which are here shipped for the London market, as are the goods for the subsistence of the town and vicinity of it from the metropolis.

 

In the return of the survey, made of the several maritime places, in this county, by order of queen Elizabeth, in her 8th year, Dartford is said to contain houses inhabited, 182; persons lacking habitation, 6; keys or landing places; 4; ships and boats, 7; three of three tons, one of six, two of ten, one of fifteen; persons for carriage from Dartford to London, and so back again, 14; Sir Thomas Walsingham, steward of the town; Mr. Asteley, keeper of the queen's house; John Beer's; the wardens of Rochester-bridge.

 

In the reign of king Henry III. the archbishop of Cologne was sent hither, with several noblemen, by the emperor Frederick, to demand Isabella, the king's sister, in marriage, which was solemnised by proxy in this town, and she was then delivered to them, to be carried over. In 1331, king Edward III. at his return from France, held a famous tornament in this town. In the 5th year of king Richard II. a great commotion of the common people begun at this place, occasioned by Wat Tyler's having beat out the brains of one of the collectors of the poll tax, on account of his insolent behaviour to his daughter. The people, who were in general discontented, being inflamed by this circumstance, broke out into open rebellion, and he soon found himself at the head of one hundred thousand men. (fn. 2)

 

Thus attended, he marched directly to London, freeing, in the mean time, the prisoners detained in the public goals; among these was a priest, in the neighbourhood of Maidstone, one John Ball, vulgarly called John Straw, who, by his seditious sermons, had raised the people's sury to the utmost heighth, insomuch that, in conformity to his maxims, they resolved to destroy all the nobility and lawyers in the realm, for he had persuaded them that all men, being the sons of Adam, there ought to be no distinction; and, confequently, it was their duty to reduce the world to a perfect equality; in consequence of which he preached to the people on these rhymes:

 

"When Adam delse, and Eve span,

"Who was then a gentleman ?"

 

The king, hearing they were advanced as far as Blackheath, sent to know their demands, to which, returning a most insolent answer, they immediately marched towards London, and took possession of the borough of Southwark; and the gates of London bridge being thrown open to them by the citizens, they entered the city, where they committed every scene of barbarity that could be expected from such a body, guided solely by their sury. They then seized on the Tower, where they sound the archbishop and the lord treasurer, whom they immediately beheaded. Upon this the king, dreading the consequences of so powerful a body, repaired to Smithfield, with some few attendants, and sent a knight to Tyler, to come there and confer with him, which this rebel, with much deliberation, at last complied with. At this conference he behaved with such insolence, that William Walworth, lord mayor of London, who attended the king, without considering the consequences that would attend it, discharged such a blow at the rebel's head with his sword, that he instantly fell dead at his feet. However, contrary to expectation, the multitude were so terrified, that they threw down their arms, and sued for mercy; and were all, in the space of a few minutes, dispersed, without the effusion of any blood, except of their leader. (fn. 3)

 

About a mile south-westward from the town is the large plain, called DARTFORD. HEATH, containing about 500 acres of land. It lies high, and on a fine gravelly soil; on it there are a great many of those pits and holes so frequent in these parts. Some of these reach below the gravel as low as the chalk, others no farther than the sand and gravel; many of them have been stopped up of late years, to prevent the frequent accidents which happen of men and cattle falling into them. The occasion of their being first dug has been already explained, under the adjoining parish of Crayford. This heath has been much noted of late, as being the spot chosen by the corps of Toxopbilites, under the appellation of the Royal Kentish Bowmen, for whose use a house has been fitted up at the western side of the heath, not far from Baldwin's, and is now distinguished by the name of the Lodge, being the scene of their exercise and recreation; at which times, on their gala days, butts, apartments, and company, have made the most splen did and costly appearance. It is as delightful and pleasant a spot as any in these parts.

 

Less than half a mile eastward from the town, the high road to Rochester crossing it, lies another heath, called DARTFORD-BRENT, vulgarly the BRIMPT. This place is famous for the encampment of the army of Richard Plantagenet, duke of York, in 1452, whilst he waited to obtain a parley with king Henry VI. who then lay encamped on Blackheath. In the year 1648 General Fairfax's army rendevouzed here.

 

The ROMAN-ROAD shews itself very conspicuously on the south side of the high road between Dartford and the Brent, and when it comes to the latter, it shapes its course more to the south south-east, leaving the high road at a greater distance, on the lefthand, and entering among the inclosures and woods, in its way to a hamlet called Stonewood, it goes on to Wingfield-bank, and thence to Shinglewell, towards Rochester.

 

At a small distance southward from the Romanroad on the Brent, close to the road to Greenstedgreen, are three small barrows, which seem to have been plundered of their contents.

 

¶The gravel-pit at the entrance of the Brent from Dartford was, whilst the affizes were held in this town, which was frequently, at the latter end of queen Elizabeth's reign, the place for the public execution of criminals; and in 1772, in digging for gravel here, eight human skeletons were sound, lying contiguous to each other; most probably the remains of some of those unhappy convicts. This spot was likewise made use of in the reign of queen Mary, for the execution of those who suffered for religion.

 

Our HERBALISTS have taken notice of several scarce plants and herbs sound here:

 

The camæpytis, herb ivy, or ground pine, not only here, but in the adjoining parishes.

 

Ruta muraria five salvia vitæ, stone rue, or rue maidenbair, on the wall of the church-yard.

 

Aphaca, small yellow fetch, in the corn-fields about this place.

 

Buckthorne, in the bedges of this place.

 

The juniper tree grows in plenty on the downs southward of Dartford-brent.

 

Mentastrum, horse mint; valde ramosum flore violaceo rubro.

 

Orchis five tragorchis max. the greatest goat stones, between Crayford and Dartford.

 

Orchis hermaphroditica, the butterfly satirion; testiculus vulpinus spegodes, the humble bee orchis; orchis melittias, the bee orchis; orchis myodes, the fly satirion; are found on the downs, southward of Dartford brent.

 

The lizard orchis, has been found in the lane between Dartford and Darent.

 

Several forts of the orchis ornithophora are found in the meadows adjoining the river Darent, southward of this town.

 

Trisolium stellatum glabrum, smooth starry headed tresoil, in Dartford salt marshes. (fn. 4)

 

Charities.

THOMAS AUDITOR, alias BARNARD, gave by will, in 1536, an annuity of 3s. to buy peas, to be distributed among the poor, in the first week in Lent, payable out of four acres of land, called Docklincrost, which bequest has not been paid for many years.

 

WILLIAM VAUGHAN gave by deed, in 1596, a rent, to be distributed quarterly to the most poor inhabitants of Dartford, out of a house and garden, vested in trustees, and of the annual produce of 13l. 4s.

 

JEROME WARRAM gave by will, in 1570, for the use of the poor, a house and garden, in the occupation of Mrs. Bugden, of the annual produce of 5s.

 

MRS. CATHARINE BAMME gave by deed, in 1572, among other charitable bequests, 20s. to the poor of this parish, to be paid out of an messuage and lands in Gillingham, vested in Edward Taylor, of the annual produce of that sum.

 

JOHN BYER gave by will, in 1572, for the habitation of the poor of this parish, nine alms houses, in Lowfield, adjoining southward to Horseman's-place, and endowed them with a barn and several pieces of land, in the occupation of Mrs. Glover and Mr. Fleet; the former at 17l. the latter at 5l. annual rent, and for the habitation of four poor aged people, and 20d. to be paid quarterly to each of them; now inhabited by paupers; annual produce 1l. 6s. 8d.

 

JOHN BARTON gave by will, in 1613, the interest of 130l. yearly, to be bestowed on bread, and distributed to the poor by the vicar and churchwardens. N. B. With this money, in 1623, the parish purchased by deed, of Francis Goldsmith and others, thirteen acres of land in Crayford parish, and a house in Dartford, the former vested in William Flint and others, at 12l. per annum rent; the latter in William Nettlefold, at 11l. 10s. per annum; on condition that 20s. should be yearly distributed to the poor on Shrove Sunday, as his gift, out of the rents of the lands purchased of him by Barton's money. He agreed to abate 15l. out of the purchase money; annual produce 1l.

 

WILLIAM REYNOLDS and WILLIAM HARRISON gave by will, in 1623, the interest of 50l. and 10l. to be laid out in bread, and distributed among the poor every Sunday in the year,

 

N. B. With these two gifts were purchased a house and piece of land belonging to it, which house has been taken down, and four new houses have been built on the ground, with monies borrowed upon them, which money the rents have discharged. The houses are let to several tenants, at the yearly rent of 5l. each; 2s. worth of bread have been yearly distributed every Sunday, out of the rent of these houses, as was stipulated when they were purchased; the annual produce 20l. per annum.

 

ROBERT ROGERS gave by deed, in 1629, rent to be distributed among the poor on Easter Monday, payable out of a house and yard, vested in Mrs. Catharine Tasker; annual produce 4l.

 

JONATHAN BRETT gave by deed, in 1629, for the relief of the poor inhabitants of this parish, four acres of land, vested in Mr. George Hardres, of the annual produce of 9l.

 

THOMAS COOPER in 1629, gave an annuity, to be distributed to the poor in bread, payable out of woodland in Bexley parish, in the occupation of James Craster, of the annual produce of 1l.

 

ANTHONY POULTER gave by will, in 1629, an annuity of 20s. to be distributed by the minister and churchwardens on Easter day, payable out of a house in Dartford, occupied by Mrs. Pettit, of the annual produce of 1l.

 

JOHN TWISLETON, esq. gave by deed, in 1660, certain rent, to be applied, one-third of it to the alms houses, and the other twothirds to be given to the poor, issuing out of three acres of land, in the occupation of Edward Rawlins, of the annual produce of 5l. 6s. 3d.

 

JOHN ROUND, in 1682, gave an annuity, to be distributed among the poor on Christmas day, payable out of the Bell inn, in Dartford, in the occupation of John Elliot, of the annual produce of 1l.

 

THE REV. CHARLES CHAMBERS gave by will, in 1745, the sum of 50l. vested in the 3 per cents. the interest to be distributed by the minister on Christmas day, among twenty-four poor persons, twenty of whom to be widows, annual produce 1l. 10s.

 

JOHN RANDALL gave by will, in 1771, 200l. now vested in the 3 per cent. the interest to be distributed among poor housekeepers and widows, at 5s. each; annual produce 7l. 8s. 6d. and he gave 100l. since, vested in like manner, the interest to be laid out in bread, and distributed to the poor on Sundays; annual produce 3l. 14s. 3d.

 

A PERSON UNKNOWN gave three houses for poor parishioners, to dwell in, now inhabited by paupers.

 

A PIECE OF LAND, on part of which the present workhouse was erected in 1728, by voluntary subscriptions; the other part, used as a garden to it, was given by a person unknown.

 

This land was let in 1720, for the use of the poor at 1l. per ann.

 

CHRISTOPHER HEATH gave lands to the next of kin of Ellen Sherrington, on condition that they should pay yearly out of them, to the use of the poor, 1l. 6s. 8d. and to the churchwardens and their successors, to the reparation of the church, 1l. 13s. 4d.

 

JOHN BEALE, of Swanscombe, devised 40s. per annum, towards the maintenance of a schoolmaster in Dartford, to be paid out of a messuage, called Hamanslay's, in Halsted, formerly occupied by William Watson.

 

THIS PARISH is within the ECCLESIASTICAL JURISDICTION of the deanry of Dartford, and diocese of Rochester. The church, which is dedicated to the Holy Trinity, stands near the east end of the town, and is a large handsome building, consisting of three isles' and two chancels. In 1793, the whole church was repaired and beautified by the parishioners, at the expence of twelve hundred pounds. The pavement within the altar rails, with the painting and gilding over it, was done at the charge of Charles Manning, gent. in 1702. The tower is at the west end of it, in which there is a clock and a good ring of bells; one of which, of the smaller size, used till of late to be constantly rung, as of old custom, at four o'clock every morning, and again at the time of curfew at night.

 

The church yard formerly surrounded it, but some few years ago that part of it, which was on the southerin side, was given to the public to make the road more commodious for passengers. There is another burying-ground belonging to this church at some little distance from it, adjoining the high London road at the top of the hill, eastward of the town, of which further mention will be made. It is situated on so high an eminence, that it overlooks even the top of the tower of the church.

 

Among other monuments and inscriptions in this church, are the following: In the great chancel, on the north side of the altar is a monument for Sir John Spilman, inclosed with iron railing; on it are his essigies in armour and that of his lady, kneeling at a desk, each with a book open, and over their heads, on a tablet of black marble, with an inscription in German text for both of them; he died in 1607; on the top of the monument his arms, Or, a serpent wreathed in pale azure, crested, gules, on a mount in base, vert, two slaunches, gules, each charged with three lions passant, or; beneath, on the tomb, are two coats, Spilman, as above, impaling argent, a man cloathed sable, with a long cap on, holding in his hand an olive branch proper, and standing on a mount, inverted, gules. On the south side of the chancel, an altar tomb, inclosed with rails, and inscription, for Clement Petit, esq. of Joyes, in this parish, whose paternal seat was at Dentelion, in Thanet, obt. 1717. Before the rails of the altar, on a grave stone, are the figures of a man and woman, in brass, under a canopy, with labels from their mouths; round the verge of the stone is an inscription in brass, in part torn away, for Richard Martyn, of Dartford, who died in 14 . . . . she died in 1402. Near it is another stone, which had the figure of a man, with a label from his mouth, and an inscription round the verge, all in brass, now lost; but an inscription in brass still remains, on a plate, for John Hornley, S. T. B. who died in 1477. On another adjoining, are the figures in brass of a woman and six children, that of the man is lost; beneath on a plate, is an inscription for capt. Arthur Bostocke, gent. who married Francis, second daughter of Francis Rogers, esq. he died in 1612. On a grave stone, before the step of the chancel, is the figure in brass, of a woman, and inscription, for Agnes, daughter of John Appleton, wife of Wm. Hesilt, one of the barons of the exchequer of Henry VI. afterwards of Robert, brother of Sir Tho. Molyngton, baron of Wemme; she died in 1454. On the south side of the chancel, a monument for Wm. Burgess, late citizen and salter of London, obt. 1640; arms, a sess sret between three rooks. On the same side, before the altar rails, a memorial for Nicholas Tooke, gent. of Dartford, obt. 1672, æt. 90; arms, Tooke, argent, on a chevton, sable, three plates of the field between three greyhounds heads erased, sable collared, or; but this is cut here very erroneous. On the north side, a memorial for Mr. Mark Fielder, 1753, æt. 91; on the south side, a memorial for Mr. Wm. Tasker, of this parish, ob. 1732; and for Wm. Tasker, jun. their second son, ob. 1733. In the south chancel, a mural monument for John Twisleton, esq. of Horseman's-place, son and heir of John Twisleton, esq. of Drax, in Yorkshire, who was uncle and heir of Sir Geo. Twisleton, bart. of Barley, in that county, the antient paternal seat of the family. A memorial for John Twisleton, esq. late of Horseman's-place, ob. 1721. At the east end an altar tomb, inclosed with wooden rails, and on the south of it an inscription for John Beer, of Dartford, who had Nicholas, Anne, and Dorothy; for Nicholas, who had Clement and Edward, and for Clement Beer, who had John and Clement, who both died, s. p. Edward Beer, their uncle, was their heir, and lived unmarried fifty-nine years, and died in 1627. On the north side, an inscription, shewing, that Christopher Twisleton, esq. of Barley, in Yorkshire, married Anne Beer, by whom he had George Twisleton, who had John Twisleton, and Edward Beer, dying, s. p. gave all his lands in Kent to John Twisleton above mentioned, who erected this monument in 1628. On the west side are two shields, one quarterly, 1st and 4th, quarterly, a canton ermine; 2d and 3d, on a fess, three garbs; the other the same arms, impaling a chevron. A grave stone, having a brass plate for John Beer, esq. of Dartford, and Alice and Joan, his wives, and also for Henry Beer, his son and heir, who married Anne Beer, widow of Rich. Howlett, gent. deceased, and had by her a son, Wm. Beer, deceased, which John Beer died in 1572, and Henry in 1574; above, are two coats in brass, both, a bear rampant, on a canton, five escallop shells. On a grave stone, the figures of a man and his two wives, with children and their shields of arms in brass, all of which are lost, excepting the second wife and four children, and a plate with the inscription, for Wm. Rothele, of Dartford, who died in 1464, and Beatrix and Joane, his wives, and their children. Another on the north side, on which were the figures of a man and woman, in brass, now lost, but part of the inscription remains, for Katryn Burlton, who died 1496, and Rich. Burlton, jantilman, her husband, who died 15 . . . the rest torn off. A mont for Margaret, relict of John Pitt, esq. predent of the S. Sea company at Vera Crux, ob. 1731, æt. 49, arms, Pitt impaling a chevron, ingrailed, betw. three eagles heads erased. In the middle isle, are several memorials of Manning; a grave stone in the south cross isle, having the figures in brass of a man between his two wives, and underneath those of fifteen children, with inscription in black letter, for Wm. Death, gent. principal of Staple's inn, who had two wives, Elizabeth and Anne, by the former he had ten sons and six daughters, ob. 1590, Elizabeth, 1582; above a shield of arms, being death, a grissin passant between three crescents, quartering four other coats. (fn. 46) In the north isle are memorials for the Round's, Woodin, Poulter, Dalling, and Chambers, all of this parish. There are many more memorials and tombs of respectable inhabitants of this populous town and parish, as well in the church as the two church yards, but they are by far too numerous for insertion in this place.

 

In the 7th year of king Edward III. Thomas de Woldham, bishop of Rochester, caused a new window to be made in the chancel of the church.

 

William the Conqueror confirmed the gift which Hamo his steward had made of the church of Tarentford, in the king's manor, to the church of St. Andrew of Rochester; (fn. 47) which king Henry I. confirmed, with the churches appendant to it, and the tithes of this parish in corn, pannage, cattle, money, and in all other things, in like manner as St. Austin held the church of Middleton, with the tithes of that parish, in the time of his father, (fn. 48) and also the tithes of his mills in Darenteford.

 

Gundulph, bishop of Rochester, who was elected to that see in the reign of the Conqueror, having recovered the manors and possessions of his church, which had been dissipated and made away with, separated his own maintenance from that of the monks, in which division he allotted this church, among others, to the support of the almonry, belonging to the convent. (fn. 49) The monks did not continue long in the possession of it, for bishop Gilbert de Glanvill, who came to the see in 1185, on pretence that his predecessor had impoverished the see by his too large donations to the priory, divested them of all right to this church, which he restored to the see of Rochester; however, he reserved and confirmed to the monks their antient pension from it. (fn. 50)

 

Laurence, bishop of Rochester, in 1253, reserving the tithes of sheaves, and of every kind of hay, demised this church, and all the small tithes, oblations, and obventions, and the tithes of sheaves arising in gardens and curtileges not being ploughed, to the convent of Rochester, at the rent of thirty-eight marcs per annum, on condition that they supplied the cure, and they were to deduct their pension of ten marcs, paid by the rector, out of it. (fn. 51) He afterwards obtained pope Innocent IV.'s leave to appropriate this church, during his life, to the use of his table, which he complained was so slenderly provided for; that he and his family had not at times common necessaries for food; the clear receipts for the bishop's table being but five hundred marcs, which were not more than sufficient for half the expence of it, and the receipts from his manors not exceeding sixty marcs per annum. (fn. 52) This was confirmed to the bishop and his successors by pope Alexander IV. and again by Clement IV. Bishop Laurence, on the appropriation, endowed the vicarage of this church, with the small tithes of it, excepting hay, with two acres of arable, and one of meadow; and also with the tithes of sheaves growing from land dug up with the foot, as well for the support of the vicar, as the discharge of the ordinary burthens of his vicarage, and the payment of the above pension to the monks, the profits of the vicarage being then sound by a jury to be worth forty marcs sterling per annum, communibis sannis; which endowment being lost, bishop Thomas de Woldham, in 1299, confirmed it; and as the vicar had no house belonging to his vicarage, he granted him one standing on the soil belonging to the church, as a vicarage house for himself and his successors; and further, the tithe of twenty-one acres of meadow, called King's-marsh, in Dartford, heretofore taken by the bishop and his predecessors, and he decreed, that the vicar and his successors should keep and maintain the books, vestments, and other ornaments of the church, in a proper state and order, and should sustain and acknowledge all other ordinary burthens of it.

 

Archbishop Robert Winchelsea further endowed this vicarage with the tithe of hay, to the value of forty shillings, in satisfaction of which the whole tithe of hay, arising from the great salt march in Dartford, (excepting to the bishop of Rochester for the time being, the yearly sum of four shillings, due from the Knights Hospitallers to the bishop, as rector of this church) was decreed to the vicar, by the desinitive sentence of Walter, archbishop of Canterbury, in the year 1315, as an augmentation of his endowment.

 

Thomas de Woldham, bishop of Rochester, in the above year, granted in mortmain, to Robert Levee, vicar of Dartford, and his successors, a messuage, with its appurtenances, in Overe-street, in Dartford, which the bishop had purchased of Robert de Levee, of Frindsbury. (fn. 53) At the dissolution of the priory of Rochester, in the 32d year of king Henry VIII. the above pension of ten marcs, or 6l. 13s. 4d. was, by the king, in his 33d year, granted, among other pre emises, to his new erected dean and chapter of Rochester, who continue possessed of it at this time. The parsonage and advowson of the vicarage still remain part of the possessions of the bishop of Rochester. (fn. 54)

 

In the antient valuation of the bishop's revenues, this church was valued at 40l. and the bishop's mill and rent here, at 100s. In the 15th year of king Edward I.'s reign, the church was valued at forty-five marcs, and the vicarage at 100s. In the 33d of king Edward III. the church was valued at the like sum. (fn. 55)

 

By virtue of a commission of enquiry, in 1650, it was returned, that Dartford was a vicarage, with a house and glebe, all worth, with the privy tithes, seventy pounds per annum, master Charnock then incumbent. (fn. 56) It is a discharged living in the king's books, of the clear yearly certified value of 45l. 5s. 10½d. the yearly tenths of which are 1l. 17s. 1½d. (fn. 57)

 

This vicarage was, in 1736, augmented by the governors of queen Anne's bounty; at which time the Rev. Mr. Charles Chambers, vicar of Dartford, contributed one hundred pounds for that purpose. (fn. 58)

 

Bishop Laurence de St. Martin seems to have purchased, in the reign of king Henry III. several of the rents which now constitute the greatest part, if not the whole of the MANOR OF DARTFORD RECTORY, from Robert and Richard de Ripa, John Badecock, William de Wilmington, and others. (fn. 59)

 

This manor extends over both sides of the Highstreet, in Dartford, from the scite of the old marketplace to the church, and southward, in Lowfield, as far as the house of correction; all which is called the Bishop's liberty. At the leet of this manor, a constable and a borsholder are annually chosen for the liberty. There are several tenants which hold of it in socage, at small quit-rents.

 

In the 21st year of king Edward I. on a Quo warranto, the jury found that the bishop was feild, in right of his church, of view of frank pledge, and assize of bread and ale of his tenants in Dartford and Stone; and that the bishops, his predecessors, had been possessed of the same beyond memory.

 

There were TWO CHANTRIES, founded for divine services, in this parish; that of St. Edmund the Martyr, and of St. Mary, otherwise called Stampit. The former stood in the upper burial ground of this parish, which was a cimetary to it, and under this building was a charnel house. This chapel was suppressed at the same time with all other such endowments, and presently sell to ruin; but the cimetary was granted to the parish, as a place of burial for the parishioners, and continues so at this time. The advowson of this chantry was granted to the prioress and convent of Dartford priory, in the 46th year of king Edward III. at their first endowment.

 

John Bykenore endowed this chapel with five marcs, payable out of lands and tenements in Dartford, for the support of the chaplain of it. This chapel was under the jurisdiction of the archdeacon of the diocese.

 

¶The latter chantry of the Blessed Virgin St. Mary was subject to the official of the diocese. (fn. 60) It was founded by Thomas de Dertford, alias Art Stampett, vicar of this parish, in 1338, for one chaplain, to celebrate divine offices daily in the parish church of Dartford, in honour of the Blessed Virgin, and for the health of his soul, &c. and he appointed Ralph de Felthorpe the first chaplain of it, and endowed it with several lands and tenements, to the amount of one hundred and twenty acres, (fn. 61) in Dartford, the chaplain paying twelve pence yearly to the vicar of Dartford and his successors; and he gave the patronage of it, and the nomination of a chaplain to it in future, to the bishop of Rochester and his successors; which was confirmed by the bishop and the prior and chapter of Rochester the same year. (fn. 62) In the year 1553, Robert Bacon, incumbent of this chantry, had a pension of six pounds per annum.

 

www.british-history.ac.uk/survey-kent/vol2/pp286-328

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Male wood ant (Formica rufa) searching for newly emerged queens. Surrey, UK.

 

Sexual pandemonium at the wood ant colony today as winged males scurried about in search of newly emerged queens.

 

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