Two men look at mobile phones atop a house in Uganda. Very soon the newly constructed house on which they are standing will be painted yellow. The other one is red. Most adjacent houses along the highways of Uganda are red and yellow - showing allegiance to either Celtel or MTN, the two main mobile telephony service providers in the country. While basic amenities such as potable water and hygiene have taken a backseat, the economic policies have created a greater rift between the rich and the poor and the country is run by corporate houses dealing in everything from oil to mobile services. One has to buy everything branded. Wonder if the roast cassava and skewers of roast beef are also going to be branded.

I had no idea that my dad kept a collection of his old cellphones and was surprised to see him bring out his stash. I don't think any of them are in working condition and if they did, there was no way to find out as their original chargers are no longer around. 😊

 

What surprised me was that he also kept my old Ericsson GH 688, which I used between 1997 and 2000. This Ericsson unit traveled with me to Malawi, Africa where I was posted for three months in June 1998. It continued to serve me faithfully until I upgraded to the unreliable Ericsson T28s flip phone in mid 2000.

 

I can however identify the earliest cellphone in this picture - the grey Nokia 101 (ETACS), which my father had back in 1992. It operated exclusively on the local Celcom ART 900 analog network and provided only basic telephony service.

 

That's right, neither the Nokia 101 phone nor the network had SMS texting or Caller ID feature! If you had a missed call, you wouldn't know who called you! The redial button was only meant for redialing the last outgoing call - not for incoming ones.

 

Its monochrome LCD display only showed the network ID, contact names, numbers, remaining battery power and signal strength, that's it.

 

I remember borrowing that Nokia 101 and stashing it in the back of my jeans pocket going to the mall. Since it was fairly long, the top part and the antenna showed above my pocket line in clear view of the public.

 

In 1992 cellphones were expensive, status symbol devices and anyone seen carrying one must be someone important like a businessman or a VIP! People often stared at me whenever I received a call on that brick, lol. 😃

 

BTW, you know you're that old if you've actually used an analog cellphone in your late twenties! 😸

  

Thanks everyone for the views and the faves, much appreciated! 😊

One of our secretive visitor this summer... lucky to snap its diving shot!

 

Peregrine Falcon (Falco peregrinus)

 

The peregrine falcon (Falco peregrinus), also known as the peregrine, and historically as the duck hawk in North America, is a widespread bird of prey in the family Falconidae. A large, crow-sized falcon, it has a blue-grey back, barred white underparts, and a black head and "moustache". As is typical of bird-eating raptors, peregrine falcons are sexually dimorphic, females being considerably larger than males. The peregrine is renowned for its speed, reaching over 322 km/h (200 mph) during its characteristic hunting stoop (high speed dive), making it the fastest member of the animal kingdom. According to a National Geographic TV programme, the highest measured speed of a peregrine falcon is 389 km/h (242 mph).

 

The peregrine's breeding range includes land regions from the Arctic tundra to the tropics. It can be found nearly everywhere on Earth, except extreme polar regions, very high mountains, and most tropical rainforests; the only major ice-free landmass from which it is entirely absent is New Zealand. This makes it the world's most widespread raptor and one of the most widely found bird species. In fact, the only land-based bird species found over a larger geographic area is not always naturally occurring but one widely introduced by humans, the rock pigeon, which in turn now supports many peregrine populations as a prey species. Both the English and scientific names of this species mean "wandering falcon", referring to the migratory habits of many northern populations. Experts recognize 17 to 19 subspecies which vary in appearance and range; there is disagreement over whether the distinctive Barbary falcon is represented by two subspecies of Falco peregrinus, or is a separate species, F. pelegrinoides. The two species' divergence is relatively recent, during the time of the last ice age, therefore the genetic differential between them (and also the difference in their appearance) is relatively small. It has been determined that they are only approximately 0.6–0.8% genetically differentiated.

 

While its diet consists almost exclusively of medium-sized birds, the peregrine will occasionally hunt small mammals, small reptiles, or even insects. Reaching sexual maturity at one year, it mates for life and nests in a scrape, normally on cliff edges or, in recent times, on tall human-made structures. The peregrine falcon became an endangered species in many areas because of the widespread use of certain pesticides, especially DDT. Since the ban on DDT from the early 1970s, populations have recovered, supported by large-scale protection of nesting places and releases to the wild.

 

The peregrine falcon is a well respected falconry bird due to its strong hunting ability, high trainability, versatility, and in recent years availability via captive breeding. It is effective on most game bird species from small to large.

  

Description

 

The peregrine falcon has a body length of 34 to 58 cm (13–23 in) and a wingspan from 74 to 120 cm (29–47 in). The male and female have similar markings and plumage, but as in many birds of prey the peregrine falcon displays marked sexual dimorphism in size, with the female measuring up to 30% larger than the male. Males weigh 330 to 1,000 g (0.73–2.20 lb) and the noticeably larger females weigh 700 to 1,500 g (1.5–3.3 lb). In most subspecies, males weigh less than 700 g (1.5 lb) and females weigh more than 800 g (1.8 lb), with cases of females weighing about 50% more than their male breeding mates not uncommon. The standard linear measurements of peregrines are: the wing chord measures 26.5–39 cm (10.4–15.4 in), the tail measures 13–19 cm (5.1–7.5 in) and the tarsus measures 4.5 to 5.6 cm (1.8 to 2.2 in).

 

The back and the long pointed wings of the adult are usually bluish black to slate grey with indistinct darker barring (see "Subspecies" below); the wingtips are black. The white to rusty underparts are barred with thin clean bands of dark brown or black. The tail, coloured like the back but with thin clean bars, is long, narrow, and rounded at the end with a black tip and a white band at the very end. The top of the head and a "moustache" along the cheeks are black, contrasting sharply with the pale sides of the neck and white throat. The cere is yellow, as are the feet, and the beak and claws are black. The upper beak is notched near the tip, an adaptation which enables falcons to kill prey by severing the spinal column at the neck. The immature bird is much browner with streaked, rather than barred, underparts, and has a pale bluish cere and orbital ring.

  

Taxonomy and systematics

 

Falco peregrinus was first described under its current binomial name by English ornithologist Marmaduke Tunstall in his 1771 work Ornithologia Britannica. The scientific name Falco peregrinus is a Medieval Latin phrase that was used by Albertus Magnus in 1225. The specific name taken from the fact that juvenile birds were taken while journeying to their breeding location rather than from the nest, as falcon nests were difficult to get at. The Latin term for falcon, falco, is related to falx, the Latin word meaning sickle, in reference to the silhouette of the falcon's long, pointed wings in flight.

 

The peregrine falcon belongs to a genus whose lineage includes the hierofalcons and the prairie falcon (F. mexicanus). This lineage probably diverged from other falcons towards the end of the Late Miocene or in the Early Pliocene, about 5–8 million years ago (mya). As the peregrine-hierofalcon group includes both Old World and North American species, it is likely that the lineage originated in western Eurasia or Africa. Its relationship to other falcons is not clear, as the issue is complicated by widespread hybridization confounding mtDNA sequence analyses. For example, a genetic lineage of the saker falcon (F. cherrug) is known which originated from a male saker producing fertile young with a female peregrine ancestor, and the descendants further breeding with sakers.

 

Today, peregrines are regularly paired in captivity with other species such as the lanner falcon (F. biarmicus) to produce the "perilanner", a somewhat popular bird in falconry as it combines the peregrine's hunting skill with the lanner's hardiness, or the gyrfalcon to produce large, strikingly coloured birds for the use of falconers. As can be seen, the peregrine is still genetically close to the hierofalcons, though their lineages diverged in the Late Pliocene (maybe some 2.5–2 mya in the Gelasian).

  

Subspecies

 

Numerous subspecies of Falco peregrinus have been described, with 19 accepted by the 1994 Handbook of the Birds of the World, which considers the Barbary falcon of the Canary Islands and coastal north Africa to be two subspecies (pelegrinoides and babylonicus) of Falco peregrinus, rather than a distinct species, F. pelegrinoides. The following map shows the general ranges of these 19 subspecies:

 

Falco peregrinus anatum

- described by Bonaparte in 1838, is known as the American peregrine falcon, or "duck hawk"; its scientific name means "duck peregrine falcon". At one time, it was partly included in leucogenys. It is mainly found in the Rocky Mountains today. It was formerly common throughout North America between the tundra and northern Mexico, where current reintroduction efforts seek to restore the population. Most mature anatum, except those that breed in more northern areas, winter in their breeding range. Most vagrants that reach western Europe seem to belong to the more northern and strongly migratory tundrius, only considered distinct since 1968. It is similar to peregrinus but is slightly smaller; adults are somewhat paler and less patterned below, but juveniles are darker and more patterned below. Males weigh 500 to 700 g (1.1–1.5 lb), while females weigh 800 to 1,100 g (1.8–2.4 lb). It has become extinct in eastern North America, and populations there are hybrids as a result of reintroductions of birds from elsewhere.

 

Falco peregrinus babylonicus

- described by P.L. Sclater in 1861, is found in eastern Iran along the Hindu Kush and Tian Shan to Mongolian Altai ranges. A few birds winter in northern and northwestern India, mainly in dry semi-desert habitats. It is paler than pelegrinoides, and somewhat similar to a small, pale lanner falcon (Falco biarmicus). Males weigh 330 to 400 grams (12 to 14 oz), while females weigh 513 to 765 grams (18.1 to 27.0 oz).

 

Falco peregrinus brookei

- described by Sharpe in 1873, is also known as the Mediterranean peregrine falcon or the Maltese falcon. It includes caucasicus and most specimens of the proposed race punicus, though others may be pelegrinoides, Barbary falcons (see also below), or perhaps the rare hybrids between these two which might occur around Algeria. They occur from the Iberian Peninsula around the Mediterranean, except in arid regions, to the Caucasus. They are non-migratory. It is smaller than the nominate subspecies, and the underside usually has rusty hue. Males weigh around 445 g (0.981 lb), while females weigh up to 920 g (2.03 lb).

 

Falco peregrinus calidus

- described by John Latham in 1790, was formerly called leucogenys and includes caeruleiceps. It breeds in the Arctic tundra of Eurasia, from Murmansk Oblast to roughly Yana and Indigirka Rivers, Siberia. It is completely migratory, and travels south in winter as far as South Asia and sub-Saharan Africa. It is often seen around wetland habitats. It is paler than peregrinus, especially on the crown. Males weigh 588 to 740 g (1.296–1.631 lb), while females weigh 925 to 1,333 g (2.039–2.939 lb).

 

Falco peregrinus cassini

- described by Sharpe in 1873, is also known as the Austral peregrine falcon. It includes kreyenborgi, the pallid falcon, a leucistic morph occurring in southernmost South America, which was long believed to be a distinct species. Its range includes South America from Ecuador through Bolivia, northern Argentina, and Chile to Tierra del Fuego and the Falkland Islands. It is non-migratory. It is similar to nominate, but slightly smaller with a black ear region. The variation kreyenborgi is medium grey above, has little barring below, and has a head pattern like the saker falcon, but the ear region is white.

 

Falco peregrinus ernesti

- described by Sharpe in 1894, is found from Indonesia to Philippines and south to Papua New Guinea and the nearby Bismarck Archipelago. Its geographical separation from nesiotes requires confirmation. It is non-migratory. It differs from the nominate subspecies in the very dark, dense barring on its underside and its black ear coverts.

 

Falco peregrinus furuitii

- described by Momiyama in 1927, is found on the Izu and Ogasawara Islands south of Honshū, Japan. It is non-migratory. It is very rare, and may only remain on a single island. It is a dark form, resembling pealei in colour, but darker, especially on tail.

 

Falco peregrinus japonensis

- described by Gmelin in 1788, includes kleinschmidti, pleskei, and harterti, and seems to refer to intergrades with calidus. It is found from northeast Siberia to Kamchatka (though it is possibly replaced by pealei on the coast there) and Japan. Northern populations are migratory, while those of Japan are resident. It is similar to peregrinus, but the young are even darker than those of anatum.

 

Falco peregrinus macropus

- described by Swainson in 1837, is the Australian peregrine falcon. It is found in Australia in all regions except the southwest. It is non-migratory. It is similar to brookei in appearance, but is slightly smaller and the ear region is entirely black. The feet are proportionally large.

 

Falco peregrinus madens

- described by Ripley and Watson in 1963, is unusual in having some sexual dichromatism. If the Barbary falcon is considered a distinct species, it is sometimes placed therein. It is found in the Cape Verde Islands, and is non-migratory; it is endangered with only six to eight pairs surviving. Males have a rufous wash on crown, nape, ears, and back; underside conspicuously washed pinkish-brown. Females are tinged rich brown overall, especially on the crown and nape.

 

Falco peregrinus minor

- first described by Bonaparte in 1850. It was formerly often perconfusus. It is sparsely and patchily distributed throughout much of sub-Saharan Africa and widespread in Southern Africa. It apparently reaches north along the Atlantic coast as far as Morocco. It is non-migratory and dark coloured. This is the smallest subspecies of peregrine, with smaller males weighing as little as approximately 300 g (11 oz).

 

Falco peregrinus nesiotes

- described by Mayr in 1941, is found in Fiji and probably also Vanuatu and New Caledonia. It is non-migratory.

 

Falco peregrinus pealei

- described by Ridgway in 1873, is also known as Peale's falcon, and includes rudolfi. It is found in the Pacific Northwest of North America, northwards from the Puget Sound along the British Columbia coast (including the Queen Charlotte Islands), along the Gulf of Alaska and the Aleutian Islands to the far eastern Bering Sea coast of Russia, and may also occur on the Kuril Islands and the coasts of Kamchatka. It is non-migratory. It is the largest subspecies, and it looks like an oversized and darker tundrius or like a strongly barred and large anatum. The bill is very wide. Juveniles occasionally have pale crowns. Males weigh 700 to 1,000 g (1.5–2.2 lb), while females weigh 1,000 to 1,500 g (2.2–3.3 lb).

 

Falco peregrinus pelegrinoides

- first described by Temminck in 1829, is found in the Canary Islands through north Africa and the Near East to Mesopotamia. It is most similar to brookei, but is markedly paler above, with a rusty neck, and is a light buff with reduced barring below. It is smaller than the nominate subspecies; females weigh around 610 g (1.34 lb).

 

Falco peregrinus peregrinator

- described by Sundevall in 1837, is known as the Indian peregrine falcon, Shaheen falcon, Indian shaheen or shaheen falcon. It was formerly sometimes known as Falco atriceps or Falco shaheen. Its range includes South Asia from Pakistan across India and Bangladesh to Sri Lanka and Southeastern China. In India, the shaheen is reported from all states except Uttar Pradesh, mainly from rocky and hilly regions. The Shaheen is also reported from the Andaman and Nicobar Islands in the Bay of Bengal. It has a clutch size of 3 to 4 eggs, with the chicks fledging time of 48 days with an average nesting success of 1.32 chicks per nest. In India, apart from nesting on cliffs, it has also been recorded as nesting on man-made structures such as buildings and cellphone transmission towers. A population estimate of 40 breeding pairs in Sri Lanka was made in 1996. It is non-migratory, and is small and dark, with rufous underparts. In Sri Lanka this species is found to favour the higher hills while the migrant calidus is more often seen along the coast.

 

Falco peregrinus peregrinus

- the nominate (first-named) subspecies, described by Tunstall in 1771, breeds over much of temperate Eurasia between the tundra in the north and the Pyrenees, Mediterranean region and Alpide belt in the south. It is mainly non-migratory in Europe, but migratory in Scandinavia and Asia. Males weigh 580 to 750 g (1.28–1.65 lb), while females weigh 925 to 1,300 g (2.039–2.866 lb). It includes brevirostris, germanicus, rhenanus, and riphaeus.

 

Falco peregrinus radama

- described by Hartlaub in 1861, is found in Madagascar and Comoros. It is non-migratory.

 

Falco peregrinus submelanogenys

- described by Mathews in 1912, is the Southwest Australian peregrine falcon. It is found in southwest Australia and is non-migratory.

 

Falco peregrinus tundrius

- described by C.M. White in 1968, was at one time included in leucogenys It is found in the Arctic tundra of North America to Greenland, and migrates to wintering grounds in Central and South America. Most vagrants that reach western Europe belong to this subspecies, which was previously united with anatum. It is the New World equivalent to calidus. It is smaller than anatum. It is also paler than anatum; most have a conspicuous white forehead and white in ear region, but the crown and "moustache" are very dark, unlike in calidus. Juveniles are browner, and less grey, than in calidus, and paler, sometimes almost sandy, than in anatum. Males weigh 500 to 700 g (1.1–1.5 lb), while females weigh 800 to 1,100 g (1.8–2.4 lb).

  

Barbary falcon (Main article: Barbary falcon)

 

Two of the subspecies listed above (Falco peregrinus pelegrinoides and F. p. babylonicus) are often instead treated together as a distinct species, Falco pelegrinoides (the Barbary falcon), although they were included within F. peregrinus in the 1994 Handbook of the Birds of the World. These birds inhabit arid regions from the Canary Islands along the rim of the Sahara through the Middle East to Central Asia and Mongolia.

 

Barbary falcons have a red neck patch but otherwise differ in appearance from the peregrine proper merely according to Gloger's Rule, relating pigmentation to environmental humidity. The Barbary falcon has a peculiar way of flying, beating only the outer part of its wings like fulmars sometimes do; this also occurs in the peregrine, but less often and far less pronounced. The Barbary falcon's shoulder and pelvis bones are stout by comparison with the peregrine, and its feet are smaller. Barbary falcons breed at different times of year than neighboring peregrine falcon subspecies, but they are capable of interbreeding. There is a 0.6–0.7% genetic distance in the peregrine-Barbary falcon ("peregrinoid") complex.

 

Another subspecies of Falco peregrinus, madens, has also sometimes been treated instead within a separately recognized F. pelegrinoides.

  

Ecology and behaviour

 

The peregrine falcon lives mostly along mountain ranges, river valleys, coastlines, and increasingly in cities. In mild-winter regions, it is usually a permanent resident, and some individuals, especially adult males, will remain on the breeding territory. Only populations that breed in Arctic climates typically migrate great distances during the northern winter.

 

The peregrine falcon reaches faster speeds than any other animal on the planet when performing the stoop, which involves soaring to a great height and then diving steeply at speeds of over 320 km/h (200 mph), hitting one wing of its prey so as not to harm itself on impact. The air pressure from such a dive could possibly damage a bird's lungs, but small bony tubercles on a falcon's nostrils guide the powerful airflow away from the nostrils, enabling the bird to breathe more easily while diving by reducing the change in air pressure. To protect their eyes, the falcons use their nictitating membranes (third eyelids) to spread tears and clear debris from their eyes while maintaining vision. A study testing the flight physics of an "ideal falcon" found a theoretical speed limit at 400 km/h (250 mph) for low altitude flight and 625 km/h (388 mph) for high altitude flight. In 2005, Ken Franklin recorded a falcon stooping at a top speed of 389 km/h (242 mph).

 

The life span of peregrine falcons in the wild is up to 15.5 years. Mortality in the first year is 59–70%, declining to 25–32% annually in adults. Apart from such anthropogenic threats as collision with human-made objects, the peregrine may be killed by larger hawks and owls.

 

The peregrine falcon is host to a range of parasites and pathogens. It is a vector for Avipoxvirus, Newcastle disease virus, Falconid herpesvirus 1 (and possibly other Herpesviridae), and some mycoses and bacterial infections. Endoparasites include Plasmodium relictum (usually not causing malaria in the peregrine falcon), Strigeidae trematodes, Serratospiculum amaculata (nematode), and tapeworms. Known peregrine falcon ectoparasites are chewing lice, Ceratophyllus garei (a flea), and Hippoboscidae flies (Icosta nigra, Ornithoctona erythrocephala).

 

Feeding

 

The peregrine falcon feeds almost exclusively on medium-sized birds such as pigeons and doves, waterfowl, songbirds, and waders. Worldwide, it is estimated that between 1,500 and 2,000 bird species (up to roughly a fifth of the world's bird species) are predated somewhere by these falcons. In North America, prey has varied in size from 3 g (0.11 oz) hummingbirds (Selasphorus and Archilochus ssp.) to a 3.1 kg (6.8 lb) sandhill crane (killed in Alaska by a peregrine in a stoop), although most prey taken by peregrines weigh from 20 g (0.71 oz) (i.e. small passerines) to 1,100 g (2.4 lb) (i.e. ducks and gulls). The peregrine falcon takes the most diverse range of bird species of any raptor in North America, with more than 300 species having fallen victim to the falcon, including nearly 100 shorebirds. Smaller hawks and owls are regularly predated, mainly smaller falcons such as the American kestrel, merlin and sharp-shinned hawks. In urban areas, the main component of the peregrine's diet is the rock or feral pigeon, which comprise 80% or more of the dietary intake for peregrines in some cities. Other common city birds are also taken regularly, including mourning doves, common wood pigeons, common swifts, northern flickers, common starlings, American robins, common blackbirds, and corvids (such as magpies or carrion, house, and American crows). Other than bats taken at night, the peregrine rarely hunts mammals, but will on occasion take small species such as rats, voles, hares, shrews, mice and squirrels. Coastal populations of the large subspecies pealei feed almost exclusively on seabirds. In the Brazilian mangrove swamp of Cubatão, a wintering falcon of the subspecies tundrius was observed while successfully hunting a juvenile scarlet ibis. Insects and reptiles make up a small proportion of the diet, which varies greatly depending on what prey is available.

 

The peregrine falcon hunts most often at dawn and dusk, when prey are most active, but also nocturnally in cities, particularly during migration periods when hunting at night may become prevalent. Nocturnal migrants taken by peregrines include species as diverse as yellow-billed cuckoo, black-necked grebe, virginia rail, and common quail. The peregrine requires open space in order to hunt, and therefore often hunts over open water, marshes, valleys, fields, and tundra, searching for prey either from a high perch or from the air. Large congregations of migrants, especially species that gather in the open like shorebirds, can be quite attractive to hunting peregrines. Once prey is spotted, it begins its stoop, folding back the tail and wings, with feet tucked. Prey is typically struck and captured in mid-air; the peregrine falcon strikes its prey with a clenched foot, stunning or killing it with the impact, then turns to catch it in mid-air. If its prey is too heavy to carry, a peregrine will drop it to the ground and eat it there. If they miss the initial strike, peregrines will chase their prey in a twisting flight. Although previously thought rare, several cases of peregrines contour-hunting, i.e. using natural contours to surprise and ambush prey on the ground, have been reported and even rare cases of prey being pursued on foot. In addition, peregrines have been documented preying on chicks in nests, from birds such as kittiwakes. Prey is plucked before consumption.

  

Reproduction

 

The peregrine falcon is sexually mature at one to three years of age, but in healthy populations they breed after two to three years of age. A pair mates for life and returns to the same nesting spot annually. The courtship flight includes a mix of aerial acrobatics, precise spirals, and steep dives. The male passes prey it has caught to the female in mid-air. To make this possible, the female actually flies upside-down to receive the food from the male's talons.

 

During the breeding season, the peregrine falcon is territorial; nesting pairs are usually more than 1 km (0.62 mi) apart, and often much farther, even in areas with large numbers of pairs. The distance between nests ensures sufficient food supply for pairs and their chicks. Within a breeding territory, a pair may have several nesting ledges; the number used by a pair can vary from one or two up to seven in a 16-year period.

 

The peregrine falcon nests in a scrape, normally on cliff edges. The female chooses a nest site, where she scrapes a shallow hollow in the loose soil, sand, gravel, or dead vegetation in which to lay eggs. No nest materials are added. Cliff nests are generally located under an overhang, on ledges with vegetation. South-facing sites are favoured. In some regions, as in parts of Australia and on the west coast of northern North America, large tree hollows are used for nesting. Before the demise of most European peregrines, a large population of peregrines in central and western Europe used the disused nests of other large birds. In remote, undisturbed areas such as the Arctic, steep slopes and even low rocks and mounds may be used as nest sites. In many parts of its range, peregrines now also nest regularly on tall buildings or bridges; these human-made structures used for breeding closely resemble the natural cliff ledges that the peregrine prefers for its nesting locations.

 

The pair defends the chosen nest site against other peregrines, and often against ravens, herons, and gulls, and if ground-nesting, also such mammals as foxes, wolverines, felids, bears, wolves, and mountain lions. Both nests and (less frequently) adults are predated by larger-bodied raptorial birds like eagles, large owls, or gyrfalcons. The most serious predators of peregrine nests in North America and Europe are the great horned owl and the Eurasian eagle owl. When reintroductions have been attempted for peregrines, the most serious impediments were these two owls routinely picking off nestlings, fledglings and adults by night. Peregrines defending their nests have managed to kill raptors as large as golden eagles and bald eagles (both of which they normally avoid as potential predators) that have come too close to the nest by ambushing them in a full stoop. In one instance, when a snowy owl killed a newly fledged peregrine, the larger owl was in turn killed by a stooping peregrine parent.

 

The date of egg-laying varies according to locality, but is generally from February to March in the Northern Hemisphere, and from July to August in the Southern Hemisphere, although the Australian subspecies macropus may breed as late as November, and equatorial populations may nest anytime between June and December. If the eggs are lost early in the nesting season, the female usually lays another clutch, although this is extremely rare in the Arctic due to the short summer season. Generally three to four eggs, but sometimes as few as one or as many as five, are laid in the scrape. The eggs are white to buff with red or brown markings. They are incubated for 29 to 33 days, mainly by the female, with the male also helping with the incubation of the eggs during the day, but only the female incubating them at night. The average number of young found in nests is 2.5, and the average number that fledge is about 1.5, due to the occasional production of infertile eggs and various natural losses of nestlings.

 

After hatching, the chicks (called "eyases") are covered with creamy-white down and have disproportionately large feet. The male (called the "tiercel") and the female (simply called the "falcon") both leave the nest to gather prey to feed the young. The hunting territory of the parents can extend a radius of 19 to 24 km (12 to 15 mi) from the nest site. Chicks fledge 42 to 46 days after hatching, and remain dependent on their parents for up to two months.

  

Relationship with humans

 

Use in falconry (Main article: Falconry)

 

The peregrine falcon is a highly admired falconry bird, and has been used in falconry for more than 3,000 years, beginning with nomads in central Asia. Its advantages in falconry include not only its athleticism and eagerness to hunt, but an equitable disposition that leads to it being one of the easier falcons to train. The peregrine falcon has the additional advantage of a natural flight style of circling above the falconer ("waiting on") for game to be flushed, and then performing an effective and exciting high speed diving stoop to take the quarry. The speed and energy of the stoop allows the falcon to catch fast flying birds, and to deliver a knock out blow with a fist-like clenched talon against game that may be much larger than itself. Additionally the versatility of the species, with agility allowing capture of smaller birds and a strength and attacking style allowing capture of game much larger than themselves, combined with the wide size range of the many peregrine subspecies, means there is a subspecies suitable to almost any size and type of game bird. This size range, evolved to fit various environments and prey species, is from the larger females of the largest subspecies to the smaller males of the smallest subspecies, approximately five to one (approximately 1500 g to 300 g). The males of smaller and medium-sized subspecies, and the females of the smaller subspecies, excel in the taking of swift and agile small game birds such as dove, quail, and smaller ducks. The females of the larger subspecies are capable of taking large and powerful game birds such as the largest of duck species, pheasant, and grouse.

 

Peregrine falcons are also occasionally used to scare away birds at airports to reduce the risk of bird-plane strikes, improving air-traffic safety, and were used to intercept homing pigeons during World War II.

 

Peregrine falcons have been successfully bred in captivity, both for falconry and for release back into the wild. Until 2004 nearly all peregrines used for falconry in the US were captive-bred from the progeny of falcons taken before the US Endangered Species Act was enacted and from those few infusions of wild genes available from Canada and special circumstances. Peregrine falcons were removed from the United States' endangered species list in 1999. The successful recovery program was aided by the effort and knowledge of falconers – in collaboration with The Peregrine Fund and state and federal agencies – through a technique called hacking. Finally, after years of close work with the US Fish and Wildlife Service, a limited take of wild peregrines was allowed in 2004, the first wild peregrines taken specifically for falconry in over 30 years.

 

The development of captive breeding methods has led to peregrines being commercially available for falconry use, thus mostly eliminating the need to capture wild birds for support of falconry. The main reason for taking wild peregrines at this point is to maintain healthy genetic diversity in the breeding lines. Hybrids of peregrines and gyrfalcons are also available that can combine the best features of both species to create what many consider to be the ultimate falconry bird for the taking of larger game such as the sage-grouse. These hybrids combine the greater size, strength, and horizontal speed of the gyrfalcon with the natural propensity to stoop and greater warm weather tolerance of the peregrine.

 

Since peregrine eggs and chicks are still often targeted by illegal collectors, it is common practice not to publicize unprotected nest locations.

  

Decline due to pesticides

 

The peregrine falcon became an endangered species because of the use of organochlorine pesticides, especially DDT, during the 1950s, '60s, and '70s. Pesticide biomagnification caused organochlorine to build up in the falcons' fat tissues, reducing the amount of calcium in their eggshells. With thinner shells, fewer falcon eggs survived to hatching. In several parts of the world, such as the eastern United States and Belgium, this species became extirpated (locally extinct) as a result. An alternate point of view is that populations in the eastern North America had vanished due to hunting and egg collection.

  

Recovery efforts

 

In the United States, Canada, Germany and Poland, wildlife services in peregrine falcon recovery teams breed the species in captivity. The chicks are usually fed through a chute or with a hand puppet mimicking a peregrine's head, so they cannot see to imprint on the human trainers. Then, when they are old enough, the rearing box is opened, allowing the bird to train its wings. As the fledgling gets stronger, feeding is reduced, forcing the bird to learn to hunt. This procedure is called hacking back to the wild. To release a captive-bred falcon, the bird is placed in a special cage at the top of a tower or cliff ledge for some days or so, allowing it to acclimate itself to its future environment.

 

Worldwide recovery efforts have been remarkably successful. The widespread restriction of DDT use eventually allowed released birds to breed successfully. The peregrine falcon was removed from the U.S. Endangered Species list on August 25, 1999.

 

Some controversy has existed over the origins of captive breeding stock used by The Peregrine Fund in the recovery of peregrine falcons throughout the contiguous United States. Several peregrine subspecies were included in the breeding stock, including birds of Eurasian origin. Due to the extirpation of the eastern anatum (Falco peregrinus anatum), the near extirpation of the anatum in the Midwest, and the limited gene pool within North American breeding stock, the inclusion of non-native subspecies was justified to optimize the genetic diversity found within the species as a whole.

  

Current status

 

Populations of the peregrine falcon have bounced back in most parts of the world. In Britain, there has been a recovery of populations since the crash of the 1960s. This has been greatly assisted by conservation and protection work led by the Royal Society for the Protection of Birds. The RSPB has estimated that there are 1,402 breeding pairs in the UK. Peregrines now breed in many mountainous and coastal areas, especially in the west and north, and nest in some urban areas, capitalising on the urban feral pigeon populations for food. In Southampton, a nest prevented restoration of mobile telephony services for several months, after Vodafone engineers despatched to repair a faulty transmitter mast discovered a nest in the mast, and were prevented by the Wildlife and Countryside Act, on pain of a possible prison sentence, from proceeding with repairs until the chicks fledged. In many parts of the world peregrine falcons have adapted to urban habitats, nesting on cathedrals, skyscraper window ledges, tower blocks, and the towers of suspension bridges. Many of these nesting birds are encouraged, sometimes gathering media attention and often monitored by cameras.

 

From an ecological perspective, raptor populations in urban areas are highly beneficial. Compared with Europe where pigeon populations have exploded to the point they are both a tourist attraction and a public nuisance. Their faeces are highly acidic causing damage to historic buildings and statues made of soft stone. They nest in bridges where it compiles and damages iron work causing rust and corrosion. In the United States, falcon and other raptors are in numbers high enough to ward off pigeon nest building in major highrises.

  

Cultural significance

 

Due to its striking hunting technique, the peregrine has often been associated with aggression and martial prowess. Native Americans of the Mississippian culture (c. 800–1500) used the peregrine, along with several other birds of prey, in imagery as a symbol of "aerial (celestial) power" and buried men of high status in costumes associating to the ferocity of "raptorial" birds. In the late Middle Ages, the Western European nobility that used peregrines for hunting, considered the bird associated with princes in formal hierarchies of birds of prey, just below the gyrfalcon associated with kings. It was considered "a royal bird, more armed by its courage than its claws". Terminology used by peregrine breeders also used the Old French term gentil, "of noble birth; aristocratic", particularly with the peregrine.

 

The peregrine falcon is the national animal of the United Arab Emirates. Since 1927, the peregrine falcon has been the official mascot of Bowling Green State University in Bowling Green, Ohio. The 2007 U.S. Idaho state quarter features a peregrine falcon. The peregrine falcon has been designated the official city bird of Chicago.

  

[Credit: en.wikipedia.org]

 

The peregrine falcon (Falco peregrinus), also known simply as the peregrine, and historically as the duck hawk in North America, is a cosmopolitan bird of prey (raptor) in the family Falconidae. A large, crow-sized falcon, it has a blue-grey back, barred white underparts, and a black head. The peregrine is renowned for its speed. It can reach over 320 km/h (200 mph) during its characteristic hunting stoop (high-speed dive), making it the fastest member of the animal kingdom. According to a National Geographic TV program, the highest measured speed of a peregrine falcon is 389 km/h (242 mph). As is typical for bird-eating (avivore) raptors, peregrine falcons are sexually dimorphic, with females being considerably larger than males.

 

The peregrine's breeding range includes land regions from the Arctic tundra to the tropics. It can be found nearly everywhere on Earth, except extreme polar regions, very high mountains, and most tropical rainforests; the only major ice-free landmass from which it is entirely absent is New Zealand. This makes it the world's most widespread raptor and one of the most widely found wild bird species. In fact, the only land-based bird species found over a larger geographic area owes its success to human-led introduction; the domestic and feral pigeons are both domesticate forms of the rock dove, which are a major prey species for Eurasian Peregrine populations. Due to their prevalence over most other bird species in cities, feral pigeons support many peregrine populations as a staple food source, especially in urban settings.

 

The peregrine is a highly successful example of urban wildlife in much of its range, taking advantage of tall buildings as nest sites and an abundance of prey such as pigeons and ducks. Both the English and scientific names of this species mean "wandering falcon", referring to the migratory habits of many northern populations. Experts recognize 17 to 19 subspecies, which vary in appearance and range; disagreement exists over whether the distinctive Barbary falcon is represented by two subspecies of Falco peregrinus or is a separate species, F. pelegrinoides. The two species' divergence is relatively recent, during the time of the last ice age, therefore the genetic differential between them (and also the difference in their appearance) is relatively tiny. They are only about 0.6–0.8% genetically differentiated.

 

Although its diet consists almost exclusively of medium-sized birds, the peregrine will sometimes hunt small mammals, small reptiles, or even insects. Reaching sexual maturity at one year, it mates for life and nests in a scrape, normally on cliff edges or, in recent times, on tall human-made structures. The peregrine falcon became an endangered species in many areas because of the widespread use of certain pesticides, especially DDT. Since the ban on DDT from the early 1970s, populations have recovered, supported by large-scale protection of nesting places and releases to the wild.

 

The peregrine falcon is a well-respected falconry bird due to its strong hunting ability, high trainability, versatility, and availability via captive breeding. It is effective on most game bird species, from small to large. It has also been used as a religious, royal, or national symbol across multiple eras and areas of human civilization.

 

Description

 

Falco peregrinus. Royal National Park, New South Wales, Australia

The peregrine falcon has a body length of 34 to 58 cm (13–23 in) and a wingspan from 74 to 120 cm (29–47 in). The male and female have similar markings and plumage but, as with many birds of prey, the peregrine falcon displays marked sexual dimorphism in size, with the female measuring up to 30% larger than the male. Males weigh 330 to 1,000 g (12–35 oz) and the noticeably larger females weigh 700 to 1,500 g (25–53 oz). In most subspecies, males weigh less than 700 g (25 oz) and females weigh more than 800 g (28 oz), and cases of females weighing about 50% more than their male breeding mates are not uncommon. The standard linear measurements of peregrines are: the wing chord measures 26.5 to 39 cm (10.4–15.4 in), the tail measures 13 to 19 cm (5.1–7.5 in) and the tarsus measures 4.5 to 5.6 cm (1.8–2.2 in).

 

The back and the long pointed wings of the adult are usually bluish black to slate grey with indistinct darker barring (see "Subspecies" below); the wingtips are black. The white to rusty underparts are barred with thin clean bands of dark brown or black. The tail, coloured like the back but with thin clean bars, is long, narrow, and rounded at the end with a black tip and a white band at the very end. The top of the head and a "moustache" along the cheeks are black, contrasting sharply with the pale sides of the neck and white throat. The cere is yellow, as are the feet, and the beak and claws are black. The upper beak is notched near the tip, an adaptation which enables falcons to kill prey by severing the spinal column at the neck. An immature bird is much browner, with streaked, rather than barred, underparts, and has a pale bluish cere and orbital ring.

 

A study shows that their black malar stripe exists to reduce glare from solar radiation, allowing them to see better. Photos from The Macaulay Library and iNaturalist showed that the malar stripe is thicker where there is more solar radiation. That supports the solar glare hypothesis.

 

Taxonomy and systematics

 

Falco peregrinus was first described under its current binomial name by English ornithologist Marmaduke Tunstall in his 1771 work Ornithologia Britannica. The scientific name Falco peregrinus is a Medieval Latin phrase that was used by Albertus Magnus in 1225. Peregrinus is Latin, meaning "one from abroad" or "coming from foreign parts". It is likely the name was used as juvenile birds were taken while journeying to their breeding location (rather than from the nest), as falcon nests are often difficult to get at. The Latin term for falcon, falco, is related to falx, meaning "sickle", in reference to the silhouette of the falcon's long, pointed wings in flight.

 

The peregrine falcon belongs to a genus whose lineage includes the hierofalcon and the prairie falcon (F. mexicanus). This lineage probably diverged from other falcons towards the end of the Late Miocene or in the Late Pliocene, about 3–8 million years ago (mya). As the peregrine-hierofalcon group includes both Old World and North American species, it is likely that the lineage originated in western Eurasia or Africa. Its relationship to other falcons is not clear, as the issue is complicated by widespread hybridization confounding mtDNA sequence analyses. One genetic lineage of the saker falcon (F. cherrug) is known to have originated from a male saker ancestor producing fertile young with a female peregrine ancestor, and the descendants further breeding with sakers.

 

Today, peregrines are regularly paired in captivity with other species such as the lanner falcon (F. biarmicus) to produce the "perilanner", a somewhat popular bird in falconry as it combines the peregrine's hunting skill with the lanner's hardiness, or the gyrfalcon to produce large, strikingly coloured birds for the use of falconers. As can be seen, the peregrine is still genetically close to the hierofalcons, though their lineages diverged in the Late Pliocene (maybe some 2.5–2 mya in the Gelasian).

 

Subspecies

Numerous subspecies of Falco peregrinus have been described, with 19 accepted by the 1994 Handbook of the Birds of the World, which considers the Barbary falcon of the Canary Islands and coastal North Africa to be two subspecies (pelegrinoides and babylonicus) of Falco peregrinus, rather than a distinct species, F. pelegrinoides. The following map shows the general ranges of these 19 subspecies.

 

A map of the world, green shows on several continents, but there are also several big bare spots marked with E for extinct

Breeding ranges of the 19 subspecies

 

Falco peregrinus anatum, described by Bonaparte in 1838, is known as the American peregrine falcon or "duck hawk"; its scientific name means "duck peregrine falcon". At one time, it was partly included in leucogenys. It is mainly found in the Rocky Mountains. It was formerly common throughout North America between the tundra and northern Mexico, where current reintroduction efforts are being made to restore the population. Most mature anatum, except those that breed in more northern areas, winter in their breeding range. Most vagrants that reach western Europe seem to belong to the more northern and strongly migratory tundrius, only considered distinct since 1968. It is similar to the nominate subspecies but is slightly smaller; adults are somewhat paler and less patterned below, but juveniles are darker and more patterned below. Males weigh 500 to 700 g (1.1–1.5 lb), while females weigh 800 to 1,100 g (1.8–2.4 lb). It has become extinct in eastern North America and populations there are hybrids as a result of reintroductions of birds from elsewhere.

Falco peregrinus babylonicus, described by P.L. Sclater in 1861, is found in eastern Iran along the Hindu Kush and the Tian Shan to the Mongolian Altai ranges. A few birds winter in northern and northwestern India, mainly in dry semi-desert habitats. It is paler than pelegrinoides and somewhat similar to a small, pale lanner falcon (Falco biarmicus). Males weigh 330 to 400 grams (12 to 14 oz), while females weigh 513 to 765 grams (18.1 to 27.0 oz).

Falco peregrinus brookei, described by Sharpe in 1873, is also known as the Mediterranean peregrine falcon or the Maltese falcon. It includes caucasicus and most specimens of the proposed race punicus, though others may be pelegrinoides (Barbary falcons), or perhaps the rare hybrids between these two which might occur around Algeria. They occur from the Iberian Peninsula around the Mediterranean, except in arid regions, to the Caucasus. They are non-migratory. It is smaller than the nominate subspecies and the underside usually has a rusty hue. Males weigh around 445 g (0.981 lb), while females weigh up to 920 g (2.03 lb).

Falco peregrinus calidus, described by John Latham in 1790, it was formerly called leucogenys and includes caeruleiceps. It breeds in the Arctic tundra of Eurasia from Murmansk Oblast to roughly Yana and Indigirka Rivers, Siberia. It is completely migratory and travels south in winter as far as South Asia and sub-Saharan Africa. It is often seen around wetland habitats. It is paler than the nominate subspecies, especially on the crown. Males weigh 588 to 740 g (1.296–1.631 lb), while females weigh 925 to 1,333 g (2.039–2.939 lb).

Falco peregrinus cassini, described by Sharpe in 1873, is also known as the austral peregrine falcon. It includes kreyenborgi, the pallid falcon, a leucistic colour morph occurring in southernmost South America, which was long believed to be a distinct species. Its range includes South America from Ecuador through Bolivia, northern Argentina and Chile to Tierra del Fuego and the Falkland Islands. It is non-migratory. It is similar to the nominate subspecies, but slightly smaller with a black ear region. The pallid falcon morph kreyenborgi is medium grey above, has little barring below and has a head pattern like the saker falcon (Falco cherrug), but the ear region is white.

Falco peregrinus ernesti, described by Sharpe in 1894, is found from the Sunda Islands to the Philippines and south to eastern New Guinea and the nearby Bismarck Archipelago. Its geographical separation from nesiotes requires confirmation. It is non-migratory. It differs from the nominate subspecies in the very dark, dense barring on its underside and its black ear coverts.

Falco peregrinus furuitii, described by Momiyama in 1927, is found on the Izu and Ogasawara Islands south of Honshū, Japan. It is non-migratory. It is very rare and may only remain on a single island. It is a dark form, resembling pealei in colour, but darker, especially on the tail.

Falco peregrinus japonensis, described by Gmelin in 1788, includes kleinschmidti, pleskei, and harterti, and seems to refer to intergrades with calidus. It is found from northeast Siberia to Kamchatka (though it is possibly replaced by pealei on the coast there) and Japan. Northern populations are migratory, while those of Japan are resident. It is similar to the nominate subspecies, but the young are even darker than those of anatum.

Falco peregrinus macropus, described by Swainson in 1837, is the Australian peregrine falcon. It is found in Australia in all regions except the southwest. It is non-migratory. It is similar to brookei in appearance, but is slightly smaller and the ear region is entirely black. The feet are proportionally large.

Falco peregrinus madens, described by Ripley and Watson in 1963, is unusual in having some sexual dichromatism. If the Barbary falcon (see below) is considered a distinct species, it is sometimes placed therein. It is found in the Cape Verde Islands and is non-migratory; it is also endangered, with only six to eight pairs surviving. Males have a rufous wash on the crown, nape, ears and back; the underside is conspicuously washed pinkish-brown. Females are tinged rich brown overall, especially on the crown and nape.

 

Falco peregrinus minor, first described by Bonaparte in 1850. It was formerly often known as perconfusus. It is sparsely and patchily distributed throughout much of sub-Saharan Africa and widespread in Southern Africa. It apparently reaches north along the Atlantic coast as far as Morocco. It is non-migratory and dark-coloured. This is the smallest subspecies, with smaller males weighing as little as approximately 300 g (11 oz).

Falco peregrinus nesiotes, described by Mayr in 1941, is found in Fiji and probably also Vanuatu and New Caledonia. It is non-migratory.

Falco peregrinus pealei, described by Ridgway in 1873, is Peale's falcon and includes rudolfi. It is found in the Pacific Northwest of North America, northwards from Puget Sound along the British Columbia coast (including the Haida Gwaii), along the Gulf of Alaska and the Aleutian Islands to the far eastern Bering Sea coast of Russia, and may also occur on the Kuril Islands and the coasts of Kamchatka. It is non-migratory. It is the largest subspecies and it looks like an oversized and darker tundrius or like a strongly barred and large anatum. The bill is very wide. Juveniles occasionally have pale crowns. Males weigh 700 to 1,000 g (1.5–2.2 lb), while females weigh 1,000 to 1,500 g (2.2–3.3 lb).

Falco peregrinus pelegrinoides, first described by Temminck in 1829, is found in the Canary Islands through North Africa and the Near East to Mesopotamia. It is most similar to brookei, but is markedly paler above, with a rusty neck, and is a light buff with reduced barring below. It is smaller than the nominate subspecies; females weigh around 610 g (1.34 lb).

Falco peregrinus peregrinator, described by Sundevall in 1837, is known as the Indian peregrine falcon, black shaheen, Indian shaheen or shaheen falcon. It was formerly sometimes known as Falco atriceps or Falco shaheen. Its range includes South Asia from across the Indian subcontinent to Sri Lanka and southeastern China. In India, the shaheen falcon is reported from all states except Uttar Pradesh, mainly from rocky and hilly regions. The shaheen falcon is also reported from the Andaman and Nicobar Islands in the Bay of Bengal. It has a clutch size of 3 to 4 eggs, with the chicks fledging time of 48 days with an average nesting success of 1.32 chicks per nest. In India, apart from nesting on cliffs, it has also been recorded as nesting on man-made structures such as buildings and cellphone transmission towers.[36] A population estimate of 40 breeding pairs in Sri Lanka was made in 1996. It is non-migratory and is small and dark, with rufous underparts. In Sri Lanka this species is found to favour the higher hills, while the migrant calidus is more often seen along the coast.

Falco peregrinus peregrinus, the nominate (first-named) subspecies, described by Tunstall in 1771, breeds over much of temperate Eurasia between the tundra in the north and the Pyrenees, Mediterranean region and Alpide belt in the south. It is mainly non-migratory in Europe, but migratory in Scandinavia and Asia. Males weigh 580 to 750 g (1.28–1.65 lb), while females weigh 925 to 1,300 g (2.039–2.866 lb). It includes brevirostris, germanicus, rhenanus and riphaeus.

Falco peregrinus radama, described by Hartlaub in 1861, is found in Madagascar and the Comoros. It is non-migratory.

Falco peregrinus submelanogenys, described by Mathews in 1912, is the Southwest Australian peregrine falcon. It is found in southwestern Australia and is non-migratory.

Falco peregrinus tundrius, described by C.M. White in 1968, was at one time included in leucogenys. It is found in the Arctic tundra of North America to Greenland, and migrates to wintering grounds in Central and South America. Most vagrants that reach western Europe belong to this subspecies, which was previously considered synonymous with anatum. It is the New World equivalent to calidus. It is smaller and paler than anatum; most have a conspicuous white forehead and white in ear region, but the crown and "moustache" are very dark, unlike in calidus. Juveniles are browner and less grey than in calidus and paler, sometimes almost sandy, than in anatum. Males weigh 500 to 700 g (1.1–1.5 lb), while females weigh 800 to 1,100 g (1.8–2.4 lb). Despite its current recognition as a valid subspecies, a population genetic study of both pre-decline (i.e., museum) and recovered contemporary populations failed to distinguish genetically the anatum and tundrius subspecies.

Barbary falcon

Main article: Barbary falcon

The Barbary falcon is a subspecies of the peregrine falcon that inhabits parts of North Africa; namely, from the Canary Islands to the Arabian Peninsula. There is discussion concerning the taxonomic status of the bird, with some considering it a subspecies of the peregrine falcon and others considering it a full species with two subspecies (White et al. 2013). Compared to the other peregrine falcon subspecies, Barbary falcons sport a slimmer body and a distinct plumage color pattern. Despite numbers and range of these birds throughout the Canary Islands generally increasing, they are considered endangered, with human interference through falconry and shooting threatening their well-being. Falconry can further complicate the speciation and genetics of these Canary Islands falcons, as the practice promotes genetic mixing between individuals from outside the islands with those originating from the islands. Population density of the Barbary falcons on Tenerife, the biggest of the seven major Canary Islands, was found to be 1.27 pairs/100 km², with the mean distance between pairs being 5869 ± 3338 m. The falcons were only observed near large and natural cliffs with a mean altitude of 697.6 m. Falcons show an affinity for tall cliffs away from human-mediated establishments and presence.

 

Barbary falcons have a red neck patch, but otherwise differ in appearance from the peregrine falcon proper merely according to Gloger's rule, relating pigmentation to environmental humidity. The Barbary falcon has a peculiar way of flying, beating only the outer part of its wings like fulmars sometimes do; this also occurs in the peregrine falcon, but less often and far less pronounced. The Barbary falcon's shoulder and pelvis bones are stout by comparison with the peregrine falcon and its feet are smaller. Barbary falcons breed at different times of year than neighboring peregrine falcon subspecies, but they are capable of interbreeding. There is a 0.6–0.7% genetic distance in the peregrine falcon-Barbary falcon ("peregrinoid") complex.

 

Ecology and behaviour

The peregrine falcon lives mostly along mountain ranges, river valleys, coastlines, and increasingly in cities. In mild-winter regions, it is usually a permanent resident, and some individuals, especially adult males, will remain on the breeding territory. Only populations that breed in Arctic climates typically migrate great distances during the northern winter.

 

The peregrine falcon reaches faster speeds than any other animal on the planet when performing the stoop, which involves soaring to a great height and then diving steeply at speeds of over 320 km/h (200 mph), hitting one wing of its prey so as not to harm itself on impact. The air pressure from such a dive could possibly damage a bird's lungs, but small bony tubercles on a falcon's nostrils are theorized to guide the powerful airflow away from the nostrils, enabling the bird to breathe more easily while diving by reducing the change in air pressure. To protect their eyes, the falcons use their nictitating membranes (third eyelids) to spread tears and clear debris from their eyes while maintaining vision. The distinctive malar stripe or 'moustache', a dark area of feathers below the eyes, is thought to reduce solar glare and improve contrast sensitivity when targeting fast moving prey in bright light condition; the malar stripe has been found to be wider and more pronounced in regions of the world with greater solar radiation supporting this solar glare hypothesis. Peregrine falcons have a flicker fusion frequency of 129 Hz (cycles per second), very fast for a bird of its size, and much faster than mammals. A study testing the flight physics of an "ideal falcon" found a theoretical speed limit at 400 km/h (250 mph) for low-altitude flight and 625 km/h (388 mph) for high-altitude flight. In 2005, Ken Franklin recorded a falcon stooping at a top speed of 389 km/h (242 mph).

 

The life span of peregrine falcons in the wild is up to 19 years 9 months. Mortality in the first year is 59–70%, declining to 25–32% annually in adults. Apart from such anthropogenic threats as collision with human-made objects, the peregrine may be killed by larger hawks and owls.

 

The peregrine falcon is host to a range of parasites and pathogens. It is a vector for Avipoxvirus, Newcastle disease virus, Falconid herpesvirus 1 (and possibly other Herpesviridae), and some mycoses and bacterial infections. Endoparasites include Plasmodium relictum (usually not causing malaria in the peregrine falcon), Strigeidae trematodes, Serratospiculum amaculata (nematode), and tapeworms. Known peregrine falcon ectoparasites are chewing lice, Ceratophyllus garei (a flea), and Hippoboscidae flies (Icosta nigra, Ornithoctona erythrocephala).

 

In the Arctic Peregrine falcons chasing away small rodent predators from their nesting territory and Rough-legged Buzzards (Buteo lagopus) could use these hot spots as a nesting territory.

 

Feeding

The peregrine falcon's diet varies greatly and is adapted to available prey in different regions. However, it typically feeds on medium-sized birds such as pigeons and doves, waterfowl, gamebirds, songbirds, parrots, seabirds, and waders. Worldwide, it is estimated that between 1,500 and 2,000 bird species, or roughly a fifth of the world's bird species, are predated somewhere by these falcons.The peregrine falcon preys on the most diverse range of bird species of any raptor in North America, with over 300 species and including nearly 100 shorebirds. Its prey can range from 3 g (0.11 oz) hummingbirds (Selasphorus and Archilochus ssp.) to the 3.1 kg (6.8 lb) sandhill crane, although most prey taken by peregrines weigh between 20 g (0.71 oz) (small passerines) and 1,100 g (2.4 lb) (ducks, geese, loons, gulls, capercaillies, ptarmigans and other grouse). Smaller hawks (such as sharp-shinned hawks) and owls are regularly predated, as well as smaller falcons such as the American kestrel, merlin and, rarely, other peregrines.

 

In urban areas, where it tends to nest on tall buildings or bridges, it subsists mostly on a variety of pigeons. Among pigeons, the rock or feral pigeon comprises 80% or more of the dietary intake of peregrines. Other common city birds are also taken regularly, including mourning doves, common wood pigeons, common swifts, northern flickers, common starlings, American robins, common blackbirds, and corvids such as magpies, jays or carrion, house, and American crows. Coastal populations of the large subspecies pealei feed almost exclusively on seabirds. In the Brazilian mangrove swamp of Cubatão, a wintering falcon of the subspecies tundrius was observed successfully hunting a juvenile scarlet ibis.

 

Among mammalian prey species, bats in the genera Eptesicus, Myotis, Pipistrellus and Tadarida are the most common prey which taken at night. Though peregrines generally do not prefer terrestrial mammalian prey, in Rankin Inlet, peregrines largely take northern collared lemmings (Dicrostonyx groenlandicus) along with a few Arctic ground squirrels (Urocitellus parryii). Other small mammals including shrews, mice, rats, voles, and squirrels are more seldom taken. Peregrines occasionally take rabbits, mainly young individuals and juvenile hares. Additionally, remains of red fox kits and adult female American marten were found among prey remains. Insects and reptiles such as small snakes make up a small proportion of the diet, and salmonid fish have been taken by peregrines.

 

The peregrine falcon hunts most often at dawn and dusk, when prey are most active, but also nocturnally in cities, particularly during migration periods when hunting at night may become prevalent. Nocturnal migrants taken by peregrines include species as diverse as yellow-billed cuckoo, black-necked grebe, virginia rail, and common quail. The peregrine requires open space in order to hunt, and therefore often hunts over open water, marshes, valleys, fields, and tundra, searching for prey either from a high perch or from the air. Large congregations of migrants, especially species that gather in the open like shorebirds, can be quite attractive to a hunting peregrine. Once prey is spotted, it begins its stoop, folding back the tail and wings, with feet tucked. Prey is typically struck and captured in mid-air; the peregrine falcon strikes its prey with a clenched foot, stunning or killing it with the impact, then turns to catch it in mid-air. If its prey is too heavy to carry, a peregrine will drop it to the ground and eat it there. If they miss the initial strike, peregrines will chase their prey in a twisting flight.

 

Although previously thought rare, several cases of peregrines contour-hunting, i.e., using natural contours to surprise and ambush prey on the ground, have been reported and even rare cases of prey being pursued on foot. In addition, peregrines have been documented preying on chicks in nests, from birds such as kittiwakes. Prey is plucked before consumption. A 2016 study showed that the presence of peregrines benefits non-preferred species while at the same time causing a decline in its preferred prey. As of 2018, the fastest recorded falcon was at 242 mph (nearly 390 km/h). Researchers at the University of Groningen in the Netherlands and at Oxford University used 3D computer simulations in 2018 to show that the high speed allows peregrines to gain better maneuverability and precision in strikes.

 

Reproduction

The peregrine falcon is sexually mature at one to three years of age, but in larger populations they breed after two to three years of age. A pair mates for life and returns to the same nesting spot annually. The courtship flight includes a mix of aerial acrobatics, precise spirals, and steep dives. The male passes prey it has caught to the female in mid-air. To make this possible, the female actually flies upside-down to receive the food from the male's talons.

 

During the breeding season, the peregrine falcon is territorial; nesting pairs are usually more than 1 km (0.62 mi) apart, and often much farther, even in areas with large numbers of pairs. The distance between nests ensures sufficient food supply for pairs and their chicks. Within a breeding territory, a pair may have several nesting ledges; the number used by a pair can vary from one or two up to seven in a 16-year period.

 

The peregrine falcon nests in a scrape, normally on cliff edges. The female chooses a nest site, where she scrapes a shallow hollow in the loose soil, sand, gravel, or dead vegetation in which to lay eggs. No nest materials are added. Cliff nests are generally located under an overhang, on ledges with vegetation. South-facing sites are favoured. In some regions, as in parts of Australia and on the west coast of northern North America, large tree hollows are used for nesting. Before the demise of most European peregrines, a large population of peregrines in central and western Europe used the disused nests of other large birds. In remote, undisturbed areas such as the Arctic, steep slopes and even low rocks and mounds may be used as nest sites. In many parts of its range, peregrines now also nest regularly on tall buildings or bridges; these human-made structures used for breeding closely resemble the natural cliff ledges that the peregrine prefers for its nesting locations.

 

The pair defends the chosen nest site against other peregrines, and often against ravens, herons, and gulls, and if ground-nesting, also such mammals as foxes, wolverines, felids, bears, wolves, and mountain lions. Both nests and (less frequently) adults are predated by larger-bodied raptorial birds like eagles, large owls, or gyrfalcons. The most serious predators of peregrine nests in North America and Europe are the great horned owl and the Eurasian eagle-owl. When reintroductions have been attempted for peregrines, the most serious impediments were these two species of owls routinely picking off nestlings, fledglings and adults by night. Peregrines defending their nests have managed to kill raptors as large as golden eagles and bald eagles (both of which they normally avoid as potential predators) that have come too close to the nest by ambushing them in a full stoop. In one instance, when a snowy owl killed a newly fledged peregrine, the larger owl was in turn killed by a stooping peregrine parent.

 

The date of egg-laying varies according to locality, but is generally from February to March in the Northern Hemisphere, and from July to August in the Southern Hemisphere, although the Australian subspecies macropus may breed as late as November, and equatorial populations may nest anytime between June and December. If the eggs are lost early in the nesting season, the female usually lays another clutch, although this is extremely rare in the Arctic due to the short summer season. Generally three to four eggs, but sometimes as few as one or as many as five, are laid in the scrape. The eggs are white to buff with red or brown markings. They are incubated for 29 to 33 days, mainly by the female, with the male also helping with the incubation of the eggs during the day, but only the female incubating them at night. The average number of young found in nests is 2.5, and the average number that fledge is about 1.5, due to the occasional production of infertile eggs and various natural losses of nestlings.

 

After hatching, the chicks (called "eyases") are covered with creamy-white down and have disproportionately large feet. The male (called the "tiercel") and the female (simply called the "falcon") both leave the nest to gather prey to feed the young. The hunting territory of the parents can extend a radius of 19 to 24 km (12 to 15 mi) from the nest site. Chicks fledge 42 to 46 days after hatching, and remain dependent on their parents for up to two months.

 

Relationship with humans

The peregrine falcon is a highly admired falconry bird, and has been used in falconry for more than 3,000 years, beginning with nomads in central Asia. Its advantages in falconry include not only its athleticism and eagerness to hunt, but an equable disposition that leads to it being one of the easier falcons to train. The peregrine falcon has the additional advantage of a natural flight style of circling above the falconer ("waiting on") for game to be flushed, and then performing an effective and exciting high-speed diving stoop to take the quarry. The speed of the stoop not only allows the falcon to catch fast flying birds, it also enhances the falcon's ability to execute maneuvers to catch highly agile prey, and allows the falcon to deliver a knockout blow with a fist-like clenched talon against game that may be much larger than itself.

 

Additionally the versatility of the species, with agility allowing capture of smaller birds and a strength and attacking style allowing capture of game much larger than themselves, combined with the wide size range of the many peregrine subspecies, means there is a subspecies suitable to almost any size and type of game bird. This size range, evolved to fit various environments and prey species, is from the larger females of the largest subspecies to the smaller males of the smallest subspecies, approximately five to one (approximately 1500 g to 300 g). The males of smaller and medium-sized subspecies, and the females of the smaller subspecies, excel in the taking of swift and agile small game birds such as dove, quail, and smaller ducks. The females of the larger subspecies are capable of taking large and powerful game birds such as the largest of duck species, pheasant, and grouse.

 

Peregrine falcons handled by falconers are also occasionally used to scare away birds at airports to reduce the risk of bird-plane strikes, improving air-traffic safety. They were also used to intercept homing pigeons during World War II.

 

Peregrine falcons have been successfully bred in captivity, both for falconry and for release into the wild. Until 2004 nearly all peregrines used for falconry in the US were captive-bred from the progeny of falcons taken before the US Endangered Species Act was enacted and from those few infusions of wild genes available from Canada and special circumstances. Peregrine falcons were removed from the United States' endangered species list in 1999. The successful recovery program was aided by the effort and knowledge of falconers – in collaboration with The Peregrine Fund and state and federal agencies – through a technique called hacking. Finally, after years of close work with the US Fish and Wildlife Service, a limited take of wild peregrines was allowed in 2004, the first wild peregrines taken specifically for falconry in over 30 years.

 

The development of captive breeding methods has led to peregrines being commercially available for falconry use, thus mostly eliminating the need to capture wild birds for support of falconry. The main reason for taking wild peregrines at this point is to maintain healthy genetic diversity in the breeding lines. Hybrids of peregrines and gyrfalcons are also available that can combine the best features of both species to create what many consider to be the ultimate falconry bird for the taking of larger game such as the sage-grouse. These hybrids combine the greater size, strength, and horizontal speed of the gyrfalcon with the natural propensity to stoop and greater warm weather tolerance of the peregrine.

 

Decline due to pesticides

The peregrine falcon became an endangered species over much of its range because of the use of organochlorine pesticides, especially DDT, during the 1950s, '60s, and '70s. Pesticide biomagnification caused organochlorine to build up in the falcons' fat tissues, reducing the amount of calcium in their eggshells. With thinner shells, fewer falcon eggs survived until hatching. In addition, the PCB concentrations found in these falcons is dependent upon the age of the falcon. While high levels are still found in young birds (only a few months old) and even higher concentrations are found in more mature falcons, further increasing in adult peregrine falcons. These pesticides caused falcon prey to also have thinner eggshells (one example of prey being the Black Petrels). In several parts of the world, such as the eastern United States and Belgium, this species became extirpated (locally extinct) as a result. An alternate point of view is that populations in the eastern North America had vanished due to hunting and egg collection. Following the ban of organochlorine pesticides, the reproductive success of Peregrines increased in Scotland in terms of territory occupancy and breeding success, although spatial variation in recovery rates indicate that in some areas Peregrines were also impacted by other factors such as persecution.

 

Recovery efforts

Peregrine falcon recovery teams breed the species in captivity. The chicks are usually fed through a chute or with a hand puppet mimicking a peregrine's head, so they cannot see to imprint on the human trainers. Then, when they are old enough, the rearing box is opened, allowing the bird to train its wings. As the fledgling gets stronger, feeding is reduced, forcing the bird to learn to hunt. This procedure is called hacking back to the wild. To release a captive-bred falcon, the bird is placed in a special cage at the top of a tower or cliff ledge for some days or so, allowing it to acclimate itself to its future environment.

 

Worldwide recovery efforts have been remarkably successful. The widespread restriction of DDT use eventually allowed released birds to breed successfully. The peregrine falcon was removed from the U.S. Endangered Species list on 25 August 1999.

 

Some controversy has existed over the origins of captive breeding stock used by the Peregrine Fund in the recovery of peregrine falcons throughout the contiguous United States. Several peregrine subspecies were included in the breeding stock, including birds of Eurasian origin. Due to the extirpation of the eastern population of Falco peregrinus anatum, the near-extirpation of anatum in the Midwest and the limited gene pool within North American breeding stock, the inclusion of non-native subspecies was justified to optimize the genetic diversity found within the species as a whole.

 

During the 1970s, peregrine falcons in Finland experienced a population bottleneck as a result of large declines associated with bio-accumulation of organochloride pesticides. However, the genetic diversity of peregrines in Finland is similar to other populations, indicating that high dispersal rates have maintained the genetic diversity of this species.

 

Since peregrine falcon eggs and chicks are still often targeted by illegal poachers, it is common practice not to publicize unprotected nest locations.

 

Current status

Populations of the peregrine falcon have bounced back in most parts of the world. In the United Kingdom, there has been a recovery of populations since the crash of the 1960s. This has been greatly assisted by conservation and protection work led by the Royal Society for the Protection of Birds. The RSPB estimated that there were 1,402 breeding pairs in the UK in 2011. In Canada, where peregrines were identified as endangered in 1978 (in the Yukon territory of northern Canada that year, only a single breeding pair was identified), the Committee on the Status of Endangered Wildlife in Canada declared the species no longer at risk in December 2017.

 

Peregrines now breed in many mountainous and coastal areas, especially in the west and north, and nest in some urban areas, capitalising on the urban feral pigeon populations for food. Additionally, falcons benefit from artificial illumination, which allows the raptors to extend their hunting periods into the dusk when natural illumination would otherwise be too low for them to pursue prey. In England, this has allowed them to prey on nocturnal migrants such as redwings, fieldfares, starlings, and woodcocks.

 

In many parts of the world peregrine falcons have adapted to urban habitats, nesting on cathedrals, skyscraper window ledges, tower blocks, and the towers of suspension bridges. Many of these nesting birds are encouraged, sometimes gathering media attention and often monitored by cameras.

 

In England, peregrine falcons have become increasingly urban in distribution, particularly in southern areas where inland cliffs suitable as nesting sites are scarce. The first recorded urban breeding pair was observed nesting on the Swansea Guildhall in the 1980s. In Southampton, a nest prevented restoration of mobile telephony services for several months in 2013, after Vodafone engineers despatched to repair a faulty transmitter mast discovered a nest in the mast, and were prevented by the Wildlife and Countryside Act – on pain of a possible prison sentence – from proceeding with repairs until the chicks fledged.

 

In Oregon, Portland houses ten percent of the state's peregrine nests, despite only covering around 0.1 percent of the state's land area.

 

Cultural significance

Due to its striking hunting technique, the peregrine has often been associated with aggression and martial prowess. The Ancient Egyptian solar deity Ra was often represented as a man with the head of a peregrine falcon adorned with the solar disk, although most Egyptologists agree that it's most likely a Lanner falcon. Native Americans of the Mississippian culture (c. 800–1500) used the peregrine, along with several other birds of prey, in imagery as a symbol of "aerial (celestial) power" and buried men of high status in costumes associating to the ferocity of raptorial birds. In the late Middle Ages, the Western European nobility that used peregrines for hunting, considered the bird associated with princes in formal hierarchies of birds of prey, just below the gyrfalcon associated with kings. It was considered "a royal bird, more armed by its courage than its claws". Terminology used by peregrine breeders also used the Old French term gentil, "of noble birth; aristocratic", particularly with the peregrine.

 

The peregrine falcon is the national animal of the United Arab Emirates. Since 1927, the peregrine falcon has been the official mascot of Bowling Green State University in Bowling Green, Ohio. The 2007 U.S. Idaho state quarter features a peregrine falcon. The peregrine falcon has been designated the official city bird of Chicago.

 

The Peregrine, by J. A. Baker, is widely regarded as one of the best nature books in English written in the twentieth century. Admirers of the book include Robert Macfarlane, Mark Cocker, who regards the book as "one of the most outstanding books on nature in the twentieth century" and Werner Herzog, who called it "the one book I would ask you to read if you want to make films", and said elsewhere "it has prose of the calibre that we have not seen since Joseph Conrad". In the book, Baker recounts, in diary form, his detailed observations of peregrines (and their interaction with other birds) near his home in Chelmsford, Essex, over a single winter from October to April.

 

An episode of the hour-long TV series Starman in 1986 titled "Peregrine" was about an injured peregrine falcon and the endangered species program. It was filmed with the assistance of the University of California's peregrine falcon project in Santa Cruz.

The peregrine falcon (Falco peregrinus), also known simply as the peregrine, and historically as the duck hawk in North America, is a cosmopolitan bird of prey (raptor) in the family Falconidae. A large, crow-sized falcon, it has a blue-grey back, barred white underparts, and a black head. The peregrine is renowned for its speed. It can reach over 320 km/h (200 mph) during its characteristic hunting stoop (high-speed dive), making it the fastest member of the animal kingdom. According to a National Geographic TV program, the highest measured speed of a peregrine falcon is 389 km/h (242 mph). As is typical for bird-eating (avivore) raptors, peregrine falcons are sexually dimorphic, with females being considerably larger than males.

 

The peregrine's breeding range includes land regions from the Arctic tundra to the tropics. It can be found nearly everywhere on Earth, except extreme polar regions, very high mountains, and most tropical rainforests; the only major ice-free landmass from which it is entirely absent is New Zealand. This makes it the world's most widespread raptor and one of the most widely found wild bird species. In fact, the only land-based bird species found over a larger geographic area owes its success to human-led introduction; the domestic and feral pigeons are both domesticate forms of the rock dove, which are a major prey species for Eurasian Peregrine populations. Due to their prevalence over most other bird species in cities, feral pigeons support many peregrine populations as a staple food source, especially in urban settings.

 

The peregrine is a highly successful example of urban wildlife in much of its range, taking advantage of tall buildings as nest sites and an abundance of prey such as pigeons and ducks. Both the English and scientific names of this species mean "wandering falcon", referring to the migratory habits of many northern populations. Experts recognize 17 to 19 subspecies, which vary in appearance and range; disagreement exists over whether the distinctive Barbary falcon is represented by two subspecies of Falco peregrinus or is a separate species, F. pelegrinoides. The two species' divergence is relatively recent, during the time of the last ice age, therefore the genetic differential between them (and also the difference in their appearance) is relatively tiny. They are only about 0.6–0.8% genetically differentiated.

 

Although its diet consists almost exclusively of medium-sized birds, the peregrine will sometimes hunt small mammals, small reptiles, or even insects. Reaching sexual maturity at one year, it mates for life and nests in a scrape, normally on cliff edges or, in recent times, on tall human-made structures. The peregrine falcon became an endangered species in many areas because of the widespread use of certain pesticides, especially DDT. Since the ban on DDT from the early 1970s, populations have recovered, supported by large-scale protection of nesting places and releases to the wild.

 

The peregrine falcon is a well-respected falconry bird due to its strong hunting ability, high trainability, versatility, and availability via captive breeding. It is effective on most game bird species, from small to large. It has also been used as a religious, royal, or national symbol across multiple eras and areas of human civilization.

 

Description

 

Falco peregrinus. Royal National Park, New South Wales, Australia

The peregrine falcon has a body length of 34 to 58 cm (13–23 in) and a wingspan from 74 to 120 cm (29–47 in). The male and female have similar markings and plumage but, as with many birds of prey, the peregrine falcon displays marked sexual dimorphism in size, with the female measuring up to 30% larger than the male. Males weigh 330 to 1,000 g (12–35 oz) and the noticeably larger females weigh 700 to 1,500 g (25–53 oz). In most subspecies, males weigh less than 700 g (25 oz) and females weigh more than 800 g (28 oz), and cases of females weighing about 50% more than their male breeding mates are not uncommon. The standard linear measurements of peregrines are: the wing chord measures 26.5 to 39 cm (10.4–15.4 in), the tail measures 13 to 19 cm (5.1–7.5 in) and the tarsus measures 4.5 to 5.6 cm (1.8–2.2 in).

 

The back and the long pointed wings of the adult are usually bluish black to slate grey with indistinct darker barring (see "Subspecies" below); the wingtips are black. The white to rusty underparts are barred with thin clean bands of dark brown or black. The tail, coloured like the back but with thin clean bars, is long, narrow, and rounded at the end with a black tip and a white band at the very end. The top of the head and a "moustache" along the cheeks are black, contrasting sharply with the pale sides of the neck and white throat. The cere is yellow, as are the feet, and the beak and claws are black. The upper beak is notched near the tip, an adaptation which enables falcons to kill prey by severing the spinal column at the neck. An immature bird is much browner, with streaked, rather than barred, underparts, and has a pale bluish cere and orbital ring.

 

A study shows that their black malar stripe exists to reduce glare from solar radiation, allowing them to see better. Photos from The Macaulay Library and iNaturalist showed that the malar stripe is thicker where there is more solar radiation. That supports the solar glare hypothesis.

 

Taxonomy and systematics

 

Falco peregrinus was first described under its current binomial name by English ornithologist Marmaduke Tunstall in his 1771 work Ornithologia Britannica. The scientific name Falco peregrinus is a Medieval Latin phrase that was used by Albertus Magnus in 1225. Peregrinus is Latin, meaning "one from abroad" or "coming from foreign parts". It is likely the name was used as juvenile birds were taken while journeying to their breeding location (rather than from the nest), as falcon nests are often difficult to get at. The Latin term for falcon, falco, is related to falx, meaning "sickle", in reference to the silhouette of the falcon's long, pointed wings in flight.

 

The peregrine falcon belongs to a genus whose lineage includes the hierofalcon and the prairie falcon (F. mexicanus). This lineage probably diverged from other falcons towards the end of the Late Miocene or in the Late Pliocene, about 3–8 million years ago (mya). As the peregrine-hierofalcon group includes both Old World and North American species, it is likely that the lineage originated in western Eurasia or Africa. Its relationship to other falcons is not clear, as the issue is complicated by widespread hybridization confounding mtDNA sequence analyses. One genetic lineage of the saker falcon (F. cherrug) is known to have originated from a male saker ancestor producing fertile young with a female peregrine ancestor, and the descendants further breeding with sakers.

 

Today, peregrines are regularly paired in captivity with other species such as the lanner falcon (F. biarmicus) to produce the "perilanner", a somewhat popular bird in falconry as it combines the peregrine's hunting skill with the lanner's hardiness, or the gyrfalcon to produce large, strikingly coloured birds for the use of falconers. As can be seen, the peregrine is still genetically close to the hierofalcons, though their lineages diverged in the Late Pliocene (maybe some 2.5–2 mya in the Gelasian).

 

Subspecies

Numerous subspecies of Falco peregrinus have been described, with 19 accepted by the 1994 Handbook of the Birds of the World, which considers the Barbary falcon of the Canary Islands and coastal North Africa to be two subspecies (pelegrinoides and babylonicus) of Falco peregrinus, rather than a distinct species, F. pelegrinoides. The following map shows the general ranges of these 19 subspecies.

 

A map of the world, green shows on several continents, but there are also several big bare spots marked with E for extinct

Breeding ranges of the 19 subspecies

 

Falco peregrinus anatum, described by Bonaparte in 1838, is known as the American peregrine falcon or "duck hawk"; its scientific name means "duck peregrine falcon". At one time, it was partly included in leucogenys. It is mainly found in the Rocky Mountains. It was formerly common throughout North America between the tundra and northern Mexico, where current reintroduction efforts are being made to restore the population. Most mature anatum, except those that breed in more northern areas, winter in their breeding range. Most vagrants that reach western Europe seem to belong to the more northern and strongly migratory tundrius, only considered distinct since 1968. It is similar to the nominate subspecies but is slightly smaller; adults are somewhat paler and less patterned below, but juveniles are darker and more patterned below. Males weigh 500 to 700 g (1.1–1.5 lb), while females weigh 800 to 1,100 g (1.8–2.4 lb). It has become extinct in eastern North America and populations there are hybrids as a result of reintroductions of birds from elsewhere.

Falco peregrinus babylonicus, described by P.L. Sclater in 1861, is found in eastern Iran along the Hindu Kush and the Tian Shan to the Mongolian Altai ranges. A few birds winter in northern and northwestern India, mainly in dry semi-desert habitats. It is paler than pelegrinoides and somewhat similar to a small, pale lanner falcon (Falco biarmicus). Males weigh 330 to 400 grams (12 to 14 oz), while females weigh 513 to 765 grams (18.1 to 27.0 oz).

Falco peregrinus brookei, described by Sharpe in 1873, is also known as the Mediterranean peregrine falcon or the Maltese falcon. It includes caucasicus and most specimens of the proposed race punicus, though others may be pelegrinoides (Barbary falcons), or perhaps the rare hybrids between these two which might occur around Algeria. They occur from the Iberian Peninsula around the Mediterranean, except in arid regions, to the Caucasus. They are non-migratory. It is smaller than the nominate subspecies and the underside usually has a rusty hue. Males weigh around 445 g (0.981 lb), while females weigh up to 920 g (2.03 lb).

Falco peregrinus calidus, described by John Latham in 1790, it was formerly called leucogenys and includes caeruleiceps. It breeds in the Arctic tundra of Eurasia from Murmansk Oblast to roughly Yana and Indigirka Rivers, Siberia. It is completely migratory and travels south in winter as far as South Asia and sub-Saharan Africa. It is often seen around wetland habitats. It is paler than the nominate subspecies, especially on the crown. Males weigh 588 to 740 g (1.296–1.631 lb), while females weigh 925 to 1,333 g (2.039–2.939 lb).

Falco peregrinus cassini, described by Sharpe in 1873, is also known as the austral peregrine falcon. It includes kreyenborgi, the pallid falcon, a leucistic colour morph occurring in southernmost South America, which was long believed to be a distinct species. Its range includes South America from Ecuador through Bolivia, northern Argentina and Chile to Tierra del Fuego and the Falkland Islands. It is non-migratory. It is similar to the nominate subspecies, but slightly smaller with a black ear region. The pallid falcon morph kreyenborgi is medium grey above, has little barring below and has a head pattern like the saker falcon (Falco cherrug), but the ear region is white.

Falco peregrinus ernesti, described by Sharpe in 1894, is found from the Sunda Islands to the Philippines and south to eastern New Guinea and the nearby Bismarck Archipelago. Its geographical separation from nesiotes requires confirmation. It is non-migratory. It differs from the nominate subspecies in the very dark, dense barring on its underside and its black ear coverts.

Falco peregrinus furuitii, described by Momiyama in 1927, is found on the Izu and Ogasawara Islands south of Honshū, Japan. It is non-migratory. It is very rare and may only remain on a single island. It is a dark form, resembling pealei in colour, but darker, especially on the tail.

Falco peregrinus japonensis, described by Gmelin in 1788, includes kleinschmidti, pleskei, and harterti, and seems to refer to intergrades with calidus. It is found from northeast Siberia to Kamchatka (though it is possibly replaced by pealei on the coast there) and Japan. Northern populations are migratory, while those of Japan are resident. It is similar to the nominate subspecies, but the young are even darker than those of anatum.

Falco peregrinus macropus, described by Swainson in 1837, is the Australian peregrine falcon. It is found in Australia in all regions except the southwest. It is non-migratory. It is similar to brookei in appearance, but is slightly smaller and the ear region is entirely black. The feet are proportionally large.

Falco peregrinus madens, described by Ripley and Watson in 1963, is unusual in having some sexual dichromatism. If the Barbary falcon (see below) is considered a distinct species, it is sometimes placed therein. It is found in the Cape Verde Islands and is non-migratory; it is also endangered, with only six to eight pairs surviving. Males have a rufous wash on the crown, nape, ears and back; the underside is conspicuously washed pinkish-brown. Females are tinged rich brown overall, especially on the crown and nape.

 

Falco peregrinus minor, first described by Bonaparte in 1850. It was formerly often known as perconfusus. It is sparsely and patchily distributed throughout much of sub-Saharan Africa and widespread in Southern Africa. It apparently reaches north along the Atlantic coast as far as Morocco. It is non-migratory and dark-coloured. This is the smallest subspecies, with smaller males weighing as little as approximately 300 g (11 oz).

Falco peregrinus nesiotes, described by Mayr in 1941, is found in Fiji and probably also Vanuatu and New Caledonia. It is non-migratory.

Falco peregrinus pealei, described by Ridgway in 1873, is Peale's falcon and includes rudolfi. It is found in the Pacific Northwest of North America, northwards from Puget Sound along the British Columbia coast (including the Haida Gwaii), along the Gulf of Alaska and the Aleutian Islands to the far eastern Bering Sea coast of Russia, and may also occur on the Kuril Islands and the coasts of Kamchatka. It is non-migratory. It is the largest subspecies and it looks like an oversized and darker tundrius or like a strongly barred and large anatum. The bill is very wide. Juveniles occasionally have pale crowns. Males weigh 700 to 1,000 g (1.5–2.2 lb), while females weigh 1,000 to 1,500 g (2.2–3.3 lb).

Falco peregrinus pelegrinoides, first described by Temminck in 1829, is found in the Canary Islands through North Africa and the Near East to Mesopotamia. It is most similar to brookei, but is markedly paler above, with a rusty neck, and is a light buff with reduced barring below. It is smaller than the nominate subspecies; females weigh around 610 g (1.34 lb).

Falco peregrinus peregrinator, described by Sundevall in 1837, is known as the Indian peregrine falcon, black shaheen, Indian shaheen or shaheen falcon. It was formerly sometimes known as Falco atriceps or Falco shaheen. Its range includes South Asia from across the Indian subcontinent to Sri Lanka and southeastern China. In India, the shaheen falcon is reported from all states except Uttar Pradesh, mainly from rocky and hilly regions. The shaheen falcon is also reported from the Andaman and Nicobar Islands in the Bay of Bengal. It has a clutch size of 3 to 4 eggs, with the chicks fledging time of 48 days with an average nesting success of 1.32 chicks per nest. In India, apart from nesting on cliffs, it has also been recorded as nesting on man-made structures such as buildings and cellphone transmission towers.[36] A population estimate of 40 breeding pairs in Sri Lanka was made in 1996. It is non-migratory and is small and dark, with rufous underparts. In Sri Lanka this species is found to favour the higher hills, while the migrant calidus is more often seen along the coast.

Falco peregrinus peregrinus, the nominate (first-named) subspecies, described by Tunstall in 1771, breeds over much of temperate Eurasia between the tundra in the north and the Pyrenees, Mediterranean region and Alpide belt in the south. It is mainly non-migratory in Europe, but migratory in Scandinavia and Asia. Males weigh 580 to 750 g (1.28–1.65 lb), while females weigh 925 to 1,300 g (2.039–2.866 lb). It includes brevirostris, germanicus, rhenanus and riphaeus.

Falco peregrinus radama, described by Hartlaub in 1861, is found in Madagascar and the Comoros. It is non-migratory.

Falco peregrinus submelanogenys, described by Mathews in 1912, is the Southwest Australian peregrine falcon. It is found in southwestern Australia and is non-migratory.

Falco peregrinus tundrius, described by C.M. White in 1968, was at one time included in leucogenys. It is found in the Arctic tundra of North America to Greenland, and migrates to wintering grounds in Central and South America. Most vagrants that reach western Europe belong to this subspecies, which was previously considered synonymous with anatum. It is the New World equivalent to calidus. It is smaller and paler than anatum; most have a conspicuous white forehead and white in ear region, but the crown and "moustache" are very dark, unlike in calidus. Juveniles are browner and less grey than in calidus and paler, sometimes almost sandy, than in anatum. Males weigh 500 to 700 g (1.1–1.5 lb), while females weigh 800 to 1,100 g (1.8–2.4 lb). Despite its current recognition as a valid subspecies, a population genetic study of both pre-decline (i.e., museum) and recovered contemporary populations failed to distinguish genetically the anatum and tundrius subspecies.

Barbary falcon

Main article: Barbary falcon

The Barbary falcon is a subspecies of the peregrine falcon that inhabits parts of North Africa; namely, from the Canary Islands to the Arabian Peninsula. There is discussion concerning the taxonomic status of the bird, with some considering it a subspecies of the peregrine falcon and others considering it a full species with two subspecies (White et al. 2013). Compared to the other peregrine falcon subspecies, Barbary falcons sport a slimmer body and a distinct plumage color pattern. Despite numbers and range of these birds throughout the Canary Islands generally increasing, they are considered endangered, with human interference through falconry and shooting threatening their well-being. Falconry can further complicate the speciation and genetics of these Canary Islands falcons, as the practice promotes genetic mixing between individuals from outside the islands with those originating from the islands. Population density of the Barbary falcons on Tenerife, the biggest of the seven major Canary Islands, was found to be 1.27 pairs/100 km², with the mean distance between pairs being 5869 ± 3338 m. The falcons were only observed near large and natural cliffs with a mean altitude of 697.6 m. Falcons show an affinity for tall cliffs away from human-mediated establishments and presence.

 

Barbary falcons have a red neck patch, but otherwise differ in appearance from the peregrine falcon proper merely according to Gloger's rule, relating pigmentation to environmental humidity. The Barbary falcon has a peculiar way of flying, beating only the outer part of its wings like fulmars sometimes do; this also occurs in the peregrine falcon, but less often and far less pronounced. The Barbary falcon's shoulder and pelvis bones are stout by comparison with the peregrine falcon and its feet are smaller. Barbary falcons breed at different times of year than neighboring peregrine falcon subspecies, but they are capable of interbreeding. There is a 0.6–0.7% genetic distance in the peregrine falcon-Barbary falcon ("peregrinoid") complex.

 

Ecology and behaviour

The peregrine falcon lives mostly along mountain ranges, river valleys, coastlines, and increasingly in cities. In mild-winter regions, it is usually a permanent resident, and some individuals, especially adult males, will remain on the breeding territory. Only populations that breed in Arctic climates typically migrate great distances during the northern winter.

 

The peregrine falcon reaches faster speeds than any other animal on the planet when performing the stoop, which involves soaring to a great height and then diving steeply at speeds of over 320 km/h (200 mph), hitting one wing of its prey so as not to harm itself on impact. The air pressure from such a dive could possibly damage a bird's lungs, but small bony tubercles on a falcon's nostrils are theorized to guide the powerful airflow away from the nostrils, enabling the bird to breathe more easily while diving by reducing the change in air pressure. To protect their eyes, the falcons use their nictitating membranes (third eyelids) to spread tears and clear debris from their eyes while maintaining vision. The distinctive malar stripe or 'moustache', a dark area of feathers below the eyes, is thought to reduce solar glare and improve contrast sensitivity when targeting fast moving prey in bright light condition; the malar stripe has been found to be wider and more pronounced in regions of the world with greater solar radiation supporting this solar glare hypothesis. Peregrine falcons have a flicker fusion frequency of 129 Hz (cycles per second), very fast for a bird of its size, and much faster than mammals. A study testing the flight physics of an "ideal falcon" found a theoretical speed limit at 400 km/h (250 mph) for low-altitude flight and 625 km/h (388 mph) for high-altitude flight. In 2005, Ken Franklin recorded a falcon stooping at a top speed of 389 km/h (242 mph).

 

The life span of peregrine falcons in the wild is up to 19 years 9 months. Mortality in the first year is 59–70%, declining to 25–32% annually in adults. Apart from such anthropogenic threats as collision with human-made objects, the peregrine may be killed by larger hawks and owls.

 

The peregrine falcon is host to a range of parasites and pathogens. It is a vector for Avipoxvirus, Newcastle disease virus, Falconid herpesvirus 1 (and possibly other Herpesviridae), and some mycoses and bacterial infections. Endoparasites include Plasmodium relictum (usually not causing malaria in the peregrine falcon), Strigeidae trematodes, Serratospiculum amaculata (nematode), and tapeworms. Known peregrine falcon ectoparasites are chewing lice, Ceratophyllus garei (a flea), and Hippoboscidae flies (Icosta nigra, Ornithoctona erythrocephala).

 

In the Arctic Peregrine falcons chasing away small rodent predators from their nesting territory and Rough-legged Buzzards (Buteo lagopus) could use these hot spots as a nesting territory.

 

Feeding

The peregrine falcon's diet varies greatly and is adapted to available prey in different regions. However, it typically feeds on medium-sized birds such as pigeons and doves, waterfowl, gamebirds, songbirds, parrots, seabirds, and waders. Worldwide, it is estimated that between 1,500 and 2,000 bird species, or roughly a fifth of the world's bird species, are predated somewhere by these falcons.The peregrine falcon preys on the most diverse range of bird species of any raptor in North America, with over 300 species and including nearly 100 shorebirds. Its prey can range from 3 g (0.11 oz) hummingbirds (Selasphorus and Archilochus ssp.) to the 3.1 kg (6.8 lb) sandhill crane, although most prey taken by peregrines weigh between 20 g (0.71 oz) (small passerines) and 1,100 g (2.4 lb) (ducks, geese, loons, gulls, capercaillies, ptarmigans and other grouse). Smaller hawks (such as sharp-shinned hawks) and owls are regularly predated, as well as smaller falcons such as the American kestrel, merlin and, rarely, other peregrines.

 

In urban areas, where it tends to nest on tall buildings or bridges, it subsists mostly on a variety of pigeons. Among pigeons, the rock or feral pigeon comprises 80% or more of the dietary intake of peregrines. Other common city birds are also taken regularly, including mourning doves, common wood pigeons, common swifts, northern flickers, common starlings, American robins, common blackbirds, and corvids such as magpies, jays or carrion, house, and American crows. Coastal populations of the large subspecies pealei feed almost exclusively on seabirds. In the Brazilian mangrove swamp of Cubatão, a wintering falcon of the subspecies tundrius was observed successfully hunting a juvenile scarlet ibis.

 

Among mammalian prey species, bats in the genera Eptesicus, Myotis, Pipistrellus and Tadarida are the most common prey which taken at night. Though peregrines generally do not prefer terrestrial mammalian prey, in Rankin Inlet, peregrines largely take northern collared lemmings (Dicrostonyx groenlandicus) along with a few Arctic ground squirrels (Urocitellus parryii). Other small mammals including shrews, mice, rats, voles, and squirrels are more seldom taken. Peregrines occasionally take rabbits, mainly young individuals and juvenile hares. Additionally, remains of red fox kits and adult female American marten were found among prey remains. Insects and reptiles such as small snakes make up a small proportion of the diet, and salmonid fish have been taken by peregrines.

 

The peregrine falcon hunts most often at dawn and dusk, when prey are most active, but also nocturnally in cities, particularly during migration periods when hunting at night may become prevalent. Nocturnal migrants taken by peregrines include species as diverse as yellow-billed cuckoo, black-necked grebe, virginia rail, and common quail. The peregrine requires open space in order to hunt, and therefore often hunts over open water, marshes, valleys, fields, and tundra, searching for prey either from a high perch or from the air. Large congregations of migrants, especially species that gather in the open like shorebirds, can be quite attractive to a hunting peregrine. Once prey is spotted, it begins its stoop, folding back the tail and wings, with feet tucked. Prey is typically struck and captured in mid-air; the peregrine falcon strikes its prey with a clenched foot, stunning or killing it with the impact, then turns to catch it in mid-air. If its prey is too heavy to carry, a peregrine will drop it to the ground and eat it there. If they miss the initial strike, peregrines will chase their prey in a twisting flight.

 

Although previously thought rare, several cases of peregrines contour-hunting, i.e., using natural contours to surprise and ambush prey on the ground, have been reported and even rare cases of prey being pursued on foot. In addition, peregrines have been documented preying on chicks in nests, from birds such as kittiwakes. Prey is plucked before consumption. A 2016 study showed that the presence of peregrines benefits non-preferred species while at the same time causing a decline in its preferred prey. As of 2018, the fastest recorded falcon was at 242 mph (nearly 390 km/h). Researchers at the University of Groningen in the Netherlands and at Oxford University used 3D computer simulations in 2018 to show that the high speed allows peregrines to gain better maneuverability and precision in strikes.

 

Reproduction

The peregrine falcon is sexually mature at one to three years of age, but in larger populations they breed after two to three years of age. A pair mates for life and returns to the same nesting spot annually. The courtship flight includes a mix of aerial acrobatics, precise spirals, and steep dives. The male passes prey it has caught to the female in mid-air. To make this possible, the female actually flies upside-down to receive the food from the male's talons.

 

During the breeding season, the peregrine falcon is territorial; nesting pairs are usually more than 1 km (0.62 mi) apart, and often much farther, even in areas with large numbers of pairs. The distance between nests ensures sufficient food supply for pairs and their chicks. Within a breeding territory, a pair may have several nesting ledges; the number used by a pair can vary from one or two up to seven in a 16-year period.

 

The peregrine falcon nests in a scrape, normally on cliff edges. The female chooses a nest site, where she scrapes a shallow hollow in the loose soil, sand, gravel, or dead vegetation in which to lay eggs. No nest materials are added. Cliff nests are generally located under an overhang, on ledges with vegetation. South-facing sites are favoured. In some regions, as in parts of Australia and on the west coast of northern North America, large tree hollows are used for nesting. Before the demise of most European peregrines, a large population of peregrines in central and western Europe used the disused nests of other large birds. In remote, undisturbed areas such as the Arctic, steep slopes and even low rocks and mounds may be used as nest sites. In many parts of its range, peregrines now also nest regularly on tall buildings or bridges; these human-made structures used for breeding closely resemble the natural cliff ledges that the peregrine prefers for its nesting locations.

 

The pair defends the chosen nest site against other peregrines, and often against ravens, herons, and gulls, and if ground-nesting, also such mammals as foxes, wolverines, felids, bears, wolves, and mountain lions. Both nests and (less frequently) adults are predated by larger-bodied raptorial birds like eagles, large owls, or gyrfalcons. The most serious predators of peregrine nests in North America and Europe are the great horned owl and the Eurasian eagle-owl. When reintroductions have been attempted for peregrines, the most serious impediments were these two species of owls routinely picking off nestlings, fledglings and adults by night. Peregrines defending their nests have managed to kill raptors as large as golden eagles and bald eagles (both of which they normally avoid as potential predators) that have come too close to the nest by ambushing them in a full stoop. In one instance, when a snowy owl killed a newly fledged peregrine, the larger owl was in turn killed by a stooping peregrine parent.

 

The date of egg-laying varies according to locality, but is generally from February to March in the Northern Hemisphere, and from July to August in the Southern Hemisphere, although the Australian subspecies macropus may breed as late as November, and equatorial populations may nest anytime between June and December. If the eggs are lost early in the nesting season, the female usually lays another clutch, although this is extremely rare in the Arctic due to the short summer season. Generally three to four eggs, but sometimes as few as one or as many as five, are laid in the scrape. The eggs are white to buff with red or brown markings. They are incubated for 29 to 33 days, mainly by the female, with the male also helping with the incubation of the eggs during the day, but only the female incubating them at night. The average number of young found in nests is 2.5, and the average number that fledge is about 1.5, due to the occasional production of infertile eggs and various natural losses of nestlings.

 

After hatching, the chicks (called "eyases") are covered with creamy-white down and have disproportionately large feet. The male (called the "tiercel") and the female (simply called the "falcon") both leave the nest to gather prey to feed the young. The hunting territory of the parents can extend a radius of 19 to 24 km (12 to 15 mi) from the nest site. Chicks fledge 42 to 46 days after hatching, and remain dependent on their parents for up to two months.

 

Relationship with humans

The peregrine falcon is a highly admired falconry bird, and has been used in falconry for more than 3,000 years, beginning with nomads in central Asia. Its advantages in falconry include not only its athleticism and eagerness to hunt, but an equable disposition that leads to it being one of the easier falcons to train. The peregrine falcon has the additional advantage of a natural flight style of circling above the falconer ("waiting on") for game to be flushed, and then performing an effective and exciting high-speed diving stoop to take the quarry. The speed of the stoop not only allows the falcon to catch fast flying birds, it also enhances the falcon's ability to execute maneuvers to catch highly agile prey, and allows the falcon to deliver a knockout blow with a fist-like clenched talon against game that may be much larger than itself.

 

Additionally the versatility of the species, with agility allowing capture of smaller birds and a strength and attacking style allowing capture of game much larger than themselves, combined with the wide size range of the many peregrine subspecies, means there is a subspecies suitable to almost any size and type of game bird. This size range, evolved to fit various environments and prey species, is from the larger females of the largest subspecies to the smaller males of the smallest subspecies, approximately five to one (approximately 1500 g to 300 g). The males of smaller and medium-sized subspecies, and the females of the smaller subspecies, excel in the taking of swift and agile small game birds such as dove, quail, and smaller ducks. The females of the larger subspecies are capable of taking large and powerful game birds such as the largest of duck species, pheasant, and grouse.

 

Peregrine falcons handled by falconers are also occasionally used to scare away birds at airports to reduce the risk of bird-plane strikes, improving air-traffic safety. They were also used to intercept homing pigeons during World War II.

 

Peregrine falcons have been successfully bred in captivity, both for falconry and for release into the wild. Until 2004 nearly all peregrines used for falconry in the US were captive-bred from the progeny of falcons taken before the US Endangered Species Act was enacted and from those few infusions of wild genes available from Canada and special circumstances. Peregrine falcons were removed from the United States' endangered species list in 1999. The successful recovery program was aided by the effort and knowledge of falconers – in collaboration with The Peregrine Fund and state and federal agencies – through a technique called hacking. Finally, after years of close work with the US Fish and Wildlife Service, a limited take of wild peregrines was allowed in 2004, the first wild peregrines taken specifically for falconry in over 30 years.

 

The development of captive breeding methods has led to peregrines being commercially available for falconry use, thus mostly eliminating the need to capture wild birds for support of falconry. The main reason for taking wild peregrines at this point is to maintain healthy genetic diversity in the breeding lines. Hybrids of peregrines and gyrfalcons are also available that can combine the best features of both species to create what many consider to be the ultimate falconry bird for the taking of larger game such as the sage-grouse. These hybrids combine the greater size, strength, and horizontal speed of the gyrfalcon with the natural propensity to stoop and greater warm weather tolerance of the peregrine.

 

Decline due to pesticides

The peregrine falcon became an endangered species over much of its range because of the use of organochlorine pesticides, especially DDT, during the 1950s, '60s, and '70s. Pesticide biomagnification caused organochlorine to build up in the falcons' fat tissues, reducing the amount of calcium in their eggshells. With thinner shells, fewer falcon eggs survived until hatching. In addition, the PCB concentrations found in these falcons is dependent upon the age of the falcon. While high levels are still found in young birds (only a few months old) and even higher concentrations are found in more mature falcons, further increasing in adult peregrine falcons. These pesticides caused falcon prey to also have thinner eggshells (one example of prey being the Black Petrels). In several parts of the world, such as the eastern United States and Belgium, this species became extirpated (locally extinct) as a result. An alternate point of view is that populations in the eastern North America had vanished due to hunting and egg collection. Following the ban of organochlorine pesticides, the reproductive success of Peregrines increased in Scotland in terms of territory occupancy and breeding success, although spatial variation in recovery rates indicate that in some areas Peregrines were also impacted by other factors such as persecution.

 

Recovery efforts

Peregrine falcon recovery teams breed the species in captivity. The chicks are usually fed through a chute or with a hand puppet mimicking a peregrine's head, so they cannot see to imprint on the human trainers. Then, when they are old enough, the rearing box is opened, allowing the bird to train its wings. As the fledgling gets stronger, feeding is reduced, forcing the bird to learn to hunt. This procedure is called hacking back to the wild. To release a captive-bred falcon, the bird is placed in a special cage at the top of a tower or cliff ledge for some days or so, allowing it to acclimate itself to its future environment.

 

Worldwide recovery efforts have been remarkably successful. The widespread restriction of DDT use eventually allowed released birds to breed successfully. The peregrine falcon was removed from the U.S. Endangered Species list on 25 August 1999.

 

Some controversy has existed over the origins of captive breeding stock used by the Peregrine Fund in the recovery of peregrine falcons throughout the contiguous United States. Several peregrine subspecies were included in the breeding stock, including birds of Eurasian origin. Due to the extirpation of the eastern population of Falco peregrinus anatum, the near-extirpation of anatum in the Midwest and the limited gene pool within North American breeding stock, the inclusion of non-native subspecies was justified to optimize the genetic diversity found within the species as a whole.

 

During the 1970s, peregrine falcons in Finland experienced a population bottleneck as a result of large declines associated with bio-accumulation of organochloride pesticides. However, the genetic diversity of peregrines in Finland is similar to other populations, indicating that high dispersal rates have maintained the genetic diversity of this species.

 

Since peregrine falcon eggs and chicks are still often targeted by illegal poachers, it is common practice not to publicize unprotected nest locations.

 

Current status

Populations of the peregrine falcon have bounced back in most parts of the world. In the United Kingdom, there has been a recovery of populations since the crash of the 1960s. This has been greatly assisted by conservation and protection work led by the Royal Society for the Protection of Birds. The RSPB estimated that there were 1,402 breeding pairs in the UK in 2011. In Canada, where peregrines were identified as endangered in 1978 (in the Yukon territory of northern Canada that year, only a single breeding pair was identified), the Committee on the Status of Endangered Wildlife in Canada declared the species no longer at risk in December 2017.

 

Peregrines now breed in many mountainous and coastal areas, especially in the west and north, and nest in some urban areas, capitalising on the urban feral pigeon populations for food. Additionally, falcons benefit from artificial illumination, which allows the raptors to extend their hunting periods into the dusk when natural illumination would otherwise be too low for them to pursue prey. In England, this has allowed them to prey on nocturnal migrants such as redwings, fieldfares, starlings, and woodcocks.

 

In many parts of the world peregrine falcons have adapted to urban habitats, nesting on cathedrals, skyscraper window ledges, tower blocks, and the towers of suspension bridges. Many of these nesting birds are encouraged, sometimes gathering media attention and often monitored by cameras.

 

In England, peregrine falcons have become increasingly urban in distribution, particularly in southern areas where inland cliffs suitable as nesting sites are scarce. The first recorded urban breeding pair was observed nesting on the Swansea Guildhall in the 1980s. In Southampton, a nest prevented restoration of mobile telephony services for several months in 2013, after Vodafone engineers despatched to repair a faulty transmitter mast discovered a nest in the mast, and were prevented by the Wildlife and Countryside Act – on pain of a possible prison sentence – from proceeding with repairs until the chicks fledged.

 

In Oregon, Portland houses ten percent of the state's peregrine nests, despite only covering around 0.1 percent of the state's land area.

 

Cultural significance

Due to its striking hunting technique, the peregrine has often been associated with aggression and martial prowess. The Ancient Egyptian solar deity Ra was often represented as a man with the head of a peregrine falcon adorned with the solar disk, although most Egyptologists agree that it's most likely a Lanner falcon. Native Americans of the Mississippian culture (c. 800–1500) used the peregrine, along with several other birds of prey, in imagery as a symbol of "aerial (celestial) power" and buried men of high status in costumes associating to the ferocity of raptorial birds. In the late Middle Ages, the Western European nobility that used peregrines for hunting, considered the bird associated with princes in formal hierarchies of birds of prey, just below the gyrfalcon associated with kings. It was considered "a royal bird, more armed by its courage than its claws". Terminology used by peregrine breeders also used the Old French term gentil, "of noble birth; aristocratic", particularly with the peregrine.

 

The peregrine falcon is the national animal of the United Arab Emirates. Since 1927, the peregrine falcon has been the official mascot of Bowling Green State University in Bowling Green, Ohio. The 2007 U.S. Idaho state quarter features a peregrine falcon. The peregrine falcon has been designated the official city bird of Chicago.

 

The Peregrine, by J. A. Baker, is widely regarded as one of the best nature books in English written in the twentieth century. Admirers of the book include Robert Macfarlane, Mark Cocker, who regards the book as "one of the most outstanding books on nature in the twentieth century" and Werner Herzog, who called it "the one book I would ask you to read if you want to make films", and said elsewhere "it has prose of the calibre that we have not seen since Joseph Conrad". In the book, Baker recounts, in diary form, his detailed observations of peregrines (and their interaction with other birds) near his home in Chelmsford, Essex, over a single winter from October to April.

 

An episode of the hour-long TV series Starman in 1986 titled "Peregrine" was about an injured peregrine falcon and the endangered species program. It was filmed with the assistance of the University of California's peregrine falcon project in Santa Cruz.

I dug into my archives and found this nearly forgotten image of two cellphones from different eras: an analog NEC Primero BX brick from the mid 1990s and an old digital HTC Incredible S - my very first smartphone from 2011.

 

The NEC brick phone belongs to my barber and is seen with an aftermarket bulbous ended antenna that was more aesthetic than it was functional. Its original antenna was actually a short, "rubber duck" type that you see on UHF walkie talkies.

 

NEC made two variants of the Primero BX: a 900 MHz ETACS unit and an 800 MHz AMPS model for two competing telecommunications companies, which used differing analog cellular network systems - the Celcom ART 900 (900 MHz) and the Mobifon 800 (800 MHz) services, which have long been defunct.

 

Between the two rivals, only the Celcom company remains in operation to this day and has been providing the digital GSM standard, wireless telephony service since 1995.

 

Mobikom, faced with a losing battle against the more established Celcom service and new, digital GSM/PCN upstarts like Maxis and DiGi cellular companies, found itself trapped with its AMPS network infrastructure and a dwindling number of subscribers. Although the company's AMPS network was ready to use the digital version (D-AMPS) the cost of dual mode (analog/digital) cellphones were too prohibitive for its consumers. Although Mobifon 800 started off in 1994 with a sizeable customer base, it ceased operations by 1996 and faded into the oblivion.

 

The two Primero BX ETACS and AMPS models were nearly indistinguishable from one another and they cannot be interchanged. In those days, if you switched services you had to sell off your existing cellphone and change to another that worked with the other service.

The peregrine falcon (Falco peregrinus), also known simply as the peregrine, and historically as the duck hawk in North America, is a cosmopolitan bird of prey (raptor) in the family Falconidae. A large, crow-sized falcon, it has a blue-grey back, barred white underparts, and a black head. The peregrine is renowned for its speed. It can reach over 320 km/h (200 mph) during its characteristic hunting stoop (high-speed dive), making it the fastest member of the animal kingdom. According to a National Geographic TV program, the highest measured speed of a peregrine falcon is 389 km/h (242 mph). As is typical for bird-eating (avivore) raptors, peregrine falcons are sexually dimorphic, with females being considerably larger than males.

 

The peregrine's breeding range includes land regions from the Arctic tundra to the tropics. It can be found nearly everywhere on Earth, except extreme polar regions, very high mountains, and most tropical rainforests; the only major ice-free landmass from which it is entirely absent is New Zealand. This makes it the world's most widespread raptor and one of the most widely found wild bird species. In fact, the only land-based bird species found over a larger geographic area owes its success to human-led introduction; the domestic and feral pigeons are both domesticate forms of the rock dove, which are a major prey species for Eurasian Peregrine populations. Due to their prevalence over most other bird species in cities, feral pigeons support many peregrine populations as a staple food source, especially in urban settings.

 

The peregrine is a highly successful example of urban wildlife in much of its range, taking advantage of tall buildings as nest sites and an abundance of prey such as pigeons and ducks. Both the English and scientific names of this species mean "wandering falcon", referring to the migratory habits of many northern populations. Experts recognize 17 to 19 subspecies, which vary in appearance and range; disagreement exists over whether the distinctive Barbary falcon is represented by two subspecies of Falco peregrinus or is a separate species, F. pelegrinoides. The two species' divergence is relatively recent, during the time of the last ice age, therefore the genetic differential between them (and also the difference in their appearance) is relatively tiny. They are only about 0.6–0.8% genetically differentiated.

 

Although its diet consists almost exclusively of medium-sized birds, the peregrine will sometimes hunt small mammals, small reptiles, or even insects. Reaching sexual maturity at one year, it mates for life and nests in a scrape, normally on cliff edges or, in recent times, on tall human-made structures. The peregrine falcon became an endangered species in many areas because of the widespread use of certain pesticides, especially DDT. Since the ban on DDT from the early 1970s, populations have recovered, supported by large-scale protection of nesting places and releases to the wild.

 

The peregrine falcon is a well-respected falconry bird due to its strong hunting ability, high trainability, versatility, and availability via captive breeding. It is effective on most game bird species, from small to large. It has also been used as a religious, royal, or national symbol across multiple eras and areas of human civilization.

 

Description

 

Falco peregrinus. Royal National Park, New South Wales, Australia

The peregrine falcon has a body length of 34 to 58 cm (13–23 in) and a wingspan from 74 to 120 cm (29–47 in). The male and female have similar markings and plumage but, as with many birds of prey, the peregrine falcon displays marked sexual dimorphism in size, with the female measuring up to 30% larger than the male. Males weigh 330 to 1,000 g (12–35 oz) and the noticeably larger females weigh 700 to 1,500 g (25–53 oz). In most subspecies, males weigh less than 700 g (25 oz) and females weigh more than 800 g (28 oz), and cases of females weighing about 50% more than their male breeding mates are not uncommon. The standard linear measurements of peregrines are: the wing chord measures 26.5 to 39 cm (10.4–15.4 in), the tail measures 13 to 19 cm (5.1–7.5 in) and the tarsus measures 4.5 to 5.6 cm (1.8–2.2 in).

 

The back and the long pointed wings of the adult are usually bluish black to slate grey with indistinct darker barring (see "Subspecies" below); the wingtips are black. The white to rusty underparts are barred with thin clean bands of dark brown or black. The tail, coloured like the back but with thin clean bars, is long, narrow, and rounded at the end with a black tip and a white band at the very end. The top of the head and a "moustache" along the cheeks are black, contrasting sharply with the pale sides of the neck and white throat. The cere is yellow, as are the feet, and the beak and claws are black. The upper beak is notched near the tip, an adaptation which enables falcons to kill prey by severing the spinal column at the neck. An immature bird is much browner, with streaked, rather than barred, underparts, and has a pale bluish cere and orbital ring.

 

A study shows that their black malar stripe exists to reduce glare from solar radiation, allowing them to see better. Photos from The Macaulay Library and iNaturalist showed that the malar stripe is thicker where there is more solar radiation. That supports the solar glare hypothesis.

 

Taxonomy and systematics

 

Falco peregrinus was first described under its current binomial name by English ornithologist Marmaduke Tunstall in his 1771 work Ornithologia Britannica. The scientific name Falco peregrinus is a Medieval Latin phrase that was used by Albertus Magnus in 1225. Peregrinus is Latin, meaning "one from abroad" or "coming from foreign parts". It is likely the name was used as juvenile birds were taken while journeying to their breeding location (rather than from the nest), as falcon nests are often difficult to get at. The Latin term for falcon, falco, is related to falx, meaning "sickle", in reference to the silhouette of the falcon's long, pointed wings in flight.

 

The peregrine falcon belongs to a genus whose lineage includes the hierofalcon and the prairie falcon (F. mexicanus). This lineage probably diverged from other falcons towards the end of the Late Miocene or in the Late Pliocene, about 3–8 million years ago (mya). As the peregrine-hierofalcon group includes both Old World and North American species, it is likely that the lineage originated in western Eurasia or Africa. Its relationship to other falcons is not clear, as the issue is complicated by widespread hybridization confounding mtDNA sequence analyses. One genetic lineage of the saker falcon (F. cherrug) is known to have originated from a male saker ancestor producing fertile young with a female peregrine ancestor, and the descendants further breeding with sakers.

 

Today, peregrines are regularly paired in captivity with other species such as the lanner falcon (F. biarmicus) to produce the "perilanner", a somewhat popular bird in falconry as it combines the peregrine's hunting skill with the lanner's hardiness, or the gyrfalcon to produce large, strikingly coloured birds for the use of falconers. As can be seen, the peregrine is still genetically close to the hierofalcons, though their lineages diverged in the Late Pliocene (maybe some 2.5–2 mya in the Gelasian).

 

Subspecies

Numerous subspecies of Falco peregrinus have been described, with 19 accepted by the 1994 Handbook of the Birds of the World, which considers the Barbary falcon of the Canary Islands and coastal North Africa to be two subspecies (pelegrinoides and babylonicus) of Falco peregrinus, rather than a distinct species, F. pelegrinoides. The following map shows the general ranges of these 19 subspecies.

 

A map of the world, green shows on several continents, but there are also several big bare spots marked with E for extinct

Breeding ranges of the 19 subspecies

 

Falco peregrinus anatum, described by Bonaparte in 1838, is known as the American peregrine falcon or "duck hawk"; its scientific name means "duck peregrine falcon". At one time, it was partly included in leucogenys. It is mainly found in the Rocky Mountains. It was formerly common throughout North America between the tundra and northern Mexico, where current reintroduction efforts are being made to restore the population. Most mature anatum, except those that breed in more northern areas, winter in their breeding range. Most vagrants that reach western Europe seem to belong to the more northern and strongly migratory tundrius, only considered distinct since 1968. It is similar to the nominate subspecies but is slightly smaller; adults are somewhat paler and less patterned below, but juveniles are darker and more patterned below. Males weigh 500 to 700 g (1.1–1.5 lb), while females weigh 800 to 1,100 g (1.8–2.4 lb). It has become extinct in eastern North America and populations there are hybrids as a result of reintroductions of birds from elsewhere.

Falco peregrinus babylonicus, described by P.L. Sclater in 1861, is found in eastern Iran along the Hindu Kush and the Tian Shan to the Mongolian Altai ranges. A few birds winter in northern and northwestern India, mainly in dry semi-desert habitats. It is paler than pelegrinoides and somewhat similar to a small, pale lanner falcon (Falco biarmicus). Males weigh 330 to 400 grams (12 to 14 oz), while females weigh 513 to 765 grams (18.1 to 27.0 oz).

Falco peregrinus brookei, described by Sharpe in 1873, is also known as the Mediterranean peregrine falcon or the Maltese falcon. It includes caucasicus and most specimens of the proposed race punicus, though others may be pelegrinoides (Barbary falcons), or perhaps the rare hybrids between these two which might occur around Algeria. They occur from the Iberian Peninsula around the Mediterranean, except in arid regions, to the Caucasus. They are non-migratory. It is smaller than the nominate subspecies and the underside usually has a rusty hue. Males weigh around 445 g (0.981 lb), while females weigh up to 920 g (2.03 lb).

Falco peregrinus calidus, described by John Latham in 1790, it was formerly called leucogenys and includes caeruleiceps. It breeds in the Arctic tundra of Eurasia from Murmansk Oblast to roughly Yana and Indigirka Rivers, Siberia. It is completely migratory and travels south in winter as far as South Asia and sub-Saharan Africa. It is often seen around wetland habitats. It is paler than the nominate subspecies, especially on the crown. Males weigh 588 to 740 g (1.296–1.631 lb), while females weigh 925 to 1,333 g (2.039–2.939 lb).

Falco peregrinus cassini, described by Sharpe in 1873, is also known as the austral peregrine falcon. It includes kreyenborgi, the pallid falcon, a leucistic colour morph occurring in southernmost South America, which was long believed to be a distinct species. Its range includes South America from Ecuador through Bolivia, northern Argentina and Chile to Tierra del Fuego and the Falkland Islands. It is non-migratory. It is similar to the nominate subspecies, but slightly smaller with a black ear region. The pallid falcon morph kreyenborgi is medium grey above, has little barring below and has a head pattern like the saker falcon (Falco cherrug), but the ear region is white.

Falco peregrinus ernesti, described by Sharpe in 1894, is found from the Sunda Islands to the Philippines and south to eastern New Guinea and the nearby Bismarck Archipelago. Its geographical separation from nesiotes requires confirmation. It is non-migratory. It differs from the nominate subspecies in the very dark, dense barring on its underside and its black ear coverts.

Falco peregrinus furuitii, described by Momiyama in 1927, is found on the Izu and Ogasawara Islands south of Honshū, Japan. It is non-migratory. It is very rare and may only remain on a single island. It is a dark form, resembling pealei in colour, but darker, especially on the tail.

Falco peregrinus japonensis, described by Gmelin in 1788, includes kleinschmidti, pleskei, and harterti, and seems to refer to intergrades with calidus. It is found from northeast Siberia to Kamchatka (though it is possibly replaced by pealei on the coast there) and Japan. Northern populations are migratory, while those of Japan are resident. It is similar to the nominate subspecies, but the young are even darker than those of anatum.

Falco peregrinus macropus, described by Swainson in 1837, is the Australian peregrine falcon. It is found in Australia in all regions except the southwest. It is non-migratory. It is similar to brookei in appearance, but is slightly smaller and the ear region is entirely black. The feet are proportionally large.

Falco peregrinus madens, described by Ripley and Watson in 1963, is unusual in having some sexual dichromatism. If the Barbary falcon (see below) is considered a distinct species, it is sometimes placed therein. It is found in the Cape Verde Islands and is non-migratory; it is also endangered, with only six to eight pairs surviving. Males have a rufous wash on the crown, nape, ears and back; the underside is conspicuously washed pinkish-brown. Females are tinged rich brown overall, especially on the crown and nape.

 

Falco peregrinus minor, first described by Bonaparte in 1850. It was formerly often known as perconfusus. It is sparsely and patchily distributed throughout much of sub-Saharan Africa and widespread in Southern Africa. It apparently reaches north along the Atlantic coast as far as Morocco. It is non-migratory and dark-coloured. This is the smallest subspecies, with smaller males weighing as little as approximately 300 g (11 oz).

Falco peregrinus nesiotes, described by Mayr in 1941, is found in Fiji and probably also Vanuatu and New Caledonia. It is non-migratory.

Falco peregrinus pealei, described by Ridgway in 1873, is Peale's falcon and includes rudolfi. It is found in the Pacific Northwest of North America, northwards from Puget Sound along the British Columbia coast (including the Haida Gwaii), along the Gulf of Alaska and the Aleutian Islands to the far eastern Bering Sea coast of Russia, and may also occur on the Kuril Islands and the coasts of Kamchatka. It is non-migratory. It is the largest subspecies and it looks like an oversized and darker tundrius or like a strongly barred and large anatum. The bill is very wide. Juveniles occasionally have pale crowns. Males weigh 700 to 1,000 g (1.5–2.2 lb), while females weigh 1,000 to 1,500 g (2.2–3.3 lb).

Falco peregrinus pelegrinoides, first described by Temminck in 1829, is found in the Canary Islands through North Africa and the Near East to Mesopotamia. It is most similar to brookei, but is markedly paler above, with a rusty neck, and is a light buff with reduced barring below. It is smaller than the nominate subspecies; females weigh around 610 g (1.34 lb).

Falco peregrinus peregrinator, described by Sundevall in 1837, is known as the Indian peregrine falcon, black shaheen, Indian shaheen or shaheen falcon. It was formerly sometimes known as Falco atriceps or Falco shaheen. Its range includes South Asia from across the Indian subcontinent to Sri Lanka and southeastern China. In India, the shaheen falcon is reported from all states except Uttar Pradesh, mainly from rocky and hilly regions. The shaheen falcon is also reported from the Andaman and Nicobar Islands in the Bay of Bengal. It has a clutch size of 3 to 4 eggs, with the chicks fledging time of 48 days with an average nesting success of 1.32 chicks per nest. In India, apart from nesting on cliffs, it has also been recorded as nesting on man-made structures such as buildings and cellphone transmission towers.[36] A population estimate of 40 breeding pairs in Sri Lanka was made in 1996. It is non-migratory and is small and dark, with rufous underparts. In Sri Lanka this species is found to favour the higher hills, while the migrant calidus is more often seen along the coast.

Falco peregrinus peregrinus, the nominate (first-named) subspecies, described by Tunstall in 1771, breeds over much of temperate Eurasia between the tundra in the north and the Pyrenees, Mediterranean region and Alpide belt in the south. It is mainly non-migratory in Europe, but migratory in Scandinavia and Asia. Males weigh 580 to 750 g (1.28–1.65 lb), while females weigh 925 to 1,300 g (2.039–2.866 lb). It includes brevirostris, germanicus, rhenanus and riphaeus.

Falco peregrinus radama, described by Hartlaub in 1861, is found in Madagascar and the Comoros. It is non-migratory.

Falco peregrinus submelanogenys, described by Mathews in 1912, is the Southwest Australian peregrine falcon. It is found in southwestern Australia and is non-migratory.

Falco peregrinus tundrius, described by C.M. White in 1968, was at one time included in leucogenys. It is found in the Arctic tundra of North America to Greenland, and migrates to wintering grounds in Central and South America. Most vagrants that reach western Europe belong to this subspecies, which was previously considered synonymous with anatum. It is the New World equivalent to calidus. It is smaller and paler than anatum; most have a conspicuous white forehead and white in ear region, but the crown and "moustache" are very dark, unlike in calidus. Juveniles are browner and less grey than in calidus and paler, sometimes almost sandy, than in anatum. Males weigh 500 to 700 g (1.1–1.5 lb), while females weigh 800 to 1,100 g (1.8–2.4 lb). Despite its current recognition as a valid subspecies, a population genetic study of both pre-decline (i.e., museum) and recovered contemporary populations failed to distinguish genetically the anatum and tundrius subspecies.

Barbary falcon

Main article: Barbary falcon

The Barbary falcon is a subspecies of the peregrine falcon that inhabits parts of North Africa; namely, from the Canary Islands to the Arabian Peninsula. There is discussion concerning the taxonomic status of the bird, with some considering it a subspecies of the peregrine falcon and others considering it a full species with two subspecies (White et al. 2013). Compared to the other peregrine falcon subspecies, Barbary falcons sport a slimmer body and a distinct plumage color pattern. Despite numbers and range of these birds throughout the Canary Islands generally increasing, they are considered endangered, with human interference through falconry and shooting threatening their well-being. Falconry can further complicate the speciation and genetics of these Canary Islands falcons, as the practice promotes genetic mixing between individuals from outside the islands with those originating from the islands. Population density of the Barbary falcons on Tenerife, the biggest of the seven major Canary Islands, was found to be 1.27 pairs/100 km², with the mean distance between pairs being 5869 ± 3338 m. The falcons were only observed near large and natural cliffs with a mean altitude of 697.6 m. Falcons show an affinity for tall cliffs away from human-mediated establishments and presence.

 

Barbary falcons have a red neck patch, but otherwise differ in appearance from the peregrine falcon proper merely according to Gloger's rule, relating pigmentation to environmental humidity. The Barbary falcon has a peculiar way of flying, beating only the outer part of its wings like fulmars sometimes do; this also occurs in the peregrine falcon, but less often and far less pronounced. The Barbary falcon's shoulder and pelvis bones are stout by comparison with the peregrine falcon and its feet are smaller. Barbary falcons breed at different times of year than neighboring peregrine falcon subspecies, but they are capable of interbreeding. There is a 0.6–0.7% genetic distance in the peregrine falcon-Barbary falcon ("peregrinoid") complex.

 

Ecology and behaviour

The peregrine falcon lives mostly along mountain ranges, river valleys, coastlines, and increasingly in cities. In mild-winter regions, it is usually a permanent resident, and some individuals, especially adult males, will remain on the breeding territory. Only populations that breed in Arctic climates typically migrate great distances during the northern winter.

 

The peregrine falcon reaches faster speeds than any other animal on the planet when performing the stoop, which involves soaring to a great height and then diving steeply at speeds of over 320 km/h (200 mph), hitting one wing of its prey so as not to harm itself on impact. The air pressure from such a dive could possibly damage a bird's lungs, but small bony tubercles on a falcon's nostrils are theorized to guide the powerful airflow away from the nostrils, enabling the bird to breathe more easily while diving by reducing the change in air pressure. To protect their eyes, the falcons use their nictitating membranes (third eyelids) to spread tears and clear debris from their eyes while maintaining vision. The distinctive malar stripe or 'moustache', a dark area of feathers below the eyes, is thought to reduce solar glare and improve contrast sensitivity when targeting fast moving prey in bright light condition; the malar stripe has been found to be wider and more pronounced in regions of the world with greater solar radiation supporting this solar glare hypothesis. Peregrine falcons have a flicker fusion frequency of 129 Hz (cycles per second), very fast for a bird of its size, and much faster than mammals. A study testing the flight physics of an "ideal falcon" found a theoretical speed limit at 400 km/h (250 mph) for low-altitude flight and 625 km/h (388 mph) for high-altitude flight. In 2005, Ken Franklin recorded a falcon stooping at a top speed of 389 km/h (242 mph).

 

The life span of peregrine falcons in the wild is up to 19 years 9 months. Mortality in the first year is 59–70%, declining to 25–32% annually in adults. Apart from such anthropogenic threats as collision with human-made objects, the peregrine may be killed by larger hawks and owls.

 

The peregrine falcon is host to a range of parasites and pathogens. It is a vector for Avipoxvirus, Newcastle disease virus, Falconid herpesvirus 1 (and possibly other Herpesviridae), and some mycoses and bacterial infections. Endoparasites include Plasmodium relictum (usually not causing malaria in the peregrine falcon), Strigeidae trematodes, Serratospiculum amaculata (nematode), and tapeworms. Known peregrine falcon ectoparasites are chewing lice, Ceratophyllus garei (a flea), and Hippoboscidae flies (Icosta nigra, Ornithoctona erythrocephala).

 

In the Arctic Peregrine falcons chasing away small rodent predators from their nesting territory and Rough-legged Buzzards (Buteo lagopus) could use these hot spots as a nesting territory.

 

Feeding

The peregrine falcon's diet varies greatly and is adapted to available prey in different regions. However, it typically feeds on medium-sized birds such as pigeons and doves, waterfowl, gamebirds, songbirds, parrots, seabirds, and waders. Worldwide, it is estimated that between 1,500 and 2,000 bird species, or roughly a fifth of the world's bird species, are predated somewhere by these falcons.The peregrine falcon preys on the most diverse range of bird species of any raptor in North America, with over 300 species and including nearly 100 shorebirds. Its prey can range from 3 g (0.11 oz) hummingbirds (Selasphorus and Archilochus ssp.) to the 3.1 kg (6.8 lb) sandhill crane, although most prey taken by peregrines weigh between 20 g (0.71 oz) (small passerines) and 1,100 g (2.4 lb) (ducks, geese, loons, gulls, capercaillies, ptarmigans and other grouse). Smaller hawks (such as sharp-shinned hawks) and owls are regularly predated, as well as smaller falcons such as the American kestrel, merlin and, rarely, other peregrines.

 

In urban areas, where it tends to nest on tall buildings or bridges, it subsists mostly on a variety of pigeons. Among pigeons, the rock or feral pigeon comprises 80% or more of the dietary intake of peregrines. Other common city birds are also taken regularly, including mourning doves, common wood pigeons, common swifts, northern flickers, common starlings, American robins, common blackbirds, and corvids such as magpies, jays or carrion, house, and American crows. Coastal populations of the large subspecies pealei feed almost exclusively on seabirds. In the Brazilian mangrove swamp of Cubatão, a wintering falcon of the subspecies tundrius was observed successfully hunting a juvenile scarlet ibis.

 

Among mammalian prey species, bats in the genera Eptesicus, Myotis, Pipistrellus and Tadarida are the most common prey which taken at night. Though peregrines generally do not prefer terrestrial mammalian prey, in Rankin Inlet, peregrines largely take northern collared lemmings (Dicrostonyx groenlandicus) along with a few Arctic ground squirrels (Urocitellus parryii). Other small mammals including shrews, mice, rats, voles, and squirrels are more seldom taken. Peregrines occasionally take rabbits, mainly young individuals and juvenile hares. Additionally, remains of red fox kits and adult female American marten were found among prey remains. Insects and reptiles such as small snakes make up a small proportion of the diet, and salmonid fish have been taken by peregrines.

 

The peregrine falcon hunts most often at dawn and dusk, when prey are most active, but also nocturnally in cities, particularly during migration periods when hunting at night may become prevalent. Nocturnal migrants taken by peregrines include species as diverse as yellow-billed cuckoo, black-necked grebe, virginia rail, and common quail. The peregrine requires open space in order to hunt, and therefore often hunts over open water, marshes, valleys, fields, and tundra, searching for prey either from a high perch or from the air. Large congregations of migrants, especially species that gather in the open like shorebirds, can be quite attractive to a hunting peregrine. Once prey is spotted, it begins its stoop, folding back the tail and wings, with feet tucked. Prey is typically struck and captured in mid-air; the peregrine falcon strikes its prey with a clenched foot, stunning or killing it with the impact, then turns to catch it in mid-air. If its prey is too heavy to carry, a peregrine will drop it to the ground and eat it there. If they miss the initial strike, peregrines will chase their prey in a twisting flight.

 

Although previously thought rare, several cases of peregrines contour-hunting, i.e., using natural contours to surprise and ambush prey on the ground, have been reported and even rare cases of prey being pursued on foot. In addition, peregrines have been documented preying on chicks in nests, from birds such as kittiwakes. Prey is plucked before consumption. A 2016 study showed that the presence of peregrines benefits non-preferred species while at the same time causing a decline in its preferred prey. As of 2018, the fastest recorded falcon was at 242 mph (nearly 390 km/h). Researchers at the University of Groningen in the Netherlands and at Oxford University used 3D computer simulations in 2018 to show that the high speed allows peregrines to gain better maneuverability and precision in strikes.

 

Reproduction

The peregrine falcon is sexually mature at one to three years of age, but in larger populations they breed after two to three years of age. A pair mates for life and returns to the same nesting spot annually. The courtship flight includes a mix of aerial acrobatics, precise spirals, and steep dives. The male passes prey it has caught to the female in mid-air. To make this possible, the female actually flies upside-down to receive the food from the male's talons.

 

During the breeding season, the peregrine falcon is territorial; nesting pairs are usually more than 1 km (0.62 mi) apart, and often much farther, even in areas with large numbers of pairs. The distance between nests ensures sufficient food supply for pairs and their chicks. Within a breeding territory, a pair may have several nesting ledges; the number used by a pair can vary from one or two up to seven in a 16-year period.

 

The peregrine falcon nests in a scrape, normally on cliff edges. The female chooses a nest site, where she scrapes a shallow hollow in the loose soil, sand, gravel, or dead vegetation in which to lay eggs. No nest materials are added. Cliff nests are generally located under an overhang, on ledges with vegetation. South-facing sites are favoured. In some regions, as in parts of Australia and on the west coast of northern North America, large tree hollows are used for nesting. Before the demise of most European peregrines, a large population of peregrines in central and western Europe used the disused nests of other large birds. In remote, undisturbed areas such as the Arctic, steep slopes and even low rocks and mounds may be used as nest sites. In many parts of its range, peregrines now also nest regularly on tall buildings or bridges; these human-made structures used for breeding closely resemble the natural cliff ledges that the peregrine prefers for its nesting locations.

 

The pair defends the chosen nest site against other peregrines, and often against ravens, herons, and gulls, and if ground-nesting, also such mammals as foxes, wolverines, felids, bears, wolves, and mountain lions. Both nests and (less frequently) adults are predated by larger-bodied raptorial birds like eagles, large owls, or gyrfalcons. The most serious predators of peregrine nests in North America and Europe are the great horned owl and the Eurasian eagle-owl. When reintroductions have been attempted for peregrines, the most serious impediments were these two species of owls routinely picking off nestlings, fledglings and adults by night. Peregrines defending their nests have managed to kill raptors as large as golden eagles and bald eagles (both of which they normally avoid as potential predators) that have come too close to the nest by ambushing them in a full stoop. In one instance, when a snowy owl killed a newly fledged peregrine, the larger owl was in turn killed by a stooping peregrine parent.

 

The date of egg-laying varies according to locality, but is generally from February to March in the Northern Hemisphere, and from July to August in the Southern Hemisphere, although the Australian subspecies macropus may breed as late as November, and equatorial populations may nest anytime between June and December. If the eggs are lost early in the nesting season, the female usually lays another clutch, although this is extremely rare in the Arctic due to the short summer season. Generally three to four eggs, but sometimes as few as one or as many as five, are laid in the scrape. The eggs are white to buff with red or brown markings. They are incubated for 29 to 33 days, mainly by the female, with the male also helping with the incubation of the eggs during the day, but only the female incubating them at night. The average number of young found in nests is 2.5, and the average number that fledge is about 1.5, due to the occasional production of infertile eggs and various natural losses of nestlings.

 

After hatching, the chicks (called "eyases") are covered with creamy-white down and have disproportionately large feet. The male (called the "tiercel") and the female (simply called the "falcon") both leave the nest to gather prey to feed the young. The hunting territory of the parents can extend a radius of 19 to 24 km (12 to 15 mi) from the nest site. Chicks fledge 42 to 46 days after hatching, and remain dependent on their parents for up to two months.

 

Relationship with humans

The peregrine falcon is a highly admired falconry bird, and has been used in falconry for more than 3,000 years, beginning with nomads in central Asia. Its advantages in falconry include not only its athleticism and eagerness to hunt, but an equable disposition that leads to it being one of the easier falcons to train. The peregrine falcon has the additional advantage of a natural flight style of circling above the falconer ("waiting on") for game to be flushed, and then performing an effective and exciting high-speed diving stoop to take the quarry. The speed of the stoop not only allows the falcon to catch fast flying birds, it also enhances the falcon's ability to execute maneuvers to catch highly agile prey, and allows the falcon to deliver a knockout blow with a fist-like clenched talon against game that may be much larger than itself.

 

Additionally the versatility of the species, with agility allowing capture of smaller birds and a strength and attacking style allowing capture of game much larger than themselves, combined with the wide size range of the many peregrine subspecies, means there is a subspecies suitable to almost any size and type of game bird. This size range, evolved to fit various environments and prey species, is from the larger females of the largest subspecies to the smaller males of the smallest subspecies, approximately five to one (approximately 1500 g to 300 g). The males of smaller and medium-sized subspecies, and the females of the smaller subspecies, excel in the taking of swift and agile small game birds such as dove, quail, and smaller ducks. The females of the larger subspecies are capable of taking large and powerful game birds such as the largest of duck species, pheasant, and grouse.

 

Peregrine falcons handled by falconers are also occasionally used to scare away birds at airports to reduce the risk of bird-plane strikes, improving air-traffic safety. They were also used to intercept homing pigeons during World War II.

 

Peregrine falcons have been successfully bred in captivity, both for falconry and for release into the wild. Until 2004 nearly all peregrines used for falconry in the US were captive-bred from the progeny of falcons taken before the US Endangered Species Act was enacted and from those few infusions of wild genes available from Canada and special circumstances. Peregrine falcons were removed from the United States' endangered species list in 1999. The successful recovery program was aided by the effort and knowledge of falconers – in collaboration with The Peregrine Fund and state and federal agencies – through a technique called hacking. Finally, after years of close work with the US Fish and Wildlife Service, a limited take of wild peregrines was allowed in 2004, the first wild peregrines taken specifically for falconry in over 30 years.

 

The development of captive breeding methods has led to peregrines being commercially available for falconry use, thus mostly eliminating the need to capture wild birds for support of falconry. The main reason for taking wild peregrines at this point is to maintain healthy genetic diversity in the breeding lines. Hybrids of peregrines and gyrfalcons are also available that can combine the best features of both species to create what many consider to be the ultimate falconry bird for the taking of larger game such as the sage-grouse. These hybrids combine the greater size, strength, and horizontal speed of the gyrfalcon with the natural propensity to stoop and greater warm weather tolerance of the peregrine.

 

Decline due to pesticides

The peregrine falcon became an endangered species over much of its range because of the use of organochlorine pesticides, especially DDT, during the 1950s, '60s, and '70s. Pesticide biomagnification caused organochlorine to build up in the falcons' fat tissues, reducing the amount of calcium in their eggshells. With thinner shells, fewer falcon eggs survived until hatching. In addition, the PCB concentrations found in these falcons is dependent upon the age of the falcon. While high levels are still found in young birds (only a few months old) and even higher concentrations are found in more mature falcons, further increasing in adult peregrine falcons. These pesticides caused falcon prey to also have thinner eggshells (one example of prey being the Black Petrels). In several parts of the world, such as the eastern United States and Belgium, this species became extirpated (locally extinct) as a result. An alternate point of view is that populations in the eastern North America had vanished due to hunting and egg collection. Following the ban of organochlorine pesticides, the reproductive success of Peregrines increased in Scotland in terms of territory occupancy and breeding success, although spatial variation in recovery rates indicate that in some areas Peregrines were also impacted by other factors such as persecution.

 

Recovery efforts

Peregrine falcon recovery teams breed the species in captivity. The chicks are usually fed through a chute or with a hand puppet mimicking a peregrine's head, so they cannot see to imprint on the human trainers. Then, when they are old enough, the rearing box is opened, allowing the bird to train its wings. As the fledgling gets stronger, feeding is reduced, forcing the bird to learn to hunt. This procedure is called hacking back to the wild. To release a captive-bred falcon, the bird is placed in a special cage at the top of a tower or cliff ledge for some days or so, allowing it to acclimate itself to its future environment.

 

Worldwide recovery efforts have been remarkably successful. The widespread restriction of DDT use eventually allowed released birds to breed successfully. The peregrine falcon was removed from the U.S. Endangered Species list on 25 August 1999.

 

Some controversy has existed over the origins of captive breeding stock used by the Peregrine Fund in the recovery of peregrine falcons throughout the contiguous United States. Several peregrine subspecies were included in the breeding stock, including birds of Eurasian origin. Due to the extirpation of the eastern population of Falco peregrinus anatum, the near-extirpation of anatum in the Midwest and the limited gene pool within North American breeding stock, the inclusion of non-native subspecies was justified to optimize the genetic diversity found within the species as a whole.

 

During the 1970s, peregrine falcons in Finland experienced a population bottleneck as a result of large declines associated with bio-accumulation of organochloride pesticides. However, the genetic diversity of peregrines in Finland is similar to other populations, indicating that high dispersal rates have maintained the genetic diversity of this species.

 

Since peregrine falcon eggs and chicks are still often targeted by illegal poachers, it is common practice not to publicize unprotected nest locations.

 

Current status

Populations of the peregrine falcon have bounced back in most parts of the world. In the United Kingdom, there has been a recovery of populations since the crash of the 1960s. This has been greatly assisted by conservation and protection work led by the Royal Society for the Protection of Birds. The RSPB estimated that there were 1,402 breeding pairs in the UK in 2011. In Canada, where peregrines were identified as endangered in 1978 (in the Yukon territory of northern Canada that year, only a single breeding pair was identified), the Committee on the Status of Endangered Wildlife in Canada declared the species no longer at risk in December 2017.

 

Peregrines now breed in many mountainous and coastal areas, especially in the west and north, and nest in some urban areas, capitalising on the urban feral pigeon populations for food. Additionally, falcons benefit from artificial illumination, which allows the raptors to extend their hunting periods into the dusk when natural illumination would otherwise be too low for them to pursue prey. In England, this has allowed them to prey on nocturnal migrants such as redwings, fieldfares, starlings, and woodcocks.

 

In many parts of the world peregrine falcons have adapted to urban habitats, nesting on cathedrals, skyscraper window ledges, tower blocks, and the towers of suspension bridges. Many of these nesting birds are encouraged, sometimes gathering media attention and often monitored by cameras.

 

In England, peregrine falcons have become increasingly urban in distribution, particularly in southern areas where inland cliffs suitable as nesting sites are scarce. The first recorded urban breeding pair was observed nesting on the Swansea Guildhall in the 1980s. In Southampton, a nest prevented restoration of mobile telephony services for several months in 2013, after Vodafone engineers despatched to repair a faulty transmitter mast discovered a nest in the mast, and were prevented by the Wildlife and Countryside Act – on pain of a possible prison sentence – from proceeding with repairs until the chicks fledged.

 

In Oregon, Portland houses ten percent of the state's peregrine nests, despite only covering around 0.1 percent of the state's land area.

 

Cultural significance

Due to its striking hunting technique, the peregrine has often been associated with aggression and martial prowess. The Ancient Egyptian solar deity Ra was often represented as a man with the head of a peregrine falcon adorned with the solar disk, although most Egyptologists agree that it's most likely a Lanner falcon. Native Americans of the Mississippian culture (c. 800–1500) used the peregrine, along with several other birds of prey, in imagery as a symbol of "aerial (celestial) power" and buried men of high status in costumes associating to the ferocity of raptorial birds. In the late Middle Ages, the Western European nobility that used peregrines for hunting, considered the bird associated with princes in formal hierarchies of birds of prey, just below the gyrfalcon associated with kings. It was considered "a royal bird, more armed by its courage than its claws". Terminology used by peregrine breeders also used the Old French term gentil, "of noble birth; aristocratic", particularly with the peregrine.

 

The peregrine falcon is the national animal of the United Arab Emirates. Since 1927, the peregrine falcon has been the official mascot of Bowling Green State University in Bowling Green, Ohio. The 2007 U.S. Idaho state quarter features a peregrine falcon. The peregrine falcon has been designated the official city bird of Chicago.

 

The Peregrine, by J. A. Baker, is widely regarded as one of the best nature books in English written in the twentieth century. Admirers of the book include Robert Macfarlane, Mark Cocker, who regards the book as "one of the most outstanding books on nature in the twentieth century" and Werner Herzog, who called it "the one book I would ask you to read if you want to make films", and said elsewhere "it has prose of the calibre that we have not seen since Joseph Conrad". In the book, Baker recounts, in diary form, his detailed observations of peregrines (and their interaction with other birds) near his home in Chelmsford, Essex, over a single winter from October to April.

 

An episode of the hour-long TV series Starman in 1986 titled "Peregrine" was about an injured peregrine falcon and the endangered species program. It was filmed with the assistance of the University of California's peregrine falcon project in Santa Cruz.

Entrance door to the previous headquarters (1923) of the Amsterdam telephony service, artwork made by Hildo Krop, supposedly depicting the 4 continents and also the Staff of Mercury, the old Roman god ruling all affairs concerning Communication.

What is SIP Trunking? What are SIP Trunks? A SIP trunk is a service offered by a supplier such as Tpad that permits businesses that have a Telephone PBX to use Voice-over-IP (VoIP), outside the enterprise network by using the Internet connection. The SIP Trunk enables connection to the standard PSTN and Mobile telephone networks via Tpad, thus eliminating the need for expensive multiple ISDN lines and also benefiting from lower cost calls. A business would keep their existing phone system but look to gain benefit from a SIP Trunk, especially if they were making a large amount of phone calls on a month by month basis and/or their call spend was high.

 

SIP trunks can offer additional significant cost-savings for enterprises, eliminating the need for local PSTN gateways, costly ISDN BRIs (Basic Rate Interfaces) or PRIs (Primary Rate Interfaces).

 

Take advantage of our Tpad's low cost SIP Trunk line rental and call charges which are compatible with all types of telephone system that can use SIP / IAX Trunks. Whether you need a VoIP telephony service for your Asterisk or VoIP Gateways connected to your Legacy (Analogue) PBX, or need to utilize the VoIP capabilities of your Panasonic, Siemens, Avaya, Samsung, LG, Toshiba or other brand of phone system then Tpad's business quality SIP trunking services are for you.

 

Even if you don't have a VoIP compatible phone system you can still take advantage of our low rates and line rental by installing a VoIP gateway which will allow you to route calls from your phone system across the internet to our SIP Trunk service. For small business users with Analogue or ISDN BRI telephone lines you can connect any popular VoIP gateway which offers VoIP G729 / G711 connectivity for SIP Trunks as well as branch office connectivity for free calls between offices.

 

Get Tpad SIP Trunking for your business now simply visit our website for a Free Quotation, www.tpadbusiness.co.uk/business/services/sip-trunks.html

About Tpad (www.tpad.com)

 

Tpad is an international telecommunications company with over 1000 customers in the UK and operating internationally from regional offices in Dubai and Cyprus. With over 20 years experience in the IT Sector, our expertise spans the development, manufacture and installation of business telephony services. Tpad offers solutions to meet all requirements ranging from Managed IP/PBX solutions to complex Contact Centre installations. The Tpad Global Phone Network that has unlimited call handling capacity and our in-house developed software provides an on-net feature rich environment with functionality that includes network based call recording, call reporting, least cost routing, Diallers, Multi-Site solutions and much more.

 

With over 100 operatives within dedicated Service, Support and Development teams, we pride ourselves on service excellence and after sales care, consistently delivering some of the highest level of support within the industry. Our services are backed by a choice of service levels to dovetail with the skill sets within your organisation.

 

The Tpad Platform and solutions are able to operate using third party connectivity. However, Tpad is able to offer a range of connectivity solutions by partnering with its sister company, Internexus. With over 12 years in the sector, that includes the provision and management of network solutions for Supanet, the largest independent Internet Service Provider in the UK with over 500 thousand customers using the a range of services from Internet access, line rental and carrier pre-select services, Tpad and Internexus boast experience and knowledge that almost is unparalleled within the industry. Indeed, it was this combined experience that led to the establishment of Tpad's On-Net Voice Platform in 2005, when users were first given the ability to log in and make phone calls over the Internet. Whilst we have come a long way since then, adding the Tpad Platform to our broad range of business IP/PBX equipment powered by industry leading connectivity make our systems a truly one-stop solution.

 

Tpad's exciting product range is constantly evolving to keep pace both with technological advancements and our commitment offering our customers new and improved features and enhanced capabilities. At the same time we remain true to our core values of providing reliable cutting edge technology with outstanding service delivery. Telecommunications are the foundations on which almost every business in the 21st century is built. Information is power and communication is king. Whatever your requirements Tpad is ideally positioned to deliver a flexible cost-effective solution to satisfy your organisation's telecommunications requirements both for today and in the years to come. Let Tpad be a valuable partner in your business.

 

About Tpad (www.tpad.com)

 

Tpad is an international telecommunications company with over 1000 customers in the UK and operating internationally from regional offices in Dubai and Cyprus. With over 20 years experience in the IT Sector, our expertise spans the development, manufacture and installation of business telephony services. Tpad offers solutions to meet all requirements ranging from Managed IP/PBX solutions to complex Contact Centre installations. The Tpad Global Phone Network that has unlimited call handling capacity and our in-house developed software provides an on-net feature rich environment with functionality that includes network based call recording, call reporting, least cost routing, Diallers, Multi-Site solutions and much more.

 

With over 100 operatives within dedicated Service, Support and Development teams, we pride ourselves on service excellence and after sales care, consistently delivering some of the highest level of support within the industry. Our services are backed by a choice of service levels to dovetail with the skill sets within your organisation.

 

The Tpad Platform and solutions are able to operate using third party connectivity. However, Tpad is able to offer a range of connectivity solutions by partnering with its sister company, Internexus. With over 12 years in the sector, that includes the provision and management of network solutions for Supanet, the largest independent Internet Service Provider in the UK with over 500 thousand customers using the a range of services from Internet access, line rental and carrier pre-select services, Tpad and Internexus boast experience and knowledge that almost is unparalleled within the industry. Indeed, it was this combined experience that led to the establishment of Tpad's On-Net Voice Platform in 2005, when users were first given the ability to log in and make phone calls over the Internet. Whilst we have come a long way since then, adding the Tpad Platform to our broad range of business IP/PBX equipment powered by industry leading connectivity make our systems a truly one-stop solution.

 

Tpad's exciting product range is constantly evolving to keep pace both with technological advancements and our commitment offering our customers new and improved features and enhanced capabilities. At the same time we remain true to our core values of providing reliable cutting edge technology with outstanding service delivery. Telecommunications are the foundations on which almost every business in the 21st century is built. Information is power and communication is king. Whatever your requirements Tpad is ideally positioned to deliver a flexible cost-effective solution to satisfy your organisation's telecommunications requirements both for today and in the years to come. Let Tpad be a valuable partner in your business.

 

Singapore Telecommunications Limited , commonly abbreviated as SingTel, is Asia's leading telecommunications company, with a combined mobile subscriber base of 249.4 million customers from its own operations and regional associates at the end of March 2009. SingTel provides ISP (SingNet), IPTV (mio TV), mobile phone and fixed line telephony services.

SingTel has expanded aggressively outside its home market and owns shares in many regional operators, including 100% of the second largest Australian telco, Optus, which was acquired in 2000 from Cable & Wireless and other shareholders of Optus, and Bharti Airtel, the largest telco in India.

SingTel is the largest company by market capitalization listed on the Singapore Exchange and is majority owned by Temasek Holdings (wiki)

About Tpad (www.tpad.com)

 

Tpad is an international telecommunications company with over 1000 customers in the UK and operating internationally from regional offices in Dubai and Cyprus. With over 20 years experience in the IT Sector, our expertise spans the development, manufacture and installation of business telephony services. Tpad offers solutions to meet all requirements ranging from Managed IP/PBX solutions to complex Contact Centre installations. The Tpad Global Phone Network that has unlimited call handling capacity and our in-house developed software provides an on-net feature rich environment with functionality that includes network based call recording, call reporting, least cost routing, Diallers, Multi-Site solutions and much more.

 

With over 100 operatives within dedicated Service, Support and Development teams, we pride ourselves on service excellence and after sales care, consistently delivering some of the highest level of support within the industry. Our services are backed by a choice of service levels to dovetail with the skill sets within your organisation.

 

The Tpad Platform and solutions are able to operate using third party connectivity. However, Tpad is able to offer a range of connectivity solutions by partnering with its sister company, Internexus. With over 12 years in the sector, that includes the provision and management of network solutions for Supanet, the largest independent Internet Service Provider in the UK with over 500 thousand customers using the a range of services from Internet access, line rental and carrier pre-select services, Tpad and Internexus boast experience and knowledge that almost is unparalleled within the industry. Indeed, it was this combined experience that led to the establishment of Tpad's On-Net Voice Platform in 2005, when users were first given the ability to log in and make phone calls over the Internet. Whilst we have come a long way since then, adding the Tpad Platform to our broad range of business IP/PBX equipment powered by industry leading connectivity make our systems a truly one-stop solution.

 

Tpad's exciting product range is constantly evolving to keep pace both with technological advancements and our commitment offering our customers new and improved features and enhanced capabilities. At the same time we remain true to our core values of providing reliable cutting edge technology with outstanding service delivery. Telecommunications are the foundations on which almost every business in the 21st century is built. Information is power and communication is king. Whatever your requirements Tpad is ideally positioned to deliver a flexible cost-effective solution to satisfy your organisation's telecommunications requirements both for today and in the years to come. Let Tpad be a valuable partner in your business.

 

Entrance door to the previous headquarters (1923) of the Amsterdam telephony service, artwork made by Hildo Krop, supposedly depicting the 4 continents and also the Staff of Mercury, the old Roman god ruling all affairs concerning Communication.

About Tpad (www.tpad.com)

 

Tpad is an international telecommunications company with over 1000 customers in the UK and operating internationally from regional offices in Dubai and Cyprus. With over 20 years experience in the IT Sector, our expertise spans the development, manufacture and installation of business telephony services. Tpad offers solutions to meet all requirements ranging from Managed IP/PBX solutions to complex Contact Centre installations. The Tpad Global Phone Network that has unlimited call handling capacity and our in-house developed software provides an on-net feature rich environment with functionality that includes network based call recording, call reporting, least cost routing, Diallers, Multi-Site solutions and much more.

 

With over 100 operatives within dedicated Service, Support and Development teams, we pride ourselves on service excellence and after sales care, consistently delivering some of the highest level of support within the industry. Our services are backed by a choice of service levels to dovetail with the skill sets within your organisation.

 

The Tpad Platform and solutions are able to operate using third party connectivity. However, Tpad is able to offer a range of connectivity solutions by partnering with its sister company, Internexus. With over 12 years in the sector, that includes the provision and management of network solutions for Supanet, the largest independent Internet Service Provider in the UK with over 500 thousand customers using the a range of services from Internet access, line rental and carrier pre-select services, Tpad and Internexus boast experience and knowledge that almost is unparalleled within the industry. Indeed, it was this combined experience that led to the establishment of Tpad's On-Net Voice Platform in 2005, when users were first given the ability to log in and make phone calls over the Internet. Whilst we have come a long way since then, adding the Tpad Platform to our broad range of business IP/PBX equipment powered by industry leading connectivity make our systems a truly one-stop solution.

 

Tpad's exciting product range is constantly evolving to keep pace both with technological advancements and our commitment offering our customers new and improved features and enhanced capabilities. At the same time we remain true to our core values of providing reliable cutting edge technology with outstanding service delivery. Telecommunications are the foundations on which almost every business in the 21st century is built. Information is power and communication is king. Whatever your requirements Tpad is ideally positioned to deliver a flexible cost-effective solution to satisfy your organisation's telecommunications requirements both for today and in the years to come. Let Tpad be a valuable partner in your business.

 

The "Rent-a-minute" philosophy that subtends the 'RAM-Exchange' Interface literally enriches a category of public domain actors often ignored when considering new urban environments: Street vendors. The mobile telephony service offered, which is increasingly present in the described vendor community, is enhanced by developing a secondary electronic market for the vendors themselves, giving their offers a competitive edge and potential returns previously curbed by subservience to the speculative strategies of telecom majors. Their services are paradoxically construed as a way to activate informal poles of urban communications in Latin American city culture. A strong political standpoint is taken in the project's endorsing and enhancing of Street Vendor activities. Their urban cultural practices are used as the pivot for undercutting and doubling up on telecom giant profit-making, by allowing them to draw their own advantage from volatile tariffs implemented by industrial consortia. New media and technology in the form of mobile telephony are situated and interpreted in the specific cultural context of a large Latin American city, characterised by intensive, gregarious, highly communicative street life.

 

Co-lateral CCS (Federico Bozo_Gustavo Muci_Armando Montilla_Alejandra Salas)

Entrance door to the previous headquarters (1923) of the Amsterdam telephony service, artwork made by Hildo Krop, supposedly depicting the 4 continents and also the Staff of Mercury, the old Roman god ruling all affairs concerning Communication.

About Tpad (www.tpad.com)

 

Tpad is an international telecommunications company with over 1000 customers in the UK and operating internationally from regional offices in Dubai and Cyprus. With over 20 years experience in the IT Sector, our expertise spans the development, manufacture and installation of business telephony services. Tpad offers solutions to meet all requirements ranging from Managed IP/PBX solutions to complex Contact Centre installations. The Tpad Global Phone Network that has unlimited call handling capacity and our in-house developed software provides an on-net feature rich environment with functionality that includes network based call recording, call reporting, least cost routing, Diallers, Multi-Site solutions and much more.

 

With over 100 operatives within dedicated Service, Support and Development teams, we pride ourselves on service excellence and after sales care, consistently delivering some of the highest level of support within the industry. Our services are backed by a choice of service levels to dovetail with the skill sets within your organisation.

 

The Tpad Platform and solutions are able to operate using third party connectivity. However, Tpad is able to offer a range of connectivity solutions by partnering with its sister company, Internexus. With over 12 years in the sector, that includes the provision and management of network solutions for Supanet, the largest independent Internet Service Provider in the UK with over 500 thousand customers using the a range of services from Internet access, line rental and carrier pre-select services, Tpad and Internexus boast experience and knowledge that almost is unparalleled within the industry. Indeed, it was this combined experience that led to the establishment of Tpad's On-Net Voice Platform in 2005, when users were first given the ability to log in and make phone calls over the Internet. Whilst we have come a long way since then, adding the Tpad Platform to our broad range of business IP/PBX equipment powered by industry leading connectivity make our systems a truly one-stop solution.

 

Tpad's exciting product range is constantly evolving to keep pace both with technological advancements and our commitment offering our customers new and improved features and enhanced capabilities. At the same time we remain true to our core values of providing reliable cutting edge technology with outstanding service delivery. Telecommunications are the foundations on which almost every business in the 21st century is built. Information is power and communication is king. Whatever your requirements Tpad is ideally positioned to deliver a flexible cost-effective solution to satisfy your organisation's telecommunications requirements both for today and in the years to come. Let Tpad be a valuable partner in your business.

 

Jeremy Darroch, chief executive of Sky, grew up in Northumberland before studying economics at Hull University Business School.

 

Having started his career at Deloitte, where he qualified as a chartered accountant, Jeremy moved to Proctor & Gamble. He remained there for 12 years, working in various roles, and eventually became European finance director for the health care business. He joined what was then Dixons Group plc as retail finance director in 2000, later becoming the group finance director of DSG International plc.

 

Jeremy joined Sky as chief financial officer in 2004 and was appointed chief executive in 2007. During his time at Sky, the company has expanded into the wider marketplace for entertainment and communications through the launch of its broadband and telephony services. Despite tough economic conditions, it has also continued to innovate through steady investment, leading the move to HD and 3D television.

 

Operational performance at Sky has been consistently strong throughout his tenure, with customer numbers on track to hit his target of 10 million by the end of 2010 and product sales growing by 1,000% over the past four years.

 

Jeremy is a non-executive director and the chairman of the audit committee of Marks & Spencer. He is also a member of the Youth Sport Trust board and of the Council for Industry and Higher Education.

About Tpad (www.tpad.com)

 

Tpad is an international telecommunications company with over 1000 customers in the UK and operating internationally from regional offices in Dubai and Cyprus. With over 20 years experience in the IT Sector, our expertise spans the development, manufacture and installation of business telephony services. Tpad offers solutions to meet all requirements ranging from Managed IP/PBX solutions to complex Contact Centre installations. The Tpad Global Phone Network that has unlimited call handling capacity and our in-house developed software provides an on-net feature rich environment with functionality that includes network based call recording, call reporting, least cost routing, Diallers, Multi-Site solutions and much more.

 

With over 100 operatives within dedicated Service, Support and Development teams, we pride ourselves on service excellence and after sales care, consistently delivering some of the highest level of support within the industry. Our services are backed by a choice of service levels to dovetail with the skill sets within your organisation.

 

The Tpad Platform and solutions are able to operate using third party connectivity. However, Tpad is able to offer a range of connectivity solutions by partnering with its sister company, Internexus. With over 12 years in the sector, that includes the provision and management of network solutions for Supanet, the largest independent Internet Service Provider in the UK with over 500 thousand customers using the a range of services from Internet access, line rental and carrier pre-select services, Tpad and Internexus boast experience and knowledge that almost is unparalleled within the industry. Indeed, it was this combined experience that led to the establishment of Tpad's On-Net Voice Platform in 2005, when users were first given the ability to log in and make phone calls over the Internet. Whilst we have come a long way since then, adding the Tpad Platform to our broad range of business IP/PBX equipment powered by industry leading connectivity make our systems a truly one-stop solution.

 

Tpad's exciting product range is constantly evolving to keep pace both with technological advancements and our commitment offering our customers new and improved features and enhanced capabilities. At the same time we remain true to our core values of providing reliable cutting edge technology with outstanding service delivery. Telecommunications are the foundations on which almost every business in the 21st century is built. Information is power and communication is king. Whatever your requirements Tpad is ideally positioned to deliver a flexible cost-effective solution to satisfy your organisation's telecommunications requirements both for today and in the years to come. Let Tpad be a valuable partner in your business.

 

SIP trunks can offer additional significant cost-savings for enterprises, eliminating the need for local PSTN gateways, costly ISDN BRIs (Basic Rate Interfaces) or PRIs (Primary Rate Interfaces).

 

Take advantage of our Tpad's low cost SIP Trunk line rental and call charges which are compatible with all types of telephone system that can use SIP / IAX Trunks. Whether you need a VoIP telephony service for your Asterisk or VoIP Gateways connected to your Legacy (Analogue) PBX, or need to utilize the VoIP capabilities of your Panasonic, Siemens, Avaya, Samsung, LG, Toshiba or other brand of phone system then Tpad's business quality SIP trunking services are for you.

 

Get Tpad SIP Trunking for your business now simply visit our website for a Free Quotation, www.tpadbusiness.co.uk/business/services/sip-trunks.html

About Tpad (www.tpad.com)

 

Tpad is an international telecommunications company with over 1000 customers in the UK and operating internationally from regional offices in Dubai and Cyprus. With over 20 years experience in the IT Sector, our expertise spans the development, manufacture and installation of business telephony services. Tpad offers solutions to meet all requirements ranging from Managed IP/PBX solutions to complex Contact Centre installations. The Tpad Global Phone Network that has unlimited call handling capacity and our in-house developed software provides an on-net feature rich environment with functionality that includes network based call recording, call reporting, least cost routing, Diallers, Multi-Site solutions and much more.

 

With over 100 operatives within dedicated Service, Support and Development teams, we pride ourselves on service excellence and after sales care, consistently delivering some of the highest level of support within the industry. Our services are backed by a choice of service levels to dovetail with the skill sets within your organisation.

 

The Tpad Platform and solutions are able to operate using third party connectivity. However, Tpad is able to offer a range of connectivity solutions by partnering with its sister company, Internexus. With over 12 years in the sector, that includes the provision and management of network solutions for Supanet, the largest independent Internet Service Provider in the UK with over 500 thousand customers using the a range of services from Internet access, line rental and carrier pre-select services, Tpad and Internexus boast experience and knowledge that almost is unparalleled within the industry. Indeed, it was this combined experience that led to the establishment of Tpad's On-Net Voice Platform in 2005, when users were first given the ability to log in and make phone calls over the Internet. Whilst we have come a long way since then, adding the Tpad Platform to our broad range of business IP/PBX equipment powered by industry leading connectivity make our systems a truly one-stop solution.

 

Tpad's exciting product range is constantly evolving to keep pace both with technological advancements and our commitment offering our customers new and improved features and enhanced capabilities. At the same time we remain true to our core values of providing reliable cutting edge technology with outstanding service delivery. Telecommunications are the foundations on which almost every business in the 21st century is built. Information is power and communication is king. Whatever your requirements Tpad is ideally positioned to deliver a flexible cost-effective solution to satisfy your organisation's telecommunications requirements both for today and in the years to come. Let Tpad be a valuable partner in your business.

 

Still have fond memories of phone booth. A good place for shelter from rain, other than telephony service that I could use to arrange of pick-up. She will know nothing about these.

 

Didn't know now that this has become an attraction of England

Tpad specializes in voice over IP telephony systems which enable an advanced communications model that can serve the needs of large and small companies worldwide. With many years of experience and expertise in the field of telecom, they offer planning, installation and servicing of office telephone systems that are not only cost-effective, but highly adaptable as well. They offer a wide selection of PBX packages that are efficient, highly upgradable and compatible with digital or IP lines. Aside from PBX systems, Tpad also provides a variety of advanced business telephony services ranging from virtual numbers, SIP trunks, and number porting.

About Tpad (http://www.tpad.com)

Tpad is an International company which has its Headquarters in Jersey, and provides products and services in many countries including the UAE, India, Hong Kong, USA, UK and Canada. In the UK, Tpad provides advanced telephony services and carrier-grade SIP trunks to businesses worldwide. Until recently the core product has been large IP PBX based business telephone systems (50 users to 500+ users and multiple sites). Tpad also has an International consumer arm providing low cost international services to all major countries. Over the past 3 years, Tpad has been investing in next generation telephone technology with the development of a powerful central telephone platform called Colossus that resides in Tpad data centres and that can be accessed from anywhere in the cloud space.

 

• Chris Kelly MP ‘cuts the ribbon’

• Modern open plan environment

• Opening hours extended by more 15 hours a week

• Full range of Post Office® services available

 

To celebrate the launch of the new modern branch Dudley South MP, Chris Kelly, ‘cut the ribbon’ at an opening ceremony at Merry Hill Centre Post Office, today Friday 12 July 2013.

 

Chris Kelly MP said: “I’m delighted to officially open the new Merry Hill Centre Post Office. Nick and his team are to be congratulated on their work to develop the new branch which will make a real difference for local residents and small business customers. I wish them every success with the venture.”

 

The new Merry Hill Centre Post Office has six serving positions, a mixture of two screened and three open plan counter positions, with a Post Office serving point provided for use at the retail counter – a new concept for delivering Post Office services alongside retail transactions during shop opening hours.

 

Opening hours at the new location have been extended by 15 hours a week to Monday to Saturday; 09:00 – 19:00 and Sunday: 11.00 – 17.00 offering customers access Post Office products and services 7 days a week.

 

Subpostmaster Nick Singh, who hosted today’s official opening, said: “I was delighted to have the opportunity to modernise the Post Office and very pleased to be re-launching it today for the community. I know customers will like the new environment and I’d like to thank Mr Kelly for coming today.”

 

Post Office Regional Manager, Richard Lynds, said: “We understand how important having a Post Office is to residents in this area and we are confident that this new modern Post Office service will meet the needs of the local community and secure services for the future.”

 

The branch relocated within the Merry Hill Centre in June 2013.

 

The Post Office, now independent of Royal Mail, is the largest retail network in the UK, with over 11,500 branches. It is also one of the fastest growing financial services companies and is developing its online and telephony services.

 

Pictured: Caroline Pritchard (Area Manager PO Midlands), Chris Kelly MP (Dudley South), Nick Singh (Postmaster) and Cllr Patrick Harley (Dudley MBC)

 

ENDS

 

Further details:

 

Post Office External Relations,

1st Floor, 148 Old Street,

LONDON

EC1V 9HQ

Tel: 020 7012 3456 (24 hours),

email: pressoffice@postoffice.co.uk

www.postoffice.co.uk

 

dudleysouth.com

dudleyconservatives.com

The peregrine falcon (Falco peregrinus), also known simply as the peregrine, and historically as the duck hawk in North America, is a cosmopolitan bird of prey (raptor) in the family Falconidae. A large, crow-sized falcon, it has a blue-grey back, barred white underparts, and a black head. The peregrine is renowned for its speed. It can reach over 320 km/h (200 mph) during its characteristic hunting stoop (high-speed dive), making it the fastest member of the animal kingdom. According to a National Geographic TV program, the highest measured speed of a peregrine falcon is 389 km/h (242 mph). As is typical for bird-eating (avivore) raptors, peregrine falcons are sexually dimorphic, with females being considerably larger than males.

 

The peregrine's breeding range includes land regions from the Arctic tundra to the tropics. It can be found nearly everywhere on Earth, except extreme polar regions, very high mountains, and most tropical rainforests; the only major ice-free landmass from which it is entirely absent is New Zealand. This makes it the world's most widespread raptor and one of the most widely found wild bird species. In fact, the only land-based bird species found over a larger geographic area owes its success to human-led introduction; the domestic and feral pigeons are both domesticate forms of the rock dove, which are a major prey species for Eurasian Peregrine populations. Due to their prevalence over most other bird species in cities, feral pigeons support many peregrine populations as a staple food source, especially in urban settings.

 

The peregrine is a highly successful example of urban wildlife in much of its range, taking advantage of tall buildings as nest sites and an abundance of prey such as pigeons and ducks. Both the English and scientific names of this species mean "wandering falcon", referring to the migratory habits of many northern populations. Experts recognize 17 to 19 subspecies, which vary in appearance and range; disagreement exists over whether the distinctive Barbary falcon is represented by two subspecies of Falco peregrinus or is a separate species, F. pelegrinoides. The two species' divergence is relatively recent, during the time of the last ice age, therefore the genetic differential between them (and also the difference in their appearance) is relatively tiny. They are only about 0.6–0.8% genetically differentiated.

 

Although its diet consists almost exclusively of medium-sized birds, the peregrine will sometimes hunt small mammals, small reptiles, or even insects. Reaching sexual maturity at one year, it mates for life and nests in a scrape, normally on cliff edges or, in recent times, on tall human-made structures. The peregrine falcon became an endangered species in many areas because of the widespread use of certain pesticides, especially DDT. Since the ban on DDT from the early 1970s, populations have recovered, supported by large-scale protection of nesting places and releases to the wild.

 

The peregrine falcon is a well-respected falconry bird due to its strong hunting ability, high trainability, versatility, and availability via captive breeding. It is effective on most game bird species, from small to large. It has also been used as a religious, royal, or national symbol across multiple eras and areas of human civilization.

 

Description

 

Falco peregrinus. Royal National Park, New South Wales, Australia

The peregrine falcon has a body length of 34 to 58 cm (13–23 in) and a wingspan from 74 to 120 cm (29–47 in). The male and female have similar markings and plumage but, as with many birds of prey, the peregrine falcon displays marked sexual dimorphism in size, with the female measuring up to 30% larger than the male. Males weigh 330 to 1,000 g (12–35 oz) and the noticeably larger females weigh 700 to 1,500 g (25–53 oz). In most subspecies, males weigh less than 700 g (25 oz) and females weigh more than 800 g (28 oz), and cases of females weighing about 50% more than their male breeding mates are not uncommon. The standard linear measurements of peregrines are: the wing chord measures 26.5 to 39 cm (10.4–15.4 in), the tail measures 13 to 19 cm (5.1–7.5 in) and the tarsus measures 4.5 to 5.6 cm (1.8–2.2 in).

 

The back and the long pointed wings of the adult are usually bluish black to slate grey with indistinct darker barring (see "Subspecies" below); the wingtips are black. The white to rusty underparts are barred with thin clean bands of dark brown or black. The tail, coloured like the back but with thin clean bars, is long, narrow, and rounded at the end with a black tip and a white band at the very end. The top of the head and a "moustache" along the cheeks are black, contrasting sharply with the pale sides of the neck and white throat. The cere is yellow, as are the feet, and the beak and claws are black. The upper beak is notched near the tip, an adaptation which enables falcons to kill prey by severing the spinal column at the neck. An immature bird is much browner, with streaked, rather than barred, underparts, and has a pale bluish cere and orbital ring.

 

A study shows that their black malar stripe exists to reduce glare from solar radiation, allowing them to see better. Photos from The Macaulay Library and iNaturalist showed that the malar stripe is thicker where there is more solar radiation. That supports the solar glare hypothesis.

 

Taxonomy and systematics

 

Falco peregrinus was first described under its current binomial name by English ornithologist Marmaduke Tunstall in his 1771 work Ornithologia Britannica. The scientific name Falco peregrinus is a Medieval Latin phrase that was used by Albertus Magnus in 1225. Peregrinus is Latin, meaning "one from abroad" or "coming from foreign parts". It is likely the name was used as juvenile birds were taken while journeying to their breeding location (rather than from the nest), as falcon nests are often difficult to get at. The Latin term for falcon, falco, is related to falx, meaning "sickle", in reference to the silhouette of the falcon's long, pointed wings in flight.

 

The peregrine falcon belongs to a genus whose lineage includes the hierofalcon and the prairie falcon (F. mexicanus). This lineage probably diverged from other falcons towards the end of the Late Miocene or in the Late Pliocene, about 3–8 million years ago (mya). As the peregrine-hierofalcon group includes both Old World and North American species, it is likely that the lineage originated in western Eurasia or Africa. Its relationship to other falcons is not clear, as the issue is complicated by widespread hybridization confounding mtDNA sequence analyses. One genetic lineage of the saker falcon (F. cherrug) is known to have originated from a male saker ancestor producing fertile young with a female peregrine ancestor, and the descendants further breeding with sakers.

 

Today, peregrines are regularly paired in captivity with other species such as the lanner falcon (F. biarmicus) to produce the "perilanner", a somewhat popular bird in falconry as it combines the peregrine's hunting skill with the lanner's hardiness, or the gyrfalcon to produce large, strikingly coloured birds for the use of falconers. As can be seen, the peregrine is still genetically close to the hierofalcons, though their lineages diverged in the Late Pliocene (maybe some 2.5–2 mya in the Gelasian).

 

Subspecies

Numerous subspecies of Falco peregrinus have been described, with 19 accepted by the 1994 Handbook of the Birds of the World, which considers the Barbary falcon of the Canary Islands and coastal North Africa to be two subspecies (pelegrinoides and babylonicus) of Falco peregrinus, rather than a distinct species, F. pelegrinoides. The following map shows the general ranges of these 19 subspecies.

 

A map of the world, green shows on several continents, but there are also several big bare spots marked with E for extinct

Breeding ranges of the 19 subspecies

 

Falco peregrinus anatum, described by Bonaparte in 1838, is known as the American peregrine falcon or "duck hawk"; its scientific name means "duck peregrine falcon". At one time, it was partly included in leucogenys. It is mainly found in the Rocky Mountains. It was formerly common throughout North America between the tundra and northern Mexico, where current reintroduction efforts are being made to restore the population. Most mature anatum, except those that breed in more northern areas, winter in their breeding range. Most vagrants that reach western Europe seem to belong to the more northern and strongly migratory tundrius, only considered distinct since 1968. It is similar to the nominate subspecies but is slightly smaller; adults are somewhat paler and less patterned below, but juveniles are darker and more patterned below. Males weigh 500 to 700 g (1.1–1.5 lb), while females weigh 800 to 1,100 g (1.8–2.4 lb). It has become extinct in eastern North America and populations there are hybrids as a result of reintroductions of birds from elsewhere.

Falco peregrinus babylonicus, described by P.L. Sclater in 1861, is found in eastern Iran along the Hindu Kush and the Tian Shan to the Mongolian Altai ranges. A few birds winter in northern and northwestern India, mainly in dry semi-desert habitats. It is paler than pelegrinoides and somewhat similar to a small, pale lanner falcon (Falco biarmicus). Males weigh 330 to 400 grams (12 to 14 oz), while females weigh 513 to 765 grams (18.1 to 27.0 oz).

Falco peregrinus brookei, described by Sharpe in 1873, is also known as the Mediterranean peregrine falcon or the Maltese falcon. It includes caucasicus and most specimens of the proposed race punicus, though others may be pelegrinoides (Barbary falcons), or perhaps the rare hybrids between these two which might occur around Algeria. They occur from the Iberian Peninsula around the Mediterranean, except in arid regions, to the Caucasus. They are non-migratory. It is smaller than the nominate subspecies and the underside usually has a rusty hue. Males weigh around 445 g (0.981 lb), while females weigh up to 920 g (2.03 lb).

Falco peregrinus calidus, described by John Latham in 1790, it was formerly called leucogenys and includes caeruleiceps. It breeds in the Arctic tundra of Eurasia from Murmansk Oblast to roughly Yana and Indigirka Rivers, Siberia. It is completely migratory and travels south in winter as far as South Asia and sub-Saharan Africa. It is often seen around wetland habitats. It is paler than the nominate subspecies, especially on the crown. Males weigh 588 to 740 g (1.296–1.631 lb), while females weigh 925 to 1,333 g (2.039–2.939 lb).

Falco peregrinus cassini, described by Sharpe in 1873, is also known as the austral peregrine falcon. It includes kreyenborgi, the pallid falcon, a leucistic colour morph occurring in southernmost South America, which was long believed to be a distinct species. Its range includes South America from Ecuador through Bolivia, northern Argentina and Chile to Tierra del Fuego and the Falkland Islands. It is non-migratory. It is similar to the nominate subspecies, but slightly smaller with a black ear region. The pallid falcon morph kreyenborgi is medium grey above, has little barring below and has a head pattern like the saker falcon (Falco cherrug), but the ear region is white.

Falco peregrinus ernesti, described by Sharpe in 1894, is found from the Sunda Islands to the Philippines and south to eastern New Guinea and the nearby Bismarck Archipelago. Its geographical separation from nesiotes requires confirmation. It is non-migratory. It differs from the nominate subspecies in the very dark, dense barring on its underside and its black ear coverts.

Falco peregrinus furuitii, described by Momiyama in 1927, is found on the Izu and Ogasawara Islands south of Honshū, Japan. It is non-migratory. It is very rare and may only remain on a single island. It is a dark form, resembling pealei in colour, but darker, especially on the tail.

Falco peregrinus japonensis, described by Gmelin in 1788, includes kleinschmidti, pleskei, and harterti, and seems to refer to intergrades with calidus. It is found from northeast Siberia to Kamchatka (though it is possibly replaced by pealei on the coast there) and Japan. Northern populations are migratory, while those of Japan are resident. It is similar to the nominate subspecies, but the young are even darker than those of anatum.

Falco peregrinus macropus, described by Swainson in 1837, is the Australian peregrine falcon. It is found in Australia in all regions except the southwest. It is non-migratory. It is similar to brookei in appearance, but is slightly smaller and the ear region is entirely black. The feet are proportionally large.

Falco peregrinus madens, described by Ripley and Watson in 1963, is unusual in having some sexual dichromatism. If the Barbary falcon (see below) is considered a distinct species, it is sometimes placed therein. It is found in the Cape Verde Islands and is non-migratory; it is also endangered, with only six to eight pairs surviving. Males have a rufous wash on the crown, nape, ears and back; the underside is conspicuously washed pinkish-brown. Females are tinged rich brown overall, especially on the crown and nape.

 

Falco peregrinus minor, first described by Bonaparte in 1850. It was formerly often known as perconfusus. It is sparsely and patchily distributed throughout much of sub-Saharan Africa and widespread in Southern Africa. It apparently reaches north along the Atlantic coast as far as Morocco. It is non-migratory and dark-coloured. This is the smallest subspecies, with smaller males weighing as little as approximately 300 g (11 oz).

Falco peregrinus nesiotes, described by Mayr in 1941, is found in Fiji and probably also Vanuatu and New Caledonia. It is non-migratory.

Falco peregrinus pealei, described by Ridgway in 1873, is Peale's falcon and includes rudolfi. It is found in the Pacific Northwest of North America, northwards from Puget Sound along the British Columbia coast (including the Haida Gwaii), along the Gulf of Alaska and the Aleutian Islands to the far eastern Bering Sea coast of Russia, and may also occur on the Kuril Islands and the coasts of Kamchatka. It is non-migratory. It is the largest subspecies and it looks like an oversized and darker tundrius or like a strongly barred and large anatum. The bill is very wide. Juveniles occasionally have pale crowns. Males weigh 700 to 1,000 g (1.5–2.2 lb), while females weigh 1,000 to 1,500 g (2.2–3.3 lb).

Falco peregrinus pelegrinoides, first described by Temminck in 1829, is found in the Canary Islands through North Africa and the Near East to Mesopotamia. It is most similar to brookei, but is markedly paler above, with a rusty neck, and is a light buff with reduced barring below. It is smaller than the nominate subspecies; females weigh around 610 g (1.34 lb).

Falco peregrinus peregrinator, described by Sundevall in 1837, is known as the Indian peregrine falcon, black shaheen, Indian shaheen or shaheen falcon. It was formerly sometimes known as Falco atriceps or Falco shaheen. Its range includes South Asia from across the Indian subcontinent to Sri Lanka and southeastern China. In India, the shaheen falcon is reported from all states except Uttar Pradesh, mainly from rocky and hilly regions. The shaheen falcon is also reported from the Andaman and Nicobar Islands in the Bay of Bengal. It has a clutch size of 3 to 4 eggs, with the chicks fledging time of 48 days with an average nesting success of 1.32 chicks per nest. In India, apart from nesting on cliffs, it has also been recorded as nesting on man-made structures such as buildings and cellphone transmission towers.[36] A population estimate of 40 breeding pairs in Sri Lanka was made in 1996. It is non-migratory and is small and dark, with rufous underparts. In Sri Lanka this species is found to favour the higher hills, while the migrant calidus is more often seen along the coast.

Falco peregrinus peregrinus, the nominate (first-named) subspecies, described by Tunstall in 1771, breeds over much of temperate Eurasia between the tundra in the north and the Pyrenees, Mediterranean region and Alpide belt in the south. It is mainly non-migratory in Europe, but migratory in Scandinavia and Asia. Males weigh 580 to 750 g (1.28–1.65 lb), while females weigh 925 to 1,300 g (2.039–2.866 lb). It includes brevirostris, germanicus, rhenanus and riphaeus.

Falco peregrinus radama, described by Hartlaub in 1861, is found in Madagascar and the Comoros. It is non-migratory.

Falco peregrinus submelanogenys, described by Mathews in 1912, is the Southwest Australian peregrine falcon. It is found in southwestern Australia and is non-migratory.

Falco peregrinus tundrius, described by C.M. White in 1968, was at one time included in leucogenys. It is found in the Arctic tundra of North America to Greenland, and migrates to wintering grounds in Central and South America. Most vagrants that reach western Europe belong to this subspecies, which was previously considered synonymous with anatum. It is the New World equivalent to calidus. It is smaller and paler than anatum; most have a conspicuous white forehead and white in ear region, but the crown and "moustache" are very dark, unlike in calidus. Juveniles are browner and less grey than in calidus and paler, sometimes almost sandy, than in anatum. Males weigh 500 to 700 g (1.1–1.5 lb), while females weigh 800 to 1,100 g (1.8–2.4 lb). Despite its current recognition as a valid subspecies, a population genetic study of both pre-decline (i.e., museum) and recovered contemporary populations failed to distinguish genetically the anatum and tundrius subspecies.

Barbary falcon

Main article: Barbary falcon

The Barbary falcon is a subspecies of the peregrine falcon that inhabits parts of North Africa; namely, from the Canary Islands to the Arabian Peninsula. There is discussion concerning the taxonomic status of the bird, with some considering it a subspecies of the peregrine falcon and others considering it a full species with two subspecies (White et al. 2013). Compared to the other peregrine falcon subspecies, Barbary falcons sport a slimmer body and a distinct plumage color pattern. Despite numbers and range of these birds throughout the Canary Islands generally increasing, they are considered endangered, with human interference through falconry and shooting threatening their well-being. Falconry can further complicate the speciation and genetics of these Canary Islands falcons, as the practice promotes genetic mixing between individuals from outside the islands with those originating from the islands. Population density of the Barbary falcons on Tenerife, the biggest of the seven major Canary Islands, was found to be 1.27 pairs/100 km², with the mean distance between pairs being 5869 ± 3338 m. The falcons were only observed near large and natural cliffs with a mean altitude of 697.6 m. Falcons show an affinity for tall cliffs away from human-mediated establishments and presence.

 

Barbary falcons have a red neck patch, but otherwise differ in appearance from the peregrine falcon proper merely according to Gloger's rule, relating pigmentation to environmental humidity. The Barbary falcon has a peculiar way of flying, beating only the outer part of its wings like fulmars sometimes do; this also occurs in the peregrine falcon, but less often and far less pronounced. The Barbary falcon's shoulder and pelvis bones are stout by comparison with the peregrine falcon and its feet are smaller. Barbary falcons breed at different times of year than neighboring peregrine falcon subspecies, but they are capable of interbreeding. There is a 0.6–0.7% genetic distance in the peregrine falcon-Barbary falcon ("peregrinoid") complex.

 

Ecology and behaviour

The peregrine falcon lives mostly along mountain ranges, river valleys, coastlines, and increasingly in cities. In mild-winter regions, it is usually a permanent resident, and some individuals, especially adult males, will remain on the breeding territory. Only populations that breed in Arctic climates typically migrate great distances during the northern winter.

 

The peregrine falcon reaches faster speeds than any other animal on the planet when performing the stoop, which involves soaring to a great height and then diving steeply at speeds of over 320 km/h (200 mph), hitting one wing of its prey so as not to harm itself on impact. The air pressure from such a dive could possibly damage a bird's lungs, but small bony tubercles on a falcon's nostrils are theorized to guide the powerful airflow away from the nostrils, enabling the bird to breathe more easily while diving by reducing the change in air pressure. To protect their eyes, the falcons use their nictitating membranes (third eyelids) to spread tears and clear debris from their eyes while maintaining vision. The distinctive malar stripe or 'moustache', a dark area of feathers below the eyes, is thought to reduce solar glare and improve contrast sensitivity when targeting fast moving prey in bright light condition; the malar stripe has been found to be wider and more pronounced in regions of the world with greater solar radiation supporting this solar glare hypothesis. Peregrine falcons have a flicker fusion frequency of 129 Hz (cycles per second), very fast for a bird of its size, and much faster than mammals. A study testing the flight physics of an "ideal falcon" found a theoretical speed limit at 400 km/h (250 mph) for low-altitude flight and 625 km/h (388 mph) for high-altitude flight. In 2005, Ken Franklin recorded a falcon stooping at a top speed of 389 km/h (242 mph).

 

The life span of peregrine falcons in the wild is up to 19 years 9 months. Mortality in the first year is 59–70%, declining to 25–32% annually in adults. Apart from such anthropogenic threats as collision with human-made objects, the peregrine may be killed by larger hawks and owls.

 

The peregrine falcon is host to a range of parasites and pathogens. It is a vector for Avipoxvirus, Newcastle disease virus, Falconid herpesvirus 1 (and possibly other Herpesviridae), and some mycoses and bacterial infections. Endoparasites include Plasmodium relictum (usually not causing malaria in the peregrine falcon), Strigeidae trematodes, Serratospiculum amaculata (nematode), and tapeworms. Known peregrine falcon ectoparasites are chewing lice, Ceratophyllus garei (a flea), and Hippoboscidae flies (Icosta nigra, Ornithoctona erythrocephala).

 

In the Arctic Peregrine falcons chasing away small rodent predators from their nesting territory and Rough-legged Buzzards (Buteo lagopus) could use these hot spots as a nesting territory.

 

Feeding

The peregrine falcon's diet varies greatly and is adapted to available prey in different regions. However, it typically feeds on medium-sized birds such as pigeons and doves, waterfowl, gamebirds, songbirds, parrots, seabirds, and waders. Worldwide, it is estimated that between 1,500 and 2,000 bird species, or roughly a fifth of the world's bird species, are predated somewhere by these falcons.The peregrine falcon preys on the most diverse range of bird species of any raptor in North America, with over 300 species and including nearly 100 shorebirds. Its prey can range from 3 g (0.11 oz) hummingbirds (Selasphorus and Archilochus ssp.) to the 3.1 kg (6.8 lb) sandhill crane, although most prey taken by peregrines weigh between 20 g (0.71 oz) (small passerines) and 1,100 g (2.4 lb) (ducks, geese, loons, gulls, capercaillies, ptarmigans and other grouse). Smaller hawks (such as sharp-shinned hawks) and owls are regularly predated, as well as smaller falcons such as the American kestrel, merlin and, rarely, other peregrines.

 

In urban areas, where it tends to nest on tall buildings or bridges, it subsists mostly on a variety of pigeons. Among pigeons, the rock or feral pigeon comprises 80% or more of the dietary intake of peregrines. Other common city birds are also taken regularly, including mourning doves, common wood pigeons, common swifts, northern flickers, common starlings, American robins, common blackbirds, and corvids such as magpies, jays or carrion, house, and American crows. Coastal populations of the large subspecies pealei feed almost exclusively on seabirds. In the Brazilian mangrove swamp of Cubatão, a wintering falcon of the subspecies tundrius was observed successfully hunting a juvenile scarlet ibis.

 

Among mammalian prey species, bats in the genera Eptesicus, Myotis, Pipistrellus and Tadarida are the most common prey which taken at night. Though peregrines generally do not prefer terrestrial mammalian prey, in Rankin Inlet, peregrines largely take northern collared lemmings (Dicrostonyx groenlandicus) along with a few Arctic ground squirrels (Urocitellus parryii). Other small mammals including shrews, mice, rats, voles, and squirrels are more seldom taken. Peregrines occasionally take rabbits, mainly young individuals and juvenile hares. Additionally, remains of red fox kits and adult female American marten were found among prey remains. Insects and reptiles such as small snakes make up a small proportion of the diet, and salmonid fish have been taken by peregrines.

 

The peregrine falcon hunts most often at dawn and dusk, when prey are most active, but also nocturnally in cities, particularly during migration periods when hunting at night may become prevalent. Nocturnal migrants taken by peregrines include species as diverse as yellow-billed cuckoo, black-necked grebe, virginia rail, and common quail. The peregrine requires open space in order to hunt, and therefore often hunts over open water, marshes, valleys, fields, and tundra, searching for prey either from a high perch or from the air. Large congregations of migrants, especially species that gather in the open like shorebirds, can be quite attractive to a hunting peregrine. Once prey is spotted, it begins its stoop, folding back the tail and wings, with feet tucked. Prey is typically struck and captured in mid-air; the peregrine falcon strikes its prey with a clenched foot, stunning or killing it with the impact, then turns to catch it in mid-air. If its prey is too heavy to carry, a peregrine will drop it to the ground and eat it there. If they miss the initial strike, peregrines will chase their prey in a twisting flight.

 

Although previously thought rare, several cases of peregrines contour-hunting, i.e., using natural contours to surprise and ambush prey on the ground, have been reported and even rare cases of prey being pursued on foot. In addition, peregrines have been documented preying on chicks in nests, from birds such as kittiwakes. Prey is plucked before consumption. A 2016 study showed that the presence of peregrines benefits non-preferred species while at the same time causing a decline in its preferred prey. As of 2018, the fastest recorded falcon was at 242 mph (nearly 390 km/h). Researchers at the University of Groningen in the Netherlands and at Oxford University used 3D computer simulations in 2018 to show that the high speed allows peregrines to gain better maneuverability and precision in strikes.

 

Reproduction

The peregrine falcon is sexually mature at one to three years of age, but in larger populations they breed after two to three years of age. A pair mates for life and returns to the same nesting spot annually. The courtship flight includes a mix of aerial acrobatics, precise spirals, and steep dives. The male passes prey it has caught to the female in mid-air. To make this possible, the female actually flies upside-down to receive the food from the male's talons.

 

During the breeding season, the peregrine falcon is territorial; nesting pairs are usually more than 1 km (0.62 mi) apart, and often much farther, even in areas with large numbers of pairs. The distance between nests ensures sufficient food supply for pairs and their chicks. Within a breeding territory, a pair may have several nesting ledges; the number used by a pair can vary from one or two up to seven in a 16-year period.

 

The peregrine falcon nests in a scrape, normally on cliff edges. The female chooses a nest site, where she scrapes a shallow hollow in the loose soil, sand, gravel, or dead vegetation in which to lay eggs. No nest materials are added. Cliff nests are generally located under an overhang, on ledges with vegetation. South-facing sites are favoured. In some regions, as in parts of Australia and on the west coast of northern North America, large tree hollows are used for nesting. Before the demise of most European peregrines, a large population of peregrines in central and western Europe used the disused nests of other large birds. In remote, undisturbed areas such as the Arctic, steep slopes and even low rocks and mounds may be used as nest sites. In many parts of its range, peregrines now also nest regularly on tall buildings or bridges; these human-made structures used for breeding closely resemble the natural cliff ledges that the peregrine prefers for its nesting locations.

 

The pair defends the chosen nest site against other peregrines, and often against ravens, herons, and gulls, and if ground-nesting, also such mammals as foxes, wolverines, felids, bears, wolves, and mountain lions. Both nests and (less frequently) adults are predated by larger-bodied raptorial birds like eagles, large owls, or gyrfalcons. The most serious predators of peregrine nests in North America and Europe are the great horned owl and the Eurasian eagle-owl. When reintroductions have been attempted for peregrines, the most serious impediments were these two species of owls routinely picking off nestlings, fledglings and adults by night. Peregrines defending their nests have managed to kill raptors as large as golden eagles and bald eagles (both of which they normally avoid as potential predators) that have come too close to the nest by ambushing them in a full stoop. In one instance, when a snowy owl killed a newly fledged peregrine, the larger owl was in turn killed by a stooping peregrine parent.

 

The date of egg-laying varies according to locality, but is generally from February to March in the Northern Hemisphere, and from July to August in the Southern Hemisphere, although the Australian subspecies macropus may breed as late as November, and equatorial populations may nest anytime between June and December. If the eggs are lost early in the nesting season, the female usually lays another clutch, although this is extremely rare in the Arctic due to the short summer season. Generally three to four eggs, but sometimes as few as one or as many as five, are laid in the scrape. The eggs are white to buff with red or brown markings. They are incubated for 29 to 33 days, mainly by the female, with the male also helping with the incubation of the eggs during the day, but only the female incubating them at night. The average number of young found in nests is 2.5, and the average number that fledge is about 1.5, due to the occasional production of infertile eggs and various natural losses of nestlings.

 

After hatching, the chicks (called "eyases") are covered with creamy-white down and have disproportionately large feet. The male (called the "tiercel") and the female (simply called the "falcon") both leave the nest to gather prey to feed the young. The hunting territory of the parents can extend a radius of 19 to 24 km (12 to 15 mi) from the nest site. Chicks fledge 42 to 46 days after hatching, and remain dependent on their parents for up to two months.

 

Relationship with humans

The peregrine falcon is a highly admired falconry bird, and has been used in falconry for more than 3,000 years, beginning with nomads in central Asia. Its advantages in falconry include not only its athleticism and eagerness to hunt, but an equable disposition that leads to it being one of the easier falcons to train. The peregrine falcon has the additional advantage of a natural flight style of circling above the falconer ("waiting on") for game to be flushed, and then performing an effective and exciting high-speed diving stoop to take the quarry. The speed of the stoop not only allows the falcon to catch fast flying birds, it also enhances the falcon's ability to execute maneuvers to catch highly agile prey, and allows the falcon to deliver a knockout blow with a fist-like clenched talon against game that may be much larger than itself.

 

Additionally the versatility of the species, with agility allowing capture of smaller birds and a strength and attacking style allowing capture of game much larger than themselves, combined with the wide size range of the many peregrine subspecies, means there is a subspecies suitable to almost any size and type of game bird. This size range, evolved to fit various environments and prey species, is from the larger females of the largest subspecies to the smaller males of the smallest subspecies, approximately five to one (approximately 1500 g to 300 g). The males of smaller and medium-sized subspecies, and the females of the smaller subspecies, excel in the taking of swift and agile small game birds such as dove, quail, and smaller ducks. The females of the larger subspecies are capable of taking large and powerful game birds such as the largest of duck species, pheasant, and grouse.

 

Peregrine falcons handled by falconers are also occasionally used to scare away birds at airports to reduce the risk of bird-plane strikes, improving air-traffic safety. They were also used to intercept homing pigeons during World War II.

 

Peregrine falcons have been successfully bred in captivity, both for falconry and for release into the wild. Until 2004 nearly all peregrines used for falconry in the US were captive-bred from the progeny of falcons taken before the US Endangered Species Act was enacted and from those few infusions of wild genes available from Canada and special circumstances. Peregrine falcons were removed from the United States' endangered species list in 1999. The successful recovery program was aided by the effort and knowledge of falconers – in collaboration with The Peregrine Fund and state and federal agencies – through a technique called hacking. Finally, after years of close work with the US Fish and Wildlife Service, a limited take of wild peregrines was allowed in 2004, the first wild peregrines taken specifically for falconry in over 30 years.

 

The development of captive breeding methods has led to peregrines being commercially available for falconry use, thus mostly eliminating the need to capture wild birds for support of falconry. The main reason for taking wild peregrines at this point is to maintain healthy genetic diversity in the breeding lines. Hybrids of peregrines and gyrfalcons are also available that can combine the best features of both species to create what many consider to be the ultimate falconry bird for the taking of larger game such as the sage-grouse. These hybrids combine the greater size, strength, and horizontal speed of the gyrfalcon with the natural propensity to stoop and greater warm weather tolerance of the peregrine.

 

Decline due to pesticides

The peregrine falcon became an endangered species over much of its range because of the use of organochlorine pesticides, especially DDT, during the 1950s, '60s, and '70s. Pesticide biomagnification caused organochlorine to build up in the falcons' fat tissues, reducing the amount of calcium in their eggshells. With thinner shells, fewer falcon eggs survived until hatching. In addition, the PCB concentrations found in these falcons is dependent upon the age of the falcon. While high levels are still found in young birds (only a few months old) and even higher concentrations are found in more mature falcons, further increasing in adult peregrine falcons. These pesticides caused falcon prey to also have thinner eggshells (one example of prey being the Black Petrels). In several parts of the world, such as the eastern United States and Belgium, this species became extirpated (locally extinct) as a result. An alternate point of view is that populations in the eastern North America had vanished due to hunting and egg collection. Following the ban of organochlorine pesticides, the reproductive success of Peregrines increased in Scotland in terms of territory occupancy and breeding success, although spatial variation in recovery rates indicate that in some areas Peregrines were also impacted by other factors such as persecution.

 

Recovery efforts

Peregrine falcon recovery teams breed the species in captivity. The chicks are usually fed through a chute or with a hand puppet mimicking a peregrine's head, so they cannot see to imprint on the human trainers. Then, when they are old enough, the rearing box is opened, allowing the bird to train its wings. As the fledgling gets stronger, feeding is reduced, forcing the bird to learn to hunt. This procedure is called hacking back to the wild. To release a captive-bred falcon, the bird is placed in a special cage at the top of a tower or cliff ledge for some days or so, allowing it to acclimate itself to its future environment.

 

Worldwide recovery efforts have been remarkably successful. The widespread restriction of DDT use eventually allowed released birds to breed successfully. The peregrine falcon was removed from the U.S. Endangered Species list on 25 August 1999.

 

Some controversy has existed over the origins of captive breeding stock used by the Peregrine Fund in the recovery of peregrine falcons throughout the contiguous United States. Several peregrine subspecies were included in the breeding stock, including birds of Eurasian origin. Due to the extirpation of the eastern population of Falco peregrinus anatum, the near-extirpation of anatum in the Midwest and the limited gene pool within North American breeding stock, the inclusion of non-native subspecies was justified to optimize the genetic diversity found within the species as a whole.

 

During the 1970s, peregrine falcons in Finland experienced a population bottleneck as a result of large declines associated with bio-accumulation of organochloride pesticides. However, the genetic diversity of peregrines in Finland is similar to other populations, indicating that high dispersal rates have maintained the genetic diversity of this species.

 

Since peregrine falcon eggs and chicks are still often targeted by illegal poachers, it is common practice not to publicize unprotected nest locations.

 

Current status

Populations of the peregrine falcon have bounced back in most parts of the world. In the United Kingdom, there has been a recovery of populations since the crash of the 1960s. This has been greatly assisted by conservation and protection work led by the Royal Society for the Protection of Birds. The RSPB estimated that there were 1,402 breeding pairs in the UK in 2011. In Canada, where peregrines were identified as endangered in 1978 (in the Yukon territory of northern Canada that year, only a single breeding pair was identified), the Committee on the Status of Endangered Wildlife in Canada declared the species no longer at risk in December 2017.

 

Peregrines now breed in many mountainous and coastal areas, especially in the west and north, and nest in some urban areas, capitalising on the urban feral pigeon populations for food. Additionally, falcons benefit from artificial illumination, which allows the raptors to extend their hunting periods into the dusk when natural illumination would otherwise be too low for them to pursue prey. In England, this has allowed them to prey on nocturnal migrants such as redwings, fieldfares, starlings, and woodcocks.

 

In many parts of the world peregrine falcons have adapted to urban habitats, nesting on cathedrals, skyscraper window ledges, tower blocks, and the towers of suspension bridges. Many of these nesting birds are encouraged, sometimes gathering media attention and often monitored by cameras.

 

In England, peregrine falcons have become increasingly urban in distribution, particularly in southern areas where inland cliffs suitable as nesting sites are scarce. The first recorded urban breeding pair was observed nesting on the Swansea Guildhall in the 1980s. In Southampton, a nest prevented restoration of mobile telephony services for several months in 2013, after Vodafone engineers despatched to repair a faulty transmitter mast discovered a nest in the mast, and were prevented by the Wildlife and Countryside Act – on pain of a possible prison sentence – from proceeding with repairs until the chicks fledged.

 

In Oregon, Portland houses ten percent of the state's peregrine nests, despite only covering around 0.1 percent of the state's land area.

 

Cultural significance

Due to its striking hunting technique, the peregrine has often been associated with aggression and martial prowess. The Ancient Egyptian solar deity Ra was often represented as a man with the head of a peregrine falcon adorned with the solar disk, although most Egyptologists agree that it's most likely a Lanner falcon. Native Americans of the Mississippian culture (c. 800–1500) used the peregrine, along with several other birds of prey, in imagery as a symbol of "aerial (celestial) power" and buried men of high status in costumes associating to the ferocity of raptorial birds. In the late Middle Ages, the Western European nobility that used peregrines for hunting, considered the bird associated with princes in formal hierarchies of birds of prey, just below the gyrfalcon associated with kings. It was considered "a royal bird, more armed by its courage than its claws". Terminology used by peregrine breeders also used the Old French term gentil, "of noble birth; aristocratic", particularly with the peregrine.

 

The peregrine falcon is the national animal of the United Arab Emirates. Since 1927, the peregrine falcon has been the official mascot of Bowling Green State University in Bowling Green, Ohio. The 2007 U.S. Idaho state quarter features a peregrine falcon. The peregrine falcon has been designated the official city bird of Chicago.

 

The Peregrine, by J. A. Baker, is widely regarded as one of the best nature books in English written in the twentieth century. Admirers of the book include Robert Macfarlane, Mark Cocker, who regards the book as "one of the most outstanding books on nature in the twentieth century" and Werner Herzog, who called it "the one book I would ask you to read if you want to make films", and said elsewhere "it has prose of the calibre that we have not seen since Joseph Conrad". In the book, Baker recounts, in diary form, his detailed observations of peregrines (and their interaction with other birds) near his home in Chelmsford, Essex, over a single winter from October to April.

 

An episode of the hour-long TV series Starman in 1986 titled "Peregrine" was about an injured peregrine falcon and the endangered species program. It was filmed with the assistance of the University of California's peregrine falcon project in Santa Cruz.

About Tpad (www.tpad.com)

 

Tpad is an international telecommunications company with over 1000 customers in the UK and operating internationally from regional offices in Dubai and Cyprus. With over 20 years experience in the IT Sector, our expertise spans the development, manufacture and installation of business telephony services. Tpad offers solutions to meet all requirements ranging from Managed IP/PBX solutions to complex Contact Centre installations. The Tpad Global Phone Network that has unlimited call handling capacity and our in-house developed software provides an on-net feature rich environment with functionality that includes network based call recording, call reporting, least cost routing, Diallers, Multi-Site solutions and much more.

 

With over 100 operatives within dedicated Service, Support and Development teams, we pride ourselves on service excellence and after sales care, consistently delivering some of the highest level of support within the industry. Our services are backed by a choice of service levels to dovetail with the skill sets within your organisation.

 

The Tpad Platform and solutions are able to operate using third party connectivity. However, Tpad is able to offer a range of connectivity solutions by partnering with its sister company, Internexus. With over 12 years in the sector, that includes the provision and management of network solutions for Supanet, the largest independent Internet Service Provider in the UK with over 500 thousand customers using the a range of services from Internet access, line rental and carrier pre-select services, Tpad and Internexus boast experience and knowledge that almost is unparalleled within the industry. Indeed, it was this combined experience that led to the establishment of Tpad's On-Net Voice Platform in 2005, when users were first given the ability to log in and make phone calls over the Internet. Whilst we have come a long way since then, adding the Tpad Platform to our broad range of business IP/PBX equipment powered by industry leading connectivity make our systems a truly one-stop solution.

 

Tpad's exciting product range is constantly evolving to keep pace both with technological advancements and our commitment offering our customers new and improved features and enhanced capabilities. At the same time we remain true to our core values of providing reliable cutting edge technology with outstanding service delivery. Telecommunications are the foundations on which almost every business in the 21st century is built. Information is power and communication is king. Whatever your requirements Tpad is ideally positioned to deliver a flexible cost-effective solution to satisfy your organisation's telecommunications requirements both for today and in the years to come. Let Tpad be a valuable partner in your business.

 

Natalia Ortiz aquí.

 

Abreviación para celular en español:

 

A mobile phone or mobile (also called cellphone and handphone) is an electronic device used for mobile telecommunications (mobile telephony, text messaging or data transmission) over a cellular network of specialized base stations known as cell sites. Mobile phones differ from cordless telephones, which only offer telephony service within a limited range, e.g. within a home or an office, through a fixed line and a base station owned by the subscriber and also from satellite phones and radio telephones. As opposed to a radio telephone, a cell phone offers full duplex communication, automatised calling to and paging from a public land mobile network (PLMN), and handoff (handover) during a phone call when the user moves from one cell (base station coverage area) to another. Most current cell phones connect to a cellular network consisting of switching points and base stations (cell sites) owned by a mobile network operator. In addition to the standard voice function, current mobile phones may support many additional services, and accessories, such as SMS for text messaging, email, packet switching for access to the Internet, gaming, Bluetooth, infrared, camera with video recorder and MMS for sending and receiving photos and video, MP3 player, radio and GPS.

The International Telecommunication Union estimated that mobile cellular subscriptions worldwide would reach approximately 4.6 billion by the end of 2009. Mobile phones have gained increased importance in the sector of Information and communication technologies for development in the 2000s and have effectively started to reach the bottom of the economic pyramid.

 

Cel, a transparent sheet on which objects are drawn or painted for animation purposes

 

Cel programming language

  

Entrance door to the previous headquarters (1923) of the Amsterdam telephony service, artwork made by Hildo Krop, supposedly depicting the 4 continents and also the Staff of Mercury, the old Roman god ruling all affairs concerning Communication.

About Tpad (http://www.tpad.com)

Tpad is an International company which has its Headquarters in Jersey, and provides products and services in many countries including the UAE, India, Hong Kong, USA, UK and Canada. In the UK, Tpad provides advanced telephony services and carrier-grade SIP trunks to businesses worldwide. Until recently the core product has been large IP PBX based business telephone systems (50 users to 500+ users and multiple sites). Tpad also has an International consumer arm providing low cost international services to all major countries. Over the past 3 years, Tpad has been investing in next generation telephone technology with the development of a powerful central telephone platform called Colossus that resides in Tpad data centres and that can be accessed from anywhere in the cloud space.

 

T24, a mobile service brand of Tata Teleservices and Future Group, has now expanded its services in Kerala, Madhya Pradesh, Rajasthan and Punjab.

 

T24 offers free talk time on shopping in Future Bazaar retail outlets such as Big Bazaar, Pantaloons, Food Bazaar, Pantaloons, Central, Brand Factory, Home Town, eZone and Aadhar, among others. It also offers discount vouchers to those buying talktime.

 

With this, the brand has reached 17 circles including Andhra Pradesh, Karnataka, Gujarat, West Bengal, Jharkhand, Orissa, Punjab, Haryana and Uttar Pradesh. Andhra Pradesh was the first telecom circle to get the services in June 2010.

 

"T24 has received tremendous response since its launch in Andhra Pradesh, Karnataka, Gujarat, West Bengal, Jharkhand, Orissa, Punjab, Haryana and Uttar Pradesh. Shopping, and talking on our mobile phones are among the two favorite activities for all of us in India," said Mayur Toshniwal, chief executive officer of T24.

 

"With T24, we have been able to develop a unique customer value proposition that combines these interests of the aspirational Indian. Customers will get shopping benefits on talking and talk-time benefits every time they shop," Toshniwal added.

 

T24 tariff plans are similar to those that Tata Docomo and Indicom offer. In addition, customers get rewarded free talk time for every purchase above Rs 350 made at any Future Group shopping outlet.

 

For example, a T24 customer buying products worth Rs 3,001 at Pantaloons or Central gets Rs 150 worth of free talk time. And a customer spending Rs 2,001 at Big Bazaar gets Rs 84 worth of free talk time.

 

According to the company, "T24 mobile telephony service will be progressively rolled out across the country in next few months, through Future Group's retail networks in over 76 cities and 65 rural destinations."

 

Customers can recharge their T24 mobile at these stores and also at any Tata Docomo retailer. Future Group retail chains attracted over 250 million customer footfalls in 2009-10.

 

The service has so far not been started in Delhi where Future Group has a sizeable presence as Tata Docomo doesn't have spectrum for the circle yet, and the company probably doesn't want to start service using its CDMA network.

On the first day of his visit, Congress General Secretary Rahul Gandhi put Sarvan village near Fursatganj in the annals of history by launching a broadband service named as “Fibre to panchayat” technically known as GPON technology. With having state-of-the-art facility, Sarvan village became the first village in India. One optic fibre enables users to make use of this facility to enjoy services like broadband, internet, telephonic service and cable-tv as well. The distribution of optic fibre in village panchayat will be done by India unit of US-based Alphion Corporation Ltd in collaboration with ITI Rae Bareli and BSNL. On this inauguration occasion, he shared the stage with Sam Pitroda, the pioneer behind the information technology revolution.

www.pressbrief.in

• Chris Kelly MP ‘cuts the ribbon’

• Modern open plan environment

• Opening hours extended by more 15 hours a week

• Full range of Post Office® services available

 

To celebrate the launch of the new modern branch Dudley South MP, Chris Kelly, ‘cut the ribbon’ at an opening ceremony at Merry Hill Centre Post Office, today Friday 12 July 2013.

 

Chris Kelly MP said: “I’m delighted to officially open the new Merry Hill Centre Post Office. Nick and his team are to be congratulated on their work to develop the new branch which will make a real difference for local residents and small business customers. I wish them every success with the venture.”

 

The new Merry Hill Centre Post Office has six serving positions, a mixture of two screened and three open plan counter positions, with a Post Office serving point provided for use at the retail counter – a new concept for delivering Post Office services alongside retail transactions during shop opening hours.

 

Opening hours at the new location have been extended by 15 hours a week to Monday to Saturday; 09:00 – 19:00 and Sunday: 11.00 – 17.00 offering customers access Post Office products and services 7 days a week.

 

Subpostmaster Nick Singh, who hosted today’s official opening, said: “I was delighted to have the opportunity to modernise the Post Office and very pleased to be re-launching it today for the community. I know customers will like the new environment and I’d like to thank Mr Kelly for coming today.”

 

Post Office Regional Manager, Richard Lynds, said: “We understand how important having a Post Office is to residents in this area and we are confident that this new modern Post Office service will meet the needs of the local community and secure services for the future.”

 

The branch relocated within the Merry Hill Centre in June 2013.

 

The Post Office, now independent of Royal Mail, is the largest retail network in the UK, with over 11,500 branches. It is also one of the fastest growing financial services companies and is developing its online and telephony services.

 

Pictured: Chris Kelly MP (Dudley South) and Nick Singh (Postmaster)

 

ENDS

 

Further details:

 

Post Office External Relations,

1st Floor, 148 Old Street,

LONDON

EC1V 9HQ

Tel: 020 7012 3456 (24 hours),

email: pressoffice@postoffice.co.uk

www.postoffice.co.uk

 

dudleysouth.com

dudleyconservatives.com

On the first day of his visit, Congress General Secretary Rahul Gandhi put Sarvan village near Fursatganj in the annals of history by launching a broadband service named as “Fibre to panchayat” technically known as GPON technology. With having state-of-the-art facility, Sarvan village became the first village in India. One optic fibre enables users to make use of this facility to enjoy services like broadband, internet, telephonic service and cable-tv as well. The distribution of optic fibre in village panchayat will be done by India unit of US-based Alphion Corporation Ltd in collaboration with ITI Rae Bareli and BSNL. On this inauguration occasion, he shared the stage with Sam Pitroda, the pioneer behind the information technology revolution.

www.pressbrief.in

Lincoln Center, more commonly known as Lincoln, is a city in and the county seat of Lincoln County, Kansas, United States. As of the 2010 census, the city population was 1,297.

 

Settler George Green founded the town of Lincoln in 1870, naming it after the county. "Center" was added to its name to indicate its location in the central part of the county. County residents initially voted to place the county seat in Lincoln Center. However, elections in November 1870 resulted in the seat relocating to nearby Abram, sparking a county seat war. Another vote in February 1872 reversed the switch, and Lincoln Center became the permanent county seat. In addition, residents relocated all the buildings in Abram to Lincoln, merging the two communities. The town was incorporated as the city of Lincoln Center in 1879, and is still officially referred to as such in the United States Census. However, both the city government and local chamber of commerce refer to the city as Lincoln, and it appears as such on state maps and signs.

 

Lincoln grew rapidly over the following decade, more than doubling in size, and the city gradually modernized. Kansas Christian College opened in Lincoln in 1885. The Union Pacific Railroad opened a branch line through the city in 1886, bringing mail service. In 1900, after the previous courthouse burned down, a new one was built out of native limestone known locally as "post rock". Rural mail delivery began in 1904, and telephony service began in 1905. In 1906, the city's first power plant opened, providing electric power.

 

In 1989, given the widespread, long-standing use of post rock limestone in local building construction, the Kansas Legislature named Lincoln County as "The Post Rock Capital of Kansas".

 

en.wikipedia.org/wiki/Lincoln_Center,_Kansas

 

en.wikipedia.org/wiki/Wikipedia:Text_of_Creative_Commons_...

On the first day of his visit, Congress General Secretary Rahul Gandhi put Sarvan village near Fursatganj in the annals of history by launching a broadband service named as “Fibre to panchayat” technically known as GPON technology. With having state-of-the-art facility, Sarvan village became the first village in India. One optic fibre enables users to make use of this facility to enjoy services like broadband, internet, telephonic service and cable-tv as well. The distribution of optic fibre in village panchayat will be done by India unit of US-based Alphion Corporation Ltd in collaboration with ITI Rae Bareli and BSNL. On this inauguration occasion, he shared the stage with Sam Pitroda, the pioneer behind the information technology revolution.

www.pressbrief.in

On the first day of his visit, Congress General Secretary Rahul Gandhi put Sarvan village near Fursatganj in the annals of history by launching a broadband service named as “Fibre to panchayat” technically known as GPON technology. With having state-of-the-art facility, Sarvan village became the first village in India. One optic fibre enables users to make use of this facility to enjoy services like broadband, internet, telephonic service and cable-tv as well. The distribution of optic fibre in village panchayat will be done by India unit of US-based Alphion Corporation Ltd in collaboration with ITI Rae Bareli and BSNL. On this inauguration occasion, he shared the stage with Sam Pitroda, the pioneer behind the information technology revolution.

www.pressbrief.in

On the first day of his visit, Congress General Secretary Rahul Gandhi put Sarvan village near Fursatganj in the annals of history by launching a broadband service named as “Fibre to panchayat” technically known as GPON technology. With having state-of-the-art facility, Sarvan village became the first village in India. One optic fibre enables users to make use of this facility to enjoy services like broadband, internet, telephonic service and cable-tv as well. The distribution of optic fibre in village panchayat will be done by India unit of US-based Alphion Corporation Ltd in collaboration with ITI Rae Bareli and BSNL. On this inauguration occasion, he shared the stage with Sam Pitroda, the pioneer behind the information technology revolution.

www.pressbrief.in

Asterisk is one of the advances technology for telephonic services that offers additionally created and business driven office phone system to endeavors. It grants to join telephones and to make calls to one another. Another reason behind Asterisk Solution prevalence is untouchable comparability with various working systems, for instance, NetBSD, OpenBSD, FreeBSD, Mac OS X, Solaris, etc. Asterisks give a broad assortment of business solutions and services.

On the first day of his visit, Congress General Secretary Rahul Gandhi put Sarvan village near Fursatganj in the annals of history by launching a broadband service named as “Fibre to panchayat” technically known as GPON technology. With having state-of-the-art facility, Sarvan village became the first village in India. One optic fibre enables users to make use of this facility to enjoy services like broadband, internet, telephonic service and cable-tv as well. The distribution of optic fibre in village panchayat will be done by India unit of US-based Alphion Corporation Ltd in collaboration with ITI Rae Bareli and BSNL. On this inauguration occasion, he shared the stage with Sam Pitroda, the pioneer behind the information technology revolution.

www.pressbrief.in

On the first day of his visit, Congress General Secretary Rahul Gandhi put Sarvan village near Fursatganj in the annals of history by launching a broadband service named as “Fibre to panchayat” technically known as GPON technology. With having state-of-the-art facility, Sarvan village became the first village in India. One optic fibre enables users to make use of this facility to enjoy services like broadband, internet, telephonic service and cable-tv as well. The distribution of optic fibre in village panchayat will be done by India unit of US-based Alphion Corporation Ltd in collaboration with ITI Rae Bareli and BSNL. On this inauguration occasion, he shared the stage with Sam Pitroda, the pioneer behind the information technology revolution.

www.pressbrief.in

Net Root Integrated Services Ltd. began provide services in October 2006. It is developed with fast rhythms in the sector of computerization of enterprises and technical support. Collaborating with the bigger companies of products of information technology, it provides services of high quality fast, responsibly and with safety.

 

Services

 

Trading of Computer Equipment

Computerization of Enterprises

Installations and Maintenance of Networks

Parcels of Technical Support

Repairs – Upgrades PC

IP Telephony Services

Web Design

Software Applications

 

48 Boumpoulinas str. , Petroupolis , 132 31, Attica - Greece

 

Tel. +30 - 210 5054358 , Fax. +30 - 210 5054648

 

Web: www.netroot.gr , e-mail: info@netroot.gr

On the first day of his visit, Congress General Secretary Rahul Gandhi put Sarvan village near Fursatganj in the annals of history by launching a broadband service named as “Fibre to panchayat” technically known as GPON technology. With having state-of-the-art facility, Sarvan village became the first village in India. One optic fibre enables users to make use of this facility to enjoy services like broadband, internet, telephonic service and cable-tv as well. The distribution of optic fibre in village panchayat will be done by India unit of US-based Alphion Corporation Ltd in collaboration with ITI Rae Bareli and BSNL. On this inauguration occasion, he shared the stage with Sam Pitroda, the pioneer behind the information technology revolution.

www.pressbrief.in

On the first day of his visit, Congress General Secretary Rahul Gandhi put Sarvan village near Fursatganj in the annals of history by launching a broadband service named as “Fibre to panchayat” technically known as GPON technology. With having state-of-the-art facility, Sarvan village became the first village in India. One optic fibre enables users to make use of this facility to enjoy services like broadband, internet, telephonic service and cable-tv as well. The distribution of optic fibre in village panchayat will be done by India unit of US-based Alphion Corporation Ltd in collaboration with ITI Rae Bareli and BSNL. On this inauguration occasion, he shared the stage with Sam Pitroda, the pioneer behind the information technology revolution.

www.pressbrief.in

On the first day of his visit, Congress General Secretary Rahul Gandhi put Sarvan village near Fursatganj in the annals of history by launching a broadband service named as “Fibre to panchayat” technically known as GPON technology. With having state-of-the-art facility, Sarvan village became the first village in India. One optic fibre enables users to make use of this facility to enjoy services like broadband, internet, telephonic service and cable-tv as well. The distribution of optic fibre in village panchayat will be done by India unit of US-based Alphion Corporation Ltd in collaboration with ITI Rae Bareli and BSNL. On this inauguration occasion, he shared the stage with Sam Pitroda, the pioneer behind the information technology revolution.

www.pressbrief.in

On the first day of his visit, Congress General Secretary Rahul Gandhi put Sarvan village near Fursatganj in the annals of history by launching a broadband service named as “Fibre to panchayat” technically known as GPON technology. With having state-of-the-art facility, Sarvan village became the first village in India. One optic fibre enables users to make use of this facility to enjoy services like broadband, internet, telephonic service and cable-tv as well. The distribution of optic fibre in village panchayat will be done by India unit of US-based Alphion Corporation Ltd in collaboration with ITI Rae Bareli and BSNL. On this inauguration occasion, he shared the stage with Sam Pitroda, the pioneer behind the information technology revolution.

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On the first day of his visit, Congress General Secretary Rahul Gandhi put Sarvan village near Fursatganj in the annals of history by launching a broadband service named as “Fibre to panchayat” technically known as GPON technology. With having state-of-the-art facility, Sarvan village became the first village in India. One optic fibre enables users to make use of this facility to enjoy services like broadband, internet, telephonic service and cable-tv as well. The distribution of optic fibre in village panchayat will be done by India unit of US-based Alphion Corporation Ltd in collaboration with ITI Rae Bareli and BSNL. On this inauguration occasion, he shared the stage with Sam Pitroda, the pioneer behind the information technology revolution.

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The peregrine falcon (Falco peregrinus), also known simply as the peregrine, and historically as the duck hawk in North America, is a cosmopolitan bird of prey (raptor) in the family Falconidae. A large, crow-sized falcon, it has a blue-grey back, barred white underparts, and a black head. The peregrine is renowned for its speed. It can reach over 320 km/h (200 mph) during its characteristic hunting stoop (high-speed dive), making it the fastest member of the animal kingdom. According to a National Geographic TV program, the highest measured speed of a peregrine falcon is 389 km/h (242 mph). As is typical for bird-eating (avivore) raptors, peregrine falcons are sexually dimorphic, with females being considerably larger than males.

 

The peregrine's breeding range includes land regions from the Arctic tundra to the tropics. It can be found nearly everywhere on Earth, except extreme polar regions, very high mountains, and most tropical rainforests; the only major ice-free landmass from which it is entirely absent is New Zealand. This makes it the world's most widespread raptor and one of the most widely found wild bird species. In fact, the only land-based bird species found over a larger geographic area owes its success to human-led introduction; the domestic and feral pigeons are both domesticate forms of the rock dove, which are a major prey species for Eurasian Peregrine populations. Due to their prevalence over most other bird species in cities, feral pigeons support many peregrine populations as a staple food source, especially in urban settings.

 

The peregrine is a highly successful example of urban wildlife in much of its range, taking advantage of tall buildings as nest sites and an abundance of prey such as pigeons and ducks. Both the English and scientific names of this species mean "wandering falcon", referring to the migratory habits of many northern populations. Experts recognize 17 to 19 subspecies, which vary in appearance and range; disagreement exists over whether the distinctive Barbary falcon is represented by two subspecies of Falco peregrinus or is a separate species, F. pelegrinoides. The two species' divergence is relatively recent, during the time of the last ice age, therefore the genetic differential between them (and also the difference in their appearance) is relatively tiny. They are only about 0.6–0.8% genetically differentiated.

 

Although its diet consists almost exclusively of medium-sized birds, the peregrine will sometimes hunt small mammals, small reptiles, or even insects. Reaching sexual maturity at one year, it mates for life and nests in a scrape, normally on cliff edges or, in recent times, on tall human-made structures. The peregrine falcon became an endangered species in many areas because of the widespread use of certain pesticides, especially DDT. Since the ban on DDT from the early 1970s, populations have recovered, supported by large-scale protection of nesting places and releases to the wild.

 

The peregrine falcon is a well-respected falconry bird due to its strong hunting ability, high trainability, versatility, and availability via captive breeding. It is effective on most game bird species, from small to large. It has also been used as a religious, royal, or national symbol across multiple eras and areas of human civilization.

 

Description

 

Falco peregrinus. Royal National Park, New South Wales, Australia

The peregrine falcon has a body length of 34 to 58 cm (13–23 in) and a wingspan from 74 to 120 cm (29–47 in). The male and female have similar markings and plumage but, as with many birds of prey, the peregrine falcon displays marked sexual dimorphism in size, with the female measuring up to 30% larger than the male. Males weigh 330 to 1,000 g (12–35 oz) and the noticeably larger females weigh 700 to 1,500 g (25–53 oz). In most subspecies, males weigh less than 700 g (25 oz) and females weigh more than 800 g (28 oz), and cases of females weighing about 50% more than their male breeding mates are not uncommon. The standard linear measurements of peregrines are: the wing chord measures 26.5 to 39 cm (10.4–15.4 in), the tail measures 13 to 19 cm (5.1–7.5 in) and the tarsus measures 4.5 to 5.6 cm (1.8–2.2 in).

 

The back and the long pointed wings of the adult are usually bluish black to slate grey with indistinct darker barring (see "Subspecies" below); the wingtips are black. The white to rusty underparts are barred with thin clean bands of dark brown or black. The tail, coloured like the back but with thin clean bars, is long, narrow, and rounded at the end with a black tip and a white band at the very end. The top of the head and a "moustache" along the cheeks are black, contrasting sharply with the pale sides of the neck and white throat. The cere is yellow, as are the feet, and the beak and claws are black. The upper beak is notched near the tip, an adaptation which enables falcons to kill prey by severing the spinal column at the neck. An immature bird is much browner, with streaked, rather than barred, underparts, and has a pale bluish cere and orbital ring.

 

A study shows that their black malar stripe exists to reduce glare from solar radiation, allowing them to see better. Photos from The Macaulay Library and iNaturalist showed that the malar stripe is thicker where there is more solar radiation. That supports the solar glare hypothesis.

 

Taxonomy and systematics

 

Falco peregrinus was first described under its current binomial name by English ornithologist Marmaduke Tunstall in his 1771 work Ornithologia Britannica. The scientific name Falco peregrinus is a Medieval Latin phrase that was used by Albertus Magnus in 1225. Peregrinus is Latin, meaning "one from abroad" or "coming from foreign parts". It is likely the name was used as juvenile birds were taken while journeying to their breeding location (rather than from the nest), as falcon nests are often difficult to get at. The Latin term for falcon, falco, is related to falx, meaning "sickle", in reference to the silhouette of the falcon's long, pointed wings in flight.

 

The peregrine falcon belongs to a genus whose lineage includes the hierofalcon and the prairie falcon (F. mexicanus). This lineage probably diverged from other falcons towards the end of the Late Miocene or in the Late Pliocene, about 3–8 million years ago (mya). As the peregrine-hierofalcon group includes both Old World and North American species, it is likely that the lineage originated in western Eurasia or Africa. Its relationship to other falcons is not clear, as the issue is complicated by widespread hybridization confounding mtDNA sequence analyses. One genetic lineage of the saker falcon (F. cherrug) is known to have originated from a male saker ancestor producing fertile young with a female peregrine ancestor, and the descendants further breeding with sakers.

 

Today, peregrines are regularly paired in captivity with other species such as the lanner falcon (F. biarmicus) to produce the "perilanner", a somewhat popular bird in falconry as it combines the peregrine's hunting skill with the lanner's hardiness, or the gyrfalcon to produce large, strikingly coloured birds for the use of falconers. As can be seen, the peregrine is still genetically close to the hierofalcons, though their lineages diverged in the Late Pliocene (maybe some 2.5–2 mya in the Gelasian).

 

Subspecies

Numerous subspecies of Falco peregrinus have been described, with 19 accepted by the 1994 Handbook of the Birds of the World, which considers the Barbary falcon of the Canary Islands and coastal North Africa to be two subspecies (pelegrinoides and babylonicus) of Falco peregrinus, rather than a distinct species, F. pelegrinoides. The following map shows the general ranges of these 19 subspecies.

 

A map of the world, green shows on several continents, but there are also several big bare spots marked with E for extinct

Breeding ranges of the 19 subspecies

 

Falco peregrinus anatum, described by Bonaparte in 1838, is known as the American peregrine falcon or "duck hawk"; its scientific name means "duck peregrine falcon". At one time, it was partly included in leucogenys. It is mainly found in the Rocky Mountains. It was formerly common throughout North America between the tundra and northern Mexico, where current reintroduction efforts are being made to restore the population. Most mature anatum, except those that breed in more northern areas, winter in their breeding range. Most vagrants that reach western Europe seem to belong to the more northern and strongly migratory tundrius, only considered distinct since 1968. It is similar to the nominate subspecies but is slightly smaller; adults are somewhat paler and less patterned below, but juveniles are darker and more patterned below. Males weigh 500 to 700 g (1.1–1.5 lb), while females weigh 800 to 1,100 g (1.8–2.4 lb). It has become extinct in eastern North America and populations there are hybrids as a result of reintroductions of birds from elsewhere.

Falco peregrinus babylonicus, described by P.L. Sclater in 1861, is found in eastern Iran along the Hindu Kush and the Tian Shan to the Mongolian Altai ranges. A few birds winter in northern and northwestern India, mainly in dry semi-desert habitats. It is paler than pelegrinoides and somewhat similar to a small, pale lanner falcon (Falco biarmicus). Males weigh 330 to 400 grams (12 to 14 oz), while females weigh 513 to 765 grams (18.1 to 27.0 oz).

Falco peregrinus brookei, described by Sharpe in 1873, is also known as the Mediterranean peregrine falcon or the Maltese falcon. It includes caucasicus and most specimens of the proposed race punicus, though others may be pelegrinoides (Barbary falcons), or perhaps the rare hybrids between these two which might occur around Algeria. They occur from the Iberian Peninsula around the Mediterranean, except in arid regions, to the Caucasus. They are non-migratory. It is smaller than the nominate subspecies and the underside usually has a rusty hue. Males weigh around 445 g (0.981 lb), while females weigh up to 920 g (2.03 lb).

Falco peregrinus calidus, described by John Latham in 1790, it was formerly called leucogenys and includes caeruleiceps. It breeds in the Arctic tundra of Eurasia from Murmansk Oblast to roughly Yana and Indigirka Rivers, Siberia. It is completely migratory and travels south in winter as far as South Asia and sub-Saharan Africa. It is often seen around wetland habitats. It is paler than the nominate subspecies, especially on the crown. Males weigh 588 to 740 g (1.296–1.631 lb), while females weigh 925 to 1,333 g (2.039–2.939 lb).

Falco peregrinus cassini, described by Sharpe in 1873, is also known as the austral peregrine falcon. It includes kreyenborgi, the pallid falcon, a leucistic colour morph occurring in southernmost South America, which was long believed to be a distinct species. Its range includes South America from Ecuador through Bolivia, northern Argentina and Chile to Tierra del Fuego and the Falkland Islands. It is non-migratory. It is similar to the nominate subspecies, but slightly smaller with a black ear region. The pallid falcon morph kreyenborgi is medium grey above, has little barring below and has a head pattern like the saker falcon (Falco cherrug), but the ear region is white.

Falco peregrinus ernesti, described by Sharpe in 1894, is found from the Sunda Islands to the Philippines and south to eastern New Guinea and the nearby Bismarck Archipelago. Its geographical separation from nesiotes requires confirmation. It is non-migratory. It differs from the nominate subspecies in the very dark, dense barring on its underside and its black ear coverts.

Falco peregrinus furuitii, described by Momiyama in 1927, is found on the Izu and Ogasawara Islands south of Honshū, Japan. It is non-migratory. It is very rare and may only remain on a single island. It is a dark form, resembling pealei in colour, but darker, especially on the tail.

Falco peregrinus japonensis, described by Gmelin in 1788, includes kleinschmidti, pleskei, and harterti, and seems to refer to intergrades with calidus. It is found from northeast Siberia to Kamchatka (though it is possibly replaced by pealei on the coast there) and Japan. Northern populations are migratory, while those of Japan are resident. It is similar to the nominate subspecies, but the young are even darker than those of anatum.

Falco peregrinus macropus, described by Swainson in 1837, is the Australian peregrine falcon. It is found in Australia in all regions except the southwest. It is non-migratory. It is similar to brookei in appearance, but is slightly smaller and the ear region is entirely black. The feet are proportionally large.

Falco peregrinus madens, described by Ripley and Watson in 1963, is unusual in having some sexual dichromatism. If the Barbary falcon (see below) is considered a distinct species, it is sometimes placed therein. It is found in the Cape Verde Islands and is non-migratory; it is also endangered, with only six to eight pairs surviving. Males have a rufous wash on the crown, nape, ears and back; the underside is conspicuously washed pinkish-brown. Females are tinged rich brown overall, especially on the crown and nape.

 

Falco peregrinus minor, first described by Bonaparte in 1850. It was formerly often known as perconfusus. It is sparsely and patchily distributed throughout much of sub-Saharan Africa and widespread in Southern Africa. It apparently reaches north along the Atlantic coast as far as Morocco. It is non-migratory and dark-coloured. This is the smallest subspecies, with smaller males weighing as little as approximately 300 g (11 oz).

Falco peregrinus nesiotes, described by Mayr in 1941, is found in Fiji and probably also Vanuatu and New Caledonia. It is non-migratory.

Falco peregrinus pealei, described by Ridgway in 1873, is Peale's falcon and includes rudolfi. It is found in the Pacific Northwest of North America, northwards from Puget Sound along the British Columbia coast (including the Haida Gwaii), along the Gulf of Alaska and the Aleutian Islands to the far eastern Bering Sea coast of Russia, and may also occur on the Kuril Islands and the coasts of Kamchatka. It is non-migratory. It is the largest subspecies and it looks like an oversized and darker tundrius or like a strongly barred and large anatum. The bill is very wide. Juveniles occasionally have pale crowns. Males weigh 700 to 1,000 g (1.5–2.2 lb), while females weigh 1,000 to 1,500 g (2.2–3.3 lb).

Falco peregrinus pelegrinoides, first described by Temminck in 1829, is found in the Canary Islands through North Africa and the Near East to Mesopotamia. It is most similar to brookei, but is markedly paler above, with a rusty neck, and is a light buff with reduced barring below. It is smaller than the nominate subspecies; females weigh around 610 g (1.34 lb).

Falco peregrinus peregrinator, described by Sundevall in 1837, is known as the Indian peregrine falcon, black shaheen, Indian shaheen or shaheen falcon. It was formerly sometimes known as Falco atriceps or Falco shaheen. Its range includes South Asia from across the Indian subcontinent to Sri Lanka and southeastern China. In India, the shaheen falcon is reported from all states except Uttar Pradesh, mainly from rocky and hilly regions. The shaheen falcon is also reported from the Andaman and Nicobar Islands in the Bay of Bengal. It has a clutch size of 3 to 4 eggs, with the chicks fledging time of 48 days with an average nesting success of 1.32 chicks per nest. In India, apart from nesting on cliffs, it has also been recorded as nesting on man-made structures such as buildings and cellphone transmission towers.[36] A population estimate of 40 breeding pairs in Sri Lanka was made in 1996. It is non-migratory and is small and dark, with rufous underparts. In Sri Lanka this species is found to favour the higher hills, while the migrant calidus is more often seen along the coast.

Falco peregrinus peregrinus, the nominate (first-named) subspecies, described by Tunstall in 1771, breeds over much of temperate Eurasia between the tundra in the north and the Pyrenees, Mediterranean region and Alpide belt in the south. It is mainly non-migratory in Europe, but migratory in Scandinavia and Asia. Males weigh 580 to 750 g (1.28–1.65 lb), while females weigh 925 to 1,300 g (2.039–2.866 lb). It includes brevirostris, germanicus, rhenanus and riphaeus.

Falco peregrinus radama, described by Hartlaub in 1861, is found in Madagascar and the Comoros. It is non-migratory.

Falco peregrinus submelanogenys, described by Mathews in 1912, is the Southwest Australian peregrine falcon. It is found in southwestern Australia and is non-migratory.

Falco peregrinus tundrius, described by C.M. White in 1968, was at one time included in leucogenys. It is found in the Arctic tundra of North America to Greenland, and migrates to wintering grounds in Central and South America. Most vagrants that reach western Europe belong to this subspecies, which was previously considered synonymous with anatum. It is the New World equivalent to calidus. It is smaller and paler than anatum; most have a conspicuous white forehead and white in ear region, but the crown and "moustache" are very dark, unlike in calidus. Juveniles are browner and less grey than in calidus and paler, sometimes almost sandy, than in anatum. Males weigh 500 to 700 g (1.1–1.5 lb), while females weigh 800 to 1,100 g (1.8–2.4 lb). Despite its current recognition as a valid subspecies, a population genetic study of both pre-decline (i.e., museum) and recovered contemporary populations failed to distinguish genetically the anatum and tundrius subspecies.

Barbary falcon

Main article: Barbary falcon

The Barbary falcon is a subspecies of the peregrine falcon that inhabits parts of North Africa; namely, from the Canary Islands to the Arabian Peninsula. There is discussion concerning the taxonomic status of the bird, with some considering it a subspecies of the peregrine falcon and others considering it a full species with two subspecies (White et al. 2013). Compared to the other peregrine falcon subspecies, Barbary falcons sport a slimmer body and a distinct plumage color pattern. Despite numbers and range of these birds throughout the Canary Islands generally increasing, they are considered endangered, with human interference through falconry and shooting threatening their well-being. Falconry can further complicate the speciation and genetics of these Canary Islands falcons, as the practice promotes genetic mixing between individuals from outside the islands with those originating from the islands. Population density of the Barbary falcons on Tenerife, the biggest of the seven major Canary Islands, was found to be 1.27 pairs/100 km², with the mean distance between pairs being 5869 ± 3338 m. The falcons were only observed near large and natural cliffs with a mean altitude of 697.6 m. Falcons show an affinity for tall cliffs away from human-mediated establishments and presence.

 

Barbary falcons have a red neck patch, but otherwise differ in appearance from the peregrine falcon proper merely according to Gloger's rule, relating pigmentation to environmental humidity. The Barbary falcon has a peculiar way of flying, beating only the outer part of its wings like fulmars sometimes do; this also occurs in the peregrine falcon, but less often and far less pronounced. The Barbary falcon's shoulder and pelvis bones are stout by comparison with the peregrine falcon and its feet are smaller. Barbary falcons breed at different times of year than neighboring peregrine falcon subspecies, but they are capable of interbreeding. There is a 0.6–0.7% genetic distance in the peregrine falcon-Barbary falcon ("peregrinoid") complex.

 

Ecology and behaviour

The peregrine falcon lives mostly along mountain ranges, river valleys, coastlines, and increasingly in cities. In mild-winter regions, it is usually a permanent resident, and some individuals, especially adult males, will remain on the breeding territory. Only populations that breed in Arctic climates typically migrate great distances during the northern winter.

 

The peregrine falcon reaches faster speeds than any other animal on the planet when performing the stoop, which involves soaring to a great height and then diving steeply at speeds of over 320 km/h (200 mph), hitting one wing of its prey so as not to harm itself on impact. The air pressure from such a dive could possibly damage a bird's lungs, but small bony tubercles on a falcon's nostrils are theorized to guide the powerful airflow away from the nostrils, enabling the bird to breathe more easily while diving by reducing the change in air pressure. To protect their eyes, the falcons use their nictitating membranes (third eyelids) to spread tears and clear debris from their eyes while maintaining vision. The distinctive malar stripe or 'moustache', a dark area of feathers below the eyes, is thought to reduce solar glare and improve contrast sensitivity when targeting fast moving prey in bright light condition; the malar stripe has been found to be wider and more pronounced in regions of the world with greater solar radiation supporting this solar glare hypothesis. Peregrine falcons have a flicker fusion frequency of 129 Hz (cycles per second), very fast for a bird of its size, and much faster than mammals. A study testing the flight physics of an "ideal falcon" found a theoretical speed limit at 400 km/h (250 mph) for low-altitude flight and 625 km/h (388 mph) for high-altitude flight. In 2005, Ken Franklin recorded a falcon stooping at a top speed of 389 km/h (242 mph).

 

The life span of peregrine falcons in the wild is up to 19 years 9 months. Mortality in the first year is 59–70%, declining to 25–32% annually in adults. Apart from such anthropogenic threats as collision with human-made objects, the peregrine may be killed by larger hawks and owls.

 

The peregrine falcon is host to a range of parasites and pathogens. It is a vector for Avipoxvirus, Newcastle disease virus, Falconid herpesvirus 1 (and possibly other Herpesviridae), and some mycoses and bacterial infections. Endoparasites include Plasmodium relictum (usually not causing malaria in the peregrine falcon), Strigeidae trematodes, Serratospiculum amaculata (nematode), and tapeworms. Known peregrine falcon ectoparasites are chewing lice, Ceratophyllus garei (a flea), and Hippoboscidae flies (Icosta nigra, Ornithoctona erythrocephala).

 

In the Arctic Peregrine falcons chasing away small rodent predators from their nesting territory and Rough-legged Buzzards (Buteo lagopus) could use these hot spots as a nesting territory.

 

Feeding

The peregrine falcon's diet varies greatly and is adapted to available prey in different regions. However, it typically feeds on medium-sized birds such as pigeons and doves, waterfowl, gamebirds, songbirds, parrots, seabirds, and waders. Worldwide, it is estimated that between 1,500 and 2,000 bird species, or roughly a fifth of the world's bird species, are predated somewhere by these falcons.The peregrine falcon preys on the most diverse range of bird species of any raptor in North America, with over 300 species and including nearly 100 shorebirds. Its prey can range from 3 g (0.11 oz) hummingbirds (Selasphorus and Archilochus ssp.) to the 3.1 kg (6.8 lb) sandhill crane, although most prey taken by peregrines weigh between 20 g (0.71 oz) (small passerines) and 1,100 g (2.4 lb) (ducks, geese, loons, gulls, capercaillies, ptarmigans and other grouse). Smaller hawks (such as sharp-shinned hawks) and owls are regularly predated, as well as smaller falcons such as the American kestrel, merlin and, rarely, other peregrines.

 

In urban areas, where it tends to nest on tall buildings or bridges, it subsists mostly on a variety of pigeons. Among pigeons, the rock or feral pigeon comprises 80% or more of the dietary intake of peregrines. Other common city birds are also taken regularly, including mourning doves, common wood pigeons, common swifts, northern flickers, common starlings, American robins, common blackbirds, and corvids such as magpies, jays or carrion, house, and American crows. Coastal populations of the large subspecies pealei feed almost exclusively on seabirds. In the Brazilian mangrove swamp of Cubatão, a wintering falcon of the subspecies tundrius was observed successfully hunting a juvenile scarlet ibis.

 

Among mammalian prey species, bats in the genera Eptesicus, Myotis, Pipistrellus and Tadarida are the most common prey which taken at night. Though peregrines generally do not prefer terrestrial mammalian prey, in Rankin Inlet, peregrines largely take northern collared lemmings (Dicrostonyx groenlandicus) along with a few Arctic ground squirrels (Urocitellus parryii). Other small mammals including shrews, mice, rats, voles, and squirrels are more seldom taken. Peregrines occasionally take rabbits, mainly young individuals and juvenile hares. Additionally, remains of red fox kits and adult female American marten were found among prey remains. Insects and reptiles such as small snakes make up a small proportion of the diet, and salmonid fish have been taken by peregrines.

 

The peregrine falcon hunts most often at dawn and dusk, when prey are most active, but also nocturnally in cities, particularly during migration periods when hunting at night may become prevalent. Nocturnal migrants taken by peregrines include species as diverse as yellow-billed cuckoo, black-necked grebe, virginia rail, and common quail. The peregrine requires open space in order to hunt, and therefore often hunts over open water, marshes, valleys, fields, and tundra, searching for prey either from a high perch or from the air. Large congregations of migrants, especially species that gather in the open like shorebirds, can be quite attractive to a hunting peregrine. Once prey is spotted, it begins its stoop, folding back the tail and wings, with feet tucked. Prey is typically struck and captured in mid-air; the peregrine falcon strikes its prey with a clenched foot, stunning or killing it with the impact, then turns to catch it in mid-air. If its prey is too heavy to carry, a peregrine will drop it to the ground and eat it there. If they miss the initial strike, peregrines will chase their prey in a twisting flight.

 

Although previously thought rare, several cases of peregrines contour-hunting, i.e., using natural contours to surprise and ambush prey on the ground, have been reported and even rare cases of prey being pursued on foot. In addition, peregrines have been documented preying on chicks in nests, from birds such as kittiwakes. Prey is plucked before consumption. A 2016 study showed that the presence of peregrines benefits non-preferred species while at the same time causing a decline in its preferred prey. As of 2018, the fastest recorded falcon was at 242 mph (nearly 390 km/h). Researchers at the University of Groningen in the Netherlands and at Oxford University used 3D computer simulations in 2018 to show that the high speed allows peregrines to gain better maneuverability and precision in strikes.

 

Reproduction

The peregrine falcon is sexually mature at one to three years of age, but in larger populations they breed after two to three years of age. A pair mates for life and returns to the same nesting spot annually. The courtship flight includes a mix of aerial acrobatics, precise spirals, and steep dives. The male passes prey it has caught to the female in mid-air. To make this possible, the female actually flies upside-down to receive the food from the male's talons.

 

During the breeding season, the peregrine falcon is territorial; nesting pairs are usually more than 1 km (0.62 mi) apart, and often much farther, even in areas with large numbers of pairs. The distance between nests ensures sufficient food supply for pairs and their chicks. Within a breeding territory, a pair may have several nesting ledges; the number used by a pair can vary from one or two up to seven in a 16-year period.

 

The peregrine falcon nests in a scrape, normally on cliff edges. The female chooses a nest site, where she scrapes a shallow hollow in the loose soil, sand, gravel, or dead vegetation in which to lay eggs. No nest materials are added. Cliff nests are generally located under an overhang, on ledges with vegetation. South-facing sites are favoured. In some regions, as in parts of Australia and on the west coast of northern North America, large tree hollows are used for nesting. Before the demise of most European peregrines, a large population of peregrines in central and western Europe used the disused nests of other large birds. In remote, undisturbed areas such as the Arctic, steep slopes and even low rocks and mounds may be used as nest sites. In many parts of its range, peregrines now also nest regularly on tall buildings or bridges; these human-made structures used for breeding closely resemble the natural cliff ledges that the peregrine prefers for its nesting locations.

 

The pair defends the chosen nest site against other peregrines, and often against ravens, herons, and gulls, and if ground-nesting, also such mammals as foxes, wolverines, felids, bears, wolves, and mountain lions. Both nests and (less frequently) adults are predated by larger-bodied raptorial birds like eagles, large owls, or gyrfalcons. The most serious predators of peregrine nests in North America and Europe are the great horned owl and the Eurasian eagle-owl. When reintroductions have been attempted for peregrines, the most serious impediments were these two species of owls routinely picking off nestlings, fledglings and adults by night. Peregrines defending their nests have managed to kill raptors as large as golden eagles and bald eagles (both of which they normally avoid as potential predators) that have come too close to the nest by ambushing them in a full stoop. In one instance, when a snowy owl killed a newly fledged peregrine, the larger owl was in turn killed by a stooping peregrine parent.

 

The date of egg-laying varies according to locality, but is generally from February to March in the Northern Hemisphere, and from July to August in the Southern Hemisphere, although the Australian subspecies macropus may breed as late as November, and equatorial populations may nest anytime between June and December. If the eggs are lost early in the nesting season, the female usually lays another clutch, although this is extremely rare in the Arctic due to the short summer season. Generally three to four eggs, but sometimes as few as one or as many as five, are laid in the scrape. The eggs are white to buff with red or brown markings. They are incubated for 29 to 33 days, mainly by the female, with the male also helping with the incubation of the eggs during the day, but only the female incubating them at night. The average number of young found in nests is 2.5, and the average number that fledge is about 1.5, due to the occasional production of infertile eggs and various natural losses of nestlings.

 

After hatching, the chicks (called "eyases") are covered with creamy-white down and have disproportionately large feet. The male (called the "tiercel") and the female (simply called the "falcon") both leave the nest to gather prey to feed the young. The hunting territory of the parents can extend a radius of 19 to 24 km (12 to 15 mi) from the nest site. Chicks fledge 42 to 46 days after hatching, and remain dependent on their parents for up to two months.

 

Relationship with humans

The peregrine falcon is a highly admired falconry bird, and has been used in falconry for more than 3,000 years, beginning with nomads in central Asia. Its advantages in falconry include not only its athleticism and eagerness to hunt, but an equable disposition that leads to it being one of the easier falcons to train. The peregrine falcon has the additional advantage of a natural flight style of circling above the falconer ("waiting on") for game to be flushed, and then performing an effective and exciting high-speed diving stoop to take the quarry. The speed of the stoop not only allows the falcon to catch fast flying birds, it also enhances the falcon's ability to execute maneuvers to catch highly agile prey, and allows the falcon to deliver a knockout blow with a fist-like clenched talon against game that may be much larger than itself.

 

Additionally the versatility of the species, with agility allowing capture of smaller birds and a strength and attacking style allowing capture of game much larger than themselves, combined with the wide size range of the many peregrine subspecies, means there is a subspecies suitable to almost any size and type of game bird. This size range, evolved to fit various environments and prey species, is from the larger females of the largest subspecies to the smaller males of the smallest subspecies, approximately five to one (approximately 1500 g to 300 g). The males of smaller and medium-sized subspecies, and the females of the smaller subspecies, excel in the taking of swift and agile small game birds such as dove, quail, and smaller ducks. The females of the larger subspecies are capable of taking large and powerful game birds such as the largest of duck species, pheasant, and grouse.

 

Peregrine falcons handled by falconers are also occasionally used to scare away birds at airports to reduce the risk of bird-plane strikes, improving air-traffic safety. They were also used to intercept homing pigeons during World War II.

 

Peregrine falcons have been successfully bred in captivity, both for falconry and for release into the wild. Until 2004 nearly all peregrines used for falconry in the US were captive-bred from the progeny of falcons taken before the US Endangered Species Act was enacted and from those few infusions of wild genes available from Canada and special circumstances. Peregrine falcons were removed from the United States' endangered species list in 1999. The successful recovery program was aided by the effort and knowledge of falconers – in collaboration with The Peregrine Fund and state and federal agencies – through a technique called hacking. Finally, after years of close work with the US Fish and Wildlife Service, a limited take of wild peregrines was allowed in 2004, the first wild peregrines taken specifically for falconry in over 30 years.

 

The development of captive breeding methods has led to peregrines being commercially available for falconry use, thus mostly eliminating the need to capture wild birds for support of falconry. The main reason for taking wild peregrines at this point is to maintain healthy genetic diversity in the breeding lines. Hybrids of peregrines and gyrfalcons are also available that can combine the best features of both species to create what many consider to be the ultimate falconry bird for the taking of larger game such as the sage-grouse. These hybrids combine the greater size, strength, and horizontal speed of the gyrfalcon with the natural propensity to stoop and greater warm weather tolerance of the peregrine.

 

Decline due to pesticides

The peregrine falcon became an endangered species over much of its range because of the use of organochlorine pesticides, especially DDT, during the 1950s, '60s, and '70s. Pesticide biomagnification caused organochlorine to build up in the falcons' fat tissues, reducing the amount of calcium in their eggshells. With thinner shells, fewer falcon eggs survived until hatching. In addition, the PCB concentrations found in these falcons is dependent upon the age of the falcon. While high levels are still found in young birds (only a few months old) and even higher concentrations are found in more mature falcons, further increasing in adult peregrine falcons. These pesticides caused falcon prey to also have thinner eggshells (one example of prey being the Black Petrels). In several parts of the world, such as the eastern United States and Belgium, this species became extirpated (locally extinct) as a result. An alternate point of view is that populations in the eastern North America had vanished due to hunting and egg collection. Following the ban of organochlorine pesticides, the reproductive success of Peregrines increased in Scotland in terms of territory occupancy and breeding success, although spatial variation in recovery rates indicate that in some areas Peregrines were also impacted by other factors such as persecution.

 

Recovery efforts

Peregrine falcon recovery teams breed the species in captivity. The chicks are usually fed through a chute or with a hand puppet mimicking a peregrine's head, so they cannot see to imprint on the human trainers. Then, when they are old enough, the rearing box is opened, allowing the bird to train its wings. As the fledgling gets stronger, feeding is reduced, forcing the bird to learn to hunt. This procedure is called hacking back to the wild. To release a captive-bred falcon, the bird is placed in a special cage at the top of a tower or cliff ledge for some days or so, allowing it to acclimate itself to its future environment.

 

Worldwide recovery efforts have been remarkably successful. The widespread restriction of DDT use eventually allowed released birds to breed successfully. The peregrine falcon was removed from the U.S. Endangered Species list on 25 August 1999.

 

Some controversy has existed over the origins of captive breeding stock used by the Peregrine Fund in the recovery of peregrine falcons throughout the contiguous United States. Several peregrine subspecies were included in the breeding stock, including birds of Eurasian origin. Due to the extirpation of the eastern population of Falco peregrinus anatum, the near-extirpation of anatum in the Midwest and the limited gene pool within North American breeding stock, the inclusion of non-native subspecies was justified to optimize the genetic diversity found within the species as a whole.

 

During the 1970s, peregrine falcons in Finland experienced a population bottleneck as a result of large declines associated with bio-accumulation of organochloride pesticides. However, the genetic diversity of peregrines in Finland is similar to other populations, indicating that high dispersal rates have maintained the genetic diversity of this species.

 

Since peregrine falcon eggs and chicks are still often targeted by illegal poachers, it is common practice not to publicize unprotected nest locations.

 

Current status

Populations of the peregrine falcon have bounced back in most parts of the world. In the United Kingdom, there has been a recovery of populations since the crash of the 1960s. This has been greatly assisted by conservation and protection work led by the Royal Society for the Protection of Birds. The RSPB estimated that there were 1,402 breeding pairs in the UK in 2011. In Canada, where peregrines were identified as endangered in 1978 (in the Yukon territory of northern Canada that year, only a single breeding pair was identified), the Committee on the Status of Endangered Wildlife in Canada declared the species no longer at risk in December 2017.

 

Peregrines now breed in many mountainous and coastal areas, especially in the west and north, and nest in some urban areas, capitalising on the urban feral pigeon populations for food. Additionally, falcons benefit from artificial illumination, which allows the raptors to extend their hunting periods into the dusk when natural illumination would otherwise be too low for them to pursue prey. In England, this has allowed them to prey on nocturnal migrants such as redwings, fieldfares, starlings, and woodcocks.

 

In many parts of the world peregrine falcons have adapted to urban habitats, nesting on cathedrals, skyscraper window ledges, tower blocks, and the towers of suspension bridges. Many of these nesting birds are encouraged, sometimes gathering media attention and often monitored by cameras.

 

In England, peregrine falcons have become increasingly urban in distribution, particularly in southern areas where inland cliffs suitable as nesting sites are scarce. The first recorded urban breeding pair was observed nesting on the Swansea Guildhall in the 1980s. In Southampton, a nest prevented restoration of mobile telephony services for several months in 2013, after Vodafone engineers despatched to repair a faulty transmitter mast discovered a nest in the mast, and were prevented by the Wildlife and Countryside Act – on pain of a possible prison sentence – from proceeding with repairs until the chicks fledged.

 

In Oregon, Portland houses ten percent of the state's peregrine nests, despite only covering around 0.1 percent of the state's land area.

 

Cultural significance

Due to its striking hunting technique, the peregrine has often been associated with aggression and martial prowess. The Ancient Egyptian solar deity Ra was often represented as a man with the head of a peregrine falcon adorned with the solar disk, although most Egyptologists agree that it's most likely a Lanner falcon. Native Americans of the Mississippian culture (c. 800–1500) used the peregrine, along with several other birds of prey, in imagery as a symbol of "aerial (celestial) power" and buried men of high status in costumes associating to the ferocity of raptorial birds. In the late Middle Ages, the Western European nobility that used peregrines for hunting, considered the bird associated with princes in formal hierarchies of birds of prey, just below the gyrfalcon associated with kings. It was considered "a royal bird, more armed by its courage than its claws". Terminology used by peregrine breeders also used the Old French term gentil, "of noble birth; aristocratic", particularly with the peregrine.

 

The peregrine falcon is the national animal of the United Arab Emirates. Since 1927, the peregrine falcon has been the official mascot of Bowling Green State University in Bowling Green, Ohio. The 2007 U.S. Idaho state quarter features a peregrine falcon. The peregrine falcon has been designated the official city bird of Chicago.

 

The Peregrine, by J. A. Baker, is widely regarded as one of the best nature books in English written in the twentieth century. Admirers of the book include Robert Macfarlane, Mark Cocker, who regards the book as "one of the most outstanding books on nature in the twentieth century" and Werner Herzog, who called it "the one book I would ask you to read if you want to make films", and said elsewhere "it has prose of the calibre that we have not seen since Joseph Conrad". In the book, Baker recounts, in diary form, his detailed observations of peregrines (and their interaction with other birds) near his home in Chelmsford, Essex, over a single winter from October to April.

 

An episode of the hour-long TV series Starman in 1986 titled "Peregrine" was about an injured peregrine falcon and the endangered species program. It was filmed with the assistance of the University of California's peregrine falcon project in Santa Cruz.

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