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Fish (Actinopterygii) by PHOTOGRAPHY by DM & DBM.

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Fish, any of approximately 34,000 species of vertebrate animals (phylum Chordata) found in the fresh and salt waters of the world. Living species range from the primitive jawless lampreys and hagfishes through the cartilaginous sharks, skates, and rays to the abundant and diverse bony fishes. Most fish species are cold-blooded; however, one species, the opah (Lampris guttatus), is warm-blooded.

The term fish is applied to a variety of vertebrates of several evolutionary lines. It describes a life-form rather than a taxonomic group. As members of the phylum Chordata, fish share certain features with other vertebrates. These features are gill slits at some point in the life cycle, a notochord, or skeletal supporting rod, a dorsal hollow nerve cord, and a tail. Living fishes represent some five classes, which are as distinct from one another as are the four classes of familiar air-breathing animals—amphibians, reptiles, birds, and mammals. For example, the jawless fishes (Agnatha) have gills in pouches and lack limb girdles. Extant agnathans are the lampreys and the hagfishes. As the name implies, the skeletons of fishes of the class Chondrichthyes (from chondr, “cartilage,” and ichthyes, “fish”) are made entirely of cartilage. Modern fish of this class lack a swim bladder, and their scales and teeth are made up of the same placoid material. Sharks, skates, and rays are examples of cartilaginous fishes. The bony fishes are by far the largest class. Examples range from the tiny seahorse to the 450-kg (1,000-pound) blue marlin, from the flattened soles and flounders to the boxy puffers and ocean sunfishes. Unlike the scales of the cartilaginous fishes, those of bony fishes, when present, grow throughout life and are made up of thin overlapping plates of bone. Bony fishes also have an operculum that covers the gill slits.

The study of fishes, the science of ichthyology, is of broad importance. Fishes are of interest to humans for many reasons, the most important being their relationship with and dependence on the environment. A more obvious reason for interest in fishes is their role as a moderate but important part of the world’s food supply. This resource, once thought unlimited, is now realized to be finite and in delicate balance with the biological, chemical, and physical factors of the aquatic environment. Overfishing, pollution, and alteration of the environment are the chief enemies of proper fisheries management, both in fresh waters and in the ocean. (For a detailed discussion of the technology and economics of fisheries, see commercial fishing.) Another practical reason for studying fishes is their use in disease control. As predators on mosquito larvae, they help curb malaria and other mosquito-borne diseases.

Fishes are valuable laboratory animals in many aspects of medical and biological research. For example, the readiness of many fishes to acclimate to captivity has allowed biologists to study behaviour, physiology, and even ecology under relatively natural conditions. Fishes have been especially important in the study of animal behaviour, where research on fishes has provided a broad base for the understanding of the more flexible behaviour of the higher vertebrates. The zebra fish is used as a model in studies of gene expression.

There are aesthetic and recreational reasons for an interest in fishes. Millions of people keep live fishes in home aquariums for the simple pleasure of observing the beauty and behaviour of animals otherwise unfamiliar to them. Aquarium fishes provide a personal challenge to many aquarists, allowing them to test their ability to keep a small section of the natural environment in their homes. Sportfishing is another way of enjoying the natural environment, also indulged in by millions of people every year. Interest in aquarium fishes and sportfishing supports multimillion-dollar industries throughout the world.

Fishes have been in existence for more than 450 million years, during which time they have evolved repeatedly to fit into almost every conceivable type of aquatic habitat. In a sense, land vertebrates are simply highly modified fishes: when fishes colonized the land habitat, they became tetrapod (four-legged) land vertebrates. The popular conception of a fish as a slippery, streamlined aquatic animal that possesses fins and breathes by gills applies to many fishes, but far more fishes deviate from that conception than conform to it. For example, the body is elongate in many forms and greatly shortened in others; the body is flattened in some (principally in bottom-dwelling fishes) and laterally compressed in many others; the fins may be elaborately extended, forming intricate shapes, or they may be reduced or even lost; and the positions of the mouth, eyes, nostrils, and gill openings vary widely. Air breathers have appeared in several evolutionary lines.

Many fishes are cryptically coloured and shaped, closely matching their respective environments; others are among the most brilliantly coloured of all organisms, with a wide range of hues, often of striking intensity, on a single individual. The brilliance of pigments may be enhanced by the surface structure of the fish, so that it almost seems to glow. A number of unrelated fishes have actual light-producing organs. Many fishes are able to alter their coloration—some for the purpose of camouflage, others for the enhancement of behavioral signals.

Fishes range in adult length from less than 10 mm (0.4 inch) to more than 20 metres (60 feet) and in weight from about 1.5 grams (less than 0.06 ounce) to many thousands of kilograms. Some live in shallow thermal springs at temperatures slightly above 42 °C (100 °F), others in cold Arctic seas a few degrees below 0 °C (32 °F) or in cold deep waters more than 4,000 metres (13,100 feet) beneath the ocean surface. The structural and, especially, the physiological adaptations for life at such extremes are relatively poorly known and provide the scientifically curious with great incentive for study.

Almost all natural bodies of water bear fish life, the exceptions being very hot thermal ponds and extremely salt-alkaline lakes, such as the Dead Sea in Asia and the Great Salt Lake in North America. The present distribution of fishes is a result of the geological history and development of Earth as well as the ability of fishes to undergo evolutionary change and to adapt to the available habitats. Fishes may be seen to be distributed according to habitat and according to geographical area. Major habitat differences are marine and freshwater. For the most part, the fishes in a marine habitat differ from those in a freshwater habitat, even in adjacent areas, but some, such as the salmon, migrate from one to the other. The freshwater habitats may be seen to be of many kinds. Fishes found in mountain torrents, Arctic lakes, tropical lakes, temperate streams, and tropical rivers will all differ from each other, both in obvious gross structure and in physiological attributes. Even in closely adjacent habitats where, for example, a tropical mountain torrent enters a lowland stream, the fish fauna will differ. The marine habitats can be divided into deep ocean floors (benthic), mid-water oceanic (bathypelagic), surface oceanic (pelagic), rocky coast, sandy coast, muddy shores, bays, estuaries, and others. Also, for example, rocky coastal shores in tropical and temperate regions will have different fish faunas, even when such habitats occur along the same coastline.

Although much is known about the present geographical distribution of fishes, far less is known about how that distribution came about. Many parts of the fish fauna of the fresh waters of North America and Eurasia are related and undoubtedly have a common origin. The faunas of Africa and South America are related, extremely old, and probably an expression of the drifting apart of the two continents. The fauna of southern Asia is related to that of Central Asia, and some of it appears to have entered Africa. The extremely large shore-fish faunas of the Indian and tropical Pacific oceans comprise a related complex, but the tropical shore fauna of the Atlantic, although containing Indo-Pacific components, is relatively limited and probably younger. The Arctic and Antarctic marine faunas are quite different from each other. The shore fauna of the North Pacific is quite distinct, and that of the North Atlantic more limited and probably younger. Pelagic oceanic fishes, especially those in deep waters, are similar the world over, showing little geographical isolation in terms of family groups. The deep oceanic habitat is very much the same throughout the world, but species differences do exist, showing geographical areas determined by oceanic currents and water masses.

All aspects of the life of a fish are closely correlated with adaptation to the total environment, physical, chemical, and biological. In studies, all the interdependent aspects of fish, such as behaviour, locomotion, reproduction, and physical and physiological characteristics, must be taken into account.

Correlated with their adaptation to an extremely wide variety of habitats is the extremely wide variety of life cycles that fishes display. The great majority hatch from relatively small eggs a few days to several weeks or more after the eggs are scattered in the water. Newly hatched young are still partially undeveloped and are called larvae until body structures such as fins, skeleton, and some organs are fully formed. Larval life is often very short, usually less than a few weeks, but it can be very long, some lampreys continuing as larvae for at least five years. Young and larval fishes, before reaching sexual maturity, must grow considerably, and their small size and other factors often dictate that they live in a habitat different than that of the adults. For example, most tropical marine shore fishes have pelagic larvae. Larval food also is different, and larval fishes often live in shallow waters, where they may be less exposed to predators.

After a fish reaches adult size, the length of its life is subject to many factors, such as innate rates of aging, predation pressure, and the nature of the local climate. The longevity of a species in the protected environment of an aquarium may have nothing to do with how long members of that species live in the wild. Many small fishes live only one to three years at the most. In some species, however, individuals may live as long as 10 or 20 or even 100 years.

Fish behaviour is a complicated and varied subject. As in almost all animals with a central nervous system, the nature of a response of an individual fish to stimuli from its environment depends upon the inherited characteristics of its nervous system, on what it has learned from past experience, and on the nature of the stimuli. Compared with the variety of human responses, however, that of a fish is stereotyped, not subject to much modification by “thought” or learning, and investigators must guard against anthropomorphic interpretations of fish behaviour.

Fishes perceive the world around them by the usual senses of sight, smell, hearing, touch, and taste and by special lateral line water-current detectors. In the few fishes that generate electric fields, a process that might best be called electrolocation aids in perception. One or another of these senses often is emphasized at the expense of others, depending upon the fish’s other adaptations. In fishes with large eyes, the sense of smell may be reduced; others, with small eyes, hunt and feed primarily by smell (such as some eels).

Specialized behaviour is primarily concerned with the three most important activities in the fish’s life: feeding, reproduction, and escape from enemies. Schooling behaviour of sardines on the high seas, for instance, is largely a protective device to avoid enemies, but it is also associated with and modified by their breeding and feeding requirements. Predatory fishes are often solitary, lying in wait to dart suddenly after their prey, a kind of locomotion impossible for beaked parrot fishes, which feed on coral, swimming in small groups from one coral head to the next. In addition, some predatory fishes that inhabit pelagic environments, such as tunas, often school.

Sleep in fishes, all of which lack true eyelids, consists of a seemingly listless state in which the fish maintains its balance but moves slowly. If attacked or disturbed, most can dart away. A few kinds of fishes lie on the bottom to sleep. Most catfishes, some loaches, and some eels and electric fishes are strictly nocturnal, being active and hunting for food during the night and retiring during the day to holes, thick vegetation, or other protective parts of the environment.

Communication between members of a species or between members of two or more species often is extremely important, especially in breeding behaviour (see below Reproduction). The mode of communication may be visual, as between the small so-called cleaner fish and a large fish of a very different species. The larger fish often allows the cleaner to enter its mouth to remove gill parasites. The cleaner is recognized by its distinctive colour and actions and therefore is not eaten, even if the larger fish is normally a predator. Communication is often chemical, signals being sent by specific chemicals called pheromones.

Many fishes have a streamlined body and swim freely in open water. Fish locomotion is closely correlated with habitat and ecological niche (the general position of the animal to its environment).

Many fishes in both marine and fresh waters swim at the surface and have mouths adapted to feed best (and sometimes only) at the surface. Often such fishes are long and slender, able to dart at surface insects or at other surface fishes and in turn to dart away from predators; needlefishes, halfbeaks, and topminnows (such as killifish and mosquito fish) are good examples. Oceanic flying fishes escape their predators by gathering speed above the water surface, with the lower lobe of the tail providing thrust in the water. They then glide hundreds of yards on enlarged, winglike pectoral and pelvic fins. South American freshwater flying fishes escape their enemies by jumping and propelling their strongly keeled bodies out of the water.

So-called mid-water swimmers, the most common type of fish, are of many kinds and live in many habitats. The powerful fusiform tunas and the trouts, for example, are adapted for strong, fast swimming, the tunas to capture prey speedily in the open ocean and the trouts to cope with the swift currents of streams and rivers. The trout body form is well adapted to many habitats. Fishes that live in relatively quiet waters such as bays or lake shores or slow rivers usually are not strong, fast swimmers but are capable of short, quick bursts of speed to escape a predator. Many of these fishes have their sides flattened, examples being the sunfish and the freshwater angelfish of aquarists. Fish associated with the bottom or substrate usually are slow swimmers. Open-water plankton-feeding fishes almost always remain fusiform and are capable of rapid, strong movement (for example, sardines and herrings of the open ocean and also many small minnows of streams and lakes).

Bottom-living fishes are of many kinds and have undergone many types of modification of their body shape and swimming habits. Rays, which evolved from strong-swimming mid-water sharks, usually stay close to the bottom and move by undulating their large pectoral fins. Flounders live in a similar habitat and move over the bottom by undulating the entire body. Many bottom fishes dart from place to place, resting on the bottom between movements, a motion common in gobies. One goby relative, the mudskipper, has taken to living at the edge of pools along the shore of muddy mangrove swamps. It escapes its enemies by flipping rapidly over the mud, out of the water. Some catfishes, synbranchid eels, the so-called climbing perch, and a few other fishes venture out over damp ground to find more promising waters than those that they left. They move by wriggling their bodies, sometimes using strong pectoral fins; most have accessory air-breathing organs. Many bottom-dwelling fishes live in mud holes or rocky crevices. Marine eels and gobies commonly are found in such habitats and for the most part venture far beyond their cavelike homes. Some bottom dwellers, such as the clingfishes (Gobiesocidae), have developed powerful adhesive disks that enable them to remain in place on the substrate in areas such as rocky coasts, where the action of the waves is great.

The methods of reproduction in fishes are varied, but most fishes lay a large number of small eggs, fertilized and scattered outside of the body. The eggs of pelagic fishes usually remain suspended in the open water. Many shore and freshwater fishes lay eggs on the bottom or among plants. Some have adhesive eggs. The mortality of the young and especially of the eggs is very high, and often only a few individuals grow to maturity out of hundreds, thousands, and in some cases millions of eggs laid.

Males produce sperm, usually as a milky white substance called milt, in two (sometimes one) testes within the body cavity. In bony fishes a sperm duct leads from each testis to a urogenital opening behind the vent or anus. In sharks and rays and in cyclostomes the duct leads to a cloaca. Sometimes the pelvic fins are modified to help transmit the milt to the eggs at the female’s vent or on the substrate where the female has placed them. Sometimes accessory organs are used to fertilize females internally—for example, the claspers of many sharks and rays.

In the females the eggs are formed in two ovaries (sometimes only one) and pass through the ovaries to the urogenital opening and to the outside. In some fishes the eggs are fertilized internally but are shed before development takes place. Members of about a dozen families each of bony fishes (teleosts) and sharks bear live young. Many skates and rays also bear live young. In some bony fishes the eggs simply develop within the female, the young emerging when the eggs hatch (ovoviviparous). Others develop within the ovary and are nourished by ovarian tissues after hatching (viviparous). There are also other methods utilized by fishes to nourish young within the female. In all live-bearers the young are born at a relatively large size and are few in number. In one family of primarily marine fishes, the surfperches from the Pacific coast of North America, Japan, and Korea, the males of at least one species are born sexually mature, although they are not fully grown.

Some fishes are hermaphroditic—an individual producing both sperm and eggs, usually at different stages of its life. Self-fertilization, however, is probably rare.

Successful reproduction and, in many cases, defense of the eggs and the young are assured by rather stereotypical but often elaborate courtship and parental behaviour, either by the male or the female or both. Some fishes prepare nests by hollowing out depressions in the sand bottom (cichlids, for example), build nests with plant materials and sticky threads excreted by the kidneys (sticklebacks), or blow a cluster of mucus-covered bubbles at the water surface (gouramis). The eggs are laid in these structures. Some varieties of cichlids and catfishes incubate eggs in their mouths.

Some fishes, such as salmon, undergo long migrations from the ocean and up large rivers to spawn in the gravel beds where they themselves hatched (anadromous fishes). Some, such as the freshwater eels (family Anguillidae), live and grow to maturity in fresh water and migrate to the sea to spawn (catadromous fishes). Other fishes undertake shorter migrations from lakes into streams, within the ocean, or enter spawning habitats that they do not ordinarily occupy in other ways.

The basic structure and function of the fish body are similar to those of all other vertebrates. The usual four types of tissues are present: surface or epithelial, connective (bone, cartilage, and fibrous tissues, as well as their derivative, blood), nerve, and muscle tissues. In addition, the fish’s organs and organ systems parallel those of other vertebrates.

The typical fish body is streamlined and spindle-shaped, with an anterior head, a gill apparatus, and a heart, the latter lying in the midline just below the gill chamber. The body cavity, containing the vital organs, is situated behind the head in the lower anterior part of the body. The anus usually marks the posterior termination of the body cavity and most often occurs just in front of the base of the anal fin. The spinal cord and vertebral column continue from the posterior part of the head to the base of the tail fin, passing dorsal to the body cavity and through the caudal (tail) region behind the body cavity. Most of the body is of muscular tissue, a high proportion of which is necessitated by swimming. In the course of evolution this basic body plan has been modified repeatedly into the many varieties of fish shapes that exist today.

The skeleton forms an integral part of the fish’s locomotion system, as well as serving to protect vital parts. The internal skeleton consists of the skull bones (except for the roofing bones of the head, which are really part of the external skeleton), the vertebral column, and the fin supports (fin rays). The fin supports are derived from the external skeleton but will be treated here because of their close functional relationship to the internal skeleton. The internal skeleton of cyclostomes, sharks, and rays is of cartilage; that of many fossil groups and some primitive living fishes is mostly of cartilage but may include some bone. In place of the vertebral column, the earliest vertebrates had a fully developed notochord, a flexible stiff rod of viscous cells surrounded by a strong fibrous sheath. During the evolution of modern fishes the rod was replaced in part by cartilage and then by ossified cartilage. Sharks and rays retain a cartilaginous vertebral column; bony fishes have spool-shaped vertebrae that in the more primitive living forms only partially replace the notochord. The skull, including the gill arches and jaws of bony fishes, is fully, or at least partially, ossified. That of sharks and rays remains cartilaginous, at times partially replaced by calcium deposits but never by true bone.

The supportive elements of the fins (basal or radial bones or both) have changed greatly during fish evolution. Some of these changes are described in the section below (Evolution and paleontology). Most fishes possess a single dorsal fin on the midline of the back. Many have two and a few have three dorsal fins. The other fins are the single tail and anal fins and paired pelvic and pectoral fins. A small fin, the adipose fin, with hairlike fin rays, occurs in many of the relatively primitive teleosts (such as trout) on the back near the base of the caudal fin.

The skin of a fish must serve many functions. It aids in maintaining the osmotic balance, provides physical protection for the body, is the site of coloration, contains sensory receptors, and, in some fishes, functions in respiration. Mucous glands, which aid in maintaining the water balance and offer protection from bacteria, are extremely numerous in fish skin, especially in cyclostomes and teleosts. Since mucous glands are present in the modern lampreys, it is reasonable to assume that they were present in primitive fishes, such as the ancient Silurian and Devonian agnathans. Protection from abrasion and predation is another function of the fish skin, and dermal (skin) bone arose early in fish evolution in response to this need. It is thought that bone first evolved in skin and only later invaded the cartilaginous areas of the fish’s body, to provide additional support and protection. There is some argument as to which came first, cartilage or bone, and fossil evidence does not settle the question. In any event, dermal bone has played an important part in fish evolution and has different characteristics in different groups of fishes. Several groups are characterized at least in part by the kind of bony scales they possess.

Scales have played an important part in the evolution of fishes. Primitive fishes usually had thick bony plates or thick scales in several layers of bone, enamel, and related substances. Modern teleost fishes have scales of bone, which, while still protective, allow much more freedom of motion in the body. A few modern teleosts (some catfishes, sticklebacks, and others) have secondarily acquired bony plates in the skin. Modern and early sharks possessed placoid scales, a relatively primitive type of scale with a toothlike structure, consisting of an outside layer of enamel-like substance (vitrodentine), an inner layer of dentine, and a pulp cavity containing nerves and blood vessels. Primitive bony fishes had thick scales of either the ganoid or the cosmoid type. Cosmoid scales have a hard, enamel-like outer layer, an inner layer of cosmine (a form of dentine), and then a layer of vascular bone (isopedine). In ganoid scales the hard outer layer is different chemically and is called ganoin. Under this is a cosminelike layer and then a vascular bony layer. The thin, translucent bony scales of modern fishes, called cycloid and ctenoid (the latter distinguished by serrations at the edges), lack enameloid and dentine layers.

Skin has several other functions in fishes. It is well supplied with nerve endings and presumably receives tactile, thermal, and pain stimuli. Skin is also well supplied with blood vessels. Some fishes breathe in part through the skin, by the exchange of oxygen and carbon dioxide between the surrounding water and numerous small blood vessels near the skin surface.

Skin serves as protection through the control of coloration. Fishes exhibit an almost limitless range of colours. The colours often blend closely with the surroundings, effectively hiding the animal. Many fishes use bright colours for territorial advertisement or as recognition marks for other members of their own species, or sometimes for members of other species. Many fishes can change their colour to a greater or lesser degree, by movement of pigment within the pigment cells (chromatophores). Black pigment cells (melanophores), of almost universal occurrence in fishes, are often juxtaposed with other pigment cells. When placed beneath iridocytes or leucophores (bearing the silvery or white pigment guanine), melanophores produce structural colours of blue and green. These colours are often extremely intense, because they are formed by refraction of light through the needlelike crystals of guanine. The blue and green refracted colours are often relatively pure, lacking the red and yellow rays, which have been absorbed by the black pigment (melanin) of the melanophores. Yellow, orange, and red colours are produced by erythrophores, cells containing the appropriate carotenoid pigments. Other colours are produced by combinations of melanophores, erythrophores, and iridocytes.

The major portion of the body of most fishes consists of muscles. Most of the mass is trunk musculature, the fin muscles usually being relatively small. The caudal fin is usually the most powerful fin, being moved by the trunk musculature. The body musculature is usually arranged in rows of chevron-shaped segments on each side. Contractions of these segments, each attached to adjacent vertebrae and vertebral processes, bends the body on the vertebral joint, producing successive undulations of the body, passing from the head to the tail, and producing driving strokes of the tail. It is the latter that provides the strong forward movement for most fishes.

The digestive system, in a functional sense, starts at the mouth, with the teeth used to capture prey or collect plant foods. Mouth shape and tooth structure vary greatly in fishes, depending on the kind of food normally eaten. Most fishes are predacious, feeding on small invertebrates or other fishes and have simple conical teeth on the jaws, on at least some of the bones of the roof of the mouth, and on special gill arch structures just in front of the esophagus. The latter are throat teeth. Most predacious fishes swallow their prey whole, and the teeth are used for grasping and holding prey, for orienting prey to be swallowed (head first) and for working the prey toward the esophagus. There are a variety of tooth types in fishes. Some fishes, such as sharks and piranhas, have cutting teeth for biting chunks out of their victims. A shark’s tooth, although superficially like that of a piranha, appears in many respects to be a modified scale, while that of the piranha is like that of other bony fishes, consisting of dentine and enamel. Parrot fishes have beaklike mouths with short incisor-like teeth for breaking off coral and have heavy pavementlike throat teeth for crushing the coral. Some catfishes have small brushlike teeth, arranged in rows on the jaws, for scraping plant and animal growth from rocks. Many fishes (such as the Cyprinidae or minnows) have no jaw teeth at all but have very strong throat teeth.

Some fishes gather planktonic food by straining it from their gill cavities with numerous elongate stiff rods (gill rakers) anchored by one end to the gill bars. The food collected on these rods is passed to the throat, where it is swallowed. Most fishes have only short gill rakers that help keep food particles from escaping out the mouth cavity into the gill chamber.

Once reaching the throat, food enters a short, often greatly distensible esophagus, a simple tube with a muscular wall leading into a stomach. The stomach varies greatly in fishes, depending upon the diet. In most predacious fishes it is a simple straight or curved tube or pouch with a muscular wall and a glandular lining. Food is largely digested there and leaves the stomach in liquid form.

Between the stomach and the intestine, ducts enter the digestive tube from the liver and pancreas. The liver is a large, clearly defined organ. The pancreas may be embedded in it, diffused through it, or broken into small parts spread along some of the intestine. The junction between the stomach and the intestine is marked by a muscular valve. Pyloric ceca (blind sacs) occur in some fishes at this junction and have a digestive or absorptive function or both.

The intestine itself is quite variable in length, depending upon the fish’s diet. It is short in predacious forms, sometimes no longer than the body cavity, but long in herbivorous forms, being coiled and several times longer than the entire length of the fish in some species of South American catfishes. The intestine is primarily an organ for absorbing nutrients into the bloodstream. The larger its internal surface, the greater its absorptive efficiency, and a spiral valve is one method of increasing its absorption surface.

Sharks, rays, chimaeras, lungfishes, surviving chondrosteans, holosteans, and even a few of the more primitive teleosts have a spiral valve or at least traces of it in the intestine. Most modern teleosts have increased the area of the intestinal walls by having numerous folds and villi (fingerlike projections) somewhat like those in humans. Undigested substances are passed to the exterior through the anus in most teleost fishes. In lungfishes, sharks, and rays, it is first passed through the cloaca, a common cavity receiving the intestinal opening and the ducts from the urogenital system.

Oxygen and carbon dioxide dissolve in water, and most fishes exchange dissolved oxygen and carbon dioxide in water by means of the gills. The gills lie behind and to the side of the mouth cavity and consist of fleshy filaments supported by the gill arches and filled with blood vessels, which give gills a bright red colour. Water taken in continuously through the mouth passes backward between the gill bars and over the gill filaments, where the exchange of gases takes place. The gills are protected by a gill cover in teleosts and many other fishes but by flaps of skin in sharks, rays, and some of the older fossil fish groups. The blood capillaries in the gill filaments are close to the gill surface to take up oxygen from the water and to give up excess carbon dioxide to the water.

Most modern fishes have a hydrostatic (ballast) organ, called the swim bladder, that lies in the body cavity just below the kidney and above the stomach and intestine. It originated as a diverticulum of the digestive canal. In advanced teleosts, especially the acanthopterygians, the bladder has lost its connection with the digestive tract, a condition called physoclistic. The connection has been retained (physostomous) by many relatively primitive teleosts. In several unrelated lines of fishes, the bladder has become specialized as a lung or, at least, as a highly vascularized accessory breathing organ. Some fishes with such accessory organs are obligate air breathers and will drown if denied access to the surface, even in well-oxygenated water. Fishes with a hydrostatic form of swim bladder can control their depth by regulating the amount of gas in the bladder. The gas, mostly oxygen, is secreted into the bladder by special glands, rendering the fish more buoyant; the gas is absorbed into the bloodstream by another special organ, reducing the overall buoyancy and allowing the fish to sink. Some deep-sea fishes may have oils, rather than gas, in the bladder. Other deep-sea and some bottom-living forms have much-reduced swim bladders or have lost the organ entirely.

The swim bladder of fishes follows the same developmental pattern as the lungs of land vertebrates. There is no doubt that the two structures have the same historical origin in primitive fishes. More or less intermediate forms still survive among the more primitive types of fishes, such as the lungfishes Lepidosiren and Protopterus.

The circulatory, or blood vascular, system consists of the heart, the arteries, the capillaries, and the veins. It is in the capillaries that the interchange of oxygen, carbon dioxide, nutrients, and other substances such as hormones and waste products takes place. The capillaries lead to the veins, which return the venous blood with its waste products to the heart, kidneys, and gills. There are two kinds of capillary beds: those in the gills and those in the rest of the body. The heart, a folded continuous muscular tube with three or four saclike enlargements, undergoes rhythmic contractions and receives venous blood in a sinus venosus. It passes the blood to an auricle and then into a thick muscular pump, the ventricle. From the ventricle the blood goes to a bulbous structure at the base of a ventral aorta just below the gills. The blood passes to the afferent (receiving) arteries of the gill arches and then to the gill capillaries. There waste gases are given off to the environment, and oxygen is absorbed. The oxygenated blood enters efferent (exuant) arteries of the gill arches and then flows into the dorsal aorta. From there blood is distributed to the tissues and organs of the body. One-way valves prevent backflow. The circulation of fishes thus differs from that of the reptiles, birds, and mammals in that oxygenated blood is not returned to the heart prior to distribution to the other parts of the body.

The primary excretory organ in fishes, as in other vertebrates, is the kidney. In fishes some excretion also takes place in the digestive tract, skin, and especially the gills (where ammonia is given off). Compared with land vertebrates, fishes have a special problem in maintaining their internal environment at a constant concentration of water and dissolved substances, such as salts. Proper balance of the internal environment (homeostasis) of a fish is in a great part maintained by the excretory system, especially the kidney.

The kidney, gills, and skin play an important role in maintaining a fish’s internal environment and checking the effects of osmosis. Marine fishes live in an environment in which the water around them has a greater concentration of salts than they can have inside their body and still maintain life. Freshwater fishes, on the other hand, live in water with a much lower concentration of salts than they require inside their bodies. Osmosis tends to promote the loss of water from the body of a marine fish and absorption of water by that of a freshwater fish. Mucus in the skin tends to slow the process but is not a sufficient barrier to prevent the movement of fluids through the permeable skin. When solutions on two sides of a permeable membrane have different concentrations of dissolved substances, water will pass through the membrane into the more concentrated solution, while the dissolved chemicals move into the area of lower concentration (diffusion).

The kidney of freshwater fishes is often larger in relation to body weight than that of marine fishes. In both groups the kidney excretes wastes from the body, but the kidney of freshwater fishes also excretes large amounts of water, counteracting the water absorbed through the skin. Freshwater fishes tend to lose salt to the environment and must replace it. They get some salt from their food, but the gills and skin inside the mouth actively absorb salt from water passed through the mouth. This absorption is performed by special cells capable of moving salts against the diffusion gradient. Freshwater fishes drink very little water and take in little water with their food.

Marine fishes must conserve water, and therefore their kidneys excrete little water. To maintain their water balance, marine fishes drink large quantities of seawater, retaining most of the water and excreting the salt. Most nitrogenous waste in marine fishes appears to be secreted by the gills as ammonia. Marine fishes can excrete salt by clusters of special cells (chloride cells) in the gills.

There are several teleosts—for example, the salmon—that travel between fresh water and seawater and must adjust to the reversal of osmotic gradients. They adjust their physiological processes by spending time (often surprisingly little time) in the intermediate brackish environment.

Marine hagfishes, sharks, and rays have osmotic concentrations in their blood about equal to that of seawater and so do not have to drink water nor perform much physiological work to maintain their osmotic balance. In sharks and rays the osmotic concentration is kept high by retention of urea in the blood. Freshwater sharks have a lowered concentration of urea in the blood.

Endocrine glands secrete their products into the bloodstream and body tissues and, along with the central nervous system, control and regulate many kinds of body functions. Cyclostomes have a well-developed endocrine system, and presumably it was well developed in the early Agnatha, ancestral to modern fishes. Although the endocrine system in fishes is similar to that of higher vertebrates, there are numerous differences in detail. The pituitary, the thyroid, the suprarenals, the adrenals, the pancreatic islets, the sex glands (ovaries and testes), the inner wall of the intestine, and the bodies of the ultimobranchial gland make up the endocrine system in fishes. There are some others whose function is not well understood. These organs regulate sexual activity and reproduction, growth, osmotic pressure, general metabolic activities such as the storage of fat and the utilization of foodstuffs, blood pressure, and certain aspects of skin colour. Many of these activities are also controlled in part by the central nervous system, which works with the endocrine system in maintaining the life of a fish. Some parts of the endocrine system are developmentally, and undoubtedly evolutionarily, derived from the nervous system.

As in all vertebrates, the nervous system of fishes is the primary mechanism coordinating body activities, as well as integrating these activities in the appropriate manner with stimuli from the environment. The central nervous system, consisting of the brain and spinal cord, is the primary integrating mechanism. The peripheral nervous system, consisting of nerves that connect the brain and spinal cord to various body organs, carries sensory information from special receptor organs such as the eyes, internal ears, nares (sense of smell), taste glands, and others to the integrating centres of the brain and spinal cord. The peripheral nervous system also carries information via different nerve cells from the integrating centres of the brain and spinal cord. This coded information is carried to the various organs and body systems, such as the skeletal muscular system, for appropriate action in response to the original external or internal stimulus. Another branch of the nervous system, the autonomic nervous system, helps to coordinate the activities of many glands and organs and is itself closely connected to the integrating centres of the brain.

The brain of the fish is divided into several anatomical and functional parts, all closely interconnected but each serving as the primary centre of integrating particular kinds of responses and activities. Several of these centres or parts are primarily associated with one type of sensory perception, such as sight, hearing, or smell (olfaction).

The sense of smell is important in almost all fishes. Certain eels with tiny eyes depend mostly on smell for location of food. The olfactory, or nasal, organ of fishes is located on the dorsal surface of the snout. The lining of the nasal organ has special sensory cells that perceive chemicals dissolved in the water, such as substances from food material, and send sensory information to the brain by way of the first cranial nerve. Odour also serves as an alarm system. Many fishes, especially various species of freshwater minnows, react with alarm to a chemical released from the skin of an injured member of their own species.

Many fishes have a well-developed sense of taste, and tiny pitlike taste buds or organs are located not only within their mouth cavities but also over their heads and parts of their body. Catfishes, which often have poor vision, have barbels (“whiskers”) that serve as supplementary taste organs, those around the mouth being actively used to search out food on the bottom. Some species of naturally blind cave fishes are especially well supplied with taste buds, which often cover most of their body surface.

Sight is extremely important in most fishes. The eye of a fish is basically like that of all other vertebrates, but the eyes of fishes are extremely varied in structure and adaptation. In general, fishes living in dark and dim water habitats have large eyes, unless they have specialized in some compensatory way so that another sense (such as smell) is dominant, in which case the eyes will often be reduced. Fishes living in brightly lighted shallow waters often will have relatively small but efficient eyes. Cyclostomes have somewhat less elaborate eyes than other fishes, with skin stretched over the eyeball perhaps making their vision somewhat less effective. Most fishes have a spherical lens and accommodate their vision to far or near subjects by moving the lens within the eyeball. A few sharks accommodate by changing the shape of the lens, as in land vertebrates. Those fishes that are heavily dependent upon the eyes have especially strong muscles for accommodation. Most fishes see well, despite the restrictions imposed by frequent turbidity of the water and by light refraction.

Fossil evidence suggests that colour vision evolved in fishes more than 300 million years ago, but not all living fishes have retained this ability. Experimental evidence indicates that many shallow-water fishes, if not all, have colour vision and see some colours especially well, but some bottom-dwelling shore fishes live in areas where the water is sufficiently deep to filter out most if not all colours, and these fishes apparently never see colours. When tested in shallow water, they apparently are unable to respond to colour differences.

Sound perception and balance are intimately associated senses in a fish. The organs of hearing are entirely internal, located within the skull, on each side of the brain and somewhat behind the eyes. Sound waves, especially those of low frequencies, travel readily through water and impinge directly upon the bones and fluids of the head and body, to be transmitted to the hearing organs. Fishes readily respond to sound; for example, a trout conditioned to escape by the approach of fishermen will take flight upon perceiving footsteps on a stream bank even if it cannot see a fisherman. Compared with humans, however, the range of sound frequencies heard by fishes is greatly restricted. Many fishes communicate with each other by producing sounds in their swim bladders, in their throats by rasping their teeth, and in other ways.

A fish or other vertebrate seldom has to rely on a single type of sensory information to determine the nature of the environment around it. A catfish uses taste and touch when examining a food object with its oral barbels. Like most other animals, fishes have many touch receptors over their body surface. Pain and temperature receptors also are present in fishes and presumably produce the same kind of information to a fish as to humans. Fishes react in a negative fashion to stimuli that would be painful to human beings, suggesting that they feel a sensation of pain.

An important sensory system in fishes that is absent in other vertebrates (except some amphibians) is the lateral line system. This consists of a series of heavily innervated small canals located in the skin and bone around the eyes, along the lower jaw, over the head, and down the mid-side of the body, where it is associated with the scales. Intermittently along these canals are located tiny sensory organs (pit organs) that apparently detect changes in pressure. The system allows a fish to sense changes in water currents and pressure, thereby helping the fish to orient itself to the various changes that occur in the physical environment.

BA License Plate Bad Aussee Austria BMW R1200 CL (c) Bernard Egger :: rumoto images 5013 by © Bernard Egger 📷

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photographer.. |►collections.. |►my sets.. |► BMW R1200 CL

BMW R 1200 CL - Woodcliff Lake, New Jersey, August 2002

Some people consider a six-day cruise as the perfect vacation. Other's might agree, as long as the days are marked by blurred fence posts and dotted lines instead of palm trees and ocean waves. For them, BMW introduces the perfect alternative to a deck chair - the R 1200 CL.

Motorcyclists were taken aback when BMW introduced its first cruiser in 1997, but the R 1200 C quickly rose to become that year's best-selling BMW. The original has since spawned several derivatives including the Phoenix, Euro, Montana and Stiletto. This year, BMW's cruiser forms the basis for the most radical departure yet, the R 1200 CL. With its standard integral hard saddlebags, top box and distinctive handlebar-mounted fairing, the CL represents twin-cylinder luxury-touring at its finest, a completely modern luxury touring-cruiser with a touch of classic BMW.

Although based on the R 1200 C, the new CL includes numerous key changes in chassis, drivetrain, equipment and appearance, specifically designed to enhance the R 1200's abilities as a long-distance mount. While it uses the same torquey, 1170cc 61-hp version of BMW's highly successful R259 twin, the CL backs it with a six-speed overdrive transmission. A reworked Telelever increases the bike's rake for more-relaxed high-speed steering, while the fork's wider spacing provides room for the sculpted double-spoke, 16-inch wheel and 150/80 front tire. Similarly, a reinforced Monolever rear suspension controls a matching 15-inch alloy wheel and 170/80 rear tire. As you'd expect, triple disc brakes featuring BMW's latest EVO front brake system and fully integrated ABS bring the bike to a halt at day's end-and set the CL apart from any other luxury cruiser on the market.

Yet despite all the chassis changes, it's the new CL's visual statement that represents the bike's biggest break with its cruiser-mates. With its grip-to-grip sweep, the handlebar-mounted fairing evokes classic touring bikes, while the CL's distinctive quad-headlamps give the bike a decidedly avant-garde look - in addition to providing standard-setting illumination. A pair of frame-mounted lowers extends the fairing's wind coverage and provides space for some of the CL's electrics and the optional stereo. The instrument panel is exceptionally clean, surrounded by a matte gray background that matches the kneepads inset in the fairing extensions. The speedometer and tachometer flank a panel of warning lights, capped by the standard analog clock. Integrated mirror/turnsignal pods extend from the fairing to provide further wind protection. Finally, fully integrated, color-matched saddlebags combine with a standard top box to provide a steamer trunk's luggage capacity.

shown in the functional details. In addition to the beautifully finished bodywork, the luxury cruiser boasts an assortment of chrome highlights, including valve covers, exhaust system, saddlebag latches and frame panels, with an optional kit to add even more brightwork. Available colors include Pearl Silver Metallic, Capri Blue Metallic and Mojave Brown Metallic, this last with a choice of black or brown saddle (other colors feature black).

The R 1200 CL Engine: Gearing For The Long Haul

BMW's newest tourer begins with a solid foundation-the 61-hp R 1200 C engine. The original, 1170cc cruiser powerplant blends a broad powerband and instantaneous response with a healthy, 72 lb.-ft. of torque. Like its forebear, the new CL provides its peak torque at 3000 rpm-exactly the kind of power delivery for a touring twin. Motronic MA 2.4 engine management ensures that this Boxer blends this accessible power with long-term reliability and minimal emissions, while at the same time eliminating the choke lever for complete push-button simplicity. Of course, the MoDiTec diagnostic feature makes maintaining the CL every bit as simple as the other members of BMW's stable.

While tourers and cruisers place similar demands on their engines, a touring bike typically operates through a wider speed range. Consequently, the R 1200 CL mates this familiar engine to a new, six-speed transmission. The first five gear ratios are similar to the original R 1200's, but the sixth gear provides a significant overdrive, which drops engine speed well under 3000 rpm at 60 mph. This range of gearing means the CL can manage either responsive in-town running or relaxed freeway cruising with equal finesse, and places the luxury cruiser right in the heart of its powerband at touring speeds for simple roll-on passes.

In addition, the new transmission has been thoroughly massaged internally, with re-angled gear teeth that provide additional overlap for quieter running. Shifting is likewise improved via a revised internal shift mechanism that produces smoother, more precise gearchanges. Finally, the new transmission design is lighter (approximately 1 kg.), which helps keep the CL's weight down to a respectable 679 lbs. (wet). The improved design of this transmission will be adopted by other Boxer-twins throughout the coming year.

The CL Chassis: Wheeled Luggage Never Worked This Well

Every bit as unique as the CL's Boxer-twin drivetrain is the bike's chassis, leading off-literally and figuratively-with BMW's standard-setting Telelever front suspension. The CL's setup is identical in concept and function to the R 1200 C's fork, but shares virtually no parts with the previous cruiser's. The tourer's wider, 16-inch front wheel called for wider-set fork tubes, so the top triple clamp, fork bridge, fork tubes and axle have all been revised, and the axle has switched to a full-floating design. The aluminum Telelever itself has been further reworked to provide a slightly more raked appearance, which also creates a more relaxed steering response for improved straight-line stability. The front shock has been re-angled and its spring and damping rates changed to accommodate the new bike's suspension geometry, but is otherwise similar to the original R 1200 C's damper.

Similarly, the R 1200 CL's Monolever rear suspension differs in detail, rather than concept, from previous BMW cruisers. Increased reinforcing provides additional strength at the shock mount, while a revised final-drive housing provides mounts for the new rear brake. But the primary rear suspension change is a switch to a shock with travel-related damping, similar to that introduced on the R 1150 GS Adventure. This new shock not only provides for a smoother, more controlled ride but also produces an additional 20mm travel compared to the other cruisers, bringing the rear suspension travel to 4.72 inches.

The Telelever and Monolever bolt to a standard R 1200 C front frame that differs only in detail from the original. The rear subframe, however, is completely new, designed to accommodate the extensive luggage system and passenger seating on the R 1200 CL. In addition to the permanently affixed saddlebags, the larger seats, floor boards, top box and new side stand all require new mounting points.

All this new hardware rolls on completely restyled double-spoke wheels (16 x 3.5 front/15 x 4.0 rear) that carry wider, higher-profile (80-series) touring tires for an extremely smooth ride. Bolted to these wheels are larger disc brakes (12.0-inch front, 11.2-inch rear), with the latest edition of BMW's standard-setting EVO brakes. A pair of four-piston calipers stop the front wheel, paired with a two-piston unit-adapted from the K 1200 LT-at the rear. In keeping with the bike's touring orientation, the new CL includes BMW's latest, fully integrated ABS, which actuates both front and rear brakes through either the front hand lever or the rear brake pedal.

The CL Bodywork: Dressed To The Nines

Although all these mechanical changes ensure that the new R 1200 CL works like no other luxury cruiser, it's the bike's styling and bodywork that really set it apart. Beginning with the bike's handlebar-mounted fairing, the CL looks like nothing else on the road, but it's the functional attributes that prove its worth. The broad sweep of the fairing emphasizes its aerodynamic shape, which provides maximum wind protection with a minimum of buffeting. Four headlamps, with their horizontal/vertical orientation, give the CL its unique face and also create the best illumination outside of a baseball stadium (the high-beams are borrowed from the GS).

The M-shaped windshield, with its dipped center section, produces exceptional wind protection yet still allows the rider to look over the clear-plastic shield when rain or road dirt obscure the view. Similarly, clear extensions at the fairing's lower edges improve wind protection even further but still allow an unobstructed view forward for maneuvering in extremely close quarters. The turnsignal pods provide further wind coverage, and at the same time the integral mirrors give a clear view to the rear.

Complementing the fairing, both visually and functionally, the frame-mounted lowers divert the wind blast around the rider to provide further weather protection. Openings vent warm air from the frame-mounted twin oil-coolers and direct the heat away from the rider. As noted earlier, the lowers also house the electronics for the bike's optional alarm system and cruise control. A pair of 12-volt accessory outlets are standard.

Like the K 1200 LT, the new R 1200 CL includes a capacious luggage system as standard, all of it color-matched and designed to accommodate rider and passenger for the long haul. The permanently attached saddlebags include clamshell lids that allow for easy loading and unloading. Chrome bumper strips protect the saddlebags from minor tipover damage. The top box provides additional secure luggage space, or it can be simply unbolted to uncover an attractive aluminum luggage rack. An optional backrest can be bolted on in place of the top box. Of course, saddlebags and top box are lockable and keyed to the ignition switch.

Options & Accessories: More Personal Than A Monogram

Given BMW's traditional emphasis on touring options and the cruiser owner's typical demands for customization, it's only logical to expect a range of accessories and options for the company's first luxury cruiser. The CL fulfills those expectations with a myriad of options and accessories, beginning with heated or velour-like Soft Touch seats and a low windshield. Electronic and communications options such as an AM/FM/CD stereo, cruise control and onboard communication can make time on the road much more pleasant, whether you're out for an afternoon ride or a cross-country trek - because after all, nobody says you have to be back in six days. Other available electronic features include an anti-theft alarm, which also disables the engine.

Accessories designed to personalize the CL even further range from cosmetic to practical, but all adhere to BMW's traditional standards for quality and fit. Chrome accessories include engine-protection and saddlebag - protection hoops. On a practical level, saddlebag and top box liners simplify packing and unpacking. In addition to the backrest, a pair of rear floorboards enhance passenger comfort even more.

- - - - -

Der Luxus-Cruiser zum genußvollen Touren.

Die Motorradwelt war überrascht, als BMW Motorrad 1997 die R 1200 C, den ersten Cruiser in der Geschichte des Hauses, vorstellte. Mit dem einzigartigen Zweizylinder-Boxermotor und einem unverwechselbar eigenständigen Design gelang es auf Anhieb, sich in diesem bis dato von BMW nicht besetzten Marktsegment erfolgreich zu positionieren. Bisher wurden neben dem Basismodell R 1200 C Classic die technisch nahezu identischen Modellvarianten Avantgarde und Independent angeboten, die sich in Farbgebung, Designelementen und Ausstattungsdetails unterscheiden.

Zur Angebotserweiterung und zur Erschließung zusätzlicher Potenziale, präsentiert BMW Motorrad für das Modelljahr 2003 ein neues Mitglied der Cruiserfamilie, den Luxus-Cruiser R 1200 CL. Er wird seine Weltpremiere im September in München auf der INTERMOT haben und voraussichtlich im Herbst 2002 auf den Markt kommen. Der Grundgedanke war, Elemente von Tourenmotorrädern auf einen Cruiser zu übertragen und ein Motorrad zu entwickeln, das Eigenschaften aus beiden Fahrzeuggattungen aufweist.

So entstand ein eigenständiges Modell, ein Cruiser zum genussvollen Touren, bei dem in Komfort und Ausstattung keine Wünsche offen bleiben.

Als technische Basis diente die R 1200 C, von der aber im wesentlichen nur der Motor, der Hinterradantrieb, der Vorderrahmen, der Tank und einige Ausstattungsumfänge übernommen wurden. Ansonsten ist das Motorrad ein völlig eigenständiger Entwurf und in weiten Teilen eine Neuentwicklung.

Fahrgestell und Design:

Einzigartiges Gesicht, optische Präsenz und Koffer integriert.

Präsenz, kraftvoller Auftritt und luxuriöser Charakter, mit diesen Worten lässt sich die Wirkung der BMW R 1200 CL kurz und treffend beschreiben. Geprägt wird dieses Motorrad von der lenkerfesten Tourenverkleidung, deren Linienführung sich in den separaten seitlichen Verkleidungsteilen am Tank fortsetzt, so dass in der Seitenansicht fast der Eindruck einer integrierten Verkleidung entsteht. Sie bietet dem Fahrer ein hohes Maß an Komfort durch guten Wind- und Wetterschutz.

Insgesamt vier in die Verkleidung integrierte Scheinwerfer, zwei für das Abblendlicht und zwei für das Fernlicht, geben dem Motorrad ein unverwechselbares, einzigartiges Gesicht und eine beeindruckende optische Wirkung, die es so bisher noch bei keinem Motorrad gab. Natürlich sorgen die vier Scheinwerfer auch für eine hervorragende Fahrbahnausleuchtung.

Besonders einfallsreich ist die aerodynamische Gestaltung der Verkleidungsscheibe mit ihrem wellenartig ausgeschnittenen oberen Rand. Sie leitet die Strömung so, dass der Fahrer wirkungsvoll geschützt wird. Gleichzeitig kann man aber wegen des Einzugs in der Mitte ungehindert über die Scheibe hinwegschauen und hat somit unabhängig von Nässe und Verschmutzung der Scheibe ein ungestörtes Sichtfeld auf die Straße.

Zur kraftvollen Erscheinung des Motorrades passt der Vorderradkotflügel, der seitlich bis tief zur Felge heruntergezogen ist. Er bietet guten Spritzschutz und unterstreicht zusammen mit dem voluminösen Vorderreifen die Dominanz der Frontpartie, die aber dennoch Gelassenheit und Eleganz ausstrahlt.

Der gegenüber den anderen Modellen flacher gestellte Telelever hebt den Cruisercharakter noch mehr hervor. Der Heckbereich wird bestimmt durch die integrierten, fest mit dem Fahrzeug verbundenen Hartschalenkoffer und das abnehmbare Topcase auf der geschwungenen Gepäckbrücke, die zugleich als Soziushaltegriff dient. Koffer und Topcase sind jeweils in Fahrzeugfarbe lackiert und bilden somit ein harmonisches Ganzes mit dem Fahrzeug.

Akzente setzen auch die stufenförmig angeordneten breiten Komfortsitze für Fahrer und Beifahrer mit der charakteristischen hinteren Abstützung. Luxus durch exklusive Farben, edle Oberflächen und Materialien.

Die R 1200 CL wird zunächst in drei exklusiven Farben angeboten: perlsilber-metallic und capriblau-metallic mit jeweils schwarzen Sitzen und mojavebraun-metallic mit braunem Sitzbezug (wahlweise auch in schwarz). Die Eleganz der Farben wird unterstützt durch sorgfältige Materialauswahl und perfektes Finish von Oberflächen und Fugen. So ist zum Beispiel die Gepäckbrücke aus Aluminium-Druckguß gefertigt und in weissaluminium lackiert, der Lenker verchromt und die obere Instrumentenabdeckung ebenfalls weissaluminiumfarben lackiert. Die Frontverkleidung ist vollständig mit einer Innenabdeckung versehen, und die Kniepads der seitlichen Verkleidungsteile sind mit dem gleichen Material wie die Sitze überzogen.

All dies unterstreicht den Anspruch auf Luxus und Perfektion.

Antrieb jetzt mit neuem, leiserem Sechsganggetriebe - Boxermotor unverändert.

Während der Boxermotor mit 1170 cm³ unverändert von der bisherigen R 1200 C übernommen wurde - auch die Leistungsdaten sind mit 45 kW (61 PS) und 98 Nm Drehmoment bei 3 000 min-1 gleich geblieben -, ist das Getriebe der R 1200 CL neu. Abgeleitet von dem bekannten Getriebe der anderen Boxermodelle hat es jetzt auch sechs Gänge und wurde grundlegend überarbeitet. Als wesentliche Neuerung kommt eine sogenannte Hochverzahnung zum Einsatz. Diese sorgt für einen "weicheren" Zahneingriff und reduziert erheblich die Laufgeräusche der Verzahnung.

Der lang übersetzte, als "overdrive" ausgelegte, sechste Gang erlaubt drehzahlschonendes Fahren auf langen Etappen in der Ebene und senkt dort Verbrauch und Geräusch. Statt eines Schalthebels gibt es eine Schaltwippe für Gangwechsel mit einem lässigen Kick. Schaltkomfort, Geräuscharmut, niedrige Drehzahlen und dennoch genügend Kraft - Eigenschaften, die zum Genusscharakter des Fahrzeugs hervorragend passen.

Dass auch die R 1200 CL, wie jedes seit 1997 neu eingeführte BMW Motorrad weltweit, serienmäßig über die jeweils modernste Abgasreinigungstechnologie mit geregeltem Drei-Wege-Katalysator verfügt, muss fast nicht mehr erwähnt werden. Es ist bei BMW zur Selbstverständlichkeit geworden.

Fahrwerkselemente für noch mehr Komfort - Telelever neu und hinteres Federbein mit wegabhängiger Dämpfung.

Ein cruisertypisches Merkmal ist die nach vorn gestreckte Vorderradführung mit flachem Winkel zur Fahrbahn und großem Nachlauf. Dazu wurde für die R 1200 CL der nach wie vor einzigartige BMW Telelever neu ausgelegt.

Die Gabelholme stehen weiter auseinander, um dem bulligen, 150 mm breiten Vorderradreifen Platz zu bieten.

Für die Hinterradfederung kommt ein Federbein mit wegabhängiger Dämpfung zum Einsatz, das sich durch hervorragende Komforteigenschaften auszeichnet. Der Gesamtfederweg wuchs um 20 mm gegenüber den anderen Cruisermodellen auf jetzt 120 mm. Die Federbasisverstellung zur Anpassung an den Beladungszustand erfolgt hydraulisch über ein bequem zugängliches Handrad.

Hinterradschwinge optimiert und Heckrahmen neu.

Die Hinterradschwinge mit Hinterachsgehäuse, der BMW Monolever, wurde verstärkt und zur Aufnahme einer größeren Hinterradbremse angepasst.

Der verstärkte Heckrahmen ist vollständig neu, um Trittbretter, Kofferhalter, Gepäckbrücke und die neuen Sitze sowie die modifizierte Seitenstütze aufnehmen zu können. Der Vorderrahmen aus Aluminiumguss wurde mit geringfügigen Modifikationen von der bisherigen R 1200 C übernommen.

Räder aus Aluminiumguss, Sitze, Trittbretter und Lenker - alles neu.

Der optische Eindruck eines Motorrades wird ganz wesentlich auch von den Rädern bestimmt. Die R 1200 CL hat avantgardistisch gestaltete neue Gussräder aus Aluminium mit 16 Zoll (vorne) beziehungsweise 15 Zoll (hinten) Felgendurchmesser, die voluminöse Reifen im Format 150/80 vorne und 170/80 hinten aufnehmen.

Die Sitze sind für Fahrer und Beifahrer getrennt ausgeführt, um den unterschiedlichen Bedürfnissen gerecht zu werden. So ist der breite Komfortsattel für den Fahrer mit einer integrierten Beckenabstützung versehen und bietet einen hervorragenden Halt. Die Sitzhöhe beträgt 745 mm. Der Sitz für den Passagier ist ebenfalls ganz auf Bequemlichkeit ausgelegt und etwas höher als der Fahrersitz angeordnet. Dadurch hat der Beifahrer einen besseren Blick am Fahrer vorbei und kann beim Cruisen die Landschaft ungestört genießen.

Großzügige cruisertypische Trittbretter für den Fahrer tragen zum entspannten Sitzen bei. Die Soziusfußrasten, die von der K 1200 LT abgeleitet sind, bieten ebenfalls sehr guten Halt und ermöglichen zusammen mit dem günstigen Kniebeugewinkel auch dem Beifahrer ein ermüdungsfreies Touren.

Der breite, verchromte Lenker vermittelt nicht nur Cruiser-Feeling; Höhe und Kröpfungswinkel sind so ausgelegt, dass auch auf langen Fahrten keine Verspannungen auftreten. Handhebel und Schalter mit der bewährten und eigenständigen BMW Bedienlogik wurden unverändert von den anderen Modellen übernommen.

HighTech bei den Bremsen - BMW EVO-Bremse und als Sonderausstattung Integral ABS.

Sicherheit hat bei BMW traditionell höchste Priorität. Deshalb kommt bei der

R 1200 CL die schon in anderen BMW Motorrädern bewährte EVO-Bremse am Vorderrad zum Einsatz, die sich durch eine verbesserte Bremsleistung auszeichnet. Auf Wunsch gibt es das einzigartige BMW Integral ABS, dem Charakter des Motorrades entsprechend in der Vollintegralversion. Das heißt, unabhängig ob der Hand- oder Fußbremshebel betätigt wird, immer wirkt die Bremskraft optimal auf beide Räder. Im Vorderrad verzögert eine Doppel-Scheibenbremse mit 305 mm Scheibendurchmesser und im Hinterrad die von der K 1200 LT übernommene Einscheiben-Bremsanlage mit einem Scheibendurchmesser von 285 mm.

Fortschrittliche Elektrik: Vierfach-Scheinwerfer, wartungsarme Batterie und elektronischer Tachometer.

Vier Scheinwerfer, je zwei für das Abblend- und Fernlicht, geben dem Motorrad von vorne ein einzigartiges prägnantes Gesicht. Durch die kreuzweise Anordnung - die Abblendscheinwerfer sitzen nebeneinander und die Fernscheinwerfer dazwischen und übereinander - wird eine hohe Signalwirkung bei Tag und eine hervorragende Fahrbahnausleuchtung bei Dunkelheit erzielt.

Neu ist die wartungsarme, komplett gekapselte Gel-Batterie, bei der kein Wasser mehr nachgefüllt werden muss. Eine zweite Steckdose ist serienmäßig. Die Instrumente sind ebenfalls neu. Drehzahlmesser und Tachometer sind elektronisch und die Zifferblätter neu gestaltetet, ebenso die Analoguhr.

Umfangreiche Sonderausstattung für Sicherheit, Komfort und individuellen Luxus.

Die Sonderausstattung der R 1200 CL ist sehr umfangreich und reicht vom BMW Integral ABS für sicheres Bremsen über Komfortausstattungen wie Temporegelung, heizbare Lenkergriffe und Sitzheizung bis hin zu luxuriöser Individualisierung mit Softtouchsitzen, Chrompaket und fernbedientem Radio mit CD-Laufwerk.

The CL's riding position blends elements of both tourer and cruiser, beginning with a reassuringly low, 29.3-inch seat height. The seat itself comprises two parts, a rider portion with an integral lower-back rest, and a taller passenger perch that includes a standard backrest built into the top box. Heated seats, first seen on the K 1200 LT, are also available for the CL to complement the standard heated grips. A broad, flat handlebar places those grips a comfortable reach away, and the CL's floorboards allow the rider to shift position easily without compromising control. Standard cruise control helps melt the miles on long highway stints. A convenient heel/toe shifter makes for effortless gearchanges while adding exactly the right classic touch.

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BA License Plate Bad Aussee Austria BMW R1200 CL (c) Bernard Egger :: rumoto images 5013 cc

Biscayne Building, 19 West Flagler Street, City of Miami, Miami-Dade County, Florida, USA / Built: 1925 / Architect: August Geiger / Floors: 14 / Architectural Style: Mediterranean Revival by Urban Florida Photographer

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The building at 19 West Flagler Street in downtown Miami is the historic Biscayne Building, a 14-story office tower constructed in 1925. Throughout its history, it has served multiple notable tenants and is now part of developer Moishe Mana's extensive revitalization project for the area.

Early history

Construction: The Biscayne Building was built in 1925, during Miami's land boom.

Original tenant: The tower was originally home to the Bank of Biscayne.

FBI offices: At one point in its history, the top three floors of the tower were leased by the FBI.

Long-term ownership: Records indicate that the building was owned by Biscayne Building Inc., managed by Dante Fiorini, since at least the early 1980s.

Recent history and current status

Moishe Mana acquisition: In August 2016, developer Moishe Mana purchased the Biscayne Building for $24.5 million. The acquisition was part of his larger strategy to buy up historic properties along Flagler Street.

Mana's vision: Mana has since acquired a "critical mass" of buildings in the downtown area, with plans to redevelop his portfolio into a mix of retail, office, and residential projects.

Ongoing investment: As a key property within Mana's assemblage, the Biscayne Building and the surrounding area are poised for significant redevelopment as part of a master plan to "return that past vibrancy to the urban core".

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This building is a family owned and operated commercial office building located in the heart of downtown Miami. What distinguishes this office business from others is the pride and detail in which the business is run.

The building is a single asset property and receives our full attention every day unlike other properties of a larger portfolio. The management team is located on site, in Suite 310 on the property. The team at the Biscayne Building has an impeccable reputation for excellence. The average tenant occupancy exceeds 17+ year.

The building is the first historic landmark to achieve the coveted Energy Star in the Central Business District of Miami in 2008. They are very proactive in maintaining and making improvements to the building keeping a high standard of excellence with upgrades in common areas, improved restrooms that are ADA compliant on every floor, adding technology to the elevators, alarm systems, sprinkler systems, HVAC units as well as back systems in place to ensure you are never without air conditioning.

Credit for the data above is given to the following websites:

www.miamidade.gov/Apps/PA/PropertySearch/#/

biscaynebuilding.com.mytempweb.com/

www.google.com/search?q=19+w+flagler+st+history+of+buildi...

floridastories.stqry.app/story/6366

www.google.com/search?q=19+w+flagler+st+history+of+buildi...

www.google.com/search?q=19+w+flagler+street+architect&amp...

Blind egyptian statue at the Turin’s Museo Egizio by Stefano Scanferla

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The afterlife in old Egypt was so true that, once a thief was violating a gravesite, he was first damaging eyes, face and sometimes braking the legs of owner’ statues. This was done to avoid he sees him from the other world and curse them, maybe reach them to duly punish. Basically they were thinking to switch off the alarm system…

Fish (Actinopterygii) by PHOTOGRAPHY by DM & DBM.

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