Narzissen enthalten wie alle Amaryllisgewächse „Amaryllidaceen-Alkaloide“. Je nach Art können dies unterschiedliche sein. Die Gelbe Narzisse beispielsweise enthält die Alkaloide Narcissin, Galantamin und Lycorin.

Für die Pflanze stellen die Alkaloide einen natürlichen Schutz gegen Parasitenbefall und weidende Pflanzenfresser dar: „Fraßschutzgifte“. Narzissen enthalten außerdem Fructane und niedermolekulare Glucomannane, die unter anderem in den Blättern und Pflanzenstängeln reichlich enthalten sind.

Zu Vergiftungen kommt es gelegentlich, weil die Zwiebeln im Ruhezustand denen der Küchenzwiebel sehr ähnlich sehen. Der Verzehr von Narzissenzwiebeln kann zu Würgereiz, Erbrechen, Diarrhoe, Schläfrigkeit, Schweißausbruch, Benommenheit, Kollaps und Lähmungserscheinungen führen. Bei sehr großen Dosen kann es auch zum Tod kommen.

 

Like all amaryllis plants, daffodils contain "amaryllidaceae alkaloids". Depending on the species, these can be different. For example, the yellow daffodil contains the alkaloids narcissine, galantamine and lycorine.

For the plant, the alkaloids represent a natural protection against parasite infestation and grazing herbivores: "anti-feeding poisons". Daffodils also contain fructans and low-molecular glucomannan, which are found in abundance in the leaves and plant stems, among other things.

Poisoning occasionally occurs because the onions look very similar to those of the kitchen onion when they are at rest. Consuming daffodil bulbs can cause retching, vomiting, diarrhea, drowsiness, sweating, drowsiness, collapse and paralysis. In very large doses, death can also occur.

anacamptis morio

 

L'orchis bouffon (Anacamptis morio), est une espèce de plantes à fleurs de la famille des Orchidaceae. C'est une orchidée terrestre eurasiatique.

L'aire de répartition, très vaste, va du sud de la Norvège à l'Iran et au bassin méditerranéen.

L'espèce est souvent victime du prélèvement systématique des tubercules pour la commercialisation du Salep. Des stations complètes sont ainsi détruites.

Le salep ou sahlep ou sohlob en Tunisie est une farine faite à partir de tubercules d'Orchis et, par extension, la boisson que l'on confectionne avec cette farine.

Durant l’Antiquité, les Romains utilisaient des bulbes d'orchidées moulus pour fabriquer des boissons auxquelles ils donnaient un certain nombre de noms, en particulier satyrion et priapiscus. Comme ces noms l'indiquent, ils considéraient également ces boissons comme de puissants aphrodisiaques

De nos jours, la boisson est souvent faite avec du lait chaud au lieu de l'eau et est saupoudrée de cannelle.

Le salep se boit dans une tasse en grès. D'autres desserts sont également fabriqués à partir de salep, notamment le pudding de salep et la crème glacée au salep.

 

Anacamptis morio, the green-winged orchid] or green-veined orchid (synonym Orchis morio), is a flowering plant of the orchid family, Orchidaceae. It usually has purple flowers, and is found in Europe and the Middle East.

The species is often the victim of systematic harvesting of tubers for the sale of Salep. Entire stations are thus destroyed.

Salep, also spelled sahlep or sahlab, is a flour made from the tubers of the orchid genus Orchis (including species Orchis mascula and Orchis militaris). These tubers contain a nutritious, starchy polysaccharide called glucomannan. Salep flour is consumed in beverages and desserts, especially in the cuisines of the former Ottoman Empire, notably in the Levant where it is a traditional winter beverage. An increase in consumption is causing local extinctions of orchids in parts of Turkey and Iran

The Ancient Romans used ground orchid bulbs to make drinks, which they called by a number of names, especially satyrion and priapiscus. As the names indicate, they considered it to be a powerful aphrodisiac.[6] Of salep, Paracelsus wrote: "behold the Satyrion root, is it not formed like the male privy parts? No one can deny this. Accordingly, magic discovered it and revealed that it can restore a man's virility and passion ».

This beverage was sold in place of tea and coffee, which were much more expensive, and was served in a similar way with milk and sugar.

 

From Wikipedia, the free encyclopedia

 

_5D42043

Narcissus is a genus of predominantly spring flowering perennial plants of the amaryllis family, Amaryllidaceae. Various common names including daffodil, narcissus, and jonquil are used to describe all or some members of the genus. Narcissus has conspicuous flowers with six petal-like tepals surmounted by a cup- or trumpet-shaped corona. The flowers are generally white and yellow (also orange or pink in garden varieties), with either uniform or contrasting coloured tepals and corona.

 

Narcissus were well known in ancient civilisation, both medicinally and botanically, but formally described by Linnaeus in his Species Plantarum (1753). The genus is generally considered to have about ten sections with approximately 50 species. The number of species has varied, depending on how they are classified, due to similarity between species and hybridisation. The genus arose some time in the Late Oligocene to Early Miocene epochs, in the Iberian peninsula and adjacent areas of southwest Europe. The exact origin of the name Narcissus is unknown, but it is often linked to a Greek word (ancient Greek ναρκῶ narkō, "to make numb") and the myth of the youth of that name who fell in love with his own reflection. The English word "daffodil" appears to be derived from "asphodel", with which it was commonly compared.

 

The species are native to meadows and woods in southern Europe and North Africa with a centre of diversity in the Western Mediterranean, particularly the Iberian peninsula. Both wild and cultivated plants have naturalised widely, and were introduced into the Far East prior to the tenth century. Narcissi tend to be long-lived bulbs, which propagate by division, but are also insect-pollinated. Known pests, diseases and disorders include viruses, fungi, the larvae of flies, mites and nematodes. Some Narcissus species have become extinct, while others are threatened by increasing urbanisation and tourism.

 

Historical accounts suggest narcissi have been cultivated from the earliest times, but became increasingly popular in Europe after the 16th century and by the late 19th century were an important commercial crop centred primarily in the Netherlands. Today narcissi are popular as cut flowers and as ornamental plants in private and public gardens. The long history of breeding has resulted in thousands of different cultivars. For horticultural purposes, narcissi are classified into divisions, covering a wide range of shapes and colours. Like other members of their family, narcissi produce a number of different alkaloids, which provide some protection for the plant, but may be poisonous if accidentally ingested. This property has been exploited for medicinal use in traditional healing and has resulted in the production of galantamine for the treatment of Alzheimer's dementia. Long celebrated in art and literature, narcissi are associated with a number of themes in different cultures, ranging from death to good fortune, and as symbols of spring.

 

The daffodil is the national flower of Wales and the symbol of cancer charities in many countries. The appearance of the wild flowers in spring is associated with festivals in many places.

 

Narcissus is a genus of perennial herbaceous bulbiferous geophytes, which die back after flowering to an underground storage bulb. They regrow in the following year from brown-skinned ovoid bulbs with pronounced necks, and reach heights of 5–80 centimetres (2.0–31.5 in) depending on the species. Dwarf species such as N. asturiensis have a maximum height of 5–8 centimetres (2.0–3.1 in), while Narcissus tazetta may grow as tall as 80 centimetres (31 in).

 

The plants are scapose, having a single central leafless hollow flower stem (scape). Several green or blue-green, narrow, strap-shaped leaves arise from the bulb. The plant stem usually bears a solitary flower, but occasionally a cluster of flowers (umbel). The flowers, which are usually conspicuous and white or yellow, sometimes both or rarely green, consist of a perianth of three parts. Closest to the stem (proximal) is a floral tube above the ovary, then an outer ring composed of six tepals (undifferentiated sepals and petals), and a central disc to conical shaped corona. The flowers may hang down (pendant), or be erect. There are six pollen bearing stamens surrounding a central style. The ovary is inferior (below the floral parts) consisting of three chambers (trilocular). The fruit consists of a dry capsule that splits (dehisces) releasing numerous black seeds.

 

The bulb lies dormant after the leaves and flower stem die back and has contractile roots that pull it down further into the soil. The flower stem and leaves form in the bulb, to emerge the following season. Most species are dormant from summer to late winter, flowering in the spring, though a few species are autumn flowering.

 

The pale brown-skinned ovoid tunicate bulbs have a membranous tunic and a corky stem (base or basal) plate from which arise the adventitious root hairs in a ring around the edge, which grow up to 40 mm in length. Above the stem plate is the storage organ consisting of bulb scales, surrounding the previous flower stalk and the terminal bud. The scales are of two types, true storage organs and the bases of the foliage leaves. These have a thicker tip and a scar from where the leaf lamina became detached. The innermost leaf scale is semicircular only partly enveloping the flower stalk (semisheathed).(see Hanks Fig 1.3). The bulb may contain a number of branched bulb units, each with two to three true scales and two to three leaf bases. Each bulb unit has a life of about four years.

 

Once the leaves die back in summer, the roots also wither. After some years, the roots shorten pulling the bulbs deeper into the ground (contractile roots). The bulbs develop from the inside, pushing the older layers outwards which become brown and dry, forming an outer shell, the tunic or skin. Up to 60 layers have been counted in some wild species. While the plant appears dormant above the ground the flower stalk which will start to grow in the following spring, develops within the bulb surrounded by two to three deciduous leaves and their sheaths. The flower stem lies in the axil of the second true leaf.

 

The single leafless Plant stem stem or scape, appearing from early to late spring depending on the species, bears from 1 to 20 blooms. Stem shape depends on the species, some are highly compressed with a visible seam, while others are rounded. The stems are upright and located at the centre of the leaves. In a few species such as N. hedraeanthus the stem is oblique. The stem is hollow in the upper portion but towards the bulb is more solid and filled with a spongy material.

 

Narcissus plants have one to several basal leaf leaves which are linear, ligulate or strap-shaped (long and narrow), sometimes channelled adaxially to semiterete, and may (pedicellate) or may not (sessile) have a petiole stalk. The leaves are flat and broad to cylindrical at the base and arise from the bulb. The emerging plant generally has two leaves, but the mature plant usually three, rarely four, and they are covered with a cutin containing cuticle, giving them a waxy appearance. Leaf colour is light green to blue-green. In the mature plant, the leaves extend higher than the flower stem, but in some species, the leaves are low-hanging. The leaf base is encased in a colorless sheath. After flowering, the leaves turn yellow and die back once the seed pod (fruit) is ripe.

 

Jonquils usually have dark green, round, rush-like leaves.

 

The inflorescence is scapose, the single stem or scape bearing either a solitary flower or forming an umbel with up to 20 blooms. Species bearing a solitary flower include section Bulbocodium and most of section Pseudonarcissus. Umbellate species have a fleshy racemose inflorescence (unbranched, with short floral stalks) with 2 to 15 or 20 flowers, such as N. papyraceus (see illustration, left) and N. tazetta (see Table I). The flower arrangement on the inflorescence may be either with (pedicellate) or without (sessile) floral stalks.

 

Prior to opening, the flower buds are enveloped and protected in a thin dry papery or membranous (scarious) spathe. The spathe consists of a singular bract that is ribbed, and which remains wrapped around the base of the open flower. As the bud grows, the spathe splits longitudinally. Bracteoles are small or absent.

 

The flowers of Narcissus are hermaphroditic (bisexual), have three parts (tripartite), and are sometimes fragrant (see Fragrances). The flower symmetry is actinomorphic (radial) to slightly zygomorphic (bilateral) due to declinate-ascending stamens (curving downwards, then bent up at the tip). Narcissus flowers are characterised by their, usually conspicuous, corona (trumpet).

 

The three major floral parts (in all species except N. cavanillesii in which the corona is virtually absent - Table I: Section Tapeinanthus) are;

 

(i) the proximal floral tube (hypanthium),

(ii) the surrounding free tepals, and

(iii) the more distal corona (paraperigon, paraperigonium).

All three parts may be considered to be components of the perianth (perigon, perigonium). The perianth arises above the apex of the inferior ovary, its base forming the hypanthial floral tube.

 

The floral tube is formed by fusion of the basal segments of the tepals (proximally connate). Its shape is from an inverted cone (obconic) to funnel-shaped (funneliform) or cylindrical, and is surmounted by the more distal corona. Floral tubes can range from long and narrow sections Apodanthi and Jonquilla to rudimentary (N. cavanillesii).

 

Surrounding the floral tube and corona and reflexed (bent back) from the rest of the perianth are the six spreading tepals or floral leaves, in two whorls which may be distally ascending, reflexed (folded back), or lanceolate. Like many monocotyledons, the perianth is homochlamydeous, which is undifferentiated into separate calyx (sepals) and corolla (petals), but rather has six tepals. The three outer tepal segments may be considered sepals, and the three inner segments petals. The transition point between the floral tube and the corona is marked by the insertion of the free tepals on the fused perianth.

 

The corona, or paracorolla, is variously described as bell-shaped (funneliform, trumpet), bowl-shaped (cupular, crateriform, cup-shaped) or disc-shaped with margins that are often frilled, and is free from the stamens. Rarely is the corona a simple callose (hardened, thickened) ring. The corona is formed during floral development as a tubular outgrowth from stamens which fuse into a tubular structure, the anthers becoming reduced. At its base, the fragrances which attract pollinators are formed. All species produce nectar at the top of the ovary. Coronal morphology varies from the tiny pigmented disk of N. serotinus (see Table I) or the rudimentary structure in N. cavanillesii to the elongated trumpets of section Pseudonarcissus (trumpet daffodils, Table I).

 

While the perianth may point forwards, in some species such as N. cyclamineus it is folded back (reflexed, see illustration, left), while in some other species such as N. bulbocodium (Table I), it is reduced to a few barely visible pointed segments with a prominent corona.

 

The colour of the perianth is white, yellow or bicoloured, with the exception of the night flowering N. viridiflorus which is green. In addition the corona of N. poeticus has a red crenulate margin (see Table I). Flower diameter varies from 12 mm (N. bulbocodium) to over 125 mm (N. nobilis=N. pseudonarcissus subsp. nobilis).

 

Flower orientation varies from pendent or deflexed (hanging down) as in N. triandrus (see illustration, left), through declinate-ascendant as in N. alpestris = N. pseudonarcissus subsp. moschatus, horizontal (patent, spreading) such as N. gaditanus or N. poeticus, erect as in N. cavanillesii, N. serotinus and N. rupicola (Table I), or intermediate between these positions (erecto-patent).

 

The flowers of Narcissus demonstrate exceptional floral diversity and sexual polymorphism, primarily by corona size and floral tube length, associated with pollinator groups (see for instance Figs. 1 and 2 in Graham and Barrett). Barrett and Harder (2005) describe three separate floral patterns;

 

"Daffodil" form

"Paperwhite" form

"Triandrus" form.

The predominant patterns are the 'daffodil' and 'paperwhite' forms, while the "triandrus" form is less common. Each corresponds to a different group of pollinators (See Pollination).

 

The "daffodil" form, which includes sections Pseudonarcissus and Bulbocodium, has a relatively short, broad or highly funnelform tube (funnel-like), which grades into an elongated corona, which is large and funnelform, forming a broad, cylindrical or trumpet-shaped perianth. Section Pseudonarcissus consists of relatively large flowers with a corolla length of around 50 mm, generally solitary but rarely in inflorescences of 2–4 flowers. They have wide greenish floral tubes with funnel-shaped bright yellow coronas. The six tepals sometimes differ in colour from the corona and may be cream coloured to pale yellow.

 

The "paperwhite" form, including sections Jonquilla, Apodanthi and Narcissus, has a relatively long, narrow tube and a short, shallow, flaring corona. The flower is horizontal and fragrant.

 

The "triandrus" form is seen in only two species, N. albimarginatus (a Moroccan endemic) and N. triandrus. It combines features of both the "daffodil" and "paperwhite" forms, with a well-developed, long, narrow tube and an extended bell-shaped corona of almost equal length. The flowers are pendent.

 

Androecium

There are six stamens in one to two rows (whorls), with the filaments separate from the corona, attached at the throat or base of the tube (epipetalous), often of two separate lengths, straight or declinate-ascending (curving downwards, then upwards). The anthers are basifixed (attached at their base).

 

Gynoecium

The ovary is inferior (below the floral parts) and trilocular (three chambered) and there is a pistil with a minutely three lobed stigma and filiform (thread like) style, which is often exserted (extending beyond the tube).

 

Fruit

The fruit consists of dehiscent loculicidal capsules (splitting between the locules) that are ellipsoid to subglobose (almost spherical) in shape and are papery to leathery in texture.

 

Seeds

The fruit contains numerous subglobose seeds which are round and swollen with a hard coat, sometimes with an attached elaiosome. The testa is black and the pericarp dry.

 

Most species have 12 ovules and 36 seeds, although some species such as N. bulbocodium have more, up to a maximum of 60. Seeds take five to six weeks to mature. The seeds of sections Jonquilla and Bulbocodium are wedge-shaped and matte black, while those of other sections are ovate and glossy black. A gust of wind or contact with a passing animal is sufficient to disperse the mature seeds.

 

Chromosomes

Chromosome numbers include 2n=14, 22, 26, with numerous aneuploid and polyploid derivatives. The basic chromosome number is 7, with the exception of N. tazetta, N. elegans and N. broussonetii in which it is 10 or 11; this subgenus (Hermione) was in fact characterised by this characteristic. Polyploid species include N. papyraceus (4x=22) and N. dubius (6x=50).

 

Phytochemistry

Alkaloids

As with all Amarylidaceae genera, Narcissus contains unique isoquinoline alkaloids. The first alkaloid to be identified was lycorine, from N. pseudonarcissus in 1877. These are considered a protective adaptation and are utilised in the classification of species. Nearly 100 alkaloids have been identified in the genus, about a third of all known Amaryllidaceae alkaloids, although not all species have been tested. Of the nine alkaloid ring types identified in the family, Narcissus species most commonly demonstrate the presence of alkaloids from within the Lycorine (lycorine, galanthine, pluviine) and Homolycorine (homolycorine, lycorenine) groups. Hemanthamine, tazettine, narciclasine, montanine and galantamine alkaloids are also represented. The alkaloid profile of any plant varies with time, location, and developmental stage. Narcissus also contain fructans and low molecular weight glucomannan in the leaves and plant stems.

 

Fragrances

Fragrances are predominantly monoterpene isoprenoids, with a small amount of benzenoids, although N. jonquilla has both equally represented. Another exception is N. cuatrecasasii which produces mainly fatty acid derivatives. The basic monoterpene precursor is geranyl pyrophosphate, and the commonest monoterpenes are limonene, myrcene, and trans-β-ocimene. Most benzenoids are non-methoxylated, while a few species contain methoxylated forms (ethers), e.g. N. bujei. Other ingredient include indole, isopentenoids and very small amounts of sesquiterpenes. Fragrance patterns can be correlated with pollinators, and fall into three main groups (see Pollination).

 

The taxonomy of Narcissus is complex, and still not fully resolved. Known to the ancients, the genus name appears in Graeco-Roman literature, although their interest was as much medicinal as botanical. It is unclear which species the ancients were familiar with. Although frequently mentioned in Mediaeval and Renaissance texts it was not formally described till the work of Linnaeus in 1753. By 1789 it had been grouped into a family (Narcissi) but shortly thereafter this was renamed Amaryllideae, from which comes the modern placement within Amaryllidaceae, although for a while it was considered part of Liliaceae.

 

Many of the species now considered to be Narcissus were in separate genera during the nineteenth century, and the situation was further confused by the inclusion of many cultivated varieties. By 1875 the current circumscription was relatively settled. By 2004 phylogenetic studies had allowed the place of Narcissus within its fairly large family to be established, nested within a series of subfamilies (Amaryllidoideae) and tribes (Narcisseae). It shares its position in the latter tribe with Sternbergia.

 

The infrageneric classification has been even more complex and many schemes of subgenera, sections, subsections and series have been proposed, although all had certain similarities. Most authorities now consider there to be 10 – 11 sections based on phylogenetic evidence. The problems have largely arisen from the diversity of the wild species, frequent natural hybridisation and extensive cultivation with escape and subsequent naturalisation. The number of species has varied anywhere from 16 to nearly 160, but is probably around 50 – 60.

 

The genus appeared some time in the Late Oligocene to Early Miocene eras, around 24 million years ago, in the Iberian peninsula. While the exact origin of the word Narcissus is unknown it is frequently linked to its fragrance which was thought to be narcotic, and to the legend of the youth of that name who fell in love with his reflection. In the English language the common name Daffodil appears to be derived from the Asphodel with which it was commonly compared.

 

Early

Narcissus was first described by Theophrastus (Θεόφραστος, c 371 - c 287 BC) in his Historia Plantarum (Greek: Περὶ φυτῶν ἱστορία) as νάρκισσος, referring to N. poeticus, but comparing it to Asphodelus (ασφοδελωδες). Theophrastus' description was frequently referred to at length by later authors writing in Latin such as Pliny the Elder (Gaius Plinius Secundus, 23 AD – 79 AD) from whom came the Latin form narcissus (see also Culture). Pliny's account is from his Natural History (Latin: Naturalis Historia). Like his contemporaries, his interests were as much therapeutic as botanical. Another much-cited Greek authority was Dioscorides (Διοσκουρίδης, 40 AD – 90 AD) in his De Materia Medica (Greek: Περὶ ὕλης ἰατρικῆς). Both authors were to remain influential until at least the Renaissance, given that their descriptions went beyond the merely botanical, to the therapeutic (see also Antiquity).

 

An early European reference is found in the work of Albert Magnus (c. 1200 – 1280), who noted in his De vegetabilibus et plantis the similarity to the leek. William Turner in his A New Herball (1551) cites all three extensively in his description of the plant and its properties.It was to remain to Linnaeus in 1753 to formally describe and name Narcissus as a genus in his Species Plantarum, at which time there were six known species (N. poeticus, N. pseudonarcissus, N. bulbocodium, N. serotinus, N. jonquilla and N. tazetta).[1] At that time, Linnaeus loosely grouped it together with 50 other genera into his Hexandria monogynia.

 

Modern

It was de Jussieu in 1789 who first formally created a 'family' (Narcissi), as the seventh 'Ordo' (Order) of the third class (Stamina epigyna) of Monocots in which Narcissus and 15 other genera were placed. The use of the term Ordo at that time was closer to what we now understand as Family, rather than Order. The family has undergone much reorganisation since then, but in 1805 it was renamed after a different genus in the family, Amaryllis, as 'Amaryllideae' by Jaume St.-Hilaire and has retained that association since. Jaume St.-Hilaire divided the family into two unnamed sections and recognised five species of Narcissus, omitting N. serotinus.

 

De Candolle brought together Linnaeus' genera and Jussieau's families into a systematic taxonomy for the first time, but included Narcissus (together with Amaryllis) in the Liliaceae in his Flore française (1805-1815) rather than Amaryllidaceae, a family he had not yet recognised. Shortly thereafter he separated the 'Amaryllidées' from 'Liliacées' (1813), though attributing the term to Brown's 'Amaryllideae' in the latter's Prodromus (1810) rather than St.-Hilaire's 'Amaryllidées'. He also provided the text to the first four volumes of Redouté illustrations in the latter's Les liliacées between 1805 and 1808 (see illustration here of N. candidissimus).

 

Historically both wide and narrow interpretations of the genus have been proposed. In the nineteenth century genus splitting was common, favouring the narrow view. Haworth (1831) using a narrow view treated many species as separate genera, as did Salisbury (1866). These authors listed various species in related genera such as Queltia (hybrids), Ajax (=Pseudonarcissus) and Hermione (=Tazettae), sixteen in all in Haworth's classification. In contrast, Herbert (1837) took a very wide view reducing Harworth's sixteen genera to six. Herbert, treating the Amaryllidacea as an 'order' as was common then, considered the narcissi to be a suborder, the Narcisseae, the six genera being Corbularia, Ajax, Ganymedes, Queltia, Narcissus and Hermione and his relatively narrow circumscription of Narcissus having only three species. Later Spach (1846) took an even wider view bringing most of Harworth's genera into the genus Narcissus, but as separate subgenera. By the time that Baker (1875) wrote his monograph all of the genera with one exception were included as Narcissus. The exception was the monotypic group Tapeinanthus which various subsequent authors have chosen to either exclude (e.g. Cullen 1986) or include (e.g. Webb 1978, 1980). Today it is nearly always included.

 

The eventual position of Narcissus within the Amaryllidaceae family only became settled in the twenty-first century with the advent of phylogenetic analysis and the Angiosperm Phylogeny Group system. The genus Narcissus belongs to the Narcisseae tribe, one of 13 within the Amaryllidoideae subfamily of the Amaryllidaceae. It is one of two sister clades corresponding to genera in the Narcisseae, being distinguished from Sternbergia by the presence of a paraperigonium, and is monophyletic

 

The infrageneric phylogeny of Narcissus still remains relatively unsettled. The taxonomy has proved very complex and difficult to resolve, particularly for the Pseudonarcissus group. This is due to a number of factors, including the diversity of the wild species, the ease with which natural hybridisation occurs, and extensive cultivation and breeding accompanied by escape and naturalisation.

 

De Candolle, in the first systematic taxonomy of Narcissus, arranged the species into named groups, and those names (Faux-Narcisse or Pseudonarcissus, Poétiques, Tazettes, Bulbocodiens, Jonquilles) have largely endured for the various subdivisions since and bear his name. The evolution of classification was confused by including many unknown or garden varieties, until Baker (1875) made the important distinction of excluding all specimens except the wild species from his system. He then grouped all of the earlier related genera as sections under one genus, Narcissus, the exception being the monotypic Tapeinanthus. Consequently, the number of accepted species has varied widely.

 

A common modern classification system has been that of Fernandes (1951, 1968, 1975) based on cytology, as modified by Blanchard (1990) and Mathew (2002), although in some countries such as Germany, the system of Meyer (1966) was preferred. Fernandes described two subgenera based on basal chromosome number, Hermione, n = 5 (11) and Narcissus, n = 7 (13). He further subdivided these into ten sections (Apodanthi, Aurelia, Bulbocodii, Ganymedes, Jonquillae, Narcissus, Pseudonarcissi, Serotini, Tapeinanthus, Tazettae), as did Blanchard later.

 

In contrast to Fernandes, Webb's treatment of the genus for the Flora Europaea (1978, 1980) prioritised morphology over genetics, and abandoned the subgenera ranks. He also restored De Candolle's original nomenclature, and made a number of changes to section Jonquilla, merging the existing subsections, reducing Apodanthi to a subsection of Jonquilla, and moving N. viridiflorus from Jonquilla to a new monotypic section of its own (Chloranthi). Finally, he divided Pseudonarcissus into two subsections. Blanchard (1990), whose Narcissus: a guide to wild daffodils has been very influential, adopted a simple approach, restoring Apodanthae, and based largely on ten sections alone.

 

The Royal Horticultural Society (RHS) currently lists ten sections, based on Fernandes (1968), three of which are monotypic (contain only one species), while two others only containing two species. Most species are placed in Pseudonarcissus While infrageneric groupings within Narcissus have been relatively constant, their status (genera, subgenera, sections, subsections, series, species) has not. Some authors treat some sections as being further subdivided into subsections, e.g. Tazettae (3 subsections). These subdivisions correspond roughly to the popular names for narcissi types, e.g. Trumpet Daffodils, Tazettas, Pheasant's Eyes, Hoop Petticoats, Jonquils.

 

While Webb had simply divided the genus into sections, Mathew found this unsatisfactory, implying every section had equal status. He adapted both Fernandes and Webb to devise a more hierarchical scheme he believed better reflected the interrelationships within the genus. Mathew's scheme consists of three subgenera (Narcissus, Hermione and Corbularia). The first two subgenera were then divided into five and two sections respectively. He then further subdivided two of the sections (subgenus Narcissus section Jonquillae, and subgenus Hermione section Hermione) into three subsections each. Finally, he divided section Hermione subsection Hermione further into two series, Hermione and Albiflorae. While lacking a phylogenetic basis, the system is still in use in horticulture. For instance the Pacific Bulb Society uses his numbering system for classifying species.

 

The phylogenetic analysis of Graham and Barrett (2004) supported the infrageneric division of Narcissus into two clades corresponding to the subgenera Hermione and Narcissus, but does not support monophyly of all sections, with only Apodanthi demonstrating clear monophyly, corresponding to Clade III of Graham and Barrett, although some other clades corresponded approximately to known sections. These authors examined 36 taxa of the 65 listed then, and a later extended analysis by Rønsted et al. (2008) with five additional taxa confirmed this pattern.

 

A very large (375 accessions) molecular analysis by Zonneveld (2008) utilising nuclear DNA content sought to reduce some of the paraphyly identified by Graham and Barrett. This led to a revision of the sectional structure, shifting some species between sections, eliminating one section and creating two new ones. In subgenus Hermione, Aurelia was merged with Tazettae. In subgenus Narcissus section Jonquillae subsection Juncifolii was elevated to sectional rank, thus resolving the paraphyly in this section observed by Graham and Barrett in Clade II due to this anomalous subsection, the remaining species being in subsection Jonquillae, which was monophyletic. The relatively large section Pseudonarcissi was divided by splitting off a new section, Nevadensis (species from southern Spain) leaving species from France, northern Spain and Portugal in the parent section. At the same time Fernández-Casas (2008) proposed a new monotypic section Angustini to accommodate Narcissus deficiens, placing it within subgenus Hermione.

 

While Graham and Barrett (2004) had determined that subgenus Hermione was monophyletic, using a much larger accession Santos-Gally et al. (2011) did not. However the former had excluded species of hybrid origins, while the latter included both N. dubius and N. tortifolius. If these two species are excluded (forming a clade with subgenus Narcissus) then Hermione can be considered monophyletic, although as a section of Hermione, Tazettae is not monophyletic. They also confirmed the monophyly of Apodanthi.

 

Some so-called nothosections have been proposed, predominantly by Fernández-Casas, to accommodate natural ('ancient') hybrids (nothospecies).

 

Subgenera and sections

Showing revisions by Zonnefeld (2008)

 

subgenus Hermione (Haw.) Spach.

(Aurelia (Gay) Baker (monotypic) - merged with Tazettae (2008)

Serotini Parlatore (2 species)

Tazettae de Candolle (16 species) syn. Hermione (Salisbury) Sprengel, in Fernandes' scheme. Incorporating Aurelia (2008)

subgenus Narcissus L.

Apodanthi A. Fernandes (6 species)

Bulbocodium de Candolle (11 species)

Ganymedes (Haworth) Schultes f. (monotypic)

Jonquillae de Candolle (8 species)

Juncifolii (A. Fern.) Zonn. sect. nov. (2008)

Narcissus L. (2 species)

Nevadensis Zonn. sect. nov. (2008)

Pseudonarcissus de Candolle (36 species) Trumpet daffodils

Tapeinanthus (Herbert) Traub (monotypic)

 

Species

Estimates of the number of species in Narcissus have varied widely, from anywhere between 16 and nearly 160, even in the modern era. Linnaeus originally included six species in 1753. By the time of the 14th edition of the Systema Naturae in 1784, there were fourteen. The 1819 Encyclopaedia Londinensis lists sixteen (see illustration here of three species) and by 1831 Adrian Haworth had described 150 species.

 

Much of the variation lies in the definition of species, and whether closely related taxa are considered separate species or subspecies. Thus, a very wide view of each species, such as Webb's results in few species, while a very narrow view such as that of Fernandes results in a larger number. Another factor is the status of hybrids, given natural hybridisation, with a distinction between 'ancient hybrids' and 'recent hybrids'. The term 'ancient hybrid' refers to hybrids found growing over a large area, and therefore now considered as separate species, while 'recent hybrid' refers to solitary plants found amongst their parents, with a more restricted range.

 

In the twentieth century Fernandes (1951) accepted 22 species, on which were based the 27 species listed by Webb in the 1980 Flora Europaea. By 1968, Fernandes had accepted 63 species, and by 1990 Blanchard listed 65 species, and Erhardt 66 in 1993. In 2006 the Royal Horticultural Society's (RHS) International Daffodil Register and Classified List listed 87 species, while Zonneveld's genetic study (2008) resulted in only 36. As of September 2014, the World Checklist of Selected Plant Families accepts 52 species, along with at least 60 hybrids, while the RHS has 81 accepted names in its October 2014 list.

 

Evolution

Within the Narcisseae, Narcissus (western Mediterranean) diverged from Sternbergia (Eurasia) some time in the Late Oligocene to Early Miocene eras, around 29.3–18.1 Ma, with a best estimate of 23.6 Ma. Later the genus divided into the two subgenera (Hermione and Narcissus) between 27.4 and 16.1 Ma (21.4 Ma). The divisions between the sections of Hermione then took place during the Miocene period 19.9–7.8 Ma.

 

Narcissus appears to have arisen in the area of the Iberian peninsula, southern France and northwestern Italy, and within this area most sections of the genus appeared, with only a few taxa being dispersed to North Africa at a time when the African and West European platforms were closer together. Subgenus Hermione in turn arose in the southwestern mediterranean and north west Africa. However, these are reconstructions, the Amaryllidaceae lacking a fossil record.

 

Names and etymology

The derivation of the Latin narcissus (Greek: νάρκισσος) is unknown. It may be a loanword from another language; for instance, it is said to be related to the Sanskrit word nark, meaning 'hell'. It is frequently linked to the Greek myth of Narcissus described by Ovid in his Metamorphoses, who became so obsessed with his own reflection that as he knelt and gazed into a pool of water, he fell into the water and drowned. In some variations, he died of starvation and thirst. In both versions, the narcissus plant sprang from where he died. Although Ovid appeared to describe the plant we now know as Narcissus there is no evidence for this popular derivation, and the person's name may have come from the flower's name. The Poet's Narcissus (N. poeticus), which grows in Greece, has a fragrance that has been described as intoxicating. This explanation is largely discredited due to lack of proof. Pliny wrote that the plant 'narce narcissum dictum, non a fabuloso puero' ('named narcissus from narce, not from the legendary youth'), i.e. that it was named for its narcotic properties (ναρκάω narkao, 'I grow numb' in Greek), not from the legend. Furthermore, there were accounts of narcissi growing, such as in the legend of Persephone, long before the story of Narcissus appeared (see Greek culture). It has also been suggested that daffodils bending over streams evoked the image of the youth admiring his own reflection in the water.

 

Linnaeus used the Latin name for the plant in formally describing the genus, although Matthias de l'Obel had previously used the name in describing various species of Narcissi in his Icones stirpium of 1591, and other publications, as had Clusius in Rariorum stirpium (1576).

 

The plural form of the common name narcissus has caused some confusion. British English sources such as the Shorter Oxford English Dictionary give two alternate forms, narcissi and narcissuses. In contrast, in American English the Merriam-Webster Dictionary provides for a third form, narcissus, used for both singular and plural. The Oxford dictionaries only list this third form under American English, although the Cambridge Dictionary allows of all three in the same order. However, Garner's Modern American Usage states that narcissi is the commonest form, narcissuses being excessively sibilant. For similar reasons, Fowler prefers narcissi in British English usage. Neither support narcissus as a plural form. Common names such as narcissus do not capitalise the first letter in contrast to the person of that name and the Latin genus name.

 

The name Narcissus (feminine Narcissa) was not uncommon in Roman times, such as Tiberius Claudius Narcissus, a Roman official in Claudius' time, an early New Testament Christian in Rome and later bishops and saints.

 

Daffodil

The word daffodil was unknown in the English language before the sixteenth century. The name is derived from an earlier affodell, a variant of asphodel. In classical Greek literature the narcissus is frequently referred to as the asphodel, such as the meadows of the Elysian fields in Homer (see Antiquity). Asphodel in turn appears to be a loanword coming from French via Mediaeval Latin affodilus from Classical Latin asphodilus and ultimately the Greek asphodelos (Greek: ἀσφόδελος). The reason for the introduction of the initial d is not known, although a probable source is an etymological merging from the Dutch article de, as in de affodil, or English the, as th'affodil or t'affodil, hence daffodil, and in French de and affodil to form fleur d'aphrodille and daphrodille.

 

From at least the 16th century, daffadown dilly and daffydowndilly have appeared as playful synonyms of the name. In common parlance and in historical documents, the term daffodil may refer specifically to populations or specimens of the wild daffodil, N. pseudonarcissus. H. N. Ellacombe suggests this may be from Saffon Lilly, citing Prior in support, though admittedly conjectural.

 

Lady Wilkinson (1858), who provides an extensive discussion of the etymology of the various names for this plant, suggests a very different origin, namely the Old English word affodyle (that which cometh early), citing a 14th-century (but likely originally much earlier) manuscript in support of this theory, and which appears to describe a plant resembling the daffodil. Ellacombe provides further support for this from a fifteenth century English translation of Palladius that also refers to it.

 

Jonquil

The name jonquil is said to be a corruption via French from the Latin juncifolius meaning 'rush-leaf' (Juncaceae) and its use is generally restricted to those species and cultivars which have rush like leaves, e.g. N. juncifolius.

 

Other

A profusion of names have attached themselves in the English language, either to the genus as a whole or to individual species or groups of species such as sections. These include narcissus, jonquil, Lent lily, Lenten lily, lide lily, yellow lily, wort or wyrt, Julians, glens, Lent cocks, corn flower, bell rose, asphodel, Solomon's lily, gracy day, haverdrils, giggary, cowslip, and crow foot.

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Narcissus is a genus of predominantly spring flowering perennial plants of the amaryllis family, Amaryllidaceae. Various common names including daffodil, narcissus, and jonquil are used to describe all or some members of the genus. Narcissus has conspicuous flowers with six petal-like tepals surmounted by a cup- or trumpet-shaped corona. The flowers are generally white and yellow (also orange or pink in garden varieties), with either uniform or contrasting coloured tepals and corona.

 

Narcissus were well known in ancient civilisation, both medicinally and botanically, but formally described by Linnaeus in his Species Plantarum (1753). The genus is generally considered to have about ten sections with approximately 50 species. The number of species has varied, depending on how they are classified, due to similarity between species and hybridisation. The genus arose some time in the Late Oligocene to Early Miocene epochs, in the Iberian peninsula and adjacent areas of southwest Europe. The exact origin of the name Narcissus is unknown, but it is often linked to a Greek word (ancient Greek ναρκῶ narkō, "to make numb") and the myth of the youth of that name who fell in love with his own reflection. The English word "daffodil" appears to be derived from "asphodel", with which it was commonly compared.

 

The species are native to meadows and woods in southern Europe and North Africa with a centre of diversity in the Western Mediterranean, particularly the Iberian peninsula. Both wild and cultivated plants have naturalised widely, and were introduced into the Far East prior to the tenth century. Narcissi tend to be long-lived bulbs, which propagate by division, but are also insect-pollinated. Known pests, diseases and disorders include viruses, fungi, the larvae of flies, mites and nematodes. Some Narcissus species have become extinct, while others are threatened by increasing urbanisation and tourism.

 

Historical accounts suggest narcissi have been cultivated from the earliest times, but became increasingly popular in Europe after the 16th century and by the late 19th century were an important commercial crop centred primarily in the Netherlands. Today narcissi are popular as cut flowers and as ornamental plants in private and public gardens. The long history of breeding has resulted in thousands of different cultivars. For horticultural purposes, narcissi are classified into divisions, covering a wide range of shapes and colours. Like other members of their family, narcissi produce a number of different alkaloids, which provide some protection for the plant, but may be poisonous if accidentally ingested. This property has been exploited for medicinal use in traditional healing and has resulted in the production of galantamine for the treatment of Alzheimer's dementia. Long celebrated in art and literature, narcissi are associated with a number of themes in different cultures, ranging from death to good fortune, and as symbols of spring.

 

The daffodil is the national flower of Wales and the symbol of cancer charities in many countries. The appearance of the wild flowers in spring is associated with festivals in many places.

 

Narcissus is a genus of perennial herbaceous bulbiferous geophytes, which die back after flowering to an underground storage bulb. They regrow in the following year from brown-skinned ovoid bulbs with pronounced necks, and reach heights of 5–80 centimetres (2.0–31.5 in) depending on the species. Dwarf species such as N. asturiensis have a maximum height of 5–8 centimetres (2.0–3.1 in), while Narcissus tazetta may grow as tall as 80 centimetres (31 in).

 

The plants are scapose, having a single central leafless hollow flower stem (scape). Several green or blue-green, narrow, strap-shaped leaves arise from the bulb. The plant stem usually bears a solitary flower, but occasionally a cluster of flowers (umbel). The flowers, which are usually conspicuous and white or yellow, sometimes both or rarely green, consist of a perianth of three parts. Closest to the stem (proximal) is a floral tube above the ovary, then an outer ring composed of six tepals (undifferentiated sepals and petals), and a central disc to conical shaped corona. The flowers may hang down (pendant), or be erect. There are six pollen bearing stamens surrounding a central style. The ovary is inferior (below the floral parts) consisting of three chambers (trilocular). The fruit consists of a dry capsule that splits (dehisces) releasing numerous black seeds.

 

The bulb lies dormant after the leaves and flower stem die back and has contractile roots that pull it down further into the soil. The flower stem and leaves form in the bulb, to emerge the following season. Most species are dormant from summer to late winter, flowering in the spring, though a few species are autumn flowering.

 

The pale brown-skinned ovoid tunicate bulbs have a membranous tunic and a corky stem (base or basal) plate from which arise the adventitious root hairs in a ring around the edge, which grow up to 40 mm in length. Above the stem plate is the storage organ consisting of bulb scales, surrounding the previous flower stalk and the terminal bud. The scales are of two types, true storage organs and the bases of the foliage leaves. These have a thicker tip and a scar from where the leaf lamina became detached. The innermost leaf scale is semicircular only partly enveloping the flower stalk (semisheathed).(see Hanks Fig 1.3). The bulb may contain a number of branched bulb units, each with two to three true scales and two to three leaf bases. Each bulb unit has a life of about four years.

 

Once the leaves die back in summer, the roots also wither. After some years, the roots shorten pulling the bulbs deeper into the ground (contractile roots). The bulbs develop from the inside, pushing the older layers outwards which become brown and dry, forming an outer shell, the tunic or skin. Up to 60 layers have been counted in some wild species. While the plant appears dormant above the ground the flower stalk which will start to grow in the following spring, develops within the bulb surrounded by two to three deciduous leaves and their sheaths. The flower stem lies in the axil of the second true leaf.

 

The single leafless Plant stem stem or scape, appearing from early to late spring depending on the species, bears from 1 to 20 blooms. Stem shape depends on the species, some are highly compressed with a visible seam, while others are rounded. The stems are upright and located at the centre of the leaves. In a few species such as N. hedraeanthus the stem is oblique. The stem is hollow in the upper portion but towards the bulb is more solid and filled with a spongy material.

 

Narcissus plants have one to several basal leaf leaves which are linear, ligulate or strap-shaped (long and narrow), sometimes channelled adaxially to semiterete, and may (pedicellate) or may not (sessile) have a petiole stalk. The leaves are flat and broad to cylindrical at the base and arise from the bulb. The emerging plant generally has two leaves, but the mature plant usually three, rarely four, and they are covered with a cutin containing cuticle, giving them a waxy appearance. Leaf colour is light green to blue-green. In the mature plant, the leaves extend higher than the flower stem, but in some species, the leaves are low-hanging. The leaf base is encased in a colorless sheath. After flowering, the leaves turn yellow and die back once the seed pod (fruit) is ripe.

 

Jonquils usually have dark green, round, rush-like leaves.

 

The inflorescence is scapose, the single stem or scape bearing either a solitary flower or forming an umbel with up to 20 blooms. Species bearing a solitary flower include section Bulbocodium and most of section Pseudonarcissus. Umbellate species have a fleshy racemose inflorescence (unbranched, with short floral stalks) with 2 to 15 or 20 flowers, such as N. papyraceus (see illustration, left) and N. tazetta (see Table I). The flower arrangement on the inflorescence may be either with (pedicellate) or without (sessile) floral stalks.

 

Prior to opening, the flower buds are enveloped and protected in a thin dry papery or membranous (scarious) spathe. The spathe consists of a singular bract that is ribbed, and which remains wrapped around the base of the open flower. As the bud grows, the spathe splits longitudinally. Bracteoles are small or absent.

 

The flowers of Narcissus are hermaphroditic (bisexual), have three parts (tripartite), and are sometimes fragrant (see Fragrances). The flower symmetry is actinomorphic (radial) to slightly zygomorphic (bilateral) due to declinate-ascending stamens (curving downwards, then bent up at the tip). Narcissus flowers are characterised by their, usually conspicuous, corona (trumpet).

 

The three major floral parts (in all species except N. cavanillesii in which the corona is virtually absent - Table I: Section Tapeinanthus) are;

 

(i) the proximal floral tube (hypanthium),

(ii) the surrounding free tepals, and

(iii) the more distal corona (paraperigon, paraperigonium).

All three parts may be considered to be components of the perianth (perigon, perigonium). The perianth arises above the apex of the inferior ovary, its base forming the hypanthial floral tube.

 

The floral tube is formed by fusion of the basal segments of the tepals (proximally connate). Its shape is from an inverted cone (obconic) to funnel-shaped (funneliform) or cylindrical, and is surmounted by the more distal corona. Floral tubes can range from long and narrow sections Apodanthi and Jonquilla to rudimentary (N. cavanillesii).

 

Surrounding the floral tube and corona and reflexed (bent back) from the rest of the perianth are the six spreading tepals or floral leaves, in two whorls which may be distally ascending, reflexed (folded back), or lanceolate. Like many monocotyledons, the perianth is homochlamydeous, which is undifferentiated into separate calyx (sepals) and corolla (petals), but rather has six tepals. The three outer tepal segments may be considered sepals, and the three inner segments petals. The transition point between the floral tube and the corona is marked by the insertion of the free tepals on the fused perianth.

 

The corona, or paracorolla, is variously described as bell-shaped (funneliform, trumpet), bowl-shaped (cupular, crateriform, cup-shaped) or disc-shaped with margins that are often frilled, and is free from the stamens. Rarely is the corona a simple callose (hardened, thickened) ring. The corona is formed during floral development as a tubular outgrowth from stamens which fuse into a tubular structure, the anthers becoming reduced. At its base, the fragrances which attract pollinators are formed. All species produce nectar at the top of the ovary. Coronal morphology varies from the tiny pigmented disk of N. serotinus (see Table I) or the rudimentary structure in N. cavanillesii to the elongated trumpets of section Pseudonarcissus (trumpet daffodils, Table I).

 

While the perianth may point forwards, in some species such as N. cyclamineus it is folded back (reflexed, see illustration, left), while in some other species such as N. bulbocodium (Table I), it is reduced to a few barely visible pointed segments with a prominent corona.

 

The colour of the perianth is white, yellow or bicoloured, with the exception of the night flowering N. viridiflorus which is green. In addition the corona of N. poeticus has a red crenulate margin (see Table I). Flower diameter varies from 12 mm (N. bulbocodium) to over 125 mm (N. nobilis=N. pseudonarcissus subsp. nobilis).

 

Flower orientation varies from pendent or deflexed (hanging down) as in N. triandrus (see illustration, left), through declinate-ascendant as in N. alpestris = N. pseudonarcissus subsp. moschatus, horizontal (patent, spreading) such as N. gaditanus or N. poeticus, erect as in N. cavanillesii, N. serotinus and N. rupicola (Table I), or intermediate between these positions (erecto-patent).

 

The flowers of Narcissus demonstrate exceptional floral diversity and sexual polymorphism, primarily by corona size and floral tube length, associated with pollinator groups (see for instance Figs. 1 and 2 in Graham and Barrett). Barrett and Harder (2005) describe three separate floral patterns;

 

"Daffodil" form

"Paperwhite" form

"Triandrus" form.

The predominant patterns are the 'daffodil' and 'paperwhite' forms, while the "triandrus" form is less common. Each corresponds to a different group of pollinators (See Pollination).

 

The "daffodil" form, which includes sections Pseudonarcissus and Bulbocodium, has a relatively short, broad or highly funnelform tube (funnel-like), which grades into an elongated corona, which is large and funnelform, forming a broad, cylindrical or trumpet-shaped perianth. Section Pseudonarcissus consists of relatively large flowers with a corolla length of around 50 mm, generally solitary but rarely in inflorescences of 2–4 flowers. They have wide greenish floral tubes with funnel-shaped bright yellow coronas. The six tepals sometimes differ in colour from the corona and may be cream coloured to pale yellow.

 

The "paperwhite" form, including sections Jonquilla, Apodanthi and Narcissus, has a relatively long, narrow tube and a short, shallow, flaring corona. The flower is horizontal and fragrant.

 

The "triandrus" form is seen in only two species, N. albimarginatus (a Moroccan endemic) and N. triandrus. It combines features of both the "daffodil" and "paperwhite" forms, with a well-developed, long, narrow tube and an extended bell-shaped corona of almost equal length. The flowers are pendent.

 

Androecium

There are six stamens in one to two rows (whorls), with the filaments separate from the corona, attached at the throat or base of the tube (epipetalous), often of two separate lengths, straight or declinate-ascending (curving downwards, then upwards). The anthers are basifixed (attached at their base).

 

Gynoecium

The ovary is inferior (below the floral parts) and trilocular (three chambered) and there is a pistil with a minutely three lobed stigma and filiform (thread like) style, which is often exserted (extending beyond the tube).

 

Fruit

The fruit consists of dehiscent loculicidal capsules (splitting between the locules) that are ellipsoid to subglobose (almost spherical) in shape and are papery to leathery in texture.

 

Seeds

The fruit contains numerous subglobose seeds which are round and swollen with a hard coat, sometimes with an attached elaiosome. The testa is black and the pericarp dry.

 

Most species have 12 ovules and 36 seeds, although some species such as N. bulbocodium have more, up to a maximum of 60. Seeds take five to six weeks to mature. The seeds of sections Jonquilla and Bulbocodium are wedge-shaped and matte black, while those of other sections are ovate and glossy black. A gust of wind or contact with a passing animal is sufficient to disperse the mature seeds.

 

Chromosomes

Chromosome numbers include 2n=14, 22, 26, with numerous aneuploid and polyploid derivatives. The basic chromosome number is 7, with the exception of N. tazetta, N. elegans and N. broussonetii in which it is 10 or 11; this subgenus (Hermione) was in fact characterised by this characteristic. Polyploid species include N. papyraceus (4x=22) and N. dubius (6x=50).

 

Phytochemistry

Alkaloids

As with all Amarylidaceae genera, Narcissus contains unique isoquinoline alkaloids. The first alkaloid to be identified was lycorine, from N. pseudonarcissus in 1877. These are considered a protective adaptation and are utilised in the classification of species. Nearly 100 alkaloids have been identified in the genus, about a third of all known Amaryllidaceae alkaloids, although not all species have been tested. Of the nine alkaloid ring types identified in the family, Narcissus species most commonly demonstrate the presence of alkaloids from within the Lycorine (lycorine, galanthine, pluviine) and Homolycorine (homolycorine, lycorenine) groups. Hemanthamine, tazettine, narciclasine, montanine and galantamine alkaloids are also represented. The alkaloid profile of any plant varies with time, location, and developmental stage. Narcissus also contain fructans and low molecular weight glucomannan in the leaves and plant stems.

 

Fragrances

Fragrances are predominantly monoterpene isoprenoids, with a small amount of benzenoids, although N. jonquilla has both equally represented. Another exception is N. cuatrecasasii which produces mainly fatty acid derivatives. The basic monoterpene precursor is geranyl pyrophosphate, and the commonest monoterpenes are limonene, myrcene, and trans-β-ocimene. Most benzenoids are non-methoxylated, while a few species contain methoxylated forms (ethers), e.g. N. bujei. Other ingredient include indole, isopentenoids and very small amounts of sesquiterpenes. Fragrance patterns can be correlated with pollinators, and fall into three main groups (see Pollination).

 

The taxonomy of Narcissus is complex, and still not fully resolved. Known to the ancients, the genus name appears in Graeco-Roman literature, although their interest was as much medicinal as botanical. It is unclear which species the ancients were familiar with. Although frequently mentioned in Mediaeval and Renaissance texts it was not formally described till the work of Linnaeus in 1753. By 1789 it had been grouped into a family (Narcissi) but shortly thereafter this was renamed Amaryllideae, from which comes the modern placement within Amaryllidaceae, although for a while it was considered part of Liliaceae.

 

Many of the species now considered to be Narcissus were in separate genera during the nineteenth century, and the situation was further confused by the inclusion of many cultivated varieties. By 1875 the current circumscription was relatively settled. By 2004 phylogenetic studies had allowed the place of Narcissus within its fairly large family to be established, nested within a series of subfamilies (Amaryllidoideae) and tribes (Narcisseae). It shares its position in the latter tribe with Sternbergia.

 

The infrageneric classification has been even more complex and many schemes of subgenera, sections, subsections and series have been proposed, although all had certain similarities. Most authorities now consider there to be 10 – 11 sections based on phylogenetic evidence. The problems have largely arisen from the diversity of the wild species, frequent natural hybridisation and extensive cultivation with escape and subsequent naturalisation. The number of species has varied anywhere from 16 to nearly 160, but is probably around 50 – 60.

 

The genus appeared some time in the Late Oligocene to Early Miocene eras, around 24 million years ago, in the Iberian peninsula. While the exact origin of the word Narcissus is unknown it is frequently linked to its fragrance which was thought to be narcotic, and to the legend of the youth of that name who fell in love with his reflection. In the English language the common name Daffodil appears to be derived from the Asphodel with which it was commonly compared.

 

Early

Narcissus was first described by Theophrastus (Θεόφραστος, c 371 - c 287 BC) in his Historia Plantarum (Greek: Περὶ φυτῶν ἱστορία) as νάρκισσος, referring to N. poeticus, but comparing it to Asphodelus (ασφοδελωδες). Theophrastus' description was frequently referred to at length by later authors writing in Latin such as Pliny the Elder (Gaius Plinius Secundus, 23 AD – 79 AD) from whom came the Latin form narcissus (see also Culture). Pliny's account is from his Natural History (Latin: Naturalis Historia). Like his contemporaries, his interests were as much therapeutic as botanical. Another much-cited Greek authority was Dioscorides (Διοσκουρίδης, 40 AD – 90 AD) in his De Materia Medica (Greek: Περὶ ὕλης ἰατρικῆς). Both authors were to remain influential until at least the Renaissance, given that their descriptions went beyond the merely botanical, to the therapeutic (see also Antiquity).

 

An early European reference is found in the work of Albert Magnus (c. 1200 – 1280), who noted in his De vegetabilibus et plantis the similarity to the leek. William Turner in his A New Herball (1551) cites all three extensively in his description of the plant and its properties.It was to remain to Linnaeus in 1753 to formally describe and name Narcissus as a genus in his Species Plantarum, at which time there were six known species (N. poeticus, N. pseudonarcissus, N. bulbocodium, N. serotinus, N. jonquilla and N. tazetta).[1] At that time, Linnaeus loosely grouped it together with 50 other genera into his Hexandria monogynia.

 

Modern

It was de Jussieu in 1789 who first formally created a 'family' (Narcissi), as the seventh 'Ordo' (Order) of the third class (Stamina epigyna) of Monocots in which Narcissus and 15 other genera were placed. The use of the term Ordo at that time was closer to what we now understand as Family, rather than Order. The family has undergone much reorganisation since then, but in 1805 it was renamed after a different genus in the family, Amaryllis, as 'Amaryllideae' by Jaume St.-Hilaire and has retained that association since. Jaume St.-Hilaire divided the family into two unnamed sections and recognised five species of Narcissus, omitting N. serotinus.

 

De Candolle brought together Linnaeus' genera and Jussieau's families into a systematic taxonomy for the first time, but included Narcissus (together with Amaryllis) in the Liliaceae in his Flore française (1805-1815) rather than Amaryllidaceae, a family he had not yet recognised. Shortly thereafter he separated the 'Amaryllidées' from 'Liliacées' (1813), though attributing the term to Brown's 'Amaryllideae' in the latter's Prodromus (1810) rather than St.-Hilaire's 'Amaryllidées'. He also provided the text to the first four volumes of Redouté illustrations in the latter's Les liliacées between 1805 and 1808 (see illustration here of N. candidissimus).

 

Historically both wide and narrow interpretations of the genus have been proposed. In the nineteenth century genus splitting was common, favouring the narrow view. Haworth (1831) using a narrow view treated many species as separate genera, as did Salisbury (1866). These authors listed various species in related genera such as Queltia (hybrids), Ajax (=Pseudonarcissus) and Hermione (=Tazettae), sixteen in all in Haworth's classification. In contrast, Herbert (1837) took a very wide view reducing Harworth's sixteen genera to six. Herbert, treating the Amaryllidacea as an 'order' as was common then, considered the narcissi to be a suborder, the Narcisseae, the six genera being Corbularia, Ajax, Ganymedes, Queltia, Narcissus and Hermione and his relatively narrow circumscription of Narcissus having only three species. Later Spach (1846) took an even wider view bringing most of Harworth's genera into the genus Narcissus, but as separate subgenera. By the time that Baker (1875) wrote his monograph all of the genera with one exception were included as Narcissus. The exception was the monotypic group Tapeinanthus which various subsequent authors have chosen to either exclude (e.g. Cullen 1986) or include (e.g. Webb 1978, 1980). Today it is nearly always included.

 

The eventual position of Narcissus within the Amaryllidaceae family only became settled in the twenty-first century with the advent of phylogenetic analysis and the Angiosperm Phylogeny Group system. The genus Narcissus belongs to the Narcisseae tribe, one of 13 within the Amaryllidoideae subfamily of the Amaryllidaceae. It is one of two sister clades corresponding to genera in the Narcisseae, being distinguished from Sternbergia by the presence of a paraperigonium, and is monophyletic

 

The infrageneric phylogeny of Narcissus still remains relatively unsettled. The taxonomy has proved very complex and difficult to resolve, particularly for the Pseudonarcissus group. This is due to a number of factors, including the diversity of the wild species, the ease with which natural hybridisation occurs, and extensive cultivation and breeding accompanied by escape and naturalisation.

 

De Candolle, in the first systematic taxonomy of Narcissus, arranged the species into named groups, and those names (Faux-Narcisse or Pseudonarcissus, Poétiques, Tazettes, Bulbocodiens, Jonquilles) have largely endured for the various subdivisions since and bear his name. The evolution of classification was confused by including many unknown or garden varieties, until Baker (1875) made the important distinction of excluding all specimens except the wild species from his system. He then grouped all of the earlier related genera as sections under one genus, Narcissus, the exception being the monotypic Tapeinanthus. Consequently, the number of accepted species has varied widely.

 

A common modern classification system has been that of Fernandes (1951, 1968, 1975) based on cytology, as modified by Blanchard (1990) and Mathew (2002), although in some countries such as Germany, the system of Meyer (1966) was preferred. Fernandes described two subgenera based on basal chromosome number, Hermione, n = 5 (11) and Narcissus, n = 7 (13). He further subdivided these into ten sections (Apodanthi, Aurelia, Bulbocodii, Ganymedes, Jonquillae, Narcissus, Pseudonarcissi, Serotini, Tapeinanthus, Tazettae), as did Blanchard later.

 

In contrast to Fernandes, Webb's treatment of the genus for the Flora Europaea (1978, 1980) prioritised morphology over genetics, and abandoned the subgenera ranks. He also restored De Candolle's original nomenclature, and made a number of changes to section Jonquilla, merging the existing subsections, reducing Apodanthi to a subsection of Jonquilla, and moving N. viridiflorus from Jonquilla to a new monotypic section of its own (Chloranthi). Finally, he divided Pseudonarcissus into two subsections. Blanchard (1990), whose Narcissus: a guide to wild daffodils has been very influential, adopted a simple approach, restoring Apodanthae, and based largely on ten sections alone.

 

The Royal Horticultural Society (RHS) currently lists ten sections, based on Fernandes (1968), three of which are monotypic (contain only one species), while two others only containing two species. Most species are placed in Pseudonarcissus While infrageneric groupings within Narcissus have been relatively constant, their status (genera, subgenera, sections, subsections, series, species) has not. Some authors treat some sections as being further subdivided into subsections, e.g. Tazettae (3 subsections). These subdivisions correspond roughly to the popular names for narcissi types, e.g. Trumpet Daffodils, Tazettas, Pheasant's Eyes, Hoop Petticoats, Jonquils.

 

While Webb had simply divided the genus into sections, Mathew found this unsatisfactory, implying every section had equal status. He adapted both Fernandes and Webb to devise a more hierarchical scheme he believed better reflected the interrelationships within the genus. Mathew's scheme consists of three subgenera (Narcissus, Hermione and Corbularia). The first two subgenera were then divided into five and two sections respectively. He then further subdivided two of the sections (subgenus Narcissus section Jonquillae, and subgenus Hermione section Hermione) into three subsections each. Finally, he divided section Hermione subsection Hermione further into two series, Hermione and Albiflorae. While lacking a phylogenetic basis, the system is still in use in horticulture. For instance the Pacific Bulb Society uses his numbering system for classifying species.

 

The phylogenetic analysis of Graham and Barrett (2004) supported the infrageneric division of Narcissus into two clades corresponding to the subgenera Hermione and Narcissus, but does not support monophyly of all sections, with only Apodanthi demonstrating clear monophyly, corresponding to Clade III of Graham and Barrett, although some other clades corresponded approximately to known sections. These authors examined 36 taxa of the 65 listed then, and a later extended analysis by Rønsted et al. (2008) with five additional taxa confirmed this pattern.

 

A very large (375 accessions) molecular analysis by Zonneveld (2008) utilising nuclear DNA content sought to reduce some of the paraphyly identified by Graham and Barrett. This led to a revision of the sectional structure, shifting some species between sections, eliminating one section and creating two new ones. In subgenus Hermione, Aurelia was merged with Tazettae. In subgenus Narcissus section Jonquillae subsection Juncifolii was elevated to sectional rank, thus resolving the paraphyly in this section observed by Graham and Barrett in Clade II due to this anomalous subsection, the remaining species being in subsection Jonquillae, which was monophyletic. The relatively large section Pseudonarcissi was divided by splitting off a new section, Nevadensis (species from southern Spain) leaving species from France, northern Spain and Portugal in the parent section. At the same time Fernández-Casas (2008) proposed a new monotypic section Angustini to accommodate Narcissus deficiens, placing it within subgenus Hermione.

 

While Graham and Barrett (2004) had determined that subgenus Hermione was monophyletic, using a much larger accession Santos-Gally et al. (2011) did not. However the former had excluded species of hybrid origins, while the latter included both N. dubius and N. tortifolius. If these two species are excluded (forming a clade with subgenus Narcissus) then Hermione can be considered monophyletic, although as a section of Hermione, Tazettae is not monophyletic. They also confirmed the monophyly of Apodanthi.

 

Some so-called nothosections have been proposed, predominantly by Fernández-Casas, to accommodate natural ('ancient') hybrids (nothospecies).

 

Subgenera and sections

Showing revisions by Zonnefeld (2008)

 

subgenus Hermione (Haw.) Spach.

(Aurelia (Gay) Baker (monotypic) - merged with Tazettae (2008)

Serotini Parlatore (2 species)

Tazettae de Candolle (16 species) syn. Hermione (Salisbury) Sprengel, in Fernandes' scheme. Incorporating Aurelia (2008)

subgenus Narcissus L.

Apodanthi A. Fernandes (6 species)

Bulbocodium de Candolle (11 species)

Ganymedes (Haworth) Schultes f. (monotypic)

Jonquillae de Candolle (8 species)

Juncifolii (A. Fern.) Zonn. sect. nov. (2008)

Narcissus L. (2 species)

Nevadensis Zonn. sect. nov. (2008)

Pseudonarcissus de Candolle (36 species) Trumpet daffodils

Tapeinanthus (Herbert) Traub (monotypic)

 

Species

Estimates of the number of species in Narcissus have varied widely, from anywhere between 16 and nearly 160, even in the modern era. Linnaeus originally included six species in 1753. By the time of the 14th edition of the Systema Naturae in 1784, there were fourteen. The 1819 Encyclopaedia Londinensis lists sixteen (see illustration here of three species) and by 1831 Adrian Haworth had described 150 species.

 

Much of the variation lies in the definition of species, and whether closely related taxa are considered separate species or subspecies. Thus, a very wide view of each species, such as Webb's results in few species, while a very narrow view such as that of Fernandes results in a larger number. Another factor is the status of hybrids, given natural hybridisation, with a distinction between 'ancient hybrids' and 'recent hybrids'. The term 'ancient hybrid' refers to hybrids found growing over a large area, and therefore now considered as separate species, while 'recent hybrid' refers to solitary plants found amongst their parents, with a more restricted range.

 

In the twentieth century Fernandes (1951) accepted 22 species, on which were based the 27 species listed by Webb in the 1980 Flora Europaea. By 1968, Fernandes had accepted 63 species, and by 1990 Blanchard listed 65 species, and Erhardt 66 in 1993. In 2006 the Royal Horticultural Society's (RHS) International Daffodil Register and Classified List listed 87 species, while Zonneveld's genetic study (2008) resulted in only 36. As of September 2014, the World Checklist of Selected Plant Families accepts 52 species, along with at least 60 hybrids, while the RHS has 81 accepted names in its October 2014 list.

 

Evolution

Within the Narcisseae, Narcissus (western Mediterranean) diverged from Sternbergia (Eurasia) some time in the Late Oligocene to Early Miocene eras, around 29.3–18.1 Ma, with a best estimate of 23.6 Ma. Later the genus divided into the two subgenera (Hermione and Narcissus) between 27.4 and 16.1 Ma (21.4 Ma). The divisions between the sections of Hermione then took place during the Miocene period 19.9–7.8 Ma.

 

Narcissus appears to have arisen in the area of the Iberian peninsula, southern France and northwestern Italy, and within this area most sections of the genus appeared, with only a few taxa being dispersed to North Africa at a time when the African and West European platforms were closer together. Subgenus Hermione in turn arose in the southwestern mediterranean and north west Africa. However, these are reconstructions, the Amaryllidaceae lacking a fossil record.

 

Names and etymology

The derivation of the Latin narcissus (Greek: νάρκισσος) is unknown. It may be a loanword from another language; for instance, it is said to be related to the Sanskrit word nark, meaning 'hell'. It is frequently linked to the Greek myth of Narcissus described by Ovid in his Metamorphoses, who became so obsessed with his own reflection that as he knelt and gazed into a pool of water, he fell into the water and drowned. In some variations, he died of starvation and thirst. In both versions, the narcissus plant sprang from where he died. Although Ovid appeared to describe the plant we now know as Narcissus there is no evidence for this popular derivation, and the person's name may have come from the flower's name. The Poet's Narcissus (N. poeticus), which grows in Greece, has a fragrance that has been described as intoxicating. This explanation is largely discredited due to lack of proof. Pliny wrote that the plant 'narce narcissum dictum, non a fabuloso puero' ('named narcissus from narce, not from the legendary youth'), i.e. that it was named for its narcotic properties (ναρκάω narkao, 'I grow numb' in Greek), not from the legend. Furthermore, there were accounts of narcissi growing, such as in the legend of Persephone, long before the story of Narcissus appeared (see Greek culture). It has also been suggested that daffodils bending over streams evoked the image of the youth admiring his own reflection in the water.

 

Linnaeus used the Latin name for the plant in formally describing the genus, although Matthias de l'Obel had previously used the name in describing various species of Narcissi in his Icones stirpium of 1591, and other publications, as had Clusius in Rariorum stirpium (1576).

 

The plural form of the common name narcissus has caused some confusion. British English sources such as the Shorter Oxford English Dictionary give two alternate forms, narcissi and narcissuses. In contrast, in American English the Merriam-Webster Dictionary provides for a third form, narcissus, used for both singular and plural. The Oxford dictionaries only list this third form under American English, although the Cambridge Dictionary allows of all three in the same order. However, Garner's Modern American Usage states that narcissi is the commonest form, narcissuses being excessively sibilant. For similar reasons, Fowler prefers narcissi in British English usage. Neither support narcissus as a plural form. Common names such as narcissus do not capitalise the first letter in contrast to the person of that name and the Latin genus name.

 

The name Narcissus (feminine Narcissa) was not uncommon in Roman times, such as Tiberius Claudius Narcissus, a Roman official in Claudius' time, an early New Testament Christian in Rome and later bishops and saints.

 

Daffodil

The word daffodil was unknown in the English language before the sixteenth century. The name is derived from an earlier affodell, a variant of asphodel. In classical Greek literature the narcissus is frequently referred to as the asphodel, such as the meadows of the Elysian fields in Homer (see Antiquity). Asphodel in turn appears to be a loanword coming from French via Mediaeval Latin affodilus from Classical Latin asphodilus and ultimately the Greek asphodelos (Greek: ἀσφόδελος). The reason for the introduction of the initial d is not known, although a probable source is an etymological merging from the Dutch article de, as in de affodil, or English the, as th'affodil or t'affodil, hence daffodil, and in French de and affodil to form fleur d'aphrodille and daphrodille.

 

From at least the 16th century, daffadown dilly and daffydowndilly have appeared as playful synonyms of the name. In common parlance and in historical documents, the term daffodil may refer specifically to populations or specimens of the wild daffodil, N. pseudonarcissus. H. N. Ellacombe suggests this may be from Saffon Lilly, citing Prior in support, though admittedly conjectural.

 

Lady Wilkinson (1858), who provides an extensive discussion of the etymology of the various names for this plant, suggests a very different origin, namely the Old English word affodyle (that which cometh early), citing a 14th-century (but likely originally much earlier) manuscript in support of this theory, and which appears to describe a plant resembling the daffodil. Ellacombe provides further support for this from a fifteenth century English translation of Palladius that also refers to it.

 

Jonquil

The name jonquil is said to be a corruption via French from the Latin juncifolius meaning 'rush-leaf' (Juncaceae) and its use is generally restricted to those species and cultivars which have rush like leaves, e.g. N. juncifolius.

 

Other

A profusion of names have attached themselves in the English language, either to the genus as a whole or to individual species or groups of species such as sections. These include narcissus, jonquil, Lent lily, Lenten lily, lide lily, yellow lily, wort or wyrt, Julians, glens, Lent cocks, corn flower, bell rose, asphodel, Solomon's lily, gracy day, haverdrils, giggary, cowslip, and crow foot.

Narcissus is a genus of predominantly spring flowering perennial plants of the amaryllis family, Amaryllidaceae. Various common names including daffodil, narcissus, and jonquil are used to describe all or some members of the genus. Narcissus has conspicuous flowers with six petal-like tepals surmounted by a cup- or trumpet-shaped corona. The flowers are generally white and yellow (also orange or pink in garden varieties), with either uniform or contrasting coloured tepals and corona.

 

Narcissus were well known in ancient civilisation, both medicinally and botanically, but formally described by Linnaeus in his Species Plantarum (1753). The genus is generally considered to have about ten sections with approximately 50 species. The number of species has varied, depending on how they are classified, due to similarity between species and hybridisation. The genus arose some time in the Late Oligocene to Early Miocene epochs, in the Iberian peninsula and adjacent areas of southwest Europe. The exact origin of the name Narcissus is unknown, but it is often linked to a Greek word (ancient Greek ναρκῶ narkō, "to make numb") and the myth of the youth of that name who fell in love with his own reflection. The English word "daffodil" appears to be derived from "asphodel", with which it was commonly compared.

 

The species are native to meadows and woods in southern Europe and North Africa with a centre of diversity in the Western Mediterranean, particularly the Iberian peninsula. Both wild and cultivated plants have naturalised widely, and were introduced into the Far East prior to the tenth century. Narcissi tend to be long-lived bulbs, which propagate by division, but are also insect-pollinated. Known pests, diseases and disorders include viruses, fungi, the larvae of flies, mites and nematodes. Some Narcissus species have become extinct, while others are threatened by increasing urbanisation and tourism.

 

Historical accounts suggest narcissi have been cultivated from the earliest times, but became increasingly popular in Europe after the 16th century and by the late 19th century were an important commercial crop centred primarily in the Netherlands. Today narcissi are popular as cut flowers and as ornamental plants in private and public gardens. The long history of breeding has resulted in thousands of different cultivars. For horticultural purposes, narcissi are classified into divisions, covering a wide range of shapes and colours. Like other members of their family, narcissi produce a number of different alkaloids, which provide some protection for the plant, but may be poisonous if accidentally ingested. This property has been exploited for medicinal use in traditional healing and has resulted in the production of galantamine for the treatment of Alzheimer's dementia. Long celebrated in art and literature, narcissi are associated with a number of themes in different cultures, ranging from death to good fortune, and as symbols of spring.

 

The daffodil is the national flower of Wales and the symbol of cancer charities in many countries. The appearance of the wild flowers in spring is associated with festivals in many places.

 

Narcissus is a genus of perennial herbaceous bulbiferous geophytes, which die back after flowering to an underground storage bulb. They regrow in the following year from brown-skinned ovoid bulbs with pronounced necks, and reach heights of 5–80 centimetres (2.0–31.5 in) depending on the species. Dwarf species such as N. asturiensis have a maximum height of 5–8 centimetres (2.0–3.1 in), while Narcissus tazetta may grow as tall as 80 centimetres (31 in).

 

The plants are scapose, having a single central leafless hollow flower stem (scape). Several green or blue-green, narrow, strap-shaped leaves arise from the bulb. The plant stem usually bears a solitary flower, but occasionally a cluster of flowers (umbel). The flowers, which are usually conspicuous and white or yellow, sometimes both or rarely green, consist of a perianth of three parts. Closest to the stem (proximal) is a floral tube above the ovary, then an outer ring composed of six tepals (undifferentiated sepals and petals), and a central disc to conical shaped corona. The flowers may hang down (pendant), or be erect. There are six pollen bearing stamens surrounding a central style. The ovary is inferior (below the floral parts) consisting of three chambers (trilocular). The fruit consists of a dry capsule that splits (dehisces) releasing numerous black seeds.

 

The bulb lies dormant after the leaves and flower stem die back and has contractile roots that pull it down further into the soil. The flower stem and leaves form in the bulb, to emerge the following season. Most species are dormant from summer to late winter, flowering in the spring, though a few species are autumn flowering.

 

The pale brown-skinned ovoid tunicate bulbs have a membranous tunic and a corky stem (base or basal) plate from which arise the adventitious root hairs in a ring around the edge, which grow up to 40 mm in length. Above the stem plate is the storage organ consisting of bulb scales, surrounding the previous flower stalk and the terminal bud. The scales are of two types, true storage organs and the bases of the foliage leaves. These have a thicker tip and a scar from where the leaf lamina became detached. The innermost leaf scale is semicircular only partly enveloping the flower stalk (semisheathed).(see Hanks Fig 1.3). The bulb may contain a number of branched bulb units, each with two to three true scales and two to three leaf bases. Each bulb unit has a life of about four years.

 

Once the leaves die back in summer, the roots also wither. After some years, the roots shorten pulling the bulbs deeper into the ground (contractile roots). The bulbs develop from the inside, pushing the older layers outwards which become brown and dry, forming an outer shell, the tunic or skin. Up to 60 layers have been counted in some wild species. While the plant appears dormant above the ground the flower stalk which will start to grow in the following spring, develops within the bulb surrounded by two to three deciduous leaves and their sheaths. The flower stem lies in the axil of the second true leaf.

 

The single leafless Plant stem stem or scape, appearing from early to late spring depending on the species, bears from 1 to 20 blooms. Stem shape depends on the species, some are highly compressed with a visible seam, while others are rounded. The stems are upright and located at the centre of the leaves. In a few species such as N. hedraeanthus the stem is oblique. The stem is hollow in the upper portion but towards the bulb is more solid and filled with a spongy material.

 

Narcissus plants have one to several basal leaf leaves which are linear, ligulate or strap-shaped (long and narrow), sometimes channelled adaxially to semiterete, and may (pedicellate) or may not (sessile) have a petiole stalk. The leaves are flat and broad to cylindrical at the base and arise from the bulb. The emerging plant generally has two leaves, but the mature plant usually three, rarely four, and they are covered with a cutin containing cuticle, giving them a waxy appearance. Leaf colour is light green to blue-green. In the mature plant, the leaves extend higher than the flower stem, but in some species, the leaves are low-hanging. The leaf base is encased in a colorless sheath. After flowering, the leaves turn yellow and die back once the seed pod (fruit) is ripe.

 

Jonquils usually have dark green, round, rush-like leaves.

 

The inflorescence is scapose, the single stem or scape bearing either a solitary flower or forming an umbel with up to 20 blooms. Species bearing a solitary flower include section Bulbocodium and most of section Pseudonarcissus. Umbellate species have a fleshy racemose inflorescence (unbranched, with short floral stalks) with 2 to 15 or 20 flowers, such as N. papyraceus (see illustration, left) and N. tazetta (see Table I). The flower arrangement on the inflorescence may be either with (pedicellate) or without (sessile) floral stalks.

 

Prior to opening, the flower buds are enveloped and protected in a thin dry papery or membranous (scarious) spathe. The spathe consists of a singular bract that is ribbed, and which remains wrapped around the base of the open flower. As the bud grows, the spathe splits longitudinally. Bracteoles are small or absent.

 

The flowers of Narcissus are hermaphroditic (bisexual), have three parts (tripartite), and are sometimes fragrant (see Fragrances). The flower symmetry is actinomorphic (radial) to slightly zygomorphic (bilateral) due to declinate-ascending stamens (curving downwards, then bent up at the tip). Narcissus flowers are characterised by their, usually conspicuous, corona (trumpet).

 

The three major floral parts (in all species except N. cavanillesii in which the corona is virtually absent - Table I: Section Tapeinanthus) are;

 

(i) the proximal floral tube (hypanthium),

(ii) the surrounding free tepals, and

(iii) the more distal corona (paraperigon, paraperigonium).

All three parts may be considered to be components of the perianth (perigon, perigonium). The perianth arises above the apex of the inferior ovary, its base forming the hypanthial floral tube.

 

The floral tube is formed by fusion of the basal segments of the tepals (proximally connate). Its shape is from an inverted cone (obconic) to funnel-shaped (funneliform) or cylindrical, and is surmounted by the more distal corona. Floral tubes can range from long and narrow sections Apodanthi and Jonquilla to rudimentary (N. cavanillesii).

 

Surrounding the floral tube and corona and reflexed (bent back) from the rest of the perianth are the six spreading tepals or floral leaves, in two whorls which may be distally ascending, reflexed (folded back), or lanceolate. Like many monocotyledons, the perianth is homochlamydeous, which is undifferentiated into separate calyx (sepals) and corolla (petals), but rather has six tepals. The three outer tepal segments may be considered sepals, and the three inner segments petals. The transition point between the floral tube and the corona is marked by the insertion of the free tepals on the fused perianth.

 

The corona, or paracorolla, is variously described as bell-shaped (funneliform, trumpet), bowl-shaped (cupular, crateriform, cup-shaped) or disc-shaped with margins that are often frilled, and is free from the stamens. Rarely is the corona a simple callose (hardened, thickened) ring. The corona is formed during floral development as a tubular outgrowth from stamens which fuse into a tubular structure, the anthers becoming reduced. At its base, the fragrances which attract pollinators are formed. All species produce nectar at the top of the ovary. Coronal morphology varies from the tiny pigmented disk of N. serotinus (see Table I) or the rudimentary structure in N. cavanillesii to the elongated trumpets of section Pseudonarcissus (trumpet daffodils, Table I).

 

While the perianth may point forwards, in some species such as N. cyclamineus it is folded back (reflexed, see illustration, left), while in some other species such as N. bulbocodium (Table I), it is reduced to a few barely visible pointed segments with a prominent corona.

 

The colour of the perianth is white, yellow or bicoloured, with the exception of the night flowering N. viridiflorus which is green. In addition the corona of N. poeticus has a red crenulate margin (see Table I). Flower diameter varies from 12 mm (N. bulbocodium) to over 125 mm (N. nobilis=N. pseudonarcissus subsp. nobilis).

 

Flower orientation varies from pendent or deflexed (hanging down) as in N. triandrus (see illustration, left), through declinate-ascendant as in N. alpestris = N. pseudonarcissus subsp. moschatus, horizontal (patent, spreading) such as N. gaditanus or N. poeticus, erect as in N. cavanillesii, N. serotinus and N. rupicola (Table I), or intermediate between these positions (erecto-patent).

 

The flowers of Narcissus demonstrate exceptional floral diversity and sexual polymorphism, primarily by corona size and floral tube length, associated with pollinator groups (see for instance Figs. 1 and 2 in Graham and Barrett). Barrett and Harder (2005) describe three separate floral patterns;

 

"Daffodil" form

"Paperwhite" form

"Triandrus" form.

The predominant patterns are the 'daffodil' and 'paperwhite' forms, while the "triandrus" form is less common. Each corresponds to a different group of pollinators (See Pollination).

 

The "daffodil" form, which includes sections Pseudonarcissus and Bulbocodium, has a relatively short, broad or highly funnelform tube (funnel-like), which grades into an elongated corona, which is large and funnelform, forming a broad, cylindrical or trumpet-shaped perianth. Section Pseudonarcissus consists of relatively large flowers with a corolla length of around 50 mm, generally solitary but rarely in inflorescences of 2–4 flowers. They have wide greenish floral tubes with funnel-shaped bright yellow coronas. The six tepals sometimes differ in colour from the corona and may be cream coloured to pale yellow.

 

The "paperwhite" form, including sections Jonquilla, Apodanthi and Narcissus, has a relatively long, narrow tube and a short, shallow, flaring corona. The flower is horizontal and fragrant.

 

The "triandrus" form is seen in only two species, N. albimarginatus (a Moroccan endemic) and N. triandrus. It combines features of both the "daffodil" and "paperwhite" forms, with a well-developed, long, narrow tube and an extended bell-shaped corona of almost equal length. The flowers are pendent.

 

Androecium

There are six stamens in one to two rows (whorls), with the filaments separate from the corona, attached at the throat or base of the tube (epipetalous), often of two separate lengths, straight or declinate-ascending (curving downwards, then upwards). The anthers are basifixed (attached at their base).

 

Gynoecium

The ovary is inferior (below the floral parts) and trilocular (three chambered) and there is a pistil with a minutely three lobed stigma and filiform (thread like) style, which is often exserted (extending beyond the tube).

 

Fruit

The fruit consists of dehiscent loculicidal capsules (splitting between the locules) that are ellipsoid to subglobose (almost spherical) in shape and are papery to leathery in texture.

 

Seeds

The fruit contains numerous subglobose seeds which are round and swollen with a hard coat, sometimes with an attached elaiosome. The testa is black and the pericarp dry.

 

Most species have 12 ovules and 36 seeds, although some species such as N. bulbocodium have more, up to a maximum of 60. Seeds take five to six weeks to mature. The seeds of sections Jonquilla and Bulbocodium are wedge-shaped and matte black, while those of other sections are ovate and glossy black. A gust of wind or contact with a passing animal is sufficient to disperse the mature seeds.

 

Chromosomes

Chromosome numbers include 2n=14, 22, 26, with numerous aneuploid and polyploid derivatives. The basic chromosome number is 7, with the exception of N. tazetta, N. elegans and N. broussonetii in which it is 10 or 11; this subgenus (Hermione) was in fact characterised by this characteristic. Polyploid species include N. papyraceus (4x=22) and N. dubius (6x=50).

 

Phytochemistry

Alkaloids

As with all Amarylidaceae genera, Narcissus contains unique isoquinoline alkaloids. The first alkaloid to be identified was lycorine, from N. pseudonarcissus in 1877. These are considered a protective adaptation and are utilised in the classification of species. Nearly 100 alkaloids have been identified in the genus, about a third of all known Amaryllidaceae alkaloids, although not all species have been tested. Of the nine alkaloid ring types identified in the family, Narcissus species most commonly demonstrate the presence of alkaloids from within the Lycorine (lycorine, galanthine, pluviine) and Homolycorine (homolycorine, lycorenine) groups. Hemanthamine, tazettine, narciclasine, montanine and galantamine alkaloids are also represented. The alkaloid profile of any plant varies with time, location, and developmental stage. Narcissus also contain fructans and low molecular weight glucomannan in the leaves and plant stems.

 

Fragrances

Fragrances are predominantly monoterpene isoprenoids, with a small amount of benzenoids, although N. jonquilla has both equally represented. Another exception is N. cuatrecasasii which produces mainly fatty acid derivatives. The basic monoterpene precursor is geranyl pyrophosphate, and the commonest monoterpenes are limonene, myrcene, and trans-β-ocimene. Most benzenoids are non-methoxylated, while a few species contain methoxylated forms (ethers), e.g. N. bujei. Other ingredient include indole, isopentenoids and very small amounts of sesquiterpenes. Fragrance patterns can be correlated with pollinators, and fall into three main groups (see Pollination).

 

The taxonomy of Narcissus is complex, and still not fully resolved. Known to the ancients, the genus name appears in Graeco-Roman literature, although their interest was as much medicinal as botanical. It is unclear which species the ancients were familiar with. Although frequently mentioned in Mediaeval and Renaissance texts it was not formally described till the work of Linnaeus in 1753. By 1789 it had been grouped into a family (Narcissi) but shortly thereafter this was renamed Amaryllideae, from which comes the modern placement within Amaryllidaceae, although for a while it was considered part of Liliaceae.

 

Many of the species now considered to be Narcissus were in separate genera during the nineteenth century, and the situation was further confused by the inclusion of many cultivated varieties. By 1875 the current circumscription was relatively settled. By 2004 phylogenetic studies had allowed the place of Narcissus within its fairly large family to be established, nested within a series of subfamilies (Amaryllidoideae) and tribes (Narcisseae). It shares its position in the latter tribe with Sternbergia.

 

The infrageneric classification has been even more complex and many schemes of subgenera, sections, subsections and series have been proposed, although all had certain similarities. Most authorities now consider there to be 10 – 11 sections based on phylogenetic evidence. The problems have largely arisen from the diversity of the wild species, frequent natural hybridisation and extensive cultivation with escape and subsequent naturalisation. The number of species has varied anywhere from 16 to nearly 160, but is probably around 50 – 60.

 

The genus appeared some time in the Late Oligocene to Early Miocene eras, around 24 million years ago, in the Iberian peninsula. While the exact origin of the word Narcissus is unknown it is frequently linked to its fragrance which was thought to be narcotic, and to the legend of the youth of that name who fell in love with his reflection. In the English language the common name Daffodil appears to be derived from the Asphodel with which it was commonly compared.

 

Early

Narcissus was first described by Theophrastus (Θεόφραστος, c 371 - c 287 BC) in his Historia Plantarum (Greek: Περὶ φυτῶν ἱστορία) as νάρκισσος, referring to N. poeticus, but comparing it to Asphodelus (ασφοδελωδες). Theophrastus' description was frequently referred to at length by later authors writing in Latin such as Pliny the Elder (Gaius Plinius Secundus, 23 AD – 79 AD) from whom came the Latin form narcissus (see also Culture). Pliny's account is from his Natural History (Latin: Naturalis Historia). Like his contemporaries, his interests were as much therapeutic as botanical. Another much-cited Greek authority was Dioscorides (Διοσκουρίδης, 40 AD – 90 AD) in his De Materia Medica (Greek: Περὶ ὕλης ἰατρικῆς). Both authors were to remain influential until at least the Renaissance, given that their descriptions went beyond the merely botanical, to the therapeutic (see also Antiquity).

 

An early European reference is found in the work of Albert Magnus (c. 1200 – 1280), who noted in his De vegetabilibus et plantis the similarity to the leek. William Turner in his A New Herball (1551) cites all three extensively in his description of the plant and its properties.It was to remain to Linnaeus in 1753 to formally describe and name Narcissus as a genus in his Species Plantarum, at which time there were six known species (N. poeticus, N. pseudonarcissus, N. bulbocodium, N. serotinus, N. jonquilla and N. tazetta).[1] At that time, Linnaeus loosely grouped it together with 50 other genera into his Hexandria monogynia.

 

Modern

It was de Jussieu in 1789 who first formally created a 'family' (Narcissi), as the seventh 'Ordo' (Order) of the third class (Stamina epigyna) of Monocots in which Narcissus and 15 other genera were placed. The use of the term Ordo at that time was closer to what we now understand as Family, rather than Order. The family has undergone much reorganisation since then, but in 1805 it was renamed after a different genus in the family, Amaryllis, as 'Amaryllideae' by Jaume St.-Hilaire and has retained that association since. Jaume St.-Hilaire divided the family into two unnamed sections and recognised five species of Narcissus, omitting N. serotinus.

 

De Candolle brought together Linnaeus' genera and Jussieau's families into a systematic taxonomy for the first time, but included Narcissus (together with Amaryllis) in the Liliaceae in his Flore française (1805-1815) rather than Amaryllidaceae, a family he had not yet recognised. Shortly thereafter he separated the 'Amaryllidées' from 'Liliacées' (1813), though attributing the term to Brown's 'Amaryllideae' in the latter's Prodromus (1810) rather than St.-Hilaire's 'Amaryllidées'. He also provided the text to the first four volumes of Redouté illustrations in the latter's Les liliacées between 1805 and 1808 (see illustration here of N. candidissimus).

 

Historically both wide and narrow interpretations of the genus have been proposed. In the nineteenth century genus splitting was common, favouring the narrow view. Haworth (1831) using a narrow view treated many species as separate genera, as did Salisbury (1866). These authors listed various species in related genera such as Queltia (hybrids), Ajax (=Pseudonarcissus) and Hermione (=Tazettae), sixteen in all in Haworth's classification. In contrast, Herbert (1837) took a very wide view reducing Harworth's sixteen genera to six. Herbert, treating the Amaryllidacea as an 'order' as was common then, considered the narcissi to be a suborder, the Narcisseae, the six genera being Corbularia, Ajax, Ganymedes, Queltia, Narcissus and Hermione and his relatively narrow circumscription of Narcissus having only three species. Later Spach (1846) took an even wider view bringing most of Harworth's genera into the genus Narcissus, but as separate subgenera. By the time that Baker (1875) wrote his monograph all of the genera with one exception were included as Narcissus. The exception was the monotypic group Tapeinanthus which various subsequent authors have chosen to either exclude (e.g. Cullen 1986) or include (e.g. Webb 1978, 1980). Today it is nearly always included.

 

The eventual position of Narcissus within the Amaryllidaceae family only became settled in the twenty-first century with the advent of phylogenetic analysis and the Angiosperm Phylogeny Group system. The genus Narcissus belongs to the Narcisseae tribe, one of 13 within the Amaryllidoideae subfamily of the Amaryllidaceae. It is one of two sister clades corresponding to genera in the Narcisseae, being distinguished from Sternbergia by the presence of a paraperigonium, and is monophyletic

 

The infrageneric phylogeny of Narcissus still remains relatively unsettled. The taxonomy has proved very complex and difficult to resolve, particularly for the Pseudonarcissus group. This is due to a number of factors, including the diversity of the wild species, the ease with which natural hybridisation occurs, and extensive cultivation and breeding accompanied by escape and naturalisation.

 

De Candolle, in the first systematic taxonomy of Narcissus, arranged the species into named groups, and those names (Faux-Narcisse or Pseudonarcissus, Poétiques, Tazettes, Bulbocodiens, Jonquilles) have largely endured for the various subdivisions since and bear his name. The evolution of classification was confused by including many unknown or garden varieties, until Baker (1875) made the important distinction of excluding all specimens except the wild species from his system. He then grouped all of the earlier related genera as sections under one genus, Narcissus, the exception being the monotypic Tapeinanthus. Consequently, the number of accepted species has varied widely.

 

A common modern classification system has been that of Fernandes (1951, 1968, 1975) based on cytology, as modified by Blanchard (1990) and Mathew (2002), although in some countries such as Germany, the system of Meyer (1966) was preferred. Fernandes described two subgenera based on basal chromosome number, Hermione, n = 5 (11) and Narcissus, n = 7 (13). He further subdivided these into ten sections (Apodanthi, Aurelia, Bulbocodii, Ganymedes, Jonquillae, Narcissus, Pseudonarcissi, Serotini, Tapeinanthus, Tazettae), as did Blanchard later.

 

In contrast to Fernandes, Webb's treatment of the genus for the Flora Europaea (1978, 1980) prioritised morphology over genetics, and abandoned the subgenera ranks. He also restored De Candolle's original nomenclature, and made a number of changes to section Jonquilla, merging the existing subsections, reducing Apodanthi to a subsection of Jonquilla, and moving N. viridiflorus from Jonquilla to a new monotypic section of its own (Chloranthi). Finally, he divided Pseudonarcissus into two subsections. Blanchard (1990), whose Narcissus: a guide to wild daffodils has been very influential, adopted a simple approach, restoring Apodanthae, and based largely on ten sections alone.

 

The Royal Horticultural Society (RHS) currently lists ten sections, based on Fernandes (1968), three of which are monotypic (contain only one species), while two others only containing two species. Most species are placed in Pseudonarcissus While infrageneric groupings within Narcissus have been relatively constant, their status (genera, subgenera, sections, subsections, series, species) has not. Some authors treat some sections as being further subdivided into subsections, e.g. Tazettae (3 subsections). These subdivisions correspond roughly to the popular names for narcissi types, e.g. Trumpet Daffodils, Tazettas, Pheasant's Eyes, Hoop Petticoats, Jonquils.

 

While Webb had simply divided the genus into sections, Mathew found this unsatisfactory, implying every section had equal status. He adapted both Fernandes and Webb to devise a more hierarchical scheme he believed better reflected the interrelationships within the genus. Mathew's scheme consists of three subgenera (Narcissus, Hermione and Corbularia). The first two subgenera were then divided into five and two sections respectively. He then further subdivided two of the sections (subgenus Narcissus section Jonquillae, and subgenus Hermione section Hermione) into three subsections each. Finally, he divided section Hermione subsection Hermione further into two series, Hermione and Albiflorae. While lacking a phylogenetic basis, the system is still in use in horticulture. For instance the Pacific Bulb Society uses his numbering system for classifying species.

 

The phylogenetic analysis of Graham and Barrett (2004) supported the infrageneric division of Narcissus into two clades corresponding to the subgenera Hermione and Narcissus, but does not support monophyly of all sections, with only Apodanthi demonstrating clear monophyly, corresponding to Clade III of Graham and Barrett, although some other clades corresponded approximately to known sections. These authors examined 36 taxa of the 65 listed then, and a later extended analysis by Rønsted et al. (2008) with five additional taxa confirmed this pattern.

 

A very large (375 accessions) molecular analysis by Zonneveld (2008) utilising nuclear DNA content sought to reduce some of the paraphyly identified by Graham and Barrett. This led to a revision of the sectional structure, shifting some species between sections, eliminating one section and creating two new ones. In subgenus Hermione, Aurelia was merged with Tazettae. In subgenus Narcissus section Jonquillae subsection Juncifolii was elevated to sectional rank, thus resolving the paraphyly in this section observed by Graham and Barrett in Clade II due to this anomalous subsection, the remaining species being in subsection Jonquillae, which was monophyletic. The relatively large section Pseudonarcissi was divided by splitting off a new section, Nevadensis (species from southern Spain) leaving species from France, northern Spain and Portugal in the parent section. At the same time Fernández-Casas (2008) proposed a new monotypic section Angustini to accommodate Narcissus deficiens, placing it within subgenus Hermione.

 

While Graham and Barrett (2004) had determined that subgenus Hermione was monophyletic, using a much larger accession Santos-Gally et al. (2011) did not. However the former had excluded species of hybrid origins, while the latter included both N. dubius and N. tortifolius. If these two species are excluded (forming a clade with subgenus Narcissus) then Hermione can be considered monophyletic, although as a section of Hermione, Tazettae is not monophyletic. They also confirmed the monophyly of Apodanthi.

 

Some so-called nothosections have been proposed, predominantly by Fernández-Casas, to accommodate natural ('ancient') hybrids (nothospecies).

 

Subgenera and sections

Showing revisions by Zonnefeld (2008)

 

subgenus Hermione (Haw.) Spach.

(Aurelia (Gay) Baker (monotypic) - merged with Tazettae (2008)

Serotini Parlatore (2 species)

Tazettae de Candolle (16 species) syn. Hermione (Salisbury) Sprengel, in Fernandes' scheme. Incorporating Aurelia (2008)

subgenus Narcissus L.

Apodanthi A. Fernandes (6 species)

Bulbocodium de Candolle (11 species)

Ganymedes (Haworth) Schultes f. (monotypic)

Jonquillae de Candolle (8 species)

Juncifolii (A. Fern.) Zonn. sect. nov. (2008)

Narcissus L. (2 species)

Nevadensis Zonn. sect. nov. (2008)

Pseudonarcissus de Candolle (36 species) Trumpet daffodils

Tapeinanthus (Herbert) Traub (monotypic)

 

Species

Estimates of the number of species in Narcissus have varied widely, from anywhere between 16 and nearly 160, even in the modern era. Linnaeus originally included six species in 1753. By the time of the 14th edition of the Systema Naturae in 1784, there were fourteen. The 1819 Encyclopaedia Londinensis lists sixteen (see illustration here of three species) and by 1831 Adrian Haworth had described 150 species.

 

Much of the variation lies in the definition of species, and whether closely related taxa are considered separate species or subspecies. Thus, a very wide view of each species, such as Webb's results in few species, while a very narrow view such as that of Fernandes results in a larger number. Another factor is the status of hybrids, given natural hybridisation, with a distinction between 'ancient hybrids' and 'recent hybrids'. The term 'ancient hybrid' refers to hybrids found growing over a large area, and therefore now considered as separate species, while 'recent hybrid' refers to solitary plants found amongst their parents, with a more restricted range.

 

In the twentieth century Fernandes (1951) accepted 22 species, on which were based the 27 species listed by Webb in the 1980 Flora Europaea. By 1968, Fernandes had accepted 63 species, and by 1990 Blanchard listed 65 species, and Erhardt 66 in 1993. In 2006 the Royal Horticultural Society's (RHS) International Daffodil Register and Classified List listed 87 species, while Zonneveld's genetic study (2008) resulted in only 36. As of September 2014, the World Checklist of Selected Plant Families accepts 52 species, along with at least 60 hybrids, while the RHS has 81 accepted names in its October 2014 list.

 

Evolution

Within the Narcisseae, Narcissus (western Mediterranean) diverged from Sternbergia (Eurasia) some time in the Late Oligocene to Early Miocene eras, around 29.3–18.1 Ma, with a best estimate of 23.6 Ma. Later the genus divided into the two subgenera (Hermione and Narcissus) between 27.4 and 16.1 Ma (21.4 Ma). The divisions between the sections of Hermione then took place during the Miocene period 19.9–7.8 Ma.

 

Narcissus appears to have arisen in the area of the Iberian peninsula, southern France and northwestern Italy, and within this area most sections of the genus appeared, with only a few taxa being dispersed to North Africa at a time when the African and West European platforms were closer together. Subgenus Hermione in turn arose in the southwestern mediterranean and north west Africa. However, these are reconstructions, the Amaryllidaceae lacking a fossil record.

 

Names and etymology

The derivation of the Latin narcissus (Greek: νάρκισσος) is unknown. It may be a loanword from another language; for instance, it is said to be related to the Sanskrit word nark, meaning 'hell'. It is frequently linked to the Greek myth of Narcissus described by Ovid in his Metamorphoses, who became so obsessed with his own reflection that as he knelt and gazed into a pool of water, he fell into the water and drowned. In some variations, he died of starvation and thirst. In both versions, the narcissus plant sprang from where he died. Although Ovid appeared to describe the plant we now know as Narcissus there is no evidence for this popular derivation, and the person's name may have come from the flower's name. The Poet's Narcissus (N. poeticus), which grows in Greece, has a fragrance that has been described as intoxicating. This explanation is largely discredited due to lack of proof. Pliny wrote that the plant 'narce narcissum dictum, non a fabuloso puero' ('named narcissus from narce, not from the legendary youth'), i.e. that it was named for its narcotic properties (ναρκάω narkao, 'I grow numb' in Greek), not from the legend. Furthermore, there were accounts of narcissi growing, such as in the legend of Persephone, long before the story of Narcissus appeared (see Greek culture). It has also been suggested that daffodils bending over streams evoked the image of the youth admiring his own reflection in the water.

 

Linnaeus used the Latin name for the plant in formally describing the genus, although Matthias de l'Obel had previously used the name in describing various species of Narcissi in his Icones stirpium of 1591, and other publications, as had Clusius in Rariorum stirpium (1576).

 

The plural form of the common name narcissus has caused some confusion. British English sources such as the Shorter Oxford English Dictionary give two alternate forms, narcissi and narcissuses. In contrast, in American English the Merriam-Webster Dictionary provides for a third form, narcissus, used for both singular and plural. The Oxford dictionaries only list this third form under American English, although the Cambridge Dictionary allows of all three in the same order. However, Garner's Modern American Usage states that narcissi is the commonest form, narcissuses being excessively sibilant. For similar reasons, Fowler prefers narcissi in British English usage. Neither support narcissus as a plural form. Common names such as narcissus do not capitalise the first letter in contrast to the person of that name and the Latin genus name.

 

The name Narcissus (feminine Narcissa) was not uncommon in Roman times, such as Tiberius Claudius Narcissus, a Roman official in Claudius' time, an early New Testament Christian in Rome and later bishops and saints.

 

Daffodil

The word daffodil was unknown in the English language before the sixteenth century. The name is derived from an earlier affodell, a variant of asphodel. In classical Greek literature the narcissus is frequently referred to as the asphodel, such as the meadows of the Elysian fields in Homer (see Antiquity). Asphodel in turn appears to be a loanword coming from French via Mediaeval Latin affodilus from Classical Latin asphodilus and ultimately the Greek asphodelos (Greek: ἀσφόδελος). The reason for the introduction of the initial d is not known, although a probable source is an etymological merging from the Dutch article de, as in de affodil, or English the, as th'affodil or t'affodil, hence daffodil, and in French de and affodil to form fleur d'aphrodille and daphrodille.

 

From at least the 16th century, daffadown dilly and daffydowndilly have appeared as playful synonyms of the name. In common parlance and in historical documents, the term daffodil may refer specifically to populations or specimens of the wild daffodil, N. pseudonarcissus. H. N. Ellacombe suggests this may be from Saffon Lilly, citing Prior in support, though admittedly conjectural.

 

Lady Wilkinson (1858), who provides an extensive discussion of the etymology of the various names for this plant, suggests a very different origin, namely the Old English word affodyle (that which cometh early), citing a 14th-century (but likely originally much earlier) manuscript in support of this theory, and which appears to describe a plant resembling the daffodil. Ellacombe provides further support for this from a fifteenth century English translation of Palladius that also refers to it.

 

Jonquil

The name jonquil is said to be a corruption via French from the Latin juncifolius meaning 'rush-leaf' (Juncaceae) and its use is generally restricted to those species and cultivars which have rush like leaves, e.g. N. juncifolius.

 

Other

A profusion of names have attached themselves in the English language, either to the genus as a whole or to individual species or groups of species such as sections. These include narcissus, jonquil, Lent lily, Lenten lily, lide lily, yellow lily, wort or wyrt, Julians, glens, Lent cocks, corn flower, bell rose, asphodel, Solomon's lily, gracy day, haverdrils, giggary, cowslip, and crow foot.

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Centro Comercial El Pueblito Suite 208

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Cartagena, Bolivar 130001 Colombia

 

email: leosmusclebeach@live.com

Phone: 011-575-665-2822

Cell: 011-57-301-353-5884

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Centro Comercial El Pueblito Suite 208

Avenida San Martin, Bocagrande

Cartagena, Bolivar 130001 Colombia

 

email: leosmusclebeach@live.com

Phone: 011-575-665-2822

Cell: 011-57-301-353-5884

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Centro Comercial El Pueblito Suite 208

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Centro Comercial El Pueblito Suite 208

Avenida San Martin, Bocagrande

Cartagena, Bolivar 130001 Colombia

 

email: leosmusclebeach@live.com

Phone: 011-575-665-2822

Cell: 011-57-301-353-5884

A thick rich drink made from a type of orchid root, flavoured with a dusting of cinnamon. Warm and comforting like a hot chocolate.

Once it's cooled a little, it becomes even thicker and even a little slimy :)

 

Sahlab سحلب - wikipedia

flour made from grinding the dried tubers of Orchis mascula, Orchis militaris and related species of orchids, which contain a nutritious starch-like polysaccharide called glucomannan

 

---

 

Craving a good Lebanese lahmacun, also known as a Lebanese pizza or Manakish, we head to A1 Bakery, home of good Lebanese bread.

 

A-1 Bakery

(03) 9386 0440

643-645 Sydney Rd

Brunswick VIC 3056

www.a1bakery.com.au/

 

Reviews:

- Breakfast for the brave - Epicure, The Age by Nina Rousseau April 8, 2008

Lebanese Pizza, $1.20-$6

join the hordes at A1 for Lebanese pizzas from the wall of ovens up the back. Try one speckled with zaatar or a meaty version with sausage and haloumi.

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Centro Comercial El Pueblito Suite 208

Avenida San Martin, Bocagrande

Cartagena, Bolivar 130001 Colombia

 

email: leosmusclebeach@live.com

Phone: 011-575-665-2822

Cell: 011-57-301-353-5884

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Centro Comercial El Pueblito Suite 208

Avenida San Martin, Bocagrande

Cartagena, Bolivar 130001 Colombia

 

email: leosmusclebeach@live.com

Phone: 011-575-665-2822

Cell: 011-57-301-353-5884

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Centro Comercial El Pueblito Suite 208

Avenida San Martin, Bocagrande

Cartagena, Bolivar 130001 Colombia

 

email: leosmusclebeach@live.com

Phone: 011-575-665-2822

Cell: 011-57-301-353-5884

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Centro Comercial El Pueblito Suite 208

Avenida San Martin, Bocagrande

Cartagena, Bolivar 130001 Colombia

 

email: leosmusclebeach@live.com

Phone: 011-575-665-2822

Cell: 011-57-301-353-5884

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Centro Comercial El Pueblito Suite 208

Avenida San Martin, Bocagrande

Cartagena, Bolivar 130001 Colombia

 

email: leosmusclebeach@live.com

Phone: 011-575-665-2822

Cell: 011-57-301-353-5884

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Centro Comercial El Pueblito Suite 208

Avenida San Martin, Bocagrande

Cartagena, Bolivar 130001 Colombia

 

email: leosmusclebeach@live.com

Phone: 011-575-665-2822

Cell: 011-57-301-353-5884

This has edible tubers which are used in Asian cooking.

 

Amorphophallus (from Ancient Greek amorphos, "without form, misshapen" + phallos, "penis", referring to the shape of the prominent spadix) is a large genus of some 170 tropical and subtropical tuberous herbaceous plants from the Arum family (Araceae). A few species are edible as "famine foods" after careful preparation to remove irritating chemicals.

 

These are typical lowland plants, growing in the tropical and subtropical zones of the paleotropics, from West Africa to the Pacific Islands. None of them are found in the Americas although a remarkably similar but not closely related genus, Dracontium, has evolved there. Most species are endemic. They grow preferentially on disturbed grounds, such as secondary forests.

 

Konjac (Amorphophallus konjac; syn. A. rivieri), also known as konjak, konjaku, devil's tongue, voodoo lily, snake palm, or elephant yam (though this name is also used for A. paeoniifolius), is a plant of the genus Amorphophallus. It is native to warm subtropical to tropical eastern Asia, from Japan and China south to Indonesia.

 

It is a perennial plant, growing from a large corm up to 25 cm in diameter. The single leaf is up to 1.3 m across, bipinnate, and divided into numerous leaflets. The flowers are produced on a spathe enclosed by a dark purple spadix up to 55 cm long.

 

The corm of the konjac is often colloquially referred to as a yam, although it bears no marked relation to tubers of the family Dioscoreaceae.

 

Konjac is grown in India, China, Japan and Korea for its large starchy corms, used to create a flour and jelly of the same name. It is also used as a vegan substitute for gelatin.

 

In Japanese cuisine, konjac (known as konnyaku) appears in dishes such as oden. It is typically mottled grey and firmer in consistency than most gelatins. It has very little taste; the common variety tastes vaguely like salt. It is valued more for its texture than flavor.

 

Ito konnyaku is a type of Japanese food consisting of konjac cut into noodle-like strips. It is usually sold in plastic bags with accompanying water. It is often used in sukiyaki and oden. The name literally means "thread-konjac."

 

Japanese konnyaku is made by mixing konjac flour with water and limewater. Hijiki is often added for the characteristic dark color and flavor. Without additives for color, konnyaku is pale white. It is then boiled and cooled to solidify. Konnyaku made in noodle form is called shirataki (see shirataki noodles) and used in foods such as sukiyaki and gyudon.

 

Konjac is consumed in parts of China's Sichuan province; the corm is called moyu (or Devil's Taro), and the jelly is called "konjac tofu" (moyu doufu) or "snow konjac" (xue moyu).

 

The dried corm of the konjac plant contains around 40% glucomannan gum. This polysaccharide makes konjac jelly highly viscous.

 

Konjac has almost 0 calories but is very high in fiber. Thus, it is often used as a diet food.It can also be used for facial massage accessories which are currently popular in Korea.

 

Konjac can also be made into a popular Asian fruit jelly snack, known variously in the United States as lychee cups (after a typical flavor and Nata de coco cube suspended in the gel) or konjac candy, usually served in bite-sized plastic cups.

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Fruttate e deliziose note di frutti di bosco (da essenza conforme al disciplinare AIAB).

 

Applicare sul viso e lasciare agire tutta la notte per un trattamento di bellezza durante il sonno. La mattina risciacquare e procedere con l’abituale skincare routine.

Da abbinare agli altri prodotti della Linea viso antiossidante Biofficina Toscana per una sinergia ottimale.

 

Aqua [Water], Caprylic/capric triglyceride, Prunus amygdalus dulcis (Sweet almond) oil, Cetearyl alcohol, Cetearyl glucoside, Glycerin, Butyrospermum parkii (Shea) butter, Argania spinosa kernel oil, Vaccinium myrtillus (Myrtle) fruit extract*, Hydrolyzed grape skin*, Olea europaea (Olive) fruit oil, Olea europaea (Olive) oil unsaponifiables, Hydrogenated olive oil, Arachidyl glucoside, Arachidyl alcohol, Behenyl alcohol, Benzyl alcohol, Sodium levulinate, Glucomannan, Lecithin, Parfum [Fragrance], Betaine, Sodium anisate, Sorbitol, Hexyl cinnamal, Citric acid, Lactic acid, Xanthan gum, Hydroxycitronellal, Linalool, Sodium benzoate, Citronellol, Potassium sorbate, Coumarin, Limonene..

 

*da agricoltura biologica

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Centro Comercial El Pueblito Suite 208

Avenida San Martin, Bocagrande

Cartagena, Bolivar 130001 Colombia

 

email: leosmusclebeach@live.com

Phone: 011-575-665-2822

Cell: 011-57-301-353-5884

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I understand that it flowers once every 10 years and then only for about 3 days. There were many photographers at the Begonia House getting snap shots of it in flower. It's name is Amorphophallus konjac and is a large, perennial exotic Asian plant. It grows a single, elongated center called a corm and a single large leaf that wraps around it. The corm is the part of the plant that is used, as well as its tuber, which is commonly called the konnyaku potato. Glucomannan is its scientific name, and its other common names include konjac mannan, devil's tongue, konjac, elephant-foot yam, snake plant and konnyaku.

 

I story about this plant featured in the DominionPost this morning - Flower pungent smell of...

 

The metre-high flower is blooming for the first time in a decade at the garden's Begonia House and was attracting a lot of attention during the Spring Festival. The flower has been in bloom for two days and is only expected to last one more day.

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Metabolic-Boosting Raspberry Ketones

 

This is a compound found in red raspberries which basically slices up fat cells, making them easier to be burned as fuel. These ketones also create a feeling of fullness and keep cravings under control. Raspberry ketones are better taken as a supplement; else you would need to eat approximately 1,000 raspberries to reap the same benefits.

 

Metabolic-Boosting White Kidney Bean Extract

 

This ingredient is a known carb blocker. White kidney bean extract blocks unwanted calories from being stored as fat while keeping your blood sugar balanced. White Kidney Bean Extract paralyzes the enzyme that turns starch into sugar in your stomach, and so your body does not absorb it.

 

Metabolic-Boosting Green Tea Extract

 

Green tea has gone from an exotic beverage to one of the most popular weight loss ingredients. Green tea slows carbohydrate absorption and increases fat burning by improving the function of insulin.

 

Metabolic-Boosting Chromium

 

The trace mineral chromium supports healthy insulin function, which helps regulate blood sugar and is necessary for proper metabolism. Keeping blood sugar levels in check also supports weight loss.

 

Metabolic-Boosting Vitamin D

 

Excess body fat absorbs and holds onto vitamin D, making it unavailable to the body. The absence of Vitamin D interferes with the functioning of leptin, a hormone which signals the brain that you are full and should stop eating.

 

Nature’s richest source of soluble fiber

 

Glucomannan is a water-soluble dietary fiber that is derived from konjac root. Glucomannan, which can absorb up to 200 times its weight in water, works by absorbing water in your stomach and forming a gel that carries moisture through your digestive system. It also helps you feel full due to its swelling action and suppresses your appetite, keeping you from overeating and having unnecessary cravings, the greatest enemies to weight loss.

 

Research shows that Glucomannan helps delay gastric emptying, slows the absorption of dietary carbohydrates to reduce insulin spiking, improves cholesterol and triglyceride levels, and acts as a prebiotic.

 

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Narcissus is a genus of predominantly spring flowering perennial plants of the amaryllis family, Amaryllidaceae. Various common names including daffodil, narcissus, and jonquil are used to describe all or some members of the genus. Narcissus has conspicuous flowers with six petal-like tepals surmounted by a cup- or trumpet-shaped corona. The flowers are generally white and yellow (also orange or pink in garden varieties), with either uniform or contrasting coloured tepals and corona.

 

Narcissus were well known in ancient civilisation, both medicinally and botanically, but formally described by Linnaeus in his Species Plantarum (1753). The genus is generally considered to have about ten sections with approximately 50 species. The number of species has varied, depending on how they are classified, due to similarity between species and hybridisation. The genus arose some time in the Late Oligocene to Early Miocene epochs, in the Iberian peninsula and adjacent areas of southwest Europe. The exact origin of the name Narcissus is unknown, but it is often linked to a Greek word (ancient Greek ναρκῶ narkō, "to make numb") and the myth of the youth of that name who fell in love with his own reflection. The English word "daffodil" appears to be derived from "asphodel", with which it was commonly compared.

 

The species are native to meadows and woods in southern Europe and North Africa with a centre of diversity in the Western Mediterranean, particularly the Iberian peninsula. Both wild and cultivated plants have naturalised widely, and were introduced into the Far East prior to the tenth century. Narcissi tend to be long-lived bulbs, which propagate by division, but are also insect-pollinated. Known pests, diseases and disorders include viruses, fungi, the larvae of flies, mites and nematodes. Some Narcissus species have become extinct, while others are threatened by increasing urbanisation and tourism.

 

Historical accounts suggest narcissi have been cultivated from the earliest times, but became increasingly popular in Europe after the 16th century and by the late 19th century were an important commercial crop centred primarily in the Netherlands. Today narcissi are popular as cut flowers and as ornamental plants in private and public gardens. The long history of breeding has resulted in thousands of different cultivars. For horticultural purposes, narcissi are classified into divisions, covering a wide range of shapes and colours. Like other members of their family, narcissi produce a number of different alkaloids, which provide some protection for the plant, but may be poisonous if accidentally ingested. This property has been exploited for medicinal use in traditional healing and has resulted in the production of galantamine for the treatment of Alzheimer's dementia. Long celebrated in art and literature, narcissi are associated with a number of themes in different cultures, ranging from death to good fortune, and as symbols of spring.

 

The daffodil is the national flower of Wales and the symbol of cancer charities in many countries. The appearance of the wild flowers in spring is associated with festivals in many places.

 

Narcissus is a genus of perennial herbaceous bulbiferous geophytes, which die back after flowering to an underground storage bulb. They regrow in the following year from brown-skinned ovoid bulbs with pronounced necks, and reach heights of 5–80 centimetres (2.0–31.5 in) depending on the species. Dwarf species such as N. asturiensis have a maximum height of 5–8 centimetres (2.0–3.1 in), while Narcissus tazetta may grow as tall as 80 centimetres (31 in).

 

The plants are scapose, having a single central leafless hollow flower stem (scape). Several green or blue-green, narrow, strap-shaped leaves arise from the bulb. The plant stem usually bears a solitary flower, but occasionally a cluster of flowers (umbel). The flowers, which are usually conspicuous and white or yellow, sometimes both or rarely green, consist of a perianth of three parts. Closest to the stem (proximal) is a floral tube above the ovary, then an outer ring composed of six tepals (undifferentiated sepals and petals), and a central disc to conical shaped corona. The flowers may hang down (pendant), or be erect. There are six pollen bearing stamens surrounding a central style. The ovary is inferior (below the floral parts) consisting of three chambers (trilocular). The fruit consists of a dry capsule that splits (dehisces) releasing numerous black seeds.

 

The bulb lies dormant after the leaves and flower stem die back and has contractile roots that pull it down further into the soil. The flower stem and leaves form in the bulb, to emerge the following season. Most species are dormant from summer to late winter, flowering in the spring, though a few species are autumn flowering.

 

The pale brown-skinned ovoid tunicate bulbs have a membranous tunic and a corky stem (base or basal) plate from which arise the adventitious root hairs in a ring around the edge, which grow up to 40 mm in length. Above the stem plate is the storage organ consisting of bulb scales, surrounding the previous flower stalk and the terminal bud. The scales are of two types, true storage organs and the bases of the foliage leaves. These have a thicker tip and a scar from where the leaf lamina became detached. The innermost leaf scale is semicircular only partly enveloping the flower stalk (semisheathed).(see Hanks Fig 1.3). The bulb may contain a number of branched bulb units, each with two to three true scales and two to three leaf bases. Each bulb unit has a life of about four years.

 

Once the leaves die back in summer, the roots also wither. After some years, the roots shorten pulling the bulbs deeper into the ground (contractile roots). The bulbs develop from the inside, pushing the older layers outwards which become brown and dry, forming an outer shell, the tunic or skin. Up to 60 layers have been counted in some wild species. While the plant appears dormant above the ground the flower stalk which will start to grow in the following spring, develops within the bulb surrounded by two to three deciduous leaves and their sheaths. The flower stem lies in the axil of the second true leaf.

 

The single leafless Plant stem stem or scape, appearing from early to late spring depending on the species, bears from 1 to 20 blooms. Stem shape depends on the species, some are highly compressed with a visible seam, while others are rounded. The stems are upright and located at the centre of the leaves. In a few species such as N. hedraeanthus the stem is oblique. The stem is hollow in the upper portion but towards the bulb is more solid and filled with a spongy material.

 

Narcissus plants have one to several basal leaf leaves which are linear, ligulate or strap-shaped (long and narrow), sometimes channelled adaxially to semiterete, and may (pedicellate) or may not (sessile) have a petiole stalk. The leaves are flat and broad to cylindrical at the base and arise from the bulb. The emerging plant generally has two leaves, but the mature plant usually three, rarely four, and they are covered with a cutin containing cuticle, giving them a waxy appearance. Leaf colour is light green to blue-green. In the mature plant, the leaves extend higher than the flower stem, but in some species, the leaves are low-hanging. The leaf base is encased in a colorless sheath. After flowering, the leaves turn yellow and die back once the seed pod (fruit) is ripe.

 

Jonquils usually have dark green, round, rush-like leaves.

 

The inflorescence is scapose, the single stem or scape bearing either a solitary flower or forming an umbel with up to 20 blooms. Species bearing a solitary flower include section Bulbocodium and most of section Pseudonarcissus. Umbellate species have a fleshy racemose inflorescence (unbranched, with short floral stalks) with 2 to 15 or 20 flowers, such as N. papyraceus (see illustration, left) and N. tazetta (see Table I). The flower arrangement on the inflorescence may be either with (pedicellate) or without (sessile) floral stalks.

 

Prior to opening, the flower buds are enveloped and protected in a thin dry papery or membranous (scarious) spathe. The spathe consists of a singular bract that is ribbed, and which remains wrapped around the base of the open flower. As the bud grows, the spathe splits longitudinally. Bracteoles are small or absent.

 

The flowers of Narcissus are hermaphroditic (bisexual), have three parts (tripartite), and are sometimes fragrant (see Fragrances). The flower symmetry is actinomorphic (radial) to slightly zygomorphic (bilateral) due to declinate-ascending stamens (curving downwards, then bent up at the tip). Narcissus flowers are characterised by their, usually conspicuous, corona (trumpet).

 

The three major floral parts (in all species except N. cavanillesii in which the corona is virtually absent - Table I: Section Tapeinanthus) are;

 

(i) the proximal floral tube (hypanthium),

(ii) the surrounding free tepals, and

(iii) the more distal corona (paraperigon, paraperigonium).

All three parts may be considered to be components of the perianth (perigon, perigonium). The perianth arises above the apex of the inferior ovary, its base forming the hypanthial floral tube.

 

The floral tube is formed by fusion of the basal segments of the tepals (proximally connate). Its shape is from an inverted cone (obconic) to funnel-shaped (funneliform) or cylindrical, and is surmounted by the more distal corona. Floral tubes can range from long and narrow sections Apodanthi and Jonquilla to rudimentary (N. cavanillesii).

 

Surrounding the floral tube and corona and reflexed (bent back) from the rest of the perianth are the six spreading tepals or floral leaves, in two whorls which may be distally ascending, reflexed (folded back), or lanceolate. Like many monocotyledons, the perianth is homochlamydeous, which is undifferentiated into separate calyx (sepals) and corolla (petals), but rather has six tepals. The three outer tepal segments may be considered sepals, and the three inner segments petals. The transition point between the floral tube and the corona is marked by the insertion of the free tepals on the fused perianth.

 

The corona, or paracorolla, is variously described as bell-shaped (funneliform, trumpet), bowl-shaped (cupular, crateriform, cup-shaped) or disc-shaped with margins that are often frilled, and is free from the stamens. Rarely is the corona a simple callose (hardened, thickened) ring. The corona is formed during floral development as a tubular outgrowth from stamens which fuse into a tubular structure, the anthers becoming reduced. At its base, the fragrances which attract pollinators are formed. All species produce nectar at the top of the ovary. Coronal morphology varies from the tiny pigmented disk of N. serotinus (see Table I) or the rudimentary structure in N. cavanillesii to the elongated trumpets of section Pseudonarcissus (trumpet daffodils, Table I).

 

While the perianth may point forwards, in some species such as N. cyclamineus it is folded back (reflexed, see illustration, left), while in some other species such as N. bulbocodium (Table I), it is reduced to a few barely visible pointed segments with a prominent corona.

 

The colour of the perianth is white, yellow or bicoloured, with the exception of the night flowering N. viridiflorus which is green. In addition the corona of N. poeticus has a red crenulate margin (see Table I). Flower diameter varies from 12 mm (N. bulbocodium) to over 125 mm (N. nobilis=N. pseudonarcissus subsp. nobilis).

 

Flower orientation varies from pendent or deflexed (hanging down) as in N. triandrus (see illustration, left), through declinate-ascendant as in N. alpestris = N. pseudonarcissus subsp. moschatus, horizontal (patent, spreading) such as N. gaditanus or N. poeticus, erect as in N. cavanillesii, N. serotinus and N. rupicola (Table I), or intermediate between these positions (erecto-patent).

 

The flowers of Narcissus demonstrate exceptional floral diversity and sexual polymorphism, primarily by corona size and floral tube length, associated with pollinator groups (see for instance Figs. 1 and 2 in Graham and Barrett). Barrett and Harder (2005) describe three separate floral patterns;

 

"Daffodil" form

"Paperwhite" form

"Triandrus" form.

The predominant patterns are the 'daffodil' and 'paperwhite' forms, while the "triandrus" form is less common. Each corresponds to a different group of pollinators (See Pollination).

 

The "daffodil" form, which includes sections Pseudonarcissus and Bulbocodium, has a relatively short, broad or highly funnelform tube (funnel-like), which grades into an elongated corona, which is large and funnelform, forming a broad, cylindrical or trumpet-shaped perianth. Section Pseudonarcissus consists of relatively large flowers with a corolla length of around 50 mm, generally solitary but rarely in inflorescences of 2–4 flowers. They have wide greenish floral tubes with funnel-shaped bright yellow coronas. The six tepals sometimes differ in colour from the corona and may be cream coloured to pale yellow.

 

The "paperwhite" form, including sections Jonquilla, Apodanthi and Narcissus, has a relatively long, narrow tube and a short, shallow, flaring corona. The flower is horizontal and fragrant.

 

The "triandrus" form is seen in only two species, N. albimarginatus (a Moroccan endemic) and N. triandrus. It combines features of both the "daffodil" and "paperwhite" forms, with a well-developed, long, narrow tube and an extended bell-shaped corona of almost equal length. The flowers are pendent.

 

Androecium

There are six stamens in one to two rows (whorls), with the filaments separate from the corona, attached at the throat or base of the tube (epipetalous), often of two separate lengths, straight or declinate-ascending (curving downwards, then upwards). The anthers are basifixed (attached at their base).

 

Gynoecium

The ovary is inferior (below the floral parts) and trilocular (three chambered) and there is a pistil with a minutely three lobed stigma and filiform (thread like) style, which is often exserted (extending beyond the tube).

 

Fruit

The fruit consists of dehiscent loculicidal capsules (splitting between the locules) that are ellipsoid to subglobose (almost spherical) in shape and are papery to leathery in texture.

 

Seeds

The fruit contains numerous subglobose seeds which are round and swollen with a hard coat, sometimes with an attached elaiosome. The testa is black and the pericarp dry.

 

Most species have 12 ovules and 36 seeds, although some species such as N. bulbocodium have more, up to a maximum of 60. Seeds take five to six weeks to mature. The seeds of sections Jonquilla and Bulbocodium are wedge-shaped and matte black, while those of other sections are ovate and glossy black. A gust of wind or contact with a passing animal is sufficient to disperse the mature seeds.

 

Chromosomes

Chromosome numbers include 2n=14, 22, 26, with numerous aneuploid and polyploid derivatives. The basic chromosome number is 7, with the exception of N. tazetta, N. elegans and N. broussonetii in which it is 10 or 11; this subgenus (Hermione) was in fact characterised by this characteristic. Polyploid species include N. papyraceus (4x=22) and N. dubius (6x=50).

 

Phytochemistry

Alkaloids

As with all Amarylidaceae genera, Narcissus contains unique isoquinoline alkaloids. The first alkaloid to be identified was lycorine, from N. pseudonarcissus in 1877. These are considered a protective adaptation and are utilised in the classification of species. Nearly 100 alkaloids have been identified in the genus, about a third of all known Amaryllidaceae alkaloids, although not all species have been tested. Of the nine alkaloid ring types identified in the family, Narcissus species most commonly demonstrate the presence of alkaloids from within the Lycorine (lycorine, galanthine, pluviine) and Homolycorine (homolycorine, lycorenine) groups. Hemanthamine, tazettine, narciclasine, montanine and galantamine alkaloids are also represented. The alkaloid profile of any plant varies with time, location, and developmental stage. Narcissus also contain fructans and low molecular weight glucomannan in the leaves and plant stems.

 

Fragrances

Fragrances are predominantly monoterpene isoprenoids, with a small amount of benzenoids, although N. jonquilla has both equally represented. Another exception is N. cuatrecasasii which produces mainly fatty acid derivatives. The basic monoterpene precursor is geranyl pyrophosphate, and the commonest monoterpenes are limonene, myrcene, and trans-β-ocimene. Most benzenoids are non-methoxylated, while a few species contain methoxylated forms (ethers), e.g. N. bujei. Other ingredient include indole, isopentenoids and very small amounts of sesquiterpenes. Fragrance patterns can be correlated with pollinators, and fall into three main groups (see Pollination).

 

The taxonomy of Narcissus is complex, and still not fully resolved. Known to the ancients, the genus name appears in Graeco-Roman literature, although their interest was as much medicinal as botanical. It is unclear which species the ancients were familiar with. Although frequently mentioned in Mediaeval and Renaissance texts it was not formally described till the work of Linnaeus in 1753. By 1789 it had been grouped into a family (Narcissi) but shortly thereafter this was renamed Amaryllideae, from which comes the modern placement within Amaryllidaceae, although for a while it was considered part of Liliaceae.

 

Many of the species now considered to be Narcissus were in separate genera during the nineteenth century, and the situation was further confused by the inclusion of many cultivated varieties. By 1875 the current circumscription was relatively settled. By 2004 phylogenetic studies had allowed the place of Narcissus within its fairly large family to be established, nested within a series of subfamilies (Amaryllidoideae) and tribes (Narcisseae). It shares its position in the latter tribe with Sternbergia.

 

The infrageneric classification has been even more complex and many schemes of subgenera, sections, subsections and series have been proposed, although all had certain similarities. Most authorities now consider there to be 10 – 11 sections based on phylogenetic evidence. The problems have largely arisen from the diversity of the wild species, frequent natural hybridisation and extensive cultivation with escape and subsequent naturalisation. The number of species has varied anywhere from 16 to nearly 160, but is probably around 50 – 60.

 

The genus appeared some time in the Late Oligocene to Early Miocene eras, around 24 million years ago, in the Iberian peninsula. While the exact origin of the word Narcissus is unknown it is frequently linked to its fragrance which was thought to be narcotic, and to the legend of the youth of that name who fell in love with his reflection. In the English language the common name Daffodil appears to be derived from the Asphodel with which it was commonly compared.

 

Early

Narcissus was first described by Theophrastus (Θεόφραστος, c 371 - c 287 BC) in his Historia Plantarum (Greek: Περὶ φυτῶν ἱστορία) as νάρκισσος, referring to N. poeticus, but comparing it to Asphodelus (ασφοδελωδες). Theophrastus' description was frequently referred to at length by later authors writing in Latin such as Pliny the Elder (Gaius Plinius Secundus, 23 AD – 79 AD) from whom came the Latin form narcissus (see also Culture). Pliny's account is from his Natural History (Latin: Naturalis Historia). Like his contemporaries, his interests were as much therapeutic as botanical. Another much-cited Greek authority was Dioscorides (Διοσκουρίδης, 40 AD – 90 AD) in his De Materia Medica (Greek: Περὶ ὕλης ἰατρικῆς). Both authors were to remain influential until at least the Renaissance, given that their descriptions went beyond the merely botanical, to the therapeutic (see also Antiquity).

 

An early European reference is found in the work of Albert Magnus (c. 1200 – 1280), who noted in his De vegetabilibus et plantis the similarity to the leek. William Turner in his A New Herball (1551) cites all three extensively in his description of the plant and its properties.It was to remain to Linnaeus in 1753 to formally describe and name Narcissus as a genus in his Species Plantarum, at which time there were six known species (N. poeticus, N. pseudonarcissus, N. bulbocodium, N. serotinus, N. jonquilla and N. tazetta).[1] At that time, Linnaeus loosely grouped it together with 50 other genera into his Hexandria monogynia.

 

Modern

It was de Jussieu in 1789 who first formally created a 'family' (Narcissi), as the seventh 'Ordo' (Order) of the third class (Stamina epigyna) of Monocots in which Narcissus and 15 other genera were placed. The use of the term Ordo at that time was closer to what we now understand as Family, rather than Order. The family has undergone much reorganisation since then, but in 1805 it was renamed after a different genus in the family, Amaryllis, as 'Amaryllideae' by Jaume St.-Hilaire and has retained that association since. Jaume St.-Hilaire divided the family into two unnamed sections and recognised five species of Narcissus, omitting N. serotinus.

 

De Candolle brought together Linnaeus' genera and Jussieau's families into a systematic taxonomy for the first time, but included Narcissus (together with Amaryllis) in the Liliaceae in his Flore française (1805-1815) rather than Amaryllidaceae, a family he had not yet recognised. Shortly thereafter he separated the 'Amaryllidées' from 'Liliacées' (1813), though attributing the term to Brown's 'Amaryllideae' in the latter's Prodromus (1810) rather than St.-Hilaire's 'Amaryllidées'. He also provided the text to the first four volumes of Redouté illustrations in the latter's Les liliacées between 1805 and 1808 (see illustration here of N. candidissimus).

 

Historically both wide and narrow interpretations of the genus have been proposed. In the nineteenth century genus splitting was common, favouring the narrow view. Haworth (1831) using a narrow view treated many species as separate genera, as did Salisbury (1866). These authors listed various species in related genera such as Queltia (hybrids), Ajax (=Pseudonarcissus) and Hermione (=Tazettae), sixteen in all in Haworth's classification. In contrast, Herbert (1837) took a very wide view reducing Harworth's sixteen genera to six. Herbert, treating the Amaryllidacea as an 'order' as was common then, considered the narcissi to be a suborder, the Narcisseae, the six genera being Corbularia, Ajax, Ganymedes, Queltia, Narcissus and Hermione and his relatively narrow circumscription of Narcissus having only three species. Later Spach (1846) took an even wider view bringing most of Harworth's genera into the genus Narcissus, but as separate subgenera. By the time that Baker (1875) wrote his monograph all of the genera with one exception were included as Narcissus. The exception was the monotypic group Tapeinanthus which various subsequent authors have chosen to either exclude (e.g. Cullen 1986) or include (e.g. Webb 1978, 1980). Today it is nearly always included.

 

The eventual position of Narcissus within the Amaryllidaceae family only became settled in the twenty-first century with the advent of phylogenetic analysis and the Angiosperm Phylogeny Group system. The genus Narcissus belongs to the Narcisseae tribe, one of 13 within the Amaryllidoideae subfamily of the Amaryllidaceae. It is one of two sister clades corresponding to genera in the Narcisseae, being distinguished from Sternbergia by the presence of a paraperigonium, and is monophyletic

 

The infrageneric phylogeny of Narcissus still remains relatively unsettled. The taxonomy has proved very complex and difficult to resolve, particularly for the Pseudonarcissus group. This is due to a number of factors, including the diversity of the wild species, the ease with which natural hybridisation occurs, and extensive cultivation and breeding accompanied by escape and naturalisation.

 

De Candolle, in the first systematic taxonomy of Narcissus, arranged the species into named groups, and those names (Faux-Narcisse or Pseudonarcissus, Poétiques, Tazettes, Bulbocodiens, Jonquilles) have largely endured for the various subdivisions since and bear his name. The evolution of classification was confused by including many unknown or garden varieties, until Baker (1875) made the important distinction of excluding all specimens except the wild species from his system. He then grouped all of the earlier related genera as sections under one genus, Narcissus, the exception being the monotypic Tapeinanthus. Consequently, the number of accepted species has varied widely.

 

A common modern classification system has been that of Fernandes (1951, 1968, 1975) based on cytology, as modified by Blanchard (1990) and Mathew (2002), although in some countries such as Germany, the system of Meyer (1966) was preferred. Fernandes described two subgenera based on basal chromosome number, Hermione, n = 5 (11) and Narcissus, n = 7 (13). He further subdivided these into ten sections (Apodanthi, Aurelia, Bulbocodii, Ganymedes, Jonquillae, Narcissus, Pseudonarcissi, Serotini, Tapeinanthus, Tazettae), as did Blanchard later.

 

In contrast to Fernandes, Webb's treatment of the genus for the Flora Europaea (1978, 1980) prioritised morphology over genetics, and abandoned the subgenera ranks. He also restored De Candolle's original nomenclature, and made a number of changes to section Jonquilla, merging the existing subsections, reducing Apodanthi to a subsection of Jonquilla, and moving N. viridiflorus from Jonquilla to a new monotypic section of its own (Chloranthi). Finally, he divided Pseudonarcissus into two subsections. Blanchard (1990), whose Narcissus: a guide to wild daffodils has been very influential, adopted a simple approach, restoring Apodanthae, and based largely on ten sections alone.

 

The Royal Horticultural Society (RHS) currently lists ten sections, based on Fernandes (1968), three of which are monotypic (contain only one species), while two others only containing two species. Most species are placed in Pseudonarcissus While infrageneric groupings within Narcissus have been relatively constant, their status (genera, subgenera, sections, subsections, series, species) has not. Some authors treat some sections as being further subdivided into subsections, e.g. Tazettae (3 subsections). These subdivisions correspond roughly to the popular names for narcissi types, e.g. Trumpet Daffodils, Tazettas, Pheasant's Eyes, Hoop Petticoats, Jonquils.

 

While Webb had simply divided the genus into sections, Mathew found this unsatisfactory, implying every section had equal status. He adapted both Fernandes and Webb to devise a more hierarchical scheme he believed better reflected the interrelationships within the genus. Mathew's scheme consists of three subgenera (Narcissus, Hermione and Corbularia). The first two subgenera were then divided into five and two sections respectively. He then further subdivided two of the sections (subgenus Narcissus section Jonquillae, and subgenus Hermione section Hermione) into three subsections each. Finally, he divided section Hermione subsection Hermione further into two series, Hermione and Albiflorae. While lacking a phylogenetic basis, the system is still in use in horticulture. For instance the Pacific Bulb Society uses his numbering system for classifying species.

 

The phylogenetic analysis of Graham and Barrett (2004) supported the infrageneric division of Narcissus into two clades corresponding to the subgenera Hermione and Narcissus, but does not support monophyly of all sections, with only Apodanthi demonstrating clear monophyly, corresponding to Clade III of Graham and Barrett, although some other clades corresponded approximately to known sections. These authors examined 36 taxa of the 65 listed then, and a later extended analysis by Rønsted et al. (2008) with five additional taxa confirmed this pattern.

 

A very large (375 accessions) molecular analysis by Zonneveld (2008) utilising nuclear DNA content sought to reduce some of the paraphyly identified by Graham and Barrett. This led to a revision of the sectional structure, shifting some species between sections, eliminating one section and creating two new ones. In subgenus Hermione, Aurelia was merged with Tazettae. In subgenus Narcissus section Jonquillae subsection Juncifolii was elevated to sectional rank, thus resolving the paraphyly in this section observed by Graham and Barrett in Clade II due to this anomalous subsection, the remaining species being in subsection Jonquillae, which was monophyletic. The relatively large section Pseudonarcissi was divided by splitting off a new section, Nevadensis (species from southern Spain) leaving species from France, northern Spain and Portugal in the parent section. At the same time Fernández-Casas (2008) proposed a new monotypic section Angustini to accommodate Narcissus deficiens, placing it within subgenus Hermione.

 

While Graham and Barrett (2004) had determined that subgenus Hermione was monophyletic, using a much larger accession Santos-Gally et al. (2011) did not. However the former had excluded species of hybrid origins, while the latter included both N. dubius and N. tortifolius. If these two species are excluded (forming a clade with subgenus Narcissus) then Hermione can be considered monophyletic, although as a section of Hermione, Tazettae is not monophyletic. They also confirmed the monophyly of Apodanthi.

 

Some so-called nothosections have been proposed, predominantly by Fernández-Casas, to accommodate natural ('ancient') hybrids (nothospecies).

 

Subgenera and sections

Showing revisions by Zonnefeld (2008)

 

subgenus Hermione (Haw.) Spach.

(Aurelia (Gay) Baker (monotypic) - merged with Tazettae (2008)

Serotini Parlatore (2 species)

Tazettae de Candolle (16 species) syn. Hermione (Salisbury) Sprengel, in Fernandes' scheme. Incorporating Aurelia (2008)

subgenus Narcissus L.

Apodanthi A. Fernandes (6 species)

Bulbocodium de Candolle (11 species)

Ganymedes (Haworth) Schultes f. (monotypic)

Jonquillae de Candolle (8 species)

Juncifolii (A. Fern.) Zonn. sect. nov. (2008)

Narcissus L. (2 species)

Nevadensis Zonn. sect. nov. (2008)

Pseudonarcissus de Candolle (36 species) Trumpet daffodils

Tapeinanthus (Herbert) Traub (monotypic)

 

Species

Estimates of the number of species in Narcissus have varied widely, from anywhere between 16 and nearly 160, even in the modern era. Linnaeus originally included six species in 1753. By the time of the 14th edition of the Systema Naturae in 1784, there were fourteen. The 1819 Encyclopaedia Londinensis lists sixteen (see illustration here of three species) and by 1831 Adrian Haworth had described 150 species.

 

Much of the variation lies in the definition of species, and whether closely related taxa are considered separate species or subspecies. Thus, a very wide view of each species, such as Webb's results in few species, while a very narrow view such as that of Fernandes results in a larger number. Another factor is the status of hybrids, given natural hybridisation, with a distinction between 'ancient hybrids' and 'recent hybrids'. The term 'ancient hybrid' refers to hybrids found growing over a large area, and therefore now considered as separate species, while 'recent hybrid' refers to solitary plants found amongst their parents, with a more restricted range.

 

In the twentieth century Fernandes (1951) accepted 22 species, on which were based the 27 species listed by Webb in the 1980 Flora Europaea. By 1968, Fernandes had accepted 63 species, and by 1990 Blanchard listed 65 species, and Erhardt 66 in 1993. In 2006 the Royal Horticultural Society's (RHS) International Daffodil Register and Classified List listed 87 species, while Zonneveld's genetic study (2008) resulted in only 36. As of September 2014, the World Checklist of Selected Plant Families accepts 52 species, along with at least 60 hybrids, while the RHS has 81 accepted names in its October 2014 list.

 

Evolution

Within the Narcisseae, Narcissus (western Mediterranean) diverged from Sternbergia (Eurasia) some time in the Late Oligocene to Early Miocene eras, around 29.3–18.1 Ma, with a best estimate of 23.6 Ma. Later the genus divided into the two subgenera (Hermione and Narcissus) between 27.4 and 16.1 Ma (21.4 Ma). The divisions between the sections of Hermione then took place during the Miocene period 19.9–7.8 Ma.

 

Narcissus appears to have arisen in the area of the Iberian peninsula, southern France and northwestern Italy, and within this area most sections of the genus appeared, with only a few taxa being dispersed to North Africa at a time when the African and West European platforms were closer together. Subgenus Hermione in turn arose in the southwestern mediterranean and north west Africa. However, these are reconstructions, the Amaryllidaceae lacking a fossil record.

 

Names and etymology

The derivation of the Latin narcissus (Greek: νάρκισσος) is unknown. It may be a loanword from another language; for instance, it is said to be related to the Sanskrit word nark, meaning 'hell'. It is frequently linked to the Greek myth of Narcissus described by Ovid in his Metamorphoses, who became so obsessed with his own reflection that as he knelt and gazed into a pool of water, he fell into the water and drowned. In some variations, he died of starvation and thirst. In both versions, the narcissus plant sprang from where he died. Although Ovid appeared to describe the plant we now know as Narcissus there is no evidence for this popular derivation, and the person's name may have come from the flower's name. The Poet's Narcissus (N. poeticus), which grows in Greece, has a fragrance that has been described as intoxicating. This explanation is largely discredited due to lack of proof. Pliny wrote that the plant 'narce narcissum dictum, non a fabuloso puero' ('named narcissus from narce, not from the legendary youth'), i.e. that it was named for its narcotic properties (ναρκάω narkao, 'I grow numb' in Greek), not from the legend. Furthermore, there were accounts of narcissi growing, such as in the legend of Persephone, long before the story of Narcissus appeared (see Greek culture). It has also been suggested that daffodils bending over streams evoked the image of the youth admiring his own reflection in the water.

 

Linnaeus used the Latin name for the plant in formally describing the genus, although Matthias de l'Obel had previously used the name in describing various species of Narcissi in his Icones stirpium of 1591, and other publications, as had Clusius in Rariorum stirpium (1576).

 

The plural form of the common name narcissus has caused some confusion. British English sources such as the Shorter Oxford English Dictionary give two alternate forms, narcissi and narcissuses. In contrast, in American English the Merriam-Webster Dictionary provides for a third form, narcissus, used for both singular and plural. The Oxford dictionaries only list this third form under American English, although the Cambridge Dictionary allows of all three in the same order. However, Garner's Modern American Usage states that narcissi is the commonest form, narcissuses being excessively sibilant. For similar reasons, Fowler prefers narcissi in British English usage. Neither support narcissus as a plural form. Common names such as narcissus do not capitalise the first letter in contrast to the person of that name and the Latin genus name.

 

The name Narcissus (feminine Narcissa) was not uncommon in Roman times, such as Tiberius Claudius Narcissus, a Roman official in Claudius' time, an early New Testament Christian in Rome and later bishops and saints.

 

Daffodil

The word daffodil was unknown in the English language before the sixteenth century. The name is derived from an earlier affodell, a variant of asphodel. In classical Greek literature the narcissus is frequently referred to as the asphodel, such as the meadows of the Elysian fields in Homer (see Antiquity). Asphodel in turn appears to be a loanword coming from French via Mediaeval Latin affodilus from Classical Latin asphodilus and ultimately the Greek asphodelos (Greek: ἀσφόδελος). The reason for the introduction of the initial d is not known, although a probable source is an etymological merging from the Dutch article de, as in de affodil, or English the, as th'affodil or t'affodil, hence daffodil, and in French de and affodil to form fleur d'aphrodille and daphrodille.

 

From at least the 16th century, daffadown dilly and daffydowndilly have appeared as playful synonyms of the name. In common parlance and in historical documents, the term daffodil may refer specifically to populations or specimens of the wild daffodil, N. pseudonarcissus. H. N. Ellacombe suggests this may be from Saffon Lilly, citing Prior in support, though admittedly conjectural.

 

Lady Wilkinson (1858), who provides an extensive discussion of the etymology of the various names for this plant, suggests a very different origin, namely the Old English word affodyle (that which cometh early), citing a 14th-century (but likely originally much earlier) manuscript in support of this theory, and which appears to describe a plant resembling the daffodil. Ellacombe provides further support for this from a fifteenth century English translation of Palladius that also refers to it.

 

Jonquil

The name jonquil is said to be a corruption via French from the Latin juncifolius meaning 'rush-leaf' (Juncaceae) and its use is generally restricted to those species and cultivars which have rush like leaves, e.g. N. juncifolius.

 

Other

A profusion of names have attached themselves in the English language, either to the genus as a whole or to individual species or groups of species such as sections. These include narcissus, jonquil, Lent lily, Lenten lily, lide lily, yellow lily, wort or wyrt, Julians, glens, Lent cocks, corn flower, bell rose, asphodel, Solomon's lily, gracy day, haverdrils, giggary, cowslip, and crow foot.

plants - aloe vera

I understand that it flowers once every 10 years and then only for about 3 days. There were many photographers at the Begonia House getting snap shots of it in flower. It's name is Amorphophallus konjac and is a large, perennial exotic Asian plant. It grows a single, elongated center called a corm and a single large leaf that wraps around it. The corm is the part of the plant that is used, as well as its tuber, which is commonly called the konnyaku potato. Glucomannan is its scientific name, and its other common names include konjac mannan, devil's tongue, konjac, elephant-foot yam, snake plant and konnyaku.

 

I story about this plant featured in the DominionPost this morning - Flower pungent smell of...

 

The metre-high flower is blooming for the first time in a decade at the garden's Begonia House and was attracting a lot of attention during the Spring Festival. The flower has been in bloom for two days and is only expected to last one more day.

Amorphophallus konjac is a large, perennial exotic Asian plant. It grows a single, elongated center called a corm and a single large leaf that wraps around it. The corm is the part of the plant that is used, as well as its tuber, which is commonly called the konnyaku potato. Glucomannan is its scientific name, and its other common names include konjac mannan, devil's tongue, konjac, elephant-foot yam, snake plant and konnyaku.

 

I story about this plant featured in the DomionPost this morning - Flower pungent smell of...

 

The metre-high flower is blooming for the first time in a decade at the garden's Begonia House and was attracting a lot of attention during the Spring Festival. The flower has been in bloom for two days and is only expected to last one more day.

    

Salep (Turkish: salep, from Arabic: سحلب saḥlab‎, Azerbaijani: səhləb, Greek: σαλέπι salepi, Macedonian: салеп)

refers to both the orchid as well as to the salep drink.

It is a flour made from grinding the dried tubers of Orchis mascula, Orchis militaris and related species of orchids, which contain a nutritious starch-like polysaccharide called glucomannan.

Glucomannan Benefits, Safety Concerns, and Weight Loss Results t.co/D18P48Pw3M t.co/x12FyW7ZR6 (via Twitter twitter.com/gottahavefitcom/status/1100920837597814784)

The ice cream is thickened by both the salep flour and finely ground mastic resin, which comes mainly from the Greek island of Chios. The main thickener is the salep, which contains a carbohydrate called glucomannan. Glucomannan's molecules get tangled and bond as the temperature of the ice cream is reduced, giving the sweet treat its unusual consistency. The ground mastic resin also helps to thicken, but primarily adds flavor and texture. Photo: Anne McBride

  

Podcast @ www.nyas.org/snc/podcasts.asp

A flour called salep or sachlav is made of the ground tubers of this or some other species of orchids.

 

It contains a nutritious starch-like polysaccharide called glucomannan. In some magical traditions, its root is called Adam and Eve Root. It is said that witches used tubers of this orchid in love potions.

  

Kebun Raya Bogor, Indonesia

(Bogor Botanical Gardens, Indonesia).

 

Note: Specimen unlabeled.

 

Amorphophallus variabilis Blume. Araceae. CN: [Malay and regional vernacular names - Walur, Acung, Iles-iles putih, Cocooan oray]. Native to native to Indonesia - Java, Madura and Kangean. A herbaceous, perennial plant producing 1 - 2 leaves from a tuberous root-stock that can be 15 cm in diameter. The leaves are up to 125 cm long on a stalk up to 120cm long. A flowering stem 15 - 80 cm tall is produced annually; Young shoots and tuber edible. Corm contains glucomannan about 50% of

fresh mass.

 

Synonym(s):

Amorphophallus variabilis var. cuspidifoliatus Engl. & Gehrm.

Amorphophallus variabilis var. immaculatus Hassk.

Amorphophallus zeylanicus Engl.

Arum caeruleopunctatum Zipp. ex Kunth

Arum stercorarium Zipp. ex Kunth

Brachyspatha variabilis (Blume) Schott

 

Ref. and suggested reading:

www.theplantlist.org/tpl/record/kew-8327

fpa.uitm.edu.my/ipc2012/images/stories/download/journalag...

id.wikipedia.org/wiki/Walur

theferns.info/tropical/viewtropical.php?id=Amorphophallus...

Narcissus is a genus of predominantly spring flowering perennial plants of the amaryllis family, Amaryllidaceae. Various common names including daffodil, narcissus, and jonquil are used to describe all or some members of the genus. Narcissus has conspicuous flowers with six petal-like tepals surmounted by a cup- or trumpet-shaped corona. The flowers are generally white and yellow (also orange or pink in garden varieties), with either uniform or contrasting coloured tepals and corona.

 

Narcissus were well known in ancient civilisation, both medicinally and botanically, but formally described by Linnaeus in his Species Plantarum (1753). The genus is generally considered to have about ten sections with approximately 50 species. The number of species has varied, depending on how they are classified, due to similarity between species and hybridisation. The genus arose some time in the Late Oligocene to Early Miocene epochs, in the Iberian peninsula and adjacent areas of southwest Europe. The exact origin of the name Narcissus is unknown, but it is often linked to a Greek word (ancient Greek ναρκῶ narkō, "to make numb") and the myth of the youth of that name who fell in love with his own reflection. The English word "daffodil" appears to be derived from "asphodel", with which it was commonly compared.

 

The species are native to meadows and woods in southern Europe and North Africa with a centre of diversity in the Western Mediterranean, particularly the Iberian peninsula. Both wild and cultivated plants have naturalised widely, and were introduced into the Far East prior to the tenth century. Narcissi tend to be long-lived bulbs, which propagate by division, but are also insect-pollinated. Known pests, diseases and disorders include viruses, fungi, the larvae of flies, mites and nematodes. Some Narcissus species have become extinct, while others are threatened by increasing urbanisation and tourism.

 

Historical accounts suggest narcissi have been cultivated from the earliest times, but became increasingly popular in Europe after the 16th century and by the late 19th century were an important commercial crop centred primarily in the Netherlands. Today narcissi are popular as cut flowers and as ornamental plants in private and public gardens. The long history of breeding has resulted in thousands of different cultivars. For horticultural purposes, narcissi are classified into divisions, covering a wide range of shapes and colours. Like other members of their family, narcissi produce a number of different alkaloids, which provide some protection for the plant, but may be poisonous if accidentally ingested. This property has been exploited for medicinal use in traditional healing and has resulted in the production of galantamine for the treatment of Alzheimer's dementia. Long celebrated in art and literature, narcissi are associated with a number of themes in different cultures, ranging from death to good fortune, and as symbols of spring.

 

The daffodil is the national flower of Wales and the symbol of cancer charities in many countries. The appearance of the wild flowers in spring is associated with festivals in many places.

 

Narcissus is a genus of perennial herbaceous bulbiferous geophytes, which die back after flowering to an underground storage bulb. They regrow in the following year from brown-skinned ovoid bulbs with pronounced necks, and reach heights of 5–80 centimetres (2.0–31.5 in) depending on the species. Dwarf species such as N. asturiensis have a maximum height of 5–8 centimetres (2.0–3.1 in), while Narcissus tazetta may grow as tall as 80 centimetres (31 in).

 

The plants are scapose, having a single central leafless hollow flower stem (scape). Several green or blue-green, narrow, strap-shaped leaves arise from the bulb. The plant stem usually bears a solitary flower, but occasionally a cluster of flowers (umbel). The flowers, which are usually conspicuous and white or yellow, sometimes both or rarely green, consist of a perianth of three parts. Closest to the stem (proximal) is a floral tube above the ovary, then an outer ring composed of six tepals (undifferentiated sepals and petals), and a central disc to conical shaped corona. The flowers may hang down (pendant), or be erect. There are six pollen bearing stamens surrounding a central style. The ovary is inferior (below the floral parts) consisting of three chambers (trilocular). The fruit consists of a dry capsule that splits (dehisces) releasing numerous black seeds.

 

The bulb lies dormant after the leaves and flower stem die back and has contractile roots that pull it down further into the soil. The flower stem and leaves form in the bulb, to emerge the following season. Most species are dormant from summer to late winter, flowering in the spring, though a few species are autumn flowering.

 

The pale brown-skinned ovoid tunicate bulbs have a membranous tunic and a corky stem (base or basal) plate from which arise the adventitious root hairs in a ring around the edge, which grow up to 40 mm in length. Above the stem plate is the storage organ consisting of bulb scales, surrounding the previous flower stalk and the terminal bud. The scales are of two types, true storage organs and the bases of the foliage leaves. These have a thicker tip and a scar from where the leaf lamina became detached. The innermost leaf scale is semicircular only partly enveloping the flower stalk (semisheathed).(see Hanks Fig 1.3). The bulb may contain a number of branched bulb units, each with two to three true scales and two to three leaf bases. Each bulb unit has a life of about four years.

 

Once the leaves die back in summer, the roots also wither. After some years, the roots shorten pulling the bulbs deeper into the ground (contractile roots). The bulbs develop from the inside, pushing the older layers outwards which become brown and dry, forming an outer shell, the tunic or skin. Up to 60 layers have been counted in some wild species. While the plant appears dormant above the ground the flower stalk which will start to grow in the following spring, develops within the bulb surrounded by two to three deciduous leaves and their sheaths. The flower stem lies in the axil of the second true leaf.

 

The single leafless Plant stem stem or scape, appearing from early to late spring depending on the species, bears from 1 to 20 blooms. Stem shape depends on the species, some are highly compressed with a visible seam, while others are rounded. The stems are upright and located at the centre of the leaves. In a few species such as N. hedraeanthus the stem is oblique. The stem is hollow in the upper portion but towards the bulb is more solid and filled with a spongy material.

 

Narcissus plants have one to several basal leaf leaves which are linear, ligulate or strap-shaped (long and narrow), sometimes channelled adaxially to semiterete, and may (pedicellate) or may not (sessile) have a petiole stalk. The leaves are flat and broad to cylindrical at the base and arise from the bulb. The emerging plant generally has two leaves, but the mature plant usually three, rarely four, and they are covered with a cutin containing cuticle, giving them a waxy appearance. Leaf colour is light green to blue-green. In the mature plant, the leaves extend higher than the flower stem, but in some species, the leaves are low-hanging. The leaf base is encased in a colorless sheath. After flowering, the leaves turn yellow and die back once the seed pod (fruit) is ripe.

 

Jonquils usually have dark green, round, rush-like leaves.

 

The inflorescence is scapose, the single stem or scape bearing either a solitary flower or forming an umbel with up to 20 blooms. Species bearing a solitary flower include section Bulbocodium and most of section Pseudonarcissus. Umbellate species have a fleshy racemose inflorescence (unbranched, with short floral stalks) with 2 to 15 or 20 flowers, such as N. papyraceus (see illustration, left) and N. tazetta (see Table I). The flower arrangement on the inflorescence may be either with (pedicellate) or without (sessile) floral stalks.

 

Prior to opening, the flower buds are enveloped and protected in a thin dry papery or membranous (scarious) spathe. The spathe consists of a singular bract that is ribbed, and which remains wrapped around the base of the open flower. As the bud grows, the spathe splits longitudinally. Bracteoles are small or absent.

 

The flowers of Narcissus are hermaphroditic (bisexual), have three parts (tripartite), and are sometimes fragrant (see Fragrances). The flower symmetry is actinomorphic (radial) to slightly zygomorphic (bilateral) due to declinate-ascending stamens (curving downwards, then bent up at the tip). Narcissus flowers are characterised by their, usually conspicuous, corona (trumpet).

 

The three major floral parts (in all species except N. cavanillesii in which the corona is virtually absent - Table I: Section Tapeinanthus) are;

 

(i) the proximal floral tube (hypanthium),

(ii) the surrounding free tepals, and

(iii) the more distal corona (paraperigon, paraperigonium).

All three parts may be considered to be components of the perianth (perigon, perigonium). The perianth arises above the apex of the inferior ovary, its base forming the hypanthial floral tube.

 

The floral tube is formed by fusion of the basal segments of the tepals (proximally connate). Its shape is from an inverted cone (obconic) to funnel-shaped (funneliform) or cylindrical, and is surmounted by the more distal corona. Floral tubes can range from long and narrow sections Apodanthi and Jonquilla to rudimentary (N. cavanillesii).

 

Surrounding the floral tube and corona and reflexed (bent back) from the rest of the perianth are the six spreading tepals or floral leaves, in two whorls which may be distally ascending, reflexed (folded back), or lanceolate. Like many monocotyledons, the perianth is homochlamydeous, which is undifferentiated into separate calyx (sepals) and corolla (petals), but rather has six tepals. The three outer tepal segments may be considered sepals, and the three inner segments petals. The transition point between the floral tube and the corona is marked by the insertion of the free tepals on the fused perianth.

 

The corona, or paracorolla, is variously described as bell-shaped (funneliform, trumpet), bowl-shaped (cupular, crateriform, cup-shaped) or disc-shaped with margins that are often frilled, and is free from the stamens. Rarely is the corona a simple callose (hardened, thickened) ring. The corona is formed during floral development as a tubular outgrowth from stamens which fuse into a tubular structure, the anthers becoming reduced. At its base, the fragrances which attract pollinators are formed. All species produce nectar at the top of the ovary. Coronal morphology varies from the tiny pigmented disk of N. serotinus (see Table I) or the rudimentary structure in N. cavanillesii to the elongated trumpets of section Pseudonarcissus (trumpet daffodils, Table I).

 

While the perianth may point forwards, in some species such as N. cyclamineus it is folded back (reflexed, see illustration, left), while in some other species such as N. bulbocodium (Table I), it is reduced to a few barely visible pointed segments with a prominent corona.

 

The colour of the perianth is white, yellow or bicoloured, with the exception of the night flowering N. viridiflorus which is green. In addition the corona of N. poeticus has a red crenulate margin (see Table I). Flower diameter varies from 12 mm (N. bulbocodium) to over 125 mm (N. nobilis=N. pseudonarcissus subsp. nobilis).

 

Flower orientation varies from pendent or deflexed (hanging down) as in N. triandrus (see illustration, left), through declinate-ascendant as in N. alpestris = N. pseudonarcissus subsp. moschatus, horizontal (patent, spreading) such as N. gaditanus or N. poeticus, erect as in N. cavanillesii, N. serotinus and N. rupicola (Table I), or intermediate between these positions (erecto-patent).

 

The flowers of Narcissus demonstrate exceptional floral diversity and sexual polymorphism, primarily by corona size and floral tube length, associated with pollinator groups (see for instance Figs. 1 and 2 in Graham and Barrett). Barrett and Harder (2005) describe three separate floral patterns;

 

"Daffodil" form

"Paperwhite" form

"Triandrus" form.

The predominant patterns are the 'daffodil' and 'paperwhite' forms, while the "triandrus" form is less common. Each corresponds to a different group of pollinators (See Pollination).

 

The "daffodil" form, which includes sections Pseudonarcissus and Bulbocodium, has a relatively short, broad or highly funnelform tube (funnel-like), which grades into an elongated corona, which is large and funnelform, forming a broad, cylindrical or trumpet-shaped perianth. Section Pseudonarcissus consists of relatively large flowers with a corolla length of around 50 mm, generally solitary but rarely in inflorescences of 2–4 flowers. They have wide greenish floral tubes with funnel-shaped bright yellow coronas. The six tepals sometimes differ in colour from the corona and may be cream coloured to pale yellow.

 

The "paperwhite" form, including sections Jonquilla, Apodanthi and Narcissus, has a relatively long, narrow tube and a short, shallow, flaring corona. The flower is horizontal and fragrant.

 

The "triandrus" form is seen in only two species, N. albimarginatus (a Moroccan endemic) and N. triandrus. It combines features of both the "daffodil" and "paperwhite" forms, with a well-developed, long, narrow tube and an extended bell-shaped corona of almost equal length. The flowers are pendent.

 

Androecium

There are six stamens in one to two rows (whorls), with the filaments separate from the corona, attached at the throat or base of the tube (epipetalous), often of two separate lengths, straight or declinate-ascending (curving downwards, then upwards). The anthers are basifixed (attached at their base).

 

Gynoecium

The ovary is inferior (below the floral parts) and trilocular (three chambered) and there is a pistil with a minutely three lobed stigma and filiform (thread like) style, which is often exserted (extending beyond the tube).

 

Fruit

The fruit consists of dehiscent loculicidal capsules (splitting between the locules) that are ellipsoid to subglobose (almost spherical) in shape and are papery to leathery in texture.

 

Seeds

The fruit contains numerous subglobose seeds which are round and swollen with a hard coat, sometimes with an attached elaiosome. The testa is black and the pericarp dry.

 

Most species have 12 ovules and 36 seeds, although some species such as N. bulbocodium have more, up to a maximum of 60. Seeds take five to six weeks to mature. The seeds of sections Jonquilla and Bulbocodium are wedge-shaped and matte black, while those of other sections are ovate and glossy black. A gust of wind or contact with a passing animal is sufficient to disperse the mature seeds.

 

Chromosomes

Chromosome numbers include 2n=14, 22, 26, with numerous aneuploid and polyploid derivatives. The basic chromosome number is 7, with the exception of N. tazetta, N. elegans and N. broussonetii in which it is 10 or 11; this subgenus (Hermione) was in fact characterised by this characteristic. Polyploid species include N. papyraceus (4x=22) and N. dubius (6x=50).

 

Phytochemistry

Alkaloids

As with all Amarylidaceae genera, Narcissus contains unique isoquinoline alkaloids. The first alkaloid to be identified was lycorine, from N. pseudonarcissus in 1877. These are considered a protective adaptation and are utilised in the classification of species. Nearly 100 alkaloids have been identified in the genus, about a third of all known Amaryllidaceae alkaloids, although not all species have been tested. Of the nine alkaloid ring types identified in the family, Narcissus species most commonly demonstrate the presence of alkaloids from within the Lycorine (lycorine, galanthine, pluviine) and Homolycorine (homolycorine, lycorenine) groups. Hemanthamine, tazettine, narciclasine, montanine and galantamine alkaloids are also represented. The alkaloid profile of any plant varies with time, location, and developmental stage. Narcissus also contain fructans and low molecular weight glucomannan in the leaves and plant stems.

 

Fragrances

Fragrances are predominantly monoterpene isoprenoids, with a small amount of benzenoids, although N. jonquilla has both equally represented. Another exception is N. cuatrecasasii which produces mainly fatty acid derivatives. The basic monoterpene precursor is geranyl pyrophosphate, and the commonest monoterpenes are limonene, myrcene, and trans-β-ocimene. Most benzenoids are non-methoxylated, while a few species contain methoxylated forms (ethers), e.g. N. bujei. Other ingredient include indole, isopentenoids and very small amounts of sesquiterpenes. Fragrance patterns can be correlated with pollinators, and fall into three main groups (see Pollination).

 

The taxonomy of Narcissus is complex, and still not fully resolved. Known to the ancients, the genus name appears in Graeco-Roman literature, although their interest was as much medicinal as botanical. It is unclear which species the ancients were familiar with. Although frequently mentioned in Mediaeval and Renaissance texts it was not formally described till the work of Linnaeus in 1753. By 1789 it had been grouped into a family (Narcissi) but shortly thereafter this was renamed Amaryllideae, from which comes the modern placement within Amaryllidaceae, although for a while it was considered part of Liliaceae.

 

Many of the species now considered to be Narcissus were in separate genera during the nineteenth century, and the situation was further confused by the inclusion of many cultivated varieties. By 1875 the current circumscription was relatively settled. By 2004 phylogenetic studies had allowed the place of Narcissus within its fairly large family to be established, nested within a series of subfamilies (Amaryllidoideae) and tribes (Narcisseae). It shares its position in the latter tribe with Sternbergia.

 

The infrageneric classification has been even more complex and many schemes of subgenera, sections, subsections and series have been proposed, although all had certain similarities. Most authorities now consider there to be 10 – 11 sections based on phylogenetic evidence. The problems have largely arisen from the diversity of the wild species, frequent natural hybridisation and extensive cultivation with escape and subsequent naturalisation. The number of species has varied anywhere from 16 to nearly 160, but is probably around 50 – 60.

 

The genus appeared some time in the Late Oligocene to Early Miocene eras, around 24 million years ago, in the Iberian peninsula. While the exact origin of the word Narcissus is unknown it is frequently linked to its fragrance which was thought to be narcotic, and to the legend of the youth of that name who fell in love with his reflection. In the English language the common name Daffodil appears to be derived from the Asphodel with which it was commonly compared.

 

Early

Narcissus was first described by Theophrastus (Θεόφραστος, c 371 - c 287 BC) in his Historia Plantarum (Greek: Περὶ φυτῶν ἱστορία) as νάρκισσος, referring to N. poeticus, but comparing it to Asphodelus (ασφοδελωδες). Theophrastus' description was frequently referred to at length by later authors writing in Latin such as Pliny the Elder (Gaius Plinius Secundus, 23 AD – 79 AD) from whom came the Latin form narcissus (see also Culture). Pliny's account is from his Natural History (Latin: Naturalis Historia). Like his contemporaries, his interests were as much therapeutic as botanical. Another much-cited Greek authority was Dioscorides (Διοσκουρίδης, 40 AD – 90 AD) in his De Materia Medica (Greek: Περὶ ὕλης ἰατρικῆς). Both authors were to remain influential until at least the Renaissance, given that their descriptions went beyond the merely botanical, to the therapeutic (see also Antiquity).

 

An early European reference is found in the work of Albert Magnus (c. 1200 – 1280), who noted in his De vegetabilibus et plantis the similarity to the leek. William Turner in his A New Herball (1551) cites all three extensively in his description of the plant and its properties.It was to remain to Linnaeus in 1753 to formally describe and name Narcissus as a genus in his Species Plantarum, at which time there were six known species (N. poeticus, N. pseudonarcissus, N. bulbocodium, N. serotinus, N. jonquilla and N. tazetta).[1] At that time, Linnaeus loosely grouped it together with 50 other genera into his Hexandria monogynia.

 

Modern

It was de Jussieu in 1789 who first formally created a 'family' (Narcissi), as the seventh 'Ordo' (Order) of the third class (Stamina epigyna) of Monocots in which Narcissus and 15 other genera were placed. The use of the term Ordo at that time was closer to what we now understand as Family, rather than Order. The family has undergone much reorganisation since then, but in 1805 it was renamed after a different genus in the family, Amaryllis, as 'Amaryllideae' by Jaume St.-Hilaire and has retained that association since. Jaume St.-Hilaire divided the family into two unnamed sections and recognised five species of Narcissus, omitting N. serotinus.

 

De Candolle brought together Linnaeus' genera and Jussieau's families into a systematic taxonomy for the first time, but included Narcissus (together with Amaryllis) in the Liliaceae in his Flore française (1805-1815) rather than Amaryllidaceae, a family he had not yet recognised. Shortly thereafter he separated the 'Amaryllidées' from 'Liliacées' (1813), though attributing the term to Brown's 'Amaryllideae' in the latter's Prodromus (1810) rather than St.-Hilaire's 'Amaryllidées'. He also provided the text to the first four volumes of Redouté illustrations in the latter's Les liliacées between 1805 and 1808 (see illustration here of N. candidissimus).

 

Historically both wide and narrow interpretations of the genus have been proposed. In the nineteenth century genus splitting was common, favouring the narrow view. Haworth (1831) using a narrow view treated many species as separate genera, as did Salisbury (1866). These authors listed various species in related genera such as Queltia (hybrids), Ajax (=Pseudonarcissus) and Hermione (=Tazettae), sixteen in all in Haworth's classification. In contrast, Herbert (1837) took a very wide view reducing Harworth's sixteen genera to six. Herbert, treating the Amaryllidacea as an 'order' as was common then, considered the narcissi to be a suborder, the Narcisseae, the six genera being Corbularia, Ajax, Ganymedes, Queltia, Narcissus and Hermione and his relatively narrow circumscription of Narcissus having only three species. Later Spach (1846) took an even wider view bringing most of Harworth's genera into the genus Narcissus, but as separate subgenera. By the time that Baker (1875) wrote his monograph all of the genera with one exception were included as Narcissus. The exception was the monotypic group Tapeinanthus which various subsequent authors have chosen to either exclude (e.g. Cullen 1986) or include (e.g. Webb 1978, 1980). Today it is nearly always included.

 

The eventual position of Narcissus within the Amaryllidaceae family only became settled in the twenty-first century with the advent of phylogenetic analysis and the Angiosperm Phylogeny Group system. The genus Narcissus belongs to the Narcisseae tribe, one of 13 within the Amaryllidoideae subfamily of the Amaryllidaceae. It is one of two sister clades corresponding to genera in the Narcisseae, being distinguished from Sternbergia by the presence of a paraperigonium, and is monophyletic

 

The infrageneric phylogeny of Narcissus still remains relatively unsettled. The taxonomy has proved very complex and difficult to resolve, particularly for the Pseudonarcissus group. This is due to a number of factors, including the diversity of the wild species, the ease with which natural hybridisation occurs, and extensive cultivation and breeding accompanied by escape and naturalisation.

 

De Candolle, in the first systematic taxonomy of Narcissus, arranged the species into named groups, and those names (Faux-Narcisse or Pseudonarcissus, Poétiques, Tazettes, Bulbocodiens, Jonquilles) have largely endured for the various subdivisions since and bear his name. The evolution of classification was confused by including many unknown or garden varieties, until Baker (1875) made the important distinction of excluding all specimens except the wild species from his system. He then grouped all of the earlier related genera as sections under one genus, Narcissus, the exception being the monotypic Tapeinanthus. Consequently, the number of accepted species has varied widely.

 

A common modern classification system has been that of Fernandes (1951, 1968, 1975) based on cytology, as modified by Blanchard (1990) and Mathew (2002), although in some countries such as Germany, the system of Meyer (1966) was preferred. Fernandes described two subgenera based on basal chromosome number, Hermione, n = 5 (11) and Narcissus, n = 7 (13). He further subdivided these into ten sections (Apodanthi, Aurelia, Bulbocodii, Ganymedes, Jonquillae, Narcissus, Pseudonarcissi, Serotini, Tapeinanthus, Tazettae), as did Blanchard later.

 

In contrast to Fernandes, Webb's treatment of the genus for the Flora Europaea (1978, 1980) prioritised morphology over genetics, and abandoned the subgenera ranks. He also restored De Candolle's original nomenclature, and made a number of changes to section Jonquilla, merging the existing subsections, reducing Apodanthi to a subsection of Jonquilla, and moving N. viridiflorus from Jonquilla to a new monotypic section of its own (Chloranthi). Finally, he divided Pseudonarcissus into two subsections. Blanchard (1990), whose Narcissus: a guide to wild daffodils has been very influential, adopted a simple approach, restoring Apodanthae, and based largely on ten sections alone.

 

The Royal Horticultural Society (RHS) currently lists ten sections, based on Fernandes (1968), three of which are monotypic (contain only one species), while two others only containing two species. Most species are placed in Pseudonarcissus While infrageneric groupings within Narcissus have been relatively constant, their status (genera, subgenera, sections, subsections, series, species) has not. Some authors treat some sections as being further subdivided into subsections, e.g. Tazettae (3 subsections). These subdivisions correspond roughly to the popular names for narcissi types, e.g. Trumpet Daffodils, Tazettas, Pheasant's Eyes, Hoop Petticoats, Jonquils.

 

While Webb had simply divided the genus into sections, Mathew found this unsatisfactory, implying every section had equal status. He adapted both Fernandes and Webb to devise a more hierarchical scheme he believed better reflected the interrelationships within the genus. Mathew's scheme consists of three subgenera (Narcissus, Hermione and Corbularia). The first two subgenera were then divided into five and two sections respectively. He then further subdivided two of the sections (subgenus Narcissus section Jonquillae, and subgenus Hermione section Hermione) into three subsections each. Finally, he divided section Hermione subsection Hermione further into two series, Hermione and Albiflorae. While lacking a phylogenetic basis, the system is still in use in horticulture. For instance the Pacific Bulb Society uses his numbering system for classifying species.

 

The phylogenetic analysis of Graham and Barrett (2004) supported the infrageneric division of Narcissus into two clades corresponding to the subgenera Hermione and Narcissus, but does not support monophyly of all sections, with only Apodanthi demonstrating clear monophyly, corresponding to Clade III of Graham and Barrett, although some other clades corresponded approximately to known sections. These authors examined 36 taxa of the 65 listed then, and a later extended analysis by Rønsted et al. (2008) with five additional taxa confirmed this pattern.

 

A very large (375 accessions) molecular analysis by Zonneveld (2008) utilising nuclear DNA content sought to reduce some of the paraphyly identified by Graham and Barrett. This led to a revision of the sectional structure, shifting some species between sections, eliminating one section and creating two new ones. In subgenus Hermione, Aurelia was merged with Tazettae. In subgenus Narcissus section Jonquillae subsection Juncifolii was elevated to sectional rank, thus resolving the paraphyly in this section observed by Graham and Barrett in Clade II due to this anomalous subsection, the remaining species being in subsection Jonquillae, which was monophyletic. The relatively large section Pseudonarcissi was divided by splitting off a new section, Nevadensis (species from southern Spain) leaving species from France, northern Spain and Portugal in the parent section. At the same time Fernández-Casas (2008) proposed a new monotypic section Angustini to accommodate Narcissus deficiens, placing it within subgenus Hermione.

 

While Graham and Barrett (2004) had determined that subgenus Hermione was monophyletic, using a much larger accession Santos-Gally et al. (2011) did not. However the former had excluded species of hybrid origins, while the latter included both N. dubius and N. tortifolius. If these two species are excluded (forming a clade with subgenus Narcissus) then Hermione can be considered monophyletic, although as a section of Hermione, Tazettae is not monophyletic. They also confirmed the monophyly of Apodanthi.

 

Some so-called nothosections have been proposed, predominantly by Fernández-Casas, to accommodate natural ('ancient') hybrids (nothospecies).

 

Subgenera and sections

Showing revisions by Zonnefeld (2008)

 

subgenus Hermione (Haw.) Spach.

(Aurelia (Gay) Baker (monotypic) - merged with Tazettae (2008)

Serotini Parlatore (2 species)

Tazettae de Candolle (16 species) syn. Hermione (Salisbury) Sprengel, in Fernandes' scheme. Incorporating Aurelia (2008)

subgenus Narcissus L.

Apodanthi A. Fernandes (6 species)

Bulbocodium de Candolle (11 species)

Ganymedes (Haworth) Schultes f. (monotypic)

Jonquillae de Candolle (8 species)

Juncifolii (A. Fern.) Zonn. sect. nov. (2008)

Narcissus L. (2 species)

Nevadensis Zonn. sect. nov. (2008)

Pseudonarcissus de Candolle (36 species) Trumpet daffodils

Tapeinanthus (Herbert) Traub (monotypic)

 

Species

Estimates of the number of species in Narcissus have varied widely, from anywhere between 16 and nearly 160, even in the modern era. Linnaeus originally included six species in 1753. By the time of the 14th edition of the Systema Naturae in 1784, there were fourteen. The 1819 Encyclopaedia Londinensis lists sixteen (see illustration here of three species) and by 1831 Adrian Haworth had described 150 species.

 

Much of the variation lies in the definition of species, and whether closely related taxa are considered separate species or subspecies. Thus, a very wide view of each species, such as Webb's results in few species, while a very narrow view such as that of Fernandes results in a larger number. Another factor is the status of hybrids, given natural hybridisation, with a distinction between 'ancient hybrids' and 'recent hybrids'. The term 'ancient hybrid' refers to hybrids found growing over a large area, and therefore now considered as separate species, while 'recent hybrid' refers to solitary plants found amongst their parents, with a more restricted range.

 

In the twentieth century Fernandes (1951) accepted 22 species, on which were based the 27 species listed by Webb in the 1980 Flora Europaea. By 1968, Fernandes had accepted 63 species, and by 1990 Blanchard listed 65 species, and Erhardt 66 in 1993. In 2006 the Royal Horticultural Society's (RHS) International Daffodil Register and Classified List listed 87 species, while Zonneveld's genetic study (2008) resulted in only 36. As of September 2014, the World Checklist of Selected Plant Families accepts 52 species, along with at least 60 hybrids, while the RHS has 81 accepted names in its October 2014 list.

 

Evolution

Within the Narcisseae, Narcissus (western Mediterranean) diverged from Sternbergia (Eurasia) some time in the Late Oligocene to Early Miocene eras, around 29.3–18.1 Ma, with a best estimate of 23.6 Ma. Later the genus divided into the two subgenera (Hermione and Narcissus) between 27.4 and 16.1 Ma (21.4 Ma). The divisions between the sections of Hermione then took place during the Miocene period 19.9–7.8 Ma.

 

Narcissus appears to have arisen in the area of the Iberian peninsula, southern France and northwestern Italy, and within this area most sections of the genus appeared, with only a few taxa being dispersed to North Africa at a time when the African and West European platforms were closer together. Subgenus Hermione in turn arose in the southwestern mediterranean and north west Africa. However, these are reconstructions, the Amaryllidaceae lacking a fossil record.

 

Names and etymology

The derivation of the Latin narcissus (Greek: νάρκισσος) is unknown. It may be a loanword from another language; for instance, it is said to be related to the Sanskrit word nark, meaning 'hell'. It is frequently linked to the Greek myth of Narcissus described by Ovid in his Metamorphoses, who became so obsessed with his own reflection that as he knelt and gazed into a pool of water, he fell into the water and drowned. In some variations, he died of starvation and thirst. In both versions, the narcissus plant sprang from where he died. Although Ovid appeared to describe the plant we now know as Narcissus there is no evidence for this popular derivation, and the person's name may have come from the flower's name. The Poet's Narcissus (N. poeticus), which grows in Greece, has a fragrance that has been described as intoxicating. This explanation is largely discredited due to lack of proof. Pliny wrote that the plant 'narce narcissum dictum, non a fabuloso puero' ('named narcissus from narce, not from the legendary youth'), i.e. that it was named for its narcotic properties (ναρκάω narkao, 'I grow numb' in Greek), not from the legend. Furthermore, there were accounts of narcissi growing, such as in the legend of Persephone, long before the story of Narcissus appeared (see Greek culture). It has also been suggested that daffodils bending over streams evoked the image of the youth admiring his own reflection in the water.

 

Linnaeus used the Latin name for the plant in formally describing the genus, although Matthias de l'Obel had previously used the name in describing various species of Narcissi in his Icones stirpium of 1591, and other publications, as had Clusius in Rariorum stirpium (1576).

 

The plural form of the common name narcissus has caused some confusion. British English sources such as the Shorter Oxford English Dictionary give two alternate forms, narcissi and narcissuses. In contrast, in American English the Merriam-Webster Dictionary provides for a third form, narcissus, used for both singular and plural. The Oxford dictionaries only list this third form under American English, although the Cambridge Dictionary allows of all three in the same order. However, Garner's Modern American Usage states that narcissi is the commonest form, narcissuses being excessively sibilant. For similar reasons, Fowler prefers narcissi in British English usage. Neither support narcissus as a plural form. Common names such as narcissus do not capitalise the first letter in contrast to the person of that name and the Latin genus name.

 

The name Narcissus (feminine Narcissa) was not uncommon in Roman times, such as Tiberius Claudius Narcissus, a Roman official in Claudius' time, an early New Testament Christian in Rome and later bishops and saints.

 

Daffodil

The word daffodil was unknown in the English language before the sixteenth century. The name is derived from an earlier affodell, a variant of asphodel. In classical Greek literature the narcissus is frequently referred to as the asphodel, such as the meadows of the Elysian fields in Homer (see Antiquity). Asphodel in turn appears to be a loanword coming from French via Mediaeval Latin affodilus from Classical Latin asphodilus and ultimately the Greek asphodelos (Greek: ἀσφόδελος). The reason for the introduction of the initial d is not known, although a probable source is an etymological merging from the Dutch article de, as in de affodil, or English the, as th'affodil or t'affodil, hence daffodil, and in French de and affodil to form fleur d'aphrodille and daphrodille.

 

From at least the 16th century, daffadown dilly and daffydowndilly have appeared as playful synonyms of the name. In common parlance and in historical documents, the term daffodil may refer specifically to populations or specimens of the wild daffodil, N. pseudonarcissus. H. N. Ellacombe suggests this may be from Saffon Lilly, citing Prior in support, though admittedly conjectural.

 

Lady Wilkinson (1858), who provides an extensive discussion of the etymology of the various names for this plant, suggests a very different origin, namely the Old English word affodyle (that which cometh early), citing a 14th-century (but likely originally much earlier) manuscript in support of this theory, and which appears to describe a plant resembling the daffodil. Ellacombe provides further support for this from a fifteenth century English translation of Palladius that also refers to it.

 

Jonquil

The name jonquil is said to be a corruption via French from the Latin juncifolius meaning 'rush-leaf' (Juncaceae) and its use is generally restricted to those species and cultivars which have rush like leaves, e.g. N. juncifolius.

 

Other

A profusion of names have attached themselves in the English language, either to the genus as a whole or to individual species or groups of species such as sections. These include narcissus, jonquil, Lent lily, Lenten lily, lide lily, yellow lily, wort or wyrt, Julians, glens, Lent cocks, corn flower, bell rose, asphodel, Solomon's lily, gracy day, haverdrils, giggary, cowslip, and crow foot.

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