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Eagle Martial

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(Polemaetus bellicosus)

 

The martial eagle (Polemaetus bellicosus) is a large eagle native to sub-Saharan Africa. It is the only member of the genus Polemaetus. A species of the booted eagle sub-family (Aquillinae), it has feathering over its tarsus. One of the largest and most powerful species of booted eagle, it is a fairly opportunistic predator that varies it prey selection between mammals, birds and reptiles. Its hunting technique is unique as it is one of few eagle species known to hunt primarily from a high soar, by stooping on its quarry. An inhabitant of wooded belts of otherwise open savanna, this species has shown a precipitous decline in the last few centuries due to a variety of factors. The martial eagle is one of the most persecuted bird species in the world. Due to its habit of taking livestock and regionally valuable game, local farmers and game wardens frequently seek to eliminate martial eagles, although the effect of eagles on this prey is almost certainly considerably exaggerated. Currently, the martial eagle is classified with the status of Vulnerable to extinction by the IUCN.

 

The martial eagle can be found in most of sub-Saharan Africa, wherever food is abundant and the environment favourable. With a total estimated distribution of about 26,000 km2 (10,000 sq mi), it is fairly widely distributed in the continent, having a somewhat broader range than species like the crowned eagle (Stephanoaetus coronatus) and the Verreaux's eagle (Aquila verreauxii). Although never common, greater population densities do exist in southern Africa and in some parts of east Africa. Martial eagles tend to be rare and irregular in west Africa but are known to reside in Senegal, The Gambia and northern Guinea-Bissau, southern Mali and the northern portions of Ivory Coast and Ghana. From southern Niger and eastern Nigeria the species is distributed spottily through Chad, Sudan and the Central African Republic as well as the northern, eastern, and southern portions of the Democratic Republic of the Congo. In east Africa, they range from northwestern Somalia and Ethiopia more or less continuously south through Kenya, Uganda, Tanzania and in southern Africa from Angola, Zambia, Malawi and southern Mozambique to South Africa. Some of the larger remaining populations are known to persist in Zimbabwe and South Africa. Generally, these birds are more abundant in protected areas such as Kruger National Park and Kgalagadi Transfrontier Park in South Africa, or Etosha National Park in Namibia.

 

The Accipitridae (hereafter accipitrids) family is by far the most diverse family of diurnal raptors in the world with more than 230 currently accepted species. As a member of the booted eagle sub-family, Aquilinae, it is one of the roughly 15% extant species of the family to have feathers covering its legs. This may be a useful feature for distinguishing these species from other eagles and raptors, as they are present even in tropical species such as the martial eagle. Under current classifications, booted eagles consist of approximately 38 living species that are distributed in every continent inhabited by the accipitrids, which excludes only the continent of Antarctica. Just under half of the living species of booted eagle are found in Africa. Studies have been conducted on the mitochondrial DNA of most booted eagle species, including the martial eagle, in order to gain insight on how the sub-family is ordered and which species bear relation to one another. DNA testing in the 1980s indicated the martial eagle was a specialized off-shoot of the small-bodied Hieraaetus eagles and one study went so far as to advocate that the martial eagle be included in the genus. However, more modern and comprehensive genetic testing has shown that the martial eagle is extremely distinct from other living booted eagles and diverged from other existing genera several million years ago. Genetically, the martial eagle fell between two other species in monotypical genera, the African long-crested eagle (Lophaetus occipitalis) and the Asian rufous-bellied eagle (Lophotriorchis kienerii), that similarly diverged long ago from other modern species. Given the disparity of this species’ unique morphology and that the two aforementioned most closely related living species are only about as large as the bigger buzzards, the unique heritage of the martial eagle is considered even superficially evident. There are no subspecies of martial eagle and the species varies little in appearance and genetic diversity across its distribution.

 

The martial eagle is a very large eagle. In total length, it can range from 78 to 96 cm (31 to 38 in), with an average of approximately 85.5 cm (33.7 in). Its total length – in comparison to its wingspan – is restricted by its relatively short tail. Nonetheless, it appears to be the sixth or seventh longest living eagle species. The wingspan of martial eagles can range from 188 to 260 cm (6 ft 2 in to 8 ft 6 in). Average wingspans have been claimed of 205 cm (6 ft 9 in) and 207.5 cm (6 ft 10 in) for the species, however ten measured martial eagles in the wild were found to average 211.9 cm (6 ft 11 in) in wingspan. Thus, the martial eagle appears to average fourth in wingspan among living eagles, behind only the Steller's sea-eagle (Haliaeetus pelagicus), the white-tailed eagle (Haliaeetus albicilla) and the wedge-tailed eagle (Aquila audax), in roughly that order. For a species that is fairly homogeneous in its genetic make-up, the body mass of martial eagles is surprisingly variable. To some extent, the variation of body masses in the species is contributable to considerable reverse sexual dimorphism as well as varying environmental conditions of various eagle populations. Unsexed martial eagles from various studies have been found to have weighed an average of 3.93 kg (8.7 lb) in 17 birds, 3.97 kg (8.8 lb) in 20 birds and 4.23 kg (9.3 lb) in 20 birds while the average weight of martial eagles shot by game wardens in the early 20th century in South Africa was listed as 4.71 kg (10.4 lb). In weight range, the martial eagle broadly overlaps in size with the golden eagle (Aquila chrysaetos) and Verreaux's eagle (and is even exceeded by them in maximum known body mass). Based on numerous studies, the martial eagle appear to average mildly heavier than the Verreaux's eagle but (derived from the globally combined body mass of its various races), the mean body masses of golden and martial eagles are identical at approximately 4.17 kg (9.2 lb). The renders the golden and martial eagles as tied as the largest African eagles (by body mass but not in total length or wingspan, in which the martial bests the golden), as well as the heaviest two species of booted eagle in the world and as tied as the sixth heaviest eagles in the world, after the three largest species of sea eagle (Steller's being the heaviest extant, the others ranking 4th and 5th), the harpy eagle (Harpia harpyja) and the Philippine eagle (Pithecophaga jefferyi). The longest African eagle (and second longest booted eagle after the wedge-tailed eagle (Aquila audax)) is the crowned eagle by virtue of its relatively longer tail, as its body weight is slightly less than these three heaviest booted eagle species.

 

The martial eagle is to some degree adaptable to varied habitats but shows an overall preference for open woods and woodland edges, wooded savannah and thornbush habitats. The martial eagle has been recorded at elevations of up to 3,000 m (9,800 ft) but is not a true mountain dwelling species and resident eagles do not usually exceed an elevation of 1,500 m (4,900 ft). These eagles also avoid closed-canopy forests and hyper-arid desert. As such it is mostly absent from Guinean and Congolian forests, despite the species’ requirement for large trees for nesting purposes. It is shown that martial eagles can inhabit forests locally in areas where openings occur. For example, in a bird atlas for the country of Kenya, perhaps surprisingly, 88% of martial eagles were found to reside in well-wooded areas and they occurred in areas where annual rainfall exceeded 250 mm (9.8 in). In southern Africa, they have adapted to seemingly more open habitats than elsewhere in their range, such as semi-desert and open savanna with scattered trees, wooded hillocks and, as a recent adaptation, around pylons. In the desert areas of Namibia, they utilize ephemeral rivers that flow occasionally and allow large trees to grow. They usually seem to prefer desolate or protected areas. In the Karoo of South Africa, they consistently avoid areas with moderate to heavy cultivation or with heavier or more consistent winter rainfall. One study on the occurrence of diurnal raptors in protected areas versus outside unprotected areas, found that martial eagle detection was nearly twice as frequent in protected areas during the dry season and more than three times as frequent during the wet season. Some assorted diurnal raptors were even relatively rarer outside of protected areas such as hooded vultures (Necrosyrtes monachus).

 

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Uploaded on March 12, 2017
Taken on September 23, 2007